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<document id="EE0421E14AAEF17AAF49AB352131E6E0" ID-DOI="10.5281/zenodo.5319032" ID-ISSN="0374-1036" ID-Zenodo-Dep="5319032" ID-ZooBank="32572CA2-D2CB-4A01-B252-CCECB5320114" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630108184603" checkinUser="marcus" docAuthor="Yin, Zi-Wei, Nomura, Shûhei &amp; Li, Li-Zhen" docDate="2015" docId="022DE650D937FFAF06FDFF70FD82FB8E" docLanguage="en" docName="ActaEntMusNatPra.2015.55.1.105-127.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 55 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Tribasodites kawadai Yin &amp; Nomura &amp; Li 2015, sp. nov." docType="treatment" docVersion="5" lastPageNumber="118" masterDocId="FE149E28D93AFFA2074AFFACFFF5FF92" masterDocTitle="Ten new species of cavernicolous Tribasodites from China and Thailand, and a list of East Asian cave-inhabiting Pselaphinae (Coleoptera: Staphylinidae)" masterLastPageNumber="127" masterPageNumber="105" pageNumber="118" updateTime="1699154406698" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="D2BB6A8FAD76FFF3A5AD5E03F33D7CFE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="94575C7964F88F316E99B7B744D9DC8D">Ten new species of cavernicolous Tribasodites from China and Thailand, and a list of East Asian cave-inhabiting Pselaphinae (Coleoptera: Staphylinidae)</mods:title>
</mods:titleInfo>
<mods:name id="030E170F2E6291B0C2BD7B30709C591A" type="personal">
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<mods:namePart id="BCC45A7811951EEF1D3F7859C95DC2B8">Yin, Zi-Wei</mods:namePart>
<mods:affiliation id="122F5A64C11C9B5BD34663D2B232FB20">Department of Biology, College of Life and Environmental Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai, 200234, P. R. China</mods:affiliation>
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<mods:namePart id="56C22D8755E1E31008D49A03F3421626">Nomura, Shûhei</mods:namePart>
<mods:affiliation id="F6E2D52D05DB720AB7116A19BFC1086F">Department of Zoology, National Museum of Nature and Science, 4 - 1 - 1, Amakubo, Tsukuba-shi, Ibaraki, 305 - 0005 Japan</mods:affiliation>
</mods:name>
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<mods:namePart id="51FB2DF1F491863F0392BE3ABC9D0F1F">Li, Li-Zhen</mods:namePart>
<mods:affiliation id="C1B2449B8B7866AD7EED494762AD0788">Department of Biology, College of Life and Environmental Sciences, Shanghai Normal University, 100 Guilin Road, Shanghai, 200234, P. R. China &amp; Corresponding author; e-mail: pselaphinae @ gmail. com</mods:affiliation>
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<mods:title id="0198674C6F6B59B7A126F955E872401B">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="62624A3F18FE755D229EC68A013A5132">2015</mods:date>
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<mods:number id="210E24AB0CD9FA2468FF8B8D9274D11D">2015-06-01</mods:number>
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<treatment id="022DE650D937FFAF06FDFF70FD82FB8E" ID-DOI="http://doi.org/10.5281/zenodo.5332493" ID-Zenodo-Dep="5332493" LSID="urn:lsid:plazi:treatment:022DE650D937FFAF06FDFF70FD82FB8E" httpUri="http://treatment.plazi.org/id/022DE650D937FFAF06FDFF70FD82FB8E" lastPageNumber="118" pageId="13" pageNumber="118">
<subSubSection id="C29E04CDD937FFAF06FDFF70FD0EFF61" box="[439,763,220,243]" pageId="13" pageNumber="118" type="nomenclature">
<paragraph id="8A3B5746D937FFAF06FDFF70FD0EFF61" blockId="13.[439,763,220,243]" box="[439,763,220,243]" pageId="13" pageNumber="118">
<heading id="D173E02AD937FFAF06FDFF70FD0EFF61" bold="true" box="[439,763,220,243]" centered="true" fontSize="8" level="2" pageId="13" pageNumber="118" reason="3">
<emphasis id="B8F08B54D937FFAF06FDFF70FD0EFF61" bold="true" box="[439,763,220,243]" pageId="13" pageNumber="118">
<taxonomicName id="4D842CC5D937FFAF06FDFF70FD6AFF61" ID-CoL="5864K" authority="Yin &amp; Nomura &amp; Li, 2015" authorityName="Yin &amp; Nomura &amp; Li" authorityYear="2015" box="[439,671,220,243]" class="Insecta" family="Staphylinidae" genus="Tribasodites" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="118" phylum="Arthropoda" rank="species" species="kawadai" status="sp. nov.">
<emphasis id="B8F08B54D937FFAF06FDFF70FD6AFF61" bold="true" box="[439,671,220,243]" italics="true" pageId="13" pageNumber="118">Tribasodites kawadai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3C3362FD937FFAF05ECFF70FD0EFF61" box="[678,763,220,243]" pageId="13" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C29E04CDD937FFAF06B7FF56FD41FE9F" box="[509,692,250,269]" pageId="13" pageNumber="118" type="description">
<paragraph id="8A3B5746D937FFAF06B7FF56FD41FE9F" blockId="13.[509,692,250,269]" box="[509,692,250,269]" pageId="13" pageNumber="118">
(
<figureCitation id="12BF4BC3D937FFAF0549FF56FDCEFE9F" box="[515,571,250,269]" captionStart="Fig" captionStartId="14.[151,182,1203,1222]" captionTargetBox="[156,1125,218,1179]" captionTargetId="figure-119@14.[151,1126,218,1179]" captionTargetPageId="14" captionText="Fig.8. Tribasodites kawadai sp. nov., male.A habitus. B left antenna. C meso- and metaventrite.D aedeagus, in lateral view. E aedeagus, in ventral view. Scales: A = 1.0 mm; C = 0.5 mm; B, D, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5363103" httpUri="https://zenodo.org/record/5363103/files/figure.png" pageId="13" pageNumber="118">Figs 8</figureCitation>
, 12PR, 13A)
</paragraph>
</subSubSection>
<subSubSection id="C29E04CDD937FFAF073BFE88FBB7FEFE" pageId="13" pageNumber="118" type="materials_examined">
<paragraph id="8A3B5746D937FFAF073BFE88FD44FEA5" blockId="13.[113,1090,292,364]" box="[113,689,292,311]" pageId="13" pageNumber="118">
<emphasis id="B8F08B54D937FFAF073BFE88FF1FFEA5" bold="true" box="[113,234,292,311]" pageId="13" pageNumber="118">
<typeStatus id="553FE9E4D937FFAF073BFE88FF6BFEA5" box="[113,158,292,311]" pageId="13" pageNumber="118">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F29317D6D937FFAF07A5FE88FED1FEA5" box="[239,292,292,311]" name="China" pageId="13" pageNumber="118">China</collectingCountry>
,
<collectingRegion id="484099A4D937FFAF0667FE88FE8CFEA5" box="[301,377,292,311]" country="China" name="Guangxi" pageId="13" pageNumber="118">Guangxi</collectingRegion>
, Leye Xian, Bagao, Tiankeng Dong.
</paragraph>
<paragraph id="8A3B5746D937FFAF073BFE92FBB7FEFE" blockId="13.[113,1090,292,364]" pageId="13" pageNumber="118">
<emphasis id="B8F08B54D937FFAF073BFE92FF02FEC3" bold="true" box="[113,247,318,337]" pageId="13" pageNumber="118">Type material.</emphasis>
<materialsCitation id="3AEC5D1BD937FFAF07B7FE92FD54FEFE" collectingDate="2004-09-12" collectingDateMax="2004-09-13" collectingDateMin="2004-09-12" collectorName="A. Kawada'" country="China" county="Chinese" location="Tiankeng Dong" municipality="Leye Xian" pageId="13" pageNumber="118" specimenCount="1" typeStatus="holotype">
<typeStatus id="553FE9E4D937FFAF07B7FE92FEA4FEC2" box="[253,337,318,337]" pageId="13" pageNumber="118" type="holotype">HOLOTYPE</typeStatus>
: J,
<location id="8F5B019DD937FFAF0631FE92FDF0FEC3" LSID="urn:lsid:plazi:treatment:022DE650D937FFAF06FDFF70FD82FB8E:8F5B019DD937FFAF0631FE92FDF0FEC3" box="[379,517,318,337]" country="China" county="Chinese" municipality="Leye Xian" name="Tiankeng Dong" pageId="13" pageNumber="118">Tiankeng Dong</location>
(cave),
<location id="8F5B019DD937FFAF0507FE92FD73FEC3" LSID="urn:lsid:plazi:treatment:022DE650D937FFAF06FDFF70FD82FB8E:8F5B019DD937FFAF0507FE92FD73FEC3" box="[589,646,318,337]" country="China" county="Chinese" municipality="Leye Xian" name="Bagao" pageId="13" pageNumber="118">Bagao</location>
,
<collectingMunicipality id="6A5FCD3CD937FFAF05C5FE92FD19FEC3" box="[655,748,318,337]" pageId="13" pageNumber="118">Leye Xian</collectingMunicipality>
, [NW-Guangxi,
<collectingCountry id="F29317D6D937FFAF0431FE92FC34FEC3" box="[891,961,318,337]" name="China" pageId="13" pageNumber="118">CHINA</collectingCountry>
] / {locality repeated in
<collectingCounty id="635A2FCAD937FFAF0789FEF5FEFDFEFE" box="[195,264,345,364]" pageId="13" pageNumber="118">Chinese</collectingCounty>
character}
<date id="FE3A7186D937FFAF0620FEF5FE11FEFE" box="[362,484,345,364]" pageId="13" pageNumber="118" value="2004-09-12" valueMax="2004-09-13" valueMin="2004-09-12">
<collectingDate id="EE7E886ED937FFAF0620FEF5FE11FEFE" box="[362,484,345,364]" pageId="13" pageNumber="118" value="2004-09-12" valueMax="2004-09-13" valueMin="2004-09-12">12-13.ix.2004</collectingDate>
</date>
,
<collectorName id="27713290D937FFAF06A1FEF5FDA5FEFE" box="[491,592,345,364]" pageId="13" pageNumber="118">A. Kawada</collectorName>
(NSMT)
</materialsCitation>
.
<typeStatus id="553FE9E4D937FFAF05E3FEF5FD0DFEF8" box="[681,760,345,364]" pageId="13" pageNumber="118" type="paratype">PARATYPE</typeStatus>
:
<specimenCount id="9C829CCFD937FFAF0449FEF5FCD7FEF9" box="[771,802,338,364]" pageId="13" pageNumber="118" type="female">1 ♀</specimenCount>
, same label as
<typeStatus id="553FE9E4D937FFAF04E8FEF5FC18FEFE" box="[930,1005,345,364]" pageId="13" pageNumber="118" type="holotype">holotype</typeStatus>
(NSMT).
</paragraph>
</subSubSection>
<subSubSection id="C29E04CDD937FFAF073BFE29FC5DFD8E" pageId="13" pageNumber="118" type="diagnosis">
<paragraph id="8A3B5746D937FFAF073BFE29FC5DFD8E" blockId="13.[113,1089,389,1052]" pageId="13" pageNumber="118">
<emphasis id="B8F08B54D937FFAF073BFE29FF13FE0E" bold="true" box="[113,230,389,412]" pageId="13" pageNumber="118">Diagnosis.</emphasis>
<emphasis id="B8F08B54D937FFAF07A1FE29FED7FE0E" box="[235,290,389,412]" italics="true" pageId="13" pageNumber="118">Male</emphasis>
: vertex simple, lacking modification; antennomeres VIIIXI modified; each eye composed of about 15 facets; pronotum lacking lateral spines; metaventrite slightly raised above metacoxae to form humps, with short row of dense setae on each hump; aedeagus with apical portion of the median lobe split to two parts, dorsal part as long as ventral part.
<emphasis id="B8F08B54D937FFAF04A3FE49FBCCFE6E" box="[1001,1081,485,508]" italics="true" pageId="13" pageNumber="118">Female</emphasis>
: Each eye with about 8 facets; may be identified by the association with male.
</paragraph>
</subSubSection>
<subSubSection id="C29E04CDD937FFAF073BFD89FEFDFC4E" pageId="13" pageNumber="118" type="description">
<paragraph id="8A3B5746D937FFAF073BFD89FE27FCAE" blockId="13.[113,1089,389,1052]" pageId="13" pageNumber="118">
<emphasis id="B8F08B54D937FFAF073BFD89FF0AFDAE" bold="true" box="[113,255,549,572]" pageId="13" pageNumber="118">Description.</emphasis>
<emphasis id="B8F08B54D937FFAF064CFD89FECBFDAE" box="[262,318,549,572]" italics="true" pageId="13" pageNumber="118">Male</emphasis>
. Body (
<figureCitation id="12BF4BC3D937FFAF06DCFD89FE19FDAE" box="[406,492,549,572]" captionStart="Fig" captionStartId="14.[151,182,1203,1222]" captionTargetBox="[156,1125,218,1179]" captionTargetId="figure-119@14.[151,1126,218,1179]" captionTargetPageId="14" captionText="Fig.8. Tribasodites kawadai sp. nov., male.A habitus. B left antenna. C meso- and metaventrite.D aedeagus, in lateral view. E aedeagus, in ventral view. Scales: A = 1.0 mm; C = 0.5 mm; B, D, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5363103" httpUri="https://zenodo.org/record/5363103/files/figure.png" pageId="13" pageNumber="118">Fig. 8A</figureCitation>
) reddish brown, mouth parts, antennae and tarsi paler; BL
<quantity id="4D7CFAA3D937FFAF07D6FDE9FEF4FDCE" box="[156,257,581,604]" metricMagnitude="-3" metricUnit="m" metricValue="2.96" pageId="13" pageNumber="118" unit="mm" value="2.96">2.96 mm</quantity>
. HL
<quantity id="4D7CFAA3D937FFAF0670FDE9FE6AFDCE" box="[314,415,581,604]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="13" pageNumber="118" unit="mm" value="0.59">0.59 mm</quantity>
, HW
<quantity id="4D7CFAA3D937FFAF06ABFDE9FDB3FDCE" box="[481,582,581,604]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="13" pageNumber="118" unit="mm" value="0.59">0.59 mm</quantity>
; median longitudinal carina vague, extending from base to level of eyes; each eye composed of about 15 facets; antennomeres VIIIIX modified (
<figureCitation id="12BF4BC3D937FFAF0795FD29FEC7FD0E" box="[223,306,645,668]" captionStart="Fig" captionStartId="14.[151,182,1203,1222]" captionTargetBox="[156,1125,218,1179]" captionTargetId="figure-119@14.[151,1126,218,1179]" captionTargetPageId="14" captionText="Fig.8. Tribasodites kawadai sp. nov., male.A habitus. B left antenna. C meso- and metaventrite.D aedeagus, in lateral view. E aedeagus, in ventral view. Scales: A = 1.0 mm; C = 0.5 mm; B, D, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5363103" httpUri="https://zenodo.org/record/5363103/files/figure.png" pageId="13" pageNumber="118">Fig. 8B</figureCitation>
). Pronotum slightly longer than wide, PL
<quantity id="4D7CFAA3D937FFAF05A4FD29FCA5FD0E" box="[750,848,645,668]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="13" pageNumber="118" unit="mm" value="0.67">0.67 mm</quantity>
, PW
<quantity id="4D7CFAA3D937FFAF04CCFD29FC12FD0E" box="[902,999,645,668]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="13" pageNumber="118" unit="mm" value="0.64">0.64 mm</quantity>
; lacking discal and lateral spines. Elytra slightly wider than long, EL
<quantity id="4D7CFAA3D937FFAF05B0FD09FCA9FD2E" box="[762,860,677,700]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="13" pageNumber="118" unit="mm" value="0.86">0.86 mm</quantity>
, EW
<quantity id="4D7CFAA3D937FFAF04DFFD09FC02FD2E" box="[917,1015,677,700]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="13" pageNumber="118" unit="mm" value="0.92">0.92 mm</quantity>
; discal striae extending to more than half elytral length. Metaventrite (
<figureCitation id="12BF4BC3D937FFAF0471FD69FC67FD4E" box="[827,914,709,732]" captionStart="Fig" captionStartId="14.[151,182,1203,1222]" captionTargetBox="[156,1125,218,1179]" captionTargetId="figure-119@14.[151,1126,218,1179]" captionTargetPageId="14" captionText="Fig.8. Tribasodites kawadai sp. nov., male.A habitus. B left antenna. C meso- and metaventrite.D aedeagus, in lateral view. E aedeagus, in ventral view. Scales: A = 1.0 mm; C = 0.5 mm; B, D, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5363103" httpUri="https://zenodo.org/record/5363103/files/figure.png" pageId="13" pageNumber="118">Fig. 8C</figureCitation>
) slightly raised above metacoxae, with pair of short rows of dense setae; metacoxae and metatrochanters simple. Abdomen narrower than elytra, AL
<quantity id="4D7CFAA3D937FFAF051EFCA9FD4DFC8E" box="[596,696,773,796]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="13" pageNumber="118" unit="mm" value="0.84">0.84 mm</quantity>
, AW
<quantity id="4D7CFAA3D937FFAF05BCFCA9FCAFFC8E" box="[758,858,773,796]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="13" pageNumber="118" unit="mm" value="0.94">0.94 mm</quantity>
. Aedeagus (
<figureCitation id="12BF4BC3D937FFAF04AAFCA9FF76FCAE" captionStart="Fig" captionStartId="14.[151,182,1203,1222]" captionTargetBox="[156,1125,218,1179]" captionTargetId="figure-119@14.[151,1126,218,1179]" captionTargetPageId="14" captionText="Fig.8. Tribasodites kawadai sp. nov., male.A habitus. B left antenna. C meso- and metaventrite.D aedeagus, in lateral view. E aedeagus, in ventral view. Scales: A = 1.0 mm; C = 0.5 mm; B, D, E = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5363103" httpUri="https://zenodo.org/record/5363103/files/figure.png" pageId="13" pageNumber="118">Figs 8D, E</figureCitation>
) asymmetric, length
<quantity id="4D7CFAA3D937FFAF0620FC89FE3BFCAE" box="[362,462,805,828]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="13" pageNumber="118" unit="mm" value="0.81">0.81 mm</quantity>
.
</paragraph>
<paragraph id="8A3B5746D937FFAF07DAFCE9FEFDFC4E" blockId="13.[113,1089,389,1052]" pageId="13" pageNumber="118">
<emphasis id="B8F08B54D937FFAF07DAFCE9FF15FCCE" box="[144,224,837,860]" italics="true" pageId="13" pageNumber="118">Female</emphasis>
. Similar to male in general; each eye with about 8 facets; antennae and metaventrite simple. Tergite VIII (Fig. 12P) and sternite VIII (Fig. 12Q) transverse; width of genital complex excluding lateral arms (Fig. 12R)
<quantity id="4D7CFAA3D937FFAF0574FC29FD55FC0E" box="[574,672,901,924]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="118" unit="mm" value="0.17">0.17 mm</quantity>
. Measurements: BL
<quantity id="4D7CFAA3D937FFAF0436FC29FC2BFC0E" box="[892,990,901,924]" metricMagnitude="-3" metricUnit="m" metricValue="2.99" pageId="13" pageNumber="118" unit="mm" value="2.99">2.99 mm</quantity>
; HL
<quantity id="4D7CFAA3D937FFAF0358FC29FF68FC2E" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="13" pageNumber="118" unit="mm" value="0.64">0.64 mm</quantity>
, HW
<quantity id="4D7CFAA3D937FFAF0797FC09FECAFC2E" box="[221,319,933,956]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="118" unit="mm" value="0.6">0.60 mm</quantity>
, PL
<quantity id="4D7CFAA3D937FFAF063BFC09FE26FC2E" box="[369,467,933,956]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="13" pageNumber="118" unit="mm" value="0.67">0.67 mm</quantity>
, PW
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, EL
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, EW
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, AL
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, AW
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.
</paragraph>
</subSubSection>
<subSubSection id="C29E04CDD937FFAF073BFC49FC28FC6E" box="[113,989,997,1020]" pageId="13" pageNumber="118" type="etymology">
<paragraph id="8A3B5746D937FFAF073BFC49FC28FC6E" blockId="13.[113,1089,389,1052]" box="[113,989,997,1020]" pageId="13" pageNumber="118">
<emphasis id="B8F08B54D937FFAF073BFC49FF07FC6E" bold="true" box="[113,242,997,1020]" pageId="13" pageNumber="118">Etymology.</emphasis>
The new species is named after A. Kawada, collector of the
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</paragraph>
</subSubSection>
<subSubSection id="C29E04CDD937FFAF073BFBA9FD82FB8E" box="[113,631,1029,1052]" pageId="13" pageNumber="118" type="distribution">
<paragraph id="8A3B5746D937FFAF073BFBA9FD82FB8E" blockId="13.[113,1089,389,1052]" box="[113,631,1029,1052]" pageId="13" pageNumber="118">
<emphasis id="B8F08B54D937FFAF073BFBA9FEF1FB8E" bold="true" box="[113,260,1029,1052]" pageId="13" pageNumber="118">Distribution.</emphasis>
South
<collectingCountry id="F29317D6D937FFAF0605FBA9FE66FB8E" box="[335,403,1029,1052]" name="China" pageId="13" pageNumber="118">China</collectingCountry>
:
<collectingRegion id="484099A4D937FFAF06D7FBA9FE0CFB8E" box="[413,505,1029,1052]" country="China" name="Guangxi" pageId="13" pageNumber="118">Guangxi</collectingRegion>
(
<figureCitation id="12BF4BC3D937FFAF054DFBA9FD9EFB8E" box="[519,619,1029,1052]" captionStart="Fig" captionStartId="20.[151,182,1225,1244]" captionTargetBox="[173,1105,219,1205]" captionTargetId="figure-10@20.[173,1106,219,1206]" captionTargetPageId="20" captionText="Fig. 13. Distribution of the Tribasodites bedosae species-group in China (A) and Thailand (B)." figureDoi="http://doi.org/10.5281/zenodo.5363109" httpUri="https://zenodo.org/record/5363109/files/figure.png" pageId="13" pageNumber="118">Fig. 13A</figureCitation>
).
</paragraph>
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</treatment>
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