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, E/
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= 45%). Its skull can be distinguished from all relatives by its relatively larger molars (Supporting Information, File S1,
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<figureCitation id="131E2A04FF88FFDCFD1765ADE0F5FC11" box="[670,751,950,974]" captionStart="Figure 5" captionStartId="9.[113,178,1006,1030]" captionTargetBox="[113,1457,145,977]" captionTargetId="figure-518@9.[112,1459,144,979]" captionTargetPageId="9" captionText="Figure 5. Morphometric variation in Hylomys. A, first and second principal components from an analysis of 16 cranial dimensions, computed with Dataset 4 (with the exclusion of external measurements); B, second and third principal components of the same analysis.Asterisks denote undamaged holotype or neotype specimens that could be included in this analysis." pageId="13" pageNumber="14">Fig. S5</figureCitation>
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<figureCitation id="131E2A04FF88FFDCFD1765ADE0F5FC11" box="[670,751,950,974]" captionStart="Figure 5" captionStartId="9.[113,178,1006,1030]" captionTargetBox="[113,1457,145,977]" captionTargetId="figure-518@9.[112,1459,144,979]" captionTargetPageId="9" captionText="Figure 5. Morphometric variation in Hylomys. A, first and second principal components from an analysis of 16 cranial dimensions, computed with Dataset 4 (with the exclusion of external measurements); B, second and third principal components of the same analysis.Asterisks denote undamaged holotype or neotype specimens that could be included in this analysis." figureDoi="http://doi.org/10.5281/zenodo.14510273" httpUri="https://zenodo.org/record/14510273/files/figure.png" pageId="13" pageNumber="14">Fig. S5</figureCitation>
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) and from all relatives but allopatric
|
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<taxonomicName id="4C254D02FF88FFDCFE7A65CDE049FC31" authorityName="Mullbo" authorityYear="1840" box="[499,595,982,1006]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="suillus">
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<emphasis id="B951EA93FF88FFDCFE7A65CDE049FC31" box="[499,595,982,1006]" italics="true" pageId="13" pageNumber="14">H. suillus</emphasis>
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by its long and relatively broad rostrum (
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<figureCitation id="131E2A04FF88FFDCFE0E65EEE3D9FBD2" box="[391,451,1013,1037]" captionStart="Figure 6" captionStartId="10.[129,194,1089,1113]" captionTargetBox="[130,1472,168,1060]" captionTargetId="figure-528@10.[129,1473,144,1061]" captionTargetPageId="10" captionText="Figure 6. Bivariate plot of rostrum length (ROL) and rostrum breadth (ROB) measurements." pageId="13" pageNumber="14">Fig. 6</figureCitation>
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<figureCitation id="131E2A04FF88FFDCFE0E65EEE3D9FBD2" box="[391,451,1013,1037]" captionStart="Figure 6" captionStartId="10.[129,194,1089,1113]" captionTargetBox="[130,1472,168,1060]" captionTargetId="figure-528@10.[129,1473,144,1061]" captionTargetPageId="10" captionText="Figure 6. Bivariate plot of rostrum length (ROL) and rostrum breadth (ROB) measurements." figureDoi="http://doi.org/10.5281/zenodo.14510275" httpUri="https://zenodo.org/record/14510275/files/figure.png" pageId="13" pageNumber="14">Fig. 6</figureCitation>
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). Its skull can also be distinguished from all other relatives by the combination of the following characters: obtuse-angled notch between anterior side of premaxilla; posterior end of nasals does not extend to the level of the antorbital rim; dorsal region of maxillary and frontal have a rugged appearance with capillary grooves and pores; prominent antorbital ridge/flange; anterior side of the antorbital rim is posterior to second cusp of M1; robust supraorbital processes and dentition; short alisphenoid canal (average
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=
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@ -1,103 +1,106 @@
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<mods:namePart id="2EBF0547E761B953855EB5C55D496A6B">Leonard, Jennifer A.</mods:namePart>
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<mods:affiliation id="4D090B85A0D95A0D365C19EC29EA4F65">Conservation & Evolutionary Genetics Group, Estación Biológica de Doñana (EBD-CSIC), Seville, Spain</mods:affiliation>
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<taxonomicName id="4C254D02FF8BFFDFFEA16375E078FA56" authority="Blyth, 1859" authorityName="Blyth" authorityYear="1859" box="[296,610,1390,1417]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="peguensis">
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|
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@ -264,9 +267,9 @@ species allopatric distribution ranges do not span this region. The specimen con
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, weight
|
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<quantity id="4CDD9B64FF8BFFDFFAD862B8E798FB64" box="[1361,1410,1187,1211]" metricMagnitude="-2" metricUnit="kg" metricValue="7.3" pageId="14" pageNumber="15" unit="g" value="73.0">73 g</quantity>
|
||||
. Skull measurements in Supporting information, Table S2. 3D surface scans of this neotype skull are available at MorphoSource: https://doi.org/10.17602/M2/M560027 (cranium); https:// doi.org/10.17602/M2/M560031 (mandible). Pictures of this holotype skin and skull are shown in
|
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<figureCitation id="131E2A04FF8BFFDFFB326324E731FA88" box="[1211,1323,1343,1367]" captionStart="Figure 7" captionStartId="16.[130,195,874,898]" captionTargetBox="[129,1471,144,847]" captionTargetId="figure-556@16.[129,1473,144,847]" captionTargetPageId="16" captionText="Figure 7. Dorsal and ventral views of the skins of (A) Hylomys peguensis (USNM583813, neotype), and (B) Hylomys macarong(USNM 320500, holotype). Photo credits:Katie Sayers." pageId="14" pageNumber="15">Figures 7A</figureCitation>
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<figureCitation id="131E2A04FF8BFFDFFB326324E731FA88" box="[1211,1323,1343,1367]" captionStart="Figure 7" captionStartId="16.[130,195,874,898]" captionTargetBox="[129,1471,144,847]" captionTargetId="figure-556@16.[129,1473,144,847]" captionTargetPageId="16" captionText="Figure 7. Dorsal and ventral views of the skins of (A) Hylomys peguensis (USNM583813, neotype), and (B) Hylomys macarong(USNM 320500, holotype). Photo credits:Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510277" httpUri="https://zenodo.org/record/14510277/files/figure.png" pageId="14" pageNumber="15">Figures 7A</figureCitation>
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and
|
||||
<figureCitation id="131E2A04FF8BFFDFFAD66324E766FA88" box="[1375,1404,1343,1367]" captionStart="Figure 8" captionStartId="17.[113,178,1610,1634]" captionTargetBox="[114,1458,144,1583]" captionTargetId="figure-160@17.[114,1458,144,1583]" captionTargetPageId="17" captionText="Figure 8. Skulls of (A) Hylomys peguensis (USNM583813, neotype) and (B) Hylomys macarong (USNM 320500, holotype).All images are at the same scale.Photo credits: Katie Sayers." pageId="14" pageNumber="15">8A</figureCitation>
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<figureCitation id="131E2A04FF8BFFDFFAD66324E766FA88" box="[1375,1404,1343,1367]" captionStart="Figure 8" captionStartId="17.[113,178,1610,1634]" captionTargetBox="[114,1458,144,1583]" captionTargetId="figure-160@17.[114,1458,144,1583]" captionTargetPageId="17" captionText="Figure 8. Skulls of (A) Hylomys peguensis (USNM583813, neotype) and (B) Hylomys macarong (USNM 320500, holotype).All images are at the same scale.Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510281" httpUri="https://zenodo.org/record/14510281/files/figure.png" pageId="14" pageNumber="15">8A</figureCitation>
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. DNA sequences associated with this neotype have been deposited in
|
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GenBank:
|
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|
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@ -351,7 +354,7 @@ It is currently thought to have an allopatric distribution with Sumatran and mai
|
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<emphasis id="B951EA93FF8AFFDEFDA1647EE063FDA3" box="[552,633,613,636]" italics="true" pageId="15" pageNumber="16">H. maxi</emphasis>
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</taxonomicName>
|
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, from which it can be distinguished based on its generally lighter dorsum (brown with yellowish tinge vs. without), venter (buff vs. greybrown) and feet coloration (light vs. dark brown), particularly on its forefeet (light brown vs. dark brown/black), presence of whitish/pale brown in ear rim (except in Cambodian population), longer tail and shorter hindfoot (Supporting information, File S1,
|
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<figureCitation id="131E2A04FF8AFFDEFF4E6524E315FC88" box="[199,271,831,855]" captionStart="Figure 6" captionStartId="10.[129,194,1089,1113]" captionTargetBox="[130,1472,168,1060]" captionTargetId="figure-528@10.[129,1473,144,1061]" captionTargetPageId="10" captionText="Figure 6. Bivariate plot of rostrum length (ROL) and rostrum breadth (ROB) measurements." pageId="15" pageNumber="16">Fig. S6</figureCitation>
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<figureCitation id="131E2A04FF8AFFDEFF4E6524E315FC88" box="[199,271,831,855]" captionStart="Figure 6" captionStartId="10.[129,194,1089,1113]" captionTargetBox="[130,1472,168,1060]" captionTargetId="figure-528@10.[129,1473,144,1061]" captionTargetPageId="10" captionText="Figure 6. Bivariate plot of rostrum length (ROL) and rostrum breadth (ROB) measurements." figureDoi="http://doi.org/10.5281/zenodo.14510275" httpUri="https://zenodo.org/record/14510275/files/figure.png" pageId="15" pageNumber="16">Fig. S6</figureCitation>
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), shorter skull length (32.3–36.0 vs.
|
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), shorter rostrum length (
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|
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@ -671,7 +674,7 @@ Abramov
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Mammal Division archives).
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<emphasis id="B951EA93FF95FFC1FF0B6571E2C0FC5C" bold="true" box="[130,218,874,899]" pageId="16" pageNumber="17">Figure 7.</emphasis>
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Dorsal and ventral views of the skins of (A)
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|
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@ -166,16 +169,16 @@ on
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, ear length
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. Skull measurements in Supporting Information, Table S2. 3D surface scans of this holotype skull are available at MorphoSource: https://doi.org/10.17602/M2/M560006 (cranium); https://doi.org/10.17602/M2/M560011 (mandible). Pictures of this holotype skin and skull are shown in
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and
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<figureCitation id="131E2A04FF94FFC0FC7060F6E60CF8DA" box="[1017,1046,1773,1797]" captionStart="Figure 8" captionStartId="17.[113,178,1610,1634]" captionTargetBox="[114,1458,144,1583]" captionTargetId="figure-160@17.[114,1458,144,1583]" captionTargetPageId="17" captionText="Figure 8. Skulls of (A) Hylomys peguensis (USNM583813, neotype) and (B) Hylomys macarong (USNM 320500, holotype).All images are at the same scale.Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510281" httpUri="https://zenodo.org/record/14510281/files/figure.png" pageId="17" pageNumber="18">8B</figureCitation>
|
||||
. DNA sequences associated with this holotype have been deposited in GenBank:
|
||||
<accessionNumber id="9476AB62FF94FFC0FA846117E798F8FB" box="[1293,1410,1804,1828]" pageId="17" pageNumber="18">OR138081</accessionNumber>
|
||||
and
|
||||
<accessionNumber id="9476AB62FF94FFC0FCA36130E1B8F89C" box="[810,930,1835,1859]" pageId="17" pageNumber="18">OR554435</accessionNumber>
|
||||
.
|
||||
</paragraph>
|
||||
<caption id="DF5A6609FF94FFC0FFF86051E01CF9A1" pageId="17" pageNumber="18" startId="17.[113,178,1610,1634]" targetBox="[114,1458,144,1583]" targetPageId="17" targetType="figure">
|
||||
<caption id="DF5A6609FF94FFC0FFF86051E01CF9A1" ID-DOI="http://doi.org/10.5281/zenodo.14510281" ID-Zenodo-Dep="14510281" httpUri="https://zenodo.org/record/14510281/files/figure.png" pageId="17" pageNumber="18" startId="17.[113,178,1610,1634]" targetBox="[114,1458,144,1583]" targetPageId="17" targetType="figure">
|
||||
<paragraph id="8B9A3681FF94FFC0FFF86051E01CF9A1" blockId="17.[113,1438,1610,1662]" pageId="17" pageNumber="18">
|
||||
<emphasis id="B951EA93FF94FFC0FFF86051E2D3F9BD" bold="true" box="[113,201,1610,1634]" pageId="17" pageNumber="18">Figure 8.</emphasis>
|
||||
Skulls of (A)
|
||||
|
|
@ -299,7 +302,7 @@ A medium-sized
|
|||
, GLS =
|
||||
<quantity id="4CDD9B64FF97FFC3FF4465BCE331FC60" box="[205,299,935,959]" metricMagnitude="-2" metricUnit="m" metricValue="3.4799999999999995" pageId="18" pageNumber="19" unit="mm" value="34.8">34.8 mm</quantity>
|
||||
) characterized by a rusty chest coloration in males (
|
||||
<figureCitation id="131E2A04FF97FFC3FF5865DCE304FC01" box="[209,286,967,991]" captionStart="Figure 7" captionStartId="16.[130,195,874,898]" captionTargetBox="[129,1471,144,847]" captionTargetId="figure-556@16.[129,1473,144,847]" captionTargetPageId="16" captionText="Figure 7. Dorsal and ventral views of the skins of (A) Hylomys peguensis (USNM583813, neotype), and (B) Hylomys macarong(USNM 320500, holotype). Photo credits:Katie Sayers." pageId="18" pageNumber="19">Fig. 7B</figureCitation>
|
||||
<figureCitation id="131E2A04FF97FFC3FF5865DCE304FC01" box="[209,286,967,991]" captionStart="Figure 7" captionStartId="16.[130,195,874,898]" captionTargetBox="[129,1471,144,847]" captionTargetId="figure-556@16.[129,1473,144,847]" captionTargetPageId="16" captionText="Figure 7. Dorsal and ventral views of the skins of (A) Hylomys peguensis (USNM583813, neotype), and (B) Hylomys macarong(USNM 320500, holotype). Photo credits:Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510277" httpUri="https://zenodo.org/record/14510277/files/figure.png" pageId="18" pageNumber="19">Fig. 7B</figureCitation>
|
||||
; possibly just present during spring/summer, since specimens collected in April and possibly winter (inferred due to long, soft, and thick pelage), lacked this coloration), bicoloured and long tail (average T =
|
||||
<quantity id="4CDD9B64FF97FFC3FE7A623EE04BFBE3" box="[499,593,1061,1085]" metricMagnitude="-2" metricUnit="m" metricValue="2.32" pageId="18" pageNumber="19" unit="mm" value="23.2">23.2 mm</quantity>
|
||||
, T/HB = 16.8%), light brown feet, short hindfoot (average HF =
|
||||
|
|
@ -329,7 +332,7 @@ species (average ASC =
|
|||
males,
|
||||
<quantity id="4CDD9B64FF97FFC3FE616001E02CF9EE" box="[488,566,1562,1586]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="18" pageNumber="19" unit="mm" value="2.9">2.9 mm</quantity>
|
||||
females; Supporting Information,
|
||||
<figureCitation id="131E2A04FF97FFC3FE8E6022E356F98E" box="[263,332,1593,1617]" captionStart="Figure 7" captionStartId="16.[130,195,874,898]" captionTargetBox="[129,1471,144,847]" captionTargetId="figure-556@16.[129,1473,144,847]" captionTargetPageId="16" captionText="Figure 7. Dorsal and ventral views of the skins of (A) Hylomys peguensis (USNM583813, neotype), and (B) Hylomys macarong(USNM 320500, holotype). Photo credits:Katie Sayers." pageId="18" pageNumber="19">Fig. S7</figureCitation>
|
||||
<figureCitation id="131E2A04FF97FFC3FE8E6022E356F98E" box="[263,332,1593,1617]" captionStart="Figure 7" captionStartId="16.[130,195,874,898]" captionTargetBox="[129,1471,144,847]" captionTargetId="figure-556@16.[129,1473,144,847]" captionTargetPageId="16" captionText="Figure 7. Dorsal and ventral views of the skins of (A) Hylomys peguensis (USNM583813, neotype), and (B) Hylomys macarong(USNM 320500, holotype). Photo credits:Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510277" httpUri="https://zenodo.org/record/14510277/files/figure.png" pageId="18" pageNumber="19">Fig. S7</figureCitation>
|
||||
), which is slightly projected towards lingual side, so that space between incisors generally narrows towards the tip; P4 labial cingulum absent or poorly developed.
|
||||
</paragraph>
|
||||
<paragraph id="8B9A3681FF97FFC3FF0860A9E180FCA7" blockId="18.[129,778,1713,1988]" lastBlockId="18.[825,1475,144,1171]" pageId="18" pageNumber="19">
|
||||
|
|
@ -343,7 +346,7 @@ In summer, easily distinguished from its closest geographical and genetic relati
|
|||
<emphasis id="B951EA93FF97FFC3FDC760EAE0D0F8D7" box="[590,714,1777,1800]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
|
||||
</taxonomicName>
|
||||
’s pale grey coloration (
|
||||
<figureCitation id="131E2A04FF97FFC3FEB96114E370F8F8" box="[304,362,1807,1831]" captionStart="Figure 7" captionStartId="16.[130,195,874,898]" captionTargetBox="[129,1471,144,847]" captionTargetId="figure-556@16.[129,1473,144,847]" captionTargetPageId="16" captionText="Figure 7. Dorsal and ventral views of the skins of (A) Hylomys peguensis (USNM583813, neotype), and (B) Hylomys macarong(USNM 320500, holotype). Photo credits:Katie Sayers." pageId="18" pageNumber="19">Fig. 7</figureCitation>
|
||||
<figureCitation id="131E2A04FF97FFC3FEB96114E370F8F8" box="[304,362,1807,1831]" captionStart="Figure 7" captionStartId="16.[130,195,874,898]" captionTargetBox="[129,1471,144,847]" captionTargetId="figure-556@16.[129,1473,144,847]" captionTargetPageId="16" captionText="Figure 7. Dorsal and ventral views of the skins of (A) Hylomys peguensis (USNM583813, neotype), and (B) Hylomys macarong(USNM 320500, holotype). Photo credits:Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510277" httpUri="https://zenodo.org/record/14510277/files/figure.png" pageId="18" pageNumber="19">Fig. 7</figureCitation>
|
||||
). White rim in ear is absent vs. present in
|
||||
<taxonomicName id="4C254D02FF97FFC3FF176134E301F899" authorityName="Blvth" authorityYear="1859" box="[158,283,1839,1862]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
|
||||
<emphasis id="B951EA93FF97FFC3FF176134E301F899" box="[158,283,1839,1862]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
|
||||
|
|
@ -402,7 +405,7 @@ Interior side of ectotympanic anterior process elongated and greatly expanded an
|
|||
<emphasis id="B951EA93FF97FFC3FAAC6777E7BAFE5C" box="[1317,1440,364,387]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
|
||||
</taxonomicName>
|
||||
, in fact, these species can be diagnosed through the combination of angular process length and rostral breadth (Supporting information, File S1,
|
||||
<figureCitation id="131E2A04FF97FFC3FC4967D2E61CFE3E" box="[960,1030,457,481]" captionStart="Figure 8" captionStartId="17.[113,178,1610,1634]" captionTargetBox="[114,1458,144,1583]" captionTargetId="figure-160@17.[114,1458,144,1583]" captionTargetPageId="17" captionText="Figure 8. Skulls of (A) Hylomys peguensis (USNM583813, neotype) and (B) Hylomys macarong (USNM 320500, holotype).All images are at the same scale.Photo credits: Katie Sayers." pageId="18" pageNumber="19">Fig. S8</figureCitation>
|
||||
<figureCitation id="131E2A04FF97FFC3FC4967D2E61CFE3E" box="[960,1030,457,481]" captionStart="Figure 8" captionStartId="17.[113,178,1610,1634]" captionTargetBox="[114,1458,144,1583]" captionTargetId="figure-160@17.[114,1458,144,1583]" captionTargetPageId="17" captionText="Figure 8. Skulls of (A) Hylomys peguensis (USNM583813, neotype) and (B) Hylomys macarong (USNM 320500, holotype).All images are at the same scale.Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510281" httpUri="https://zenodo.org/record/14510281/files/figure.png" pageId="18" pageNumber="19">Fig. S8</figureCitation>
|
||||
). Coronoid process is broader vs. narrower in
|
||||
<taxonomicName id="4C254D02FF97FFC3FCD867F2E1D2FDDF" authorityName="Blvth" authorityYear="1859" box="[849,968,489,512]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="peguensis">
|
||||
<emphasis id="B951EA93FF97FFC3FCD867F2E1D2FDDF" box="[849,968,489,512]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
|
||||
|
|
@ -416,7 +419,7 @@ than in
|
|||
<emphasis id="B951EA93FF97FFC3FB826433E69EFDE0" box="[1035,1156,552,575]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="131E2A04FF97FFC3FB1D643CE6D7FDE0" box="[1172,1229,551,575]" captionStart="Figure 8" captionStartId="17.[113,178,1610,1634]" captionTargetBox="[114,1458,144,1583]" captionTargetId="figure-160@17.[114,1458,144,1583]" captionTargetPageId="17" captionText="Figure 8. Skulls of (A) Hylomys peguensis (USNM583813, neotype) and (B) Hylomys macarong (USNM 320500, holotype).All images are at the same scale.Photo credits: Katie Sayers." pageId="18" pageNumber="19">Fig. 8</figureCitation>
|
||||
<figureCitation id="131E2A04FF97FFC3FB1D643CE6D7FDE0" box="[1172,1229,551,575]" captionStart="Figure 8" captionStartId="17.[113,178,1610,1634]" captionTargetBox="[114,1458,144,1583]" captionTargetId="figure-160@17.[114,1458,144,1583]" captionTargetPageId="17" captionText="Figure 8. Skulls of (A) Hylomys peguensis (USNM583813, neotype) and (B) Hylomys macarong (USNM 320500, holotype).All images are at the same scale.Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510281" httpUri="https://zenodo.org/record/14510281/files/figure.png" pageId="18" pageNumber="19">Fig. 8</figureCitation>
|
||||
). I1 sexually dimorphic and longer (males: 3.6,
|
||||
<quantity id="4CDD9B64FF97FFC3FBAF645DE6B9FD81" box="[1062,1187,582,606]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="4.5" metricValueMin="2.6" pageId="18" pageNumber="19" unit="mm" value="3.55" valueMax="4.5" valueMin="2.6">2.6–4.5 mm</quantity>
|
||||
, females: 2.9,
|
||||
|
|
@ -464,9 +467,9 @@ due to its generally longer first incisor and through the combination of angular
|
|||
<emphasis id="B951EA93FF97FFC3FAB66247E7A1FBAC" box="[1343,1467,1116,1139]" italics="true" pageId="18" pageNumber="19">H. peguensis</emphasis>
|
||||
</taxonomicName>
|
||||
; Supporting information, File S1,
|
||||
<figureCitation id="131E2A04FF97FFC3FB006261E6C8FB4C" box="[1161,1234,1146,1171]" captionStart="Figure 7" captionStartId="16.[130,195,874,898]" captionTargetBox="[129,1471,144,847]" captionTargetId="figure-556@16.[129,1473,144,847]" captionTargetPageId="16" captionText="Figure 7. Dorsal and ventral views of the skins of (A) Hylomys peguensis (USNM583813, neotype), and (B) Hylomys macarong(USNM 320500, holotype). Photo credits:Katie Sayers." pageId="18" pageNumber="19">Figs S7</figureCitation>
|
||||
<figureCitation id="131E2A04FF97FFC3FB006261E6C8FB4C" box="[1161,1234,1146,1171]" captionStart="Figure 7" captionStartId="16.[130,195,874,898]" captionTargetBox="[129,1471,144,847]" captionTargetId="figure-556@16.[129,1473,144,847]" captionTargetPageId="16" captionText="Figure 7. Dorsal and ventral views of the skins of (A) Hylomys peguensis (USNM583813, neotype), and (B) Hylomys macarong(USNM 320500, holotype). Photo credits:Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510277" httpUri="https://zenodo.org/record/14510277/files/figure.png" pageId="18" pageNumber="19">Figs S7</figureCitation>
|
||||
, S
|
||||
<figureCitation id="131E2A04FF97FFC3FB626260E6EDFB4C" box="[1259,1271,1147,1171]" captionStart="Figure 8" captionStartId="17.[113,178,1610,1634]" captionTargetBox="[114,1458,144,1583]" captionTargetId="figure-160@17.[114,1458,144,1583]" captionTargetPageId="17" captionText="Figure 8. Skulls of (A) Hylomys peguensis (USNM583813, neotype) and (B) Hylomys macarong (USNM 320500, holotype).All images are at the same scale.Photo credits: Katie Sayers." pageId="18" pageNumber="19">8</figureCitation>
|
||||
<figureCitation id="131E2A04FF97FFC3FB626260E6EDFB4C" box="[1259,1271,1147,1171]" captionStart="Figure 8" captionStartId="17.[113,178,1610,1634]" captionTargetBox="[114,1458,144,1583]" captionTargetId="figure-160@17.[114,1458,144,1583]" captionTargetPageId="17" captionText="Figure 8. Skulls of (A) Hylomys peguensis (USNM583813, neotype) and (B) Hylomys macarong (USNM 320500, holotype).All images are at the same scale.Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510281" httpUri="https://zenodo.org/record/14510281/files/figure.png" pageId="18" pageNumber="19">8</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
|
|
|
|||
|
|
@ -1,103 +1,106 @@
|
|||
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<mods:namePart id="160EE409FA0BCFF01958D878FBF85332">Hinckley, Arlo</mods:namePart>
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<mods:affiliation id="6A0E76565970C7FA3620FB476DDFA37D">Division of Mammals, Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, District of Columbia, Washington, USA</mods:affiliation>
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<mods:nameIdentifier id="96D8C46265BC3706CEB459B2A2D3057B" type="email">hinckleya@si.edu</mods:nameIdentifier>
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<mods:namePart id="4961BD9F675EA2FA36FB0C7C21E09666">Hinckley, Arlo</mods:namePart>
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<mods:affiliation id="24883ABC8CEC246C1D9F9D5CE74B119B">Division of Mammals, Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, District of Columbia, Washington, USA</mods:affiliation>
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<mods:nameIdentifier id="AD04514E6385CAA5396AC0F4C73203F6" type="email">hinckleya@si.edu</mods:nameIdentifier>
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<treatment id="038C8797FF96FFC4FF16631EE76CFD80" LSID="urn:lsid:plazi:treatment:038C8797FF96FFC4FF16631EE76CFD80" httpUri="http://treatment.plazi.org/id/038C8797FF96FFC4FF16631EE76CFD80" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
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<treatment id="038C8797FF96FFC4FF16631EE76CFD80" ID-DOI="http://doi.org/10.5281/zenodo.14510289" ID-Zenodo-Dep="14510289" LSID="urn:lsid:plazi:treatment:038C8797FF96FFC4FF16631EE76CFD80" httpUri="http://treatment.plazi.org/id/038C8797FF96FFC4FF16631EE76CFD80" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
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<subSubSection id="C33F650AFF96FFC2FF16631EE0D6FAFF" box="[159,716,1285,1312]" pageId="19" pageNumber="20" type="nomenclature">
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<paragraph id="8B9A3681FF96FFC2FF16631EE0D6FAFF" blockId="19.[113,762,1285,1442]" box="[159,716,1285,1312]" pageId="19" pageNumber="20">
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@ -166,9 +169,9 @@ on
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, weight
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<quantity id="4CDD9B64FF96FFC2FE6A60E7E039F8CA" box="[483,547,1788,1813]" metricMagnitude="-2" metricUnit="kg" metricValue="4.25" pageId="19" pageNumber="20" unit="g" value="42.5">42.5 g</quantity>
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. Skull measurements in Supporting Information, Table S2. 3D surface scans of this holotype skull are available at: https://doi.org/10.17602/ M2/M560018 (cranium); https://doi.org/10.17602/M2/ M560022 (mandible). Pictures of this holotype skull and skin are shown in
|
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<figureCitation id="131E2A04FF96FFC2FF706182E344F86E" box="[249,350,1945,1969]" captionStart="Figure 9" captionStartId="22.[129,194,1582,1606]" captionTargetBox="[129,1473,144,1554]" captionTargetId="figure-201@22.[129,1473,144,1554]" captionTargetPageId="22" captionText="Figure 9. Skulls of (A) Hylomys oorax (USNM 271034, holotype) and (B) Hylomys maxi (USNM 487905, standard specimen since holotype is damaged). All images are at the same scale. Photo credits: Katie Sayers." pageId="19" pageNumber="20">Figure 9A</figureCitation>
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<figureCitation id="131E2A04FF96FFC2FF706182E344F86E" box="[249,350,1945,1969]" captionStart="Figure 9" captionStartId="22.[129,194,1582,1606]" captionTargetBox="[129,1473,144,1554]" captionTargetId="figure-201@22.[129,1473,144,1554]" captionTargetPageId="22" captionText="Figure 9. Skulls of (A) Hylomys oorax (USNM 271034, holotype) and (B) Hylomys maxi (USNM 487905, standard specimen since holotype is damaged). All images are at the same scale. Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510285" httpUri="https://zenodo.org/record/14510285/files/figure.png" pageId="19" pageNumber="20">Figure 9A</figureCitation>
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and the Supporting Information,
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<figureCitation id="131E2A04FF96FFC2FD316182E15FFF77" box="[696,837,144,1969]" captionStart="Figure 9" captionStartId="22.[129,194,1582,1606]" captionTargetBox="[129,1473,144,1554]" captionTargetId="figure-201@22.[129,1473,144,1554]" captionTargetPageId="22" captionText="Figure 9. Skulls of (A) Hylomys oorax (USNM 271034, holotype) and (B) Hylomys maxi (USNM 487905, standard specimen since holotype is damaged). All images are at the same scale. Photo credits: Katie Sayers." pageId="19" pageNumber="20">Figure S9</figureCitation>
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<figureCitation id="131E2A04FF96FFC2FD316182E15FFF77" box="[696,837,144,1969]" captionStart="Figure 9" captionStartId="22.[129,194,1582,1606]" captionTargetBox="[129,1473,144,1554]" captionTargetId="figure-201@22.[129,1473,144,1554]" captionTargetPageId="22" captionText="Figure 9. Skulls of (A) Hylomys oorax (USNM 271034, holotype) and (B) Hylomys maxi (USNM 487905, standard specimen since holotype is damaged). All images are at the same scale. Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510285" httpUri="https://zenodo.org/record/14510285/files/figure.png" pageId="19" pageNumber="20">Figure S9</figureCitation>
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, respectively. DNA sequences associated with this holotype have been deposited in GenBank:
|
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<accessionNumber id="9476AB62FF96FFC2FB0C66B4E6E2FF18" box="[1157,1272,175,199]" pageId="19" pageNumber="20">OR138102</accessionNumber>
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and
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|
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@ -269,7 +272,7 @@ a.s.l.
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was illustrated by
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<collectorName id="26D05357FF96FFC2FADA65E8E192FBF5" pageId="19" pageNumber="20">Frederick A. Ulmer</collectorName>
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(Supporting Information, File S1,
|
||||
<figureCitation id="131E2A04FF96FFC2FB6D6209E722FBF5" box="[1252,1336,1042,1066]" captionStart="Figure 11" captionStartId="24.[128,193,1291,1315]" captionTargetBox="[129,1473,154,1263]" captionTargetId="figure-340@24.[129,1473,144,1263]" captionTargetPageId="24" captionText="Figure 11. A, trapline set by Frederick A. Ulmer and other collectors of the George Vanderbilt Expedition. B, habitat of H. oorax in Blangbeke (2073 m a.s.l.), Aceh, Sumatra.C, habitat of H.oorax, view from the summit of Gunung Leuser looking eastward.D, paratype of Hylomys oorax (ANSP20377) collected on 12 April 1939 by Fred Ulmer in Blangbeke, Aceh, Sumatra, and illustrated by him (Supporting Information, Fig. S11). Picture credits:ANSP Archives Collection (unaccessioned Vertebrate Zoology 2012-01: George Vanderbilt Sumatra Expedition in 1939: photographs and images by Frederick A. Ulmer, Jr)." pageId="19" pageNumber="20">Fig. S11</figureCitation>
|
||||
<figureCitation id="131E2A04FF96FFC2FB6D6209E722FBF5" box="[1252,1336,1042,1066]" captionStart="Figure 11" captionStartId="24.[128,193,1291,1315]" captionTargetBox="[129,1473,154,1263]" captionTargetId="figure-340@24.[129,1473,144,1263]" captionTargetPageId="24" captionText="Figure 11. A, trapline set by Frederick A. Ulmer and other collectors of the George Vanderbilt Expedition. B, habitat of H. oorax in Blangbeke (2073 m a.s.l.), Aceh, Sumatra.C, habitat of H.oorax, view from the summit of Gunung Leuser looking eastward.D, paratype of Hylomys oorax (ANSP20377) collected on 12 April 1939 by Fred Ulmer in Blangbeke, Aceh, Sumatra, and illustrated by him (Supporting Information, Fig. S11). Picture credits:ANSP Archives Collection (unaccessioned Vertebrate Zoology 2012-01: George Vanderbilt Sumatra Expedition in 1939: photographs and images by Frederick A. Ulmer, Jr)." figureDoi="http://doi.org/10.5281/zenodo.14510291" httpUri="https://zenodo.org/record/14510291/files/figure.png" pageId="19" pageNumber="20">Fig. S11</figureCitation>
|
||||
)
|
||||
</materialsCitation>
|
||||
</paragraph>
|
||||
|
|
@ -344,7 +347,7 @@ is distinguished by its smaller (average
|
|||
; LMA =
|
||||
<quantity id="4CDD9B64FF91FFC5FE86659AE344FC47" box="[271,350,897,921]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="20" pageNumber="21" unit="mm" value="24.0">24 mm</quantity>
|
||||
;
|
||||
<figureCitation id="131E2A04FF91FFC5FEE2659AE3AFFC46" box="[363,437,897,921]" captionStart="Figure 9" captionStartId="22.[129,194,1582,1606]" captionTargetBox="[129,1473,144,1554]" captionTargetId="figure-201@22.[129,1473,144,1554]" captionTargetPageId="22" captionText="Figure 9. Skulls of (A) Hylomys oorax (USNM 271034, holotype) and (B) Hylomys maxi (USNM 487905, standard specimen since holotype is damaged). All images are at the same scale. Photo credits: Katie Sayers." pageId="20" pageNumber="21">Fig. 9A</figureCitation>
|
||||
<figureCitation id="131E2A04FF91FFC5FEE2659AE3AFFC46" box="[363,437,897,921]" captionStart="Figure 9" captionStartId="22.[129,194,1582,1606]" captionTargetBox="[129,1473,144,1554]" captionTargetId="figure-201@22.[129,1473,144,1554]" captionTargetPageId="22" captionText="Figure 9. Skulls of (A) Hylomys oorax (USNM 271034, holotype) and (B) Hylomys maxi (USNM 487905, standard specimen since holotype is damaged). All images are at the same scale. Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510285" httpUri="https://zenodo.org/record/14510285/files/figure.png" pageId="20" pageNumber="21">Fig. 9A</figureCitation>
|
||||
) vs. larger size (
|
||||
<collectionCode id="ED34AE44FF91FFC5FDEC659AE096FC46" box="[613,652,897,921]" country="Brazil" lsid="urn:lsid:biocol.org:col:15564" name="Herbarium Bradeanum" pageId="20" pageNumber="21" type="Herbarium">HB</collectionCode>
|
||||
=
|
||||
|
|
@ -358,7 +361,7 @@ is distinguished by its smaller (average
|
|||
; LMA =
|
||||
<quantity id="4CDD9B64FF91FFC5FD9D65BBE06EFC68" box="[532,628,928,952]" metricMagnitude="-2" metricUnit="m" metricValue="2.63" pageId="20" pageNumber="21" unit="mm" value="26.3">26.3 mm</quantity>
|
||||
;
|
||||
<figureCitation id="131E2A04FF91FFC5FD0965BBE0D2FC67" box="[640,712,928,952]" captionStart="Figure 9" captionStartId="22.[129,194,1582,1606]" captionTargetBox="[129,1473,144,1554]" captionTargetId="figure-201@22.[129,1473,144,1554]" captionTargetPageId="22" captionText="Figure 9. Skulls of (A) Hylomys oorax (USNM 271034, holotype) and (B) Hylomys maxi (USNM 487905, standard specimen since holotype is damaged). All images are at the same scale. Photo credits: Katie Sayers." pageId="20" pageNumber="21">Fig. 9B</figureCitation>
|
||||
<figureCitation id="131E2A04FF91FFC5FD0965BBE0D2FC67" box="[640,712,928,952]" captionStart="Figure 9" captionStartId="22.[129,194,1582,1606]" captionTargetBox="[129,1473,144,1554]" captionTargetId="figure-201@22.[129,1473,144,1554]" captionTargetPageId="22" captionText="Figure 9. Skulls of (A) Hylomys oorax (USNM 271034, holotype) and (B) Hylomys maxi (USNM 487905, standard specimen since holotype is damaged). All images are at the same scale. Photo credits: Katie Sayers." figureDoi="http://doi.org/10.5281/zenodo.14510285" httpUri="https://zenodo.org/record/14510285/files/figure.png" pageId="20" pageNumber="21">Fig. 9B</figureCitation>
|
||||
), uniformly black and relatively longer (average
|
||||
<collectionCode id="ED34AE44FF91FFC5FDC765DBE07AFC08" box="[590,608,960,983]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="21">T</collectionCode>
|
||||
=
|
||||
|
|
@ -374,13 +377,13 @@ is distinguished by its smaller (average
|
|||
,
|
||||
<collectionCode id="ED34AE44FF91FFC5FD6265E5E0E5FBC9" box="[747,767,1022,1046]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="21">T</collectionCode>
|
||||
/ HB = 5.11–16.9%) (Supporting Information, File S1,
|
||||
<figureCitation id="131E2A04FF91FFC5FD6A6206E2A2FB8A" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." pageId="20" pageNumber="21">Fig. S10A</figureCitation>
|
||||
<figureCitation id="131E2A04FF91FFC5FD6A6206E2A2FB8A" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." figureDoi="http://doi.org/10.5281/zenodo.14510287" httpUri="https://zenodo.org/record/14510287/files/figure.png" pageId="20" pageNumber="21">Fig. S10A</figureCitation>
|
||||
), and a relatively longer (E/
|
||||
<collectionCode id="ED34AE44FF91FFC5FD806226E027FB8A" box="[521,573,1085,1109]" pageId="20" pageNumber="21">GLS</collectionCode>
|
||||
= 54.8–55.9%) vs. shorter ear (E/
|
||||
<collectionCode id="ED34AE44FF91FFC5FE966247E349FBAB" box="[287,339,1116,1140]" pageId="20" pageNumber="21">GLS</collectionCode>
|
||||
= 39.3–48.2%) (Supporting Information, File S1,
|
||||
<figureCitation id="131E2A04FF91FFC5FF526260E35AFB4C" box="[219,320,1147,1171]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." pageId="20" pageNumber="21">Fig. S10B</figureCitation>
|
||||
<figureCitation id="131E2A04FF91FFC5FF526260E35AFB4C" box="[219,320,1147,1171]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." figureDoi="http://doi.org/10.5281/zenodo.14510287" httpUri="https://zenodo.org/record/14510287/files/figure.png" pageId="20" pageNumber="21">Fig. S10B</figureCitation>
|
||||
). Its fur is softer, denser, and longer (
|
||||
<quantity id="4CDD9B64FF91FFC5FD566260E2B3FB6D" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="20" pageNumber="21" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 mm</quantity>
|
||||
at axial region) than
|
||||
|
|
@ -417,7 +420,7 @@ has a narrower and more gracile appearance than
|
|||
<emphasis id="B951EA93FF91FFC5FEE8634CE3AEFAB1" box="[353,436,1367,1390]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="131E2A04FF91FFC5FE4E634CE014FAB0" box="[455,526,1367,1391]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." pageId="20" pageNumber="21">Fig. 10</figureCitation>
|
||||
<figureCitation id="131E2A04FF91FFC5FE4E634CE014FAB0" box="[455,526,1367,1391]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." figureDoi="http://doi.org/10.5281/zenodo.14510287" httpUri="https://zenodo.org/record/14510287/files/figure.png" pageId="20" pageNumber="21">Fig. 10</figureCitation>
|
||||
). Its rostrum breadth is narrower (4.5–5.0 mm) vs. broader (
|
||||
<quantity id="4CDD9B64FF91FFC5FD88636DE09FFA51" box="[513,645,1398,1422]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" metricValueMax="6.4" metricValueMin="5.2" pageId="20" pageNumber="21" unit="mm" value="5.8" valueMax="6.4" valueMin="5.2">5.2–6.4 mm</quantity>
|
||||
) in
|
||||
|
|
@ -437,7 +440,7 @@ has a narrower and more gracile appearance than
|
|||
/
|
||||
<collectionCode id="ED34AE44FF91FFC5FE7B63AEE03EFA12" box="[498,548,1461,1485]" pageId="20" pageNumber="21">GLS</collectionCode>
|
||||
= 32.4%; Supporting Information, File S1,
|
||||
<figureCitation id="131E2A04FF91FFC5FED463CFE3D9FA33" box="[349,451,1492,1516]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." pageId="20" pageNumber="21">Fig. S10C</figureCitation>
|
||||
<figureCitation id="131E2A04FF91FFC5FED463CFE3D9FA33" box="[349,451,1492,1516]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." figureDoi="http://doi.org/10.5281/zenodo.14510287" httpUri="https://zenodo.org/record/14510287/files/figure.png" pageId="20" pageNumber="21">Fig. S10C</figureCitation>
|
||||
). Antorbital ridge/flange is undeveloped vs. well developed in
|
||||
<taxonomicName id="4C254D02FF91FFC5FE5C63EFE032F9D4" authorityName="Sody" authorityYear="1933" box="[469,552,1524,1547]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
|
||||
<emphasis id="B951EA93FF91FFC5FE5C63EFE032F9D4" box="[469,552,1524,1547]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
|
||||
|
|
@ -452,7 +455,7 @@ Its interorbital constriction is proportionately broader than
|
|||
<emphasis id="B951EA93FF91FFC5FEC3606AE384F957" box="[330,414,1649,1672]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
|
||||
</taxonomicName>
|
||||
(Supporting Information, File S1,
|
||||
<figureCitation id="131E2A04FF91FFC5FF08608BE2FCF977" box="[129,230,1680,1704]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." pageId="20" pageNumber="21">Fig. S10D</figureCitation>
|
||||
<figureCitation id="131E2A04FF91FFC5FF08608BE2FCF977" box="[129,230,1680,1704]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." figureDoi="http://doi.org/10.5281/zenodo.14510287" httpUri="https://zenodo.org/record/14510287/files/figure.png" pageId="20" pageNumber="21">Fig. S10D</figureCitation>
|
||||
). Zygomata and supraorbital processes are less robust than in
|
||||
<taxonomicName id="4C254D02FF91FFC5FF5C60ABE333F918" authorityName="Sody" authorityYear="1933" box="[213,297,1712,1735]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="maxi">
|
||||
<emphasis id="B951EA93FF91FFC5FF5C60ABE333F918" box="[213,297,1712,1735]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
|
||||
|
|
@ -490,7 +493,7 @@ Its interorbital constriction is proportionately broader than
|
|||
<emphasis id="B951EA93FF91FFC5FAF9668AE7D8FF77" box="[1392,1474,145,168]" italics="true" pageId="20" pageNumber="21">H. maxi</emphasis>
|
||||
</taxonomicName>
|
||||
(
|
||||
<figureCitation id="131E2A04FF91FFC5FCCD66ABE18DFF18" box="[836,919,175,199]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." pageId="20" pageNumber="21">
|
||||
<figureCitation id="131E2A04FF91FFC5FCCD66ABE18DFF18" box="[836,919,175,199]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." figureDoi="http://doi.org/10.5281/zenodo.14510287" httpUri="https://zenodo.org/record/14510287/files/figure.png" pageId="20" pageNumber="21">
|
||||
<emphasis id="B951EA93FF91FFC5FCCD66ABE178FF18" box="[836,866,176,199]" italics="true" pageId="20" pageNumber="21">Fig</emphasis>
|
||||
.
|
||||
<emphasis id="B951EA93FF91FFC5FCE766B4E18DFF18" box="[878,919,175,199]" italics="true" pageId="20" pageNumber="21">10B</emphasis>
|
||||
|
|
@ -680,7 +683,7 @@ a more elongated appearance. Toothrow length is longer (
|
|||
<emphasis id="B951EA93FF91FFC5FAE962A1E7D8FB0E" box="[1376,1474,1210,1233]" italics="true" pageId="20" pageNumber="21">H. parvus</emphasis>
|
||||
</taxonomicName>
|
||||
(Supporting Information, File S1,
|
||||
<figureCitation id="131E2A04FF91FFC5FB1A62C2E6E8FB2E" box="[1171,1266,1241,1265]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." pageId="20" pageNumber="21">Fig. S10E</figureCitation>
|
||||
<figureCitation id="131E2A04FF91FFC5FB1A62C2E6E8FB2E" box="[1171,1266,1241,1265]" captionStart="Figure 10" captionStartId="23.[114,179,712,736]" captionTargetBox="[114,1458,144,684]" captionTargetId="figure-659@23.[114,1458,144,684]" captionTargetPageId="23" captionText="Figure 10. A, dorsal view of the rostrum of two H.oorax specimens (left; ANSP 20374 and 20377) and two H. maxi specimens from Sumatra (right; ANSP 20372 and ZRC4.5051), which illustrates the narrow, notched space between premaxillary tips of H.oorax (vs. broad/obtuse in H. maxi). B, lateral view of the bullae of H. oorax (left; ANSP 20377) and H. maxi (right; USNM487909), which illustrates the robust ectotympanic process that thickens distally in H. oorax (vs. more gracile and dorsoventrally flattened in H. maxi)." figureDoi="http://doi.org/10.5281/zenodo.14510287" httpUri="https://zenodo.org/record/14510287/files/figure.png" pageId="20" pageNumber="21">Fig. S10E</figureCitation>
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||||
). First upper incisor is more procumbent than
|
||||
<taxonomicName id="4C254D02FF91FFC5FBC962E2E6BAFACF" baseAuthorityName="Jbnkins and Robinson" baseAuthorityYear="2002" box="[1088,1184,1273,1296]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="parous">
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||||
<emphasis id="B951EA93FF91FFC5FBC962E2E6BAFACF" box="[1088,1184,1273,1296]" italics="true" pageId="20" pageNumber="21">H. parvus</emphasis>
|
||||
|
|
@ -708,7 +711,7 @@ Currently known from montane forest from
|
|||
a.s.l. on Mount Leuser, but it seems to be most abundant at elevations ranging from
|
||||
<quantity id="4CDD9B64FF91FFC5FA0763D5E1BFF9DA" metricMagnitude="3" metricUnit="m" metricValue="2.2405" metricValueMax="2.408" metricValueMin="2.073" pageId="20" pageNumber="21" unit="m" value="2240.5" valueMax="2408.0" valueMin="2073.0">2073 to 2408 m</quantity>
|
||||
a.s.l. (
|
||||
<figureCitation id="131E2A04FF91FFC5FC6163F6E635F9DA" box="[1000,1071,1517,1541]" captionStart="Figure 11" captionStartId="24.[128,193,1291,1315]" captionTargetBox="[129,1473,154,1263]" captionTargetId="figure-340@24.[129,1473,144,1263]" captionTargetPageId="24" captionText="Figure 11. A, trapline set by Frederick A. Ulmer and other collectors of the George Vanderbilt Expedition. B, habitat of H. oorax in Blangbeke (2073 m a.s.l.), Aceh, Sumatra.C, habitat of H.oorax, view from the summit of Gunung Leuser looking eastward.D, paratype of Hylomys oorax (ANSP20377) collected on 12 April 1939 by Fred Ulmer in Blangbeke, Aceh, Sumatra, and illustrated by him (Supporting Information, Fig. S11). Picture credits:ANSP Archives Collection (unaccessioned Vertebrate Zoology 2012-01: George Vanderbilt Sumatra Expedition in 1939: photographs and images by Frederick A. Ulmer, Jr)." pageId="20" pageNumber="21">Fig. 11</figureCitation>
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<figureCitation id="131E2A04FF91FFC5FC6163F6E635F9DA" box="[1000,1071,1517,1541]" captionStart="Figure 11" captionStartId="24.[128,193,1291,1315]" captionTargetBox="[129,1473,154,1263]" captionTargetId="figure-340@24.[129,1473,144,1263]" captionTargetPageId="24" captionText="Figure 11. A, trapline set by Frederick A. Ulmer and other collectors of the George Vanderbilt Expedition. B, habitat of H. oorax in Blangbeke (2073 m a.s.l.), Aceh, Sumatra.C, habitat of H.oorax, view from the summit of Gunung Leuser looking eastward.D, paratype of Hylomys oorax (ANSP20377) collected on 12 April 1939 by Fred Ulmer in Blangbeke, Aceh, Sumatra, and illustrated by him (Supporting Information, Fig. S11). Picture credits:ANSP Archives Collection (unaccessioned Vertebrate Zoology 2012-01: George Vanderbilt Sumatra Expedition in 1939: photographs and images by Frederick A. Ulmer, Jr)." figureDoi="http://doi.org/10.5281/zenodo.14510291" httpUri="https://zenodo.org/record/14510291/files/figure.png" pageId="20" pageNumber="21">Fig. 11</figureCitation>
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||||
). This habitat is characterized by ‘trees averaging only 15 to 40 feet in height, very hard, knotted and twisted. Everywhere the ground and the branches of the trees were covered with a deep carpet of moss and ferns’ (Ulmer’s field notes,
|
||||
<bibRefCitation id="EFB44B70FF91FFC5FC266070E632F95C" author="Millbo GS" box="[943,1064,1643,1667]" pageId="20" pageNumber="21" pagination="107 - 65" refId="ref34041" refString="Millbo GS. Zoological obsults of thb Gboogb Vandbobilt Sumatoan Expbdition, 1936 - 1939 paot V: mammals collbctbd bv Fobdboick A Ulmbo, Jo on Sumatoa and Nias. Proceedings of the Academy of Natural Sciences of Philadelphia 1942; 9: 107 - 65." type="journal article" year="1942">Miller 1942</bibRefCitation>
|
||||
). However, one specimen was recorded at
|
||||
|
|
|
|||
|
|
@ -1,103 +1,106 @@
|
|||
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<mods:title id="77FF40480B206693E2AC065AB5219380">Zoological Journal of the Linnean Society</mods:title>
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<mods:title id="F33E912228C673BBD38E0F2250366A81">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="2F745EDCE799ABAA603BEC5FA4632D7A">
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<mods:date id="05525E03A93B7F4B1906E108E4DF9BAA">2024</mods:date>
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<mods:number id="A7DCD3EB616893D97AF31A4211A3E9B1">2023-12-21</mods:number>
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<mods:date id="14B3527A7EDAA70A87252980E6687507">2024</mods:date>
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<mods:number id="CD35CE07D56CB30B5C7932A174B5EED2">202</mods:number>
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<mods:number id="5EA7E4857A5679EE425E56BFD46FEC21">202</mods:number>
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<mods:url id="8FE3A407CABE7D4225D312B289CD3DD2">https://doi.org/10.1093/zoolinnean/zlad177</mods:url>
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<mods:location id="7FBB4B64588DFC6AE431B5DACBCEEE47">
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<mods:url id="6A384695CC27351987EE1461B40DD581">https://doi.org/10.1093/zoolinnean/zlad177</mods:url>
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</mods:mods>
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<treatment id="038C8797FF9CFFC8FEAB64C1E788F92C" LSID="urn:lsid:plazi:treatment:038C8797FF9CFFC8FEAB64C1E788F92C" httpUri="http://treatment.plazi.org/id/038C8797FF9CFFC8FEAB64C1E788F92C" lastPageNumber="26" pageId="25" pageNumber="26">
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<treatment id="038C8797FF9CFFC8FEAB64C1E788F92C" ID-DOI="http://doi.org/10.5281/zenodo.14510297" ID-Zenodo-Dep="14510297" LSID="urn:lsid:plazi:treatment:038C8797FF9CFFC8FEAB64C1E788F92C" httpUri="http://treatment.plazi.org/id/038C8797FF9CFFC8FEAB64C1E788F92C" lastPageNumber="26" pageId="25" pageNumber="26">
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<subSubSection id="C33F650AFF9CFFC8FEAB64C1E050FD2B" box="[290,586,730,757]" pageId="25" pageNumber="26" type="nomenclature">
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<paragraph id="8B9A3681FF9CFFC8FEAB64C1E050FD2B" blockId="25.[113,700,730,793]" box="[290,586,730,757]" pageId="25" pageNumber="26">
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<taxonomicName id="4C254D02FF9CFFC8FEAB64C1E050FD2B" ID-CoL="6MNPV" authority="Muller, 1841" authorityName="Muller" authorityYear="1841" box="[290,586,730,757]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suillus">
|
||||
|
|
@ -299,7 +302,7 @@ based on its bicoloured vs. monocoloured tail, shorter hindfoot (
|
|||
+ nail = 23–26 vs.
|
||||
<quantity id="4CDD9B64FF9CFFC8FBF66757E703FEBB" box="[1151,1305,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="2.75" metricValueMin="2.55" pageId="25" pageNumber="26" unit="mm" value="26.5" valueMax="27.5" valueMin="25.5">25.5–27.5 mm</quantity>
|
||||
), and relatively broader rostrum (
|
||||
<figureCitation id="131E2A04FF9CFFC8FC6C6770E605FE5C" box="[997,1055,363,387]" captionStart="Figure 6" captionStartId="10.[129,194,1089,1113]" captionTargetBox="[130,1472,168,1060]" captionTargetId="figure-528@10.[129,1473,144,1061]" captionTargetPageId="10" captionText="Figure 6. Bivariate plot of rostrum length (ROL) and rostrum breadth (ROB) measurements." pageId="25" pageNumber="26">Fig. 6</figureCitation>
|
||||
<figureCitation id="131E2A04FF9CFFC8FC6C6770E605FE5C" box="[997,1055,363,387]" captionStart="Figure 6" captionStartId="10.[129,194,1089,1113]" captionTargetBox="[130,1472,168,1060]" captionTargetId="figure-528@10.[129,1473,144,1061]" captionTargetPageId="10" captionText="Figure 6. Bivariate plot of rostrum length (ROL) and rostrum breadth (ROB) measurements." figureDoi="http://doi.org/10.5281/zenodo.14510275" httpUri="https://zenodo.org/record/14510275/files/figure.png" pageId="25" pageNumber="26">Fig. 6</figureCitation>
|
||||
). It has a larger braincase (
|
||||
<collectionCode id="ED34AE44FF9CFFC8FAB06770E746FE5C" box="[1337,1372,363,387]" country="0" name="Buffalo Bill Museum" pageId="25" pageNumber="26">BB</collectionCode>
|
||||
= 13.8– 15.6 vs.
|
||||
|
|
@ -403,7 +406,7 @@ Ruedi
|
|||
<paragraph id="8B9A3681FF9CFFC8FCA36004E788F92C" blockId="25.[810,1459,1567,1779]" pageId="25" pageNumber="26">
|
||||
<emphasis id="B951EA93FF9CFFC8FCA36004E180F9E9" box="[810,922,1567,1590]" italics="true" pageId="25" pageNumber="26">Comments:</emphasis>
|
||||
This species exhibits a high degree of morphological variation, spanning about twice the morphospace compared to its relatives in the craniodental PCA (
|
||||
<figureCitation id="131E2A04FF9CFFC8FB5B6045E717F9A9" box="[1234,1293,1630,1654]" captionStart="Figure 5" captionStartId="9.[113,178,1006,1030]" captionTargetBox="[113,1457,145,977]" captionTargetId="figure-518@9.[112,1459,144,979]" captionTargetPageId="9" captionText="Figure 5. Morphometric variation in Hylomys. A, first and second principal components from an analysis of 16 cranial dimensions, computed with Dataset 4 (with the exclusion of external measurements); B, second and third principal components of the same analysis.Asterisks denote undamaged holotype or neotype specimens that could be included in this analysis." pageId="25" pageNumber="26">Fig. 5</figureCitation>
|
||||
<figureCitation id="131E2A04FF9CFFC8FB5B6045E717F9A9" box="[1234,1293,1630,1654]" captionStart="Figure 5" captionStartId="9.[113,178,1006,1030]" captionTargetBox="[113,1457,145,977]" captionTargetId="figure-518@9.[112,1459,144,979]" captionTargetPageId="9" captionText="Figure 5. Morphometric variation in Hylomys. A, first and second principal components from an analysis of 16 cranial dimensions, computed with Dataset 4 (with the exclusion of external measurements); B, second and third principal components of the same analysis.Asterisks denote undamaged holotype or neotype specimens that could be included in this analysis." figureDoi="http://doi.org/10.5281/zenodo.14510273" httpUri="https://zenodo.org/record/14510273/files/figure.png" pageId="25" pageNumber="26">Fig. 5</figureCitation>
|
||||
). This variation seems somewhat geographically structured (east and west Java), although there are several outliers that break this pattern. Future studies should examine species limits within
|
||||
<taxonomicName id="4C254D02FF9CFFC8FB6360A7E759F90B" authorityName="Mullbo" authorityYear="1841" box="[1258,1347,1724,1748]" class="Mammalia" family="Erinaceidae" genus="Hylomys" kingdom="Animalia" order="Erinaceomorpha" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="suillus">
|
||||
<emphasis id="B951EA93FF9CFFC8FB6360A7E759F90B" box="[1258,1347,1724,1748]" italics="true" pageId="25" pageNumber="26">H. suillus</emphasis>
|
||||
|
|
|
|||
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Reference in a new issue