treatments-xml/data/03/E1/98/03E1981EB537FFEBFF59F895044EECA5.xml

<document id="35E3F1601BA3E67D801C044C11F711DA" ID-DOI="10.11646/zootaxa.3948.1.2" ID-GBIF-Dataset="9e109059-3d75-4e2c-997e-6f4217d201d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="244003" ID-ZooBank="9A0330AD-EEF7-4A80-8D80-1B6116BD856F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461042933937" checkinUser="plazi" docAuthor="Filippini, Valentina, Galante, Eduardo &amp; Micó, Estefanía" docDate="2015" docId="03E1981EB537FFEBFF59F895044EECA5" docLanguage="en" docName="zt03948p040.pdf" docOrigin="Zootaxa 3948 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anomala estrella Filippini, Galante" docType="treatment" docVersion="7" masterDocId="FFD8E066B530FFE2FFCEFFD3014CEC5D" masterDocTitle="Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae)" updateTime="1698576078296" updateUser="plazi">
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<mods:title id="B1DEA7DCFCD60ACB1898B1A2EC4258BB">Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae)</mods:title>
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<mods:namePart id="66F35FBCDA0FC4AFEE239FE651FBFB31">Filippini, Valentina</mods:namePart>
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<mods:namePart id="25069EF8F6683B8FA6C43F7D1C218BEB">Galante, Eduardo</mods:namePart>
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<mods:namePart id="252FE92282C1E34827FDAF1475B147AA">Micó, Estefanía</mods:namePart>
</mods:name>
<mods:typeOfResource id="FD0E40DD054F7282DDBBA6ECABD115A0">text</mods:typeOfResource>
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<mods:title id="22636CFB36FBB30A1A82BAC2F2A34890">Zootaxa</mods:title>
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<mods:part id="89FE05F8956F399583026E22510F9C28">
<mods:date id="F46B7D535A7AA0E1C8BE298D5B8EE9B7">2015</mods:date>
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<treatment id="03E1981EB537FFEBFF59F895044EECA5" ID-DOI="http://doi.org/10.5281/zenodo.6122700" ID-GBIF-Taxon="119609992" ID-Zenodo-Dep="6122700" LSID="urn:lsid:plazi:treatment:03E1981EB537FFEBFF59F895044EECA5" httpUri="http://treatment.plazi.org/id/03E1981EB537FFEBFF59F895044EECA5" lastPageId="9" pageId="7">
<subSubSection id="C3527A83B537FFE5FF59F8950195EBDF" pageId="7" type="nomenclature">
<paragraph id="8BF72908B537FFE5FF59F895021AEB3C" blockId="7.[151,854,1862,1922]" box="[151,854,1862,1889]" pageId="7">
<heading id="D0BF9E64B537FFE5FF59F895021AEB3C" bold="true" box="[151,854,1862,1889]" fontSize="11" level="1" pageId="7" reason="1">
<emphasis id="B93CF51AB537FFE5FF59F895021AEB3C" bold="true" box="[151,854,1862,1889]" pageId="7">
<taxonomicName id="4C48528BB537FFE5FF59F8950307EB3C" authority="Filippini, Galante" authorityName="Filippini, Galante" box="[151,587,1862,1889]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="estrella">
<emphasis id="B93CF51AB537FFE5FF59F8950029EB3D" bold="true" box="[151,357,1862,1888]" italics="true" pageId="7">Anomala estrella</emphasis>
<collectingCountry id="F35F6998B537FFE5FEA2F8940095EB3C" box="[364,473,1863,1889]" name="Philippines" pageId="7">Filippini</collectingCountry>
, Galante
</taxonomicName>
, &amp; Micó, new species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF72908B537FFE5FF59F8BA0195EBDF" blockId="7.[151,854,1862,1922]" box="[151,217,1897,1922]" pageId="7">
<figureCitation id="1373358DB537FFE5FF59F8BA0195EBDF" box="[151,217,1897,1922]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1977,2000]" captionTargetBox="[186,1400,705,1952]" captionTargetId="figure@4.[179,1408,697,1957]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Habitus. 1: Anomala aglaos (holotype). 2: A. estrella (holotype). 3: A. inbio (holotype). 4: A. pincelada (holotype). 5: Callistethus ruteloides (holotype). 6: C. yalizo (holotype). Scale = 5 mm." httpUri="https://zenodo.org/record/244004/files/figure.png" pageId="7">Fig. 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3527A83B537FFEAFF59F8620515E999" lastPageId="8" pageId="7" type="materials_examined">
<paragraph id="8BF72908B537FFE5FF59F86205CCEBB3" blockId="7.[151,1436,1969,2030]" pageId="7">
<emphasis id="B93CF51AB537FFE5FF59F86200CDEB97" bold="true" box="[151,385,1969,1994]" pageId="7">Material examined.</emphasis>
<typeStatus id="54F397AAB537FFE5FE47F86200BAEB97" box="[393,502,1969,1994]" pageId="7" type="holotype">Holotype</typeStatus>
: ♂ &quot;Est. La Casona, 
<quantity id="4CB084EDB537FFE5FD2DF862027FEB97" box="[739,819,1969,1994]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="7" unit="m" value="1520.0">1520m</quantity>
, Res. Biol. Monteverde, Prov. Puntarenas, 
<collectingCountry id="F35F6998B537FFE5FAD4F86104D4EB97" box="[1306,1432,1969,1994]" name="Costa Rica" pageId="7">Costa Rica</collectingCountry>
, N. Obando, 
<date id="FFF60FC8B537FFE5FEEEF80500C1EBB3" box="[288,397,2006,2030]" pageId="7" value="1992-03">Mar 1992</date>
L-N 253250,449700 / INBIOCRI000898153&quot; deposited in 
<taxonomicName id="4C48528BB537FFE5FBE5F8050530EBB3" box="[1067,1148,2006,2030]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED59B1CDB537FFE5FBE5F8050530EBB3" box="[1067,1148,2006,2030]" pageId="7">INBIO</collectionCode>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF72908B538FFEAFF09FF4B0521EDA8" blockId="8.[151,1437,151,2015]" pageId="8">
<typeStatus id="54F397AAB538FFEAFF09FF4B0078ECED" box="[199,308,152,176]" pageId="8" type="paratype">Paratypes</typeStatus>
(8): 1♀ &quot;Estac. Maritza, 
<specimenCount id="9D4EE281B538FFEAFD90FF4403E3ECF2" box="[606,687,151,176]" pageId="8" type="generic">
<quantity id="4CB084EDB538FFEAFD90FF4403E7ECF2" box="[606,683,151,176]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="8" unit="m" value="600.0">600 m</quantity>
.
</specimenCount>
W side Volcan Orosi Guanac. Pr. 
<collectingCountry id="F35F6998B538FFEAFB8CFF4B05A1ECED" box="[1090,1261,152,176]" name="Costa Rica" pageId="8">COSTA RICA</collectingCountry>
. 
<date id="FFF60FC8B538FFEAFB34FF4B04D4ECED" box="[1274,1432,152,176]" pageId="8" value="1988-05-17">17 May 1988</date>
. Janzen &amp; Hallwachs 
<geoCoordinate id="EE7C4FCFB538FFEAFE49FF6F0345EC89" box="[391,521,188,213]" direction="west" orientation="longitude" pageId="8" precision="15" value="-85.49361">W85 29 37</geoCoordinate>
&quot;, 
<geoCoordinate id="EE7C4FCFB538FFEAFDEEFF6F03D6EC89" box="[544,666,188,213]" direction="north" orientation="latitude" pageId="8" precision="15" value="10.960834">N10 57 39</geoCoordinate>
&quot; / INBIOCRI002517167&quot;; 
<specimenCount id="9D4EE281B538FFEAFC1EFF6F02BFEC89" box="[976,1011,188,212]" pageId="8" type="generic">1♂</specimenCount>
&quot;San Jose. 
<collectingCountry id="F35F6998B538FFEAFBB6FF6F05BFEC88" box="[1144,1267,188,213]" name="Costa Rica" pageId="8">Costa Rica</collectingCountry>
Hda. Tiquires. 
<quantity id="4CB084EDB538FFEAFF59FF0C01A9ECA5" box="[151,229,223,248]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="8" unit="m" value="1500.0">1500m</quantity>
Rio Tiquires 
<date id="FFF60FC8B538FFEAFE43FF330372ECA5" box="[397,574,224,248]" pageId="8" value="1988-03-28">28 marzo 1988</date>
Col.A.Solis / 
<collectionCode id="ED59B1CDB538FFEAFD23FF0C0270ECA5" box="[749,828,223,248]" pageId="8">CEUA</collectionCode>
00106164&quot;; 2♀ &quot;Est. La Casona, 
<quantity id="4CB084EDB538FFEAFB09FF0C045BECA5" box="[1223,1303,223,248]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="8" unit="m" value="1520.0">1520m</quantity>
, Res. Biol. Monteverde, Prov. Puntarenas, 
<collectingCountry id="F35F6998B538FFEAFE30FED70332ED40" box="[510,638,260,285]" name="Costa Rica" pageId="8">Costa Rica</collectingCountry>
, N. Obando, 
<date id="FFF60FC8B538FFEAFCD4FED602C6ED41" box="[794,906,260,284]" pageId="8" value="1992-03">Mar 1992</date>
L-N 253250,449700&quot; / INBIOCRI000898149 and INBIOCRI000898152; 
<specimenCount id="9D4EE281B538FFEAFE01FEFB00BEED1D" box="[463,498,296,320]" pageId="8" type="generic">1♂</specimenCount>
&quot;Est. Las Pailas, 
<quantity id="4CB084EDB538FFEAFD78FEFB03B3ED62" box="[694,767,296,320]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" unit="m" value="800.0">800 m</quantity>
, P.N. Rincon de la Vieja, Prov. Guan., 
<collectingCountry id="F35F6998B538FFEAFB61FEFB041BED1D" box="[1199,1367,296,320]" name="Costa Rica" pageId="8">COSTA RICA</collectingCountry>
. 
<date id="FFF60FC8B538FFEAFAAFFEF4007BED39" pageId="8" value="1993-01-06" valueMax="1993-04-13" valueMin="1993-01-06">6 ene a 13 abr 1993</date>
, J. Sihezar, G. Rodriguez, L- N 306300_388600 / INBIOCRI001331532&quot;; 1♀ &quot;Est. Las Pailas, 
<quantity id="4CB084EDB538FFEAFABCFE9F01E3EDDA" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" unit="m" value="800.0">800 m</quantity>
, P.N. Rincon de la Vieja, Prov. Guan., 
<collectingCountry id="F35F6998B538FFEAFDB6FEA3026FEDD5" box="[632,803,368,392]" name="Costa Rica" pageId="8">COSTA RICA</collectingCountry>
. 
<date id="FFF60FC8B538FFEAFCFEFEA3054AEDD5" box="[816,1030,368,392]" pageId="8" value="1993-03-10" valueMax="1993-03-27" valueMin="1993-03-10">10 a 27 Mar 1993</date>
, K. Taylor, L- N 306300_388600 / INBIOCRI001355780&quot;; 
<specimenCount id="9D4EE281B538FFEAFE63FE47009CEDF1" box="[429,464,404,428]" pageId="8" type="generic">1♂</specimenCount>
&quot;Est. Maritza, 
<quantity id="4CB084EDB538FFEAFDB3FE47038BEDF1" box="[637,711,404,429]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="8" unit="m" value="600.0">600 m</quantity>
, lado 
<collectingCountry id="F35F6998B538FFEAFCC3FE47026DEDF1" box="[781,801,404,428]" name="Iceland" pageId="8">O</collectingCountry>
Vol. Orosi, Prov. Guan. 
<collectingCountry id="F35F6998B538FFEAFBF2FE4705A9EDF1" box="[1084,1253,404,428]" name="Costa Rica" pageId="8">COSTA RICA</collectingCountry>
. R. Blanco, 
<date id="FFF60FC8B538FFEAFABFFE47019EED8D" pageId="8" value="1990-04">Abr 1990</date>
, L- N 326900_373000 / INBIOCRI000238024&quot;; 1♀ &quot;Los Almendros, P. N. Guanacaste, Prov. Guanacaste, 
<collectingCountry id="F35F6998B538FFEAFF59FE0F005FEDA8" box="[151,275,476,501]" name="Costa Rica" pageId="8">Costa Rica</collectingCountry>
, 
<date id="FFF60FC8B538FFEAFED1FE0F035DEDA9" box="[287,529,476,501]" pageId="8" value="1992-03-28" valueMax="1992-04-24" valueMin="1992-03-28">28 mar a 24 abr 1992</date>
, M. Reyes, L- N 334800_369800 / 
<collectionCode id="ED59B1CDB538FFEAFC53FE0F02A0EDA8" box="[925,1004,476,501]" pageId="8">CEUA</collectionCode>
00105806&quot;.
</paragraph>
<paragraph id="8BF72908B538FFEAFF09FE2C053BEE4A" blockId="8.[151,1437,151,2015]" box="[199,1143,511,536]" pageId="8">
<emphasis id="B93CF51AB538FFEAFF09FE2C0344EE45" bold="true" box="[199,520,511,536]" pageId="8">
<typeStatus id="54F397AAB538FFEAFF09FE2C0078EE45" box="[199,308,511,536]" pageId="8" type="holotype">Holotype</typeStatus>
male description.
</emphasis>
Body shape elongate. Length 
<quantity id="4CB084EDB538FFEAFC93FDD30288EE4A" box="[861,964,512,536]" metricMagnitude="-2" metricUnit="m" metricValue="1.49" pageId="8" unit="mm" value="14.9">14.9 mm</quantity>
. Width 
<quantity id="4CB084EDB538FFEAFBD4FDD3053FEE4A" box="[1050,1139,512,536]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="8" unit="mm" value="8.4">8.4 mm</quantity>
.
</paragraph>
<paragraph id="8BF72908B538FFEAFF09FDF202FAEE97" blockId="8.[151,1437,151,2015]" pageId="8">Color. Head dark brown. Pronotum orange ochre with 1 large irregular dark brown macula on disc, not reaching basal margin, crossed by a lighter weak longitudinal line; small maculae on lateral foveae; surface with green luster. Scutellum light brown with lighter center. Elytra orange ochre. Pygidium ochre. Legs reddish brown; protibiae with darker margins, metafemura ochre, metatibiae and metatarsi dark brown with green luster. Venter, sternum reddish brown with darker margins, abdominal sternites ochre.</paragraph>
<paragraph id="8BF72908B538FFEAFF09FD0603BFEF6B" blockId="8.[151,1437,151,2015]" pageId="8">Clypeus trapezoidal, surface densely punctate-reticulate, flat. Anterior margin straight, clypeus anteriorly thin. Clypeus ratio width/length 2.3. Frons densely punctate, flat. Ocular canthum long, thin, with rounded apex. Interocular ratio (interocular width/width of eye): 2.9.</paragraph>
<paragraph id="8BF72908B538FFEAFF09FC9101B1EFFC" blockId="8.[151,1437,151,2015]" pageId="8">Pronotum subtrapezoidal. Ratio width/length 1.7. Lateral margins angulated at 1/3 of pronotum length. Apical angles acute, sharp; basal angles obtuse, blunt. Basal margin sinuate, subapical bead complete. Surface densely punctate.</paragraph>
<paragraph id="8BF72908B538FFEAFF09FC7E01B1EFB4" blockId="8.[151,1437,151,2015]" pageId="8">Scutellum. Shape subtriangular, with rounded sides, apex acute. Ratio width/length 1.5. Surface densely punctate.</paragraph>
<paragraph id="8BF72908B538FFEAFF09FC2B05F6E86A" blockId="8.[151,1437,151,2015]" pageId="8">Elytra with costae defined by regular rows of punctures. Subsutural interstice with 3–5 rows of punctures, 2nd– 4th interstices with 1–2 irregular rows of punctures. Surface covered with secondary punctures.</paragraph>
<paragraph id="8BF72908B538FFEAFF09FB90052BE806" blockId="8.[151,1437,151,2015]" box="[199,1127,1091,1116]" pageId="8">Pygidium finely granulate with long setae at apical margin. Ratio width/length 1.4.</paragraph>
<paragraph id="8BF72908B538FFEAFF09FBB40230E89A" blockId="8.[151,1437,151,2015]" pageId="8">Space between the mesocoxae narrow, flat. Mesometasternal suture well defined, at base of mesocoxae. Abdominal sternites with 1 row of sparse setae and 5–6 rows of punctures per sternite. Terminal sternite punctate, with subapical bead narrow and sinuate, apical membrane narrow.</paragraph>
<paragraph id="8BF72908B538FFEAFF09FB00040EE90A" blockId="8.[151,1437,151,2015]" pageId="8">Protibia (Fig. 8) bidentate; apical tooth long and curved; basal tooth below internal apex of protibia, triangular in shape, acute. Metatibia stout, slightly narrower subapically. Ratio length/width 2.6. First external carina obliterated; surface densely punctate above second external carina, rugose below. Protarsal claws: internal claw bifurcate, with upper branch of same length and 2/3 the width of the lower branch; inferior margin sinuate.</paragraph>
<paragraph id="8BF72908B538FFEAFF09FAB00515E999" blockId="8.[151,1437,151,2015]" pageId="8">
Aedeagus (
<figureCitation id="1373358DB538FFEAFE83FAB000E5E926" box="[333,425,1379,1404]" captionStart="FIGURES 13 – 18" captionStartId="6.[151,264,1885,1908]" captionTargetBox="[153,1429,1020,1837]" captionTargetId="figure@6.[151,1436,1010,1864]" captionTargetPageId="6" captionText="FIGURES 13 – 18. Shape of aedeagus, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 13: Anomala aglaos (paratype, Isla Bonita, Costa Rica). 14: A. estrella (holotype). 15: A. inbio (holotype). 16: A. pincelada (holotype). 17: Callistethus ruteloides (holotype). 18: C. yalizo (Cerro Chompipe, Heredia). Scale = 1 mm." httpUri="https://zenodo.org/record/244005/files/figure.png" pageId="8">Fig. 14</figureCitation>
): parameres with sinuate dorsal and ventral margins, apex acute in a lateral view. Endophallus (
<figureCitation id="1373358DB538FFEAFEFAFA5500CBE9C2" box="[308,391,1414,1439]" captionStart="FIGURES 19 – 22" captionStartId="7.[151,264,1709,1732]" captionTargetBox="[155,1421,200,1673]" captionTargetId="figure@7.[151,1436,193,1688]" captionTargetPageId="7" captionText="FIGURES 19 – 22. Endophallus of 19: Anomala aglaos, (a) particular in frontal view, (b) lateral view (paratype, Isla Bonita, Costa Rica). 20: A. estrella (paratype, Hacienda Tiquires, Costa Rica). 21: A. inbio (holotype). 22: A. pincelada (paratype, Cuajiniquil, Costa Rica). Scale = 1 mm." httpUri="https://zenodo.org/record/244006/files/figure.png" pageId="8">Fig. 20</figureCitation>
) with 1 long sacculus with wide apex, covered with setae medially and with a strip of long and short, thick spines at apex. Ejaculatory duct opening placed on a ventral inflation.
</paragraph>
</subSubSection>
<subSubSection id="C3527A83B538FFEAFF09FA1D00DBEAC1" pageId="8" type="description">
<paragraph id="8BF72908B538FFEAFF09FA1D04D4EA72" blockId="8.[151,1437,151,2015]" pageId="8">
<emphasis id="B93CF51AB538FFEAFF09FA1D0077E9BA" bold="true" box="[199,315,1486,1511]" pageId="8">Variation.</emphasis>
Macula on pronotum with variable extension either fusing with maculae on foveae, forming a starshaped macula or reaching the posterior margin; elytra ochre to red. Female. Similar to male, protibia (Fig. 8) with longer and wider apical tooth; basal tooth above internal apex; internal protarsal claw with curved inferior margin.
</paragraph>
<paragraph id="8BF72908B538FFEAFF09F9E800DBEAC1" blockId="8.[151,1437,151,2015]" pageId="8">
Body length 
<quantity id="4CB084EDB538FFEAFEACF9E80340EA0E" box="[354,524,1595,1620]" metricMagnitude="-2" metricUnit="m" metricValue="1.525" metricValueMax="1.61" metricValueMin="1.44" pageId="8" unit="mm" value="15.25" valueMax="16.1" valueMin="14.4">14.4–16.1 mm</quantity>
, body width 
<quantity id="4CB084EDB538FFEAFD63F9E80270EA0E" box="[685,828,1595,1620]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" metricValueMax="9.6" metricValueMin="7.9" pageId="8" unit="mm" value="8.75" valueMax="9.6" valueMin="7.9">7.9–9.6 mm</quantity>
. Clypeus w/l: 2.2–2.3. Interocular ratio (interocular width/width of eye): 2.5–3.0. Antenna: ratio funiculus/club 0.6–0.7. Scutellum w/l: 1.2–1.5. Pygidium w/l: 1.4–1.6. Metatibia l/w: 2.5–2.8.
</paragraph>
</subSubSection>
<subSubSection id="C3527A83B538FFEAFF09F97503BBEB82" pageId="8" type="diagnosis">
<paragraph id="8BF72908B538FFEAFF09F97505F3EABE" blockId="8.[151,1437,151,2015]" pageId="8">
<emphasis id="B93CF51AB538FFEAFF09F975000DEAE2" bold="true" box="[199,321,1702,1727]" pageId="8">Diagnosis.</emphasis>
Size large; macula on pronotum irregular, dark; elytra ochre; parameres with sinuate dorsal and ventral margins in lateral view; endophallus with a strip of long and short, thick spines at apex.
</paragraph>
<paragraph id="8BF72908B538FFEAFF09F93C03BBEB82" blockId="8.[151,1437,151,2015]" pageId="8">
<taxonomicName id="4C48528BB538FFEAFF09F93C0356EB5A" authority="Ohaus, 1897" authorityName="Ohaus" authorityYear="1897" box="[199,538,1774,1799]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="jansoni">
<emphasis id="B93CF51AB538FFEAFF09F93C00C9EB5A" box="[199,389,1775,1799]" italics="true" pageId="8">Anomala jansoni</emphasis>
Ohaus, 1897
</taxonomicName>
is a similar species of similar size but with two small maculae on pronotum and parameres with long, narrow, curved apex (
<figureCitation id="1373358DB538FFEAFD58F8C003A0EB76" box="[662,748,1811,1836]" captionStart="FIGURES 27 – 33. 27" captionStartId="14.[151,264,1539,1562]" captionTargetBox="[156,1433,641,1510]" captionTargetId="figure@14.[151,1436,630,1517]" captionTargetPageId="14" captionText="FIGURES 27 – 33. 27: Aedeagus of Anomala obovata, lateral view (Quebrada Kuisa, Costa Rica, INBIO). 28: Endophallus of A. obovata (Cerro Chompipe, Costa Rica, INBIO). 29: Aedeagus of A. jansoni, lateral view (Monte Rotondo, Costa Rica, syntype, MNHUB). 30: Aedeagus of A. antica, lateral view (Estación Palo Verde, Costa Rica, INBIO). 31: Aedeagus of A. luciae, lateral view (Santa Lucia, Antilles, lectotype, MNHN). 32: Aedeagus of A. quiche, lateral view (Estación Maritza, Costa Rica, INBIO). 33: Aedeagus of A. inconstans, (a) lateral view, (b) other side of parameres (Bogotá, Colombia, syntype, MLUH). Scale = 1 mm." httpUri="https://zenodo.org/record/244009/files/figure.png" pageId="8">Fig. 29</figureCitation>
) (Monte Rotondo, 
<collectingCountry id="F35F6998B538FFEAFC1EF8C0051EEB71" box="[976,1106,1811,1836]" name="Costa Rica" pageId="8">Costa Rica</collectingCountry>
; MNHUB). 
<taxonomicName id="4C48528BB538FFEAFB2BF8C0006BEB12" authority="Ohaus, 1897" authorityName="Ohaus" authorityYear="1897" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="antica">
<emphasis id="B93CF51AB538FFEAFB2BF8C004D0EB76" box="[1253,1436,1811,1835]" italics="true" pageId="8">Anomala antica</emphasis>
Ohaus, 1897
</taxonomicName>
, 
<taxonomicName id="4C48528BB538FFEAFEFCF8E4031BEB12" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[306,599,1846,1871]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="luciae">
<emphasis id="B93CF51AB538FFEAFEFCF8E400DAEB12" box="[306,406,1847,1871]" italics="true" pageId="8">A. luciae</emphasis>
Blanchard, 1851
</taxonomicName>
, and 
<taxonomicName id="4C48528BB538FFEAFD5CF8E402DCEB12" authority="Ohaus, 1897" authorityName="Ohaus" authorityYear="1897" box="[658,912,1846,1871]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="quiche">
<emphasis id="B93CF51AB538FFEAFD5CF8E403B0EB12" box="[658,764,1847,1871]" italics="true" pageId="8">A. quiche</emphasis>
Ohaus, 1897
</taxonomicName>
are similar but smaller species, with one small and pentagonal macula on pronotum (
<taxonomicName id="4C48528BB538FFEAFD82F88803F8EB2E" box="[588,692,1883,1907]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="antica">
<emphasis id="B93CF51AB538FFEAFD82F88803F8EB2E" box="[588,692,1883,1907]" italics="true" pageId="8">A. antica</emphasis>
</taxonomicName>
), pronotum with sinuate lateral margins (
<taxonomicName id="4C48528BB538FFEAFB5DF888044CEB2E" box="[1171,1280,1883,1907]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="quiche">
<emphasis id="B93CF51AB538FFEAFB5DF888044CEB2E" box="[1171,1280,1883,1907]" italics="true" pageId="8">A. quiche</emphasis>
</taxonomicName>
), maculae on elytra (
<taxonomicName id="4C48528BB538FFEAFF28F8AC0005EBCA" box="[230,329,1919,1943]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="luciae">
<emphasis id="B93CF51AB538FFEAFF28F8AC0005EBCA" box="[230,329,1919,1943]" italics="true" pageId="8">A. luciae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C48528BB538FFEAFEB1F8AC00A5EBCA" box="[383,489,1919,1943]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="quiche">
<emphasis id="B93CF51AB538FFEAFEB1F8AC00A5EBCA" box="[383,489,1919,1943]" italics="true" pageId="8">A. quiche</emphasis>
</taxonomicName>
), and differences in the parameres: 
<taxonomicName id="4C48528BB538FFEAFCB6F8AC0291EBCA" box="[888,989,1919,1943]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="antica">
<emphasis id="B93CF51AB538FFEAFCB6F8AC0291EBCA" box="[888,989,1919,1943]" italics="true" pageId="8">A. antica</emphasis>
</taxonomicName>
has a long thin apex (
<figureCitation id="1373358DB538FFEAFB1CF8AD0469EBCA" box="[1234,1317,1918,1943]" captionStart="FIGURES 27 – 33. 27" captionStartId="14.[151,264,1539,1562]" captionTargetBox="[156,1433,641,1510]" captionTargetId="figure@14.[151,1436,630,1517]" captionTargetPageId="14" captionText="FIGURES 27 – 33. 27: Aedeagus of Anomala obovata, lateral view (Quebrada Kuisa, Costa Rica, INBIO). 28: Endophallus of A. obovata (Cerro Chompipe, Costa Rica, INBIO). 29: Aedeagus of A. jansoni, lateral view (Monte Rotondo, Costa Rica, syntype, MNHUB). 30: Aedeagus of A. antica, lateral view (Estación Palo Verde, Costa Rica, INBIO). 31: Aedeagus of A. luciae, lateral view (Santa Lucia, Antilles, lectotype, MNHN). 32: Aedeagus of A. quiche, lateral view (Estación Maritza, Costa Rica, INBIO). 33: Aedeagus of A. inconstans, (a) lateral view, (b) other side of parameres (Bogotá, Colombia, syntype, MLUH). Scale = 1 mm." httpUri="https://zenodo.org/record/244009/files/figure.png" pageId="8">Fig. 30</figureCitation>
), 
<taxonomicName id="4C48528BB538FFEAFAF7F8AC04D0EBCA" box="[1337,1436,1919,1943]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="luciae">
<emphasis id="B93CF51AB538FFEAFAF7F8AC04D0EBCA" box="[1337,1436,1919,1943]" italics="true" pageId="8">A. luciae</emphasis>
</taxonomicName>
has lobes on anterior portion of inferior margins (
<figureCitation id="1373358DB538FFEAFD16F8700261EBE6" box="[728,813,1955,1980]" captionStart="FIGURES 27 – 33. 27" captionStartId="14.[151,264,1539,1562]" captionTargetBox="[156,1433,641,1510]" captionTargetId="figure@14.[151,1436,630,1517]" captionTargetPageId="14" captionText="FIGURES 27 – 33. 27: Aedeagus of Anomala obovata, lateral view (Quebrada Kuisa, Costa Rica, INBIO). 28: Endophallus of A. obovata (Cerro Chompipe, Costa Rica, INBIO). 29: Aedeagus of A. jansoni, lateral view (Monte Rotondo, Costa Rica, syntype, MNHUB). 30: Aedeagus of A. antica, lateral view (Estación Palo Verde, Costa Rica, INBIO). 31: Aedeagus of A. luciae, lateral view (Santa Lucia, Antilles, lectotype, MNHN). 32: Aedeagus of A. quiche, lateral view (Estación Maritza, Costa Rica, INBIO). 33: Aedeagus of A. inconstans, (a) lateral view, (b) other side of parameres (Bogotá, Colombia, syntype, MLUH). Scale = 1 mm." httpUri="https://zenodo.org/record/244009/files/figure.png" pageId="8">Fig. 31</figureCitation>
), and 
<taxonomicName id="4C48528BB538FFEAFCB6F87002A9EBE6" box="[888,997,1955,1979]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="quiche">
<emphasis id="B93CF51AB538FFEAFCB6F87002A9EBE6" box="[888,997,1955,1979]" italics="true" pageId="8">A. quiche</emphasis>
</taxonomicName>
has wide rounded parameres, and the ventral plate curved at apex in a lateral view (
<figureCitation id="1373358DB538FFEAFD58F81503A7EB82" box="[662,747,1990,2015]" captionStart="FIGURES 27 – 33. 27" captionStartId="14.[151,264,1539,1562]" captionTargetBox="[156,1433,641,1510]" captionTargetId="figure@14.[151,1436,630,1517]" captionTargetPageId="14" captionText="FIGURES 27 – 33. 27: Aedeagus of Anomala obovata, lateral view (Quebrada Kuisa, Costa Rica, INBIO). 28: Endophallus of A. obovata (Cerro Chompipe, Costa Rica, INBIO). 29: Aedeagus of A. jansoni, lateral view (Monte Rotondo, Costa Rica, syntype, MNHUB). 30: Aedeagus of A. antica, lateral view (Estación Palo Verde, Costa Rica, INBIO). 31: Aedeagus of A. luciae, lateral view (Santa Lucia, Antilles, lectotype, MNHN). 32: Aedeagus of A. quiche, lateral view (Estación Maritza, Costa Rica, INBIO). 33: Aedeagus of A. inconstans, (a) lateral view, (b) other side of parameres (Bogotá, Colombia, syntype, MLUH). Scale = 1 mm." httpUri="https://zenodo.org/record/244009/files/figure.png" pageId="8">Fig. 32</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3527A83B539FFEBFF09FF440093EC88" pageId="9" type="etymology">
<paragraph id="8BF72908B539FFEBFF09FF440093EC88" blockId="9.[151,1435,151,248]" pageId="9">
<emphasis id="B93CF51AB539FFEBFF09FF440002ECED" bold="true" box="[199,334,151,176]" pageId="9">Etymology.</emphasis>
Latinized from the Spanish noun 
<taxonomicName id="4C48528BB539FFEBFD1BFF4B0264ECED" box="[725,808,152,176]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="estrella">
<emphasis id="B93CF51AB539FFEBFD1BFF4B0264ECED" box="[725,808,152,176]" italics="true" pageId="9">estrella</emphasis>
</taxonomicName>
, star, for the star-shaped macula on pronotum. Treated here as a noun in apposition,.
</paragraph>
</subSubSection>
<subSubSection id="C3527A83B539FFEBFF09FF0C044EECA5" box="[199,1282,223,248]" pageId="9" type="distribution">
<paragraph id="8BF72908B539FFEBFF09FF0C044EECA5" blockId="9.[151,1435,151,248]" box="[199,1282,223,248]" pageId="9">
<emphasis id="B93CF51AB539FFEBFF09FF0C002EECA5" bold="true" box="[199,354,223,248]" pageId="9">Distribution:</emphasis>
Guanacaste and Tilarán mountain ranges, 
<collectingCountry id="F35F6998B539FFEBFC8EFF3302F1ECA5" box="[832,957,223,248]" name="Costa Rica" pageId="9">Costa Rica</collectingCountry>
, from 
<quantity id="4CB084EDB539FFEBFBCBFF0C05DEECAA" box="[1029,1170,223,248]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="15.2" metricValueMin="2.8" pageId="9" unit="m" value="900.0" valueMax="1520.0" valueMin="280.0">280–1520 m</quantity>
(
<figureCitation id="1373358DB539FFEBFB6FFF0C05BAECA5" box="[1185,1270,223,248]" captionStart="FIGURE 34 – 37" captionStartId="15.[151,250,1947,1969]" captionTargetBox="[158,1427,933,1891]" captionTargetId="figure@15.[151,1436,925,1898]" captionTargetPageId="15" captionText="FIGURE 34 – 37. Distribution of 34: Anomala aglaos and A. estrella. 35: A. inbio and A. pincelada. 36: Callistethus ruteloides. 37: C. yalizo." httpUri="https://zenodo.org/record/244010/files/figure.png" pageId="9">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>