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<document id="52542D30AD0D807037C9665C2C125E2E" ID-CLB-Dataset="306597" ID-DOI="10.1093/zoolinnean/zlad177" ID-GBIF-Dataset="dcaf5aa0-c7c6-4dc1-871a-8815b1aabb88" ID-ISSN="0024-4082" ID-Zenodo-Dep="14510261" ID-ZooBank="80D1924-E984-4900-88E5-85FE2C5688D8" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1734027918295" checkinUser="plazi" docAuthor="Hinckley, Arlo, Camacho-Sanchez, Miguel, Chua, Marcus A. H., Ruedi, Manuel, Lunde, Darrin, Maldonado, Jesús E., Omar, Hasmahzaiti, Leonard, Jennifer A. &amp; Hawkins, Melissa T. R." docDate="2024" docId="038C8797FF95FFC2FCC66386E3F1FB39" docLanguage="en" docName="zlad177.pdf" docOrigin="Zoological Journal of the Linnean Society 202 (2)" docSource="https://doi.org/10.1093/zoolinnean/zlad177" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Hylomys macarong Hinckley, Lunde &amp; Hawkins 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="20" masterDocId="FFB5FFEFFF85FFD1FF89661BE21AFFDF" masterDocTitle="An intbgoativb taxonomic obvision of lbssbo gvmnuobs (Eulipotvphla: Hylomys) obvbals fivb nbw spbcibs and bmboging paưbons of local bndbmism in Toopical East Asia" masterLastPageNumber="33" masterPageNumber="1" pageNumber="17" updateTime="1734460109358" updateUser="ExternalLinkService">
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<mods:title id="DCB381347F6305B1A1C86C33EA53AD7D">An intbgoativb taxonomic obvision of lbssbo gvmnuobs (Eulipotvphla: Hylomys) obvbals fivb nbw spbcibs and bmboging paưbons of local bndbmism in Toopical East Asia</mods:title>
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<mods:namePart id="9F3D976ABDA4B5D080A8297CF71F8188">Hinckley, Arlo</mods:namePart>
<mods:affiliation id="C33D5568A088F107CF2411C7C0F5F8EC">Division of Mammals, Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, District of Columbia, Washington, USA</mods:affiliation>
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<mods:namePart id="A1F80E2CF1A778B523BEB756AC07C458">Camacho-Sanchez, Miguel</mods:namePart>
<mods:affiliation id="4079AB3363A5C8E8E5C0CA64A2C25FEA">Conservation &amp; Evolutionary Genetics Group, Estación Biológica de Doñana (EBD-CSIC), Seville, Spain</mods:affiliation>
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<mods:namePart id="A14C021F10D8D9095917609B39CCC306">Chua, Marcus A. H.</mods:namePart>
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<mods:namePart id="4DE044D2742BB05024C2DF2AEE286112">Ruedi, Manuel</mods:namePart>
<mods:affiliation id="CD78138DA5F82886107787C89A7C3A1A">Natural History Museum of Geneva, Geneva, Switzerland</mods:affiliation>
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<mods:namePart id="87746543FF215FFA280A53C3326A0968">Lunde, Darrin</mods:namePart>
<mods:affiliation id="529470AEC880AB5ACF70A9BB9E90DD7D">Division of Mammals, Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, District of Columbia, Washington, USA</mods:affiliation>
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<mods:namePart id="8EDBA2385D7DD2BF75561ABEA930CEF7">Maldonado, Jesús E.</mods:namePart>
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<mods:namePart id="3686770460D6A31A594E6C1C5F679777">Omar, Hasmahzaiti</mods:namePart>
<mods:affiliation id="69052BF5EE3A8903D2595A577988BEEA">Museum of Zoology, Institute of Biological Sciences, Faculty of Science, Universiti Malaya, Kuala Lumpur, Malaysia</mods:affiliation>
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<mods:namePart id="ED5A69064941F416956EB4F74F073520">Hawkins, Melissa T. R.</mods:namePart>
<mods:affiliation id="3C61CF0E03D8371E41CE6C2F8B8494C4">Division of Mammals, Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, District of Columbia, Washington, USA</mods:affiliation>
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<mods:date id="5311980939F6E724ABD05C985E4A9630">2024</mods:date>
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<treatment id="038C8797FF95FFC2FCC66386E3F1FB39" ID-DOI="http://doi.org/10.5281/zenodo.14510283" ID-Zenodo-Dep="14510283" LSID="urn:lsid:plazi:treatment:038C8797FF95FFC2FCC66386E3F1FB39" httpUri="http://treatment.plazi.org/id/038C8797FF95FFC2FCC66386E3F1FB39" lastPageId="19" lastPageNumber="20" pageId="16" pageNumber="17">
<subSubSection id="C33F650AFF95FFC1FCC66386E7B6FA67" box="[847,1452,1437,1464]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8B9A3681FF95FFC1FCC66386E7B6FA67" blockId="16.[825,1474,1437,1594]" box="[847,1452,1437,1464]" pageId="16" pageNumber="17">
<emphasis id="B951EA93FF95FFC1FCC66386E7B6FA67" bold="true" box="[847,1452,1437,1464]" pageId="16" pageNumber="17">
<taxonomicName id="4C254D02FF95FFC1FCC66386E74CFA68" authority="Hinckley, Lunde &amp; Hawkins" authorityName="Hinckley, Lunde &amp; Hawkins" authorityYear="2024" box="[847,1366,1437,1464]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="macarong" status="sp. nov.">
<emphasis id="B951EA93FF95FFC1FCC66386E63DFA67" bold="true" box="[847,1063,1437,1464]" italics="true" pageId="16" pageNumber="17">Hylomys macarong</emphasis>
Hinckley, Lunde &amp; Hawkins
</taxonomicName>
,
<taxonomicNameLabel id="A26257E8FF95FFC1FAE96385E7B6FA67" box="[1376,1452,1438,1464]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8B9A3681FF95FFC1FCB063DFE775FA03" blockId="16.[825,1474,1437,1594]" box="[825,1391,1476,1500]" pageId="16" pageNumber="17">
<taxonomicName id="4C254D02FF95FFC1FCB063DFE1B5FA04" box="[825,943,1476,1500]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B951EA93FF95FFC1FCB063DFE194FA03" box="[825,910,1476,1500]" italics="true" pageId="16" pageNumber="17">Hylomys</emphasis>
sp.
</taxonomicName>
Bannikova et al., 2014: 501. Informal name.
</paragraph>
<paragraph id="8B9A3681FF95FFC1FCB063F8E6F1FA24" blockId="16.[825,1474,1437,1594]" box="[825,1259,1507,1531]" pageId="16" pageNumber="17">
<taxonomicName id="4C254D02FF95FFC1FCB063F8E6FCFA24" authority=", Bbst, 2018: 328" authorityName="Bbst" authorityPageNumber="328" authorityYear="2018" box="[825,1254,1507,1531]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="subSpecies" species="suillus" subSpecies="microtinus">
<emphasis id="B951EA93FF95FFC1FCB063F8E621FA24" box="[825,1083,1507,1531]" italics="true" pageId="16" pageNumber="17">Hylomys suillus microtinus</emphasis>
, Best, 2018: 328
</taxonomicName>
.
</paragraph>
<paragraph id="8B9A3681FF95FFC1FCB06019E162F9E5" blockId="16.[825,1474,1437,1594]" pageId="16" pageNumber="17">
<taxonomicName id="4C254D02FF95FFC1FCB06019E1CBF9C5" authorityName="Mullbo" authorityYear="1840" box="[825,977,1538,1562]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="suillus">
<emphasis id="B951EA93FF95FFC1FCB06019E1CBF9C5" box="[825,977,1538,1562]" italics="true" pageId="16" pageNumber="17">Hylomys suillus</emphasis>
</taxonomicName>
ssp. 2 Zeng et al., 2022: preprint fig. 3. Informal name.
</paragraph>
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<subSubSection id="C33F650AFF95FFC1FCB06040E6ACF94D" pageId="16" pageNumber="17" type="description">
<paragraph id="8B9A3681FF95FFC1FCB06040E6ACF94D" blockId="16.[825,1474,1627,1682]" pageId="16" pageNumber="17">
<emphasis id="B951EA93FF95FFC1FCB06040E639F9AC" box="[825,1059,1627,1651]" italics="true" pageId="16" pageNumber="17">Zoobank registration:</emphasis>
urn:lsid:zoobank.org:act:
<uuid id="FF830C54FF95FFC1FACF6040E6A8F94D" pageId="16" pageNumber="17">A1305A1B-AA64-479F-9D88-59EAE5CCE2F0</uuid>
.
</paragraph>
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<subSubSection id="C33F650AFF95FFC3FCB060AFE36EFD86" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8B9A3681FF95FFC0FCB060AFE1BCF89C" blockId="16.[825,1474,1716,1959]" lastBlockId="17.[810,1459,1710,1859]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<materialsCitation id="3B4D3CDCFF95FFC1FCB060AFE671F934" accessionNumber="OR138081, OR554435" collectingDate="1961-07-21" collectionCode="USNM" collectorName="Bernard R. Feinstein" country="Vietnam'" elevation="1200" latitude="11.88" location="Phu Son" longLatPrecision="5464" longitude="108.2" municipality="Lam Ha District" pageId="16" pageNumber="17" specimenCode="USNM 320500" specimenCount="1" typeStatus="holotype">
<emphasis id="B951EA93FF95FFC1FCB060AFE186F913" box="[825,924,1716,1740]" italics="true" pageId="16" pageNumber="17">
<typeStatus id="549E8823FF95FFC1FCB060AFE18CF913" box="[825,918,1716,1740]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="DB839EFAFF95FFC1FC2160AFE656F913" box="[936,1100,1716,1740]" pageId="16" pageNumber="17">USNM 320500</specimenCode>
, an
<specimenCount id="9D23FD08FF95FFC1FBFF60AFE6F9F913" box="[1142,1251,1716,1740]" pageId="16" pageNumber="17" type="adult">adult male</specimenCount>
collected by
<collectorName id="26D05357FF95FFC1FAE760AFE1A3F934" pageId="16" pageNumber="17">Bernard R. Feinstein</collectorName>
on
<date id="FF9B1041FF95FFC1FC6A60C8E671F934" box="[995,1131,1747,1771]" pageId="16" pageNumber="17" value="1961-07-21">
<collectingDate id="EFDFE9A9FF95FFC1FC6A60C8E671F934" box="[995,1131,1747,1771]" pageId="16" pageNumber="17" value="1961-07-21">21 July 1961</collectingDate>
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. The specimen consists of a museum study skin with accompanying cranium and mandible; skin is in relatively good condition, but is partially open along the suture line, and has a twisted right forelimb with the most external phalanx missing (clipped for DNA extraction), right ankle skin is pierced by tibia or calcaneus, partially chewed ears (possibly by ants, as reported for other specimens of the type series), and broken proximal and distal sections of tail, exposing the internal wire; skull is well prepared but has damaged pterygoid processes, left zygomatic arch, dorsal side of angular process and tips of left i1 and i2; and left P1 and occipital condyle are missing (unerupted and broken, respectively). Total length
<quantity id="4CDD9B64FF94FFC0FF356150E30DF8BC" box="[188,279,1867,1891]" metricMagnitude="-1" metricUnit="m" metricValue="1.65" pageId="17" pageNumber="18" unit="mm" value="165.0">165 mm</quantity>
, tail
<quantity id="4CDD9B64FF94FFC0FEC56150E38DF8BC" box="[332,407,1867,1891]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="18" unit="mm" value="25.0">25 mm</quantity>
(18% HB), hindfoot
<quantity id="4CDD9B64FF94FFC0FD086150E0D4F8BC" box="[641,718,1867,1891]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="17" pageNumber="18" unit="mm" value="27.0">27 mm</quantity>
, ear length
<quantity id="4CDD9B64FF94FFC0FF306171E31EF85D" box="[185,260,1898,1922]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="17" pageNumber="18" unit="mm" value="18.0">18 mm</quantity>
. Skull measurements in Supporting Information, Table S2. 3D surface scans of this holotype skull are available at MorphoSource: https://doi.org/10.17602/M2/M560006 (cranium); https://doi.org/10.17602/M2/M560011 (mandible). Pictures of this holotype skin and skull are shown in
<figureCitation id="131E2A04FF94FFC0FCC060F6E1A4F8DB" box="[841,958,1773,1797]" captionStart="Figure 7" captionStartId="16.[130,195,874,898]" captionTargetBox="[129,1471,144,847]" captionTargetId="figure-556@16.[129,1473,144,847]" captionTargetPageId="16" captionText="Figure 7. Dorsal and ventral views of the skins of (A) Hylomys peguensis (USNM583813, neotype), and (B) Hylomys macarong(USNM 320500, holotype). Photo credits:Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510277" httpUri="https://zenodo.org/record/14510277/files/figure.png" pageId="17" pageNumber="18">Figures 7B</figureCitation>
and
<figureCitation id="131E2A04FF94FFC0FC7060F6E60CF8DA" box="[1017,1046,1773,1797]" captionStart="Figure 8" captionStartId="17.[113,178,1610,1634]" captionTargetBox="[114,1458,144,1583]" captionTargetId="figure-160@17.[114,1458,144,1583]" captionTargetPageId="17" captionText="Figure 8. Skulls of (A) Hylomys peguensis (USNM583813, neotype) and (B) Hylomys macarong (USNM 320500, holotype).All images are at the same scale.Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510281" httpUri="https://zenodo.org/record/14510281/files/figure.png" pageId="17" pageNumber="18">8B</figureCitation>
. DNA sequences associated with this holotype have been deposited in GenBank:
<accessionNumber id="9476AB62FF94FFC0FA846117E798F8FB" box="[1293,1410,1804,1828]" pageId="17" pageNumber="18">OR138081</accessionNumber>
and
<accessionNumber id="9476AB62FF94FFC0FCA36130E1B8F89C" box="[810,930,1835,1859]" pageId="17" pageNumber="18">OR554435</accessionNumber>
.
</paragraph>
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<paragraph id="8B9A3681FF94FFC0FFF86051E01CF9A1" blockId="17.[113,1438,1610,1662]" pageId="17" pageNumber="18">
<emphasis id="B951EA93FF94FFC0FFF86051E2D3F9BD" bold="true" box="[113,201,1610,1634]" pageId="17" pageNumber="18">Figure 8.</emphasis>
Skulls of (A)
<taxonomicName id="4C254D02FF94FFC0FEC26051E3E9F9BD" authorityName="Blvth" authorityYear="1859" box="[331,499,1610,1634]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF94FFC0FEC26051E3E9F9BD" box="[331,499,1610,1634]" italics="true" pageId="17" pageNumber="18">Hylomys peguensis</emphasis>
</taxonomicName>
(USNM583813, neotype) and (B)
<taxonomicName id="4C254D02FF94FFC0FCCD6051E1EAF9BD" authorityName="Hinckley, Lunde &amp; Hawkins" authorityYear="2024" box="[836,1008,1610,1634]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="macarong">
<emphasis id="B951EA93FF94FFC0FCCD6051E1EAF9BD" box="[836,1008,1610,1634]" italics="true" pageId="17" pageNumber="18">Hylomys macarong</emphasis>
</taxonomicName>
(USNM 320500, holotype). All images are at the same scale. Photo credits: Katie Sayers.
</paragraph>
</caption>
<paragraph id="8B9A3681FF94FFC0FCA36177E6AEF81D" blockId="17.[810,1459,1899,1986]" pageId="17" pageNumber="18">
<emphasis id="B951EA93FF94FFC0FCA36177E1D9F85D" box="[810,963,1899,1923]" italics="true" pageId="17" pageNumber="18">Type locality: ‘</emphasis>
Fyan, Lam Dong,
<collectingCountry id="F3327611FF94FFC0FB096170E6C6F85C" box="[1152,1244,1899,1923]" pageId="17" pageNumber="18">Vietnam’</collectingCountry>
(now known as
<location id="8EFA605AFF94FFC0FA006170E14EF87D" LSID="urn:lsid:plazi:treatment:038C8797FF95FFC2FCC66386E3F1FB39:8EFA605AFF94FFC0FA006170E14EF87D" country="Vietnam'" latitude="11.88" longLatPrecision="5464" longitude="108.2" municipality="Lam Ha District" name="Phu Son" pageId="17" pageNumber="18">Phú Sơn</location>
,
<collectingMunicipality id="6BFEACFBFF94FFC0FCD76191E613F87D" box="[862,1033,1930,1954]" pageId="17" pageNumber="18">Lâm Hà District</collectingMunicipality>
, Viet Nam,
<geoCoordinate id="EE115046FF94FFC0FB0B6191E6FBF87D" box="[1154,1249,1930,1954]" degrees="11.88" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="555" value="11.88">11.88º N</geoCoordinate>
,
<geoCoordinate id="EE115046FF94FFC0FB626191E75FF87D" box="[1259,1349,1930,1954]" degrees="108.2" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="5555" value="108.2">108.2º E</geoCoordinate>
,
<emphasis id="B951EA93FF94FFC0FAC66190E745F87D" box="[1359,1375,1931,1954]" italics="true" pageId="17" pageNumber="18">c.</emphasis>
<elevation id="0008D1B2FF94FFC0FAED6191E142F81D" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="18" unit="m" value="1200.0">
<quantity id="4CDD9B64FF94FFC0FAED6191E7A9F87D" box="[1380,1459,1930,1954]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="17" pageNumber="18" unit="m" value="1200.0">1200 m</quantity>
a.s.l.
</elevation>
; source: US military intelligence).
</paragraph>
<paragraph id="8B9A3681FF97FFC3FF08668AE3BFFED9" blockId="18.[129,778,144,262]" pageId="18" pageNumber="19">
<materialsCitation id="3B4D3CDCFF97FFC3FF08668AE11CFF77" box="[129,774,144,168]" collectionCode="USNM" pageId="18" pageNumber="19" specimenCode="USNM 320492, 320495" specimenCount="2" typeStatus="paratype">
<emphasis id="B951EA93FF97FFC3FF08668AE30BFF77" box="[129,273,144,168]" italics="true" pageId="18" pageNumber="19">
<typeStatus id="549E8823FF97FFC3FF08668AE2FAFF77" box="[129,224,145,168]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D23FD08FF97FFC3FF78668BE31AFF77" box="[241,256,144,168]" pageId="18" pageNumber="19">2</specimenCount>
):
</emphasis>
<specimenCode id="DB839EFAFF97FFC3FEA9668BE3D9FF77" box="[288,451,144,168]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">USNM 320492</specimenCode>
,
<specimenCode id="DB839EFAFF97FFC3FE47668BE03AFF77" box="[462,544,144,168]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">320495</specimenCode>
: same as type locality
</materialsCitation>
;
<materialsCitation id="3B4D3CDCFF97FFC3FF0866B4E31FFF38" collectionCode="USNM" country="Vietnam" county="Nac Datan La" elevation="1450" location="Thac Datan La" municipality="Dalat" pageId="18" pageNumber="19" specimenCode="USNM 320485" specimenCount="1" stateProvince="Lam Dong" typeStatus="paratype">
<specimenCode id="DB839EFAFF97FFC3FF0866B4E33EFF18" box="[129,292,175,199]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">USNM 320485</specimenCode>
:
<location id="8EFA605AFF97FFC3FEA466ABE3DFFF18" LSID="urn:lsid:plazi:treatment:038C8797FF95FFC2FCC66386E3F1FB39:8EFA605AFF97FFC3FEA466ABE3DFFF18" box="[301,453,176,200]" country="Vietnam" county="Nac Datan La" municipality="Dalat" name="Thac Datan La" pageId="18" pageNumber="19" stateProvince="Lam Dong">Thac Datan La</location>
,
<collectingMunicipality id="6BFEACFBFF97FFC3FE4666B4E012FF18" box="[463,520,175,199]" pageId="18" pageNumber="19">Dalat</collectingMunicipality>
,
<collectingRegion id="49E1F863FF97FFC3FD9866ABE099FF17" box="[529,643,176,200]" country="Vietnam" name="Lam Dong" pageId="18" pageNumber="19">Lam Dong</collectingRegion>
,
<collectingCountry id="F3327611FF97FFC3FD0566B4E0EBFF18" box="[652,753,175,199]" name="Vietnam" pageId="18" pageNumber="19">Viet Nam</collectingCountry>
,
<emphasis id="B951EA93FF97FFC3FD7266ABE110FF18" box="[763,778,176,199]" italics="true" pageId="18" pageNumber="19">c.</emphasis>
<elevation id="0008D1B2FF97FFC3FF0866D5E31FFF38" box="[129,261,206,231]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="18" pageNumber="19" unit="m" value="1450.0">
<quantity id="4CDD9B64FF97FFC3FF0866D5E2CAFF39" box="[129,208,206,230]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="18" pageNumber="19" unit="m" value="1450.0">1450 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
;
<materialsCitation id="3B4D3CDCFF97FFC3FE9966D4E3BFFED9" collectionCode="USNM" elevation="500" location="Poste De Mdrak" municipality="Dac Lak" pageId="18" pageNumber="19" specimenCode="USNM 320490" specimenCount="1" stateProvince="Lam Dong" typeStatus="paratype">
<specimenCode id="DB839EFAFF97FFC3FE9966D4E3AFFF38" box="[272,437,207,231]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">USNM 320490</specimenCode>
:
<locationDeviation id="9A282FA3FF97FFC3FE3666D4E013FF39" box="[447,521,207,231]" pageId="18" pageNumber="19">
<quantity id="4CDD9B64FF97FFC3FE3666D4E3E9FF38" box="[447,499,207,231]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="km" value="1.0">1 km</quantity>
E
</locationDeviation>
of
<location id="8EFA605AFF97FFC3FDA266D4E0CCFF38" LSID="urn:lsid:plazi:treatment:038C8797FF95FFC2FCC66386E3F1FB39:8EFA605AFF97FFC3FDA266D4E0CCFF38" box="[555,726,207,231]" municipality="Dac Lak" name="Poste De Mdrak" pageId="18" pageNumber="19" stateProvince="Lam Dong">Poste De Mdrak</location>
,
<collectingMunicipality id="6BFEACFBFF97FFC3FD6966D4E2B0FED9" pageId="18" pageNumber="19">Dac Lak</collectingMunicipality>
,
<location id="8EFA605AFF97FFC3FF3A66F5E30CFEDA" LSID="urn:lsid:plazi:treatment:038C8797FF95FFC2FCC66386E3F1FB39:8EFA605AFF97FFC3FF3A66F5E30CFEDA" box="[179,278,238,262]" municipality="Dac Lak" name="Viet Nam" pageId="18" pageNumber="19" stateProvince="Lam Dong">Viet Nam</location>
,
<emphasis id="B951EA93FF97FFC3FEA966F4E335FED9" box="[288,303,239,262]" italics="true" pageId="18" pageNumber="19">c.</emphasis>
<elevation id="0008D1B2FF97FFC3FEBA66F5E3BFFED9" box="[307,421,238,262]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="m" value="500.0">
<quantity id="4CDD9B64FF97FFC3FEBA66F5E36FFEDA" box="[307,373,238,262]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="m" value="500.0">500 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
</paragraph>
<paragraph id="8B9A3681FF97FFC3FF086733E36EFD86" blockId="18.[129,778,295,601]" pageId="18" pageNumber="19">
<emphasis id="B951EA93FF97FFC3FF086733E3A5FEE0" box="[129,447,296,319]" italics="true" pageId="18" pageNumber="19">Representative DNA sequences:</emphasis>
Deposited (or already available) in GenBank with the following accession numbers: KF783150, KF783151 (cytochrome
<emphasis id="B951EA93FF97FFC3FD87677DE000FEA1" box="[526,538,358,382]" italics="true" pageId="18" pageNumber="19">b</emphasis>
, two representatives), (mitochondrial genomes, two representatives). New sequences: OR138079-OR138081 (mitochondrial genomes, three representatives), OR554432-OR554435(
<emphasis id="B951EA93FF97FFC3FE7B67DFE032FE04" box="[498,552,452,475]" italics="true" pageId="18" pageNumber="19">GHR</emphasis>
, four representatives). DNA sequences associated with the following voucher specimens: USNM 320485, 320490, 320495, 320500 (this study); USNM 320485, 320501 (Zeng
<emphasis id="B951EA93FF97FFC3FE406438E3E3FDE5" box="[457,505,546,570]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2022); ZMMU S-190307 (
<bibRefCitation id="EFB44B70FF97FFC3FF02645AE37EFD86" author="Bannikova AA &amp; Aboamov AV" box="[139,356,577,601]" pageId="18" pageNumber="19" pagination="499 - 519" refId="ref30844" refString="Bannikova AA, Lbbbdbv VS, Aboamov AV et al. Contoasting bvolutionaov histoov of hbdgbhogs and gvmnuobs (Mammalia: Eoinacbomoopha) as infboobd foom a multigbnb studv: Evolutionaov Histoov of Eoinacbidab. Biological Journal of the Linnean Society 2014; 112: 499 - 519. hups: // doi. oog / 10.1111 / bij. 12299" type="journal article" year="2014">
Bannikova
<emphasis id="B951EA93FF97FFC3FF746459E331FD86" box="[253,299,577,601]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF97FFC3FF086460E350FCB2" pageId="18" pageNumber="19" type="etymology">
<paragraph id="8B9A3681FF97FFC3FF086460E350FCB2" blockId="18.[129,778,635,878]" pageId="18" pageNumber="19">
<emphasis id="B951EA93FF97FFC3FF086460E2EAFD4C" box="[129,240,635,659]" italics="true" pageId="18" pageNumber="19">Etymology:</emphasis>
The specific name ‘macarong’, which means vampire in Vietnamese (Ma cà rồng), acknowledges the long fangs (first upper incisors) that characterize mature males of this species. We suggest the common names Dalat Gymnure, Chuột Voi Đà Lạt, and Gimnuro de Dalat, in English, Vietnamese and Spanish, respectively, given that this species is just currently known from the Da Lat and
<collectingRegion id="49E1F863FF97FFC3FEAA652DE361FC91" box="[291,379,822,846]" country="Vietnam" name="Dak Lak" pageId="18" pageNumber="19">Dak Lak</collectingRegion>
Plateaus and surrounding lowlands, in southern Viet Nam.
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF97FFC3FF086592E71DFB4C" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="8B9A3681FF97FFC3FF086592E0B4F94F" blockId="18.[128,779,904,1680]" pageId="18" pageNumber="19">
<emphasis id="B951EA93FF97FFC3FF086592E2FDFC7F" box="[129,231,905,928]" italics="true" pageId="18" pageNumber="19">Diagnosis:</emphasis>
A medium-sized
<taxonomicName id="4C254D02FF97FFC3FE2B6593E3EDFC7F" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[418,503,904,928]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B951EA93FF97FFC3FE2B6593E3EDFC7F" box="[418,503,904,928]" italics="true" pageId="18" pageNumber="19">Hylomys</emphasis>
</taxonomicName>
(average HB =
<quantity id="4CDD9B64FF97FFC3FD156593E11DFC40" box="[668,775,904,928]" metricMagnitude="-1" metricUnit="m" metricValue="1.384" pageId="18" pageNumber="19" unit="mm" value="138.4">138.4 mm</quantity>
, GLS =
<quantity id="4CDD9B64FF97FFC3FF4465BCE331FC60" box="[205,299,935,959]" metricMagnitude="-2" metricUnit="m" metricValue="3.4799999999999995" pageId="18" pageNumber="19" unit="mm" value="34.8">34.8 mm</quantity>
) characterized by a rusty chest coloration in males (
<figureCitation id="131E2A04FF97FFC3FF5865DCE304FC01" box="[209,286,967,991]" captionStart="Figure 7" captionStartId="16.[130,195,874,898]" captionTargetBox="[129,1471,144,847]" captionTargetId="figure-556@16.[129,1473,144,847]" captionTargetPageId="16" captionText="Figure 7. Dorsal and ventral views of the skins of (A) Hylomys peguensis (USNM583813, neotype), and (B) Hylomys macarong(USNM 320500, holotype). Photo credits:Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510277" httpUri="https://zenodo.org/record/14510277/files/figure.png" pageId="18" pageNumber="19">Fig. 7B</figureCitation>
; possibly just present during spring/summer, since specimens collected in April and possibly winter (inferred due to long, soft, and thick pelage), lacked this coloration), bicoloured and long tail (average T =
<quantity id="4CDD9B64FF97FFC3FE7A623EE04BFBE3" box="[499,593,1061,1085]" metricMagnitude="-2" metricUnit="m" metricValue="2.32" pageId="18" pageNumber="19" unit="mm" value="23.2">23.2 mm</quantity>
, T/HB = 16.8%), light brown feet, short hindfoot (average HF =
<quantity id="4CDD9B64FF97FFC3FD17625FE11CFB84" box="[670,774,1092,1116]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="18" pageNumber="19" unit="mm" value="23.5">23.5 mm</quantity>
, HF + nail =
<quantity id="4CDD9B64FF97FFC3FE8A6278E37DFBA4" box="[259,359,1123,1147]" metricMagnitude="-2" metricUnit="m" metricValue="2.57" pageId="18" pageNumber="19" unit="mm" value="25.7">25.7 mm</quantity>
), ears are uniformly brown-coloured and relatively long (average E =
<quantity id="4CDD9B64FF97FFC3FE566298E058FB45" box="[479,578,1155,1179]" metricMagnitude="-2" metricUnit="m" metricValue="1.81" pageId="18" pageNumber="19" unit="mm" value="18.1">18.1 mm</quantity>
, E/GLS = 52.6%). Its skull is characterized by: obtuse-angled notch between premaxilla tips; narrow rostrum (average ROB =
<quantity id="4CDD9B64FF97FFC3FDF162DAE0DEFB06" box="[632,708,1217,1241]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="18" pageNumber="19" unit="mm" value="5.2">5.2 mm</quantity>
); long nasals (average NL =
<quantity id="4CDD9B64FF97FFC3FEEB62FAE3AAFB27" box="[354,432,1249,1273]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="18" pageNumber="19" unit="mm" value="13.0">13 mm</quantity>
; NL/GLS = 37%) that extend to the level of the antorbital rim; highly to moderately developed supraorbital processes; conspicuous and posteriorly expanded horizontal crest behind infraorbital canal; alisphenoid canal intermediate in length among other
<taxonomicName id="4C254D02FF97FFC3FD8A6345E042FAA9" authorityName="Hincklbv &amp; Camacho-Sanchbz &amp; Chua &amp; Rubdi &amp; Lundb &amp; Maldonado &amp; Omao &amp; Lbonaod &amp; Hawkins" authorityYear="2024" box="[515,600,1374,1398]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B951EA93FF97FFC3FD8A6345E042FAA9" box="[515,600,1374,1398]" italics="true" pageId="18" pageNumber="19">Hylomys</emphasis>
</taxonomicName>
species (average ASC =
<quantity id="4CDD9B64FF97FFC3FF466366E301FA4A" box="[207,283,1405,1429]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="mm" value="0.5">0.5 mm</quantity>
); interior side of ectotympanic anterior process elongated and greatly expanded anteriorly; deep braincase at the occipitum (average BD =
<quantity id="4CDD9B64FF97FFC3FE1B63A7E3F9FA0C" box="[402,483,1468,1492]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="18" pageNumber="19" unit="mm" value="9.7">9.7 mm</quantity>
); robust mandibular ramus (average LAP =
<quantity id="4CDD9B64FF97FFC3FEA563C0E390FA2C" box="[300,394,1499,1523]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="18" pageNumber="19" unit="mm" value="10.1">10.1 mm</quantity>
) with broad coronoid process; very long first upper incisor (average I1 =
<quantity id="4CDD9B64FF97FFC3FD8463E0E044F9CD" box="[525,606,1531,1555]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="18" pageNumber="19" unit="mm" value="3.3">3.3 mm</quantity>
), particularly in males (average I1 =
<quantity id="4CDD9B64FF97FFC3FEC76001E386F9EE" box="[334,412,1562,1586]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="18" pageNumber="19" unit="mm" value="3.6">3.6 mm</quantity>
males,
<quantity id="4CDD9B64FF97FFC3FE616001E02CF9EE" box="[488,566,1562,1586]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="18" pageNumber="19" unit="mm" value="2.9">2.9 mm</quantity>
females; Supporting Information,
<figureCitation id="131E2A04FF97FFC3FE8E6022E356F98E" box="[263,332,1593,1617]" captionStart="Figure 7" captionStartId="16.[130,195,874,898]" captionTargetBox="[129,1471,144,847]" captionTargetId="figure-556@16.[129,1473,144,847]" captionTargetPageId="16" captionText="Figure 7. Dorsal and ventral views of the skins of (A) Hylomys peguensis (USNM583813, neotype), and (B) Hylomys macarong(USNM 320500, holotype). Photo credits:Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510277" httpUri="https://zenodo.org/record/14510277/files/figure.png" pageId="18" pageNumber="19">Fig. S7</figureCitation>
), which is slightly projected towards lingual side, so that space between incisors generally narrows towards the tip; P4 labial cingulum absent or poorly developed.
</paragraph>
<paragraph id="8B9A3681FF97FFC3FF0860A9E180FCA7" blockId="18.[129,778,1713,1988]" lastBlockId="18.[825,1475,144,1171]" pageId="18" pageNumber="19">
<emphasis id="B951EA93FF97FFC3FF0860A9E312F916" box="[129,264,1714,1737]" italics="true" pageId="18" pageNumber="19">Comparisons:</emphasis>
In summer, easily distinguished from its closest geographical and genetic relative,
<taxonomicName id="4C254D02FF97FFC3FE5D60CAE056F937" authorityName="Blvth" authorityYear="1859" box="[468,588,1745,1768]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF97FFC3FE5D60CAE056F937" box="[468,588,1745,1768]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
</taxonomicName>
, by its rusty venter coloration (in males), which contrasts with
<taxonomicName id="4C254D02FF97FFC3FDC760EAE0D0F8D7" authorityName="Blvth" authorityYear="1859" box="[590,714,1777,1800]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF97FFC3FDC760EAE0D0F8D7" box="[590,714,1777,1800]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
</taxonomicName>
’s pale grey coloration (
<figureCitation id="131E2A04FF97FFC3FEB96114E370F8F8" box="[304,362,1807,1831]" captionStart="Figure 7" captionStartId="16.[130,195,874,898]" captionTargetBox="[129,1471,144,847]" captionTargetId="figure-556@16.[129,1473,144,847]" captionTargetPageId="16" captionText="Figure 7. Dorsal and ventral views of the skins of (A) Hylomys peguensis (USNM583813, neotype), and (B) Hylomys macarong(USNM 320500, holotype). Photo credits:Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510277" httpUri="https://zenodo.org/record/14510277/files/figure.png" pageId="18" pageNumber="19">Fig. 7</figureCitation>
). White rim in ear is absent vs. present in
<taxonomicName id="4C254D02FF97FFC3FF176134E301F899" authorityName="Blvth" authorityYear="1859" box="[158,283,1839,1862]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF97FFC3FF176134E301F899" box="[158,283,1839,1862]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
</taxonomicName>
(except in Cambodian specimen). Rostrum is longer (average =
<quantity id="4CDD9B64FF97FFC3FEB56155E382F8BA" box="[316,408,1870,1894]" metricMagnitude="-2" metricUnit="m" metricValue="1.51" pageId="18" pageNumber="19" unit="mm" value="15.1">15.1 mm</quantity>
) and narrower (average =
<quantity id="4CDD9B64FF97FFC3FD266155E0E5F8BA" box="[687,767,1870,1894]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="18" pageNumber="19" unit="mm" value="5.2">5.2 mm</quantity>
) than
<taxonomicName id="4C254D02FF97FFC3FF326175E320F85A" authorityName="Blvth" authorityYear="1859" box="[187,314,1902,1925]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF97FFC3FF326175E320F85A" box="[187,314,1902,1925]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
</taxonomicName>
(average =
<quantity id="4CDD9B64FF97FFC3FE316176E006F85A" box="[440,540,1901,1925]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="18" pageNumber="19" unit="mm" value="14.6">14.6 mm</quantity>
,
<quantity id="4CDD9B64FF97FFC3FDA56176E065F85A" box="[556,639,1901,1925]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="18" pageNumber="19" unit="mm" value="5.5">5.5 mm</quantity>
). Nasals are longer (average =
<quantity id="4CDD9B64FF97FFC3FEB46196E392F87B" box="[317,392,1933,1957]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="18" pageNumber="19" unit="mm" value="13.0">13 mm</quantity>
) vs. shorter (average =
<quantity id="4CDD9B64FF97FFC3FD0D6196E0F8F87B" box="[644,738,1933,1957]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="18" pageNumber="19" unit="mm" value="11.8">11.8 mm</quantity>
) in
<taxonomicName id="4C254D02FF97FFC3FF0861B6E2E0F81B" authorityName="Blvth" authorityYear="1859" box="[129,250,1965,1988]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF97FFC3FF0861B6E2E0F81B" box="[129,250,1965,1988]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
</taxonomicName>
. Conspicuous and posteriorly expanded horizontal crest behind infraorbital canal vs. less prominent and expanded in
<taxonomicName id="4C254D02FF97FFC3FCDF66ABE1CBFF18" authorityName="Blvth" authorityYear="1859" box="[854,977,176,199]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF97FFC3FCDF66ABE1CBFF18" box="[854,977,176,199]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
</taxonomicName>
. Alisphenoid canal shorter (average =
<quantity id="4CDD9B64FF97FFC3FAEE66B4E7ADFF18" box="[1383,1463,175,199]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="mm" value="0.5">0.5 mm</quantity>
) vs. longer (average =
<quantity id="4CDD9B64FF97FFC3FB9466D4E647FF39" box="[1053,1117,207,231]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="mm" value="1.0">1 mm</quantity>
) in
<emphasis id="B951EA93FF97FFC3FB1B66D4E70CFF39" box="[1170,1302,207,230]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C254D02FF97FFC3FB1B66D4E708FF39" authorityName="Blvth" authorityYear="1859" box="[1170,1298,207,230]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">H. peguensis</taxonomicName>
.
</emphasis>
Interior side of ectotympanic anterior process elongated and greatly expanded anteriorly vs. not expanded in
<taxonomicName id="4C254D02FF97FFC3FB016715E71CFEFA" authorityName="Blvth" authorityYear="1859" box="[1160,1286,270,293]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF97FFC3FB016715E71CFEFA" box="[1160,1286,270,293]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
</taxonomicName>
. Braincase at the occipitum is deeper (
<quantity id="4CDD9B64FF97FFC3FB936736E6B1FE9A" box="[1050,1195,301,325]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" metricValueMax="10.2" metricValueMin="9.4" pageId="18" pageNumber="19" unit="mm" value="9.8" valueMax="10.2" valueMin="9.4">9.4–10.2 mm</quantity>
) vs. flatter (
<quantity id="4CDD9B64FF97FFC3FAA26736E787FE9A" box="[1323,1437,301,325]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" metricValueMax="9.6" metricValueMin="9.0" pageId="18" pageNumber="19" unit="mm" value="9.3" valueMax="9.6" valueMin="9.0">9–9.6 mm</quantity>
) in
<taxonomicName id="4C254D02FF97FFC3FCB06756E1ACFEBB" authorityName="Blvth" authorityYear="1859" box="[825,950,333,356]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF97FFC3FCB06756E1ACFEBB" box="[825,950,333,356]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
</taxonomicName>
. Mandibular ramus broader (average LAP =
<quantity id="4CDD9B64FF97FFC3FA1C6757E17EFE5C" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="18" pageNumber="19" unit="mm" value="10.1">10.1 mm</quantity>
) vs. narrower (average LAP =
<quantity id="4CDD9B64FF97FFC3FB2F6770E6ECFE5C" box="[1190,1270,363,387]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="18" pageNumber="19" unit="mm" value="9.1">9.1 mm</quantity>
) in
<taxonomicName id="4C254D02FF97FFC3FAAC6777E7BAFE5C" authorityName="Blvth" authorityYear="1859" box="[1317,1440,364,387]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF97FFC3FAAC6777E7BAFE5C" box="[1317,1440,364,387]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
</taxonomicName>
, in fact, these species can be diagnosed through the combination of angular process length and rostral breadth (Supporting information, File S1,
<figureCitation id="131E2A04FF97FFC3FC4967D2E61CFE3E" box="[960,1030,457,481]" captionStart="Figure 8" captionStartId="17.[113,178,1610,1634]" captionTargetBox="[114,1458,144,1583]" captionTargetId="figure-160@17.[114,1458,144,1583]" captionTargetPageId="17" captionText="Figure 8. Skulls of (A) Hylomys peguensis (USNM583813, neotype) and (B) Hylomys macarong (USNM 320500, holotype).All images are at the same scale.Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510281" httpUri="https://zenodo.org/record/14510281/files/figure.png" pageId="18" pageNumber="19">Fig. S8</figureCitation>
). Coronoid process is broader vs. narrower in
<taxonomicName id="4C254D02FF97FFC3FCD867F2E1D2FDDF" authorityName="Blvth" authorityYear="1859" box="[849,968,489,512]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF97FFC3FCD867F2E1D2FDDF" box="[849,968,489,512]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
</taxonomicName>
, making the space between the distal region of coronoid and condylar processes frequently more narrow/acute in
<taxonomicName id="4C254D02FF97FFC3FCB06433E1ADFDE0" authorityName="Hinckley, Lunde &amp; Hawkins" authorityYear="2024" box="[825,951,552,575]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="macarong">
<emphasis id="B951EA93FF97FFC3FCB06433E1ADFDE0" box="[825,951,552,575]" italics="true" pageId="18" pageNumber="19">H. macarong</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C254D02FF97FFC3FB826433E69EFDE0" authorityName="Blvth" authorityYear="1859" box="[1035,1156,552,575]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF97FFC3FB826433E69EFDE0" box="[1035,1156,552,575]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
</taxonomicName>
(
<figureCitation id="131E2A04FF97FFC3FB1D643CE6D7FDE0" box="[1172,1229,551,575]" captionStart="Figure 8" captionStartId="17.[113,178,1610,1634]" captionTargetBox="[114,1458,144,1583]" captionTargetId="figure-160@17.[114,1458,144,1583]" captionTargetPageId="17" captionText="Figure 8. Skulls of (A) Hylomys peguensis (USNM583813, neotype) and (B) Hylomys macarong (USNM 320500, holotype).All images are at the same scale.Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510281" httpUri="https://zenodo.org/record/14510281/files/figure.png" pageId="18" pageNumber="19">Fig. 8</figureCitation>
). I1 sexually dimorphic and longer (males: 3.6,
<quantity id="4CDD9B64FF97FFC3FBAF645DE6B9FD81" box="[1062,1187,582,606]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="4.5" metricValueMin="2.6" pageId="18" pageNumber="19" unit="mm" value="3.55" valueMax="4.5" valueMin="2.6">2.6–4.5 mm</quantity>
, females: 2.9,
<quantity id="4CDD9B64FF97FFC3FAB9645DE7A0FD80" box="[1328,1466,582,607]" metricMagnitude="-3" metricUnit="m" metricValue="2.875" metricValueMax="3.15" metricValueMin="2.6" pageId="18" pageNumber="19" unit="mm" value="2.875" valueMax="3.15" valueMin="2.6">2.6–3.15 mm</quantity>
) vs. sexually monomorphic and shorter (males: 2.6,
<quantity id="4CDD9B64FF97FFC3FAD8647DE7A4FDA2" box="[1361,1470,614,638]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="3.0" metricValueMin="2.1" pageId="18" pageNumber="19" unit="mm" value="2.55" valueMax="3.0" valueMin="2.1">2.1–3 mm</quantity>
, females: 2.6,
<quantity id="4CDD9B64FF97FFC3FC4A649EE659FD42" box="[963,1091,645,669]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.1" metricValueMin="2.4" pageId="18" pageNumber="19" unit="mm" value="2.75" valueMax="3.1" valueMin="2.4">2.4–3.1 mm</quantity>
) in
<emphasis id="B951EA93FF97FFC3FBE7649DE6F5FD42" box="[1134,1263,646,669]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C254D02FF97FFC3FBE7649DE6F1FD42" authorityName="Blvth" authorityYear="1859" box="[1134,1259,646,669]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">H. peguensis</taxonomicName>
.
</emphasis>
I1 generally slightly projected towards lingual side, so that space between incisors generally narrows towards the tip vs. I1 slightly projected towards labial side, so that space between incisors does not narrow towards the tip in
<taxonomicName id="4C254D02FF97FFC3FC726518E66CFCC5" authorityName="Blvth" authorityYear="1859" box="[1019,1142,771,794]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF97FFC3FC726518E66CFCC5" box="[1019,1142,771,794]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
</taxonomicName>
. I2 projected posterolabially vs. posteriorly in
<taxonomicName id="4C254D02FF97FFC3FC586538E654FCE5" authorityName="Blvth" authorityYear="1859" box="[977,1102,803,826]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF97FFC3FC586538E654FCE5" box="[977,1102,803,826]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
</taxonomicName>
. P4 labial cingulum absent/poorly developed vs. generally well developed/conspicuous in
<emphasis id="B951EA93FF97FFC3FA206559E180FCA7" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C254D02FF97FFC3FA206559E18FFCA7" authorityName="Blvth" authorityYear="1859" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">H. peguensis</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8B9A3681FF97FFC3FCDC659BE71DFB4C" blockId="18.[825,1475,144,1171]" pageId="18" pageNumber="19">
It is currently thought to have a highly disjunct allopatric distribution with Malayan
<taxonomicName id="4C254D02FF97FFC3FBC465BBE685FC68" authorityName="Sody" authorityYear="1933" box="[1101,1183,928,951]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF97FFC3FBC465BBE685FC68" box="[1101,1183,928,951]" italics="true" pageId="18" pageNumber="19">H. maxi</emphasis>
</taxonomicName>
, from which it can be easily distinguished based on its shorter rostrum (RL: 14.6–15.7 vs.
<quantity id="4CDD9B64FF97FFC3FCE965C5E61AFC29" box="[864,1024,990,1014]" metricMagnitude="-2" metricUnit="m" metricValue="1.7149999999999999" metricValueMax="1.8199999999999998" metricValueMin="1.61" pageId="18" pageNumber="19" unit="mm" value="17.15" valueMax="18.2" valueMin="16.1">16.1–18.2 mm</quantity>
), upper toothrow (IM3SB: 16.1–17.4 vs.
<quantity id="4CDD9B64FF97FFC3FCB065E6E1CEFBCA" box="[825,980,1021,1045]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" metricValueMax="1.96" metricValueMin="1.76" pageId="18" pageNumber="19" unit="mm" value="18.6" valueMax="19.6" valueMin="17.6">17.6–19.6 mm</quantity>
) and molars (P4M3: 7.6–8.3 vs.
<quantity id="4CDD9B64FF97FFC3FABD65E6E7ADFBCA" box="[1332,1463,1021,1045]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" metricValueMax="9.9" metricValueMin="8.3" pageId="18" pageNumber="19" unit="mm" value="9.1" valueMax="9.9" valueMin="8.3">8.3–9.9 mm</quantity>
). Additionally, it can also be diagnosed from
<taxonomicName id="4C254D02FF97FFC3FA8A6206E74CFBEB" authorityName="Sody" authorityYear="1933" box="[1283,1366,1053,1076]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="maxi">
<emphasis id="B951EA93FF97FFC3FA8A6206E74CFBEB" box="[1283,1366,1053,1076]" italics="true" pageId="18" pageNumber="19">H. maxi</emphasis>
</taxonomicName>
due to its generally longer first incisor and through the combination of angular process length and rostral breadth (as with
<taxonomicName id="4C254D02FF97FFC3FAB66247E7A1FBAC" authorityName="Blvth" authorityYear="1859" box="[1343,1467,1116,1139]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF97FFC3FAB66247E7A1FBAC" box="[1343,1467,1116,1139]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
</taxonomicName>
; Supporting information, File S1,
<figureCitation id="131E2A04FF97FFC3FB006261E6C8FB4C" box="[1161,1234,1146,1171]" captionStart="Figure 7" captionStartId="16.[130,195,874,898]" captionTargetBox="[129,1471,144,847]" captionTargetId="figure-556@16.[129,1473,144,847]" captionTargetPageId="16" captionText="Figure 7. Dorsal and ventral views of the skins of (A) Hylomys peguensis (USNM583813, neotype), and (B) Hylomys macarong(USNM 320500, holotype). Photo credits:Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510277" httpUri="https://zenodo.org/record/14510277/files/figure.png" pageId="18" pageNumber="19">Figs S7</figureCitation>
, S
<figureCitation id="131E2A04FF97FFC3FB626260E6EDFB4C" box="[1259,1271,1147,1171]" captionStart="Figure 8" captionStartId="17.[113,178,1610,1634]" captionTargetBox="[114,1458,144,1583]" captionTargetId="figure-160@17.[114,1458,144,1583]" captionTargetPageId="17" captionText="Figure 8. Skulls of (A) Hylomys peguensis (USNM583813, neotype) and (B) Hylomys macarong (USNM 320500, holotype).All images are at the same scale.Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510281" httpUri="https://zenodo.org/record/14510281/files/figure.png" pageId="18" pageNumber="19">8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF97FFC2FCB062AFE0C6FDDE" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8B9A3681FF97FFC3FCB062AFE180F8A2" blockId="18.[825,1476,1204,1980]" pageId="18" pageNumber="19">
<emphasis id="B951EA93FF97FFC3FCB062AFE6D4FB13" box="[825,1230,1204,1228]" italics="true" pageId="18" pageNumber="19">Distribution, habitat, and natural history:</emphasis>
Currently known from the Langbian/Da Lat and
<collectingRegion id="49E1F863FF97FFC3FBDA62C8E6B4FB34" box="[1107,1198,1235,1259]" country="Vietnam" name="Dak Lak" pageId="18" pageNumber="19">Dak Lak</collectingRegion>
Plateaus and surrounding lowlands, in southern Viet Nam. Collected at
<emphasis id="B951EA93FF97FFC3FA8162E8E70DFAD5" box="[1288,1303,1267,1290]" italics="true" pageId="18" pageNumber="19">c.</emphasis>
<quantity id="4CDD9B64FF97FFC3FA9562E9E762FAD5" box="[1308,1400,1266,1290]" metricMagnitude="1" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="18" pageNumber="19" unit="m" value="65.0" valueMax="80.0" valueMin="50.0">50–80 m</quantity>
a.s.l. in semi-deciduous dipterocarp forest with extremely dense understory with shrubs, saplings, and lianas; at a site at
<quantity id="4CDD9B64FF97FFC3FAB6632AE78AFA96" box="[1343,1424,1329,1353]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="18" pageNumber="19" unit="m" value="1700.0">1700 m</quantity>
a.s.l. containing both pine savanna (dominated by
<taxonomicName id="4C254D02FF97FFC3FA82634AE799FAB7" box="[1291,1411,1360,1384]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="kesiya">
<emphasis id="B951EA93FF97FFC3FA82634AE799FAB7" box="[1291,1411,1360,1384]" italics="true" pageId="18" pageNumber="19">Pinus kesiya</emphasis>
</taxonomicName>
) with a dense grass cover and a tract of primary evergreen hill forest (
<bibRefCitation id="EFB44B70FF97FFC3FCCD6394E1F4FA78" author="Adlbo GH &amp; Suntsova NI &amp; Suntsov VV" box="[836,1006,1423,1447]" pageId="18" pageNumber="19" pagination="210 - 3" refId="ref30485" refString="Adlbo GH, Suntsova NI, Suntsov VV et al. Flbas (Siphonaptboa) collbctbd foom small mammals in Southbon Vibt Nam in 1997 - 1998. Journal of Medical Entomology 2001; 38: 210 - 3. hups: // doi. oog / 10.1603 / 0022 - 2585 - 38.2.210" type="journal article" year="2001">
Adler
<emphasis id="B951EA93FF97FFC3FC0D638BE1AEFA78" box="[900,948,1423,1447]" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2001
</bibRefCitation>
); and at a Dalat pine forest at
<emphasis id="B951EA93FF97FFC3FAA2638BE720FA78" box="[1323,1338,1424,1447]" italics="true" pageId="18" pageNumber="19">c.</emphasis>
<quantity id="4CDD9B64FF97FFC3FAC96394E78AFA78" box="[1344,1424,1423,1447]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="18" pageNumber="19" unit="m" value="1520.0">1520 m</quantity>
a.s.l. (USNM 269789; J.F. Rock). Specimens were collected under log in grass (
<emphasis id="B951EA93FF97FFC3FC1D63D5E1BDFA3A" box="[916,935,1486,1509]" italics="true" pageId="18" pageNumber="19">N</emphasis>
= 6), grass (
<emphasis id="B951EA93FF97FFC3FBA163D4E621FA39" box="[1064,1083,1487,1510]" italics="true" pageId="18" pageNumber="19">N</emphasis>
= 5), grass area in wood edge (
<emphasis id="B951EA93FF97FFC3FAFD63D4E79DFA39" box="[1396,1415,1487,1510]" italics="true" pageId="18" pageNumber="19">N</emphasis>
= 5), under log (
<emphasis id="B951EA93FF97FFC3FC2763F5E1DBF9DA" box="[942,961,1518,1541]" italics="true" pageId="18" pageNumber="19">N</emphasis>
= 1), under broadleaf herbaceous shrub near wet spot (
<emphasis id="B951EA93FF97FFC3FCFE6016E190F9FB" box="[887,906,1549,1572]" italics="true" pageId="18" pageNumber="19">N</emphasis>
= 1), and at base of
<taxonomicName id="4C254D02FF97FFC3FBDB6016E6A8F9FA" box="[1106,1202,1549,1573]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="genus">
<emphasis id="B951EA93FF97FFC3FBDB6016E6A8F9FA" box="[1106,1202,1549,1573]" italics="true" pageId="18" pageNumber="19">Pandanus</emphasis>
</taxonomicName>
in moist area (
<emphasis id="B951EA93FF97FFC3FACE6016E740F9FB" box="[1351,1370,1549,1572]" italics="true" pageId="18" pageNumber="19">N</emphasis>
= 1). Not recorded in bamboo patches or inside fagaceous forest, as with some sympatric rats, squirrels, and treeshrews (Feinstein’s field notes, USNM Mammal Division archives). Shares its habitat with
<taxonomicName id="4C254D02FF97FFC3FCE76091E61AF97E" box="[878,1024,1674,1698]" class="Mammalia" family="Soricidae" genus="Crocidura" kingdom="Animalia" order="Soricomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B951EA93FF97FFC3FCE76091E1CAF97D" box="[878,976,1674,1698]" italics="true" pageId="18" pageNumber="19">Crocidura</emphasis>
spp.
</taxonomicName>
,
<taxonomicName id="4C254D02FF97FFC3FB836091E6BEF97D" baseAuthorityName="Linnaeus" baseAuthorityYear="1766" box="[1034,1188,1674,1698]" class="Mammalia" family="Soricidae" genus="Suncus" kingdom="Animalia" order="Soricomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="murinus">
<emphasis id="B951EA93FF97FFC3FB836091E6BEF97D" box="[1034,1188,1674,1698]" italics="true" pageId="18" pageNumber="19">Suncus murinus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF97FFC3FB396090E74EF97D" baseAuthorityName="Wagner" baseAuthorityYear="1841" box="[1200,1364,1674,1698]" class="Mammalia" family="Tupaiidae" genus="Tupaia" kingdom="Animalia" order="Scandentia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="belangeri">
<emphasis id="B951EA93FF97FFC3FB396090E74EF97D" box="[1200,1364,1674,1698]" italics="true" pageId="18" pageNumber="19">Tupaia belangeri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF97FFC3FAD66091E16FF91E" class="Mammalia" family="Muridae" genus="Bandicota" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="saoilei">
<emphasis id="B951EA93FF97FFC3FAD66091E16FF91E" italics="true" pageId="18" pageNumber="19">Bandicota savilei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF97FFC3FC0160B2E64AF91E" baseAuthorityName="Blyth" baseAuthorityYear="1851" box="[904,1104,1705,1729]" class="Mammalia" family="Muridae" genus="Berylmys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="B951EA93FF97FFC3FC0160B2E64AF91E" box="[904,1104,1705,1729]" italics="true" pageId="18" pageNumber="19">Berylmys berdmorei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF97FFC3FBEB60B2E72EF91E" baseAuthorityName="Thomas" baseAuthorityYear="1916" box="[1122,1332,1705,1729]" class="Mammalia" family="Muridae" genus="Berylmys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="mackenziei">
<emphasis id="B951EA93FF97FFC3FBEB60B2E72EF91E" box="[1122,1332,1705,1729]" italics="true" pageId="18" pageNumber="19">Berylmys mackenziei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF97FFC3FACE60B2E186F93E" baseAuthorityName="H.C.Robinson &amp; Kloss" baseAuthorityYear="1922" class="Mammalia" family="Muridae" genus="Chiromyscus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="langbianis">
<emphasis id="B951EA93FF97FFC3FACE60B2E186F93E" italics="true" pageId="18" pageNumber="19">Chiromyscus langbianis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF97FFC3FC2360D2E675F93E" baseAuthorityName="Robinson and Kloss" baseAuthorityYear="1922" box="[938,1135,1737,1761]" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="milleti">
<emphasis id="B951EA93FF97FFC3FC2360D2E675F93E" box="[938,1135,1737,1761]" italics="true" pageId="18" pageNumber="19">Leopoldamys milleti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF97FFC3FBF460D2E710F93F" baseAuthorityName="Robinson &amp; Kloss" baseAuthorityYear="1922" box="[1149,1290,1737,1760]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="moi">
<emphasis id="B951EA93FF97FFC3FBF460D2E710F93F" box="[1149,1290,1737,1760]" italics="true" pageId="18" pageNumber="19">Maxomys moi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF97FFC3FA9E60D2E7A6F93E" box="[1303,1468,1737,1761]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="surifer">
<emphasis id="B951EA93FF97FFC3FA9E60D2E7A6F93E" box="[1303,1468,1737,1761]" italics="true" pageId="18" pageNumber="19">Maxomys surifer</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF97FFC3FCB060F2E1B9F8DF" authorityName="Bonhote" authorityYear="1902" box="[825,931,1768,1792]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="carolii">
<emphasis id="B951EA93FF97FFC3FCB060F2E1B9F8DF" box="[825,931,1768,1792]" italics="true" pageId="18" pageNumber="19">Mus carolii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF97FFC3FC2560F2E693F8DF" authority=", Nioioenter" authorityName="Nioioenter" box="[940,1161,1768,1792]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="pahari">
<emphasis id="B951EA93FF97FFC3FC2560F2E606F8DF" box="[940,1052,1768,1792]" italics="true" pageId="18" pageNumber="19">Mus pahari</emphasis>
,
<emphasis id="B951EA93FF97FFC3FBAD60F2E6FCF8DF" box="[1060,1254,1768,1792]" italics="true" pageId="18" pageNumber="19">
<emphasis id="B951EA93FF97FFC3FBAD60F2E693F8DF" box="[1060,1161,1769,1792]" italics="true" pageId="18" pageNumber="19">Niviventer</emphasis>
<emphasis id="B951EA93FF97FFC3FB0560F3E6FCF8DF" box="[1164,1254,1768,1792]" italics="true" pageId="18" pageNumber="19">mekongis</emphasis>
</emphasis>
</taxonomicName>
,
<taxonomicName id="4C254D02FF97FFC3FB6660F2E163F8C0" authority=", Ratus" authorityName="Ratus" genus="Ratus" pageId="18" pageNumber="19" rank="species" species="andamanensis">
<emphasis id="B951EA93FF97FFC3FB6660F2E7A6F8DF" box="[1263,1468,1768,1792]" italics="true" pageId="18" pageNumber="19">Rattus andamanensis</emphasis>
,
<emphasis id="B951EA93FF97FFC3FCB06113E1E0F8C0" box="[825,1018,1800,1823]" italics="true" pageId="18" pageNumber="19">
<emphasis id="B951EA93FF97FFC3FCB06113E163F8C0" box="[825,889,1800,1823]" italics="true" pageId="18" pageNumber="19">Rattus</emphasis>
<emphasis id="B951EA93FF97FFC3FCF46113E1E0F8C0" box="[893,1018,1800,1823]" italics="true" pageId="18" pageNumber="19">argentiventer</emphasis>
</emphasis>
</taxonomicName>
,
<emphasis id="B951EA93FF97FFC3FB8D6113E69AF8C0" box="[1028,1152,1800,1823]" italics="true" pageId="18" pageNumber="19">Rattus rattus</emphasis>
,
<taxonomicName id="4C254D02FF97FFC3FB00611CE755F8C0" baseAuthorityName="Bennett" baseAuthorityYear="1832" box="[1161,1359,1799,1823]" class="Mammalia" family="Muridae" genus="Vandeleuria" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="oleracea">
<emphasis id="B951EA93FF97FFC3FB00611CE755F8C0" box="[1161,1359,1799,1823]" italics="true" pageId="18" pageNumber="19">Vandeleuria oleracea</emphasis>
</taxonomicName>
,
<emphasis id="B951EA93FF97FFC3FAD06113E1A8F8E0" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C254D02FF97FFC3FAD06113E190F8E0" baseAuthorityName="Blanford" baseAuthorityYear="1878" class="Mammalia" family="Sciuridae" genus="Dremomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" sensu="lato" species="rufigenis">Dremomys rufigenis</taxonomicName>
<taxonomicNameLabel id="A26257E8FF97FFC3FC10613CE1A8F8E0" box="[921,946,1831,1855]" pageId="18" pageNumber="19" sensu="lato">s.l.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C254D02FF97FFC3FC73613CE6A3F8E0" baseAuthorityName="Blyth" baseAuthorityYear="1849" box="[1018,1209,1831,1855]" class="Mammalia" family="Sciuridae" genus="Menetes" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="B951EA93FF97FFC3FC73613CE6A3F8E0" box="[1018,1209,1831,1855]" italics="true" pageId="18" pageNumber="19">Menetes berdmorei</emphasis>
</taxonomicName>
(Feinstein’s field notes, USNM Mammal Division archives; USNM collection;
<bibRefCitation id="EFB44B70FF97FFC3FAF8615DE190F8A2" author="Adlbo GH &amp; Suntsova NI &amp; Suntsov VV" pageId="18" pageNumber="19" pagination="210 - 3" refId="ref30485" refString="Adlbo GH, Suntsova NI, Suntsov VV et al. Flbas (Siphonaptboa) collbctbd foom small mammals in Southbon Vibt Nam in 1997 - 1998. Journal of Medical Entomology 2001; 38: 210 - 3. hups: // doi. oog / 10.1603 / 0022 - 2585 - 38.2.210" type="journal article" year="2001">
Adler
<emphasis id="B951EA93FF97FFC3FA38615CE148F8A2" italics="true" pageId="18" pageNumber="19">et al.</emphasis>
2001
</bibRefCitation>
).
</paragraph>
<paragraph id="8B9A3681FF97FFC2FCDC619EE0C6FDDE" blockId="18.[825,1476,1204,1980]" lastBlockId="19.[113,763,144,513]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
A juvenile female and adult male were collected in the same trap during two consecutive days, perhaps suggesting parental care and/or male territory overlap with offspring at least until adulthood.Two juvenile female specimens were also collected
<quantity id="4CDD9B64FF96FFC2FD2966B4E0DCFF18" box="[672,710,175,199]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="19" pageNumber="20" unit="m" value="3.0">3 m</quantity>
away from each other during two consecutive days. Mature males have a reddish ventral coloration in July, which subadults and females lack, suggesting it might represent a sexual character. This coloration might last until late September, since one specimen (USNM 320485) was moulting it but still had enlarged testes (13 ×
<quantity id="4CDD9B64FF96FFC2FD316757E0E9FEBC" box="[696,755,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="20" unit="mm" value="6.0">6 mm</quantity>
; using
<taxonomicName id="4C254D02FF96FFC2FF276777E33CFE5C" authorityName="Blvth" authorityYear="1859" box="[174,294,364,387]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF96FFC2FF276777E33CFE5C" box="[174,294,364,387]" italics="true" pageId="19" pageNumber="20">H. peguensis</emphasis>
</taxonomicName>
specimens as a reference), perhaps suggesting end of breeding season. One female with six mammae was recorded by mid-December (USNM 320490), and juveniles have been recorded from mid-September to mid-October. Specimens were collected with snap traps baited with banana and manioc.
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF96FFC2FFF86439E0F5FBF0" pageId="19" pageNumber="20" type="conservation">
<paragraph id="8B9A3681FF96FFC2FFF86439E0F5FBF0" blockId="19.[113,763,546,1071]" pageId="19" pageNumber="20">
<emphasis id="B951EA93FF96FFC2FFF86439E2E3FDE6" box="[113,249,546,569]" italics="true" pageId="19" pageNumber="20">Conservation:</emphasis>
Recorded in Bu Gia Map and Cat Tien National Parks (
<bibRefCitation id="EFB44B70FF96FFC2FF35645AE381FD86" author="Bannikova AA &amp; Aboamov AV" box="[188,411,577,601]" pageId="19" pageNumber="20" pagination="499 - 519" refId="ref30844" refString="Bannikova AA, Lbbbdbv VS, Aboamov AV et al. Contoasting bvolutionaov histoov of hbdgbhogs and gvmnuobs (Mammalia: Eoinacbomoopha) as infboobd foom a multigbnb studv: Evolutionaov Histoov of Eoinacbidab. Biological Journal of the Linnean Society 2014; 112: 499 - 519. hups: // doi. oog / 10.1111 / bij. 12299" type="journal article" year="2014">
Bannikova
<emphasis id="B951EA93FF96FFC2FEA66459E37AFD86" box="[303,352,577,601]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2014
</bibRefCitation>
) and in proximity to Kalon Song Mao and Hon Ba Nature Reserves (Supporting information,
<tableCitation id="C6A7033AFF96FFC2FFF8649BE2D2FD47" box="[113,200,640,664]" captionStart="Table 1" captionStartId="11.[113,168,143,167]" captionTargetPageId="11" captionText="Table 1. Craniodental and selected external measurements (mm), weight (g) of adult Hylomys, including new species from Mount Leuser and the Da Lat–Dak Lak Plateaus. Mean, SD (w), ranges, and sample size (N) are specified for each measurement." httpUri="http://table.plazi.org/id/DF5A6609FF8EFFDAFFF86694E662FF1C" pageId="19" pageNumber="20" tableUuid="DF5A6609FF8EFFDAFFF86694E662FF1C">Table S1</tableCitation>
). Viet Nam now has the second-highest rate of deforestation in primary forests (
<bibRefCitation id="EFB44B70FF96FFC2FEFE6484E050FD68" author="Povaokov N &amp; Nguvbn T &amp; Popov E" box="[375,586,671,695]" pageId="19" pageNumber="20" pagination="255 - 476" refId="ref34856" refString="Povaokov N, Nguvbn T, Popov E et al. Rbcbnt poogobss in taxonomic studibs, biogbogoaphic analvsis, and obvisbd chbcklist of Rbptilbs in Indochina. Russian Journal of Herpetology 2023; 30: 255 - 476." type="journal article" year="2023">
Poyarkov
<emphasis id="B951EA93FF96FFC2FE5764BBE013FD68" box="[478,521,671,695]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
. 2023
</bibRefCitation>
). Co-distributed species with a potential similar extent of occurrence than that currently known for
<taxonomicName id="4C254D02FF96FFC2FEDD64C4E3C2FD29" authorityName="Hinckley, Lunde &amp; Hawkins" authorityYear="2024" box="[340,472,735,758]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="macarong">
<emphasis id="B951EA93FF96FFC2FEDD64C4E3C2FD29" box="[340,472,735,758]" italics="true" pageId="19" pageNumber="20">H. macarong</emphasis>
</taxonomicName>
, such as
<taxonomicName id="4C254D02FF96FFC2FDCA64C5E37AFCCA" authority="(Milnb-Edwaods 1871)" baseAuthorityName="Milnb-Edwaods" baseAuthorityYear="1871" class="Mammalia" family="Cercopithecidae" genus="Pygathrix" kingdom="Animalia" order="Primates" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="nigripes">
<emphasis id="B951EA93FF96FFC2FDCA64C5E0E0FD29" box="[579,762,734,758]" italics="true" pageId="19" pageNumber="20">Pygathrix nigripes</emphasis>
(Milne-Edwards 1871)
</taxonomicName>
,
<taxonomicName id="4C254D02FF96FFC2FEE264E6E0ECFCCA" authority="(Elliot 1909)" baseAuthorityName="Elliot" baseAuthorityYear="1909" box="[363,758,765,789]" class="Mammalia" family="Cercopithecidae" genus="Trachypithecus" kingdom="Animalia" order="Primates" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="margarita">
<emphasis id="B951EA93FF96FFC2FEE264E6E07FFCCA" box="[363,613,765,789]" italics="true" pageId="19" pageNumber="20">Trachypithecus margarita</emphasis>
(Elliot 1909)
</taxonomicName>
,
<taxonomicName id="4C254D02FF96FFC2FFF86506E3F5FCEA" authority="(Nomas 1909)" baseAuthorityName="Nomas" baseAuthorityYear="1909" box="[113,495,797,821]" class="Mammalia" family="Hylobatidae" genus="Nomascus" kingdom="Animalia" order="Primates" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="gabriellae">
<emphasis id="B951EA93FF96FFC2FFF86506E326FCEA" box="[113,316,797,821]" italics="true" pageId="19" pageNumber="20">Nomascus gabriellae</emphasis>
(Thomas 1909)
</taxonomicName>
, and
<taxonomicName id="4C254D02FF96FFC2FDA76506E392FC8B" authority="(Kouskop and Egbo 2008)" baseAuthorityName="Kouskop and Egbo" baseAuthorityYear="2008" class="Mammalia" family="Vespertilionidae" genus="Murina" kingdom="Animalia" order="Chiroptera" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="harpioloides">
<emphasis id="B951EA93FF96FFC2FDA76506E0E0FCEA" box="[558,762,797,821]" italics="true" pageId="19" pageNumber="20">Murina harpioloides</emphasis>
(Kruskop and Eger 2008)
</taxonomicName>
, are listed as endangered or critically endangered (IUCN Red List of Threatened Species 2023). Further research is needed to assess its tolerance to habitat degradation, and to evaluate whether this species is present in other potentially suitable protected areas, such as Ea So, Krong Trai, Nam Nung, and Nam Kar Nature Reserves, and Lo Go Xa Mat and Ta Dung National Parks, in Viet Nam, and Lumphat, Phnum Prech, and Phnom Namlear Wildlife Sanctuaries in
<collectingCountry id="F3327611FF96FFC2FDF5620CE0F0FBF0" box="[636,746,1047,1071]" name="Cambodia" pageId="19" pageNumber="20">Cambodia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33F650AFF96FFC2FFF8624AE3F1FB39" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8B9A3681FF96FFC2FFF8624AE3F1FB39" blockId="19.[113,763,1104,1254]" pageId="19" pageNumber="20">
<emphasis id="B951EA93FF96FFC2FFF8624AE2F8FBB7" box="[113,226,1105,1128]" italics="true" pageId="19" pageNumber="20">Comments:</emphasis>
Specimens from
<collectingRegion id="49E1F863FF96FFC2FE48624AE034FBB7" box="[449,558,1105,1128]" country="Vietnam" name="Kon Tum" pageId="19" pageNumber="20">Kon Tum</collectingRegion>
province, Central Highlands region of Viet Nam (MNHN1929320–5) have not been sequenced but were clustered in the morphospace of
<taxonomicName id="4C254D02FF96FFC2FD68628BE2D1FB19" authorityName="Blvth" authorityYear="1859" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF96FFC2FD68628BE2D1FB19" italics="true" pageId="19" pageNumber="20">H. peguensis</emphasis>
</taxonomicName>
and were thus assigned to
<taxonomicName id="4C254D02FF96FFC2FE5062B4E04BFB19" authorityName="Blvth" authorityYear="1859" box="[473,593,1199,1222]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B951EA93FF96FFC2FE5062B4E04BFB19" box="[473,593,1199,1222]" italics="true" pageId="19" pageNumber="20">H. peguensis</emphasis>
</taxonomicName>
in univariate descriptive statistics and bivariate plots.
</paragraph>
</subSubSection>
</treatment>
</document>
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