672 lines
127 KiB
XML
672 lines
127 KiB
XML
<document id="573A23F97EF8CB799AD32B628631FA01" ID-CLB-Dataset="46773" ID-DOI="10.11646/zootaxa.4078.1.6" ID-GBIF-Dataset="598cb955-c4b7-4c2b-8c74-6ee0348c7f7e" ID-ISSN="1175-5326" ID-Zenodo-Dep="270853" ID-ZooBank="3D0D79C2-2583-47F4-BC01-5CF347614B44" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460362976381" checkinUser="plazi" docAuthor="Pinto, Ângelo Parise & Almeida, Marcus Vinícius Oliveira De" docDate="2016" docId="495E274CFFCEFF8AFF43FB81EAB8F9D1" docLanguage="en" docName="zootaxa.4078.1.6.pdf" docOrigin="Zootaxa 4078 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyanogomphus angelomachadoi Pinto & Almeida, 2016, sp. nov." docType="treatment" docUuid="61BDDFBA-202E-41CD-AA6D-2E65CD171CF4" docUuidSource="ZooBank" docVersion="9" masterDocId="B5675F34FFC9FF87FFD4FFFCE97EFD39" masterDocTitle="A taxonomic synopsis of South American Cyanogomphini Carle with description of Cyanogomphus angelomachadoi sp. nov. from the Cerrado of Brazil (Odonata: Gomphidae)" updateTime="1698294756334" updateUser="ExternalLinkService">
|
||
<mods:mods id="173A379CA3431B23A58AC06AF3164F11" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="A21F6320DDADF3C24A05F45ABEF1BC5F">
|
||
<mods:title id="6B1E98D290FF2B182C4BF6B05E36BBAA">A taxonomic synopsis of South American Cyanogomphini Carle with description of Cyanogomphus angelomachadoi sp. nov. from the Cerrado of Brazil (Odonata: Gomphidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="094B51EB758785A6DAD330F9D0113FA6" type="personal">
|
||
<mods:role id="77C0543ADFB40A5C8D82F582215BDA0D">
|
||
<mods:roleTerm id="35238EF1878270CE54E7B4B2D053C221">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6DB4C4B582780B393C857E3BF3719109">Pinto, Ângelo Parise</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="E427A935ED089807C00F47A044CCF7CF" type="personal">
|
||
<mods:role id="BEF6D0E4B39678B238E16DB51AF0A90E">
|
||
<mods:roleTerm id="B8FD51A1067DE62496252B18E71B2E82">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="757B3B6547BCDCFBC76749C308436904">Almeida, Marcus Vinícius Oliveira De</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="E0D02044AC47A64CBE8A2A42A15007B2">text</mods:typeOfResource>
|
||
<mods:relatedItem id="989BBB565E1013D3D3595725A8705946" type="host">
|
||
<mods:titleInfo id="4F026658A741C0A3A0373FB4948578E2">
|
||
<mods:title id="FE4F7618924AE9C4249C6109E4EBFDC5">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="8B33C60375ACBC5E1760190F3F3603BC">
|
||
<mods:date id="2543B35715B54948D80C9B0A05A09848">2016</mods:date>
|
||
<mods:detail id="4E9D88E56187A8355BAD01A6183C56BD" type="volume">
|
||
<mods:number id="F4B7E458542736C9C6DB46B8EA3243D8">4078</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="6DC3EE682D674A412402FD25338DC796" type="issue">
|
||
<mods:number id="7F3B578809F11C96CA0705A14F6FFA49">1</mods:number>
|
||
</mods:detail>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="CB71EACFDCBF383394E51B3BC5A55828">journal volume</mods:classification>
|
||
<mods:identifier id="253A3151EFC227A906E300DE95F554EC" type="CLB-Dataset">46773</mods:identifier>
|
||
<mods:identifier id="5214100F5C97B8FAC66C4D4A9AA53725" type="DOI">10.11646/zootaxa.4078.1.6</mods:identifier>
|
||
<mods:identifier id="B45C83356175B662B1B84F24BF0B1962" type="GBIF-Dataset">598cb955-c4b7-4c2b-8c74-6ee0348c7f7e</mods:identifier>
|
||
<mods:identifier id="3069996611C090C4EECA471CD62AED58" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="60BA9A27A4E41A16F373425A94B1E396" type="Zenodo-Dep">270853</mods:identifier>
|
||
<mods:identifier id="D675D2C19174186B7E64339990D5A88C" type="ZooBank">3D0D79C2-2583-47F4-BC01-5CF347614B44</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="495E274CFFCEFF8AFF43FB81EAB8F9D1" ID-DOI="http://doi.org/10.5281/zenodo.6077749" ID-GBIF-Taxon="119410156" ID-Zenodo-Dep="6077749" LSID="urn:lsid:zoobank.org:act:61BDDFBA-202E-41CD-AA6D-2E65CD171CF4" httpUri="http://treatment.plazi.org/id/495E274CFFCEFF8AFF43FB81EAB8F9D1" lastPageId="13" pageId="7">
|
||
<subSubSection id="89EDC5D1FFCEFF80FF43FB81E810F805" pageId="7" type="nomenclature">
|
||
<paragraph id="C148965AFFCEFF80FF43FB81EBF6F9AE" blockId="7.[151,1024,1149,1240]" box="[151,648,1149,1175]" pageId="7">
|
||
<heading id="9A002136FFCEFF80FF43FB81EBF6F9AE" bold="true" box="[151,648,1149,1175]" fontSize="11" level="1" pageId="7" reason="1">
|
||
<emphasis id="F3834A48FFCEFF80FF43FB81EBF6F9AE" bold="true" box="[151,648,1149,1175]" pageId="7">
|
||
<taxonomicName id="06F7EDD9FFCEFF80FF43FB81EB5DF9AE" ID-CoL="9WYQQ" box="[151,547,1149,1175]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="7" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
|
||
<emphasis id="F3834A48FFCEFF80FF43FB81EB5DF9AE" bold="true" box="[151,547,1149,1175]" italics="true" pageId="7">Cyanogomphus angelomachadoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E8B0F733FFCEFF80FDFFFB81EBF6F9AE" box="[555,648,1149,1175]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C148965AFFCEFF80FF43FB63EA93F9E1" blockId="7.[151,1024,1149,1240]" pageId="7">
|
||
LSID urn:lsid:zoobank.org:act:61BDDFBA-202E-41CD-AA6D-2E65CD171CF4 (
|
||
<figureCitation id="59CC8ADFFFCEFF80FF4AFB43E877F9E1" box="[158,265,1215,1240]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="7">Figures 1</figureCitation>
|
||
,
|
||
<figureCitation id="59CC8ADFFFCEFF80FEC3FB3CE85BF9E1" box="[279,293,1216,1240]" captionStart="FIGURE 2" captionStartId="6.[151,250,1639,1661]" captionTargetBox="[151,1436,193,1617]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURE 2. Wings of Cyanogomphini in ventral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449), B. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448); C. Cyanogomphus comparabilis Belle, 1994, Holotype ♂ (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, ♀ (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), ♀ (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), ♂ (Brazil. SC: Nova Teutônia village, MZSP)." httpUri="https://zenodo.org/record/270855/files/figure.png" pageId="7">2</figureCitation>
|
||
A–B, 3A, 4A–B, 6A–C, 7A–C, 8A–B, 9C, 10A–B, 11A–B, 12)
|
||
</paragraph>
|
||
<paragraph id="C148965AFFCEFF80FF43FAFBE810F805" blockId="7.[151,1435,1287,1340]" pageId="7">
|
||
<treatmentCitationGroup id="E1E7B174FFCEFF80FF43FAFBE810F805" pageId="7">
|
||
<taxonomicName id="06F7EDD9FFCEFF80FF43FAFBE84BF824" ID-CoL="3XNW" box="[151,309,1287,1309]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="7" phylum="Arthropoda" rank="genus" status="sp. nov.">
|
||
<emphasis id="F3834A48FFCEFF80FF43FAFBE84BF824" box="[151,309,1287,1309]" italics="true" pageId="7">Cyanogomphus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E8B0F733FFCEFF80FEE9FAF4E8F2F824" box="[317,396,1288,1309]" pageId="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
—
|
||
<treatmentCitation id="4056B04BFFCEFF80FE72FAF4EB9CF824" author="" box="[422,738,1287,1310]" page="418" pageId="7">
|
||
<bibRefCitation id="A566EBABFFCEFF80FE72FAF4EBCEF824" box="[422,688,1287,1310]" pageId="7" refString="De Almeida, M. V. O., Pinto, A. P., Carvalho, A. L. & Takiya, D. M. (2013) When rare is just a matter of sampling: Unexpected dominance of clubtail dragonflies (Odonata, Gomphidae) through different collecting methods at Parque Nacional da Serra do Cipo, Minas Gerais State, Brazil. Revista Brasileira de Entomologia, 57 (4), 417 - 423. http: // dx. doi. org / 10.1590 / S 0085 - 56262013005000042" type="journal article">
|
||
<collectingCountry id="B9E0D6CAFFCEFF80FE72FAF4E8BAF824" box="[422,452,1288,1309]" name="South Georgia and the South Sandwich Islands" pageId="7">De</collectingCountry>
|
||
Almeida
|
||
<emphasis id="F3834A48FFCEFF80FDF8FAF5EB1FF824" box="[556,609,1287,1310]" italics="true" pageId="7">et al.</emphasis>
|
||
(2013)
|
||
</bibRefCitation>
|
||
: 418
|
||
</treatmentCitation>
|
||
(mention of the material described here from “Parque Nacional da Serra do Cipó”).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89EDC5D1FFCEFF80FF43FA95EC19FA37" pageId="7" type="materials_examined">
|
||
<paragraph id="C148965AFFCEFF80FF43FA95EC19FA37" blockId="7.[151,1437,1385,2022]" pageId="7">
|
||
<emphasis id="F3834A48FFCEFF80FF43FA95EBEDF8BB" bold="true" box="[151,659,1385,1410]" pageId="7">
|
||
Material examined (
|
||
<specimenCount id="D7F15DD3FFCEFF80FE46FA95E8CBF8BB" box="[402,437,1385,1410]" pageId="7" type="generic">4♂</specimenCount>
|
||
, 2♀).
|
||
<typeStatus id="1E4C28F8FFCEFF80FDD2FA95EB0DF8BB" box="[518,627,1385,1410]" pageId="7" type="holotype">Holotype</typeStatus>
|
||
♂
|
||
</emphasis>
|
||
.
|
||
<collectingCountry id="B9E0D6CAFFCEFF80FD72FA95EA71F8B8" box="[678,783,1385,1409]" name="Brazil" pageId="7">BRAZIL</collectingCountry>
|
||
. Minas Gerais State: Jaboticatubas municipality, Parque Nacional da Serra do Cipó (
|
||
<collectionCode id="A7E60E9FFFCEFF80FE0AFA71EB57F89C" box="[478,553,1421,1445]" pageId="7">PNSC</collectionCode>
|
||
),, light trap using a white sheet with mixed mercury vapor lights (250W) at Córrego das Pedras stream (
|
||
<geoCoordinate id="A4C3F09DFFCEFF80FE0EFA4DEB1FF8F0" box="[474,609,1457,1481]" direction="south" orientation="latitude" pageId="7" precision="15" value="-19.37139">19°22’17”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A4C3F09DFFCEFF80FDBBFA4DEB80F8F3" box="[623,766,1457,1482]" direction="west" orientation="longitude" pageId="7" precision="15" value="-43.60083">43°36’03”W</geoCoordinate>
|
||
,
|
||
<quantity id="060F3BBFFFCEFF80FCD8FA4DEA2AF8F0" box="[780,852,1457,1482]" metricMagnitude="2" metricUnit="m" metricValue="7.659999999999999" pageId="7" unit="m" value="766.0">766 m</quantity>
|
||
a.s.l.),
|
||
<date id="B549B09AFFCEFF80FC7FFA4DED4BF8F0" box="[939,1077,1457,1481]" pageId="7" value="2011-12-12">12.XII.2011</date>
|
||
, A.P.M. Santos & D.M. Takiya leg. (
|
||
<collectionCode id="A7E60E9FFFCEFF80FF1BFA29E86AF8D4" box="[207,276,1493,1517]" pageId="7">DZRJ</collectionCode>
|
||
1449);
|
||
<emphasis id="F3834A48FFCEFF80FEBEFA29EB7FF8D7" bold="true" box="[362,513,1493,1518]" pageId="7">
|
||
1♀
|
||
<typeStatus id="1E4C28F8FFCEFF80FE40FA29EB7FF8D7" box="[404,513,1493,1518]" pageId="7" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
, same data as
|
||
<typeStatus id="1E4C28F8FFCEFF80FD77FA29EA7BF8D7" box="[675,773,1493,1518]" pageId="7" type="holotype">holotype</typeStatus>
|
||
but 6 meter-long Malaise trap crossing Córrego das Pedras stream,
|
||
<date id="B549B09AFFCEFF80FF21FA05E8E2FB28" box="[245,412,1529,1553]" pageId="7" value="2011-12-09" valueMax="2011-12-13" valueMin="2011-12-09">9–13.XII.2011</date>
|
||
, A.P.M. Santos, D.M. Takiya, M.L. Monné & R.R. Cavichioli leg. (
|
||
<collectionCode id="A7E60E9FFFCEFF80FB06FA05EC69FB28" box="[1234,1303,1529,1553]" pageId="7">DZRJ</collectionCode>
|
||
1448);
|
||
<emphasis id="F3834A48FFCEFF80FAADFA05E87AFB0F" bold="true" pageId="7">
|
||
<specimenCount id="D7F15DD3FFCEFF80FAADFA05ECE2FB2B" box="[1401,1436,1529,1554]" pageId="7" type="generic">1♂</specimenCount>
|
||
<typeStatus id="1E4C28F8FFCEFF80FF43F9E1E87AFB0F" box="[151,260,1565,1590]" pageId="7" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
, [Lagoa Santa municipality], Estr[ada] B[elo] Horizonte, Serra do Cipó, Km 46–47 [
|
||
<geoCoordinate id="A4C3F09DFFCEFF80FB3FF9E1ECE9FB0F" box="[1259,1431,1565,1590]" direction="south" orientation="latitude" pageId="7" precision="1" value="-19.561266">19°33’40.56”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A4C3F09DFFCEFF80FF43F9BDE837FB63" box="[151,329,1601,1626]" direction="west" orientation="longitude" pageId="7" precision="1" value="-43.913116">43°54’47.21”W</geoCoordinate>
|
||
,
|
||
<quantity id="060F3BBFFFCEFF80FE81F9BDE8E0FB60" box="[341,414,1601,1626]" metricMagnitude="2" metricUnit="m" metricValue="6.64" pageId="7" unit="m" value="664.0">664 m</quantity>
|
||
a.s.l.], Exc[ursão] 221, Col[eta] 30,
|
||
<date id="B549B09AFFCEFF80FCE9F9BDEAB9FB63" box="[829,967,1601,1626]" pageId="7" value="1963-12-01">01.XII.1963</date>
|
||
, N.[D.] Santos, [J.P.] Machado & Borges leg. (
|
||
<collectionCode id="A7E60E9FFFCEFF80FF1BF999E85EFB44" box="[207,288,1637,1661]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="7">MNRJ</collectionCode>
|
||
);
|
||
<emphasis id="F3834A48FFCEFF80FEE6F999E8B7FB47" bold="true" box="[306,457,1637,1662]" pageId="7">
|
||
<specimenCount id="D7F15DD3FFCEFF80FEE6F999E828FB47" box="[306,342,1637,1662]" pageId="7" type="generic">1♂</specimenCount>
|
||
<typeStatus id="1E4C28F8FFCEFF80FE88F999E8B7FB47" box="[348,457,1637,1662]" pageId="7" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
, Serra do Salitre municipality, river at riparian Forest,
|
||
<date id="B549B09AFFCEFF80FBFFF999EDE7FB44" box="[1067,1177,1637,1661]" pageId="7" value="2014-01-13">13.I.2014</date>
|
||
, J.E. Santos Junior leg. (
|
||
<collectionCode id="A7E60E9FFFCEFF80FF4BF975E980FB98" box="[159,254,1673,1697]" pageId="7">ABMM</collectionCode>
|
||
); São Paulo State:
|
||
<emphasis id="F3834A48FFCEFF80FE1BF975EB18FB9B" bold="true" box="[463,614,1673,1698]" pageId="7">
|
||
<specimenCount id="D7F15DD3FFCEFF80FE1BF975E88CFB9B" box="[463,498,1673,1698]" pageId="7" type="generic">1♂</specimenCount>
|
||
<typeStatus id="1E4C28F8FFCEFF80FE2DF975EB18FB9B" box="[505,614,1673,1698]" pageId="7" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
, Brotas municipality, Fazenda
|
||
<collectingCountry id="B9E0D6CAFFCEFF80FC16F975ED39FB9B" box="[962,1095,1673,1698]" name="Saint Lucia" pageId="7">Santa Lúcia</collectingCountry>
|
||
da Boa Vista (
|
||
<geoCoordinate id="A4C3F09DFFCEFF80FB39F975ECE6FB98" box="[1261,1432,1673,1697]" direction="south" orientation="latitude" pageId="7" precision="1" value="-22.2108">22°12’38.88”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A4C3F09DFFCEFF80FF43F951E837FBFF" box="[151,329,1709,1734]" direction="west" orientation="longitude" pageId="7" precision="1" value="-48.2433">48°14’35.88”W</geoCoordinate>
|
||
,
|
||
<quantity id="060F3BBFFFCEFF80FE8DF951E8DAFBFC" box="[345,420,1709,1734]" metricMagnitude="2" metricUnit="m" metricValue="4.85" pageId="7" unit="m" value="485.0">485 m</quantity>
|
||
a.s.l.),
|
||
<date id="B549B09AFFCEFF80FDD4F951EBFCFBFC" box="[512,642,1709,1733]" pageId="7" value="2001-11-02">02.XI.2001</date>
|
||
, F.A.A. Lencioni leg. (RWG, not examined).
|
||
<emphasis id="F3834A48FFCEFF80FB48F951ECE2FBFF" bold="true" box="[1180,1436,1709,1734]" pageId="7">Additional specimen.</emphasis>
|
||
<collectingCountry id="B9E0D6CAFFCEFF80FF43F92DE87EFBD0" box="[151,256,1745,1769]" name="Brazil" pageId="7">BRAZIL</collectingCountry>
|
||
. Minas Gerais State: 1♀, Serra do Salitre municipality, river at riparian forest,
|
||
<bibRefCitation id="A566EBABFFCEFF80FBA7F92DEDB9FBD0" author="II" box="[1139,1223,1745,1769]" pageId="7" refString="GEOnet Names Server (GNS), National Geospatial-Intelligence Agency (2014) U. S. National Imagery and Mapping Agency's (NIMA) database of foreign geographic feature names. Available from: http: // geonames. nga. mil / gns / html / (accessed October, 2014). [Argentina gazetteer available from: http: // biogeo. ucdavis. edu / data / diva / gaz / ARG _ gaz. zip; Uruguay gazetteer available from: http: // biogeo. ucdavis. edu / data / diva / gaz / URY _ gaz. zip]" type="book chapter" year="2014">
|
||
<date id="B549B09AFFCEFF80FBA7F92DEDB9FBD0" box="[1139,1223,1745,1769]" pageId="7" value="2014-02">II.2014</date>
|
||
</bibRefCitation>
|
||
, J.E. Santos Junior leg. (
|
||
<collectionCode id="A7E60E9FFFCEFF80FF1BF909E850FA34" box="[207,302,1781,1805]" pageId="7">ABMM</collectionCode>
|
||
). Specimens in
|
||
<collectionCode id="A7E60E9FFFCEFF80FE0AF90AEB5FFA34" box="[478,545,1782,1805]" pageId="7">DZRJ</collectionCode>
|
||
collected under ICMBIO/SISBIO licenses numbers 14591-8 and 23620-6.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89EDC5D1FFCEFF8BFF13F8E5E8EAF8DE" lastPageId="12" pageId="7" type="description">
|
||
<paragraph id="C148965AFFCEFF8FFF13F8E5ED1EFCCC" blockId="7.[151,1437,1385,2022]" lastBlockId="8.[151,1437,151,2016]" lastPageId="8" pageId="7">
|
||
<emphasis id="F3834A48FFCEFF80FF13F8E5EBF4FA0B" bold="true" box="[199,650,1817,1842]" pageId="7">
|
||
Description of male
|
||
<typeStatus id="1E4C28F8FFCEFF80FE1AF8E5EB49FA0B" box="[462,567,1817,1842]" pageId="7" type="holotype">holotype</typeStatus>
|
||
. Head
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="59CC8ADFFFCEFF80FD75F8E5EB92FA08" box="[673,748,1817,1842]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="7">Fig. 4</figureCitation>
|
||
A). Labium greenish-yellow, visible parts of maxilla and mandible greenish-yellow, except dark-brown to glossy black movable hook, extreme apex of maxillary outer lobe and mandibular incisor lobe. Genae brown. Labrum greenish-yellow, with a narrow straight dark brown to black stripe at ventral margin, one lateral margin with a narrow dark brown line from ventral margin to level of clypeolabral suture, clypeolabral suture light brown connected to a small mesal rounded spot over labrum occupying about half of its total height. Anteclypeus brownish-yellow, clypeus greenish-yellow. Frons yellowishgreen, darker than remainder of face, with dark brown bar at posterior 0.4, extended laterally. Antenna, vertex and occiput dark brown to black, except yellowish membranous part of scape and area close to postocellar suture noticeably lighter than adjacent areas of cranium. Posterior region of cranium (“postgena” plus “occiput”) gradually lightening ventrally with irregular dark areas, especially close to postoccipital suture. Ventral margin of labrum with a wide concavity almost mesal, giving to sclerite a slight asymmetric shape; ante- and postfrons division carinated, dorsal margin of frons sinuous in anterior view with a small mesal concavity, anterior margin regularly convex and with a shallow mesal furrow in dorsal view. Posterior portion of vertex with a ridge (postocellar ridge
|
||
<emphasis id="F3834A48FFC1FF8FFEBCFE69E8D8FC95" box="[360,422,405,428]" italics="true" pageId="8">sensu</emphasis>
|
||
Belle 1973) slightly projecting anteriorly over lateral ocelli, occiput (occipital plate) in dorsal view with a distinct ridge along its entire width (
|
||
<figureCitation id="59CC8ADFFFC1FF8FFCF2FE4BEA13FCE9" box="[806,877,439,464]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="8">Fig. 4</figureCitation>
|
||
A), area posterior to ridge distinctly higher than anterior, posterior margin of occiput with scattered setae, not forming a distinct fringe.
|
||
</paragraph>
|
||
<paragraph id="C148965AFFC1FF8FFF13FE03ED79F86D" blockId="8.[151,1437,151,2016]" pageId="8">
|
||
<emphasis id="F3834A48FFC1FF8FFF13FE03E858FF21" bold="true" box="[199,294,511,536]" pageId="8">Thorax.</emphasis>
|
||
Prothorax greenish-yellow, lightening ventrally to basisternum with dark brown spots or stripes on anterior margin of anterior lobe, an ill-defined spot on dorsum of median lobe and distinct stripes surrounding central pale area of posterior lobe. Synthorax (
|
||
<figureCitation id="59CC8ADFFFC1FF8FFD6AFDBBEA7AFF59" box="[702,772,583,608]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="8">Figs 1</figureCitation>
|
||
A–B) largely pale, patterned with dark brown spots and greenish-yellow stripes dorsally, almost entirely greenish-yellow laterally; dorsal carina dark on anterior third, then with narrow yellowish line; mesepisternum with first and second pale antehumeral stripes distinct, first antehumeral stripe joined to pale mesothoracic collar, to second antehumeral stripe at about half length, and at level of antealar sinus (
|
||
<figureCitation id="59CC8ADFFFC1FF8FFE89FD2BE8DEFFC9" box="[349,416,727,752]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="8">Figs 1</figureCitation>
|
||
A–B); pale areas enclosing three large dark spots, a triangular dark brown mesal spot from collar to antealar sinus, at maximum half pleurite width, a barely defined rounded brown spot from mesinfraepisternum to about of posterior 0.4, and an ellipsoid dark brown spot posteriorly; a narrow brown stripe parallel to the pale mesopleural suture (humeral) beginning at 0.2 height of epimeron and extending to near antealar sinus, enlarged posteriorly; mesepisternum with a dark brown stripe parallel to mesopleural suture darkened and wider ventrally close to mesinfraepisternum (
|
||
<figureCitation id="59CC8ADFFFC1FF8FFD7AFC70EB88FE9D" box="[686,758,908,933]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="8">Figs 1</figureCitation>
|
||
A–B); metepisternum with a barely defined brown stripe dorsally from metapleural antealar process to intersegmental suture, remainder of synthorax greenish-yellow lightening to light yellow ventrally, with irregular ill-defined dark areas laterally (
|
||
<figureCitation id="59CC8ADFFFC1FF8FFBBAFC28EDCAFED5" box="[1134,1204,980,1005]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="8">Fig. 1</figureCitation>
|
||
B). Legs, coxa and trochanter yellowish-brown, ventral and posteroventral surfaces of femur light yellow, anteroventral and dorsal surfaces dark brown to black in pro- and mesothoracic legs, in metathoracic leg yellowish-brown with irregular dark areas and distal 0.1 black; tibia black with dorsal surface yellow to brownish-yellow (
|
||
<figureCitation id="59CC8ADFFFC1FF8FFB4AFBC3ED9FF961" box="[1182,1249,1087,1112]" captionStart="FIGURE 3" captionStartId="7.[151,250,894,916]" captionTargetBox="[151,1436,193,871]" captionTargetId="figure@7.[151,1436,193,873]" captionTargetPageId="7" captionText="FIGURE 3. Details of metathoracic tibia and tarsus of males of Cyanogomphini in lateral view. A. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); C. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); D. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270856/files/figure.png" pageId="8">Fig. 3</figureCitation>
|
||
A); tarsus black, pretarsal claws dark brown with distinct supplementary inferior tooth. Proximal portion of trochanter (1
|
||
<collectingCountry id="B9E0D6CAFFC1FF8FFAE2FB98EC2BF945" box="[1334,1365,1124,1148]" name="Chile" pageId="8">Tr</collectingCountry>
|
||
<emphasis id="F3834A48FFC1FF8FFA8AFB99ECE2F945" box="[1374,1436,1125,1148]" italics="true" pageId="8">sensu</emphasis>
|
||
<bibRefCitation id="A566EBABFFC1FF8FFF43FB7BE835F999" author="Snodgrass" box="[151,331,1159,1184]" pageId="8" refString="Snodgrass, R. E. (1935) Principles of insect morphology. McGraw-Hill Book Company, New York, 667 pp." type="book" year="1935">Snodgrass 1935</bibRefCitation>
|
||
) armed with a row of small black spurs only in the mesothoracic leg, distal portion (2
|
||
<collectingCountry id="B9E0D6CAFFC1FF8FFADCFB74EC5BF999" box="[1288,1317,1160,1184]" name="Chile" pageId="8">Tr</collectingCountry>
|
||
) armed on all legs, most numerous on mesothoracic leg. Femur with rows of numerous small black spurs, metathoracic femur slight convex anteriorly in lateral view, anteroventral surface of prothoracic tibia armed with 10–12 spurs (including scale-like spurs of tibial comb), 11–12 on mesothoracic, 14–15 on metathoracic, posteroventral surface with 14–15 on prothoracic, 14–15 on mesothoracic, 13–14 on metathoracic, usually at least two times longer than intervening space. Tibial and tarsal spurs modified, basally enlarged (
|
||
<figureCitation id="59CC8ADFFFC1FF8FFC74FAC0EA9DF86D" box="[928,995,1340,1365]" captionStart="FIGURE 3" captionStartId="7.[151,250,894,916]" captionTargetBox="[151,1436,193,871]" captionTargetId="figure@7.[151,1436,193,873]" captionTargetPageId="7" captionText="FIGURE 3. Details of metathoracic tibia and tarsus of males of Cyanogomphini in lateral view. A. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); C. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); D. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270856/files/figure.png" pageId="8">Fig. 3</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<paragraph id="C148965AFFC1FF8FFF13FAA3EBF5FA3E" blockId="8.[151,1437,151,2016]" pageId="8">
|
||
Wings (
|
||
<figureCitation id="59CC8ADFFFC1FF8FFEF0FAA3E812F841" box="[292,364,1375,1400]" captionStart="FIGURE 2" captionStartId="6.[151,250,1639,1661]" captionTargetBox="[151,1436,193,1617]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURE 2. Wings of Cyanogomphini in ventral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449), B. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448); C. Cyanogomphus comparabilis Belle, 1994, Holotype ♂ (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, ♀ (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), ♀ (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), ♂ (Brazil. SC: Nova Teutônia village, MZSP)." httpUri="https://zenodo.org/record/270855/files/figure.png" pageId="8">Fig. 2</figureCitation>
|
||
A). Membrane hyaline; venation dark-brown to black; pt brown, contrasting with black of marginal veins; one basal subcostal crossvein in all wings, membranula strongly reduced. Venation as follows: 12–13 Ax in Fw, 9–10 Ax in Hw, Ax2 the 5th in all wings; 9 Px all wings; 8–9 postsubnodals (including brace vein) in Fw,
|
||
<quantity id="060F3BBFFFC1FF8FFF3CFA33E868F8DE" box="[232,278,1486,1511]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="8" unit="in" value="8.0">8 in</quantity>
|
||
Hw; 5–6 bridge crossveins in Fw,
|
||
<quantity id="060F3BBFFFC1FF8FFD79FA33EBA4F8DE" box="[685,730,1486,1511]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="8" unit="in" value="4.0">4 in</quantity>
|
||
Hw; arc at or slightly distal to second Ax; sectors of arc not stalked, origin located 0.5 basal of arc in all wings; 3–4 crossveins in the space between arc and RP-midfork in Fw,
|
||
<quantity id="060F3BBFFFC1FF8FFF43F9EBE99FFB16" box="[151,225,1558,1583]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="8" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">2–3 in</quantity>
|
||
Hw; Rspl indistinct in all wings; discoidal triangles, supratriangles and subtriangles not crossed in four wings; space between CuP-crossing and PsA with one crossvein in all wings; Fw discoidal field with two rows of cells up to level of nodus, beyond that three and increasing to 9–11 cells at wing margin; Hw discoidal field divergent, with 4–5 rows of two cells, followed by 1–2 rows of three cells, then broadening to 9–11 cells towards wing margin; Mspl indistinct in the four wings; anal loop rectangular with four cells; seven paranal cells from wing base to apex of subtriangle in Fw, five in Hw; anal triangle distinct, three-celled, with seven strong and short spines along the dorso-proximal margin (AP&AA).
|
||
</paragraph>
|
||
<paragraph id="C148965AFFC1FF8EFF13F8EEE81FF841" blockId="8.[151,1437,151,2016]" lastBlockId="9.[151,1437,151,1977]" lastPageId="9" pageId="8">
|
||
<emphasis id="F3834A48FFC1FF8FFF13F8EEE842FA12" bold="true" box="[199,316,1810,1835]" pageId="8">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="59CC8ADFFFC1FF8FFE99F8EEE8DFFA12" box="[333,417,1810,1835]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="8">Fig. 10</figureCitation>
|
||
A). Dark brown with pale greenish-yellow spots. Tergite of S1 almost entirely greenishyellow expect the dark brown rounded process (swelling of abdominal articulation
|
||
<emphasis id="F3834A48FFC1FF8FFB9AF8C4EDF2FA76" box="[1102,1164,1848,1871]" italics="true" pageId="8">sensu</emphasis>
|
||
<bibRefCitation id="A566EBABFFC1FF8FFB41F8CAEC4DFA76" author="Asahina" box="[1173,1331,1846,1871]" pageId="8" refString="Asahina, S. (1954) A morphological study of a relic dragonfly Epiophlebia superstes. The Japan Society for the promotion of science publ., Tokyo, iv + 153 pp., 71 pls." type="book" year="1954">Asahina 1954</bibRefCitation>
|
||
) close to posterior carina; S2 with a large pale spot on anterior 0.6, narrowing posteriad to 0.33 of segment height in lateral view, a small pale spot close to posterior carina and a middorsal pale line along entire segment; S3–7 with a pale proximal ring up to transverse carina, darkened in its middle, remainder of segment posteriorly dark brown to black; S8 almost entirely dark brown with an irregular lateroventral pale spot, enlarged at both ends of segment; S9 dark brown with ill-defined lateral lighter areas; S10 dark brown above, with pale lateral spot extended dorsally in a narrow ring at 0.66 length separating a rounded basal spot from the dark posterior end (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFB1CFF40EC6CFDED" box="[1224,1298,188,213]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Figs 7</figureCitation>
|
||
A–C, 10A); sternites dark brown to black; ventral carina of S2–8 and anterior 0.5 of S9 greenish-yellow; S1–7 cylindrical, S8–10 distinctly wider than previous segments, maximum width at S8. Auricles rounded, positioned obliquely to S2 longitudinal axis, length about one-third height of S2; posterior margin with two patches of 5–6 minute dark spines; posterior carina of S9 armed with small spines dorsally and with V-shaped concavity. Secondary genitalia typical of the
|
||
<taxonomicName id="06F7EDD9FFC0FF8EFEE0FE93E8A7FCB1" box="[308,473,367,392]" class="Insecta" family="Gomphidae" genus="Agriogomphus" kingdom="Animalia" order="Odonata" pageId="9" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3834A48FFC0FF8EFEE0FE93E8A7FCB1" box="[308,473,367,392]" italics="true" pageId="9">Agriogomphus</emphasis>
|
||
</taxonomicName>
|
||
-complex, anterior lamina not visible in lateral view, in ventral view posterior margin with a wide U-shaped ridge, anterior hamule (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFD60FE68EB84FC95" box="[692,762,404,429]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="9">Fig. 4</figureCitation>
|
||
B) yellowish-brown, quadrangular in lateral view, anterior margin strongly ridged, sickle-shaped, posteriorly prolonged into a concave laminar projection with scattered hairlike setae denser posteriorly, a small ventral V-shaped concavity between anterior ridge and posterior laminar projection; in ventral view posterior margin convergent, the limit between anterior ridge and posterior laminar projection concave in a wide V-shape; posterior hamule (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFCCAFDD8EA1DFF05" box="[798,867,548,573]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="9">Fig. 4</figureCitation>
|
||
B) light brown, except black stout acute anteriorly directed apex, distinctly longer than other structures of secondary genitalia in lateral view, anterior margin slightly sinuous, slightly convex at proximal 0.4 then gently concave, with small deep concavity before extreme distal; two distinct sets of pale hair-like long setae, a basal line with 8–10 setae at anterior margin and a distal tuft with about a dozen setae close to acute apex (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFDF5FD48EB14FFF5" box="[545,618,692,717]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="9">Fig. 4</figureCitation>
|
||
B); in ventral view regularly curved toward convergent black apexes, somewhat flattened; without a distinct genital lobe. Vesica spermalis (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFC18FD2BED6AFFC9" box="[972,1044,727,752]" captionStart="FIGURE 4" captionStartId="10.[151,250,1598,1620]" captionTargetBox="[154,1436,151,1575]" captionTargetId="figure@10.[154,1439,151,1576]" captionTargetPageId="10" captionText="FIGURE 4. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B – F). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="9">Fig. 4</figureCitation>
|
||
B) with
|
||
<date id="B549B09AFFC0FF8EFBA3FD24EDE7FFC9" box="[1143,1177,728,752]" pageId="9">V1</date>
|
||
black, in lateral view slightly higher than level of anterior hamule, scoop-like process on
|
||
<date id="B549B09AFFC0FF8EFC5AFD00EACEFE2D" box="[910,944,764,788]" pageId="9">V1</date>
|
||
(median cleft
|
||
<emphasis id="F3834A48FFC0FF8EFB87FD01EDEFFE2D" box="[1107,1169,765,788]" italics="true" pageId="9">sensu</emphasis>
|
||
<bibRefCitation id="A566EBABFFC0FF8EFB4CFD00EC6AFE2D" author="Belle" box="[1176,1300,764,789]" pageId="9" refString="Belle, J. (1988) A synopsis of the species of Phyllocycla Calvert, with descriptions of four new taxa and a key to the genera of Neotropical Gomphidae (Odonata, Gomphidae). Tijdschrift voor Entomologie, 131, 73 - 102." type="journal article" year="1988">Belle 1988</bibRefCitation>
|
||
) rounded in front view, strongly concave and covered with hair-like yellow setae internally; see
|
||
<typeStatus id="1E4C28F8FFC0FF8EFB82FCDCEDC8FE01" box="[1110,1206,800,824]" pageId="9" type="paratype">paratype</typeStatus>
|
||
for full description. Cercus (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFF21FCB8E842FE65" box="[245,316,836,861]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Figs 7</figureCitation>
|
||
A–C, 8A–B) greenish-yellow spotted with brown by numerous tubercular setal alveoli proximally, darkening to black apex, ventral branch dark brown (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFD3EFC9BEA50FEB9" box="[746,814,871,896]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Fig. 7</figureCitation>
|
||
A); in lateral view cercus strongly curved at half-length at level of epiproct posteriorly, directed obliquely dorsad (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFCB4FC70EAD0FE9D" box="[864,942,908,933]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Figs 7</figureCitation>
|
||
A, 8A); apex acute and slightly directed posteriorly; posterior margin flattened, forming a distinct rounded keel (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFC6FFC53EA83FEF1" box="[955,1021,943,968]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Fig. 7</figureCitation>
|
||
A); ventral branch strongly curved in lateral view with apex directed anteriorly (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFDAEFC28EBBCFED5" box="[634,706,980,1005]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Figs 7</figureCitation>
|
||
A, 8A); cercus in dorsal view cylindrical, tapering posteriorly to acute apex, slightly divergent (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFE2EFC0BEB3EF929" box="[506,576,1015,1040]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Fig. 7</figureCitation>
|
||
B); mesal margin with a rounded basal projection, then gradually tapering, external margin almost straight (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFDC0FBE0EB22F90D" box="[532,604,1052,1077]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Fig. 7</figureCitation>
|
||
B); cerci in ventral view touching each other proximally and gradually diverging distally, apexes almost parallel, ventral branch strongly directing parallel toward anteriorly, apex rounded, slightly flattened (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFE1AFB98EB6CF945" box="[462,530,1124,1149]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Fig. 7</figureCitation>
|
||
C). Epiproct dark brown, largely bifid, posteriorly at level of cercus (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFACDFB98EC1EF945" box="[1305,1376,1124,1149]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Figs 7</figureCitation>
|
||
A, C, 8A–B); almost straight in lateral view, with a dorsal and ventral ridge at proximal 0.25, a small tubercle-like rounded process on dorsal margin at ca. distal 0.70 (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFD39FB50EA4BF9FD" box="[749,821,1196,1221]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Figs 7</figureCitation>
|
||
A, 8A–B); laterodistal projection forefinger-like, apex extending obliquely upward, posterior margin slightly concave (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFC72FB33EA88F9D1" box="[934,1014,1231,1256]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Figs 7</figureCitation>
|
||
A, 8A–B); in dorsal view slightly convergent, dilated at about posterior 0.70 arms surrounding a deep quadrangular concavity (hereafter distal concavity,
|
||
<figureCitation id="59CC8ADFFFC0FF8EFEC5FAEBE82DF809" box="[273,339,1303,1328]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="9">Fig. 8</figureCitation>
|
||
B); in dorsolateral view with a strong tubercle-like process anterior to the concavity (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFACCFAEBEC25F809" box="[1304,1371,1303,1328]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="9">Fig. 8</figureCitation>
|
||
B); in ventral view almost straight, laterodistal projection distinctly posterior to internal margin, posterior margin slightly sinuous (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFF28FAA3E841F841" box="[252,319,1375,1400]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="9">Fig. 7</figureCitation>
|
||
C).
|
||
</paragraph>
|
||
<paragraph id="C148965AFFC0FF8EFF13FA78EBB4FB31" blockId="9.[151,1437,151,1977]" pageId="9">
|
||
<emphasis id="F3834A48FFC0FF8EFF13FA78E803F8A4" bold="true" box="[199,381,1412,1437]" pageId="9">Measurements.</emphasis>
|
||
Total length (including caudal appendages) 40.5; abdomen length (excluding caudal appendages) 29.2; head maximum width 5.7; Fw length 25; Hw length 23.5; Fw maximum width 5.6, in Hw 7.2; pt length
|
||
<quantity id="060F3BBFFFC0FF8EFF32FA30E856F8DC" box="[230,296,1484,1509]" metricMagnitude="-2" metricUnit="m" metricValue="7.111999999999999" pageId="9" unit="in" value="2.8">2.8 in</quantity>
|
||
Fw,
|
||
<quantity id="060F3BBFFFC0FF8EFEB6FA30E8DAF8DC" box="[354,420,1484,1509]" metricMagnitude="-2" metricUnit="m" metricValue="7.366" pageId="9" unit="in" value="2.9">2.9 in</quantity>
|
||
Hw; length of metathoracic femur 6.8; metathoracic tibia 5.4; length of S9+
|
||
<quantity id="060F3BBFFFC0FF8EFAC7FA30EC30F8DC" box="[1299,1358,1484,1509]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="9" unit="in" value="10.0">10 in</quantity>
|
||
lateral view 3.3; total length of cercus in lateral view 1.4.
|
||
</paragraph>
|
||
<paragraph id="C148965AFFC0FF8EFF13F9E8E82CFBD9" blockId="9.[151,1437,151,1977]" pageId="9">
|
||
<emphasis id="F3834A48FFC0FF8EFF13F9E8EB31FB14" bold="true" box="[199,591,1556,1581]" pageId="9">
|
||
Description of female
|
||
<typeStatus id="1E4C28F8FFC0FF8EFE09F9E8EB34FB14" box="[477,586,1556,1581]" pageId="9" type="paratype">paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Similar to
|
||
<typeStatus id="1E4C28F8FFC0FF8EFD08F9E8EA3FFB14" box="[732,833,1556,1581]" pageId="9" type="holotype">holotype</typeStatus>
|
||
, only the differences described here.
|
||
<emphasis id="F3834A48FFC0FF8EFAD5F9E8EC36FB14" bold="true" box="[1281,1352,1556,1581]" pageId="9">Head.</emphasis>
|
||
Genae greenish-yellow. Dark brown to black stripe at ventral margin of labrum prolonged laterally to dorsal 0.5. Frons uniformly light brown, darker than remainder of face. Vertex and occiput light brown, the latter slightly paler. Posterior region of cranium (“postgena” plus “occiput”) light brown with irregular pale areas. Ante- and postfrons division slightly carinated. Occiput (occipital plate) behind carina clearly swollen, posterior margin sinuous with a mesal concavity.
|
||
</paragraph>
|
||
<paragraph id="C148965AFFC0FF8EFF13F910E81DFA81" blockId="9.[151,1437,151,1977]" pageId="9">
|
||
<emphasis id="F3834A48FFC0FF8EFF13F910E858FA3C" bold="true" box="[199,294,1772,1797]" pageId="9">Thorax.</emphasis>
|
||
Prothorax with posterior lobe entirely greenish-yellow. Synthorax with dark areas slightly paler than in
|
||
<typeStatus id="1E4C28F8FFC0FF8EFF60F8F3E868FA11" box="[180,278,1807,1832]" pageId="9" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="59CC8ADFFFC0FF8EFEF1F8F3E819FA11" box="[293,359,1807,1832]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="9">Figs 1</figureCitation>
|
||
E–F, 11A); pale area over dorsal carina larger. Legs with dark areas on anteroventral and dorsal surfaces more extensive; tibia dark brown to black with dorsal surface yellow (
|
||
<figureCitation id="59CC8ADFFFC0FF8EFBDAF8C8ED20FA75" box="[1038,1118,1844,1869]" captionStart="FIGURE 11" captionStartId="23.[151,250,1163,1185]" captionTargetBox="[151,1436,193,1140]" captionTargetId="figure@23.[151,1436,193,1141]" captionTargetPageId="23" captionText="FIGURE 11. Female habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Paratype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448), B. Additional specimen (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); D. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270864/files/figure.png" pageId="9">Fig. 11</figureCitation>
|
||
A). Anteroventral surface of prothoracic tibia armed with 11 spurs, 12–13 on mesothoracic, 16 on metathoracic, posteroventral surface of prothoracic tibia with 14, 16 on mesothoracic, 13–15 on metathoracic, usually more than two times longer than intervening space.
|
||
</paragraph>
|
||
<caption id="9588C6D2FFC3FF8DFF43F9C2EC55FBE3" httpUri="https://zenodo.org/record/270857/files/figure.png" pageId="10" targetBox="[154,1436,151,1575]" targetPageId="10">
|
||
<paragraph id="C148965AFFC3FF8DFF43F9C2EC55FBE3" blockId="10.[151,1436,1598,1754]" pageId="10">
|
||
<emphasis id="F3834A48FFC3FF8DFF43F9C2E86DFB6A" bold="true" box="[151,275,1598,1620]" pageId="10">FIGURE 4</emphasis>
|
||
. Males of Cyanogomphini, head in dorsal view (A) and secondary genitalia in lateral view (B–F). A–B.
|
||
<taxonomicName id="06F7EDD9FFC3FF8DFF43F99CE894FB4F" box="[151,490,1632,1654]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
|
||
<emphasis id="F3834A48FFC3FF8DFF43F99CE894FB4F" box="[151,490,1632,1654]" italics="true" pageId="10">Cyanogomphus angelomachadoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F3834A48FFC3FF8DFE22F99CEB32FB4C" bold="true" box="[502,588,1632,1653]" pageId="10">
|
||
<taxonomicNameLabel id="E8B0F733FFC3FF8DFE22F99CEB32FB4C" box="[502,588,1632,1653]" pageId="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C.
|
||
<taxonomicName id="06F7EDD9FFC3FF8DFF43F97DEB39FBAE" authority="Belle, 1994" authorityName="Belle" authorityYear="1994" box="[151,583,1665,1687]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="comparabilis">
|
||
<emphasis id="F3834A48FFC3FF8DFF43F97DE8BAFBAE" box="[151,452,1665,1687]" italics="true" pageId="10">Cyanogomphus comparabilis</emphasis>
|
||
Belle, 1994
|
||
</taxonomicName>
|
||
, Holotype (Brazil. MT: Diamantino, ABMM); D.
|
||
<taxonomicName id="06F7EDD9FFC3FF8DFB8EF97DE9B4FB80" authority="Selys, 1873" authorityName="Selys" authorityYear="1873" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="waltheri">
|
||
<emphasis id="F3834A48FFC3FF8DFB8EF97DEC2BFBAE" box="[1114,1365,1665,1687]" italics="true" pageId="10">Cyanogomphus waltheri</emphasis>
|
||
Selys, 1873
|
||
</taxonomicName>
|
||
(Brazil. RJ: Santana de Japuíba district, MNRJ); E.
|
||
<taxonomicName id="06F7EDD9FFC3FF8DFCC4F95FEC6FFB80" authority="Rodrigues Capitulo, 1985" authorityName="Rodrigues Capitulo" authorityYear="1985" box="[784,1297,1699,1721]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="noval">
|
||
<emphasis id="F3834A48FFC3FF8DFCC4F95FEA9DFB80" box="[784,995,1699,1721]" italics="true" pageId="10">Tibiagomphus noval</emphasis>
|
||
(Rodrigues Capítulo, 1985)
|
||
</taxonomicName>
|
||
(Brazil. RS: Jaguarão, MNRJ); F.
|
||
<taxonomicName id="06F7EDD9FFC3FF8DFEBBF938EB96FBE3" authority="Fraser, 1947" authorityName="Fraser" authorityYear="1947" box="[367,744,1732,1754]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="10" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFC3FF8DFEBBF938EB2AFBE3" box="[367,596,1732,1754]" italics="true" pageId="10">Tibiagomphus uncatus</emphasis>
|
||
(Fraser, 1947)
|
||
</taxonomicName>
|
||
(Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C148965AFFC3FF8DFF13F8F8EB62FACC" blockId="10.[151,1436,1796,2037]" pageId="10">
|
||
Wings (
|
||
<figureCitation id="59CC8ADFFFC3FF8DFECAF8F8E81DFA24" box="[286,355,1796,1821]" captionStart="FIGURE 2" captionStartId="6.[151,250,1639,1661]" captionTargetBox="[151,1436,193,1617]" captionTargetId="figure@6.[151,1436,193,1618]" captionTargetPageId="6" captionText="FIGURE 2. Wings of Cyanogomphini in ventral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449), B. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448); C. Cyanogomphus comparabilis Belle, 1994, Holotype ♂ (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, ♀ (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), ♀ (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), ♂ (Brazil. SC: Nova Teutônia village, MZSP)." httpUri="https://zenodo.org/record/270855/files/figure.png" pageId="10">Fig. 2</figureCitation>
|
||
B). Venation as follows: 14 Ax in Fw,
|
||
<quantity id="060F3BBFFFC3FF8DFCC7F8F9EA32FA24" box="[787,844,1796,1821]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="10" unit="in" value="10.0">10 in</quantity>
|
||
Hw, 9 Px in Fw,
|
||
<quantity id="060F3BBFFFC3FF8DFBC4F8F9ED0CFA24" box="[1040,1138,1796,1821]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.794" metricValueMin="2.54" pageId="10" unit="in" value="10.5" valueMax="11.0" valueMin="10.0">10–11 in</quantity>
|
||
Hw; 10–11 postsubnodals (including brace vein) in Fw,
|
||
<quantity id="060F3BBFFFC3FF8DFE30F8D4EB44FA78" box="[484,570,1832,1857]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.54" metricValueMin="2.286" pageId="10" unit="in" value="9.5" valueMax="10.0" valueMin="9.0">9–10 in</quantity>
|
||
Hw; 7 bridge crossveins in Fw,
|
||
<quantity id="060F3BBFFFC3FF8DFC73F8D4EA90FA78" box="[935,1006,1832,1857]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.524" metricValueMin="1.016" pageId="10" unit="in" value="5.0" valueMax="6.0" valueMin="4.0">4–6 in</quantity>
|
||
Hw; arc at or slightly distal to second Ax; 4 crossveins in space between arc and RP-midfork in Fw,
|
||
<quantity id="060F3BBFFFC3FF8DFC9AF8B1EA07FA5C" box="[846,889,1868,1893]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="10" unit="in" value="2.0">2 in</quantity>
|
||
Hw; Fw discoidal field increasing to 10–11 cells at wing margin; Hw discoidal field with 4–6 rows of two cells, followed by 1–2 rows of three cells, then broadening to 11 cells toward wing margin; anal loop barely defined with 6 cells; 6–7 paranal cells from wing base to apex of subtriangle in Fw; anal wing base rounded without distinct anal triangle and strong spines along of the dorso-proximal margin (AP&AA).
|
||
</paragraph>
|
||
<caption id="9588C6D2FFC2FF8CFF43FA03EA75FB48" httpUri="https://zenodo.org/record/270858/files/figure.png" pageId="11" targetBox="[151,1436,195,1507]" targetPageId="11">
|
||
<paragraph id="C148965AFFC2FF8CFF43FA03EA75FB48" blockId="11.[151,1436,1535,1650]" pageId="11">
|
||
<emphasis id="F3834A48FFC2FF8CFF43FA03E870FB2D" bold="true" box="[151,270,1535,1557]" pageId="11">FIGURE 5</emphasis>
|
||
. Males of Cyanogomphini, details of variations of postfrons in dorsal view (A–D) and posterior hamule in lateral view (E–L). A–B, F–H.
|
||
<taxonomicName id="06F7EDD9FFC2FF8CFE64F9E1EAC2FB0A" authority="Rodrigues Capitulo, 1985" authorityName="Rodrigues Capitulo" authorityYear="1985" box="[432,956,1565,1587]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="11" phylum="Arthropoda" rank="species" species="noval">
|
||
<emphasis id="F3834A48FFC2FF8CFE64F9E1EBF8FB0A" box="[432,646,1565,1587]" italics="true" pageId="11">Tibiagomphus noval</emphasis>
|
||
(Rodrigues Capítulo, 1985)
|
||
</taxonomicName>
|
||
(Brazil. RS: Jaguarão, MNRJ); C–D, I–L.
|
||
<taxonomicName id="06F7EDD9FFC2FF8CFF43F9C0EB5EFB6B" authority="Fraser, 1947" authorityName="Fraser" authorityYear="1947" box="[151,544,1596,1618]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="11" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="F3834A48FFC2FF8CFF43F9C0E8FFFB6B" box="[151,385,1596,1618]" italics="true" pageId="11">Tibiagomphus uncatus</emphasis>
|
||
(Fraser, 1947)
|
||
</taxonomicName>
|
||
(Brazil. SC: Nova Teutônia, MZSP). E.
|
||
<taxonomicName id="06F7EDD9FFC2FF8CFC35F9C0ECE6FB6B" authority="Belle, 1994" authorityName="Belle" authorityYear="1994" box="[993,1432,1596,1618]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="11" phylum="Arthropoda" rank="species" species="comparabilis">
|
||
<emphasis id="F3834A48FFC2FF8CFC35F9C0EC71FB6B" box="[993,1295,1596,1618]" italics="true" pageId="11">Cyanogomphus comparabilis</emphasis>
|
||
Belle, 1994
|
||
</taxonomicName>
|
||
, Paratype (Brazil. SP: Igarapava, ABMM). Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C148965AFFC2FF8CFF13F964E8C1FA5C" blockId="11.[151,1436,1688,2037]" pageId="11">
|
||
<emphasis id="F3834A48FFC2FF8CFF13F964E842FB88" bold="true" box="[199,316,1688,1713]" pageId="11">Abdomen</emphasis>
|
||
(only S1–4). Largely pale, greenish-yellow with irregular dark brown transverse stripes and irregular spots, transverse carina ill defined (
|
||
<figureCitation id="59CC8ADFFFC2FF8CFD5BF940EBA1FBEC" box="[655,735,1724,1749]" captionStart="FIGURE 11" captionStartId="23.[151,250,1163,1185]" captionTargetBox="[151,1436,193,1140]" captionTargetId="figure@23.[151,1436,193,1141]" captionTargetPageId="23" captionText="FIGURE 11. Female habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Paratype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448), B. Additional specimen (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); D. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270864/files/figure.png" pageId="11">Fig. 11</figureCitation>
|
||
A). S2 yellowish lateroventrally in anterior 0.5, light brown at level of transverse carina (not carinated) and dark brown at posterior field, S3–4 with a dark brown stripe on posterior 0.8 adjacent to ventral carina, enlarging and darkening posteriorly, a transverse dark brown stripe at level of transverse carina and at posterior field, and a dorsolateral oval dark brown spot not connected dorsally, similar to PD spot of
|
||
<bibRefCitation id="A566EBABFFC2FF8CFEC1F8B0E8C4FA5C" author="Walker" box="[277,442,1868,1893]" pageId="11" refString="Walker, E. M. (1912) The North American dragonflies of the genus Aeshna. University of Toronto Studies, Biological Series, 11, 1 - 213, pls. 1 - 28." type="journal article" year="1912">Walker (1912)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="C148965AFFC2FF8CFF13F88CEC77FA94" blockId="11.[151,1436,1688,2037]" pageId="11">
|
||
<emphasis id="F3834A48FFC2FF8CFF13F88CE803FAB0" bold="true" box="[199,381,1904,1929]" pageId="11">Measurements.</emphasis>
|
||
Head maximum width 5.7; Fw length 25.8–26.0; Hw length 24.4–24.5; Fw maximum width 5.7, in Hw 7.2; pt length
|
||
<quantity id="060F3BBFFFC2FF8CFE64F869E88EFA94" box="[432,496,1940,1965]" metricMagnitude="-2" metricUnit="m" metricValue="7.111999999999999" pageId="11" unit="in" value="2.8">2.8 in</quantity>
|
||
Fw, 3.0 in Hw; length of metathoracic femur 6.3; metathoracic tibia 5.
|
||
</paragraph>
|
||
<paragraph id="C148965AFFC2FF8BFF13F844EC43F836" blockId="11.[151,1436,1688,2037]" lastBlockId="12.[151,1437,151,2016]" lastPageId="12" pageId="11">
|
||
<emphasis id="F3834A48FFC2FF8CFF13F844EB2CFAE8" bold="true" box="[199,594,1976,2001]" pageId="11">
|
||
Variation in the male
|
||
<typeStatus id="1E4C28F8FFC2FF8CFE01F844EB30FAE8" box="[469,590,1976,2001]" pageId="11" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
The general appearance in size and color of the male from Lagoa Santa (MNRJ) is very similar to that of the
|
||
<typeStatus id="1E4C28F8FFC2FF8CFDEBF820EBDAFACC" box="[575,676,2012,2037]" pageId="11" type="holotype">holotype</typeStatus>
|
||
, though slightly paler, while that from Serra do Salitre (ABMM) is smaller and distinctly darker with first and second humeral stripes clearly defined and limited by dark ground color (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFF4BFF40E99FFDED" box="[159,225,188,213]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="12">Figs 1</figureCitation>
|
||
C–D, 10B). They agree with the
|
||
<typeStatus id="1E4C28F8FFC5FF8BFD80FF40EBC8FDEC" box="[596,694,188,213]" pageId="12" type="holotype">holotype</typeStatus>
|
||
except as follows (characters observed in one or both specimens):
|
||
<emphasis id="F3834A48FFC5FF8BFF43FF23E9AAFDC1" box="[151,212,223,248]" italics="true" pageId="12">Head</emphasis>
|
||
. Labrum with the black distal stripe almost restricted to ventral margin, mesal dark spot longer, occupying 0.66 of its total length (height). Anteclypeus greenish-yellow. Frontoclypeal suture with a distinct brownish stripe widening dorsolaterally over frons.
|
||
<emphasis id="F3834A48FFC5FF8BFDFCFEDBEB07FC79" box="[552,633,295,320]" italics="true" pageId="12">Thorax</emphasis>
|
||
. Prothorax with dark brown areas better defined. Synthorax with larger and clearly better defined dark areas; mesepisternum at level of junction between first and second pale antehumeral stripes only slightly paler than anterior and posterior dark areas, pale areas not enclosing dark spots, showing a regular gomphid pattern with defined mesepisternal stripes (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFC97FE68EAF9FC95" box="[835,903,404,429]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="12">Figs 1</figureCitation>
|
||
C–D); metepisternal dark stripe better defined. Legs, coxa and trochanter greenish-yellow; tibia dark brown to black with dorsal surface yellow. Anteroventral surface of prothoracic tibia armed with 10–11 spurs, 11–12 on mesothoracic, 14–18 on metathoracic, posteroventral surface 13–14 on prothoracic, 13 on mesothoracic, 14–16 on metathoracic. Wings (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFAF0FE03EC05FF21" box="[1316,1403,511,536]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="12">Fig. 10</figureCitation>
|
||
B). Membrane hyaline, very slightly smoked with light brown. Venation as follows: one Hw lacking basal subcostal crossvein; 11–13 Ax in Fw, 10 Ax in Hw, Ax2 the 4th in one Fw and one Hw; 7–10 Px in Fw,
|
||
<quantity id="060F3BBFFFC5FF8BFB32FDB7EC51FF5A" box="[1254,1327,586,611]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.778" pageId="12" unit="in" value="7.5" valueMax="8.0" valueMin="7.0">7–8 in</quantity>
|
||
Hw; 8–9 postsubnodals (including brace vein) in Fw,
|
||
<quantity id="060F3BBFFFC5FF8BFD4AFD93EB89FFBE" box="[670,759,622,647]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.54" metricValueMin="2.286" pageId="12" unit="in" value="9.5" valueMax="10.0" valueMin="9.0">9–10 in</quantity>
|
||
Hw; 5–7 bridge crossveins in Fw,
|
||
<quantity id="060F3BBFFFC5FF8BFB43FD92EDB8FFBE" box="[1175,1222,622,647]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="12" unit="in" value="5.0">5 in</quantity>
|
||
Hw; arc at or 0.3 proximal to second Ax; 3–5 crossveins in the space between arc and RP-midfork in Fw,
|
||
<quantity id="060F3BBFFFC5FF8BFB57FD6FEDB5FF92" box="[1155,1227,658,683]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="12" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">2–3 in</quantity>
|
||
Hw; Fw discoidal field 9–12 cells at wing margin; Hw discoidal field with 2–5 rows of two cells, followed by 1–2 rows of three cells, then broadening to 9–13 cells towards wing margin; anal loop barely defined with five cells; 6 paranal cells from wing base to apex of subtriangle in Fw,
|
||
<quantity id="060F3BBFFFC5FF8BFDACFD03EBD9FE2E" box="[632,679,766,791]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="12" unit="in" value="4.0">4 in</quantity>
|
||
Hw; 6–9 spines along of the dorso-proximal margin (AP&AA).
|
||
<emphasis id="F3834A48FFC5FF8BFF43FCDEE87FFE02" box="[151,257,802,827]" italics="true" pageId="12">Abdomen</emphasis>
|
||
. S2 with the large pale spot with irregular margin; S7 with a pale spot entirely greenish-yellow; S8 with a round pale spot at anterior 0.33; S10 distinctly paler than former segments with ill-defined paler areas. Posterior margin of auricles with irregular rows containing 8–14 minute spines, 5 setae at the line of basal hair-like setae on anterior margin of posterior hamule. Vesica spermalis (based on
|
||
<typeStatus id="1E4C28F8FFC5FF8BFC7AFC6CED71FE91" box="[942,1039,912,936]" pageId="12" type="paratype">paratype</typeStatus>
|
||
from Lagoa Santa,
|
||
<figureCitation id="59CC8ADFFFC5FF8BFADEFC72EC29FE9E" box="[1290,1367,910,935]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
|
||
A–C). Sclerotized parts dark brown to black; ventral surface of
|
||
<date id="B549B09AFFC5FF8BFCC9FC4FEA41FEF2" box="[797,831,947,971]" pageId="12">V1</date>
|
||
with two processes, a small rounded one at proximal third and a large strong scoop-like process at distal third (median cleft
|
||
<emphasis id="F3834A48FFC5FF8BFC01FC24ED6DFED6" box="[981,1043,984,1007]" italics="true" pageId="12">sensu</emphasis>
|
||
<bibRefCitation id="A566EBABFFC5FF8BFBCAFC2AEDE0FED6" author="Belle" box="[1054,1182,982,1007]" pageId="12" refString="Belle, J. (1988) A synopsis of the species of Phyllocycla Calvert, with descriptions of four new taxa and a key to the genera of Neotropical Gomphidae (Odonata, Gomphidae). Tijdschrift voor Entomologie, 131, 73 - 102." type="journal article" year="1988">Belle 1988</bibRefCitation>
|
||
), the latter laminar in lateral view, in anteroposterior view with converging rounded apexes and covered with hair-like yellow setae internally (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFEC5FBE2E827F90E" box="[273,345,1054,1079]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
|
||
A, C);
|
||
<date id="B549B09AFFC5FF8BFE72FBE3E8B7F90E" box="[422,457,1055,1079]" pageId="12">V2</date>
|
||
tubular, lacking a defined H;
|
||
<date id="B549B09AFFC5FF8BFCCDFBE3EA45F90E" box="[793,827,1055,1079]" pageId="12">V3</date>
|
||
globose in ventral view, dorsal surface (process ‘f’ of
|
||
<bibRefCitation id="A566EBABFFC5FF8BFF43FBBEE875F962" author="Pfau" box="[151,267,1090,1115]" pageId="12" refString="Pfau, H. K. (2011). Functional morphology and evolution of the male secondary copulatory apparatus of the Anisoptera (Insecta: Odonata). Zoologica, 156 (1), 1 - 103." type="journal article" year="2011">Pfau 2011</bibRefCitation>
|
||
) with a rounded, slightly sclerotized distal process (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFC8EFBBEEADCF962" box="[858,930,1090,1115]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
|
||
B, h-f) prolonged over joint between
|
||
<date id="B549B09AFFC5FF8BFA9DFBBFE9C7F946" pageId="12">V3 and V4</date>
|
||
(J
|
||
<date id="B549B09AFFC5FF8BFF08FB9BE85EF946" box="[220,288,1127,1151]" pageId="12">v3-v4</date>
|
||
), ventrally with a distinct proximal sclerite (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFCC6FB9AEA27F946" box="[786,857,1126,1151]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
|
||
A, p-ps), and a membranous-like hood distally over
|
||
<date id="B549B09AFFC5FF8BFF43FB77E9C2F99A" box="[151,188,1163,1187]" pageId="12">V4</date>
|
||
, tapering into a tongue-like apex ventrally (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFD10FB76EA71F99A" box="[708,783,1162,1187]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
|
||
A, “mh”);
|
||
<date id="B549B09AFFC5FF8BFC5BFB77EACFF99A" box="[911,945,1163,1187]" pageId="12">V4</date>
|
||
cornet-like, with a pair of lateroproximal sclerotized projections (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFE72FB52E891F9FE" box="[422,495,1198,1223]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
|
||
A-B, “lsp”), curved anteriorly in lateral view (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFBD5FB52ED38F9FE" box="[1025,1094,1198,1223]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Fig. 6</figureCitation>
|
||
A), pair of long distal flagella somewhat forcipate in ventral view (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFDFBFB2EEB0CF9D2" box="[559,626,1234,1259]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Fig. 6</figureCitation>
|
||
B, d-fl), in lateral view as long as
|
||
<date id="B549B09AFFC5FF8BFC3FFB2FED70F9D2" box="[1003,1038,1235,1259]" pageId="12">V4</date>
|
||
length (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFBBDFB2EEDD2F9D2" box="[1129,1196,1234,1259]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Fig. 6</figureCitation>
|
||
A). Cercus brownishyellow to dark brown to black, ventral branch light brown to dark-brown. Epiproct brown to almost black.
|
||
</paragraph>
|
||
<paragraph id="C148965AFFC5FF8BFF13FAE6E98CF8FA" blockId="12.[151,1437,151,2016]" pageId="12">
|
||
<emphasis id="F3834A48FFC5FF8BFF13FAE6E803F80A" bold="true" box="[199,381,1306,1331]" pageId="12">Measurements.</emphasis>
|
||
Abdomen length (excluding caudal appendages) 27.5–29.2; head maximum width 6.0 (specimen from Serra do Salitre not measured, eyes smashed); Fw length 23.4–25.0; Hw length 22.2–23.8; Fw maximum width 5.1–5.2, in Hw 6.4–6.7; pt length
|
||
<quantity id="060F3BBFFFC5FF8BFD33FA9EEA25F842" box="[743,859,1378,1403]" metricMagnitude="-2" metricUnit="m" metricValue="6.731" metricValueMax="6.858" metricValueMin="6.604" pageId="12" unit="in" value="2.65" valueMax="2.7" valueMin="2.6">2.6–2.7 in</quantity>
|
||
Fw,
|
||
<quantity id="060F3BBFFFC5FF8BFC43FA9EED75F842" box="[919,1035,1378,1403]" metricMagnitude="-2" metricUnit="m" metricValue="6.858" metricValueMax="7.111999999999999" metricValueMin="6.604" pageId="12" unit="in" value="2.7" valueMax="2.8" valueMin="2.6">2.6–2.8 in</quantity>
|
||
Hw; length of metathoracic femur 6.0–6.3; metathoracic tibia 4.6–4.8; length of S9+
|
||
<quantity id="060F3BBFFFC5FF8BFD0AFA7BEA62F8A6" box="[734,796,1414,1439]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="12" unit="in" value="10.0">10 in</quantity>
|
||
lateral view 2.8; total length of cercus in lateral view 1.2–1.3.
|
||
</paragraph>
|
||
<paragraph id="C148965AFFC5FF8BFF13FA32E8EAF8DE" blockId="12.[151,1437,151,2016]" box="[199,404,1486,1511]" pageId="12">
|
||
<emphasis id="F3834A48FFC5FF8BFF13FA32E869F8DE" bold="true" box="[199,279,1486,1511]" pageId="12">Larva.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89EDC5D1FFC5FF8AFF13FA0EED66FCE9" lastPageId="13" pageId="12" type="diagnosis">
|
||
<paragraph id="C148965AFFC5FF8BFF13FA0EEB3FFB86" blockId="12.[151,1437,151,2016]" pageId="12">
|
||
<emphasis id="F3834A48FFC5FF8BFF13FA0EE83FFB32" bold="true" box="[199,321,1522,1547]" pageId="12">Diagnosis.</emphasis>
|
||
The wide cornet-like distal portion of
|
||
<date id="B549B09AFFC5FF8BFD27FA0FEA6BFB32" box="[755,789,1523,1547]" pageId="12">V4</date>
|
||
(
|
||
<figureCitation id="59CC8ADFFFC5FF8BFCF0FA0EEA15FB32" box="[804,875,1522,1547]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
|
||
A–B), upturned tapered apex of cylindrical cercus (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFF4BF9EAE998FB16" box="[159,230,1558,1583]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="12">Figs 7</figureCitation>
|
||
A–C, 8A–B), distal concavity from posterior 0.7 of epiproct of males (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFBDBF9EAED2DFB16" box="[1039,1107,1558,1583]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="12">Fig. 8</figureCitation>
|
||
B) and short subgenital plate (ratio between length of plate and S9 <0.4) with a wide concavity of females (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFB50F9C6EDAAFB6A" box="[1156,1236,1594,1619]" captionStart="FIGURE 9" captionStartId="18.[151,250,1160,1182]" captionTargetBox="[151,1434,193,1138]" captionTargetId="figure@18.[151,1436,193,1139]" captionTargetPageId="18" captionText="FIGURE 9. Female S 8 – 10 of Cyanogomphini in ventral, inset an outline of subgenital plate under camera lucida. A – B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ, Cachoeiras de Macacu), A. Reserva Ecológica de Guapiaçu (DZRJ 2495), B. Boca do Mato (DZRJ 2494); C. Cyanogomphus angelomachadoi sp. nov. additional specimen (Brazil. MG: Serra do Salitre, ABMM); D. Cyanogomphus sp. (Brazil. GO: Jataí, ABMM); E. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars for photographs = 1 mm." httpUri="https://zenodo.org/record/270862/files/figure.png" pageId="12">Figs 9</figureCitation>
|
||
C), will separate
|
||
<taxonomicName id="06F7EDD9FFC5FF8BFF43F9A2EB79FB4E" box="[151,519,1630,1655]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="12" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
|
||
<emphasis id="F3834A48FFC5FF8BFF43F9A2EB79FB4E" box="[151,519,1630,1655]" italics="true" pageId="12">Cyanogomphus angelomachadoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F3834A48FFC5FF8BFDD9F9A2EB19FB4E" bold="true" box="[525,615,1630,1655]" pageId="12">
|
||
<taxonomicNameLabel id="E8B0F733FFC5FF8BFDD9F9A2EB19FB4E" box="[525,615,1630,1655]" pageId="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from
|
||
<taxonomicName id="06F7EDD9FFC5FF8BFD7FF9A2EA34FB4E" box="[683,842,1630,1655]" class="Insecta" family="Gomphidae" genus="Tibiagomphus" kingdom="Animalia" order="Odonata" pageId="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3834A48FFC5FF8BFD7FF9A2EA34FB4E" box="[683,842,1630,1655]" italics="true" pageId="12">Tibiagomphus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<date id="B549B09AFFC5FF8BFC8DF9A3EA02FB4E" box="[857,892,1631,1655]" pageId="12">V4</date>
|
||
distal portion narrow, apex of cercus posteriorly directed, epiproct dorsally smooth, lacking concavity, and female subgenital plate large and with a narrow mesal incision,
|
||
<figureCitation id="59CC8ADFFFC5FF8BFF29F95AE83CFB86" box="[253,322,1702,1727]" captionStart="FIGURE 6" captionStartId="14.[151,250,1371,1393]" captionTargetBox="[155,1434,153,1337]" captionTargetId="figure@14.[154,1439,151,1349]" captionTargetPageId="14" captionText="FIGURE 6. Male vesica spermalis (VS) of Cyanogomphini. A – C. Cyanogomphus angelomachadoi sp. nov., Paratype (Brazil. MG: Lagoa Santa, MNRJ); D – E. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); F – G. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270859/files/figure.png" pageId="12">Figs 6</figureCitation>
|
||
F–G, 7J–
|
||
<collectingCountry id="B9E0D6CAFFC5FF8BFE7CF95BE8C3FB86" box="[424,445,1703,1727]" name="Iceland" pageId="12">O</collectingCountry>
|
||
, 8G–J, 9E).
|
||
</paragraph>
|
||
<paragraph id="C148965AFFC5FF8AFF13F936ED66FCE9" blockId="12.[151,1437,151,2016]" lastBlockId="13.[151,1437,151,1257]" lastPageId="13" pageId="12">
|
||
The dark mesal part of the mesepisternum close to the dorsal carina (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFC2CF936ED40FBDA" box="[1016,1086,1738,1763]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="12">Fig. 1</figureCitation>
|
||
), epiproct with a larger distal concavity at posterior 0.70 (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFE0BF912EB5DFA3E" box="[479,547,1774,1799]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="12">Fig. 8</figureCitation>
|
||
B) and dark abdomen with pale spots (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFC38F912ED3DFA3E" box="[1004,1091,1774,1799]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="12">Figs 10</figureCitation>
|
||
A–B, 11A–B) distinguish this species from
|
||
<taxonomicName id="06F7EDD9FFC5FF8BFEF9F8EFE89CFA12" box="[301,482,1810,1835]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="12" phylum="Arthropoda" rank="species" species="comparabilis">
|
||
<emphasis id="F3834A48FFC5FF8BFEF9F8EFE89CFA12" box="[301,482,1810,1835]" italics="true" pageId="12">C. comparabilis</emphasis>
|
||
</taxonomicName>
|
||
(mesal part of mesepisternum pale, epiproct with distal concavity reduced to a small posterodorsal area, abdomen almost uniform in color,
|
||
<figureCitation id="59CC8ADFFFC5FF8BFD23F8CAEA42FA76" box="[759,828,1846,1871]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="12">Figs 7</figureCitation>
|
||
E, 8F, 10C).
|
||
<taxonomicName id="06F7EDD9FFC5FF8BFC1FF8CAEC45FA76" box="[971,1339,1846,1871]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="12" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
|
||
<emphasis id="F3834A48FFC5FF8BFC1FF8CAEC45FA76" box="[971,1339,1846,1871]" italics="true" pageId="12">Cyanogomphus angelomachadoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F3834A48FFC5FF8BFA96F8CAECE2FA76" bold="true" box="[1346,1436,1846,1871]" pageId="12">
|
||
<taxonomicNameLabel id="E8B0F733FFC5FF8BFA96F8CAECE2FA76" box="[1346,1436,1846,1871]" pageId="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar in general appearance to
|
||
<taxonomicName id="06F7EDD9FFC5FF8BFD83F8A7EBA1FA4A" box="[599,735,1882,1907]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="12" phylum="Arthropoda" rank="species" species="waltheri">
|
||
<emphasis id="F3834A48FFC5FF8BFD83F8A7EBA1FA4A" box="[599,735,1882,1907]" italics="true" pageId="12">C. waltheri</emphasis>
|
||
</taxonomicName>
|
||
, with which the smallest specimens might be confused.
|
||
<taxonomicName id="06F7EDD9FFC5FF8BFF43F882EB75FAAE" box="[151,523,1918,1943]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="12" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
|
||
<emphasis id="F3834A48FFC5FF8BFF43F882EB75FAAE" box="[151,523,1918,1943]" italics="true" pageId="12">Cyanogomphus angelomachadoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F3834A48FFC5FF8BFDC1F882EB0CFAAE" bold="true" box="[533,626,1918,1943]" pageId="12">
|
||
<taxonomicNameLabel id="E8B0F733FFC5FF8BFDC1F882EB0CFAAE" box="[533,626,1918,1943]" pageId="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished by its smaller size (length of Hw 22.2–24.5 vs. 25.8–31.6 for
|
||
<taxonomicName id="06F7EDD9FFC5FF8BFE90F85FE8B7FA82" box="[324,457,1954,1979]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="12" phylum="Arthropoda" rank="species" species="waltheri">
|
||
<emphasis id="F3834A48FFC5FF8BFE90F85FE8B7FA82" box="[324,457,1954,1979]" italics="true" pageId="12">C. waltheri</emphasis>
|
||
</taxonomicName>
|
||
); general paler color (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFD34F85EEA45FA82" box="[736,827,1954,1979]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="12">Figs 10</figureCitation>
|
||
A–B, 11A–B), with large pale areas connected on mesepisternum (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFE85F83AE8E8FAE6" box="[337,406,1990,2015]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="12">Figs 1</figureCitation>
|
||
A–B, E–F), except the
|
||
<typeStatus id="1E4C28F8FFC5FF8BFD77F834EA7AFAD9" box="[675,772,1992,2016]" pageId="12" type="paratype">paratype</typeStatus>
|
||
male from Serra do Salitre (
|
||
<figureCitation id="59CC8ADFFFC5FF8BFB99F83AEDDFFAE6" box="[1101,1185,1990,2015]" captionStart="FIGURE 10" captionStartId="21.[151,250,1646,1668]" captionTargetBox="[151,1436,195,1620]" captionTargetId="figure@21.[151,1436,195,1622]" captionTargetPageId="21" captionText="FIGURE 10. Male habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Paratype (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); D. Cyanogomphus waltheri Selys, 1873, (Brazil. RJ: Santana de Japuíba district, MNRJ); E. Tibiagomphus noval (Rodrigues Capítulo, 1985), (Brazil. RS: Jaguarão, MNRJ); F. Tibiagomphus uncatus (Fraser, 1947), (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270863/files/figure.png" pageId="12">Fig. 10</figureCitation>
|
||
B), which is distinctly darker with pale areas only partially connected and showing well-defined mesepisternal stripes limited by dark ground color (
|
||
<figureCitation id="59CC8ADFFFC4FF8AFEE2FF40E806FDED" box="[310,376,188,213]" captionStart="FIGURE 1" captionStartId="5.[151,250,1937,1959]" captionTargetBox="[151,1433,196,1895]" captionTargetId="figure@5.[151,1436,195,1913]" captionTargetPageId="5" captionText="FIGURE 1. Synthorax of Cyanogomphus angelomachadoi sp. nov. in dorsal (A, C, E) and lateral (B, D, F) views. A – B. Holotype ♂ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Paratype ♂ (Brazil. MG: Serra do Salitre, ABMM); E – F. Paratype ♀ (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448)." httpUri="https://zenodo.org/record/270854/files/figure.png" pageId="13">Figs 1</figureCitation>
|
||
C–D), much like some
|
||
<taxonomicName id="06F7EDD9FFC4FF8AFD55FF41EB81FDEC" box="[641,767,188,213]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="waltheri">
|
||
<emphasis id="F3834A48FFC4FF8AFD55FF41EB81FDEC" box="[641,767,188,213]" italics="true" pageId="13">C. waltheri</emphasis>
|
||
</taxonomicName>
|
||
specimens (
|
||
<taxonomicName id="06F7EDD9FFC4FF8AFC58FF41ED74FDEC" box="[908,1034,188,213]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="waltheri">
|
||
<emphasis id="F3834A48FFC4FF8AFC58FF41ED74FDEC" box="[908,1034,188,213]" italics="true" pageId="13">C. waltheri</emphasis>
|
||
</taxonomicName>
|
||
pale areas less broad, with first and second antehumeral stripes well defined); dorsal surface of metathoracic tibia yellow to brownish-yellow (
|
||
<figureCitation id="59CC8ADFFFC4FF8AFAEBFF23ECFEFDC1" box="[1343,1408,223,248]" captionStart="FIGURE 3" captionStartId="7.[151,250,894,916]" captionTargetBox="[151,1436,193,871]" captionTargetId="figure@7.[151,1436,193,873]" captionTargetPageId="7" captionText="FIGURE 3. Details of metathoracic tibia and tarsus of males of Cyanogomphini in lateral view. A. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); C. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); D. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270856/files/figure.png" pageId="13">Fig. 3</figureCitation>
|
||
A, black in
|
||
<taxonomicName id="06F7EDD9FFC4FF8AFF2DFEF9E809FC24" box="[249,375,260,285]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="waltheri">
|
||
<emphasis id="F3834A48FFC4FF8AFF2DFEF9E809FC24" box="[249,375,260,285]" italics="true" pageId="13">C. waltheri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<figureCitation id="59CC8ADFFFC4FF8AFE52FEF8E8B6FC25" box="[390,456,260,285]" captionStart="FIGURE 3" captionStartId="7.[151,250,894,916]" captionTargetBox="[151,1436,193,871]" captionTargetId="figure@7.[151,1436,193,873]" captionTargetPageId="7" captionText="FIGURE 3. Details of metathoracic tibia and tarsus of males of Cyanogomphini in lateral view. A. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); B. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Santana de Japuíba district, MNRJ); C. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); D. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270856/files/figure.png" pageId="13">Fig. 3</figureCitation>
|
||
B), distal concavity on epiproct larger (beginning at <0.76 length of epiproct,
|
||
<figureCitation id="59CC8ADFFFC4FF8AFA94FEF8ECFCFC25" box="[1344,1410,260,285]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="13">Fig. 8</figureCitation>
|
||
B; while in
|
||
<taxonomicName id="06F7EDD9FFC4FF8AFED5FED4E8FDFC79" box="[257,387,295,320]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="waltheri">
|
||
<emphasis id="F3834A48FFC4FF8AFED5FED4E8FDFC79" box="[257,387,295,320]" italics="true" pageId="13">C. waltheri</emphasis>
|
||
</taxonomicName>
|
||
at> 0.80 length,
|
||
<figureCitation id="59CC8ADFFFC4FF8AFD88FEDBEBDFFC79" box="[604,673,295,320]" captionStart="FIGURE 8" captionStartId="17.[151,250,1860,1882]" captionTargetBox="[151,1435,197,1827]" captionTargetId="figure@17.[151,1436,195,1835]" captionTargetPageId="17" captionText="FIGURE 8. Male caudal appendages of Cyanogomphini in lateral view (A, C, E, G, I), and detail of epiproct in laterodorsal view (B, D, F, H, J). A – B. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); C – D. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato, DZRJ 2493); E – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – H. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); I – J. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270861/files/figure.png" pageId="13">Fig. 8</figureCitation>
|
||
D) with laterodistal projection in lateral view longer and slender, forefinger-shaped (
|
||
<figureCitation id="59CC8ADFFFC4FF8AFEBBFEB0E8C6FC5D" box="[367,440,332,357]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="13">Figs 7</figureCitation>
|
||
A, 8A; in
|
||
<taxonomicName id="06F7EDD9FFC4FF8AFDE4FEB1EBCFFC5C" box="[560,689,332,357]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="waltheri">
|
||
<emphasis id="F3834A48FFC4FF8AFDE4FEB1EBCFFC5C" box="[560,689,332,357]" italics="true" pageId="13">C. waltheri</emphasis>
|
||
</taxonomicName>
|
||
epiproct laterodistal projection smaller and stout, thumb-shaped,
|
||
<figureCitation id="59CC8ADFFFC4FF8AFF43FE93E9A4FCB1" box="[151,218,367,392]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="13">Figs 7</figureCitation>
|
||
G, 8C); and cercus upright (
|
||
<figureCitation id="59CC8ADFFFC4FF8AFDDAFE93EB2AFCB1" box="[526,596,367,392]" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="13">Figs 7</figureCitation>
|
||
A, 8A) will distinguish it from
|
||
<taxonomicName id="06F7EDD9FFC4FF8AFC7BFE8CED52FCB1" box="[943,1068,367,392]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="waltheri">
|
||
<emphasis id="F3834A48FFC4FF8AFC7BFE8CED52FCB1" box="[943,1068,367,392]" italics="true" pageId="13">C. waltheri</emphasis>
|
||
</taxonomicName>
|
||
(cercus less curved upright,
|
||
<figureCitation id="59CC8ADFFFC4FF8AFABFFE93E9DAFC95" captionStart="FIGURE 7" captionStartId="15.[151,250,1768,1790]" captionTargetBox="[151,1436,197,1742]" captionTargetId="figure@15.[151,1436,195,1743]" captionTargetPageId="15" captionText="FIGURE 7. Male caudal appendages of Cyanogomphini in lateral (A, D, G, J, M), dorsal (B, E, H, K, N), and ventral (C, F, I, L, O) views. A – C. Cyanogomphus angelomachadoi sp. nov., Holotype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1449); D – F. Cyanogomphus comparabilis Belle, 1994, Holotype (Brazil. MT: Diamantino, ABMM); G – I. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Cachoeiras de Macacu, Boca do Mato DZRJ 2493); J – L. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ); M – O. Tibiagomphus uncatus (Fraser, 1947) (Brazil. SC: Nova Teutônia, MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/270860/files/figure.png" pageId="13">Figs 7</figureCitation>
|
||
G, 8C). Both species appear to be allopatric with
|
||
<taxonomicName id="06F7EDD9FFC4FF8AFD1BFE69EAD2FC94" box="[719,940,404,429]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="angelomachadoi" status="sp. nov.">
|
||
<emphasis id="F3834A48FFC4FF8AFD1BFE69EAD2FC94" box="[719,940,404,429]" italics="true" pageId="13">C. angelomachadoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="F3834A48FFC4FF8AFC67FE68ED73FC94" bold="true" box="[947,1037,404,429]" pageId="13">
|
||
<taxonomicNameLabel id="E8B0F733FFC4FF8AFC67FE68ED73FC94" box="[947,1037,404,429]" pageId="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, thus far known as a representative of the Cerrado, and
|
||
<taxonomicName id="06F7EDD9FFC4FF8AFEA2FE44E88AFCE9" box="[374,500,439,464]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="waltheri">
|
||
<emphasis id="F3834A48FFC4FF8AFEA2FE44E88AFCE9" box="[374,500,439,464]" italics="true" pageId="13">C. waltheri</emphasis>
|
||
</taxonomicName>
|
||
, a representative of the Atlantic Forest (
|
||
<figureCitation id="59CC8ADFFFC4FF8AFC6CFE4BED75FCE9" box="[952,1035,439,464]" captionStart="FIGURE 12" captionStartId="24.[151,250,1240,1262]" captionTargetBox="[151,1436,215,1198]" captionTargetId="figure@24.[151,1436,195,1215]" captionTargetPageId="24" captionText="FIGURE 12. Map of southeastern South America with distribution records of Cyanogomphus Selys, 1873, species. Colored areas correspond to the Neotropical regionalization of biotic elements by Morrone (2014). Abbreviations: MI, Misiones Province, Argentina; BA, Bahia; GO, Goiás; MG, Minas Gerais; MT, Mato Grosso; RJ, Rio de Janeiro; RS, Rio Grande do Sul; SC, Santa Catarina, and SP, São Paulo for Brazilian States." httpUri="https://zenodo.org/record/270865/files/figure.png" pageId="13">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89EDC5D1FFC4FF8AFF13FE20EC02FE64" pageId="13" type="distribution">
|
||
<paragraph id="C148965AFFC4FF8AFF13FE20ECE2FCCD" blockId="13.[151,1437,151,1257]" box="[199,1436,476,501]" pageId="13">
|
||
<emphasis id="F3834A48FFC4FF8AFF13FE20E81EFCCC" bold="true" box="[199,352,476,501]" pageId="13">Distribution.</emphasis>
|
||
States of Minas Gerais and São Paulo,
|
||
<collectingCountry id="B9E0D6CAFFC4FF8AFCC3FE20EA20FCCC" box="[791,862,476,501]" name="Brazil" pageId="13">Brazil</collectingCountry>
|
||
, in the Cerrado biogeographical province (
|
||
<figureCitation id="59CC8ADFFFC4FF8AFAE8FE20ECF1FCCD" box="[1340,1423,476,501]" captionStart="FIGURE 12" captionStartId="24.[151,250,1240,1262]" captionTargetBox="[151,1436,215,1198]" captionTargetId="figure@24.[151,1436,195,1215]" captionTargetPageId="24" captionText="FIGURE 12. Map of southeastern South America with distribution records of Cyanogomphus Selys, 1873, species. Colored areas correspond to the Neotropical regionalization of biotic elements by Morrone (2014). Abbreviations: MI, Misiones Province, Argentina; BA, Bahia; GO, Goiás; MG, Minas Gerais; MT, Mato Grosso; RJ, Rio de Janeiro; RS, Rio Grande do Sul; SC, Santa Catarina, and SP, São Paulo for Brazilian States." httpUri="https://zenodo.org/record/270865/files/figure.png" pageId="13">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C148965AFFC4FF8AFF13FE03EC02FE64" blockId="13.[151,1437,151,1257]" pageId="13">
|
||
<emphasis id="F3834A48FFC4FF8AFF13FE03EB41FF21" bold="true" box="[199,575,511,536]" pageId="13">Biological and ecological data.</emphasis>
|
||
Adults were collected at lotic systems, near rivers, streams, and creeks, surrounded by forests (riparian) in rupestrian formations over ironstone outcrops known as “canga”, at 664–766 meters in the core of Serra do Espinhaço mountain range (PNSC and Lagoa Santa sites) and at 1205 meters of elevation in the Serra da Canastra formation (Serra do Salitre site) in the Brazilian Cerrado domain. The pair from PNSC was caught using a Malaise trap and light sheet, non-traditional methods to capture dragonflies (
|
||
<bibRefCitation id="A566EBABFFC4FF8AFAC7FD6CE86DFFF5" author="De" pageId="13" refString="De Almeida, M. V. O., Pinto, A. P., Carvalho, A. L. & Takiya, D. M. (2013) When rare is just a matter of sampling: Unexpected dominance of clubtail dragonflies (Odonata, Gomphidae) through different collecting methods at Parque Nacional da Serra do Cipo, Minas Gerais State, Brazil. Revista Brasileira de Entomologia, 57 (4), 417 - 423. http: // dx. doi. org / 10.1590 / S 0085 - 56262013005000042" type="journal article" year="2013">
|
||
<collectingCountry id="B9E0D6CAFFC4FF8AFAC7FD6CEC4BFF91" box="[1299,1333,656,680]" name="South Georgia and the South Sandwich Islands" pageId="13">De</collectingCountry>
|
||
Almeida
|
||
<emphasis id="F3834A48FFC4FF8AFF43FD49E9B1FFF4" box="[151,207,692,717]" italics="true" pageId="13">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
). Despite more than
|
||
<quantity id="060F3BBFFFC4FF8AFE2FFD48EB2EFFF4" box="[507,592,692,717]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="13" unit="km" value="300.0">300 km</quantity>
|
||
between sites in PNSC and Lagoa Santa (Espinhaço mountain range) from that in Serra do Salitre (Canastra mountain range), they are similar environmentally, typical of the Brazilian Cerrado in that region; temperature, altitude, vegetation and rock outcrops are common to both regions. However, the
|
||
<typeStatus id="1E4C28F8FFC4FF8AFF17FCDCE85DFE01" box="[195,291,800,824]" pageId="13" type="paratype">paratype</typeStatus>
|
||
from São Paulo was collected at 485 meters in an apparently different environment, a sedimentary basin located in the north of Serra Geral rather than rupestrian formation where other specimens were collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89EDC5D1FFC4FF8AFF13FC9BE984FEF1" pageId="13" type="etymology">
|
||
<paragraph id="C148965AFFC4FF8AFF13FC9BE984FEF1" blockId="13.[151,1437,151,1257]" pageId="13">
|
||
<emphasis id="F3834A48FFC4FF8AFF13FC9BE830FEB9" bold="true" box="[199,334,871,896]" pageId="13">Etymology.</emphasis>
|
||
<taxonomicName id="06F7EDD9FFC4FF8AFE80FC9BEB71FEB9" box="[340,527,871,896]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="species" species="angelomachadoi">
|
||
<emphasis id="F3834A48FFC4FF8AFE80FC9BEB71FEB9" box="[340,527,871,896]" italics="true" pageId="13">angelomachadoi</emphasis>
|
||
</taxonomicName>
|
||
(noun phrase in genitive singular formed by a given and surname) specific name in honor of our friend and eminent Brazilian odonatologist Angelo B. M. Machado on the occasion of his 80th birthday.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89EDC5D1FFC4FF8AFF13FC28EAB8F9D1" pageId="13" type="discussion">
|
||
<paragraph id="C148965AFFC4FF8AFF13FC28EAB8F9D1" blockId="13.[151,1437,151,1257]" pageId="13">
|
||
<emphasis id="F3834A48FFC4FF8AFF13FC28E84AFED4" bold="true" box="[199,308,980,1005]" pageId="13">Remarks</emphasis>
|
||
. The only bona fide female from PNSC (DZRJ) is damaged, lacking S5–10 (
|
||
<figureCitation id="59CC8ADFFFC4FF8AFB42FC28ED9BFED5" box="[1174,1253,980,1005]" captionStart="FIGURE 11" captionStartId="23.[151,250,1163,1185]" captionTargetBox="[151,1436,193,1140]" captionTargetId="figure@23.[151,1436,193,1141]" captionTargetPageId="23" captionText="FIGURE 11. Female habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Paratype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448), B. Additional specimen (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); D. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270864/files/figure.png" pageId="13">Fig. 11</figureCitation>
|
||
A), precluding a full description, while the complete female collected at same site of
|
||
<typeStatus id="1E4C28F8FFC4FF8AFC43FC04EA89F929" box="[919,1015,1016,1040]" pageId="13" type="paratype">paratype</typeStatus>
|
||
male from Serra do Salitre (ABMM) is poorly preserved as to general color pattern (
|
||
<figureCitation id="59CC8ADFFFC4FF8AFD72FBE0EB89F90D" box="[678,759,1052,1077]" captionStart="FIGURE 11" captionStartId="23.[151,250,1163,1185]" captionTargetBox="[151,1436,193,1140]" captionTargetId="figure@23.[151,1436,193,1141]" captionTargetPageId="23" captionText="FIGURE 11. Female habitus of Cyanogomphini in lateral view. A – B. Cyanogomphus angelomachadoi sp. nov., A. Paratype (Brazil. MG: Parque Nacional da Serra do Cipó, DZRJ 1448), B. Additional specimen (Brazil. MG: Serra do Salitre, ABMM); C. Cyanogomphus waltheri Selys, 1873 (Brazil. RJ: Reserva Ecológica de Guapiaçu, DZRJ 2495); D. Tibiagomphus noval (Rodrigues Capítulo, 1985) (Brazil. RS: Jaguarão, MNRJ). Scale bars = 10 mm." httpUri="https://zenodo.org/record/270864/files/figure.png" pageId="13">Fig. 11</figureCitation>
|
||
B), leaving in some doubt an association with this species. It was for this reason excluded from the
|
||
<typeStatus id="1E4C28F8FFC4FF8AFDB5FBBCEBEFF961" box="[609,657,1088,1112]" pageId="13">type</typeStatus>
|
||
series. The Lagoa Santa
|
||
<typeStatus id="1E4C28F8FFC4FF8AFC14FBBCED5EF961" box="[960,1056,1088,1112]" pageId="13" type="paratype">paratype</typeStatus>
|
||
(MNRJ) is damaged, with head detached, synthorax crushed, and S1–6 varying from slightly to strongly damaged, with the left auricle broken. The male
|
||
<typeStatus id="1E4C28F8FFC4FF8AFF00FB75E84BF998" box="[212,309,1161,1185]" pageId="13" type="paratype">paratype</typeStatus>
|
||
from São Paulo State in RWG collection was not examined; however, it agrees fairly well with other specimens, showing the mesepisternal stripes divided as in the
|
||
<typeStatus id="1E4C28F8FFC4FF8AFCB9FB50EAB1F9FC" box="[877,975,1196,1221]" pageId="13" type="holotype">holotype</typeStatus>
|
||
(RWG pers. comm.). The females from Goiás (ABMM) may also be this species (see under
|
||
<taxonomicName id="06F7EDD9FFC4FF8AFD34FB33EAF0F9D1" box="[736,910,1231,1256]" class="Insecta" family="Gomphidae" genus="Cyanogomphus" kingdom="Animalia" order="Odonata" pageId="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F3834A48FFC4FF8AFD34FB33EAF0F9D1" box="[736,910,1231,1256]" italics="true" pageId="13">Cyanogomphus</emphasis>
|
||
</taxonomicName>
|
||
sp.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |