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<document id="F5A154B23FDDB3B1F19573D2FC6AA98C" ID-DOI="10.11646/phytotaxa.375.4.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13728284" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725680695066" checkinUser="felipe" docAuthor="Tagane, Shuichiro, Dang, Van-Son, Souladeth, Phetlasy, Nagamasu, Hidetoshi, Toyama, Hironori, Naiki, Akiyo, Fuse, Kengo, Tran, Hop, Yang, Ceng-Jui, Prajaksood, Amornrat &amp; Yahara, Tetsukazu" docDate="2018" docId="039D87D0FF908D1AFF45CF25DB8BF89A" docLanguage="en" docName="phytotaxa.375.4.1.pdf" docOrigin="Phytotaxa 375 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.375.4.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Syzygium scabrum Tagane, Soulad. &amp; Yahara. A 2018, sp. nov." docType="treatment" docVersion="2" lastPageNumber="255" masterDocId="FFA4FFA8FF968D12FFCDC80ADB71FFCF" masterDocTitle="Five new species of Syzygium (Myrtaceae) from Indochina and Thailand" masterLastPageNumber="260" masterPageNumber="247" pageNumber="253" updateTime="1725692588566" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3400AE2AEE60114EC3C5F5AAE9AFEED8">Five new species of Syzygium (Myrtaceae) from Indochina and Thailand</mods:title>
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<mods:namePart id="985B0C524A4C7A35A110AD8979BE15AB">Tagane, Shuichiro</mods:namePart>
<mods:affiliation id="608FE925A74B5B04EA69FAD848429AC1">Center for Asian Conservation Ecology, Kyushu University, 744 Motooka, Fukuoka, 819 - 0395, Japan. &amp; The Kagoshima University Museum, Kagoshima University, 1 - 21 - 30, Korimoto, Kagoshima, 890 - 0065, Japan.</mods:affiliation>
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<mods:namePart id="1AC8640DFBADE4B0154C19291892DE72">Dang, Van-Son</mods:namePart>
<mods:affiliation id="78CD8B728F3CD4E569AD8A8DDD516FC6">The VNM Herbarium, Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85 Tran Quoc Toan Street, District 3, Hochi Minh City, Vietnam.</mods:affiliation>
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<mods:namePart id="7C93EDBB72FDB4840847524F80BD218D">Souladeth, Phetlasy</mods:namePart>
<mods:affiliation id="E66C19EFC0B89B56BBF4F69E26E4B1C8">Faculty of Forest Science, National University of Laos, Dongdok campus, P. O. Box 7322, Vientiane, Lao PDR.</mods:affiliation>
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<mods:namePart id="C25E6A47E24DF08F5BC3C7DA6003897D">Nagamasu, Hidetoshi</mods:namePart>
<mods:affiliation id="124B32855DEE01339F3643AE318BD92A">The Kyoto University Museum, Kyoto University, Yoshida Honmachi, Sakyo-ku, Kyoto, 606 - 8501, Japan.</mods:affiliation>
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<mods:namePart id="C746573A8D7D5E2EB18C842721E72857">Toyama, Hironori</mods:namePart>
<mods:affiliation id="EE6774524E85685964491EDE8A028C21">Center for Asian Conservation Ecology, Kyushu University, 744 Motooka, Fukuoka, 819 - 0395, Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan.</mods:affiliation>
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<mods:namePart id="653572A25F47D2F9140F906EEE9FE8C7">Naiki, Akiyo</mods:namePart>
<mods:affiliation id="9A8B9A0EAADB87C9E62090765605A891">Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan.</mods:affiliation>
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<mods:namePart id="4FBDC16B3CB5647193413FB5EAB11E82">Fuse, Kengo</mods:namePart>
<mods:affiliation id="FA1DE71B43A794FCCB15FEF8C7B94C2F">Center for Asian Conservation Ecology, Kyushu University, 744 Motooka, Fukuoka, 819 - 0395, Japan.</mods:affiliation>
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<mods:namePart id="ACDEC97643096456FCCA95D977BA2CF0">Tran, Hop</mods:namePart>
<mods:affiliation id="97B0B0AE451CB1505C681340738F27F7">University of Science Ho Chi Minh City, 227 Nguyen Van Cu Street, District 5, Ho Chi Minh City, Vietnam.</mods:affiliation>
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<mods:namePart id="1D46FA07CC2DA379A0B59C4EAB343A01">Yang, Ceng-Jui</mods:namePart>
<mods:affiliation id="6118FE076D02AFB4127213BFCCA31330">Institute of Ecology and Evolutionary Biology, National Taiwan University, Taipei, Taiwan.</mods:affiliation>
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<mods:namePart id="33F80E56AE27C832061E1BA00C372A84">Prajaksood, Amornrat</mods:namePart>
<mods:affiliation id="9D38F3ADAB68C1EDEE4A59E72FD6C17C">Department of Biology, Khon Kaen University, Khon Kaen 40002, Thailand.</mods:affiliation>
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<mods:namePart id="535B8F59AA20A0D98D90DDCCED8DB475">Yahara, Tetsukazu</mods:namePart>
<mods:affiliation id="FAD1FF104DE0E018DBCE9A467242B6C2">Center for Asian Conservation Ecology, Kyushu University, 744 Motooka, Fukuoka, 819 - 0395, Japan.</mods:affiliation>
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<mods:date id="F0C6C9F33A85636067B8FC9034B1190A">2018</mods:date>
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<taxonomicName id="4C344D45FF908D14FF45CF25D9FEF886" ID-CoL="546SH" authority="Tagane, Soulad. &amp; Yahara" authorityName="Tagane, Soulad. &amp; Yahara. A" authorityYear="2018" box="[136,655,1838,1865]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="scabrum" status="sp. nov.">
<emphasis id="B940EAD4FF908D14FF45CF25DA13F886" bold="true" box="[136,354,1838,1865]" italics="true" pageId="6" pageNumber="253">Syzygium scabrum</emphasis>
Tagane, Soulad. &amp; Yahara
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,
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</emphasis>
(
<figureCitation id="130F2A43FF908D14FD30CF3AD82EF886" box="[765,863,1840,1865]" captionStart="FIGURE 5" captionStartId="7.[136,229,1938,1959]" captionTargetBox="[151,1436,190,1913]" captionTargetId="figure-14@7.[151,1436,190,1913]" captionTargetPageId="7" captionText="FIGURE 5. Syzygium scabrum Tagane, Soulad. &amp; Yahara. A) Leafy twig, B) Portion of lower leaf surface, C) Inflorescence, D) Flowers and flower buds, E) Young fruit, F) Fruit and seed. Photographs:A &amp; B for Yahara et al. L1727, taken by S. Tagane on Dec. 2017; C &amp; D for Souladeth 86, taken by P. Sutthisaksopon on 24 May 2011; E &amp; F for Phonsena et al. 7280 taken by P. Phonsena on 22 Nov. 2015." figureDoi="http://doi.org/10.5281/zenodo.13728294" httpUri="https://zenodo.org/record/13728294/files/figure.png" pageId="6" pageNumber="253">Figure 5</figureCitation>
).
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TYPE:
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<collectingCountry id="F3237656FF908D14FF3ECF5EDAF6F8A2" box="[243,391,1876,1901]" name="Thailand" pageId="6" pageNumber="253">THAILAND</collectingCountry>
. Buengkan Province, Bung Khla, Phu Wua Wildlife Sanctuary,
<date id="FF8A1006FF908D14FB9FCF5EDF9CF8A2" box="[1106,1261,1876,1901]" pageId="6" pageNumber="253" value="2015-11-22">22 Nov. 2015</date>
,
<emphasis id="B940EAD4FF908D14FB35CF5ED957F85E" italics="true" pageId="6" pageNumber="253">Phonsena P., de Wilde W.J.J.O &amp; Duyfjes B.E.E 7280</emphasis>
[fl. &amp; fr.] (
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BKF!,
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<collectionCode id="ED25AE03FF908D14FC73CF72D899F85E" box="[958,1000,1912,1937]" country="Thailand" lsid="urn:lsid:biocol.org:col:12928" name="Department of Agriculture" pageId="6" pageNumber="253" type="Herbarium">BK</collectionCode>
, KKU).
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<taxonomicName id="4C344D45FF908D14FF70CF96DAFEF87A" ID-CoL="546SH" authorityName="Tagane, Soulad. &amp; Yahara. A" authorityYear="2018" box="[189,399,1948,1973]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="scabrum">
<emphasis id="B940EAD4FF908D14FF70CF96DAFEF87A" box="[189,399,1948,1973]" italics="true" pageId="6" pageNumber="253">Syzygium scabrum</emphasis>
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is similar to
<taxonomicName id="4C344D45FF908D14FDE8CF96D892F87A" ID-CoL="546ZQ" authority="Merrill &amp; Perry (1938: 110)" authorityName="Merrill &amp; Perry" authorityPageNumber="110" authorityYear="1938" box="[549,995,1948,1973]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="vestitum">
<emphasis id="B940EAD4FF908D14FDE8CF96D9EFF87A" box="[549,670,1948,1973]" italics="true" pageId="6" pageNumber="253">S. vestitum</emphasis>
<bibRefCitation id="EFA54B37FF908D14FD6ACF96D892F87A" author="Merrill, E. D. &amp; Perry, L. M." box="[679,995,1948,1973]" pageId="6" pageNumber="253" pagination="99 - 116" refId="ref7876" refString="Merrill, E. D. &amp; Perry, L. M. (1938) On the Indo-Chinese species of Syzygium Gaertner. Journal of Arnold Arboretum 19: 99 - 116." type="journal article" year="1938">Merrill &amp; Perry (1938: 110)</bibRefCitation>
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in having reddish brown hairs on twigs, leaves and hypanthium, but differs in having more or less cordate leaf base (vs. broadly cuneate to rounded in
<taxonomicName id="4C344D45FF908D14FA5BCFCADB94F832" ID-CoL="546ZQ" authorityName="Merrill &amp; Perry" authorityYear="1938" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="vestitum">
<emphasis id="B940EAD4FF908D14FA5BCFCADB94F832" italics="true" pageId="6" pageNumber="253">S. vestitum</emphasis>
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), scabrid on both sides of leaf surfaces (vs. glabrous except on veins on abaxial side, never scabrid on adaxial side), more secondary veins (1630 pairs vs. 1016 pair) and longer styles (
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long vs.
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long).
</paragraph>
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<emphasis id="B940EAD4FF918D15FF45CF98DB89F868" bold="true" box="[136,248,1938,1959]" pageId="7" pageNumber="254">FIGURE 5</emphasis>
.
<taxonomicName id="4C344D45FF918D15FEC9CF98D9A5F868" authority="Tagane, Soulad. &amp; Yahara. A" authorityName="Tagane, Soulad. &amp; Yahara. A" authorityYear="2018" box="[260,724,1938,1959]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="254" phylum="Tracheophyta" rank="species" species="scabrum">
<emphasis id="B940EAD4FF918D15FEC9CF98DAC6F868" box="[260,439,1938,1959]" italics="true" pageId="7" pageNumber="254">Syzygium scabrum</emphasis>
Tagane, Soulad. &amp; Yahara. A
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) Leafy twig, B) Portion of lower leaf surface, C) Inflorescence, D) Flowers and flower buds, E) Young fruit, F) Fruit and seed. Photographs: A &amp; B for
<emphasis id="B940EAD4FF918D15FC96CFBCDF69F804" box="[859,1048,1974,1995]" italics="true" pageId="7" pageNumber="254">Yahara et al. L1727</emphasis>
, taken by S. Tagane on Dec. 2017; C &amp; D for
<emphasis id="B940EAD4FF918D15FF67CFD0DA59F820" box="[170,296,2010,2031]" italics="true" pageId="7" pageNumber="254">Souladeth 86</emphasis>
, taken by P. Sutthisaksopon on 24 May 2011; E &amp; F for
<emphasis id="B940EAD4FF918D15FC85CFD0DF63F820" box="[840,1042,2010,2031]" italics="true" pageId="7" pageNumber="254">Phonsena et al. 7280</emphasis>
taken by P. Phonsena on 22 Nov. 2015.
</paragraph>
</caption>
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Tree,
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tall. Twigs terete,
<quantity id="4CCC9B23FF9E8D1AFDDEC8AAD9F6FF76" box="[531,647,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="8" pageNumber="255" unit="mm" value="3.25" valueMax="3.5" valueMin="3.0">33.5 mm</quantity>
in diam., densely covered with yellowish brown, dark grayish brown, reddish brown hairs, internode
<quantity id="4CCC9B23FF9E8D1AFE2DC8CED92DFF12" box="[480,604,196,221]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="6.5" metricValueMin="1.9" pageId="8" pageNumber="255" unit="cm" value="4.2" valueMax="6.5" valueMin="1.9">1.96.5 cm</quantity>
long. Leaves opposite, blade ovate-elliptic, ovate-oblong, (6.3)11.519.5 × (2.2)
<quantity id="4CCC9B23FF9E8D1AFF06C8E2DA3DFECE" box="[203,332,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="4.35" metricValueMax="5.6" metricValueMin="3.1" pageId="8" pageNumber="255" unit="cm" value="4.35" valueMax="5.6" valueMin="3.1">3.15.6 cm</quantity>
, 2.73.8 times longer than wide, chartaceous, apex acuminate, acumen to
<quantity id="4CCC9B23FF9E8D1AFB41C8E2DFA9FECE" box="[1164,1240,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="255" unit="cm" value="1.2">1.2 cm</quantity>
long, base slightly cordate or rounded, margin entire, recurved, dark dull green to dark yellow-green abaxially, dull brownish yellow to brown abaxially, coarsely reddish brown hairy and scabrid on both surfaces, especially denser on midrib and secondary veins, midrib sunken adaxially, prominent abaxially, secondary veins (16)2126(30) pairs, (2)
<quantity id="4CCC9B23FF9E8D1AFB75C95EDE65FEA2" box="[1208,1300,340,365]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="8" pageNumber="255" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">47 mm</quantity>
apart, sunken adaxially, prominent abaxially, intersecondary veins more or less distinct, prominent abaxially, intramarginal veins 1 or 2, (0.6)
<quantity id="4CCC9B23FF9E8D1AFF27C996DA34FE7A" box="[234,325,412,437]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="8" pageNumber="255" unit="mm" value="2.5" valueMax="4.0" valueMin="1.0">14 mm</quantity>
from margin, prominent abaxially, outer ones (0.1)
<quantity id="4CCC9B23FF9E8D1AFC5FC996DF75FE7A" box="[914,1028,412,437]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="8" pageNumber="255" unit="mm" value="0.8" valueMax="1.0" valueMin="0.6">0.61 mm</quantity>
from margin, prominent to indistinct; petiole (2)
<quantity id="4CCC9B23FF9E8D1AFEC7C9CADA0EFE16" box="[266,383,448,473]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="8" pageNumber="255" unit="mm" value="3.75" valueMax="4.5" valueMin="3.0">34.5 mm</quantity>
long, densely scabrid as in twig. Inflorescences terminal, paniculate,
<quantity id="4CCC9B23FF9E8D1AFB57C9CADF8DFE16" box="[1178,1276,448,473]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="8" pageNumber="255" unit="cm" value="10.5" valueMax="15.0" valueMin="6.0">615 cm</quantity>
long, peduncle
<quantity id="4CCC9B23FF9E8D1AFF45C9EEDBAAFE32" box="[136,219,484,509]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="8" pageNumber="255" unit="cm" value="5.0" valueMax="8.0" valueMin="2.0">28 cm</quantity>
long, densely scabrid with reddish brown hairs. Flower buds ca.
<quantity id="4CCC9B23FF9E8D1AFC7EC9EEDF78FE32" box="[947,1033,484,509]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="255" unit="mm" value="3.5">3.5 mm</quantity>
in diam. Hypanthium funnel-shaped,
<quantity id="4CCC9B23FF9E8D1AFF45CA02DB94FDEE" box="[136,229,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="8" pageNumber="255" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
long, densely covered with brown hairs, pseudostipe
<quantity id="4CCC9B23FF9E8D1AFCF2CA02D8EDFDEE" box="[831,924,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="8" pageNumber="255" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long, bracteoles linear, ca.
<quantity id="4CCC9B23FF9E8D1AFB02CA02DE61FDEE" box="[1231,1296,520,545]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="255" unit="mm" value="4.0">4 mm</quantity>
long, densely brown hairy on both surfaces. Sepals 4, semi-circular, ca.
<quantity id="4CCC9B23FF9E8D1AFCE1CA26D8F7FD8A" box="[812,902,556,581]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="255" unit="mm" value="1.1">1.1 mm</quantity>
long, petals 4, orbicular,
<quantity id="4CCC9B23FF9E8D1AFB7FCA26DE4DFD8A" box="[1202,1340,556,581]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="8" pageNumber="255" unit="mm" value="3.15" valueMax="3.5" valueMin="2.8">2.83.5 mm</quantity>
in diam., membranous, with 3050 gland dots per petal. Stamens many, to
<quantity id="4CCC9B23FF9E8D1AFCB4CA5AD8BAFDA6" box="[889,971,592,617]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="255" unit="mm" value="12.0">12 mm</quantity>
long, anthers
<quantity id="4CCC9B23FF9E8D1AFBA2CA5ADF88FDA6" box="[1135,1273,592,617]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="8" pageNumber="255" unit="mm" value="0.3" valueMax="0.4" valueMin="0.2">0.20.4 mm</quantity>
long, filaments
<quantity id="4CCC9B23FF9E8D1AFF45CA7EDA6DFD42" box="[136,284,628,653]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" metricValueMax="11.6" metricValueMin="0.8" pageId="8" pageNumber="255" unit="mm" value="6.2" valueMax="11.6" valueMin="0.8">0.811.6 mm</quantity>
long, white (
<emphasis id="B940EAD4FF9E8D1AFE7FCA7EDA8FFD42" box="[434,510,628,653]" italics="true" pageId="8" pageNumber="255">in vivo</emphasis>
), glabrous. Style (8)
<quantity id="4CCC9B23FF9E8D1AFD3CCA7ED8F0FD42" box="[753,897,628,653]" metricMagnitude="-2" metricUnit="m" metricValue="1.425" metricValueMax="1.55" metricValueMin="1.3" pageId="8" pageNumber="255" unit="mm" value="14.25" valueMax="15.5" valueMin="13.0">1315.5 mm</quantity>
long, glabrous. Ovary 2-locular, 510 per locule. Fruits (sub)globose, ca.
<quantity id="4CCC9B23FF9E8D1AFE58CA92DA90FD7E" box="[405,481,664,689]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="255" unit="cm" value="2.2">2.2 cm</quantity>
in diam., hairy when young, later glabrous except near calyx ring, calyx ring ca.
<quantity id="4CCC9B23FF9E8D1AFAA6CA92DEDDFD7E" box="[1387,1452,664,689]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="255" unit="mm" value="4.0">4 mm</quantity>
in diam., rim ca.
<quantity id="4CCC9B23FF9E8D1AFE8ACAB6DAECFD1A" box="[327,413,700,725]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="255" unit="mm" value="0.5">0.5 mm</quantity>
high. Seed 1, subglobose,
<quantity id="4CCC9B23FF9E8D1AFD04CAB6D864FD1A" box="[713,789,700,725]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="255" unit="cm" value="1.7">1.7 cm</quantity>
high,
<quantity id="4CCC9B23FF9E8D1AFC91CAB6D8D9FD1A" box="[860,936,700,725]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="255" unit="cm" value="1.5">1.5 cm</quantity>
in diam., dark brown, glabrous.
</paragraph>
</subSubSection>
<subSubSection id="C32E654DFF9E8D1AFF70CAEADB93FBD6" pageId="8" pageNumber="255" type="description">
<paragraph id="8B8B36C6FF9E8D1AFF70CAEADB93FBD6" blockId="8.[136,1452,160,1877]" pageId="8" pageNumber="255">
<materialsCitation id="3B5C3C9BFF9E8D1AFF70CAEADE17FCD2" collectingDate="2011-05-22" collectorName="Souladeth P." country="Laos" elevation="769" latitude="18.354395" location="Ban Tha Heua" longLatPrecision="1" longitude="102.80205" municipality="Thoulakhom District" pageId="8" pageNumber="255" specimenCount="1" stateProvince="Viangchan" typeStatus="paratype">
<emphasis id="B940EAD4FF9E8D1AFF70CAEADA2BFD36" bold="true" box="[189,346,736,761]" pageId="8" pageNumber="255">
<typeStatus id="548F8864FF9E8D1AFF70CAEADA49FD36" box="[189,312,736,761]" pageId="8" pageNumber="255" type="paratype">Paratypes</typeStatus>
:—
</emphasis>
<collectingCountry id="F3237656FF9E8D1AFE97CAEADAD6FD36" box="[346,423,736,761]" name="Laos" pageId="8" pageNumber="255">LAOS</collectingCountry>
.
<collectingRegion id="49F0F824FF9E8D1AFE78CAEAD9E2FD36" box="[437,659,736,761]" country="Laos" name="Viangchan" pageId="8" pageNumber="255">Vientiane Province</collectingRegion>
:
<collectingMunicipality id="6BEFACBCFF9E8D1AFD6CCAEAD8E3FD36" box="[673,914,736,761]" pageId="8" pageNumber="255">Thoulakhom District</collectingMunicipality>
, Phou Khao Khouay National Protected Area,
<location id="8EEB601DFF9E8D1AFF45CB0EDA5CFCD2" LSID="urn:lsid:plazi:treatment:039D87D0FF908D1AFF45CF25DB8BF89A:8EEB601DFF9E8D1AFF45CB0EDA5CFCD2" box="[136,301,772,797]" country="Laos" latitude="18.354395" longLatPrecision="1" longitude="102.80205" municipality="Thoulakhom District" name="Ban Tha Heua" pageId="8" pageNumber="255" stateProvince="Viangchan">Ban Tha Heua</location>
,
<geoCoordinate id="EE005001FF9E8D1AFEF5CB0EDA92FCD2" box="[312,483,772,797]" degrees="18" direction="north" minutes="21" orientation="latitude" pageId="8" pageNumber="255" precision="1" seconds="15.82" value="18.354395">18°2115.82″N</geoCoordinate>
,
<geoCoordinate id="EE005001FF9E8D1AFE23CB0ED9D2FCD2" box="[494,675,772,797]" degrees="102" direction="east" minutes="48" orientation="longitude" pageId="8" pageNumber="255" precision="1" seconds="07.35" value="102.80205">102°4807.35″E</geoCoordinate>
,
<quantity id="4CCC9B23FF9E8D1AFD7DCB0ED986FCD2" box="[688,759,772,797]" metricMagnitude="2" metricUnit="m" metricValue="7.69" pageId="8" pageNumber="255" unit="m" value="769.0">
<elevation id="0019D1F5FF9E8D1AFD7DCB0ED986FCD2" box="[688,759,772,797]" metricMagnitude="2" metricUnit="m" metricValue="7.69" pageId="8" pageNumber="255" unit="m" value="769.0">769 m</elevation>
</quantity>
elev.,
<date id="FF8A1006FF9E8D1AFC8DCB0ED8A9FCD2" box="[832,984,772,797]" pageId="8" pageNumber="255" value="2011-05-22">
<collectingDate id="EFCEE9EEFF9E8D1AFC8DCB0ED8A9FCD2" box="[832,984,772,797]" pageId="8" pageNumber="255" value="2011-05-22">22 May 2011</collectingDate>
</date>
,
<emphasis id="B940EAD4FF9E8D1AFC2ECB0EDFE3FCD2" box="[995,1170,772,797]" italics="true" pageId="8" pageNumber="255">
<collectorName id="26C15310FF9E8D1AFC2ECB0EDF1EFCD2" box="[995,1135,772,797]" pageId="8" pageNumber="255">Souladeth P.</collectorName>
91
</emphasis>
[fl.] (FOF, KKU!)
</materialsCitation>
;
<materialsCitation id="3B5C3C9BFF9E8D1AFABFCB0EDACFFCAA" collectingDate="2017-12-26" collectorName="Yahara T. &amp; Tagane S. &amp; Souladeth P. &amp; Nagamasu &amp; Naiki &amp; Chayer &amp; Kongxaisavath" country="Laos" elevation="769" latitude="18.354395" location="Ban Tha Heua" longLatPrecision="1" longitude="102.80205" municipality="Thoulakhom District" pageId="8" pageNumber="255" specimenCount="1" stateProvince="Viangchan" typeStatus="paratype">
ibid.,
<date id="FF8A1006FF9E8D1AFF45CB22DA50FC8E" box="[136,289,808,833]" pageId="8" pageNumber="255" value="2017-12-26">
<collectingDate id="EFCEE9EEFF9E8D1AFF45CB22DA50FC8E" box="[136,289,808,833]" pageId="8" pageNumber="255" value="2017-12-26">26 Dec. 2017</collectingDate>
</date>
,
<emphasis id="B940EAD4FF9E8D1AFEE0CB22DE08FC8E" box="[301,1401,808,833]" italics="true" pageId="8" pageNumber="255">
<collectorName id="26C15310FF9E8D1AFEE0CB22DAEAFC8E" box="[301,411,808,833]" pageId="8" pageNumber="255">Yahara T.</collectorName>
,
<collectorName id="26C15310FF9E8D1AFE6BCB22D964FC8E" box="[422,533,808,833]" pageId="8" pageNumber="255">Tagane S.</collectorName>
,
<collectorName id="26C15310FF9E8D1AFDEFCB22D9C0FC8E" box="[546,689,808,833]" pageId="8" pageNumber="255">Souladeth P.</collectorName>
,
<collectorName id="26C15310FF9E8D1AFD71CB22D845FC8E" box="[700,820,808,833]" pageId="8" pageNumber="255">Nagamasu</collectorName>
H.,
<collectorName id="26C15310FF9E8D1AFCA9CB22D8D0FC8E" box="[868,929,808,833]" pageId="8" pageNumber="255">Naiki</collectorName>
A.,
<collectorName id="26C15310FF9E8D1AFC03CB22DF51FC8E" box="[974,1056,808,833]" pageId="8" pageNumber="255">Chayer</collectorName>
S. &amp;
<collectorName id="26C15310FF9E8D1AFBACCB22DE78FC8E" box="[1121,1289,808,833]" pageId="8" pageNumber="255">Kongxaisavath</collectorName>
D. L1438
</emphasis>
[fr.] (BKF, FOF!, KYO!, NHL)
</materialsCitation>
;
<materialsCitation id="3B5C3C9BFF9E8D1AFE07CB46D892FC46" collectingDate="2017-12-26" collectorName="Yahara T. &amp; Tagane S. &amp; Souladeth &amp; Nagamasu &amp; Naiki &amp; Chayer &amp; Kongxaisavath D. L" country="Laos" elevation="695" latitude="18.388634" location="Ban Tha Heua" longLatPrecision="1" longitude="102.911285" municipality="Thoulakhom District" pageId="8" pageNumber="255" specimenCount="1" stateProvince="Viangchan" typeStatus="paratype">
ibid.,
<geoCoordinate id="EE005001FF9E8D1AFDC0CB46D9C9FCAA" box="[525,696,844,869]" degrees="18" direction="north" minutes="23" orientation="latitude" pageId="8" pageNumber="255" precision="1" seconds="19.08" value="18.388634">18°2319.08″N</geoCoordinate>
,
<geoCoordinate id="EE005001FF9E8D1AFD08CB46D80BFCAA" box="[709,890,844,869]" degrees="102" direction="east" minutes="54" orientation="longitude" pageId="8" pageNumber="255" precision="1" seconds="40.61" value="102.911285">102°5440.61″E</geoCoordinate>
, elev.
<quantity id="4CCC9B23FF9E8D1AFC09CB46DF7FFCAA" box="[964,1038,844,869]" metricMagnitude="2" metricUnit="m" metricValue="6.95" pageId="8" pageNumber="255" unit="m" value="695.0">
<elevation id="0019D1F5FF9E8D1AFC09CB46DF7FFCAA" box="[964,1038,844,869]" metricMagnitude="2" metricUnit="m" metricValue="6.95" pageId="8" pageNumber="255" unit="m" value="695.0">695 m</elevation>
</quantity>
,
<emphasis id="B940EAD4FF9E8D1AFBD1CB46D833FC46" italics="true" pageId="8" pageNumber="255">
<collectorName id="26C15310FF9E8D1AFBD1CB46DFFAFCAA" box="[1052,1163,844,869]" pageId="8" pageNumber="255">Yahara T.</collectorName>
,
<collectorName id="26C15310FF9E8D1AFB55CB46DE78FCAA" box="[1176,1289,844,869]" pageId="8" pageNumber="255">Tagane S.</collectorName>
,
<collectorName id="26C15310FF9E8D1AFADACB46DEF6FCAA" box="[1303,1415,844,869]" pageId="8" pageNumber="255">Souladeth</collectorName>
P.,
<collectorName id="26C15310FF9E8D1AFF45CB7ADA71FC46" box="[136,256,880,905]" pageId="8" pageNumber="255">Nagamasu</collectorName>
H.,
<collectorName id="26C15310FF9E8D1AFEE2CB7ADA1DFC46" box="[303,364,880,905]" pageId="8" pageNumber="255">Naiki</collectorName>
A.,
<collectorName id="26C15310FF9E8D1AFE55CB7ADA9BFC46" box="[408,490,880,905]" pageId="8" pageNumber="255">Chayer</collectorName>
S. &amp;
<collectorName id="26C15310FF9E8D1AFDE4CB7AD87BFC46" box="[553,778,880,905]" pageId="8" pageNumber="255">Kongxaisavath D. L</collectorName>
1727
</emphasis>
(FOF, KYO!)
</materialsCitation>
;
<materialsCitation id="3B5C3C9BFF9E8D1AFC23CB7ADA59FC62" collectingDate="2011-04-23" collectionCode="KKU" collectorName="Souladeth P." country="Laos" elevation="695" latitude="18.388634" location="Ban Tha Heua" longLatPrecision="1" longitude="102.911285" municipality="Thoulakhom District" pageId="8" pageNumber="255" specimenCount="1" stateProvince="Viangchan" typeStatus="paratype">
ibid.,
<date id="FF8A1006FF9E8D1AFBE2CB7ADFB0FC46" box="[1071,1217,880,905]" pageId="8" pageNumber="255" value="2011-04-23">
<collectingDate id="EFCEE9EEFF9E8D1AFBE2CB7ADFB0FC46" box="[1071,1217,880,905]" pageId="8" pageNumber="255" value="2011-04-23">23 Apr. 2011</collectingDate>
</date>
,
<emphasis id="B940EAD4FF9E8D1AFB01CB7ADE0BFC46" box="[1228,1402,880,905]" italics="true" pageId="8" pageNumber="255">
<collectorName id="26C15310FF9E8D1AFB01CB7ADE29FC46" box="[1228,1368,880,905]" pageId="8" pageNumber="255">Souladeth P.</collectorName>
86
</emphasis>
[fr.] (FOF!,
<collectionCode id="ED25AE03FF9E8D1AFF12CB9EDA51FC62" box="[223,288,916,941]" country="Thailand" name="Herbarium, Department of Biology, Khon Kaen University" pageId="8" pageNumber="255">KKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5C3C9BFF9E8D1AFEFBCB9EDFD4FC62" box="[310,1189,916,941]" collectingDate="1971-10-29" collectorName="Vidal" country="Laos" location="Phou Khao Khouay" pageId="8" pageNumber="255" specimenCount="1" typeStatus="paratype">
<location id="8EEB601DFF9E8D1AFEFBCB9ED968FC62" LSID="urn:lsid:plazi:treatment:039D87D0FF908D1AFF45CF25DB8BF89A:8EEB601DFF9E8D1AFEFBCB9ED968FC62" box="[310,537,916,941]" country="Laos" name="Phou Khao Khouay" pageId="8" pageNumber="255">Phou Khao Khouay</location>
,
<date id="FF8A1006FF9E8D1AFDEBCB9ED9CFFC62" box="[550,702,916,941]" pageId="8" pageNumber="255" value="1971-10-29">
<collectingDate id="EFCEE9EEFF9E8D1AFDEBCB9ED9CFFC62" box="[550,702,916,941]" pageId="8" pageNumber="255" value="1971-10-29">29 Oct. 1971</collectingDate>
</date>
,
<emphasis id="B940EAD4FF9E8D1AFD00CB9ED838FC62" box="[717,841,916,941]" italics="true" pageId="8" pageNumber="255">
<collectorName id="26C15310FF9E8D1AFD00CB9ED879FC62" box="[717,776,916,941]" pageId="8" pageNumber="255">Vidal</collectorName>
5969
</emphasis>
[fr.] (P [P00589453], image!)
</materialsCitation>
.
<materialsCitation id="3B5C3C9BFF9E8D1AFB7FCB9EDBAFFBD6" collectingDate="2001-08-25" collectorName="Pooma R. &amp; de Wilde W. J. J. O. &amp; Duyfjes B. E. E. &amp; Chamchumroon V. &amp; Phattarahirankanok" country="Thailand" county="Ban Phaeng" elevation="150" latitude="17.95" location="Tat Kham Fall" longLatPrecision="21" longitude="104.162224" municipality="Phu Langka National Park" pageId="8" pageNumber="255" specimenCount="1" stateProvince="Nakhon Phanom" typeStatus="paratype">
<collectingCountry id="F3237656FF9E8D1AFB7FCB9EDE37FC62" box="[1202,1350,916,941]" name="Thailand" pageId="8" pageNumber="255">THAILAND</collectingCountry>
.
<collectingRegion id="49F0F824FF9E8D1AFA9ECB9EDA22FC1E" country="Thailand" name="Nakhon Phanom" pageId="8" pageNumber="255">Nakhon Phanom Province</collectingRegion>
,
<collectingCounty id="62EA4E4AFF9E8D1AFEADCBB2DA9AFC1E" box="[352,491,952,977]" pageId="8" pageNumber="255">Ban Phaeng</collectingCounty>
,
<collectingMunicipality id="6BEFACBCFF9E8D1AFE35CBB2D858FC1E" box="[504,809,952,977]" pageId="8" pageNumber="255">Phu Langka National Park</collectingMunicipality>
,
<location id="8EEB601DFF9E8D1AFCF8CBB2D8ACFC1E" LSID="urn:lsid:plazi:treatment:039D87D0FF908D1AFF45CF25DB8BF89A:8EEB601DFF9E8D1AFCF8CBB2D8ACFC1E" box="[821,989,952,977]" country="Thailand" county="Ban Phaeng" latitude="17.95" longLatPrecision="21" longitude="104.162224" municipality="Phu Langka National Park" name="Tat Kham Fall" pageId="8" pageNumber="255" stateProvince="Nakhon Phanom">Tat Kham Fall</location>
,
<geoCoordinate id="EE005001FF9E8D1AFC27CBB2DF03FC1E" box="[1002,1138,952,977]" degrees="17" direction="north" minutes="57" orientation="latitude" pageId="8" pageNumber="255" precision="15" seconds="00" value="17.95">17°5700″N</geoCoordinate>
,
<geoCoordinate id="EE005001FF9E8D1AFBB2CBB2DE60FC1E" box="[1151,1297,952,977]" degrees="104" direction="east" minutes="09" orientation="longitude" pageId="8" pageNumber="255" precision="15" seconds="44" value="104.162224">104°0944″E</geoCoordinate>
, elev.
<quantity id="4CCC9B23FF9E8D1AFA91CBB2DED6FC1E" box="[1372,1447,952,977]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="255" unit="m" value="150.0">
<elevation id="0019D1F5FF9E8D1AFA91CBB2DED6FC1E" box="[1372,1447,952,977]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="255" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<date id="FF8A1006FF9E8D1AFF45CBD6DA54FC3A" box="[136,293,988,1013]" pageId="8" pageNumber="255" value="2001-08-25">
<collectingDate id="EFCEE9EEFF9E8D1AFF45CBD6DA54FC3A" box="[136,293,988,1013]" pageId="8" pageNumber="255" value="2001-08-25">25 Aug. 2001</collectingDate>
</date>
,
<emphasis id="B940EAD4FF9E8D1AFEFCCBD6DE09FC3A" box="[305,1400,988,1013]" italics="true" pageId="8" pageNumber="255">
<collectorName id="26C15310FF9E8D1AFEFCCBD6DAD2FC3A" box="[305,419,988,1013]" pageId="8" pageNumber="255">Pooma R.</collectorName>
,
<collectorName id="26C15310FF9E8D1AFE7DCBD6D90AFC3A" box="[432,635,988,1013]" pageId="8" pageNumber="255">de Wilde W.J.J.O.</collectorName>
,
<collectorName id="26C15310FF9E8D1AFD45CBD6D841FC3A" box="[648,816,988,1013]" pageId="8" pageNumber="255">Duyfjes B.E.E.</collectorName>
,
<collectorName id="26C15310FF9E8D1AFCF0CBD6DF7FFC3A" box="[829,1038,988,1013]" pageId="8" pageNumber="255">Chamchumroon V.</collectorName>
&amp;
<collectorName id="26C15310FF9E8D1AFBFECBD6DE67FC3A" box="[1075,1302,988,1013]" pageId="8" pageNumber="255">Phattarahirankanok</collectorName>
K. 2613
</emphasis>
[fl.] (BKF!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32E654DFF9E8D1AFF70CC2EDFAFFB3E" pageId="8" pageNumber="255" type="distribution">
<paragraph id="8B8B36C6FF9E8D1AFF70CC2EDFBEFBAE" blockId="8.[136,1452,160,1877]" pageId="8" pageNumber="255">
<emphasis id="B940EAD4FF9E8D1AFF70CC2EDA04FBF2" bold="true" box="[189,373,1060,1085]" pageId="8" pageNumber="255">Distribution:—</emphasis>
<collectingCountry id="F3237656FF9E8D1AFEB8CC2EDADAFBF2" box="[373,427,1060,1085]" name="Laos" pageId="8" pageNumber="255">Laos</collectingCountry>
(
<collectingRegion id="49F0F824FF9E8D1AFE72CC2ED9D0FBF2" box="[447,673,1060,1085]" country="Laos" name="Viangchan" pageId="8" pageNumber="255">Vientiane Province</collectingRegion>
: Phou Khao Khouay National Protected Area),
<collectingCountry id="F3237656FF9E8D1AFB14CC2EDE4CFBF2" box="[1241,1341,1060,1085]" name="Thailand" pageId="8" pageNumber="255">Thailand</collectingCountry>
(
<collectingRegion id="49F0F824FF9E8D1AFA9CCC2EDA22FBAE" country="Thailand" name="Nakhon Phanom" pageId="8" pageNumber="255">Nakhon Phanom Province</collectingRegion>
: Phu Langka National Park, Buengkan Province: Phu Wua Wildlife Sanctuary).
</paragraph>
<paragraph id="8B8B36C6FF9E8D1AFF70CC66DA89FB66" blockId="8.[136,1452,160,1877]" pageId="8" pageNumber="255">
<emphasis id="B940EAD4FF9E8D1AFF70CC66DAA6FB4A" bold="true" box="[189,471,1132,1157]" pageId="8" pageNumber="255">Habitat and Ecology:—</emphasis>
In hill evergreen forest, at altitudes of
<quantity id="4CCC9B23FF9E8D1AFC4CCC66D88EFB4A" box="[897,1023,1132,1157]" metricMagnitude="2" metricUnit="m" metricValue="7.3" metricValueMax="7.7" metricValueMin="6.9" pageId="8" pageNumber="255" unit="m" value="730.0" valueMax="770.0" valueMin="690.0">690770 m</quantity>
in
<collectingCountry id="F3237656FF9E8D1AFBEFCC66DF2AFB4A" box="[1058,1115,1132,1157]" name="Laos" pageId="8" pageNumber="255">Laos</collectingCountry>
, and in dry evergreen forest at an altitude of
<quantity id="4CCC9B23FF9E8D1AFEEECC9ADA1BFB66" box="[291,362,1168,1193]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="255" unit="m" value="150.0">150 m</quantity>
in
<collectingCountry id="F3237656FF9E8D1AFE40CC9ADA85FB66" box="[397,500,1168,1193]" name="Thailand" pageId="8" pageNumber="255">Thailand</collectingCountry>
.
</paragraph>
<paragraph id="8B8B36C6FF9E8D1AFF70CCBEDE27FB02" blockId="8.[136,1452,160,1877]" box="[189,1366,1204,1229]" pageId="8" pageNumber="255">
<emphasis id="B940EAD4FF9E8D1AFF70CCBEDA2EFB02" bold="true" box="[189,351,1204,1229]" pageId="8" pageNumber="255">Phenology:—</emphasis>
Flowering specimens collected in August and November, fruiting in October to December.
</paragraph>
<paragraph id="8B8B36C6FF9E8D1AFF70CCD2DFAFFB3E" blockId="8.[136,1452,160,1877]" box="[189,1246,1240,1265]" pageId="8" pageNumber="255">
<emphasis id="B940EAD4FF9E8D1AFF70CCD2DAFEFB3E" bold="true" box="[189,399,1240,1265]" pageId="8" pageNumber="255">DNA barcodes:—</emphasis>
LC381850 (
<emphasis id="B940EAD4FF9E8D1AFDD5CCD2D93CFB3E" box="[536,589,1240,1265]" italics="true" pageId="8" pageNumber="255">rbcL</emphasis>
), LC381853 (
<emphasis id="B940EAD4FF9E8D1AFD21CCD2D858FB3E" box="[748,809,1240,1265]" italics="true" pageId="8" pageNumber="255">matK</emphasis>
); sequenced from
<emphasis id="B940EAD4FF9E8D1AFC37CCD2DFA6FB3E" box="[1018,1239,1240,1265]" italics="true" pageId="8" pageNumber="255">Yahara et al. L1438</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C32E654DFF9E8D1AFF70CCF6DB8BF89A" pageId="8" pageNumber="255" type="etymology">
<paragraph id="8B8B36C6FF9E8D1AFF70CCF6DA41FAF6" blockId="8.[136,1452,160,1877]" pageId="8" pageNumber="255">
<emphasis id="B940EAD4FF9E8D1AFF70CCF6DA15FADA" bold="true" box="[189,356,1276,1301]" pageId="8" pageNumber="255">Etymology:—</emphasis>
The species epithet refer to its nature of roughened (scabrid) surfaces of twigs and leaves caused by dense hairs.
</paragraph>
<paragraph id="8B8B36C6FF9E8D1AFF70CD4EDF28FA92" blockId="8.[136,1452,160,1877]" box="[189,1113,1343,1373]" pageId="8" pageNumber="255">
<emphasis id="B940EAD4FF9E8D1AFF70CD4ED975FA92" bold="true" box="[189,516,1343,1373]" pageId="8" pageNumber="255">Vernacular name:—หวาขน</emphasis>
(Wa Khon) (
<emphasis id="B940EAD4FF9E8D1AFD55CD4ED80FFA92" bold="true" box="[664,894,1343,1373]" pageId="8" pageNumber="255">
<collectingCountry id="F3237656FF9E8D1AFD55CD4ED87BFA92" box="[664,778,1348,1373]" name="Thailand" pageId="8" pageNumber="255">Thailand</collectingCountry>
); ຫວາຂນ
</emphasis>
(Wa Khon) (Laos).
</paragraph>
<paragraph id="8B8B36C6FF9E8D1AFF70CD62D971F9DE" blockId="8.[136,1452,160,1877]" pageId="8" pageNumber="255">
<emphasis id="B940EAD4FF9E8D1AFF70CD62D9EFFA4E" bold="true" box="[189,670,1384,1409]" pageId="8" pageNumber="255">Preliminary conservation assessment:—</emphasis>
Endangered (EN). Based on collections and our field observations,
<taxonomicName id="4C344D45FF9E8D1AFA5BCD62DB99FA6A" authorityName="Tagane, Soulad. &amp; Yahara. A" authorityYear="2018" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="255" phylum="Tracheophyta" rank="species" species="scabrum">
<emphasis id="B940EAD4FF9E8D1AFA5BCD62DB99FA6A" italics="true" pageId="8" pageNumber="255">S. scabrum</emphasis>
</taxonomicName>
is considered to be widely distributed in forests at lower to high elevations (
<quantity id="4CCC9B23FF9E8D1AFBE0CD86DFDCFA6A" box="[1069,1197,1420,1445]" metricMagnitude="2" metricUnit="m" metricValue="4.75" metricValueMax="8.0" metricValueMin="1.5" pageId="8" pageNumber="255" unit="m" value="475.0" valueMax="800.0" valueMin="150.0">150800 m</quantity>
) in Northeast
<collectingCountry id="F3237656FF9E8D1AFA85CD86DEDDFA6A" box="[1352,1452,1420,1445]" name="Thailand" pageId="8" pageNumber="255">Thailand</collectingCountry>
and Central
<collectingCountry id="F3237656FF9E8D1AFEDCCDBADA3BFA06" box="[273,330,1456,1481]" name="Laos" pageId="8" pageNumber="255">Laos</collectingCountry>
, and the area of occupancy (AOO) of this species calculated based on the collection localities is less than
<quantity id="4CCC9B23FF9E8D1AFF72CDDEDA53FA22" box="[191,290,1492,1517]" metricMagnitude="6" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="255" unit="km" value="2200.0">2200 km</quantity>
<superScript id="7C419B8EFF9E8D1AFEEFCDD8DA5BFA2F" attach="left" box="[290,298,1490,1504]" fontSize="6" pageId="8" pageNumber="255">2</superScript>
with some disturbed localities in
<collectingCountry id="F3237656FF9E8D1AFD6ECDDED9ACFA22" box="[675,733,1492,1517]" name="Laos" pageId="8" pageNumber="255">Laos</collectingCountry>
. Therefore we qualify this species as EN according to the IUCN criteria of B1a &amp; b (
<bibRefCitation id="EFA54B37FF9E8D1AFEA7CDF2DA85F9DE" author="IUCN" box="[362,500,1528,1553]" pageId="8" pageNumber="255" refId="ref7560" refString="IUCN (2012) IUCN Red List Categories and Criteria. Version 3.1. Second edition. IUCN, Switzerland and Cambridge, Gland, 32 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
<paragraph id="8B8B36C6FF9E8D1AFF70CE16DF13F8C2" blockId="8.[136,1452,160,1877]" pageId="8" pageNumber="255">
<emphasis id="B940EAD4FF9E8D1AFF70CE16DA64F9FA" bold="true" box="[189,277,1564,1589]" pageId="8" pageNumber="255">Note 1:</emphasis>
—This species has been confused with
<taxonomicName id="4C344D45FF9E8D1AFD15CE16D822F9FA" authorityName="Merrill &amp; Perry" authorityYear="1938" box="[728,851,1564,1589]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="255" phylum="Tracheophyta" rank="species" species="vestitum">
<emphasis id="B940EAD4FF9E8D1AFD15CE16D822F9FA" box="[728,851,1564,1589]" italics="true" pageId="8" pageNumber="255">S. vestitum</emphasis>
</taxonomicName>
(
<typeStatus id="548F8864FF9E8D1AFCA9CE16D8EBF9FA" box="[868,922,1564,1589]" pageId="8" pageNumber="255">type</typeStatus>
: Mt. Bana,
<collectingCountry id="F3237656FF9E8D1AFBD2CE16DFF0F9FA" box="[1055,1153,1564,1589]" name="Vietnam" pageId="8" pageNumber="255">Vietnam</collectingCountry>
,
<emphasis id="B940EAD4FF9E8D1AFB40CE16DED5F9FA" box="[1165,1444,1564,1589]" italics="true" pageId="8" pageNumber="255">J. &amp; M.S. Clemens 3296</emphasis>
, K, image!) by
<bibRefCitation id="EFA54B37FF9E8D1AFEFBCE4AD9EBF996" author="Souladeth, P. &amp; Meesawat, A." box="[310,666,1600,1625]" pageId="8" pageNumber="255" pagination="927 - 931" refId="ref8794" refString="Souladeth, P. &amp; Meesawat, A. (2012) Myrtaceae in Phou Khao Khouay National Protected Area, Lao PDR. Proceedings: 1 st ASEAN Plus Three Graduate Research Congress 2012: 927 - 931." type="journal article" year="2012">Souladeth &amp; Meesawat (2012)</bibRefCitation>
,
<bibRefCitation id="EFA54B37FF9E8D1AFD64CE4AD8DDF996" author="Chantaranothai, P." box="[681,940,1600,1625]" pageId="8" pageNumber="255" pagination="75 - 80" refId="ref6955" refString="Chantaranothai, P. (2014) Three new records of Syzygium Gaertn. in Thailand and lectotypification of 19 taxa of Eugenia L. (Myrtaceae). Thai Forest Bulletin (Botany) 42: 75 - 80." type="book chapter" year="2014">Chantaranothai (2014)</bibRefCitation>
, and
<bibRefCitation id="EFA54B37FF9E8D1AFC23CE4ADF9AF996" author="Soh, W. - K. &amp; Parnell, J." box="[1006,1259,1600,1625]" pageId="8" pageNumber="255" pagination="179 - 275" refId="ref8731" refString="Soh, W. - K. &amp; Parnell, J. (2015) A revision of Syzygium Gaertn. (Myrtaceae) in Indochina (Cambodia, Laos and Vietnam). Adansonia Ser. 3, 37 (2): 179 - 275. https: // doi. org / 10.5252 / a 2015 n 2 a 1" type="journal article" year="2015">Soh &amp; Parnell (2015)</bibRefCitation>
, but it is clearly distinguished from
<taxonomicName id="4C344D45FF9E8D1AFEADCE6EDAA9F9B2" authorityName="Merrill &amp; Perry" authorityYear="1938" box="[352,472,1636,1661]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="255" phylum="Tracheophyta" rank="species" species="vestitum">
<emphasis id="B940EAD4FF9E8D1AFEADCE6EDAA9F9B2" box="[352,472,1636,1661]" italics="true" pageId="8" pageNumber="255">S. vestitum</emphasis>
</taxonomicName>
by the above diagnostic characters.
<taxonomicName id="4C344D45FF9E8D1AFCA4CE6EDF47F9B2" authorityName="Merrill &amp; Perry" authorityYear="1938" box="[873,1078,1636,1661]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="255" phylum="Tracheophyta" rank="species" species="vestitum">
<emphasis id="B940EAD4FF9E8D1AFCA4CE6EDF47F9B2" box="[873,1078,1636,1661]" italics="true" pageId="8" pageNumber="255">Syzygium vestitum</emphasis>
</taxonomicName>
is restricted to northern to central
<collectingCountry id="F3237656FF9E8D1AFF45CE82DB96F96E" box="[136,231,1672,1697]" name="Vietnam" pageId="8" pageNumber="255">Vietnam</collectingCountry>
and southern
<collectingCountry id="F3237656FF9E8D1AFE45CE82DABAF96E" box="[392,459,1672,1697]" name="China" pageId="8" pageNumber="255">China</collectingCountry>
(southeast
<collectingRegion id="49F0F824FF9E8D1AFD81CE82D9D5F96E" box="[588,676,1672,1697]" country="China" name="Yunnan" pageId="8" pageNumber="255">Yunnan</collectingRegion>
) (
<bibRefCitation id="EFA54B37FF9E8D1AFD76CE82D853F96E" author="Ho, P. H." box="[699,802,1672,1697]" pageId="8" pageNumber="255" refId="ref7530" refString="Ho, P. H. (2003) An Illustrated Flora of Vietnam. Vol. 2. published by author, Ho Chi Minh, 951 pp." type="book" year="2003">Hô 2003</bibRefCitation>
,
<bibRefCitation id="EFA54B37FF9E8D1AFCE3CE82DF52F96E" author="Chen, J. &amp; Craven, L. A." box="[814,1059,1672,1697]" pageId="8" pageNumber="255" refId="ref6996" refString="Chen, J. &amp; Craven, L. A. (2007) Myrtaceae. In: Wu, Z. Y., Raven, P. H. &amp; Hong, D. Y. (Eds.) Flora of China. Vol. 13. Science Press, Beijing and Missouri Botanical Garden Press, St Louis, Missouri, pp. 321 - 359." type="book" year="2007">Chen &amp; Craven 2007</bibRefCitation>
, from our field observations in SE Asia). The sequence of
<emphasis id="B940EAD4FF9E8D1AFE43CEA6DABAF90A" box="[398,459,1708,1733]" italics="true" pageId="8" pageNumber="255">matK</emphasis>
region of
<taxonomicName id="4C344D45FF9E8D1AFDF0CEA6D9C9F90A" authorityName="Tagane, Soulad. &amp; Yahara. A" authorityYear="2018" box="[573,696,1708,1733]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="255" phylum="Tracheophyta" rank="species" species="scabrum">
<emphasis id="B940EAD4FF9E8D1AFDF0CEA6D9C9F90A" box="[573,696,1708,1733]" italics="true" pageId="8" pageNumber="255">S. scabrum</emphasis>
</taxonomicName>
(GenBank accession no. LC381853) differs 7 bp of the total 760 bp from the
<taxonomicName id="4C344D45FF9E8D1AFF23CEDADA17F926" authorityName="Merrill &amp; Perry" authorityYear="1938" box="[238,358,1744,1769]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="255" phylum="Tracheophyta" rank="species" species="vestitum">
<emphasis id="B940EAD4FF9E8D1AFF23CEDADA17F926" box="[238,358,1744,1769]" italics="true" pageId="8" pageNumber="255">S. vestitum</emphasis>
</taxonomicName>
(LC381852:
<emphasis id="B940EAD4FF9E8D1AFE30CEDAD9ADF926" box="[509,732,1744,1769]" italics="true" pageId="8" pageNumber="255">
Tagane et al.
<date id="FF8A1006FF9E8D1AFD5ECEDAD9ADF926" box="[659,732,1744,1769]" pageId="8" pageNumber="255" value="2522-05">V2522</date>
</emphasis>
(FU!) from Bach Ma National Park, Central
<collectingCountry id="F3237656FF9E8D1AFB1FCEDADE45F926" box="[1234,1332,1744,1769]" name="Vietnam" pageId="8" pageNumber="255">Vietnam</collectingCountry>
, ca.
<quantity id="4CCC9B23FF9E8D1AFAA8CEDADEDDF926" box="[1381,1452,1744,1769]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="255" unit="km" value="25.0">25 km</quantity>
apart from the
<typeStatus id="548F8864FF9E8D1AFEE3CEFEDA2FF8C2" box="[302,350,1780,1805]" pageId="8" pageNumber="255">type</typeStatus>
locality of
<taxonomicName id="4C344D45FF9E8D1AFE13CEFED926F8C2" authorityName="Merrill &amp; Perry" authorityYear="1938" box="[478,599,1780,1805]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="255" phylum="Tracheophyta" rank="species" species="vestitum">
<emphasis id="B940EAD4FF9E8D1AFE13CEFED926F8C2" box="[478,599,1780,1805]" italics="true" pageId="8" pageNumber="255">S. vestitum</emphasis>
</taxonomicName>
), supporting the separation of the two species.
</paragraph>
<paragraph id="8B8B36C6FF9E8D1AFF70CF12DB8BF89A" blockId="8.[136,1452,160,1877]" pageId="8" pageNumber="255">
<emphasis id="B940EAD4FF9E8D1AFF70CF12DA64F8FE" bold="true" box="[189,277,1816,1841]" pageId="8" pageNumber="255">Note 2:</emphasis>
—The specimens
<emphasis id="B940EAD4FF9E8D1AFE29CF12D829F8FE" box="[484,856,1816,1841]" italics="true" pageId="8" pageNumber="255">Souladeth P. 86, Souladeth P. 91</emphasis>
and
<emphasis id="B940EAD4FF9E8D1AFC5ECF12DFE2F8FE" box="[915,1171,1816,1841]" italics="true" pageId="8" pageNumber="255">Yahara T. et al. L1438</emphasis>
were collected from the same tree.
</paragraph>
</subSubSection>
</treatment>
</document>