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<document id="F20091D40D57622A55523A8021BBBB96" ID-CLB-Dataset="50795" ID-DOI="10.5281/zenodo.171623" ID-GBIF-Dataset="e1d6871f-7dfa-444b-8eaa-fc65a409abf7" ID-ISSN="1175­5326" ID-Zenodo-Dep="171623" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459755598005" checkinUser="plazi" docAuthor="Keable, Stephen J. &amp; Wilson, George D. F." docDate="2006" docId="02358794FFAAFF93C13BB560FDDCFBAD" docLanguage="en" docName="zt01116p027.pdf" docOrigin="Zootaxa 1116" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Pygolabis Wilson 2003" docType="treatment" docVersion="11" lastPageNumber="4" masterDocId="FE0CFFECFFA8FF90C033B00CFF89FF83" masterDocTitle="New species of Pygolabis Wilson, 2003 (Isopoda, Tainisopidae, Crustacea) from Western Australia" masterLastPageNumber="27" masterPageNumber="1" pageNumber="3" updateTime="1698217552216" updateUser="ExternalLinkService">
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<mods:title id="D0BB9837F23CAF130E9BBF5954965585">New species of Pygolabis Wilson, 2003 (Isopoda, Tainisopidae, Crustacea) from Western Australia</mods:title>
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<mods:namePart id="1E3B81464042D0E8EFCF7A5406E31251">Keable, Stephen J.</mods:namePart>
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<mods:namePart id="5878B83C58028638CB4B8DE0D8685972">Wilson, George D. F.</mods:namePart>
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<mods:date id="BC9A67DE751DF9FD4259E751C63DAD01">2006</mods:date>
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<treatment id="02358794FFAAFF93C13BB560FDDCFBAD" ID-DOI="http://doi.org/10.5281/zenodo.3511195" ID-GBIF-Taxon="119343536" ID-Zenodo-Dep="3511195" LSID="urn:lsid:plazi:treatment:02358794FFAAFF93C13BB560FDDCFBAD" httpUri="http://treatment.plazi.org/id/02358794FFAAFF93C13BB560FDDCFBAD" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<subSubSection id="C2866509FFAAFF92C13BB560FDACFA05" box="[264,549,1388,1414]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8A233682FFAAFF92C13BB560FDACFA05" blockId="2.[264,549,1388,1414]" box="[264,549,1388,1414]" pageId="2" pageNumber="3">
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<taxonomicName id="4D9C4D01FFAAFF92C13BB560FDACFA05" ID-CoL="7PX54" authority="Wilson, 2003" authorityName="Wilson" authorityYear="2003" box="[264,549,1388,1414]" class="Malacostraca" family="Tainisopidae" genus="Pygolabis" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B8E8EA90FFAAFF92C13BB560FDACFA05" bold="true" box="[264,549,1388,1414]" pageId="2" pageNumber="3">
<emphasis id="B8E8EA90FFAAFF92C13BB560FEF4FA05" bold="true" box="[264,381,1388,1414]" italics="true" pageId="2" pageNumber="3">Pygolabis</emphasis>
<bibRefCitation id="EE0D4B73FFAAFF92C1B6B560FDACFA05" author="Wilson" box="[389,549,1388,1414]" pageId="2" pageNumber="27" refString="Wilson, G. D. F. (2003) A new genus of Tainisopidae fam. nov. (Crustacea: Isopoda) from the Pilbara, Western Australia. Zootaxa, 245, 1 - 20." type="journal article" year="2003">Wilson, 2003</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<paragraph id="8A233682FFAAFF92C13BB5BBFDA3FA4E" blockId="2.[264,554,1462,1486]" box="[264,554,1462,1486]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="AA8C11ACFFAAFF92C13BB5BBFDA3FA4E" box="[264,554,1462,1486]" pageId="2" pageNumber="3">
<taxonomicName id="4D9C4D01FFAAFF92C13BB5BBFDAFFA4E" ID-CoL="7PX54" authority="Wilson, 2003: 7" authorityName="Wilson" authorityPageNumber="7" authorityYear="2003" box="[264,550,1462,1486]" class="Malacostraca" family="Tainisopidae" genus="Pygolabis" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B8E8EA90FFAAFF92C13BB5BBFEFAFA4D" box="[264,371,1463,1486]" italics="true" pageId="2" pageNumber="3">Pygolabis</emphasis>
<treatmentCitation id="0B3D1093FFAAFF92C14AB5BAFDAFFA4E" ID-DOI="http://doi.org/10.5281/zenodo.3508981" ID-Zenodo-Dep="3508981" author="Wilson" box="[377,550,1462,1485]" httpUri="http://treatment.plazi.org/id/4911878AFFEFFFCBA506F8D0FE9005EC" page="7" pageId="2" pageNumber="3" year="2003">
<bibRefCitation id="EE0D4B73FFAAFF92C14AB5BAFD85FA4E" author="Wilson" box="[377,524,1462,1485]" pageId="2" pageNumber="27" refString="Wilson, G. D. F. (2003) A new genus of Tainisopidae fam. nov. (Crustacea: Isopoda) from the Pilbara, Western Australia. Zootaxa, 245, 1 - 20." type="journal article" year="2003">Wilson, 2003</bibRefCitation>
: 7
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
</subSubSection>
<subSubSection id="C2866509FFAAFF92C13BB60DFB09F998" box="[264,1152,1537,1563]" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8A233682FFAAFF92C13BB60DFB09F998" blockId="2.[264,1152,1537,1563]" box="[264,1152,1537,1563]" pageId="2" pageNumber="3">
<emphasis id="B8E8EA90FFAAFF92C13BB60DFE23F998" bold="true" box="[264,426,1537,1563]" pageId="2" pageNumber="3">
<typeStatus id="55278820FFAAFF92C13BB60DFECCF998" box="[264,325,1537,1563]" pageId="2" pageNumber="3">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4D9C4D01FFAAFF92C181B60DFCD7F998" authority="Wilson, 2003" authorityName="Wilson" authorityYear="2003" box="[434,862,1537,1563]" class="Malacostraca" family="Tainisopidae" genus="Pygolabis" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="humphreysi">
<emphasis id="B8E8EA90FFAAFF92C181B60DFD3EF999" box="[434,695,1537,1562]" italics="true" pageId="2" pageNumber="3">Pygolabis humphreysi</emphasis>
<bibRefCitation id="EE0D4B73FFAAFF92C28EB60DFCD7F998" author="Wilson" box="[701,862,1537,1563]" pageId="2" pageNumber="27" refString="Wilson, G. D. F. (2003) A new genus of Tainisopidae fam. nov. (Crustacea: Isopoda) from the Pilbara, Western Australia. Zootaxa, 245, 1 - 20." type="journal article" year="2003">Wilson, 2003</bibRefCitation>
</taxonomicName>
; by original designation.
</paragraph>
</subSubSection>
<subSubSection id="C2866509FFAAFF93C13BB65DFE20FC95" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="discussion">
<paragraph id="8A233682FFAAFF93C13BB65DFC56FDCD" blockId="2.[264,1324,1617,1883]" lastBlockId="3.[264,1325,284,1070]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis id="B8E8EA90FFAAFF92C13BB65DFEF3F9E8" bold="true" box="[264,378,1617,1643]" pageId="2" pageNumber="3">Remarks</emphasis>
. The species described below conform to the original diagnosis and description of the genus (
<bibRefCitation id="EE0D4B73FFAAFF92C187B675FDD2F910" author="Wilson" box="[436,603,1657,1683]" pageId="2" pageNumber="27" refString="Wilson, G. D. F. (2003) A new genus of Tainisopidae fam. nov. (Crustacea: Isopoda) from the Pilbara, Western Australia. Zootaxa, 245, 1 - 20." type="journal article" year="2003">Wilson, 2003</bibRefCitation>
), so we do not revise the family or the generic diagnoses.
<bibRefCitation id="EE0D4B73FFAAFF92C13BB6ADFE30F938" author="Wilson" box="[264,441,1697,1723]" pageId="2" pageNumber="27" refString="Wilson, G. D. F. (2003) A new genus of Tainisopidae fam. nov. (Crustacea: Isopoda) from the Pilbara, Western Australia. Zootaxa, 245, 1 - 20." type="journal article" year="2003">Wilson (2003)</bibRefCitation>
indicated that the group is clearly a member of the higher isopods, informally referred to as “Flabellifera
<emphasis id="B8E8EA90FFAAFF92C2C4B6C5FCFFF961" box="[759,886,1737,1762]" italics="true" pageId="2" pageNumber="3">sensu lato</emphasis>
,” the classification of which was substantially reorganised by
<bibRefCitation id="EE0D4B73FFAAFF92C256B6FDFC0FF888" author="Brandt" box="[613,902,1777,1803]" pageId="2" pageNumber="26" refString="Brandt, A. &amp; Poore, G. C. B. (2003) Higher classification of the flabelliferan and related Isopoda based on a reappraisal of relationships. Invertebrate Systematics, 17, 893 - 923." type="journal article" year="2003">Brandt &amp; Poore (2003)</bibRefCitation>
. Although
<bibRefCitation id="EE0D4B73FFAAFF92C423B6FDFAA5F888" author="Brandt" box="[1040,1324,1777,1803]" pageId="2" pageNumber="26" refString="Brandt, A. &amp; Poore, G. C. B. (2003) Higher classification of the flabelliferan and related Isopoda based on a reappraisal of relationships. Invertebrate Systematics, 17, 893 - 923." type="journal article" year="2003">Brandt &amp; Poore (2003)</bibRefCitation>
introduced a subordinal name for the
<taxonomicName id="4D9C4D01FFAAFF92C2EFB715FCFDF8B0" box="[732,884,1817,1843]" class="Malacostraca" family="Tainisopidae" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="3" phylum="Arthropoda" rank="family">Tainisopidae</taxonomicName>
, their analysis had no well–defined basal structure for the higher isopods, and did not establish a sister group relationship between the
<taxonomicName id="4D9C4D01FFABFF93C197B110FDB0FEB5" box="[420,569,284,310]" class="Malacostraca" family="Tainisopidae" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Tainisopidae</taxonomicName>
and any other clade. The overall classification introduced by
<bibRefCitation id="EE0D4B73FFABFF93C13BB148FD9FFEDD" author="Brandt" box="[264,534,324,350]" pageId="3" pageNumber="26" refString="Brandt, A. &amp; Poore, G. C. B. (2003) Higher classification of the flabelliferan and related Isopoda based on a reappraisal of relationships. Invertebrate Systematics, 17, 893 - 923." type="journal article" year="2003">Brandt &amp; Poore (2003)</bibRefCitation>
disagrees with other recent phylogenetic studies and classifications (e.g.,
<bibRefCitation id="EE0D4B73FFABFF93C162B160FD34FE05" author="Tabacaru" box="[337,701,364,390]" pageId="3" pageNumber="26" refString="Tabacaru, I. &amp; Danielopol, D. L. (1999) Contribution a la connaissance de la phylogenie des Isopoda (Crustacea). Vie et Milieu, 49, 163 - 176." type="journal article" year="1999">Tabacaru &amp; Danielopol 1999</bibRefCitation>
;
<bibRefCitation id="EE0D4B73FFABFF93C2E3B160FC6BFE05" author="Martin" box="[720,994,364,390]" pageId="3" pageNumber="26" refString="Martin, J. W. &amp; Davis, G. E. (2001) An Updated Classification of the Recent Crustacea. Natural History Museum of Los Angeles County Science Series, 39, 1 - 124." type="journal article" year="2001">Martin &amp; Davis 2001</bibRefCitation>
;
<bibRefCitation id="EE0D4B73FFABFF93C3C6B160FAA8FE05" author="Dreyer" box="[1013,1313,364,390]" pageId="3" pageNumber="26" refString="Dreyer, H. &amp; Wagele, J. - W. (2002) The Scutocoxifera tax. nov. and the information content of nuclear ssu rDNA sequences for reconstruction of isopod phylogeny (Peracarida: Isopoda). Journal of Crustacean Biology, 22, 217 - 234." type="journal article" year="2002">Dreyer &amp; Wägele 2002</bibRefCitation>
). Recent genetic analyses (C. Francis, personal communication), although only using similarity procedure, found a species of
<taxonomicName id="4D9C4D01FFABFF93C2DEB1B0FCEBFE56" box="[749,866,444,469]" class="Malacostraca" family="Tainisopidae" genus="Pygolabis" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B8E8EA90FFABFF93C2DEB1B0FCEBFE56" box="[749,866,444,469]" italics="true" pageId="3" pageNumber="4">Pygolabis</emphasis>
</taxonomicName>
and an idoteid isopod to have similar sequences. The relationships of the higher isopods apparently require further research, so the subordinal status of the
<taxonomicName id="4D9C4D01FFABFF93C26EB200FD7BFDA5" box="[605,754,524,550]" class="Malacostraca" family="Tainisopidae" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Tainisopidae</taxonomicName>
within the isopods should be left open until a better corroborated analysis of the higher isopods is available.
</paragraph>
<paragraph id="8A233682FFABFF93C10BB250FE20FC95" blockId="3.[264,1325,284,1070]" pageId="3" pageNumber="4">
The sexually dimorphic pleopod 1 endopod is an additional character of generic importance. In males of
<taxonomicName id="4D9C4D01FFABFF93C210B288FD11FD1E" box="[547,664,644,669]" class="Malacostraca" family="Tainisopidae" genus="Pygolabis" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B8E8EA90FFABFF93C210B288FD11FD1E" box="[547,664,644,669]" italics="true" pageId="3" pageNumber="4">Pygolabis</emphasis>
</taxonomicName>
, this limb is twisted, usually around the elongated penes that extend through it, and has several long denticulate setae on the distal margin (PLP
<quantity id="4D649B67FFABFF93C530B2A0FAA2FD45" box="[1283,1323,684,710]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="4" unit="in" value="1.0">1 in</quantity>
<figureCitation id="12A72A07FFABFF93C13BB2D8FED0FD6D" box="[264,345,724,750]" captionStart="FIGURE 7. P" captionStartId="12.[264,367,1844,1868]" captionTargetBox="[289,1315,326,1789]" captionTargetId="figure@12.[272,1315,326,1820]" captionTargetPageId="12" captionText="FIGURE 7. P. eberhardi sp. nov. Pleopods in dorsal view. Male 1 (AM P 70408); male 2 (WAM C 34942); female (WAM C 34944). Scale bar 1 mm." httpUri="https://zenodo.org/record/171630/files/figure.png" pageId="3" pageNumber="4">Figs. 7</figureCitation>
,
<figureCitation id="12A72A07FFABFF93C155B2D8FEFEFD6D" box="[358,375,724,750]" captionStart="FIGURE 9. P" captionStartId="17.[264,367,1844,1868]" captionTargetBox="[279,1297,339,1804]" captionTargetId="figure@17.[272,1315,326,1820]" captionTargetPageId="17" captionText="FIGURE 9. P. weeliwolli sp. nov. Holotype male (WAM C 34948); paratype male 2 (AM P 70413). Scale bar: pereopods, 1 mm; pleopods 0.5 mm." httpUri="https://zenodo.org/record/171632/files/figure.png" pageId="3" pageNumber="4">9</figureCitation>
,
<figureCitation id="12A72A07FFABFF93C1B0B2D8FE28FD6D" box="[387,417,724,750]" captionStart="FIGURE 11. P" captionStartId="20.[264,367,1832,1856]" captionTargetBox="[287,1311,339,1788]" captionTargetId="figure@20.[276,1311,326,1808]" captionTargetPageId="20" captionText="FIGURE 11. P. paraburdoo sp. nov. Paratype male 2 (AM P 70416); holotype male (WAM C 34951). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/171634/files/figure.png" pageId="3" pageNumber="4">11</figureCitation>
,
<figureCitation id="12A72A07FFABFF93C19DB2D8FE47FD6D" box="[430,462,724,750]" captionStart="FIGURE 13. P" captionStartId="23.[264,367,1844,1868]" captionTargetBox="[279,1296,326,1811]" captionTargetId="figure@23.[272,1315,326,1820]" captionTargetPageId="23" captionText="FIGURE 13. P. gascoyne sp. nov. Holotype male (WAM C 34953). Scale bar 0.5 mm." httpUri="https://zenodo.org/record/171636/files/figure.png" pageId="3" pageNumber="4">13</figureCitation>
). In females, pleopod 1 is flattened with a few short plumose setae on the distal margin.
</paragraph>
</subSubSection>
<subSubSection id="C2866509FFABFF93C10BB328FDDCFBAD" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="8A233682FFABFF93C10BB328FDDCFBAD" blockId="3.[264,1325,284,1070]" pageId="3" pageNumber="4">
The
<taxonomicName id="4D9C4D01FFABFF93C143B328FD8CFCBD" ID-CoL="7NL8J" box="[368,517,804,830]" class="Malacostraca" family="Tainisopidae" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="family">Tainisopidae</taxonomicName>
show two distinctly different morphologies for the pleotelson and the position of the anus (
<bibRefCitation id="EE0D4B73FFABFF93C21CB340FCCEFCE5" author="Wilson" box="[559,839,844,870]" pageId="3" pageNumber="27" refString="Wilson, G. D .. F. &amp; Ponder, W. F. (1992) Extraordinary new subterranean isopods (Peracarida, Crustacea) from the Kimberley Region, Western Australia. Records of the Australian Museum, 44, 279 - 298." type="journal article" year="1992">Wilson &amp; Ponder 1992</bibRefCitation>
: fig. 3A;
<bibRefCitation id="EE0D4B73FFABFF93C38BB340FBDDFCE5" author="Wilson" box="[952,1108,844,870]" pageId="3" pageNumber="27" refString="Wilson, G. D. F. (2003) A new genus of Tainisopidae fam. nov. (Crustacea: Isopoda) from the Pilbara, Western Australia. Zootaxa, 245, 1 - 20." type="journal article" year="2003">Wilson 2003</bibRefCitation>
: fig. 6F). Because the section of the pleotelson anterior to the insertion of the uropods is elongate in
<taxonomicName id="4D9C4D01FFABFF93C13BB390FEF4FC36" ID-CoL="7PX54" box="[264,381,924,949]" class="Malacostraca" family="Tainisopidae" genus="Pygolabis" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B8E8EA90FFABFF93C13BB390FEF4FC36" box="[264,381,924,949]" italics="true" pageId="3" pageNumber="4">Pygolabis</emphasis>
</taxonomicName>
, the anus is separated from the pleopodal region by a small ridge and faces posteroventrally. In
<taxonomicName id="4D9C4D01FFABFF93C1CBB3C8FDFFFC5E" ID-CoL="7Q3KH" box="[504,630,964,989]" class="Malacostraca" family="Tainisopidae" genus="Tainisopus" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B8E8EA90FFABFF93C1CBB3C8FDFFFC5E" box="[504,630,964,989]" italics="true" pageId="3" pageNumber="4">Tainisopus</emphasis>
</taxonomicName>
species, where the anterior part of the pleotelson is short relative to the posterior part, the anus is covered by the pleopods, faces ventrally and is not marked by an anterior ridge.
</paragraph>
</subSubSection>
</treatment>
</document>
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