Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae) Author Filippini, Valentina Author Galante, Eduardo Author Micó, Estefanía text Zootaxa 2015 3948 1 journal volume 10.11646/zootaxa.3948.1.2 9e109059-3d75-4e2c-997e-6f4217d201d0 1175-5326 244003 9A0330AD-EEF7-4A80-8D80-1B6116BD856F Anomala estrella Filippini , Galante , & Micó, new species Fig. 2 Material examined. Holotype : ♂ "Est. La Casona, 1520m , Res. Biol. Monteverde, Prov. Puntarenas, Costa Rica , N. Obando, Mar 1992 L-N 253250,449700 / INBIOCRI000898153" deposited in INBIO . Paratypes (8): 1♀ "Estac. Maritza, 600 m . W side Volcan Orosi Guanac. Pr. COSTA RICA . 17 May 1988 . Janzen & Hallwachs W85 29 37 ", N10 57 39 " / INBIOCRI002517167"; 1♂ "San Jose. Costa Rica Hda. Tiquires. 1500m Rio Tiquires 28 marzo 1988 Col.A.Solis / CEUA 00106164"; 2♀ "Est. La Casona, 1520m , Res. Biol. Monteverde, Prov. Puntarenas, Costa Rica , N. Obando, Mar 1992 L-N 253250,449700" / INBIOCRI000898149 and INBIOCRI000898152; 1♂ "Est. Las Pailas, 800 m , P.N. Rincon de la Vieja, Prov. Guan., COSTA RICA . 6 ene a 13 abr 1993 , J. Sihezar, G. Rodriguez, L- N 306300_388600 / INBIOCRI001331532"; 1♀ "Est. Las Pailas, 800 m , P.N. Rincon de la Vieja, Prov. Guan., COSTA RICA . 10 a 27 Mar 1993 , K. Taylor, L- N 306300_388600 / INBIOCRI001355780"; 1♂ "Est. Maritza, 600 m , lado O Vol. Orosi, Prov. Guan. COSTA RICA . R. Blanco, Abr 1990 , L- N 326900_373000 / INBIOCRI000238024"; 1♀ "Los Almendros, P. N. Guanacaste, Prov. Guanacaste, Costa Rica , 28 mar a 24 abr 1992 , M. Reyes, L- N 334800_369800 / CEUA 00105806". Holotype male description. Body shape elongate. Length 14.9 mm . Width 8.4 mm . Color. Head dark brown. Pronotum orange ochre with 1 large irregular dark brown macula on disc, not reaching basal margin, crossed by a lighter weak longitudinal line; small maculae on lateral foveae; surface with green luster. Scutellum light brown with lighter center. Elytra orange ochre. Pygidium ochre. Legs reddish brown; protibiae with darker margins, metafemura ochre, metatibiae and metatarsi dark brown with green luster. Venter, sternum reddish brown with darker margins, abdominal sternites ochre. Clypeus trapezoidal, surface densely punctate-reticulate, flat. Anterior margin straight, clypeus anteriorly thin. Clypeus ratio width/length 2.3. Frons densely punctate, flat. Ocular canthum long, thin, with rounded apex. Interocular ratio (interocular width/width of eye): 2.9. Pronotum subtrapezoidal. Ratio width/length 1.7. Lateral margins angulated at 1/3 of pronotum length. Apical angles acute, sharp; basal angles obtuse, blunt. Basal margin sinuate, subapical bead complete. Surface densely punctate. Scutellum. Shape subtriangular, with rounded sides, apex acute. Ratio width/length 1.5. Surface densely punctate. Elytra with costae defined by regular rows of punctures. Subsutural interstice with 3–5 rows of punctures, 2nd– 4th interstices with 1–2 irregular rows of punctures. Surface covered with secondary punctures. Pygidium finely granulate with long setae at apical margin. Ratio width/length 1.4. Space between the mesocoxae narrow, flat. Mesometasternal suture well defined, at base of mesocoxae. Abdominal sternites with 1 row of sparse setae and 5–6 rows of punctures per sternite. Terminal sternite punctate, with subapical bead narrow and sinuate, apical membrane narrow. Protibia (Fig. 8) bidentate; apical tooth long and curved; basal tooth below internal apex of protibia, triangular in shape, acute. Metatibia stout, slightly narrower subapically. Ratio length/width 2.6. First external carina obliterated; surface densely punctate above second external carina, rugose below. Protarsal claws: internal claw bifurcate, with upper branch of same length and 2/3 the width of the lower branch; inferior margin sinuate. Aedeagus ( Fig. 14 ): parameres with sinuate dorsal and ventral margins, apex acute in a lateral view. Endophallus ( Fig. 20 ) with 1 long sacculus with wide apex, covered with setae medially and with a strip of long and short, thick spines at apex. Ejaculatory duct opening placed on a ventral inflation. Variation. Macula on pronotum with variable extension either fusing with maculae on foveae, forming a starshaped macula or reaching the posterior margin; elytra ochre to red. Female. Similar to male, protibia (Fig. 8) with longer and wider apical tooth; basal tooth above internal apex; internal protarsal claw with curved inferior margin. Body length 14.4–16.1 mm , body width 7.9–9.6 mm . Clypeus w/l: 2.2–2.3. Interocular ratio (interocular width/width of eye): 2.5–3.0. Antenna: ratio funiculus/club 0.6–0.7. Scutellum w/l: 1.2–1.5. Pygidium w/l: 1.4–1.6. Metatibia l/w: 2.5–2.8. Diagnosis. Size large; macula on pronotum irregular, dark; elytra ochre; parameres with sinuate dorsal and ventral margins in lateral view; endophallus with a strip of long and short, thick spines at apex. Anomala jansoni Ohaus, 1897 is a similar species of similar size but with two small maculae on pronotum and parameres with long, narrow, curved apex ( Fig. 29 ) (Monte Rotondo, Costa Rica ; MNHUB). Anomala antica Ohaus, 1897 , A. luciae Blanchard, 1851 , and A. quiche Ohaus, 1897 are similar but smaller species, with one small and pentagonal macula on pronotum ( A. antica ), pronotum with sinuate lateral margins ( A. quiche ), maculae on elytra ( A. luciae and A. quiche ), and differences in the parameres: A. antica has a long thin apex ( Fig. 30 ), A. luciae has lobes on anterior portion of inferior margins ( Fig. 31 ), and A. quiche has wide rounded parameres, and the ventral plate curved at apex in a lateral view ( Fig. 32 ). Etymology. Latinized from the Spanish noun estrella , star, for the star-shaped macula on pronotum. Treated here as a noun in apposition,. Distribution: Guanacaste and Tilarán mountain ranges, Costa Rica , from 280–1520 m ( Fig. 34 ).