497 lines
60 KiB
XML
497 lines
60 KiB
XML
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<mods:title id="DF30377FB663080E53D7290A14A34B3A">Tityopsis rolandoi sp. n. (Scorpiones: Buthidae) from Cuba</mods:title>
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<taxonomicName id="4C40E5A48B45966EFC3DFB4BFB21CEF1" ID-CoL="63WHJ" authority="Armas, 1974" authorityName="Armas" authorityYear="1974" box="[1013,1278,1246,1273]" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="B93442358B45966EFC3DFB4BFB80CEF0" bold="true" box="[1013,1119,1246,1272]" italics="true" pageId="2" pageNumber="1">Tityopsis</emphasis>
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<bibRefCitation id="EFD1E3D68B45966EFBAEFB4AFB21CEF1" author="ARMAS" box="[1126,1278,1247,1273]" pageId="2" pageNumber="1" pagination="1 - 15" refId="ref4497" refString="ARMAS, L. F. DE. 1974. Escorpiones del archipielago cubano. 3. Genero Tityus C. L. Koch, 1836 (Scorpionida: Buthidae). Poeyana, 135: 1 - 15." type="journal article" year="1974">Armas, 1974</bibRefCitation>
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<paragraph id="8BFF9E278B45966EFC34FA95FB29CF10" blockId="2.[979,1311,1211,1304]" box="[1020,1270,1280,1304]" pageId="2" pageNumber="1">
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(
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<figureCitation id="137B82A28B45966EFBCCFA95FB4BCF10" box="[1028,1172,1280,1304]" captionStart-0="Figure 1" captionStart-1="Figures 2–5" captionStart-10="Figures 57–58" captionStart-2="Figures 6–13" captionStart-3="Figures 14–15" captionStart-4="Figures 16–17" captionStart-5="Figures 18–25" captionStart-6="Figures 26–47" captionStart-7="Figures 48–50" captionStart-8="Figures 51–54" captionStart-9="Figures 55–56" captionStartId-0="3.[114,182,2016,2038]" captionStartId-1="5.[113,190,2013,2035]" captionStartId-10="15.[114,191,2021,2043]" captionStartId-2="7.[113,190,2002,2024]" captionStartId-3="8.[115,192,1980,2002]" captionStartId-4="9.[116,193,1998,2020]" captionStartId-5="10.[114,191,2011,2033]" captionStartId-6="11.[113,190,1999,2021]" captionStartId-7="12.[115,192,2034,2056]" captionStartId-8="13.[115,192,1245,1267]" captionStartId-9="14.[111,188,2004,2026]" captionTargetBox-0="[111,1472,234,1995]" captionTargetBox-1="[162,1360,222,1996]" captionTargetBox-10="[115,1476,227,2004]" captionTargetBox-2="[115,1461,219,1980]" captionTargetBox-3="[116,1477,226,1971]" captionTargetBox-4="[113,1474,223,1989]" captionTargetBox-5="[119,1474,223,2002]" captionTargetBox-6="[141,1458,226,1984]" captionTargetBox-7="[113,1474,223,2019]" captionTargetBox-8="[113,1474,237,1233]" captionTargetBox-9="[113,1474,246,1981]" captionTargetId-0="figure-25@3.[111,1472,159,1998]" captionTargetId-1="figure-52@5.[113,1474,198,2011]" captionTargetId-10="figure-31@15.[115,1476,202,2016]" captionTargetId-2="figure-72@7.[113,1474,204,2008]" captionTargetId-3="figure-50@8.[116,1477,210,1986]" captionTargetId-4="figure-45@9.[113,1474,193,2004]" captionTargetId-5="figure-61@10.[113,1474,223,2014]" captionTargetId-6="figure-119@11.[113,1474,214,2009]" captionTargetId-7="figure-36@12.[113,1474,223,2019]" captionTargetId-8="figure-460@13.[113,1474,234,1233]" captionTargetId-9="figure-37@14.[113,1474,246,1985]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-10="15" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionTargetPageId-8="13" captionTargetPageId-9="14" captionText-0="Figure 1. Paratypes male (left) and female of Tityopsis rolandoi sp. n. in vivo habitus." captionText-1="Figures 2–5: Tityopsis rolandoi sp. n. Figures 2–3. Male holotype in dorsal (2) and ventral (3) views. Figures 4–5. Female paratype in dorsal (4) and ventral (5) views. Scale bar: 10 mm." captionText-10="Figures 57–58. Tityopsis rolandoi sp. n., female paratype with newborns (57) and juveniles after the first ecdysis (58)." captionText-2="Figures 6–13: Tityopsis rolandoi sp. n. Figures 6, 11–13. Male holotype, telson in lateral view (6), metasoma and telson lateral (11), dorsal (12), and ventral (13) views. Figures 7–10. Female paratype, telson in lateral view (7), metasoma and telson lateral (8), dorsal (9), and ventral (10) views. Scale bar: 10 mm (8–10, 11–13)." captionText-3="Figures 14–15: Tityopsis rolandoi sp. n., male holotype. Figure 14. Chelicera, carapace, tergites and part of right legs under UV fluorescence. Figure 15. Coxosternal area and sternites under UV fluorescence." captionText-4="Figures 16–17: Tityopsis rolandoi sp. n., female paratype. Figure 16. Chelicera, carapace, tergites and part of right legs under UV fluorescence. Figure 17. Coxosternal area and sternites under UV fluorescence." captionText-5="Figures 18–25: Tityopsis rolandoi sp. n. Figures 18–19, 22–25. Male holotype, chelicera, carapace, tergites I–III (18), coxosternal area and sternites (19), left legs I–IV in retrolateral aspect (respectively) (22–25). Figures 20–21. Female paratype, chelicera, carapace, tergites I–III (20), coxosternal area and sternites (21)." captionText-6="Figures 26–47: Tityopsis rolandoi sp. n., pedipalp segments under white light. Figures 26–36. Male holotype, chela dorsal (26), external (27), ventral (28) views, patella dorsal (29), external (30), and ventral (31) views, femur and trochanter external (32), dorsal (33), and ventral (34) views, movable (35) and fixed (36) finger dentition. Figures 37–47. Female paratype, chela dorsal (37), external (38), ventral (39) views, patella dorsal (40), external (41), and ventral (42) views, femur and trochanter external (43), dorsal (44), and ventral (45) views, movable (46) and fixed (47) finger dentition. Trichobothrial pattern is indicated by white circles in Figures 38–41, 43–44." captionText-7="Figures 48–50. Tityopsis rolandoi sp. n., male holotype, pedipalp under UV fluorescence, dorsal (48), external (49), and ventral (50)." captionText-8="Figures 51–54: The chromosomes of Tityopsis sheylae (2n=22) (51, 52) and T. rolandoi sp. n. (2n=20) (53, 54). Postpachytene (51, 53) and ideograms (52, 54) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values). Scale bar: 5 μm." captionText-9="Figures 55–56: Figure 55. Type localities of Tityopsis species. Figure 56. Type locality of Tityopsis rolandoi sp. n." figureDoi-0="http://doi.org/10.5281/zenodo.14660691" figureDoi-1="http://doi.org/10.5281/zenodo.14660693" figureDoi-10="http://doi.org/10.5281/zenodo.14660711" figureDoi-2="http://doi.org/10.5281/zenodo.14660695" figureDoi-3="http://doi.org/10.5281/zenodo.14660697" figureDoi-4="http://doi.org/10.5281/zenodo.14660699" figureDoi-5="http://doi.org/10.5281/zenodo.14660701" figureDoi-6="http://doi.org/10.5281/zenodo.14660703" figureDoi-7="http://doi.org/10.5281/zenodo.14660705" figureDoi-8="http://doi.org/10.5281/zenodo.14660707" figureDoi-9="http://doi.org/10.5281/zenodo.14660709" httpUri-0="https://zenodo.org/record/14660691/files/figure.png" httpUri-1="https://zenodo.org/record/14660693/files/figure.png" httpUri-10="https://zenodo.org/record/14660711/files/figure.png" httpUri-2="https://zenodo.org/record/14660695/files/figure.png" httpUri-3="https://zenodo.org/record/14660697/files/figure.png" httpUri-4="https://zenodo.org/record/14660699/files/figure.png" httpUri-5="https://zenodo.org/record/14660701/files/figure.png" httpUri-6="https://zenodo.org/record/14660703/files/figure.png" httpUri-7="https://zenodo.org/record/14660705/files/figure.png" httpUri-8="https://zenodo.org/record/14660707/files/figure.png" httpUri-9="https://zenodo.org/record/14660709/files/figure.png" pageId="2" pageNumber="1">Figures 1–58</figureCitation>
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,
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<tableCitation id="C6C2AB9C8B45966EFB56FA95FB33CF10" box="[1182,1260,1280,1304]" captionStart="Table 1" captionStartId="4.[115,171,1000,1021]" captionText="Table 1. Comparative measurements of holotype male of Tityopsis rolandoi sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="2" pageNumber="1">Table 1</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35ACDAC8B45966EFCF8FAD4FC02CC10" pageId="2" pageNumber="1" type="reference_group">
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<paragraph id="8BFF9E278B45966EFCF8FAD4FB8ECF70" blockId="2.[816,1474,1344,1560]" pageId="2" pageNumber="1">
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<treatmentCitationGroup id="AB50B9098B45966EFCF8FAD4FB8ECF70" pageId="2" pageNumber="1">
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<treatmentCitation id="0AE1B8368B45966EFCF8FAD4FB6ACF50" author="ARMAS" box="[816,1205,1344,1368]" page="8" pageId="2" pageNumber="1" year="1974">
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<taxonomicName id="4C40E5A48B45966EFCF8FAD4FB6ACF50" authority="Armas, 1974: 8" authorityName="Armas" authorityPageNumber="8" authorityYear="1974" box="[816,1205,1344,1368]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Tityopsis">
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<emphasis id="B93442358B45966EFCF8FAD4FCAFCF50" box="[816,880,1345,1368]" italics="true" pageId="2" pageNumber="1">Tityus</emphasis>
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(
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<emphasis id="B93442358B45966EFC40FAD4FC38CF50" box="[904,999,1345,1368]" italics="true" pageId="2" pageNumber="1">Tityopsis</emphasis>
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)
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<bibRefCitation id="EFD1E3D68B45966EFC35FAD5FB6ACF50" author="ARMAS" box="[1021,1205,1344,1368]" pageId="2" pageNumber="1" pagination="1 - 15" refId="ref4497" refString="ARMAS, L. F. DE. 1974. Escorpiones del archipielago cubano. 3. Genero Tityus C. L. Koch, 1836 (Scorpionida: Buthidae). Poeyana, 135: 1 - 15." type="journal article" year="1974">Armas, 1974: 8</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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<treatmentCitation id="0AE1B8368B45966EFB00FAD5FA53CF50" author="ARMAS" box="[1224,1420,1344,1368]" page="11" pageId="2" pageNumber="1" year="1982">
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<bibRefCitation id="EFD1E3D68B45966EFB00FAD5FA53CF50" author="ARMAS" box="[1224,1420,1344,1368]" pageId="2" pageNumber="1" pagination="1 - 17" refId="ref4537" refString="ARMAS, L. F. DE. 1982. Algunos aspectos zoogeogrAficos de la escorpiofauna antillana. Poeyana, 238: 1 - 17." type="journal article" year="1982">Armas, 1982: 11</bibRefCitation>
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</treatmentCitation>
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, 13;
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<treatmentCitation id="0AE1B8368B45966EFCA8FAF5FBFBCF70" author="FRANCKE" box="[864,1060,1376,1400]" page="16" pageId="2" pageNumber="1" year="1985">
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<bibRefCitation id="EFD1E3D68B45966EFCA8FAF5FBFBCF70" author="FRANCKE" box="[864,1060,1376,1400]" pageId="2" pageNumber="1" pagination="1 - 32" refId="ref4705" refString="FRANCKE, O. F. 1985. Conspectus genericus scorpionorum 1758 - 1982 (Arachnida: Scorpiones). Occasional Papers of the Museum, Texas Tech University, 98: 1 - 32." type="journal article" year="1985">Francke, 1985: 16</bibRefCitation>
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</treatmentCitation>
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, 19.
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</paragraph>
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<paragraph id="8BFF9E278B45966EFCF8FA14FC02CC10" blockId="2.[816,1474,1344,1560]" pageId="2" pageNumber="1">
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<taxonomicName id="4C40E5A48B45966EFCF8FA14FC50CF90" ID-CoL="63WHJ" authorityName="Armas" authorityYear="1974" box="[816,911,1409,1432]" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis id="B93442358B45966EFCF8FA14FC50CF90" box="[816,911,1409,1432]" italics="true" pageId="2" pageNumber="1">Tityopsis</emphasis>
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</taxonomicName>
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||
:
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<bibRefCitation id="EFD1E3D68B45966EFC68FA15FB81CF90" author="ARMAS" box="[928,1118,1408,1432]" pageId="2" pageNumber="1" pagination="1 - 37" refId="ref4563" refString="ARMAS, L. F. DE. 1984. Escorpiones del archipielago cubano. VIII. Adiciones y enmiendas (Scorpiones, Buthidae, Diplocentridae). Poeyana, 275: 1 - 37." type="journal article" year="1984">Armas, 1984: 29</bibRefCitation>
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; tab. 3; Fet & Lowe, 2000: 58, 227–228; Teruel & Kovařík, 2012: 143–151, figs. 7, 30, 42–43, 49, 306–322, 570–577; Teruel & Rodríguez-Cabrera, 2022: 1–40, figs. 1–4 (complete references list until 2021).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35ACDAC8B45966EFCF8F9D4FC31CC90" pageId="2" pageNumber="1" type="materials_examined">
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<paragraph id="8BFF9E278B45966EFCF8F9D4FC31CC90" blockId="2.[816,1474,1600,1688]" pageId="2" pageNumber="1">
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<collectionCode id="ED5106E28B45966EFCF8F9D4FC1BCC50" box="[816,964,1600,1624]" pageId="2" pageNumber="1">
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<typeStatus id="54FB20858B45966EFCF8F9D4FCB9CC50" box="[816,870,1601,1624]" pageId="2" pageNumber="1">TYPE</typeStatus>
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SPECIES
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</collectionCode>
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.
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<taxonomicName id="4C40E5A48B45966EFC1EF9D4FA92CC50" authority="Moreno, 1940" authorityName="Moreno" authorityYear="1940" box="[982,1357,1600,1624]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="inexpectatus">
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<emphasis id="B93442358B45966EFC1EF9D4FB79CC50" box="[982,1190,1601,1624]" italics="true" pageId="2" pageNumber="1">Tityus inexpectatus</emphasis>
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Moreno, 1940
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</taxonomicName>
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[currently
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<taxonomicName id="4C40E5A48B45966EFCF8F9F4FB1FCC70" authority="(Moreno, 1940)" baseAuthorityName="Moreno" baseAuthorityYear="1940" box="[816,1216,1632,1656]" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="inexpectata">
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<emphasis id="B93442358B45966EFCF8F9F4FBD0CC70" box="[816,1039,1633,1656]" italics="true" pageId="2" pageNumber="1">Tityopsis inexpectata</emphasis>
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||
(Moreno, 1940)
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||
</taxonomicName>
|
||
], by original designation (
|
||
<bibRefCitation id="EFD1E3D68B45966EFCF0F915FC3FCC90" author="ARMAS" box="[824,992,1664,1688]" pageId="2" pageNumber="1" pagination="1 - 15" refId="ref4497" refString="ARMAS, L. F. DE. 1974. Escorpiones del archipielago cubano. 3. Genero Tityus C. L. Koch, 1836 (Scorpionida: Buthidae). Poeyana, 135: 1 - 15." type="journal article" year="1974">Armas, 1974: 8</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35ACDAC8B45966AFCF8F955FDEEC82B" lastPageId="6" lastPageNumber="5" pageId="2" pageNumber="1" type="diagnosis">
|
||
<paragraph id="8BFF9E278B459668FCF8F955FB91CC91" blockId="2.[816,1474,1728,2041]" lastBlockId="4.[818,1477,1089,1689]" lastPageId="4" lastPageNumber="3" pageId="2" pageNumber="1">
|
||
DIAGNOSIS (after Teruel & Rodríguez-Cabrera, 2022). Adult size small for the family (
|
||
<quantity id="4CB833C28B45966EFB90F975FB0FCCF0" box="[1112,1232,1760,1784]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="2" pageNumber="1" unit="mm" value="22.5" valueMax="30.0" valueMin="15.0">15–30 mm</quantity>
|
||
in males,
|
||
<quantity id="4CB833C28B45966EFA83F975FA1ECCF0" box="[1355,1473,1760,1784]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="2" pageNumber="1" unit="mm" value="30.0" valueMax="40.0" valueMin="20.0">20–40 mm</quantity>
|
||
in females), with males slightly smaller than females within the same size-class. Coloration light yellowish to blackish, immaculate to densely patterned all over, especially on carapace, legs and metasoma. Entire body and appendages covered with modified macrosetae: brightly UV-fluorescent, short to medium-sized, thick, translucent, with tip truncate to crown-shaped. Cheliceral fixed finger with one ventral denticle, movable finger with two. Pedipalp very similar in both sexes: not especially robust nor attenuate and strongly carinate; trichobothrial pattern A-α orthobothriotaxic, without clearly defined petite trichobothria (e. g., chela
|
||
<emphasis id="B93442358B439668FD28FB14FCDACE97" box="[736,773,1153,1183]" italics="true" pageId="4" pageNumber="3">
|
||
Eb
|
||
<subScript id="17C49C628B439668FD35FB04FCDACE97" attach="left" box="[765,773,1169,1183]" fontSize="6" pageId="4" pageNumber="3">3</subScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B93442358B439668FF6BFB34FF14CEB1" box="[163,203,1185,1209]" italics="true" pageId="4" pageNumber="3">Esb</emphasis>
|
||
and fixed finger
|
||
<emphasis id="B93442358B439668FE44FB34FE6FCEB1" box="[396,432,1185,1209]" italics="true" pageId="4" pageNumber="3">esb</emphasis>
|
||
are somewhat smaller, but not as disproportionately reduced as a typical petite); femur with
|
||
<emphasis id="B93442358B439668FF65FB74FEC3CEF7" box="[173,284,1249,1279]" italics="true" pageId="4" pageNumber="3">
|
||
d
|
||
<subScript id="17C49C628B439668FF72FB64FF1DCEF7" attach="left" box="[186,194,1265,1279]" fontSize="6" pageId="4" pageNumber="3">1</subScript>
|
||
-d
|
||
<subScript id="17C49C628B439668FF10FB64FF3FCEF7" attach="left" box="[216,224,1265,1279]" fontSize="6" pageId="4" pageNumber="3">3</subScript>
|
||
-d
|
||
<subScript id="17C49C628B439668FF3EFB64FF21CEF7" attach="left" box="[246,254,1265,1279]" fontSize="6" pageId="4" pageNumber="3">4</subScript>
|
||
-d
|
||
<subScript id="17C49C628B439668FEDCFB64FEC3CEF7" attach="left" box="[276,284,1265,1279]" fontSize="6" pageId="4" pageNumber="3">5</subScript>
|
||
</emphasis>
|
||
regularly spaced along basal half,
|
||
<emphasis id="B93442358B439668FD6BFB77FD68CEF7" box="[675,695,1250,1279]" italics="true" pageId="4" pageNumber="3">
|
||
e
|
||
<subScript id="17C49C628B439668FD67FB64FD68CEF7" attach="left" box="[687,695,1265,1279]" fontSize="6" pageId="4" pageNumber="3">1</subScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B93442358B439668FD39FB77FCDACEF7" box="[753,773,1250,1279]" italics="true" pageId="4" pageNumber="3">
|
||
e
|
||
<subScript id="17C49C628B439668FD35FB64FCDACEF7" attach="left" box="[765,773,1265,1279]" fontSize="6" pageId="4" pageNumber="3">2</subScript>
|
||
</emphasis>
|
||
located on subbasal and median position, respectively; patella with
|
||
<emphasis id="B93442358B439668FF64FAB4FF1ECF37" box="[172,193,1313,1343]" italics="true" pageId="4" pageNumber="3">
|
||
d
|
||
<subScript id="17C49C628B439668FF71FAA4FF1ECF37" attach="left" box="[185,193,1329,1343]" fontSize="6" pageId="4" pageNumber="3">3</subScript>
|
||
</emphasis>
|
||
external to dorsal median carina and
|
||
<emphasis id="B93442358B439668FDAAFAB7FDB6CF31" box="[610,617,1314,1337]" italics="true" pageId="4" pageNumber="3">i</emphasis>
|
||
slightly distal to midpoint; manus with
|
||
<emphasis id="B93442358B439668FE45FAD7FE79CF57" box="[397,422,1346,1375]" italics="true" pageId="4" pageNumber="3">
|
||
V
|
||
<subScript id="17C49C628B439668FE56FAC4FE79CF57" attach="left" box="[414,422,1361,1375]" fontSize="6" pageId="4" pageNumber="3">2</subScript>
|
||
</emphasis>
|
||
clearly closer to ventroexternal carina than
|
||
<emphasis id="B93442358B439668FF24FAF7FEDBCF77" box="[236,260,1378,1407]" italics="true" pageId="4" pageNumber="3">
|
||
V
|
||
<subScript id="17C49C628B439668FF34FAE4FEDBCF77" attach="left" box="[252,260,1393,1407]" fontSize="6" pageId="4" pageNumber="3">1</subScript>
|
||
</emphasis>
|
||
; fixed finger with
|
||
<emphasis id="B93442358B439668FE0AFAF4FD8ACF71" box="[450,597,1377,1401]" italics="true" pageId="4" pageNumber="3">est-db-et-dt-it</emphasis>
|
||
located on distal half, with
|
||
<emphasis id="B93442358B439668FF17FA17FF31CF91" box="[223,238,1410,1433]" italics="true" pageId="4" pageNumber="3">it</emphasis>
|
||
midway between
|
||
<emphasis id="B93442358B439668FE67FA14FE1BCF91" box="[431,452,1409,1433]" italics="true" pageId="4" pageNumber="3">dt</emphasis>
|
||
and fingertip. Pedipalp chelae robust, wider than patella and strongly carinate; fixed and movable fingers with 11–13 principal rows of denticles which are short, oblique, subequal and flanked by very large internal and external accessory denticles, apical subrow oblique and composed of four denticles, basal lobe/notch combination absent. Carapace trapezoidal, essentially flat and with carinae distinct but variably fused and poorly defined from intercarinal granulation, which is variable but always well developed; 3–6 pairs of lateral eyes, not concealed below anterolateral margin. Legs without tibial spurs, both pedal spurs present; telotarsi ventrally with two well-defined, parallel, longitudinal rows of thin spiniform setae. Male genital papillae lip-like, not protruding from genital operculum and each with a distinct fleshy point, hardly visible by being located extremely basal and almost entirely concealed beneath pre-pectinal plate. Pre-pectinal plate well developed, heavily sclerotized and medially invaginate, not well visible by being largely concealed beneath genital operculum. Pectines with 11–16 teeth in males,
|
||
<quantity id="4CB833C28B439668FEDDF874FEACCDF1" box="[277,371,2017,2041]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.81" metricValueMin="2.54" pageId="4" pageNumber="3" unit="in" value="12.5" valueMax="15.0" valueMin="10.0">10–15 in</quantity>
|
||
females, fulcra well developed; basal middle lamella modified, slightly to moderately enlarged and angulose in males, moderately to remarkably enlarged and oval to round in females; basal plate unmodified, but in females with a large, whitish discal area of presumably glandular function. Tergites heavily granulose, monocarinate, with median carina and coarse granulation projecting over posterior margin. Sternites with small, round to short slit-like spiracles; III without stridulatory organ or deep furrows but with a raised, granulose, median triangular area flanked by two lateral depressions that fit pectines for protection or rest, V–VII with paired submedian and lateral keels, V with three smooth patches (one median and two laterals, all conspicuously larger, bulkier and also usually paler in males). Metasoma very similar in both sexes: short, robust and very strongly carinate, segment V with lateral median carinae irregularly defined but clearly present. Telson vesicle globose; subaculear tubercle vestigial to moderate, blunt, irregular to conical and widely separate from aculeus, which is long, very sharp and strongly curved.
|
||
</paragraph>
|
||
<caption id="DF3FCEAF8B44966FFFBAF875FC6FCDFE" ID-DOI="http://doi.org/10.5281/zenodo.14660691" ID-Zenodo-Dep="14660691" box="[114,944,2016,2038]" httpUri="https://zenodo.org/record/14660691/files/figure.png" pageId="3" pageNumber="2" startId="3.[114,182,2016,2038]" targetBox="[111,1472,234,1995]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BFF9E278B44966FFFBAF875FC6FCDFE" blockId="3.[114,944,2016,2038]" box="[114,944,2016,2038]" pageId="3" pageNumber="2">
|
||
<emphasis id="B93442358B44966FFFBAF875FF17CDFD" bold="true" box="[114,200,2016,2038]" pageId="3" pageNumber="2">Figure 1</emphasis>
|
||
. Paratypes male (left) and female of
|
||
<taxonomicName id="4C40E5A48B44966FFDE3F875FD06CDFD" authorityName="Kovařík & Stockmann & Šťáhlavský & Yong" authorityYear="2024" box="[555,729,2016,2037]" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="rolandoi" status="sp. nov.">
|
||
<emphasis id="B93442358B44966FFDE3F875FD06CDFD" box="[555,729,2016,2037]" italics="true" pageId="3" pageNumber="2">Tityopsis rolandoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207FF4E8B44966FFD17F875FCCACDFD" box="[735,789,2016,2037]" pageId="3" pageNumber="2" rank="species">
|
||
<emphasis id="B93442358B44966FFD17F875FD29CDFD" bold="true" box="[735,758,2016,2037]" pageId="3" pageNumber="2">sp</emphasis>
|
||
.
|
||
<emphasis id="B93442358B44966FFCC9F875FCD1CDFD" bold="true" box="[769,782,2016,2037]" pageId="3" pageNumber="2">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
in vivo habitus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFF9E278B439668FD35FF62FAFCC9C0" pageId="4" pageNumber="3">
|
||
<table id="F9406C878B436993FFBCFF62FA52C9C3" box="[116,1421,247,971]" gridcols="4" gridrows="19" pageId="4" pageNumber="3">
|
||
<tr id="35709C658B436993FFBCFF62FA52CB18" box="[116,1421,247,272]" gridrow="0" pageId="4" pageNumber="3" rowspan-0="1" rowspan-1="1">
|
||
<th id="76A1F5198B436993FD35FF62FBD4CB18" box="[765,1035,247,272]" gridcol="2" gridrow="0" pageId="4" pageNumber="3">
|
||
<taxonomicName id="4C40E5A48B439668FD35FF62FC1DCB07" authorityName="Kovařík & Stockmann & Šťáhlavský & Yong" authorityYear="2024" box="[765,962,247,271]" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="rolandoi" status="sp. nov.">
|
||
<emphasis id="B93442358B439668FD35FF62FC1DCB07" bold="true" box="[765,962,247,271]" italics="true" pageId="4" pageNumber="3">Tityopsis rolandoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207FF4E8B439668FC06FF6DFBD4CB07" box="[974,1035,248,272]" pageId="4" pageNumber="3" rank="species">
|
||
<emphasis id="B93442358B439668FC06FF6DFC38CB18" bold="true" box="[974,999,248,272]" pageId="4" pageNumber="3">sp</emphasis>
|
||
.
|
||
<emphasis id="B93442358B439668FC3DFF6DFBDBCB18" bold="true" box="[1013,1028,248,272]" pageId="4" pageNumber="3">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</th>
|
||
<th id="76A1F5198B436993FB48FF62FA52CB18" box="[1152,1421,247,272]" gridcol="3" gridrow="0" pageId="4" pageNumber="3">
|
||
<taxonomicName id="4C40E5A48B439668FB48FF62FA9ACB07" authorityName="Kovařík & Stockmann & Šťáhlavský & Yong" authorityYear="2024" box="[1152,1349,247,271]" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="rolandoi" status="sp. nov.">
|
||
<emphasis id="B93442358B439668FB48FF62FA9ACB07" bold="true" box="[1152,1349,247,271]" italics="true" pageId="4" pageNumber="3">Tityopsis rolandoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207FF4E8B439668FA99FF6DFA52CB07" box="[1361,1421,248,272]" pageId="4" pageNumber="3" rank="species">
|
||
<emphasis id="B93442358B439668FA99FF6DFAB5CB18" bold="true" box="[1361,1386,248,272]" pageId="4" pageNumber="3">sp</emphasis>
|
||
.
|
||
<emphasis id="B93442358B439668FAB0FF6DFA58CB18" bold="true" box="[1400,1415,248,272]" pageId="4" pageNumber="3">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</th>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFEB5FA52CB31" box="[116,1421,288,313]" gridrow="1" pageId="4" pageNumber="3" rowspan-1="1">
|
||
<th id="76A1F5198B436993FFBCFEB5FE9ECB31" box="[116,321,288,313]" gridcol="0" gridrow="1" pageId="4" pageNumber="3">
|
||
<emphasis id="B93442358B439668FFBCFEB5FEE5CB30" bold="true" box="[116,314,288,312]" pageId="4" pageNumber="3">Dimensions (mm)</emphasis>
|
||
</th>
|
||
<td id="76A1F5198B436993FD35FEB5FBD4CB31" box="[765,1035,288,313]" gridcol="2" gridrow="1" pageId="4" pageNumber="3">♂ holotype</td>
|
||
<td id="76A1F5198B436993FB48FEB5FA52CB31" box="[1152,1421,288,313]" gridcol="3" gridrow="1" pageId="4" pageNumber="3">♀ paratype</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFED0FA52CB57" box="[116,1421,325,351]" gridrow="2" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFED0FE9ECB57" box="[116,321,325,351]" gridcol="0" gridrow="2" pageId="4" pageNumber="3">Carapace</th>
|
||
<td id="76A1F5198B436993FDE0FED0FD51CB57" box="[552,654,325,351]" gridcol="1" gridrow="2" pageId="4" pageNumber="3">L / W</td>
|
||
<td id="76A1F5198B436993FD35FED0FBD4CB57" box="[765,1035,325,351]" gridcol="2" gridrow="2" pageId="4" pageNumber="3">2.62 / 2.65</td>
|
||
<td id="76A1F5198B436993FB48FED0FA52CB57" box="[1152,1421,325,351]" gridcol="3" gridrow="2" pageId="4" pageNumber="3">3.13 / 3.45</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFEFEFA52CB8D" box="[116,1421,363,389]" gridrow="3" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFEFEFE9ECB8D" box="[116,321,363,389]" gridcol="0" gridrow="3" pageId="4" pageNumber="3">Mesosoma</th>
|
||
<td id="76A1F5198B436993FDE0FEFEFD51CB8D" box="[552,654,363,389]" gridcol="1" gridrow="3" pageId="4" pageNumber="3">L</td>
|
||
<td id="76A1F5198B436993FD35FEFEFBD4CB8D" box="[765,1035,363,389]" gridcol="2" gridrow="3" pageId="4" pageNumber="3">7.25</td>
|
||
<td id="76A1F5198B436993FB48FEFEFA52CB8D" box="[1152,1421,363,389]" gridcol="3" gridrow="3" pageId="4" pageNumber="3">9.75</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFE07FA52CBA4" box="[116,1421,402,428]" gridrow="4" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFE07FE9ECBA4" box="[116,321,402,428]" gridcol="0" gridrow="4" pageId="4" pageNumber="3">Tergite VII</th>
|
||
<td id="76A1F5198B436993FDE0FE07FD51CBA4" box="[552,654,402,428]" gridcol="1" gridrow="4" pageId="4" pageNumber="3">L / W</td>
|
||
<td id="76A1F5198B436993FD35FE07FBD4CBA4" box="[765,1035,402,428]" gridcol="2" gridrow="4" pageId="4" pageNumber="3">1.83 / 2.62</td>
|
||
<td id="76A1F5198B436993FB48FE07FA52CBA4" box="[1152,1421,402,428]" gridcol="3" gridrow="4" pageId="4" pageNumber="3">2.27 / 3.62</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFE2CFA52CBDB" box="[116,1421,441,467]" gridrow="5" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFE2CFE9ECBDB" box="[116,321,441,467]" gridcol="0" gridrow="5" pageId="4" pageNumber="3">Metasoma + telson</th>
|
||
<td id="76A1F5198B436993FDE0FE2CFD51CBDB" box="[552,654,441,467]" gridcol="1" gridrow="5" pageId="4" pageNumber="3">L</td>
|
||
<td id="76A1F5198B436993FD35FE2CFBD4CBDB" box="[765,1035,441,467]" gridcol="2" gridrow="5" pageId="4" pageNumber="3">13.45</td>
|
||
<td id="76A1F5198B436993FB48FE2CFA52CBDB" box="[1152,1421,441,467]" gridcol="3" gridrow="5" pageId="4" pageNumber="3">15.67</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFE75FA52CBF2" box="[116,1421,480,506]" gridrow="6" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFE75FE9ECBF2" box="[116,321,480,506]" gridcol="0" gridrow="6" pageId="4" pageNumber="3">Segment I</th>
|
||
<td id="76A1F5198B436993FDE0FE75FD51CBF2" box="[552,654,480,506]" gridcol="1" gridrow="6" pageId="4" pageNumber="3">L / W / D</td>
|
||
<td id="76A1F5198B436993FD35FE75FBD4CBF2" box="[765,1035,480,506]" gridcol="2" gridrow="6" pageId="4" pageNumber="3">1.64 / 1.47 / 1.33</td>
|
||
<td id="76A1F5198B436993FB48FE75FA52CBF2" box="[1152,1421,480,506]" gridcol="3" gridrow="6" pageId="4" pageNumber="3">1.96 / 1.71 / 1.63</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFD93FA52C829" box="[116,1421,518,545]" gridrow="7" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFD93FE9EC829" box="[116,321,518,545]" gridcol="0" gridrow="7" pageId="4" pageNumber="3">Segment II</th>
|
||
<td id="76A1F5198B436993FDE0FD93FD51C829" box="[552,654,518,545]" gridcol="1" gridrow="7" pageId="4" pageNumber="3">L / W / D</td>
|
||
<td id="76A1F5198B436993FD35FD93FBD4C829" box="[765,1035,518,545]" gridcol="2" gridrow="7" pageId="4" pageNumber="3">1.98 / 1.37 / 1.23</td>
|
||
<td id="76A1F5198B436993FB48FD93FA52C829" box="[1152,1421,518,545]" gridcol="3" gridrow="7" pageId="4" pageNumber="3">2.36 / 1.64 / 1.56</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFDB8FA52C84F" box="[116,1421,557,583]" gridrow="8" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFDB8FE9EC84F" box="[116,321,557,583]" gridcol="0" gridrow="8" pageId="4" pageNumber="3">Segment III</th>
|
||
<td id="76A1F5198B436993FDE0FDB8FD51C84F" box="[552,654,557,583]" gridcol="1" gridrow="8" pageId="4" pageNumber="3">L / W / D</td>
|
||
<td id="76A1F5198B436993FD35FDB8FBD4C84F" box="[765,1035,557,583]" gridcol="2" gridrow="8" pageId="4" pageNumber="3">2.21 / 1.36 / 1.22</td>
|
||
<td id="76A1F5198B436993FB48FDB8FA52C84F" box="[1152,1421,557,583]" gridcol="3" gridrow="8" pageId="4" pageNumber="3">2.46 / 1.59 / 1.53</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFDC1FA52C866" box="[116,1421,596,622]" gridrow="9" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFDC1FE9EC866" box="[116,321,596,622]" gridcol="0" gridrow="9" pageId="4" pageNumber="3">Segment IV</th>
|
||
<td id="76A1F5198B436993FDE0FDC1FD51C866" box="[552,654,596,622]" gridcol="1" gridrow="9" pageId="4" pageNumber="3">L / W / D</td>
|
||
<td id="76A1F5198B436993FD35FDC1FBD4C866" box="[765,1035,596,622]" gridcol="2" gridrow="9" pageId="4" pageNumber="3">2.33 / 1.28 / 1.19</td>
|
||
<td id="76A1F5198B436993FB48FDC1FA52C866" box="[1152,1421,596,622]" gridcol="3" gridrow="9" pageId="4" pageNumber="3">2.66 / 1.56 / 1.43</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFDEEFA52C89D" box="[116,1421,635,661]" gridrow="10" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFDEEFE9EC89D" box="[116,321,635,661]" gridcol="0" gridrow="10" pageId="4" pageNumber="3">Segment V</th>
|
||
<td id="76A1F5198B436993FDE0FDEEFD51C89D" box="[552,654,635,661]" gridcol="1" gridrow="10" pageId="4" pageNumber="3">L / W / D</td>
|
||
<td id="76A1F5198B436993FD35FDEEFBD4C89D" box="[765,1035,635,661]" gridcol="2" gridrow="10" pageId="4" pageNumber="3">2.70 / 1.29 / 1.16</td>
|
||
<td id="76A1F5198B436993FB48FDEEFA52C89D" box="[1152,1421,635,661]" gridcol="3" gridrow="10" pageId="4" pageNumber="3">3.24 / 1.58 / 1.40</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFD34FA52C8B4" box="[116,1421,673,700]" gridrow="11" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFD34FE9EC8B4" box="[116,321,673,700]" gridcol="0" gridrow="11" pageId="4" pageNumber="3">Telson</th>
|
||
<td id="76A1F5198B436993FDE0FD34FD51C8B4" box="[552,654,673,700]" gridcol="1" gridrow="11" pageId="4" pageNumber="3">L / W / D</td>
|
||
<td id="76A1F5198B436993FD35FD34FBD4C8B4" box="[765,1035,673,700]" gridcol="2" gridrow="11" pageId="4" pageNumber="3">2.59 / 1.13 / 1.00</td>
|
||
<td id="76A1F5198B436993FB48FD34FA52C8B4" box="[1152,1421,673,700]" gridcol="3" gridrow="11" pageId="4" pageNumber="3">2.99 / 1.27 / 1.19</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFD5DFA52C8EA" box="[116,1421,712,738]" gridrow="12" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFD5DFE9EC8EA" box="[116,321,712,738]" gridcol="0" gridrow="12" pageId="4" pageNumber="3">Pedipalp</th>
|
||
<td id="76A1F5198B436993FDE0FD5DFD51C8EA" box="[552,654,712,738]" gridcol="1" gridrow="12" pageId="4" pageNumber="3">L</td>
|
||
<td id="76A1F5198B436993FD35FD5DFBD4C8EA" box="[765,1035,712,738]" gridcol="2" gridrow="12" pageId="4" pageNumber="3">9.59</td>
|
||
<td id="76A1F5198B436993FB48FD5DFA52C8EA" box="[1152,1421,712,738]" gridcol="3" gridrow="12" pageId="4" pageNumber="3">11.-29</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFD7AFA52C901" box="[116,1421,751,777]" gridrow="13" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFD7AFE9EC901" box="[116,321,751,777]" gridcol="0" gridrow="13" pageId="4" pageNumber="3">Femur</th>
|
||
<td id="76A1F5198B436993FDE0FD7AFD51C901" box="[552,654,751,777]" gridcol="1" gridrow="13" pageId="4" pageNumber="3">L / W</td>
|
||
<td id="76A1F5198B436993FD35FD7AFBD4C901" box="[765,1035,751,777]" gridcol="2" gridrow="13" pageId="4" pageNumber="3">2.34 / 0.81</td>
|
||
<td id="76A1F5198B436993FB48FD7AFA52C901" box="[1152,1421,751,777]" gridcol="3" gridrow="13" pageId="4" pageNumber="3">2.73 / 1.06</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFC80FA52C938" box="[116,1421,789,816]" gridrow="14" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFC80FE9EC938" box="[116,321,789,816]" gridcol="0" gridrow="14" pageId="4" pageNumber="3">Patella</th>
|
||
<td id="76A1F5198B436993FDE0FC80FD51C938" box="[552,654,789,816]" gridcol="1" gridrow="14" pageId="4" pageNumber="3">L / W</td>
|
||
<td id="76A1F5198B436993FD35FC80FBD4C938" box="[765,1035,789,816]" gridcol="2" gridrow="14" pageId="4" pageNumber="3">2.82 / 1.05</td>
|
||
<td id="76A1F5198B436993FB48FC80FA52C938" box="[1152,1421,789,816]" gridcol="3" gridrow="14" pageId="4" pageNumber="3">3.30 / 1.31</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFCA9FA52C95E" box="[116,1421,828,854]" gridrow="15" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFCA9FE9EC95E" box="[116,321,828,854]" gridcol="0" gridrow="15" pageId="4" pageNumber="3">Chela</th>
|
||
<td id="76A1F5198B436993FDE0FCA9FD51C95E" box="[552,654,828,854]" gridcol="1" gridrow="15" pageId="4" pageNumber="3">L</td>
|
||
<td id="76A1F5198B436993FD35FCA9FBD4C95E" box="[765,1035,828,854]" gridcol="2" gridrow="15" pageId="4" pageNumber="3">4.43</td>
|
||
<td id="76A1F5198B436993FB48FCA9FA52C95E" box="[1152,1421,828,854]" gridcol="3" gridrow="15" pageId="4" pageNumber="3">5.26</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFCF6FA52C975" box="[116,1421,867,893]" gridrow="16" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFCF6FE9EC975" box="[116,321,867,893]" gridcol="0" gridrow="16" pageId="4" pageNumber="3">Manus</th>
|
||
<td id="76A1F5198B436993FDE0FCF6FD51C975" box="[552,654,867,893]" gridcol="1" gridrow="16" pageId="4" pageNumber="3">W / D</td>
|
||
<td id="76A1F5198B436993FD35FCF6FBD4C975" box="[765,1035,867,893]" gridcol="2" gridrow="16" pageId="4" pageNumber="3">1.23 / 1.11</td>
|
||
<td id="76A1F5198B436993FB48FCF6FA52C975" box="[1152,1421,867,893]" gridcol="3" gridrow="16" pageId="4" pageNumber="3">1.55 / 1.50</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFC1FFA52C9AC" box="[116,1421,906,932]" gridrow="17" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFC1FFE9EC9AC" box="[116,321,906,932]" gridcol="0" gridrow="17" pageId="4" pageNumber="3">Movable finger</th>
|
||
<td id="76A1F5198B436993FDE0FC1FFD51C9AC" box="[552,654,906,932]" gridcol="1" gridrow="17" pageId="4" pageNumber="3">L</td>
|
||
<td id="76A1F5198B436993FD35FC1FFBD4C9AC" box="[765,1035,906,932]" gridcol="2" gridrow="17" pageId="4" pageNumber="3">2.56</td>
|
||
<td id="76A1F5198B436993FB48FC1FFA52C9AC" box="[1152,1421,906,932]" gridcol="3" gridrow="17" pageId="4" pageNumber="3">3.04</td>
|
||
</tr>
|
||
<tr id="35709C658B436993FFBCFC25FA52C9C3" box="[116,1421,944,971]" gridrow="18" pageId="4" pageNumber="3">
|
||
<th id="76A1F5198B436993FFBCFC25FE9EC9C3" box="[116,321,944,971]" gridcol="0" gridrow="18" pageId="4" pageNumber="3">Total</th>
|
||
<td id="76A1F5198B436993FDE0FC25FD51C9C3" box="[552,654,944,971]" gridcol="1" gridrow="18" pageId="4" pageNumber="3">L</td>
|
||
<td id="76A1F5198B436993FD35FC25FBD4C9C3" box="[765,1035,944,971]" gridcol="2" gridrow="18" pageId="4" pageNumber="3">23.32</td>
|
||
<td id="76A1F5198B436993FB48FC25FA52C9C3" box="[1152,1421,944,971]" gridcol="3" gridrow="18" pageId="4" pageNumber="3">28.55</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF3FCEAF8B439668FFBBFC7DFDD0CE12" pageId="4" pageNumber="3" startId="4.[115,171,1000,1021]" targetType="table">
|
||
<paragraph id="8BFF9E278B439668FFBBFC7DFDD0CE12" blockId="4.[115,1473,1000,1051]" pageId="4" pageNumber="3">
|
||
<emphasis id="B93442358B439668FFBBFC7DFF1DC9F5" bold="true" box="[115,194,1000,1021]" pageId="4" pageNumber="3">Table 1</emphasis>
|
||
. Comparative measurements of holotype male of
|
||
<taxonomicName id="4C40E5A48B439668FD77FC7CFCAFC9F5" authorityName="Kovařík & Stockmann & Šťáhlavský & Yong" authorityYear="2024" box="[703,880,1000,1022]" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="rolandoi" status="sp. nov.">
|
||
<emphasis id="B93442358B439668FD77FC7CFCAFC9F5" box="[703,880,1000,1022]" italics="true" pageId="4" pageNumber="3">Tityopsis rolandoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207FF4E8B439668FCB2FC7DFC6BC9F5" box="[890,948,1000,1021]" pageId="4" pageNumber="3" rank="species">
|
||
<emphasis id="B93442358B439668FCB2FC7DFC4EC9F5" bold="true" box="[890,913,1000,1021]" pageId="4" pageNumber="3">sp</emphasis>
|
||
.
|
||
<emphasis id="B93442358B439668FC69FC7DFC71C9F5" bold="true" box="[929,942,1000,1021]" pageId="4" pageNumber="3">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFF9E278B43966AFCFAF954FDEEC82B" blockId="4.[818,1476,1729,2041]" lastBlockId="6.[113,774,235,547]" lastPageId="6" lastPageNumber="5" pageId="4" pageNumber="3">
|
||
<emphasis id="B93442358B439668FCFAF954FC69CCD1" bold="true" box="[818,950,1729,1753]" pageId="4" pageNumber="3">Karyotypes</emphasis>
|
||
(
|
||
<figureCitation id="137B82A28B439668FC1BF954FBBACCD1" box="[979,1125,1729,1753]" captionStart="Figures 51–54" captionStartId="13.[115,192,1245,1267]" captionTargetBox="[113,1474,237,1233]" captionTargetId="figure-460@13.[113,1474,234,1233]" captionTargetPageId="13" captionText="Figures 51–54: The chromosomes of Tityopsis sheylae (2n=22) (51, 52) and T. rolandoi sp. n. (2n=20) (53, 54). Postpachytene (51, 53) and ideograms (52, 54) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values). Scale bar: 5 μm." figureDoi="http://doi.org/10.5281/zenodo.14660707" httpUri="https://zenodo.org/record/14660707/files/figure.png" pageId="4" pageNumber="3">Figs. 51–54</figureCitation>
|
||
). In total, we analyzed the chromosomes of
|
||
<specimenCount id="9D4655AE8B439668FC30F974FBBFCCF1" box="[1016,1120,1761,1785]" count="1" pageId="4" pageNumber="3" type="male">one male</specimenCount>
|
||
<taxonomicName id="4C40E5A48B439668FBA5F977FA1BCCF1" authority="(DNA 2826)" baseAuthorityName="DNA" baseAuthorityYear="2826" box="[1133,1476,1761,1785]" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="sheylae">
|
||
<emphasis id="B93442358B439668FBA5F977FAF6CCF1" box="[1133,1321,1761,1785]" italics="true" pageId="4" pageNumber="3">Tityopsis sheylae</emphasis>
|
||
(DNA 2826)
|
||
</taxonomicName>
|
||
from Cueva del Indio (GPS:
|
||
<geoCoordinate id="EE74F8E08B439668FBA4F894FB3FCD11" box="[1132,1248,1793,1817]" degrees="23.017507" direction="north" orientation="latitude" pageId="4" pageNumber="3" precision="1" value="23.017508">23.017507</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE74F8E08B439668FB23F894FAB8CD11" box="[1259,1383,1793,1817]" degrees="82.088714" direction="west" orientation="longitude" pageId="4" pageNumber="3" precision="1" value="-82.088715">-82.088714</geoCoordinate>
|
||
) and
|
||
<specimenCount id="9D4655AE8B439668FA6DF894FC0ACD31" count="6" pageId="4" pageNumber="3" type="male" typeStatus="paratype">six paratype males</specimenCount>
|
||
of
|
||
<taxonomicName id="4C40E5A48B439668FC34F8B7FBACCD31" authorityName="Kovařík & Stockmann & Šťáhlavský & Yong" authorityYear="2024" box="[1020,1139,1825,1849]" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="rolandoi" status="sp. n.">
|
||
<emphasis id="B93442358B439668FC34F8B7FBACCD31" box="[1020,1139,1825,1849]" italics="true" pageId="4" pageNumber="3">T. rolandoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207FF4E8B439668FBB3F8B4FB66CD30" box="[1147,1209,1825,1849]" pageId="4" pageNumber="3" rank="species">
|
||
<emphasis id="B93442358B439668FBB3F8B4FB4BCD31" bold="true" box="[1147,1172,1825,1849]" pageId="4" pageNumber="3">sp</emphasis>
|
||
.
|
||
<emphasis id="B93442358B439668FB6CF8B4FB6CCD31" bold="true" box="[1188,1203,1825,1849]" pageId="4" pageNumber="3">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(DNA 2829-2834) from the type locality. The chromosomes of both species display typical characteristics for members of the family
|
||
<taxonomicName id="4C40E5A48B439668FA95F8F4FA1FCD71" authorityName="C. L. Koch" authorityYear="1837" box="[1373,1472,1889,1913]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
|
||
, including holocentric organization, achiasmatic meiosis, and no differentiated sex chromosomes in males. Our initial findings on chromosomes within the genus
|
||
<taxonomicName id="4C40E5A48B439668FAE4F857FA54CDD1" authorityName="Armas" authorityYear="1974" box="[1324,1419,1986,2009]" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93442358B439668FAE4F857FA54CDD1" box="[1324,1419,1986,2009]" italics="true" pageId="4" pageNumber="3">Tityopsis</emphasis>
|
||
</taxonomicName>
|
||
also confirm the lower chromosome numbers commonly observed in
|
||
<taxonomicName id="4C40E5A48B41966AFF5DFF7EFF2ACB0B" authorityName="C. L. Koch" authorityYear="1837" box="[149,245,235,259]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="family">Buthidae</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EFD1E3D68B41966AFEF7FF7EFD99CB0B" author="SCHNEIDER M. C., V." box="[319,582,235,259]" pageId="6" pageNumber="5" refId="ref4938" refString="SCHNEIDER M. C., V. F. MATTOS & D. M. CELLA. 2024. The Scorpion Cytogenetic Database. Current version: 12 (October, 2024) https: // arthropodacytogenetics. bio. br / scorpiondatabase /" type="journal volume" year="2024">Schneider et al., 2024</bibRefCitation>
|
||
). The karyotype of
|
||
<taxonomicName id="4C40E5A48B41966AFF5CFE9EFEDBCB2B" baseAuthorityName="DNA" baseAuthorityYear="2826" box="[148,260,267,291]" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="sheylae">
|
||
<emphasis id="B93442358B41966AFF5CFE9EFEDBCB2B" box="[148,260,267,291]" italics="true" pageId="6" pageNumber="5">T. sheylae</emphasis>
|
||
</taxonomicName>
|
||
consists of 22 chromosomes (
|
||
<figureCitation id="137B82A28B41966AFDADFE9EFD65CB2A" box="[613,698,266,291]" captionStart="Figures 51–54" captionStartId="13.[115,192,1245,1267]" captionTargetBox="[113,1474,237,1233]" captionTargetId="figure-460@13.[113,1474,234,1233]" captionTargetPageId="13" captionText="Figures 51–54: The chromosomes of Tityopsis sheylae (2n=22) (51, 52) and T. rolandoi sp. n. (2n=20) (53, 54). Postpachytene (51, 53) and ideograms (52, 54) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values). Scale bar: 5 μm." figureDoi="http://doi.org/10.5281/zenodo.14660707" httpUri="https://zenodo.org/record/14660707/files/figure.png" pageId="6" pageNumber="5">Fig. 51</figureCitation>
|
||
), with chromosome lengths gradually decreasing from 6.98% to 2.32% of the diploid set (
|
||
<figureCitation id="137B82A28B41966AFE7FFEDEFDD3CB6A" box="[439,524,330,355]" captionStart="Figures 51–54" captionStartId="13.[115,192,1245,1267]" captionTargetBox="[113,1474,237,1233]" captionTargetId="figure-460@13.[113,1474,234,1233]" captionTargetPageId="13" captionText="Figures 51–54: The chromosomes of Tityopsis sheylae (2n=22) (51, 52) and T. rolandoi sp. n. (2n=20) (53, 54). Postpachytene (51, 53) and ideograms (52, 54) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values). Scale bar: 5 μm." figureDoi="http://doi.org/10.5281/zenodo.14660707" httpUri="https://zenodo.org/record/14660707/files/figure.png" pageId="6" pageNumber="5">Fig. 52</figureCitation>
|
||
). The karyotype of
|
||
<taxonomicName id="4C40E5A48B41966AFD27FEDEFF13CB8B" authorityName="Kovařík & Stockmann & Šťáhlavský & Yong" authorityYear="2024" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="rolandoi" status="sp. n.">
|
||
<emphasis id="B93442358B41966AFD27FEDEFF13CB8B" italics="true" pageId="6" pageNumber="5">T. rolandoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207FF4E8B41966AFF10FEFEFEC6CB8A" box="[216,281,363,387]" pageId="6" pageNumber="5" rank="species">
|
||
<emphasis id="B93442358B41966AFF10FEFEFF2ECB8B" bold="true" box="[216,241,363,387]" pageId="6" pageNumber="5">sp</emphasis>
|
||
.
|
||
<emphasis id="B93442358B41966AFECBFEFEFECDCB8B" bold="true" box="[259,274,363,387]" pageId="6" pageNumber="5">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
includes 20 chromosomes (
|
||
<figureCitation id="137B82A28B41966AFD9EFEFEFD76CB8A" box="[598,681,362,387]" captionStart="Figures 51–54" captionStartId="13.[115,192,1245,1267]" captionTargetBox="[113,1474,237,1233]" captionTargetId="figure-460@13.[113,1474,234,1233]" captionTargetPageId="13" captionText="Figures 51–54: The chromosomes of Tityopsis sheylae (2n=22) (51, 52) and T. rolandoi sp. n. (2n=20) (53, 54). Postpachytene (51, 53) and ideograms (52, 54) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values). Scale bar: 5 μm." figureDoi="http://doi.org/10.5281/zenodo.14660707" httpUri="https://zenodo.org/record/14660707/files/figure.png" pageId="6" pageNumber="5">Fig. 53</figureCitation>
|
||
), where the first three pairs appear slightly longer than the remaining chromosomes (
|
||
<figureCitation id="137B82A28B41966AFED1FE3EFEB2CBCA" box="[281,365,426,451]" captionStart="Figures 51–54" captionStartId="13.[115,192,1245,1267]" captionTargetBox="[113,1474,237,1233]" captionTargetId="figure-460@13.[113,1474,234,1233]" captionTargetPageId="13" captionText="Figures 51–54: The chromosomes of Tityopsis sheylae (2n=22) (51, 52) and T. rolandoi sp. n. (2n=20) (53, 54). Postpachytene (51, 53) and ideograms (52, 54) (y axis - % of the chromosome length of the diploid set, lines indicate min.-max. values). Scale bar: 5 μm." figureDoi="http://doi.org/10.5281/zenodo.14660707" httpUri="https://zenodo.org/record/14660707/files/figure.png" pageId="6" pageNumber="5">Fig. 54</figureCitation>
|
||
). We observed differences in the 2n and chromosome lengths between the two
|
||
<taxonomicName id="4C40E5A48B41966AFD85FE5EFCDCCBEB" box="[589,771,459,483]" pageId="6" pageNumber="5">
|
||
<emphasis id="B93442358B41966AFD85FE5EFD73CBEA" box="[589,684,459,482]" italics="true" pageId="6" pageNumber="5">Tityopsis</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
analyzed, suggesting that cytogenetic data may be valuable for taxonomic purposes within this genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3FCEAF8B429669FFB9F848FDC5C218" ID-DOI="http://doi.org/10.5281/zenodo.14660693" ID-Zenodo-Dep="14660693" httpUri="https://zenodo.org/record/14660693/files/figure.png" pageId="5" pageNumber="4" startId="5.[113,190,2013,2035]" targetBox="[162,1360,222,1996]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BFF9E278B429669FFB9F848FDC5C218" blockId="5.[113,1471,2013,2064]" pageId="5" pageNumber="4">
|
||
<emphasis id="B93442358B429669FFB9F848FF38CDFB" bold="true" box="[113,231,2013,2035]" pageId="5" pageNumber="4">Figures 2–5</emphasis>
|
||
:
|
||
<taxonomicName id="4C40E5A48B429669FF3CF84BFE7ECDFB" authorityName="Kovařík & Stockmann & Šťáhlavský & Yong" authorityYear="2024" box="[244,417,2014,2035]" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="rolandoi" status="sp. nov.">
|
||
<emphasis id="B93442358B429669FF3CF84BFE7ECDFB" box="[244,417,2014,2035]" italics="true" pageId="5" pageNumber="4">Tityopsis rolandoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207FF4E8B429669FE6EF84BFE03CDFB" box="[422,476,2014,2035]" pageId="5" pageNumber="4" rank="species">
|
||
<emphasis id="B93442358B429669FE6EF84BFE62CDFB" bold="true" box="[422,445,2014,2035]" pageId="5" pageNumber="4">sp</emphasis>
|
||
.
|
||
<emphasis id="B93442358B429669FE01F84BFE09CDFB" bold="true" box="[457,470,2014,2035]" pageId="5" pageNumber="4">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<emphasis id="B93442358B429669FE2AF848FD87CDFA" bold="true" box="[482,600,2013,2035]" pageId="5" pageNumber="4">Figures 2–3</emphasis>
|
||
. Male holotype in dorsal (2) and ventral (3) views.
|
||
<emphasis id="B93442358B429669FB8DF848FB63CDFB" bold="true" box="[1093,1212,2013,2035]" pageId="5" pageNumber="4">Figures 4–5</emphasis>
|
||
. Female paratype in dorsal (4) and ventral (5) views. Scale bar: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35ACDAC8B41966AFFB9FDDEFF71C90B" pageId="6" pageNumber="5" type="description">
|
||
<paragraph id="8BFF9E278B41966AFFB9FDDEFF71C90B" blockId="6.[113,773,587,771]" pageId="6" pageNumber="5">
|
||
SUBORDINATE TAXA.
|
||
<taxonomicName id="4C40E5A48B41966AFEA4FDDEFE55C88B" authority="Teruel & Rodriguez-Cabrera, 2022" authorityName="Teruel & Rodriguez-Cabrera" authorityYear="2022" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="canizaresorum">
|
||
<emphasis id="B93442358B41966AFEA4FDDEFDA0C86A" box="[364,639,587,610]" italics="true" pageId="6" pageNumber="5">Tityopsis canizaresorum</emphasis>
|
||
Teruel & Rodríguez-Cabrera, 2022
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C40E5A48B41966AFE53FDFEFD01C88B" authority="(Armas, 1974)" baseAuthorityName="Armas" baseAuthorityYear="1974" box="[411,734,619,643]" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="inaequalis">
|
||
<emphasis id="B93442358B41966AFE53FDFEFDF4C88B" box="[411,555,619,643]" italics="true" pageId="6" pageNumber="5">T. inaequalis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD1E3D68B41966AFD88FDFEFD09C88B" author="ARMAS" box="[576,726,619,643]" pageId="6" pageNumber="5" pagination="1 - 15" refId="ref4497" refString="ARMAS, L. F. DE. 1974. Escorpiones del archipielago cubano. 3. Genero Tityus C. L. Koch, 1836 (Scorpionida: Buthidae). Poeyana, 135: 1 - 15." type="journal article" year="1974">Armas, 1974</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C40E5A48B41966AFD27FDFEFE7CC8AB" authority="(Moreno, 1940)" baseAuthorityName="Moreno" baseAuthorityYear="1940" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="inexpectata">
|
||
<emphasis id="B93442358B41966AFD27FDFEFF33C8AA" italics="true" pageId="6" pageNumber="5">T. inexpectata</emphasis>
|
||
(Moreno, 1940)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C40E5A48B41966AFE79FD1EFED3C8CB" authority="Teruel & Rodriguez-Cabrera, 2022" authorityName="Teruel & Rodriguez-Cabrera" authorityYear="2022" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="mulata">
|
||
<emphasis id="B93442358B41966AFE79FD1EFDC8C8AB" box="[433,535,651,675]" italics="true" pageId="6" pageNumber="5">T. mulata</emphasis>
|
||
Teruel & Rodríguez-Cabrera, 2022
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C40E5A48B41966AFED0FD3EFD20C8CB" authority="Teruel & Rodriguez-Cabrera, 2022" authorityName="Teruel & Rodriguez-Cabrera" authorityYear="2022" box="[280,767,683,707]" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="pumila">
|
||
<emphasis id="B93442358B41966AFED0FD3EFEA2C8CB" box="[280,381,683,707]" italics="true" pageId="6" pageNumber="5">T. pumila</emphasis>
|
||
Teruel & Rodríguez-Cabrera, 2022
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C40E5A48B41966AFFB9FD5EFF3AC8EB" authorityName="Kovařík & Stockmann & Šťáhlavský & Yong" authorityYear="2024" box="[113,229,715,739]" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="rolandoi" status="sp. n.">
|
||
<emphasis id="B93442358B41966AFFB9FD5EFF3AC8EB" box="[113,229,715,739]" italics="true" pageId="6" pageNumber="5">T. rolandoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A207FF4E8B41966AFF23FD5EFEFAC8EA" box="[235,293,715,739]" pageId="6" pageNumber="5" rank="species">
|
||
<emphasis id="B93442358B41966AFF23FD5EFEDBC8EB" bold="true" box="[235,260,715,739]" pageId="6" pageNumber="5">sp</emphasis>
|
||
.
|
||
<emphasis id="B93442358B41966AFED8FD5EFEC0C8EB" bold="true" box="[272,287,715,739]" pageId="6" pageNumber="5">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C40E5A48B41966AFE96FD5EFF75C90B" authority="Teruel & Rodriguez-Cabrera, 2022" authorityName="Teruel & Rodriguez-Cabrera" authorityYear="2022" class="Arachnida" family="Buthidae" genus="Tityopsis" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="sheylae">
|
||
<emphasis id="B93442358B41966AFE96FD5EFE18C8EB" box="[350,455,715,739]" italics="true" pageId="6" pageNumber="5">T. sheylae</emphasis>
|
||
Teruel & Rodríguez-Cabrera, 2022
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35ACDAC8B41966AFFB9FCBEFD64C94B" box="[113,699,811,835]" pageId="6" pageNumber="5" type="distribution">
|
||
<paragraph id="8BFF9E278B41966AFFB9FCBEFD64C94B" blockId="6.[113,699,811,835]" box="[113,699,811,835]" pageId="6" pageNumber="5">
|
||
DISTRIBUTION. This genus is endemic to western
|
||
<collectingCountry id="F357DEB78B41966AFDB3FCBEFD68C94B" box="[635,695,811,835]" name="Cuba" pageId="6" pageNumber="5">Cuba</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |