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<document id="21A719E941D97E5373CB3B00647AC741" ID-DOI="10.50826/bnmnszool.47.2-65" ID-ISSN="2434-091X" ID-Zenodo-Dep="12759907" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721224081901" checkinUser="felipe" docAuthor="Takeda, Masatsune, Ohtsuchi, Naoya &amp; Komatsu, Hironori" docDate="2021" docId="03CF87EEFF85D64EB3AEFC0AFC4EBE61" docLanguage="en" docName="BullNatlMusNatSciSerA.47.2.65-97.pdf" docOrigin="Bulletin of the National Museum of Nature and Science. Series A, Zoology 47 (2)" docStyle="DocumentStyle:A6FFB80BFE6276686563735C80A89639.2:BullNatlMusNatSciSerA.2021-.journal_article.english.affilindex" docStyleId="A6FFB80BFE6276686563735C80A89639" docStyleName="BullNatlMusNatSciSerA.2021-.journal_article.english.affilindex" docStyleVersion="2" docTitle="Ethusa sexdentata" docType="treatment" docVersion="2" lastPageNumber="80" masterDocId="FFF6FF96FF8AD641B361FFB6FF99BC46" masterDocTitle="Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage" masterLastPageNumber="97" masterPageNumber="65" pageNumber="80" updateTime="1721224661766" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="393A0E0413D6C63A9D2555D4543A2560">Crabs (Crustacea, Decapoda) from the Sea off East and Southeast Asia collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 1. Sulu Sea and Sibutu Passage</mods:title>
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<mods:namePart id="CF0B4D5C356BA9C59F8E36F03E20E8FC">Takeda, Masatsune</mods:namePart>
<mods:affiliation id="8E83A43102F1D1E139A09625E3BDC528">Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan E-mail: takeda @ kahaku. go. jp (MT) / h-komatu @ kahaku. go. jp (HK)</mods:affiliation>
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<mods:namePart id="B33011EADF692B33B50A02E35BDDA5E6">Ohtsuchi, Naoya</mods:namePart>
<mods:affiliation id="3A22FB9D711C9E94C143E4498B843D5A">International Coastal Research Center, Atmosphere and Ocean Research Institute, The University of Tokyo, 1 - 19 - 8 Akahama, Otsuchi, Iwate 028 - 1102, Japan E-mail: ohtsuchi @ g. ecc. u-tokyo. ac. jp</mods:affiliation>
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<mods:namePart id="432936383E5C58D0D7A354C2F5E3FE3D">Komatsu, Hironori</mods:namePart>
<mods:affiliation id="6E22969F678429923E6C872F72E7382B">Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan E-mail: takeda @ kahaku. go. jp (MT) / h-komatu @ kahaku. go. jp (HK)</mods:affiliation>
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<mods:date id="982A74EFB816856B4E8270A815CC0E40">2021</mods:date>
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<taxonomicName id="4C664D7BFF85D64EB3AEFC0AFDC3BF92" ID-CoL="6H4W3" authority="(Stimpson, 1858)" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[207,602,956,981]" class="Malacostraca" family="Ethusidae" genus="Ethusa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="80" phylum="Arthropoda" rank="species" species="sexdentata">
<emphasis id="B912EAEAFF85D64EB3AEFC0AFE03BF92" bold="true" box="[207,410,956,980]" italics="true" pageId="15" pageNumber="80">Ethusa sexdentata</emphasis>
(
<bibRefCitation id="EFF74B09FF85D64EB2C9FC0BFDCBBF92" author="Stimpson, W." box="[424,594,956,981]" pageId="15" pageNumber="80" pagination="159 - 163" refId="ref19550" refString="Stimpson, W. 1858. Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Paci- ficum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars VI. Crustacea Oxystomata. Proceedings of the Academy of Natural Sciences of Philadelphia 1858: 159 - 163." type="book chapter" year="1858">Stimpson, 1858</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C37C6573FF85D64EB3C6FC43FC4EBE61" pageId="15" pageNumber="80" type="materials_examined">
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<emphasis id="B912EAEAFF85D64EB3C6FC43FE08B84B" box="[167,401,1013,1037]" italics="true" pageId="15" pageNumber="80">Material examined.</emphasis>
<materialsCitation id="3B0E3CA5FF85D64EB2CAFC43FEF3B8D1" collectionCode="RV, CB" elevation="498" latitude="8.3116665" location="Sea" longLatPrecision="129" longitude="118.14167" municipality="Hakuho Maru" pageId="15" pageNumber="80" specimenCount="1" stateProvince="Sulu">
<collectionCode id="ED77AE3DFF85D64EB2CAFC43FE49B84A" box="[427,464,1013,1036]" country="Italy" name="Collection of Leptospira Strains" pageId="15" pageNumber="80">RV</collectionCode>
<emphasis id="B912EAEAFF85D64EB28CFC43FD04B84A" box="[493,669,1013,1037]" italics="true" pageId="15" pageNumber="80">
<collectingMunicipality id="6BBDAC82FF85D64EB28CFC43FD04B84A" box="[493,669,1013,1037]" pageId="15" pageNumber="80">Hakuhō Maru</collectingMunicipality>
</emphasis>
KH-72-1 cruise, sta. 12 (
<collectingRegion id="49A2F81AFF85D64EB2CEFBA1FE78B869" box="[431,481,1047,1071]" country="Philippines" name="Sulu" pageId="15" pageNumber="80">Sulu</collectingRegion>
<location id="8EB96023FF85D64EB28EFBA1FD80B869" LSID="urn:lsid:plazi:treatment:03CF87EEFF85D64EB3AEFC0AFC4EBE61:8EB96023FF85D64EB28EFBA1FD80B869" box="[495,537,1047,1071]" latitude="8.3116665" longLatPrecision="129" longitude="118.14167" municipality="Hakuho Maru" name="Sea" pageId="15" pageNumber="80" stateProvince="Sulu">Sea</location>
;
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,
<geoCoordinate id="EE52503FFF85D64EB3EDFB8CFE98B817" box="[140,257,1081,1106]" degrees="118" direction="east" minutes="09.1" orientation="longitude" pageId="15" pageNumber="80" precision="92" value="118.151665">118°09.1E</geoCoordinate>
<geoCoordinate id="EE52503FFF85D64EB241FB8CFE16B817" box="[288,399,1081,1106]" degrees="08" direction="north" minutes="18.7" orientation="latitude" pageId="15" pageNumber="80" precision="92" value="8.3116665">08°18.7N</geoCoordinate>
,
<geoCoordinate id="EE52503FFF85D64EB2FEFB8CFD8EB817" box="[415,535,1081,1106]" degrees="118" direction="east" minutes="08.5" orientation="longitude" pageId="15" pageNumber="80" precision="92" value="118.14167">118°08.5E</geoCoordinate>
;
<quantity id="4C9E9B1DFF85D64EB149FB8CFD04B817" box="[552,669,1082,1106]" metricMagnitude="2" metricUnit="m" metricValue="4.975" metricValueMax="5.0" metricValueMin="4.95" pageId="15" pageNumber="80" unit="m" value="497.5" valueMax="500.0" valueMin="495.0">
<elevation id="004BD1CBFF85D64EB149FB8CFD04B817" box="[552,669,1082,1106]" metricMagnitude="2" metricUnit="m" metricValue="4.975" metricValueMax="5.0" metricValueMin="4.95" pageId="15" pageNumber="80" unit="m" value="497.5" valueMax="500.0" valueMin="495.0">495500 m</elevation>
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deep);
<quantity id="4C9E9B1DFF85D64EB3B9FBEBFF67B832" box="[216,254,1117,1141]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="80" unit="m" value="3.0">
<elevation id="004BD1CBFF85D64EB3B9FBEBFF67B832" box="[216,254,1117,1141]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="80" unit="m" value="3.0">3 m</elevation>
</quantity>
beam trawl; 26 May, 1972; 1 ˁ (
<collectionCode id="ED77AE3DFF85D64EB119FBEBFD04B833" box="[632,669,1117,1141]" country="Australia" name="The CB Rhizobium Collection" pageId="15" pageNumber="80">CB</collectionCode>
<quantity id="4C9E9B1DFF85D64EB3EDFBC9FF47B8D1" box="[140,222,1151,1175]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="15" pageNumber="80" unit="mm" value="5.5">5.5 mm</quantity>
; CL 6.0 mm)
</materialsCitation>
, NSMT-Cr 28970.
</paragraph>
<paragraph id="8BD936F8FF85D64EB3C6FB14FF43BB41" blockId="15.[140,670,1013,1799]" pageId="15" pageNumber="80">
<emphasis id="B912EAEAFF85D64EB3C6FB14FE9DB8FC" box="[167,260,1186,1210]" italics="true" pageId="15" pageNumber="80">Remarks</emphasis>
. The male specimen at hand agrees well with the males recorded by
<bibRefCitation id="EFF74B09FF85D64EB28DFB73FD00B89A" author="Stimpson, W." box="[492,665,1220,1245]" pageId="15" pageNumber="80" pagination="159 - 163" refId="ref19550" refString="Stimpson, W. 1858. Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Paci- ficum Septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers Ducibus, observavit et descripsit. Pars VI. Crustacea Oxystomata. Proceedings of the Academy of Natural Sciences of Philadelphia 1858: 159 - 163." type="book chapter" year="1858">Stimpson (1858</bibRefCitation>
,
<bibRefCitation id="EFF74B09FF85D64EB3EDFB51FF5AB8B9" author="Stimpson, W." box="[140,195,1255,1279]" pageId="15" pageNumber="80" pagination="1 - 240" refId="ref19607" refString="Stimpson, W. 1907. Report on the Crustacea (Brachyura and Anomura) collected by the North Pacific Exploring Expedition, 1853 - 1856. Smithsonian Miscellaneous Collections 49: 1 - 240, pls. 1 - 26." type="journal article" year="1907">1907</bibRefCitation>
) in having the sharp external orbital tooth directed obliquely outward, and differs from the females illustrated by
<bibRefCitation id="EFF74B09FF85D64EB2E1FA9BFD9FB903" author="Sakai, T." box="[384,518,1325,1349]" pageId="15" pageNumber="80" pagination="67 - 192" refId="ref19056" refString="Sakai, T. 1937. Studies on the crabs of Japan. II. Oxystomata. Science Report of Tokyo Bunrika Daigaku (B) 3, Supplement 2: 67 - 192, pls. 10 - 19." type="journal article" year="1937">Sakai (1937</bibRefCitation>
, 1965, 1976), in which the external orbital tooth is stout and not tuberculated. In the present male, four frontal teeth are spiniform, weakly directed obliquely outward, and similar to, but shorter than the external orbital tooth; the tip of the frontal inner tooth slightly exceeds the tip of the frontal outer tooth; the tip of the external orbital tooth does not reach the basal part of the frontal teeth. The lateral margin of the carapace is directed obliquely outward, with the laterally convex branchial margin. The distal part of the whip-like G2 is sticking out from the subterminal part of the G1.
</paragraph>
<paragraph id="8BD936F8FF85D64EB1BCFF6EFC5CBDF9" blockId="15.[707,1236,215,551]" pageId="15" pageNumber="80">
In the revision of the subfamily
<taxonomicName id="4C664D7BFF85D64EB705FF61FCC5BD54" authority=", Castro (2005)" authorityName="Castro" authorityYear="2005" class="Malacostraca" family="Oregoniidae" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="80" phylum="Arthropoda" rank="subFamily" subFamily="Ethusinae">
Ethusinae,
<bibRefCitation id="EFF74B09FF85D64EB1A2FF4CFCC5BD54" author="Castro, P." box="[707,860,250,274]" pageId="15" pageNumber="80" pagination="499 - 600" refId="ref14364" refString="Castro, P. 2005. Crabs of the subfamily Ethusinae Guinot, 1977 (Crustacea, Decapoda, Brachyura, Dorippidae) of the Indo-West Pacific region. Zoosystema 27: 499 - 600." type="journal article" year="2005">Castro (2005)</bibRefCitation>
</taxonomicName>
extensively studied all the Indo-West Pacific species of
<taxonomicName id="4C664D7BFF85D64EB0A9FEABFB8BBD73" authorityName="Roux" authorityYear="1830" box="[968,1042,285,309]" class="Malacostraca" family="Ethusidae" genus="Ethusa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="B912EAEAFF85D64EB0A9FEABFB8BBD73" box="[968,1042,285,309]" italics="true" pageId="15" pageNumber="80">Ethusa</emphasis>
</taxonomicName>
including
<taxonomicName id="4C664D7BFF85D64EB7E8FEABFC8ABD11" baseAuthorityName="Stimpson" baseAuthorityYear="1858" class="Malacostraca" family="Ethusidae" genus="Ethusa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="80" phylum="Arthropoda" rank="species" species="sexdentata">
<emphasis id="B912EAEAFF85D64EB7E8FEABFC8ABD11" italics="true" pageId="15" pageNumber="80">E. sexdentata</emphasis>
</taxonomicName>
. According to the monograph,
<taxonomicName id="4C664D7BFF85D64EB705FEF6FD75BD3C" baseAuthorityName="Stimpson" baseAuthorityYear="1858" class="Malacostraca" family="Ethusidae" genus="Ethusa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="80" phylum="Arthropoda" rank="species" species="sexdentata">
<emphasis id="B912EAEAFF85D64EB705FEF6FD75BD3C" italics="true" pageId="15" pageNumber="80">E. sexdentata</emphasis>
</taxonomicName>
reported by
<bibRefCitation id="EFF74B09FF85D64EB0E1FED4FB93BD3C" author="Chen, H." box="[896,1034,354,378]" pageId="15" pageNumber="80" pagination="179 - 203" refId="ref14588" refString="Chen, H. 1986. Decapod Crustacea: Dorippidae. In Forest, J. (ed.): Resultats des Campagnes MUSORSTOM. I &amp; II. Philippines, Tome 2. Memoires du Museum national d`Histoire naturelle, Paris (A) 133: 179 - 203." type="journal article" year="1986">Chen (1986)</bibRefCitation>
was misidentified and represents a new species, named by
<bibRefCitation id="EFF74B09FF85D64EB7EFFE33FC93BDF9" author="Castro, P." pageId="15" pageNumber="80" pagination="499 - 600" refId="ref14364" refString="Castro, P. 2005. Crabs of the subfamily Ethusinae Guinot, 1977 (Crustacea, Decapoda, Brachyura, Dorippidae) of the Indo-West Pacific region. Zoosystema 27: 499 - 600." type="journal article" year="2005">Castro (2005)</bibRefCitation>
as
<taxonomicName id="4C664D7BFF85D64EB04CFE1EFC27BDF9" authorityName="Castro" authorityYear="2005" box="[813,958,423,447]" class="Malacostraca" family="Ethusidae" genus="Ethusa" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="80" phylum="Arthropoda" rank="species" species="abbreviata">
<emphasis id="B912EAEAFF85D64EB04CFE1EFC27BDF9" box="[813,958,423,447]" italics="true" pageId="15" pageNumber="80">E. abbreviata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD936F8FF85D64EB1BCFE7CFC4EBE61" blockId="15.[707,1236,215,551]" pageId="15" pageNumber="80">
<emphasis id="B912EAEAFF85D64EB1BCFE7CFCC6BDA4" box="[733,863,458,482]" italics="true" pageId="15" pageNumber="80">Distribution</emphasis>
. Known from
<collectingCountry id="F3717668FF85D64EB748FE7CFBF1BDA7" box="[1065,1128,458,481]" name="Japan" pageId="15" pageNumber="80">Japan</collectingCountry>
,
<collectingCountry id="F3717668FF85D64EB71EFE7CFB49BDA4" box="[1151,1232,458,482]" name="Taiwan" pageId="15" pageNumber="80">Taiwan</collectingCountry>
, the East and South
<collectingCountry id="F3717668FF85D64EB0C3FE5BFC7BBE43" box="[930,994,493,517]" name="China" pageId="15" pageNumber="80">China</collectingCountry>
Seas, and the
<collectingCountry id="F3717668FF85D64EB7EAFE5BFD66BE61" name="Philippines" pageId="15" pageNumber="80">Philippines</collectingCountry>
; 30ca.
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deep.
</paragraph>
</subSubSection>
</treatment>
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