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<document id="BA3FF600ABC539B1D2D06AE14F05579B" ID-DOI="10.11646/zootaxa.5540.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14240496" ID-ZooBank="4DBD3A92-F14B-4C5B-95B3-2B430EC197DD" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1732832094216" checkinUser="plazi" docAuthor="Wu, Jingyu &amp; Liu, Xingyue" docDate="2024" docId="03B88790FFAFFFAC9EE7C8109ABA4804" docLanguage="en" docName="zootaxa.5540.1.1.pdf" docOrigin="Zootaxa 5540 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5540.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Ankylopterygini Navas 1910" docType="treatment" docVersion="2" lastPageNumber="10" masterDocId="FF81FFE8FFA7FFA59E70CE029F264D33" masterDocTitle="Systematics of the green lacewing tribe Ankylopterygini Navás, 1910 (Neuroptera: Chrysopidae: Chrysopinae) from China" masterLastPageNumber="169" masterPageNumber="1" pageNumber="9" updateTime="1733249290003" updateUser="carolina">
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<mods:title id="8B22BCCC9B6420BCBF500DBDB0D14D70">Systematics of the green lacewing tribe Ankylopterygini Navás, 1910 (Neuroptera: Chrysopidae: Chrysopinae) from China</mods:title>
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<mods:namePart id="D510FE87118000039F8336308702D9FB">Wu, Jingyu</mods:namePart>
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<mods:affiliation id="966567BD7826F3354F7BB969C7ED103E">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659</mods:affiliation>
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<mods:affiliation id="1F2C7931B906991B81A89B7CDB4BFD15">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659</mods:affiliation>
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<subSubSection id="C30B650DFFAFFFAD9EE7C8109D6A4B1F" box="[151,588,1554,1581]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BAE3686FFAFFFAD9EE7C8109D6A4B1F" blockId="8.[151,588,1554,1581]" box="[151,588,1554,1581]" pageId="8" pageNumber="9">
<heading id="D0E681EAFFAFFFAD9EE7C8109D6A4B1F" bold="true" box="[151,588,1554,1581]" fontSize="11" level="2" pageId="8" pageNumber="9" reason="3">
<emphasis id="B965EA94FFAFFFAD9EE7C8109D6A4B1F" bold="true" box="[151,588,1554,1581]" pageId="8" pageNumber="9">
Tribe
<taxonomicName id="4C114D05FFAFFFAD9E91C8109D6A4B1F" authority="Navas, 1910" authorityName="Navas" authorityYear="1910" box="[225,588,1554,1581]" class="Insecta" family="Chrysopidae" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="tribe" tribe="Ankylopterygini">Ankylopterygini Navás, 1910</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30B650DFFAFFFAD9EE7C8559ED84BBD" pageId="8" pageNumber="9" type="reference_group">
<paragraph id="8BAE3686FFAFFFAD9EE7C8559EDD4B5C" blockId="8.[151,510,1623,1678]" box="[151,507,1623,1647]" pageId="8" pageNumber="9">
Ancilopterygino
<bibRefCitation id="EF804B77FFAFFFAD9F34C8559ED14B5C" author="Navas, L." box="[324,503,1623,1647]" pageId="8" pageNumber="9" pagination="38 - 59" refId="ref90149" refString="Navas, L. (1910 a) Crisopidos (Ins. Neur.) nuevos. Broteria, Zoologica, 9, 38 - 59." type="journal article" year="1910">Navás, 1910a: 59</bibRefCitation>
.
</paragraph>
<paragraph id="8BAE3686FFAFFFAD9EE7C8749ED84BBD" blockId="8.[151,510,1623,1678]" box="[151,510,1654,1678]" pageId="8" pageNumber="9">
<taxonomicName id="4C114D05FFAFFFAD9EE7C8749EA14BBD" authority="Navas" authorityName="Navas" authorityYear="1910" box="[151,391,1654,1678]" class="Insecta" family="Chrysopidae" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="tribe" tribe="Ankylopterygini">Ankylopterygini Navás</taxonomicName>
, 1913: 293.
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFAFFFAC9EE7C8BE9D1B4CBA" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8BAE3686FFAFFFAC9EE7C8BE9D1B4CBA" blockId="8.[151,1437,1724,2039]" lastBlockId="9.[151,1437,151,753]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="B965EA94FFAFFFAD9EE7C8BE9E344BE5" bold="true" box="[151,274,1724,1750]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
Body small to large. Body length 5.621.0 mm, forewing length 7.527.0 mm, hindwing length
<quantity id="4CE99B63FFAFFFAD9B4AC8BE9FE04BC8" metricMagnitude="-2" metricUnit="m" metricValue="1.61" metricValueMax="2.55" metricValueMin="0.67" pageId="8" pageNumber="9" unit="mm" value="16.1" valueMax="25.5" valueMin="6.7">6.725.5 mm</quantity>
. This tribe is characterized by the following characters: the head sometimes with spots on the frons, clypeus and gena, the large prominent compound eyes, the antennal scape sometimes with brown stripe on the outer or inner side, the scape wider than pedicel and flagellum, the sickle-shaped mandibles without basal teeth, the maxillary and labial palpi usually marked, the terminal palpomere pointed or not, the pronotum sometimes with small brown spots anterolaterally, the mesonotum and metanotum with brown markings or not, the legs sometimes marked with black spots on tibia and with long pale setae, the pretarsal claws with basal dilation, the forewing sometimes with brown shadings on gradates, CuP, etc., the costal area strongly or feebly broadened basally, the dark or indistinct pterostigma, the cell
<emphasis id="B965EA94FFAFFFAD9FF2C9DE9E834AC5" box="[386,421,2012,2038]" italics="true" pageId="8" pageNumber="9">m1</emphasis>
smaller than
<emphasis id="B965EA94FFAFFFAD9C4CC9DE9D784AC5" box="[572,606,2012,2038]" italics="true" pageId="8" pageNumber="9">m2</emphasis>
, the present or absent cell
<emphasis id="B965EA94FFAFFFAD9DFAC9DF9C804AC5" box="[906,934,2013,2038]" italics="true" pageId="8" pageNumber="9">im</emphasis>
, the gradates in two or more divergent rows, the basal crossvein of inner gradate series meeting Psm or not, the cell
<emphasis id="B965EA94FFAEFFAC9DDECE959CEF4D82" box="[942,969,151,177]" italics="true" pageId="9" pageNumber="10">c1</emphasis>
smaller than
<emphasis id="B965EA94FFAEFFAC9A2FCE959B5F4D82" box="[1119,1145,151,177]" italics="true" pageId="9" pageNumber="10">c2</emphasis>
, the opened or closed cell
<emphasis id="B965EA94FFAEFFAC9EE7CEB99F984DE6" box="[151,190,187,213]" italics="true" pageId="9" pageNumber="10">dcc</emphasis>
, the hindwing usually without shadings, the abdomen with dense setae, the male tergum IX fused with ectoproct, the male sternum VIII fused with sternum IX or not, the ovoid callus cerci, the male genitalia without tignum and gonapsis, the arcuate gonarcus, the entoprocessus always attached with gonarcus, the pseudopenis not attached with entoprocessus or the arcessus attached to the entoprocessus, the female subgenitale distinctly or slightly bilobed apically, the thick or thin spermatheca, the long or short or absent vela, the deep or shallow or absent ventral impression, and the long curved duct.
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFAEFFAC9EB7CF909B7C4C9E" box="[199,1114,402,429]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BAE3686FFAEFFAC9EB7CF909B7C4C9E" blockId="9.[151,1437,151,753]" box="[199,1114,402,429]" pageId="9" pageNumber="10">
<emphasis id="B965EA94FFAEFFAC9EB7CF909E474C9F" bold="true" box="[199,353,402,428]" pageId="9" pageNumber="10">Distribution.</emphasis>
Palaearctic, Afrotropical, Australian, Oriental and Nearctic regions.
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFAEFFAC9EB7CFB59B444FC2" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BAE3686FFAEFFAC9EB7CFB59B444FC2" blockId="9.[151,1437,151,753]" pageId="9" pageNumber="10">
<emphasis id="B965EA94FFAEFFAC9EB7CFB59E1D4CE2" bold="true" box="[199,315,439,465]" pageId="9" pageNumber="10">Remarks.</emphasis>
In
<bibRefCitation id="EF804B77FFAEFFAC9F16CFB59DB74CE2" author="Brooks, S. J. &amp; Barnard, P. C." box="[358,657,439,465]" pageId="9" pageNumber="10" pagination="117 - 286" refId="ref88564" refString="Brooks, S. J. &amp; Barnard, P. C. (1990) The green lacewings of the world: a generic review (Neuroptera: Chrysopidae). Bulletin of the British Museum (Natural History), Entomology Series, 59, 117 - 286." type="journal article" year="1990">Brooks &amp; Barnard (1990)</bibRefCitation>
, based on the male genitalia being without tignum and gonapsis as well as being pointed terminal palpomere of maxillary and labial palpi, the tribe
<taxonomicName id="4C114D05FFAEFFAC9A4ECFD99BD14CC6" authorityName="Navas" authorityYear="1910" box="[1086,1271,475,501]" class="Insecta" family="Chrysopidae" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="tribe" tribe="Ankylopterygini">Ankylopterygini</taxonomicName>
included only five genera:
<taxonomicName id="4C114D05FFAEFFAC9F5ECFFD9D4E4F2A" authority="Brauer, 1864" authorityName="Brauer" authorityYear="1864" box="[302,616,511,537]" class="Insecta" family="Chrysopidae" genus="Ankylopteryx" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9F5ECFFD9EE44F2A" box="[302,450,511,537]" italics="true" pageId="9" pageNumber="10">Ankylopteryx</emphasis>
<bibRefCitation id="EF804B77FFAEFFAC9FBFCFFD9D4E4F2A" author="Brauer, F." box="[463,616,511,537]" pageId="9" pageNumber="10" pagination="891 - 902" refId="ref88294" refString="Brauer, F. (1864) Entomologische Beitrage. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien, 14, 891 - 902." type="journal article" year="1864">Brauer, 1864</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C114D05FFAEFFAC9C09CFFD9CB04F2A" authority="Brooks, 1986" authorityName="Brooks" authorityYear="1986" box="[633,918,511,537]" class="Insecta" family="Chrysopidae" genus="Retipenna" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9C09CFFD9DCD4F2B" box="[633,747,511,536]" italics="true" pageId="9" pageNumber="10">Retipenna</emphasis>
<bibRefCitation id="EF804B77FFAEFFAC9C87CFFD9CB04F2A" author="Brooks, S. J." box="[759,918,511,537]" pageId="9" pageNumber="10" pagination="35 - 48" refId="ref88537" refString="Brooks, S. J. (1986) A new genus of Ankylopterygini (Chrysopidae). Neuroptera International, 4, 35 - 48." type="journal article" year="1986">Brooks, 1986</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C114D05FFAEFFAC9DD7CFFD9A224F2A" authority="Tjeder, 1966" authorityName="Tjeder" authorityYear="1966" box="[935,1284,511,537]" class="Insecta" family="Chrysopidae" genus="Parankylopteryx" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9DD7CFFD9B444F2A" box="[935,1122,511,537]" italics="true" pageId="9" pageNumber="10">Parankylopteryx</emphasis>
<bibRefCitation id="EF804B77FFAEFFAC9A1ECFFD9A224F2A" author="Tjeder, B." box="[1134,1284,511,537]" pageId="9" pageNumber="10" pagination="228 - 534" refId="ref91317" refString="Tjeder, B. (1966) Neuroptera-Planipennia. The Lace-wings of Southern Africa. 5. Family Chrysopidae. South African Animal Life, 12, 228 - 534." type="journal article" year="1966">Tjeder, 1966</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C114D05FFAEFFAC9B65CFFD9E094F0E" authority="Brooks, 1983" authorityName="Brooks" authorityYear="1983" class="Insecta" family="Chrysopidae" genus="Semachrysa" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9B65CFFD9ABA4F2A" box="[1301,1436,511,537]" italics="true" pageId="9" pageNumber="10">Semachrysa</emphasis>
<bibRefCitation id="EF804B77FFAEFFAC9EE7CC219E094F0E" author="Brooks, S. J." box="[151,303,547,573]" pageId="9" pageNumber="10" pagination="1 - 26" refId="ref88497" refString="Brooks, S. J. (1983) A new genus of oriental lacewings (Neuroptera: Chrysopidae). Bulletin of the British Museum (Natural History), Entomology Series, 47, 1 - 26." type="journal article" year="1983">Brooks, 1983</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C114D05FFAEFFAC9F17CC219C354F0E" authority="Brooks &amp; Barnard, 1990" authorityName="Brooks &amp; Barnard" authorityYear="1990" box="[359,787,547,573]" class="Insecta" family="Chrysopidae" genus="Signochrysa" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9F17CC219ED74F0E" box="[359,497,547,573]" italics="true" pageId="9" pageNumber="10">Signochrysa</emphasis>
<bibRefCitation id="EF804B77FFAEFFAC9F89CC219C354F0E" author="Brooks, S. J. &amp; Barnard, P. C." box="[505,787,547,573]" pageId="9" pageNumber="10" pagination="117 - 286" refId="ref88564" refString="Brooks, S. J. &amp; Barnard, P. C. (1990) The green lacewings of the world: a generic review (Neuroptera: Chrysopidae). Bulletin of the British Museum (Natural History), Entomology Series, 59, 117 - 286." type="journal article" year="1990">Brooks &amp; Barnard, 1990</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="EF804B77FFAEFFAC9D6FCC219B444F0E" author="Garzon-Orduna, I. J. &amp; Winterton, S. L. &amp; Jiang, Y. L. &amp; Breitkreuz, L. C. V. &amp; Duelli, P. &amp; Engel, M. S. &amp; Penny, N. D. &amp; Tauber, C. A. &amp; Mochizuki, A. &amp; Liu, X. Y." box="[799,1122,547,573]" pageId="9" pageNumber="10" pagination="499 - 513" refId="ref88713" refString="Garzon-Orduna, I. J., Winterton, S. L., Jiang, Y. L., Breitkreuz, L. C. V., Duelli, P., Engel, M. S., Penny, N. D., Tauber, C. A., Mochizuki, A. &amp; Liu, X. Y. (2019) Evolution of green lacewings (Neuroptera: Chrysopidae): a molecular supermatrix approach. Systematic Entomology, 44, 499 - 513. https: // doi. org / 10.1111 / syen. 12339" type="journal article" year="2019">
Garzón-Orduña
<emphasis id="B965EA94FFAEFFAC9DA6CC219B294F0E" box="[982,1039,547,573]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
(2019)
</bibRefCitation>
and
<bibRefCitation id="EF804B77FFAEFFAC9AE9CC219ABA4F0E" author="Winterton, S. L. &amp; Gillung, J. P. &amp; Garzon-Orduna, I. J. &amp; Badano, D. &amp; Breitkreuz, L. C. V. &amp; Duelli, P. &amp; Engel, M. S. &amp; Liu, X. &amp; Machado, R. J. P. &amp; Mansell, M. &amp; Mochizuki, A. &amp; Penny, N. D. &amp; Tauber, C. A. &amp; Oswald, J. D." box="[1177,1436,547,573]" pageId="9" pageNumber="10" pagination="514 - 526" refId="ref91432" refString="Winterton, S. L., Gillung, J. P., Garzon-Orduna, I. J., Badano, D., Breitkreuz, L. C. V., Duelli, P., Engel, M. S., Liu, X., Machado, R. J. P., Mansell, M., Mochizuki, A., Penny, N. D., Tauber, C. A. and Oswald, J. D. (2019) Evolution of green lacewings (Neuroptera: Chrysopidae): an anchored phylogenomics approach. Systematic Entomology, 44, 514 - 526. https: // doi. org / 10.1111 / syen. 12347" type="journal article" year="2019">
Winterton
<emphasis id="B965EA94FFAEFFAC9B61CC219A6C4F0E" box="[1297,1354,547,573]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
(2019)
</bibRefCitation>
reconstructed the phylogeny of
<taxonomicName id="4C114D05FFAEFFAC9C79CC459DB14F52" box="[521,663,583,609]" class="Insecta" family="Chrysopidae" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Chrysopidae</taxonomicName>
and the results showed that
<taxonomicName id="4C114D05FFAEFFAC9D9BCC459A654F52" authority="Needham, 1909" authorityName="Needham" authorityYear="1909" box="[1003,1347,583,609]" class="Insecta" family="Chrysopidae" genus="Tumeochrysa" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9D9BCC459BA64F52" box="[1003,1152,583,609]" italics="true" pageId="9" pageNumber="10">Tumeochrysa</emphasis>
<bibRefCitation id="EF804B77FFAEFFAC9AFBCC459A654F52" author="Needham, J. G." box="[1163,1347,583,609]" pageId="9" pageNumber="10" pagination="185 - 210" refId="ref90677" refString="Needham, J. G. (1909) Notes on the Neuroptera in the collection of the Indian Museum. Records of the Indian Museum, Calcutta, 3, 185 - 210. https: // doi. org / 10.26515 / rzsi / v 3 / i 3 / 1909 / 163272" type="journal article" year="1909">Needham, 1909</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C114D05FFAEFFAC9B22CC459E004FB6" authority="Navas, 1912" authorityName="Navas" authorityYear="1912" class="Insecta" family="Chrysopidae" genus="Nineta" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9B22CC459ABB4F53" box="[1362,1437,583,608]" italics="true" pageId="9" pageNumber="10">Nineta</emphasis>
<bibRefCitation id="EF804B77FFAEFFAC9EE7CC699E004FB6" author="Navas, L." box="[151,294,619,645]" pageId="9" pageNumber="10" pagination="98 - 113" refId="ref90273" refString="Navas, L. (1912) Crisopidos y Hemerobidos (Ins. Neur.) nuevos o criticos. Broteria, Zoologica, 10, 98 - 113." type="journal article" year="1912">Navás, 1912</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C114D05FFAEFFAC9F11CC699DA24FB6" authority="Navas, 1910" authorityName="Navas" authorityYear="1910" box="[353,644,619,645]" class="Insecta" family="Chrysopidae" genus="Chrysopidia" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9F11CC699ECD4FB6" box="[353,491,619,645]" italics="true" pageId="9" pageNumber="10">Chrysopidia</emphasis>
Navás, 1910
</taxonomicName>
are closely related to
<taxonomicName id="4C114D05FFAEFFAC9DF6CC699B654FB6" authorityName="Navas" authorityYear="1910" box="[902,1091,619,645]" class="Insecta" family="Chrysopidae" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="tribe" tribe="Ankylopterygini">Ankylopterygini</taxonomicName>
. The species of
<taxonomicName id="4C114D05FFAEFFAC9B70CC699AB34FB6" authorityName="Needham" authorityYear="1909" box="[1280,1429,619,645]" class="Insecta" family="Chrysopidae" genus="Tumeochrysa" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9B70CC699AB34FB6" box="[1280,1429,619,645]" italics="true" pageId="9" pageNumber="10">Tumeochrysa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C114D05FFAEFFAC9EE7CC8D9FC44F9B" authorityName="Navas" authorityYear="1912" box="[151,226,655,680]" class="Insecta" family="Chrysopidae" genus="Nineta" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9EE7CC8D9FC44F9B" box="[151,226,655,680]" italics="true" pageId="9" pageNumber="10">Nineta</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D05FFAEFFAC9F68CC8D9E844F9A" authorityName="Navas" authorityYear="1910" box="[280,418,655,681]" class="Insecta" family="Chrysopidae" genus="Chrysopidia" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9F68CC8D9E844F9A" box="[280,418,655,681]" italics="true" pageId="9" pageNumber="10">Chrysopidia</emphasis>
</taxonomicName>
do not possess a pointed terminal palpomere of the maxillary and labial palpi, but the male genitalia in these species also lack tignum and gonapsis as in typical
<taxonomicName id="4C114D05FFAEFFAC9DCCCCB19B534FFE" authorityName="Navas" authorityYear="1910" box="[956,1141,691,717]" class="Insecta" family="Chrysopidae" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="tribe" tribe="Ankylopterygini">Ankylopterygini</taxonomicName>
species.
<bibRefCitation id="EF804B77FFAEFFAC9A91CCB19FC44FC2" author="Breitkreuz, L. C. V. &amp; Garzon-Orduna, I. J. &amp; Winterton, S. L. &amp; Engel, M. S." pageId="9" pageNumber="10" pagination="1374 - 1395" refId="ref88364" refString="Breitkreuz, L. C. V., Garzon-Orduna, I. J., Winterton, S. L. &amp; Engel, M. S. (2022) Phylogeny of Chrysopidae (Neuroptera), with emphasis on morphological trait evolution. Zoological Journal of the Linnean Society, 194, 1374 - 1395. https: // doi. org / 10.1093 / zoolinnean / zlab 024" type="journal article" year="2022">
Breitkreuz
<emphasis id="B965EA94FFAEFFAC9B11CCB19ABB4FFE" box="[1377,1437,691,717]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
(2022)
</bibRefCitation>
formally transferred
<taxonomicName id="4C114D05FFAEFFAC9FA0CCD59D3D4FC3" authorityName="Navas" authorityYear="1912" box="[464,539,727,752]" class="Insecta" family="Chrysopidae" genus="Nineta" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9FA0CCD59D3D4FC3" box="[464,539,727,752]" italics="true" pageId="9" pageNumber="10">Nineta</emphasis>
</taxonomicName>
,
<taxonomicName id="4C114D05FFAEFFAC9C59CCD59D984FC2" authorityName="Needham" authorityYear="1909" box="[553,702,727,753]" class="Insecta" family="Chrysopidae" genus="Tumeochrysa" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9C59CCD59D984FC2" box="[553,702,727,753]" italics="true" pageId="9" pageNumber="10">Tumeochrysa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D05FFAEFFAC9C85CCD59C594FC2" authorityName="Navas" authorityYear="1910" box="[757,895,727,753]" class="Insecta" family="Chrysopidae" genus="Chrysopidia" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9C85CCD59C594FC2" box="[757,895,727,753]" italics="true" pageId="9" pageNumber="10">Chrysopidia</emphasis>
</taxonomicName>
to
<taxonomicName id="4C114D05FFAEFFAC9DD2CCD59B794FC2" authorityName="Navas" authorityYear="1910" box="[930,1119,727,753]" class="Insecta" family="Chrysopidae" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="tribe" tribe="Ankylopterygini">Ankylopterygini</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFAEFFAC9EE7CD419ABA4804" pageId="9" pageNumber="10" type="key">
<paragraph id="8BAE3686FFAEFFAC9EE7CD419D1E4E6F" blockId="9.[151,568,834,861]" box="[151,568,834,861]" pageId="9" pageNumber="10">
<heading id="D0E681EAFFAEFFAC9EE7CD419D1E4E6F" bold="true" box="[151,568,834,861]" fontSize="11" level="2" pageId="9" pageNumber="10" reason="3">
<emphasis id="B965EA94FFAEFFAC9EE7CD419D1E4E6F" bold="true" box="[151,568,834,861]" pageId="9" pageNumber="10">
Key to genera of
<taxonomicName id="4C114D05FFAEFFAC9F1BCD409D1E4E6F" authorityName="Navas" authorityYear="1910" box="[363,568,834,860]" class="Insecta" family="Chrysopidae" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="tribe" tribe="Ankylopterygini">Ankylopterygini</taxonomicName>
</emphasis>
</heading>
</paragraph>
<key id="FF8061B7FFAEFFAC9EE7CD8A9ABA4804" pageId="9" pageNumber="10">
<paragraph id="8BAE3686FFAEFFAC9EE7CD8A9ABA4E89" blockId="9.[151,1437,904,1336]" pageId="9" pageNumber="10">
<keyLead id="30E095B3FFAEFFAC9EE7CD8A9ABA4E89" pageId="9" pageNumber="10">
<keyStep id="30E52E23FFAEFFAC9EE7CD8A9ABA4E89" pageId="9" pageNumber="10">1. Terminal palpomere of maxillary and labial palpi finely pointed................................................ 2 Terminal palpomere of maxillary and labial palpi flat and obtuse................................................ 3</keyStep>
</keyLead>
</paragraph>
<paragraph id="8BAE3686FFAEFFAC9EE7CDC09ABA4EC6" blockId="9.[151,1437,904,1336]" pageId="9" pageNumber="10">
<keyLead id="30E095B3FFAEFFAC9EE7CDC09ABA4EC6" pageId="9" pageNumber="10">
<keyStep id="30E52E23FFAEFFAC9EE7CDC09ABA4EC6" pageId="9" pageNumber="10">
2. Body small; gradates of forewing and hindwing in two or three divergent rows; meso- and metatibia without spur
<taxonomicName id="4C114D05FFAEFFAC9B56CDC09E304EC5" authority="Body" authorityName="Body" class="Insecta" family="Chrysopidae" genus="Chrysopidia" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9B56CDC09ABB4EE4" box="[1318,1437,962,983]" italics="true" pageId="9" pageNumber="10">Chrysopidia</emphasis>
Body
</taxonomicName>
large; gradates of forewing and hindwing in three to five divergent rows; meso- and metatibia with a spur
<taxonomicName id="4C114D05FFAEFFAC9B6CCDE29ABA4EC6" authorityName="Needham" authorityYear="1909" box="[1308,1436,992,1013]" class="Insecta" family="Chrysopidae" genus="Tumeochrysa" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9B6CCDE29ABA4EC6" box="[1308,1436,992,1013]" italics="true" pageId="9" pageNumber="10">Tumeochrysa</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8BAE3686FFAEFFAC9EE7CDFF9ABA4903" blockId="9.[151,1437,904,1336]" pageId="9" pageNumber="10">
<keyLead id="30E095B3FFAEFFAC9EE7CDFF9ABA4903" pageId="9" pageNumber="10">
<keyStep id="30E52E23FFAEFFAC9EE7CDFF9ABA4903" pageId="9" pageNumber="10">3. Costal area of forewing obviously widened................................................................. 4 Costal area of forewing not obviously widened.............................................................. 5</keyStep>
</keyLead>
</paragraph>
<paragraph id="8BAE3686FFAEFFAC9EE7CA3A9ABA4996" blockId="9.[151,1437,904,1336]" pageId="9" pageNumber="10">
<keyLead id="30E095B3FFAEFFAC9EE7CA3A9ABA4996" pageId="9" pageNumber="10">
<keyStep id="30E52E23FFAEFFAC9EE7CA3A9ABA4996" pageId="9" pageNumber="10">
4. Pterostigma of forewing brown or black; male entoprocessus fused apically; pseudopenis not attached with entoprocessus or gonarcus...................................................................................
<taxonomicName id="4C114D05FFAEFFAC9B6ECA579ABB4959" authorityName="Brauer" authorityYear="1864" box="[1310,1437,1109,1130]" class="Insecta" family="Chrysopidae" genus="Ankylopteryx" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9B6ECA579ABB4959" box="[1310,1437,1109,1130]" italics="true" pageId="9" pageNumber="10">Ankylopteryx</emphasis>
</taxonomicName>
Pterostigma of forewing indistinct; male entoprocessus not fused; arcessus attached with entoprocessus and gonarcus..............................................................................................
<taxonomicName id="4C114D05FFAEFFAC9A8CCA929ABA4996" authorityName="Tjeder" authorityYear="1966" box="[1276,1436,1168,1189]" class="Insecta" family="Chrysopidae" genus="Parankylopteryx" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9A8CCA929ABA4996" box="[1276,1436,1168,1189]" italics="true" pageId="9" pageNumber="10">Parankylopteryx</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8BAE3686FFAEFFAC9EE7CAAF9ABA49CD" blockId="9.[151,1437,904,1336]" pageId="9" pageNumber="10">
<keyLead id="30E095B3FFAEFFAC9EE7CAAF9ABA49CD" pageId="9" pageNumber="10">
<keyStep id="30E52E23FFAEFFAC9EE7CAAF9ABA49CD" pageId="9" pageNumber="10">
5. One black spot present between antennae; mesopleura usually with black spots; male sternum IX elongated and exceeded ectoproct; ectoproct with two clusters of setae......................................................
<taxonomicName id="4C114D05FFAEFFAC9B56CAC89FD949CE" authority="No" authorityName="No" class="Insecta" family="Chrysopidae" genus="Signochrysa" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9B56CAC89ABB49EC" box="[1318,1437,1226,1247]" italics="true" pageId="9" pageNumber="10">Signochrysa</emphasis>
No
</taxonomicName>
interantennal spot; mesopleura without spots; male sternum IX not elongated; setae on ectoproct not concentrated..... 6
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8BAE3686FFAEFFAC9EE7CB079ABA4804" blockId="9.[151,1437,904,1336]" pageId="9" pageNumber="10">
<keyLead id="30E095B3FFAEFFAC9EE7CB079ABA4804" pageId="9" pageNumber="10">
<keyStep id="30E52E23FFAEFFAC9EE7CB079ABA4804" pageId="9" pageNumber="10">
6. Frons usually without spots; arcessus covered with membrane, without dense setae..........................
<taxonomicName id="4C114D05FFAEFFAC9B4BCB049ABA4828" authorityName="Brooks" authorityYear="1986" box="[1339,1436,1286,1307]" class="Insecta" family="Chrysopidae" genus="Retipenna" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9B4BCB049ABA4828" box="[1339,1436,1286,1307]" italics="true" pageId="9" pageNumber="10">Retipenna</emphasis>
</taxonomicName>
Frons always with three dark spots in a row; arcessus not covered and with four setae......................
<taxonomicName id="4C114D05FFAEFFAC9B58CB209ABA4804" authorityName="Brooks" authorityYear="1983" box="[1320,1436,1314,1335]" class="Insecta" family="Chrysopidae" genus="Semachrysa" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EA94FFAEFFAC9B58CB209ABA4804" box="[1320,1436,1314,1335]" italics="true" pageId="9" pageNumber="10">Semachrysa</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
</key>
</subSubSection>
</treatment>
</document>