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<document id="D1525910A4C0A1E5B95C09E45A889091" ID-CLB-Dataset="86221" ID-DOI="10.11646/zootaxa.5159.3.1" ID-GBIF-Dataset="e19dda1c-86af-404d-9595-c08d8c848abd" ID-ISSN="1175-5326" ID-Zenodo-Dep="6781669" ID-ZooBank="13308111-EF49-4710-9C45-CF69DABE2C5D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656566366326" checkinUser="plazi" docAuthor="Kataev, Boris M., Hongbin, Liang &amp; Wrase, David W." docDate="2022" docId="D364737A241B170DFF60B32F161E1BC3" docLanguage="en" docName="zootaxa.5159.3.1.pdf" docOrigin="Zootaxa 5159 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichotichnus doiinthanonensis Ito 1997" docType="treatment" docVersion="7" lastPageNumber="326" masterDocId="2F5D0B02240C1714FFF7B578133B1A36" masterDocTitle="New data on carabid beetles of Trichotichnus s. str. (Coleoptera: Carabidae) of Yunnan (China) and adjacent areas, with description of six new species and two new subspecies" masterLastPageNumber="353" masterPageNumber="301" pageNumber="324" updateTime="1699353733348" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="50AE458027B24A4BE3378FBD4461C2D1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DDB9B74C0119AA5A3EFA5ABB2355A544">
<mods:title id="E9A5BC75F910269FDD7147A498236AB6">New data on carabid beetles of Trichotichnus s. str. (Coleoptera: Carabidae) of Yunnan (China) and adjacent areas, with description of six new species and two new subspecies</mods:title>
</mods:titleInfo>
<mods:name id="1CDBB31C0F595F7C3A3B16B2B1950BA5" type="personal">
<mods:role id="7956F3916723E5058C02DEA42B0AA36E">
<mods:roleTerm id="FF507E5CA1DD4944E53B080E69601E21">Author</mods:roleTerm>
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<mods:namePart id="7B3796143CE22D43FA861C67F546F00F">Kataev, Boris M.</mods:namePart>
<mods:affiliation id="42378E276E0C48962620E97E5E0AE75C">Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St. Petersburg 199034, Russia.</mods:affiliation>
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<mods:role id="2D62D082838384EA04585406094402D7">
<mods:roleTerm id="863DEF9DA85164BD853FA4C11F5FC040">Author</mods:roleTerm>
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<mods:namePart id="0233EFE31E7F4DE434C344064DC63CD3">Hongbin, Liang</mods:namePart>
<mods:nameIdentifier id="AF7DFDB049C2A94EC532ADA801D21381" type="ORCID">0000-0002-9668-1167</mods:nameIdentifier>
<mods:affiliation id="0C10C1882E8CD5DE3DBE224BAE752661">Key laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China. lianghb @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 9668 - 1167</mods:affiliation>
<mods:nameIdentifier id="5856D9389E0460BC84EDF75AD9565F71" type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
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<mods:name id="03833BA82A7FEA6FF4127E3AE31212A6" type="personal">
<mods:role id="EF0B03D6B2C33DA5C7942ABEAC09AAD9">
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<mods:namePart id="13503F03C7F3FF3CF07B28879274D736">Wrase, David W.</mods:namePart>
<mods:affiliation id="1C3F4AD5825BA47FE7D38A304211C274">Oderstr. 2, D- 15306 Gusow-Platkow, Germany.</mods:affiliation>
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<mods:typeOfResource id="D26A80AEE647FA8C6EAAF6F06D5B78DD">text</mods:typeOfResource>
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<mods:title id="8464BA25C8304B8522437D223F33EB71">Zootaxa</mods:title>
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<mods:part id="EE76338796538080B68647ADBC184C80">
<mods:date id="54D2620ED2F0AF292752747D6638D074">2022</mods:date>
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<mods:number id="715DEC0A2C3F7308D5BF1D676EA571E8">2022-06-30</mods:number>
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<mods:number id="35A87D8288EBFF81CE37F69396872451">5159</mods:number>
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<mods:classification id="2683FF12779EF2744A976BB1B791DA3F">journal article</mods:classification>
<mods:identifier id="8832058D3F461BAB563A07B3E9A1589D" type="CLB-Dataset">86221</mods:identifier>
<mods:identifier id="98251467851453AA70B20FB652987E5F" type="DOI">10.11646/zootaxa.5159.3.1</mods:identifier>
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<treatment id="D364737A241B170DFF60B32F161E1BC3" ID-DOI="http://doi.org/10.5281/zenodo.6787405" ID-GBIF-Taxon="196373191" ID-Zenodo-Dep="6787405" LSID="urn:lsid:plazi:treatment:D364737A241B170DFF60B32F161E1BC3" httpUri="http://treatment.plazi.org/id/D364737A241B170DFF60B32F161E1BC3" lastPageId="25" lastPageNumber="326" pageId="23" pageNumber="324">
<subSubSection id="13D791E7241B1703FF60B32F11D11C44" box="[151,746,1623,1650]" pageId="23" pageNumber="324" type="nomenclature">
<paragraph id="5B72C26C241B1703FF60B32F11D11C44" blockId="23.[151,746,1623,1687]" box="[151,746,1623,1650]" pageId="23" pageNumber="324">
<heading id="003A7500241B1703FF60B32F11D11C44" bold="true" box="[151,746,1623,1650]" fontSize="11" level="1" pageId="23" pageNumber="324" reason="1">
<taxonomicName id="9CCDB9EF241B1703FF60B32F11D11C44" ID-CoL="58LKV" authority="Ito, 1997" authorityName="Ito" authorityYear="1997" box="[151,746,1623,1650]" class="Insecta" family="Carabidae" genus="Trichotichnus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="doiinthanonensis">
<emphasis id="69B91E7E241B1703FF60B32F11D11C44" bold="true" box="[151,746,1623,1650]" pageId="23" pageNumber="324">
<emphasis id="69B91E7E241B1703FF60B32F127A1C44" bold="true" box="[151,321,1623,1650]" italics="true" pageId="23" pageNumber="324">Trichotichnus</emphasis>
(s. str.)
<emphasis id="69B91E7E241B1703FE53B32F114E1C44" bold="true" box="[420,629,1623,1650]" italics="true" pageId="23" pageNumber="324">doiinthanonensis</emphasis>
<bibRefCitation id="3F5CBF9D241B1703FD8BB32011D11C44" author="Ito, N." box="[636,746,1624,1650]" pageId="23" pageNumber="324" pagination="45 - 49" refId="ref38089" refString="Ito, N. (1997) Two new species of the genus Trichotichnus from Mt. Doi Inthanon in North Thailand (Coleoptera, Carabidae, Harpalini). The Entomological Review of Japan, 52 (1), 45 - 49." type="journal article" year="1997">Ito, 1997</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="13D791E7241B1703FF60B30412A31CA0" box="[151,408,1660,1687]" pageId="23" pageNumber="324" type="description">
<paragraph id="5B72C26C241B1703FF60B30412A31CA0" blockId="23.[151,746,1623,1687]" box="[151,408,1660,1687]" pageId="23" pageNumber="324">
(
<figureCitation id="C3F6DEE9241B1703FF68B30413C61CA0" box="[159,253,1660,1686]" captionStart="FIGURES 35–37" captionStartId="20.[151,264,1335,1359]" captionTargetBox="[158,1428,536,1307]" captionTargetId="figure-178@20.[150,1437,525,1312]" captionTargetPageId="20" captionText="FIGURES 35–37. Trichotichnus spp., habitus. 35, T. cyrtops (Tschitschérine, 1906) (Yunnan); 36, T. doiinthanonensis Ito, 1997 (Dulongjiang); 37, T. fedorenkoi sp. n. (Tam Dao). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781715" httpUri="https://zenodo.org/record/6781715/files/figure.png" pageId="23" pageNumber="324">Figs. 36</figureCitation>
,
<figureCitation id="C3F6DEE9241B1703FEFEB30412741CA1" box="[265,335,1660,1687]" captionStart="FIGURES 47–52" captionStartId="24.[151,264,1260,1284]" captionTargetBox="[158,1430,570,1230]" captionTargetId="figure-200@24.[151,1436,561,1237]" captionTargetPageId="24" captionText="FIGURES 47–52. Trichotichnus doiinthanonensis Ito, 1997 (47–49, Yunnan, Xishuangbanna; 50–52, Yunnan, Dulongjiang). 47 &amp; 50, terminal lamella of aedeagus; 48, 49, 51 &amp; 52, median lobe of aedeagus. Dorsal (47, 49, 50 &amp; 52) and lateral (48 &amp; 51) views. Scale bars: A = 0.5 mm (47 &amp; 50), B = 1.0 mm (48, 49, 51 &amp; 52)." figureDoi="http://doi.org/10.5281/zenodo.6781725" httpUri="https://zenodo.org/record/6781725/files/figure.png" pageId="23" pageNumber="324">47–52</figureCitation>
&amp;
<figureCitation id="C3F6DEE9241B1703FE84B30412AB1CA0" box="[371,400,1660,1686]" captionStart="FIGURE 73" captionStartId="31.[151,250,1841,1865]" captionTargetBox="[151,1435,181,1817]" captionTargetId="figure-19@31.[151,1435,181,1817]" captionTargetPageId="31" captionText="FIGURE 73. Trichotichnus spp., distribution: T. doiinthanonensis Ito, 1997 (circles); T. fedorenkoi sp. n. (squares)." figureDoi="http://doi.org/10.5281/zenodo.6781743" httpUri="https://zenodo.org/record/6781743/files/figure.png" pageId="23" pageNumber="324">73</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="13D791E7241B1703FF60B3B8118A1CEF" box="[151,689,1728,1753]" pageId="23" pageNumber="324" type="reference_group">
<paragraph id="5B72C26C241B1703FF60B3B8118A1CEF" blockId="23.[151,689,1728,1753]" box="[151,689,1728,1753]" pageId="23" pageNumber="324">
<treatmentCitationGroup id="7BDDE542241B1703FF60B3B8118A1CEF" box="[151,689,1728,1753]" pageId="23" pageNumber="324">
<treatmentCitation id="DA6CE47D241B1703FF60B3B811961CEF" author="Ito, N." box="[151,685,1728,1753]" page="48" pageId="23" pageNumber="324" year="1997">
<taxonomicName id="9CCDB9EF241B1703FF60B3B811961CEF" ID-CoL="58LKV" authority="Ito, 1997: 48" authorityName="Ito" authorityPageNumber="48" authorityYear="1997" box="[151,685,1728,1753]" class="Insecta" family="Carabidae" genus="Trichotichnus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="doiinthanonensis">
<emphasis id="69B91E7E241B1703FF60B3B812191CEE" box="[151,290,1728,1752]" italics="true" pageId="23" pageNumber="324">Trichotichnus</emphasis>
(s. str.)
<emphasis id="69B91E7E241B1703FE82B3B811191CEE" box="[373,546,1728,1752]" italics="true" pageId="23" pageNumber="324">doiinthanonensis</emphasis>
<bibRefCitation id="3F5CBF9D241B1703FDDFB3B911961CEF" author="Ito, N." box="[552,685,1729,1753]" pageId="23" pageNumber="324" pagination="45 - 49" refId="ref38089" refString="Ito, N. (1997) Two new species of the genus Trichotichnus from Mt. Doi Inthanon in North Thailand (Coleoptera, Carabidae, Harpalini). The Entomological Review of Japan, 52 (1), 45 - 49." type="journal article" year="1997">Ito, 1997: 48</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="13D791E7241B170CFF60B27E176C1C13" lastPageId="24" lastPageNumber="325" pageId="23" pageNumber="324" type="materials_examined">
<paragraph id="5B72C26C241B1703FF60B27E16091D5F" blockId="23.[151,1437,1798,2041]" pageId="23" pageNumber="324">
<emphasis id="69B91E7E241B1703FF60B27E12B91D16" bold="true" box="[151,386,1798,1824]" pageId="23" pageNumber="324">Material examined.</emphasis>
<materialsCitation id="EBA5C831241B1703FE7DB27E10BE1D73" ID-GBIF-Occurrence="3821894404" collectingDate="2006-08-29" collectionCode="IOZ" collectorName="D. Kavanaugh" country="China" elevation="1845" latitude="27.6831" location="Dulongjiang" longLatPrecision="1" longitude="98.30038" municipality="Gongshan" pageId="23" pageNumber="324" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<emphasis id="69B91E7E241B1703FE7DB27E12E11D16" bold="true" box="[394,474,1798,1824]" pageId="23" pageNumber="324">
<collectingCountry id="23DA82FC241B1703FE7DB27E12EE1D16" box="[394,469,1798,1824]" name="China" pageId="23" pageNumber="324">China</collectingCountry>
.
</emphasis>
<collectingRegion id="99090C8E241B1703FE15B27F11601D17" box="[482,603,1799,1825]" country="China" name="Yunnan" pageId="23" pageNumber="324">YUNNAN</collectingRegion>
:
<specimenCount id="4DCB09E5241B1703FD9CB27F11811D17" box="[619,698,1799,1825]" pageId="23" pageNumber="324" type="male">1 male</specimenCount>
,
<collectingMunicipality id="BB165816241B1703FD30B27F10001D17" box="[711,827,1799,1825]" pageId="23" pageNumber="324">Gongshan</collectingMunicipality>
,
<location id="5E1294B7241B1703FCBFB27F10EC1D17" LSID="urn:lsid:plazi:treatment:D364737A241B170DFF60B32F161E1BC3:5E1294B7241B1703FCBFB27F10EC1D17" box="[840,983,1799,1825]" country="China" latitude="27.6831" longLatPrecision="1" longitude="98.30038" municipality="Gongshan" name="Dulongjiang" pageId="23" pageNumber="324" stateProvince="Yunnan">Dulongjiang</location>
,
<locationDeviation id="4AC0DB4E241B1703FC14B27E17F01D17" box="[995,1227,1798,1825]" location="Maku" pageId="23" pageNumber="324">
<quantity id="9C356F89241B1703FC14B27E17081D17" box="[995,1075,1798,1825]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="324" unit="km" value="0.5">0.5 km</quantity>
WSW Maku
</locationDeviation>
, trail,
<geoCoordinate id="3EF9A4AB241B1703FAE3B27F16A71D17" box="[1300,1436,1799,1825]" degrees="27.68310" direction="north" orientation="latitude" pageId="23" pageNumber="324" precision="1" value="27.6831">27.68310°N</geoCoordinate>
<geoCoordinate id="3EF9A4AB241B1703FF60B253121B1D73" box="[151,288,1835,1861]" degrees="98.30038" direction="east" orientation="longitude" pageId="23" pageNumber="324" precision="1" value="98.30038">98.30038°E</geoCoordinate>
,
<quantity id="9C356F89241B1703FEDCB25212B81D73" box="[299,387,1834,1861]" metricMagnitude="3" metricUnit="m" metricValue="1.845" pageId="23" pageNumber="324" unit="m" value="1845.0">
<elevation id="D0E0255F241B1703FEDCB25212B81D73" box="[299,387,1834,1861]" metricMagnitude="3" metricUnit="m" metricValue="1.845" pageId="23" pageNumber="324" unit="m" value="1845.0">1845 m</elevation>
</quantity>
, day,
<date id="2F73E4AC241B1703FE32B25311611D73" box="[453,602,1835,1861]" pageId="23" pageNumber="324" value="2006-08-29">
<collectingDate id="3F371D44241B1703FE32B25311611D73" box="[453,602,1835,1861]" pageId="23" pageNumber="324" value="2006-08-29">29.VIII.2006</collectingDate>
</date>
,
<collectorName id="F638A7BA241B1703FD91B25310321D73" box="[614,777,1835,1861]" pageId="23" pageNumber="324">D. Kavanaugh</collectorName>
leg. (
<collectionCode id="3DDC5AA9241B1703FCBEB25310461D73" box="[841,893,1835,1861]" pageId="23" pageNumber="324">IOZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="EBA5C831241B1703FC65B25312E61D5F" ID-GBIF-Occurrence="3821894503" collectionCode="NNNR" collectorName="Xishuangbanna &amp; Guo Men Shan" country="China" location="Jinghong" pageId="23" pageNumber="324" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<specimenCount id="4DCB09E5241B1703FC65B25310DB1D73" box="[914,992,1835,1861]" pageId="23" pageNumber="324" type="male">1 male</specimenCount>
,
<collectorName id="F638A7BA241B1703FC1AB25317A51D73" box="[1005,1182,1835,1861]" pageId="23" pageNumber="324">Xishuangbanna</collectorName>
,
<quantity id="9C356F89241B1703FB5EB25217CA1D73" box="[1193,1265,1834,1861]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="23" pageNumber="324" unit="km" value="35.0">35 km</quantity>
NW
<location id="5E1294B7241B1703FAD8B25316A21D73" LSID="urn:lsid:plazi:treatment:D364737A241B170DFF60B32F161E1BC3:5E1294B7241B1703FAD8B25316A21D73" box="[1327,1433,1835,1861]" country="China" name="Jinghong" pageId="23" pageNumber="324" stateProvince="Yunnan">Jinghong</location>
, vic.
<collectorName id="F638A7BA241B1703FF30B237124A1D5F" box="[199,369,1871,1897]" pageId="23" pageNumber="324">Guo Men Shan</collectorName>
(
<collectionCode id="3DDC5AA9241B1703FE77B23712EE1D5F" box="[384,469,1871,1897]" pageId="23" pageNumber="324">NNNR</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="EBA5C831241B1703FE1FB23716151D5F" ID-GBIF-Occurrence="3821894417" box="[488,1326,1871,1899]" collectingDate="2019-06-18" collectionCode="NME" collectorName="A. Weigel" country="China" elevation="1110" latitude="22.2405" longLatPrecision="12" longitude="100.602" pageId="23" pageNumber="324" specimenCount="1" stateProvince="Yunnan">
<quantity id="9C356F89241B1703FE1FB23711061D5F" box="[488,573,1871,1897]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="23" pageNumber="324" unit="m" value="1110.0">
<elevation id="D0E0255F241B1703FE1FB23711061D5F" box="[488,573,1871,1897]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="23" pageNumber="324" unit="m" value="1110.0">1110 m</elevation>
</quantity>
,
<geoCoordinate id="3EF9A4AB241B1703FDBEB22811F21D5D" box="[585,713,1872,1899]" degrees="22" direction="north" minutes="14.43" orientation="latitude" pageId="23" pageNumber="324" precision="9" value="22.2405">22°14.43′N</geoCoordinate>
<geoCoordinate id="3EF9A4AB241B1703FD27B22810651D5D" box="[720,862,1872,1899]" degrees="100" direction="east" minutes="36.12" orientation="longitude" pageId="23" pageNumber="324" precision="9" value="100.602">100°36.12′E</geoCoordinate>
,
<date id="2F73E4AC241B1703FC9DB23710D71D5F" box="[874,1004,1871,1897]" pageId="23" pageNumber="324" value="2019-06-18">
<collectingDate id="3F371D44241B1703FC9DB23710D71D5F" box="[874,1004,1871,1897]" pageId="23" pageNumber="324" value="2019-06-18">18.VI.2019</collectingDate>
</date>
, LFF,
<collectorName id="F638A7BA241B1703FBC5B23717991D5F" box="[1074,1186,1871,1897]" pageId="23" pageNumber="324">A. Weigel</collectorName>
leg. (
<collectionCode id="3DDC5AA9241B1703FB16B237161E1D5F" box="[1249,1317,1871,1897]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="23" pageNumber="324">NME</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5B72C26C241B1703FF30B20A12B11D87" blockId="23.[151,1437,1798,2041]" pageId="23" pageNumber="324">
<materialsCitation id="EBA5C831241B1703FF30B20A12BD1D87" ID-GBIF-Occurrence="3821894385" collectingDate="1997-08-05" collectingDateMax="1997-08-26" collectingDateMin="1997-08-05" collectionCode="SIEE" collectorName="M. Kalyakin" country="Vietnam" elevation="700" location="Nature Reserve" pageId="23" pageNumber="324" specimenCount="1" specimenCount-male="1" stateProvince="Ha Tinh">
<emphasis id="69B91E7E241B1703FF30B20A120F1DBA" bold="true" box="[199,308,1906,1932]" pageId="23" pageNumber="324">
<collectingCountry id="23DA82FC241B1703FF30B20A12141DBA" box="[199,303,1906,1932]" name="Vietnam" pageId="23" pageNumber="324">Vietnam</collectingCountry>
.
</emphasis>
<collectingRegion id="99090C8E241B1703FECCB20B12941DBB" box="[315,431,1907,1933]" country="Vietnam" name="Ha Tinh" pageId="23" pageNumber="324">HA TINH</collectingRegion>
:
<specimenCount id="4DCB09E5241B1703FE4DB20B11331DBB" box="[442,520,1907,1933]" pageId="23" pageNumber="324" type="male">1 male</specimenCount>
,
<locationDeviation id="4AC0DB4E241B1703FDE4B20B10231DBB" box="[531,792,1907,1933]" location="Vu Quang" pageId="23" pageNumber="324">
<quantity id="9C356F89241B1703FDE4B20B11611DBB" box="[531,602,1907,1933]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="23" pageNumber="324" unit="km" value="30.0">30 km</quantity>
SE of Vu Quang
</locationDeviation>
(Phu Quon),
<location id="5E1294B7241B1703FC47B20B175A1DBB" LSID="urn:lsid:plazi:treatment:D364737A241B170DFF60B32F161E1BC3:5E1294B7241B1703FC47B20B175A1DBB" box="[944,1121,1907,1933]" country="Vietnam" name="Nature Reserve" pageId="23" pageNumber="324" stateProvince="Ha Tinh">Nature Reserve</location>
,
<quantity id="9C356F89241B1703FB9BB20B178E1DBB" box="[1132,1205,1907,1933]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="23" pageNumber="324" unit="m" value="700.0">
<elevation id="D0E0255F241B1703FB9BB20B178E1DBB" box="[1132,1205,1907,1933]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="23" pageNumber="324" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<date id="2F73E4AC241B1703FB37B20A164A1DBB" box="[1216,1393,1906,1933]" pageId="23" pageNumber="324" value="1997-08-05" valueMax="1997-08-26" valueMin="1997-08-05">
<collectingDate id="3F371D44241B1703FB37B20A164A1DBB" box="[1216,1393,1906,1933]" pageId="23" pageNumber="324" value="1997-08-05" valueMax="1997-08-26" valueMin="1997-08-05">5–26.VIII.1997</collectingDate>
</date>
,
<collectorName id="F638A7BA241B1703FA8BB20B13C51D87" pageId="23" pageNumber="324">M. Kalyakin</collectorName>
leg. (
<collectionCode id="3DDC5AA9241B1703FECAB2EF12451D87" box="[317,382,1943,1969]" pageId="23" pageNumber="324">SIEE</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5B72C26C241B1703FF30B2C217801DCF" blockId="23.[151,1437,1798,2041]" pageId="23" pageNumber="324">
<emphasis id="69B91E7E241B1703FF30B2C212451DE2" bold="true" box="[199,382,1978,2004]" pageId="23" pageNumber="324">Re-description.</emphasis>
Body length in the examined specimens
<quantity id="9C356F89241B1703FCACB2C210DC1DE3" box="[859,999,1978,2005]" metricMagnitude="-3" metricUnit="m" metricValue="8.35" metricValueMax="8.5" metricValueMin="8.2" pageId="23" pageNumber="324" unit="mm" value="8.35" valueMax="8.5" valueMin="8.2">8.2–8.5 mm</quantity>
. Habitus as in
<figureCitation id="C3F6DEE9241B1703FB63B2C317D21DE3" box="[1172,1257,1979,2005]" captionStart="FIGURES 35–37" captionStartId="20.[151,264,1335,1359]" captionTargetBox="[158,1428,536,1307]" captionTargetId="figure-178@20.[150,1437,525,1312]" captionTargetPageId="20" captionText="FIGURES 35–37. Trichotichnus spp., habitus. 35, T. cyrtops (Tschitschérine, 1906) (Yunnan); 36, T. doiinthanonensis Ito, 1997 (Dulongjiang); 37, T. fedorenkoi sp. n. (Tam Dao). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.6781715" httpUri="https://zenodo.org/record/6781715/files/figure.png" pageId="23" pageNumber="324">Fig. 36</figureCitation>
. Morphological character states are similar to those as described for
<taxonomicName id="9CCDB9EF241B1703FD28B2A710761DCE" baseAuthorityName="Tschitscherine" baseAuthorityYear="1906" box="[735,845,2015,2040]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="cyrtops">
<emphasis id="69B91E7E241B1703FD28B2A710761DCE" box="[735,845,2015,2040]" italics="true" pageId="23" pageNumber="324">T. cyrtops</emphasis>
</taxonomicName>
, but differing mostly as follows.
</paragraph>
<paragraph id="5B72C26C2414170CFF30B5EF12A31B53" blockId="24.[151,1437,151,537]" pageId="24" pageNumber="325">
Body slenderer and flatter. Supraorbital setigerous pores situated slightly before level of posterior margin of eye. Genae very narrow, narrower than antennomere 3 basally. Paraglossae markedly longer than ligular sclerite. Elytra with distinct but rather shallow preapical sinuation (as in
<figureCitation id="C3F6DEE92414170CFC8EB5A710F51ACF" box="[889,974,223,249]" captionStart="FIGURES 38–39" captionStartId="21.[151,264,1041,1065]" captionTargetBox="[151,1436,669,1016]" captionTargetId="figure-229@21.[150,1437,669,1018]" captionTargetPageId="21" captionText="FIGURES 38–39. Trichotichnus spp., elytral apex. 38, T. cyrtops (Tschitschérine, 1906); 39, T. schmidti sp. n. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6781717" httpUri="https://zenodo.org/record/6781717/files/figure.png" pageId="24" pageNumber="325">Fig. 39</figureCitation>
). Humeri without denticle. Protibia with longitudinal sulcus on dorsal side. Metatarsus about as long as HWmax. Mesotarsomere
<quantity id="9C356F892414170CFB89B47B17911B2B" box="[1150,1194,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="24" pageNumber="325" unit="in" value="1.0">1 in</quantity>
both sexes at least as long as mesotarsomeres 2 and 3 combined, in male without adhesive scales. Pro- and mesotarsomeres 2 and 3 of male longer than wide.
</paragraph>
<paragraph id="5B72C26C2414170CFF30B41710F5182F" blockId="24.[151,1437,151,537]" pageId="24" pageNumber="325">
Median lobe of aedeagus (
<figureCitation id="C3F6DEE92414170CFE0FB41711BA1BBF" box="[504,641,366,393]" captionStart="FIGURES 47–52" captionStartId="24.[151,264,1260,1284]" captionTargetBox="[158,1430,570,1230]" captionTargetId="figure-200@24.[151,1436,561,1237]" captionTargetPageId="24" captionText="FIGURES 47–52. Trichotichnus doiinthanonensis Ito, 1997 (47–49, Yunnan, Xishuangbanna; 50–52, Yunnan, Dulongjiang). 47 &amp; 50, terminal lamella of aedeagus; 48, 49, 51 &amp; 52, median lobe of aedeagus. Dorsal (47, 49, 50 &amp; 52) and lateral (48 &amp; 51) views. Scale bars: A = 0.5 mm (47 &amp; 50), B = 1.0 mm (48, 49, 51 &amp; 52)." figureDoi="http://doi.org/10.5281/zenodo.6781725" httpUri="https://zenodo.org/record/6781725/files/figure.png" pageId="24" pageNumber="325">Figs. 47–52</figureCitation>
) in lateral view arcuate, narrow, angularly bent ventrally just at apex and slightly swollen at tip forming a small apical capitulum; in dorsal view comparatively wide, somewhat parallel-sided, with terminal lamella (
<figureCitation id="C3F6DEE92414170CFDF5B4CF11BC1BE7" box="[514,647,438,465]" captionStart="FIGURES 47–52" captionStartId="24.[151,264,1260,1284]" captionTargetBox="[158,1430,570,1230]" captionTargetId="figure-200@24.[151,1436,561,1237]" captionTargetPageId="24" captionText="FIGURES 47–52. Trichotichnus doiinthanonensis Ito, 1997 (47–49, Yunnan, Xishuangbanna; 50–52, Yunnan, Dulongjiang). 47 &amp; 50, terminal lamella of aedeagus; 48, 49, 51 &amp; 52, median lobe of aedeagus. Dorsal (47, 49, 50 &amp; 52) and lateral (48 &amp; 51) views. Scale bars: A = 0.5 mm (47 &amp; 50), B = 1.0 mm (48, 49, 51 &amp; 52)." figureDoi="http://doi.org/10.5281/zenodo.6781725" httpUri="https://zenodo.org/record/6781725/files/figure.png" pageId="24" pageNumber="325">Figs. 47, 50</figureCitation>
) comparatively short, strongly narrowed posteriorly; its apex narrowly rounded. Ventral side in apical portion edged laterally and depressed between margins. Apical orifice in dorsal position, wide apically. Internal sac without spines and distinct spiny patches.
</paragraph>
<caption id="0FB292E42414170CFF60B19410421F7A" ID-DOI="http://doi.org/10.5281/zenodo.6781725" ID-Zenodo-Dep="6781725" httpUri="https://zenodo.org/record/6781725/files/figure.png" pageId="24" pageNumber="325" startId="24.[151,264,1260,1284]" targetBox="[158,1430,570,1230]" targetPageId="24">
<paragraph id="5B72C26C2414170CFF60B19410421F7A" blockId="24.[151,1437,1260,1356]" pageId="24" pageNumber="325">
<emphasis id="69B91E7E2414170CFF60B194126D1F32" bold="true" box="[151,342,1260,1284]" pageId="24" pageNumber="325">FIGURES 47–52.</emphasis>
<taxonomicName id="9CCDB9EF2414170CFEAAB19410391F32" authority="Ito, 1997" authorityName="Ito" authorityYear="1997" box="[349,770,1260,1284]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="doiinthanonensis">
<emphasis id="69B91E7E2414170CFEAAB19411A61F32" box="[349,669,1260,1284]" italics="true" pageId="24" pageNumber="325">Trichotichnus doiinthanonensis</emphasis>
<bibRefCitation id="3F5CBF9D2414170CFD52B19510391F32" author="Ito, N." box="[677,770,1260,1284]" pageId="24" pageNumber="325" pagination="45 - 49" refId="ref38089" refString="Ito, N. (1997) Two new species of the genus Trichotichnus from Mt. Doi Inthanon in North Thailand (Coleoptera, Carabidae, Harpalini). The Entomological Review of Japan, 52 (1), 45 - 49." type="journal article" year="1997">Ito, 1997</bibRefCitation>
</taxonomicName>
(47–49, Yunnan, Xishuangbanna; 50–52, Yunnan, Dulongjiang). 47 &amp; 50, terminal lamella of aedeagus; 48, 49, 51 &amp; 52, median lobe of aedeagus. Dorsal (47, 49, 50 &amp; 52) and lateral (48 &amp; 51) views. Scale bars: A = 0.5 mm (47 &amp; 50), B = 1.0 mm (48, 49, 51 &amp; 52).
</paragraph>
</caption>
<paragraph id="5B72C26C2414170CFF30B00211481FEB" blockId="24.[151,1437,1402,2041]" pageId="24" pageNumber="325">Proportions (in males from Xishuangbanna and Phu Quon): HWmax/PWmax 0.69–0.73; HWmin/PWmax 0.53–0.54; HWmax/HWmin 1.29–1.36; PWmax/PL 1.45–1.47; PWmax/PWmin 1.19–1.23; EL/EW 1.53–1.57; EL/PL 3.02–3.18; EW/PWmax 1.34–1.40.</paragraph>
<paragraph id="5B72C26C2414170CFF30B09E176C1C13" blockId="24.[151,1437,1402,2041]" pageId="24" pageNumber="325">Proportions (in male from Dulongjiang): HWmax/PWmax 0.69; HWmin/PWmax 0.51; HWmax/HWmin 1.36; PWmax/PL 1.50; PWmax/PWmin 1.15; EL/EW 1.55; EL/PL 3.14; EW/PWmax 1.35.</paragraph>
</subSubSection>
<subSubSection id="13D791E72414170CFF30B35610DB1C5A" pageId="24" pageNumber="325" type="distribution">
<paragraph id="5B72C26C2414170CFF30B35610DB1C5A" blockId="24.[151,1437,1402,2041]" pageId="24" pageNumber="325">
<emphasis id="69B91E7E2414170CFF30B35612621C7E" bold="true" box="[199,345,1582,1608]" pageId="24" pageNumber="325">Distribution</emphasis>
(
<figureCitation id="C3F6DEE92414170CFE9EB35612841C7F" box="[361,447,1582,1609]" captionStart="FIGURE 73" captionStartId="31.[151,250,1841,1865]" captionTargetBox="[151,1435,181,1817]" captionTargetId="figure-19@31.[151,1435,181,1817]" captionTargetPageId="31" captionText="FIGURE 73. Trichotichnus spp., distribution: T. doiinthanonensis Ito, 1997 (circles); T. fedorenkoi sp. n. (squares)." figureDoi="http://doi.org/10.5281/zenodo.6781743" httpUri="https://zenodo.org/record/6781743/files/figure.png" pageId="24" pageNumber="325">Fig. 73</figureCitation>
). Described from
<specimenCount id="4DCB09E52414170CFD7DB356100E1C7E" box="[650,821,1582,1608]" pageId="24" pageNumber="325" type="generic">four specimens</specimenCount>
collected in Doi Inthanon,
<collectingRegion id="99090C8E2414170CFB9CB35617C81C7E" box="[1131,1267,1582,1608]" country="Thailand" name="Chiang Mai" pageId="24" pageNumber="325">Chiang Mai</collectingRegion>
,
<collectingCountry id="23DA82FC2414170CFB08B356165D1C7E" box="[1279,1382,1582,1608]" name="Thailand" pageId="24" pageNumber="325">Thailand</collectingCountry>
. It is recorded here for
<collectingCountry id="23DA82FC2414170CFEA8B32A12991C5A" box="[351,418,1618,1644]" name="China" pageId="24" pageNumber="325">China</collectingCountry>
(
<collectingRegion id="99090C8E2414170CFE47B32B11321C5B" box="[432,521,1619,1645]" country="China" name="Yunnan" pageId="24" pageNumber="325">Yunnan</collectingRegion>
) and
<collectingCountry id="23DA82FC2414170CFDB0B32A119D1C5A" box="[583,678,1618,1644]" name="Vietnam" pageId="24" pageNumber="325">Vietnam</collectingCountry>
(
<collectingRegion id="99090C8E2414170CFD43B32B10281C5A" box="[692,787,1618,1645]" country="Vietnam" name="Ha Tinh" pageId="24" pageNumber="325">Ha Tinh</collectingRegion>
) for the first time.
</paragraph>
</subSubSection>
<subSubSection id="13D791E72414170DFF30B30F161E1BC3" lastPageId="25" lastPageNumber="326" pageId="24" pageNumber="325" type="discussion">
<paragraph id="5B72C26C2414170CFF30B30F12CB1D5E" blockId="24.[151,1437,1402,2041]" pageId="24" pageNumber="325">
<emphasis id="69B91E7E2414170CFF30B30F12001CA7" bold="true" box="[199,315,1655,1681]" pageId="24" pageNumber="325">Remarks.</emphasis>
In the male from southern Yunnan (Xishuangbanna) and
<collectingCountry id="23DA82FC2414170CFC3FB30E171C1CA6" box="[968,1063,1654,1680]" name="Vietnam" pageId="24" pageNumber="325">Vietnam</collectingCountry>
(Phu Quon), the aedeagus (
<figureCitation id="C3F6DEE92414170CFA93B30E13E51C83" captionStart="FIGURES 47–52" captionStartId="24.[151,264,1260,1284]" captionTargetBox="[158,1430,570,1230]" captionTargetId="figure-200@24.[151,1436,561,1237]" captionTargetPageId="24" captionText="FIGURES 47–52. Trichotichnus doiinthanonensis Ito, 1997 (47–49, Yunnan, Xishuangbanna; 50–52, Yunnan, Dulongjiang). 47 &amp; 50, terminal lamella of aedeagus; 48, 49, 51 &amp; 52, median lobe of aedeagus. Dorsal (47, 49, 50 &amp; 52) and lateral (48 &amp; 51) views. Scale bars: A = 0.5 mm (47 &amp; 50), B = 1.0 mm (48, 49, 51 &amp; 52)." figureDoi="http://doi.org/10.5281/zenodo.6781725" httpUri="https://zenodo.org/record/6781725/files/figure.png" pageId="24" pageNumber="325">Figs. 47–49</figureCitation>
) agrees with the original description (
<bibRefCitation id="3F5CBF9D2414170CFD7DB3E310251C83" author="Ito, N." box="[650,798,1691,1717]" pageId="24" pageNumber="325" pagination="45 - 49" refId="ref38089" refString="Ito, N. (1997) Two new species of the genus Trichotichnus from Mt. Doi Inthanon in North Thailand (Coleoptera, Carabidae, Harpalini). The Entomological Review of Japan, 52 (1), 45 - 49." type="journal article" year="1997">Ito, 1997: 48</bibRefCitation>
), and the last visible abdominal sternite (VII) is subtruncate at apex, but in the male from northwestern
<collectingRegion id="99090C8E2414170CFD52B3C711C01CEF" box="[677,763,1727,1753]" country="China" name="Yunnan" pageId="24" pageNumber="325">Yunnan</collectingRegion>
(Dulongjiang), the median lobe (
<figureCitation id="C3F6DEE92414170CFB9BB3C617CA1CEF" box="[1132,1265,1726,1753]" captionStart="FIGURES 47–52" captionStartId="24.[151,264,1260,1284]" captionTargetBox="[158,1430,570,1230]" captionTargetId="figure-200@24.[151,1436,561,1237]" captionTargetPageId="24" captionText="FIGURES 47–52. Trichotichnus doiinthanonensis Ito, 1997 (47–49, Yunnan, Xishuangbanna; 50–52, Yunnan, Dulongjiang). 47 &amp; 50, terminal lamella of aedeagus; 48, 49, 51 &amp; 52, median lobe of aedeagus. Dorsal (47, 49, 50 &amp; 52) and lateral (48 &amp; 51) views. Scale bars: A = 0.5 mm (47 &amp; 50), B = 1.0 mm (48, 49, 51 &amp; 52)." figureDoi="http://doi.org/10.5281/zenodo.6781725" httpUri="https://zenodo.org/record/6781725/files/figure.png" pageId="24" pageNumber="325">Figs. 50–52</figureCitation>
) is slightly less arcuate and with shorter terminal lamella, and the apex of the last visible abdominal sternite is rounded. The male from Dulongjiang can belong to a separate taxon (subspecies or species) distributed in northwestern
<collectingRegion id="99090C8E2414170CFAE4B27F16571D17" box="[1299,1388,1799,1825]" country="China" name="Yunnan" pageId="24" pageNumber="325">Yunnan</collectingRegion>
, but since only
<specimenCount id="4DCB09E52414170CFEE5B25312431D72" box="[274,376,1834,1861]" pageId="24" pageNumber="325" type="male">one male</specimenCount>
is available, additional material from
<collectingCountry id="23DA82FC2414170CFCD0B25210511D72" box="[807,874,1834,1860]" name="China" pageId="24" pageNumber="325">China</collectingCountry>
is needed to determine its taxonomical status and the nature of these differences.
</paragraph>
<paragraph id="5B72C26C2414170DFF30B20A16B51B9B" blockId="24.[151,1437,1402,2041]" lastBlockId="25.[151,1437,151,501]" lastPageId="25" lastPageNumber="326" pageId="24" pageNumber="325">
This and the following four species (
<taxonomicName id="9CCDB9EF2414170CFD9DB20B11DD1DBA" authority="Kataev &amp; Hongbin &amp; Wrase, 2022" authorityName="Kataev &amp; Hongbin &amp; Wrase" authorityYear="2022" box="[618,742,1906,1932]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="schmidti" status="sp. nov.">
<emphasis id="69B91E7E2414170CFD9DB20B11DD1DBA" box="[618,742,1906,1932]" italics="true" pageId="24" pageNumber="325">T. schmidti</emphasis>
</taxonomicName>
<emphasis id="69B91E7E2414170CFD19B20B10151DBB" bold="true" box="[750,814,1907,1933]" pageId="24" pageNumber="325">
<taxonomicNameLabel id="728AA3052414170CFD19B20B10151DBB" box="[750,814,1907,1933]" pageId="24" pageNumber="325" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="9CCDB9EF2414170CFCCAB20B10EA1DBA" authority="Kataev &amp; Hongbin &amp; Wrase, 2022" authorityName="Kataev &amp; Hongbin &amp; Wrase" authorityYear="2022" box="[829,977,1906,1932]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="fedorenkoi" status="sp. nov.">
<emphasis id="69B91E7E2414170CFCCAB20B10EA1DBA" box="[829,977,1906,1932]" italics="true" pageId="24" pageNumber="325">T. fedorenkoi</emphasis>
</taxonomicName>
<emphasis id="69B91E7E2414170CFC2FB20B17221DBB" bold="true" box="[984,1049,1907,1933]" pageId="24" pageNumber="325">
<taxonomicNameLabel id="728AA3052414170CFC2FB20B17221DBB" box="[984,1049,1907,1933]" pageId="24" pageNumber="325" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="9CCDB9EF2414170CFBDFB20B16701DBB" authority="Ito, 2021" authorityName="Ito" authorityYear="2021" box="[1064,1355,1906,1933]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="subangulatus">
<emphasis id="69B91E7E2414170CFBDFB20B17E21DBA" box="[1064,1241,1906,1932]" italics="true" pageId="24" pageNumber="325">T. subangulatus</emphasis>
Ito, 2021
</taxonomicName>
, and
<taxonomicName id="9CCDB9EF2414170CFA7FB20B12871D87" authority="Habu, 1954" authorityName="Habu" authorityYear="1954" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="noctuabundus">
<emphasis id="69B91E7E2414170CFA7FB20B120F1D86" italics="true" pageId="24" pageNumber="325">T. noctuabundus</emphasis>
<bibRefCitation id="3F5CBF9D2414170CFECDB2EE12871D87" author="Habu, A." box="[314,444,1942,1969]" pageId="24" pageNumber="325" pagination="53 - 59" refId="ref37858" refString="Habu, A. (1954) On four new species of Trichotichnus from Mt. Hiko, Kyushu (Coleoptera) (The Carabidae fauna of Mt. Hiko, II). Mushi, 26, 53 - 59." type="journal article" year="1954">Habu, 1954</bibRefCitation>
</taxonomicName>
) are very similar in external morphology and differing mostly in the male genitalia. These species apparently are closely related and represent a natural group (named here from the oldest nominal species as the
<taxonomicName id="9CCDB9EF2414170CFF15B2A612441DCE" authorityName="Habu" authorityYear="1954" box="[226,383,2014,2040]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="noctuabundus">
<emphasis id="69B91E7E2414170CFF15B2A612441DCE" box="[226,383,2014,2040]" italics="true" pageId="24" pageNumber="325">noctuabundus</emphasis>
</taxonomicName>
species group) with the following distinctive features: appendages brownish yellow, femora not darker than tibiae; pronotum with sides not or only hardly sinuate basally and with basal angles obtuse, without prominent denticle; protibia on dorsal side with longitudinal sulcus; elytra with distinct but rather shallow preapical sinuation; tarsi without dense setae on ventral side; mesotarsomere 1 elongate, about as long as mesotarsomeres 2 and 3 combined, in male without adhesive scales ventrally; last visible abdominal sternite in male with one pair of marginal setae, and aedeagus with short and wide terminal lamella and without individual spines in internal sac. All the species of this group are winged and their ranges are at least partly overlapping. In external features, these species are very similar to
<taxonomicName id="9CCDB9EF2415170DFE48B41711101BBE" baseAuthorityName="Tschitscherine" baseAuthorityYear="1906" box="[447,555,367,392]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="cyrtops">
<emphasis id="69B91E7E2415170DFE48B41711101BBE" box="[447,555,367,392]" italics="true" pageId="25" pageNumber="326">T. cyrtops</emphasis>
</taxonomicName>
, but the latter species easily differs from them in having elytra with rather deep preapical sinuation, protibia not sulcate on dorsal side and mesotarsomere 1 of male with adhesive scales at apex.
</paragraph>
<paragraph id="5B72C26C2415170DFF30B4CF161E1BC3" blockId="25.[151,1437,151,501]" pageId="25" pageNumber="326">
<bibRefCitation id="3F5CBF9D2415170DFF30B4CF12761BE7" author="Habu, A." box="[199,333,439,465]" pageId="25" pageNumber="326" pagination="127 - 169" refId="ref37923" refString="Habu, A. (1961) Revisional study of the species of the Trichotichni, the subtribe of the tribe Harpalini from Japan. Bulletin of the National Institute of Agricultural Sciences (Series C), 13, 127 - 169." type="journal article" year="1961">Habu (1961</bibRefCitation>
,
<bibRefCitation id="3F5CBF9D2415170DFEAFB4CF12AA1BE7" author="Habu, A." box="[344,401,439,465]" pageId="25" pageNumber="326" refId="ref37967" refString="Habu, A. (1973) Fauna Japonica. Carabidae: Harpalini (Insecta, Coleoptera). Keigaku Publishing Co., Tokyo, xiii + 430 pp." type="book" year="1973">1973</bibRefCitation>
) included
<taxonomicName id="9CCDB9EF2415170DFDF1B4CF11851BE7" authorityName="Habu" authorityYear="1954" box="[518,702,439,465]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="noctuabundus">
<emphasis id="69B91E7E2415170DFDF1B4CF11851BE7" box="[518,702,439,465]" italics="true" pageId="25" pageNumber="326">T. noctuabundus</emphasis>
</taxonomicName>
in the
<emphasis id="69B91E7E2415170DFCFFB4CF10BB1BE7" box="[776,896,439,465]" italics="true" pageId="25" pageNumber="326">longitarsis</emphasis>
species group, but it differs from other members at least in having legs relatively shorter and pro- and mesotarsi of male with sparser ventro-lateral setae.
</paragraph>
</subSubSection>
</treatment>
</document>
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