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<document id="9D2FE43D7BE5A447FBC005B4F97A64C1" ID="10.11646/zootaxa.4237.2.1SLASH10325" ID-CLB-Dataset="36432" ID-DOI="10.11646/zootaxa.4237.2.1" ID-GBIF-Dataset="e3be2901-f4eb-49ac-8625-78cb1238372a" ID-ISSN="1175-5326" ID-Zenodo-Dep="343756" ID-ZooBank="494C759E-107E-4C5E-A8DC-3CE3DED035AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1488270235565" checkinUser="plazi" docAuthor="Martín, Guillermo San, Álvarez-Campos, Patricia &amp; Hutchings, Pat" docDate="2017" docId="A36887B95A43FFDCFF12D0BD1C46F95D" docLanguage="en" docName="zootaxa.4237.2.1SLASH10325.pdf" docOrigin="Zootaxa 4237 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Syllis monilaris Savigny in Lamarck 1818" docType="treatment" docVersion="9" lastPageNumber="228" masterDocId="5F51FFC15A5BFFC7FF85D2321D2DFFC6" masterDocTitle="The genus Syllis Savigny in Lamarck, 1818 (Annelida: Syllidae: Syllinae) from Australia (second part): four new species and re-description of twelve previously described species" masterLastPageNumber="243" masterPageNumber="201" pageNumber="225" updateTime="1698437396435" updateUser="ExternalLinkService">
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<mods:title id="F01E29D7FCA793E2022E645D257CB97F">The genus Syllis Savigny in Lamarck, 1818 (Annelida: Syllidae: Syllinae) from Australia (second part): four new species and re-description of twelve previously described species</mods:title>
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<taxonomicName id="ECC14D2C5A43FFDFFF12D0BD1F8DFD6C" ID-CoL="7B73R" authority="Savigny in Lamarck, 1818" authorityName="Savigny in Lamarck" authorityYear="1818" box="[151,672,655,682]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="24" pageNumber="225" phylum="Annelida" rank="species" species="monilaris">
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<emphasis id="19B5EABD5A43FFDFFF12D0BD1C7FFD6F" bold="true" box="[151,338,655,681]" italics="true" pageId="24" pageNumber="225">Syllis monilaris</emphasis>
Savigny
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<figureCitation id="B3FA2A2A5A43FFDFFF12D0801C3DFD0D" box="[151,272,690,715]" captionStart="FIGURE 14" captionStartId="25.[151,250,1894,1916]" captionTargetBox="[163,1429,317,1857]" captionTargetId="figure@25.[151,1436,301,1872]" captionTargetPageId="25" captionText="FIGURE 14. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, anterior end, dorsal view. B, anterior parapodium, lateral view. C, midbody parapodium, lateral view. D, posterior parapodium, lateral view. Scale bars: A: 0.4 mm. B – D: 0.2 mm." httpUri="https://zenodo.org/record/343770/files/figure.png" pageId="24" pageNumber="225">Figures 14</figureCitation>
,
<figureCitation id="B3FA2A2A5A43FFDFFE99D0801C17FD0D" box="[284,314,690,715]" captionStart="FIGURE 15" captionStartId="26.[151,250,1939,1961]" captionTargetBox="[301,1281,200,1885]" captionTargetId="figure@26.[291,1288,193,1894]" captionTargetPageId="26" captionText="FIGURE 15. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, compound chaetae, anterior parapodium. B, compound chaetae, midbody parapodium. C, compound chaetae, posterior parapodium. D, aciculae, anterior parapodium. E, aciculae, midbody parapodium. F, aciculae, posterior parapodium. Scale bars: A – F: 20 µm." httpUri="https://zenodo.org/record/343771/files/figure.png" pageId="24" pageNumber="225">15</figureCitation>
,
<figureCitation id="B3FA2A2A5A43FFDFFEC3D0801C4FFD0D" box="[326,354,690,715]" captionStart="FIGURE 16" captionStartId="27.[151,250,1436,1458]" captionTargetBox="[235,1357,193,1408]" captionTargetId="figure@27.[229,1357,193,1414]" captionTargetPageId="27" captionText="FIGURE 16. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, complete specimen. B, midbody, dorsal view. C, posterior end, dorsal view. Syllis nigropunctata Haswell, 1886 (AM W. 29505). D, anterior end, dorsal view. E, prostomium and most anterior segments." httpUri="https://zenodo.org/record/343772/files/figure.png" pageId="24" pageNumber="225">16</figureCitation>
A–C
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<taxonomicName id="ECC14D2C5A43FFDFFF12D0C81FA0FCD6" ID-CoL="7B73R" authority="Savigny in Lamarck, 1818: 318" authorityName="Savigny in Lamarck" authorityPageNumber="318" authorityYear="1818" box="[151,653,762,784]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="24" pageNumber="225" phylum="Annelida" rank="species" species="monilaris">
<emphasis id="19B5EABD5A43FFDFFF12D0C81C1AFCD6" box="[151,311,762,784]" italics="true" pageId="24" pageNumber="225">Syllis monilaris</emphasis>
Savigny
<emphasis id="19B5EABD5A43FFDFFE19D0C91C9DFCD6" box="[412,432,763,784]" italics="true" pageId="24" pageNumber="225">in</emphasis>
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: 318
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;
<treatmentCitation id="AA6010BE5A43FFDFFD1FD0C81E7EFCD6" author="Savigny" box="[666,851,762,784]" page="44" pageId="24" pageNumber="225" year="1822">
<bibRefCitation id="4F504B5E5A43FFDFFD1FD0C81E01FCD6" author="Savigny" box="[666,812,762,784]" pageId="24" pageNumber="225" refString="Savigny, J. C. (1822) Systemes de diverses classes d'animaux sans vertebres. Systeme des annelides, principalement de celles des cotes de l'Egypte et de la Syrie, offrant les caracteres tant distinctifs que naturels des ordres, familles et genres, avec la description des especes. Description de l'Egypte ou Recueil des Observations et des Recherches qui on ete faites en Egypte pendant l'Expedition de l'Armee Francaise, publie par les Ordres de sa Majeste l'Empereur Napoleon le Grand, Histoire Naturelle, Paris, 1 (3), 1 - 128." type="journal article" year="1822">Savigny 1822</bibRefCitation>
: 44
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–45, pl. 4, fig. 3;
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<bibRefCitation id="4F504B5E5A43FFDFFB88D0C819B2FCD6" author="Savigny" box="[1037,1183,762,784]" pageId="24" pageNumber="225" refString="Savigny, J. C. (1826) Systeme des annelides, principalment des celles des cotes de l'Egypte et de la Syrie, offrant des caracteres tant distintifs que naturels des ordres, familles et genres, avec la description des especes. Description de l'Egypte ou Recueil des Observations et des Recherches qui on ete faites en Egypte pendant l'Expedition de l'Armee Francaise. Seconde Edition dediee au Roi publie par C. L. F. Panckoucke, Histoire Naturelle. Mineralogie. Zoologie, 21, 325 - 482." type="journal article" year="1826">Savigny 1826</bibRefCitation>
: 372
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–373, pl. IV, fig. 3;
<treatmentCitation id="AA6010BE5A43FFDFFF42D12B1C50FCE9" author="Licher" box="[199,381,793,815]" page="292" pageId="24" pageNumber="225" year="1999">
<bibRefCitation id="4F504B5E5A43FFDFFF42D12B1C67FCE9" author="Licher" box="[199,330,793,815]" pageId="24" pageNumber="225" refString="Licher, F. (1999) Revision der Gattung Typosyllis Langerhans, 1879 (Polychaeta: Syllidae). Morphologie, Taxonomie und Phylogenie. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 551, 1 - 336." type="journal article" year="1999">Licher, 1999</bibRefCitation>
: 292
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;
<treatmentCitation id="AA6010BE5A43FFDFFE0DD12B1FCAFCE9" ID-DOI="http://doi.org/10.5281/zenodo.3507808" ID-Zenodo-Dep="3507808" author="Alvarez-Campos" box="[392,743,793,815]" httpUri="http://treatment.plazi.org/id/03BB87D0FFCC0725FF4E8386D6AFFADD" page="322" pageId="24" pageNumber="225" year="2015">
<bibRefCitation id="4F504B5E5A43FFDFFE0DD12B1F98FCE9" author="Alvarez-Campos" box="[392,693,793,815]" pageId="24" pageNumber="225" refString="Alvarez-Campos, P., Gil, J. &amp; San Martin, G. (2015 b) Unveiling the Rosetta Stone of syllids: redescription and neotype designation of Syllis monilaris Savigny in Lamarck, 1818, type species of type genus of family Syllidae Grube, 1850 (Annelida). Zootaxa, 4040 (3), 317 - 330." type="journal article" year="2015" yearSuffix="b">
Álvarez-Campos
<emphasis id="19B5EABD5A43FFDFFDBFD1281F4BFCE9" box="[570,614,793,815]" italics="true" pageId="24" pageNumber="225">et al</emphasis>
. 2015b
</bibRefCitation>
: 322
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, figs 1–5.
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</paragraph>
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<taxonomicName id="ECC14D2C5A43FFDFFF12D10A1C7BFC88" ID-CoL="85S3S" box="[151,342,824,846]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="24" pageNumber="225" phylum="Annelida" rank="species" species="moniliformis">
<emphasis id="19B5EABD5A43FFDFFF12D10A1C7BFC88" box="[151,342,824,846]" italics="true" pageId="24" pageNumber="225">Syllis moniliformis</emphasis>
</taxonomicName>
[sic; lapsus calami]
<treatmentCitation id="AA6010BE5A43FFDFFDA9D10A1FFAFC88" author="Grube" box="[556,727,824,846]" page="502" pageId="24" pageNumber="225" year="1869">
<bibRefCitation id="4F504B5E5A43FFDFFDA9D10A1F8AFC88" author="Grube" box="[556,679,824,846]" pageId="24" pageNumber="225" refString="Grube, A. E. (1869) Bechreibungen neuer oder weniger bekannter von Hrn. Prof. Ehrenberg gesammelter Anneliden des rothen Meeres. Monatsbericht der Koniglich Preussischen Akademie der Wissenschaften zu Berlin, Juni 1869, 484 - 521." type="journal article" year="1869">Grube 1869</bibRefCitation>
: 502
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;
<treatmentCitation id="AA6010BE5A43FFDFFD6ED10A1EB1FC88" author="Licher" box="[747,924,824,846]" page="306" pageId="24" pageNumber="225" year="1999">
<bibRefCitation id="4F504B5E5A43FFDFFD6ED10A1E47FC88" author="Licher" box="[747,874,824,846]" pageId="24" pageNumber="225" refString="Licher, F. (1999) Revision der Gattung Typosyllis Langerhans, 1879 (Polychaeta: Syllidae). Morphologie, Taxonomie und Phylogenie. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 551, 1 - 336." type="journal article" year="1999">Licher 1999</bibRefCitation>
: 306
</treatmentCitation>
.
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</paragraph>
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<collectingCounty id="C21F4E235A43FFDFFF12D1481DD2FC55" box="[151,255,890,915]" pageId="24" pageNumber="225">Material</collectingCounty>
examined.
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<collectingCountry id="53D6763F5A43FFDFFE09D1491F03FC55" box="[396,558,891,915]" name="Australia" pageId="24" pageNumber="225">AUSTRALIA</collectingCountry>
,
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AUSTRALIA
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:
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entrance
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,
<location id="2E1E60745A43FFDFFF12D1AD1C16FC71" LSID="urn:lsid:plazi:treatment:A36887B95A43FFDCFF12D0BD1C46F95D:2E1E60745A43FFDFFF12D1AD1C16FC71" box="[151,315,926,951]" country="Australia" county="Material" latitude="-28.465" longLatPrecision="20" longitude="113.778336" municipality="Goss Passage" name="Beacon Island" pageId="24" pageNumber="225" stateProvince="WESTERN AUSTRALIA">Beacon Island</location>
,
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, AM W.29492,
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, coll.
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,
<collectingDate id="4F3BE9875A43FFDFFBF0D1AC183FFC71" box="[1141,1298,926,951]" pageId="24" pageNumber="225" value="1994-05-25">
<date id="5F7F106F5A43FFDFFBF0D1AC183FFC71" box="[1141,1298,926,951]" pageId="24" pageNumber="225" value="1994-05-25">25 May 1994</date>
</collectingDate>
, underneath boulders embedded in coral sand, by
<collectorName id="863453795A43FFDFFDBDD1F11FE3FC1D" box="[568,718,962,987]" pageId="24" pageNumber="225">P. Hutchings.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="2B7E36AF5A43FFDFFF42D1D419F5FBAD" blockId="24.[151,1437,890,2031]" pageId="24" pageNumber="225">
<materialsCitation id="9BA93CF25A43FFDFFF42D1D41F7EFBE5" ID-GBIF-Occurrence="1437751728" collectionCode="ZMB" collectorName="El Tor &amp; C. G. Ehrenberg &amp; A. E. Grube" country="Egypt" location="El Tor" municipality="Comparative" pageId="24" pageNumber="225" specimenCount="2" stateProvince="Sinai Peninsula" typeStatus="neotype">
<emphasis id="19B5EABD5A43FFDFFF42D1D41CFDFC39" bold="true" box="[199,464,998,1023]" pageId="24" pageNumber="225">
<collectingMunicipality id="CB1AACD55A43FFDFFF42D1D41C49FC39" box="[199,356,998,1023]" pageId="24" pageNumber="225">Comparative</collectingMunicipality>
material
</emphasis>
.
<collectingCountry id="53D6763F5A43FFDFFE5BD1D51F16FC39" box="[478,571,999,1023]" name="Egypt" pageId="24" pageNumber="225">EGYPT</collectingCountry>
:
<collectorName id="863453795A43FFDFFDC0D1D41FA6FC38" box="[581,651,998,1023]" pageId="24" pageNumber="225">
<location id="2E1E60745A43FFDFFDC0D1D41FA6FC38" LSID="urn:lsid:plazi:treatment:A36887B95A43FFDCFF12D0BD1C46F95D:2E1E60745A43FFDFFDC0D1D41FA6FC38" box="[581,651,998,1023]" country="Egypt" municipality="Comparative" name="El Tor" pageId="24" pageNumber="225" stateProvince="Sinai Peninsula">El Tor</location>
</collectorName>
(
<collectingRegion id="E905F84D5A43FFDFFD1ED1D41E60FC39" box="[667,845,998,1023]" country="Egypt" name="Shamal Sina'" pageId="24" pageNumber="225">Sinai Peninsula</collectingRegion>
, Gulf of
<collectingRegion id="E905F84D5A43FFDFFC35D1D51EC7FC39" box="[944,1002,999,1023]" country="Egypt" name="As Suways" pageId="24" pageNumber="225">Suez</collectingRegion>
),
<collectorName id="863453795A43FFDFFC7BD1D51986FC39" box="[1022,1195,998,1023]" pageId="24" pageNumber="225">C.G. Ehrenberg</collectorName>
coll., id.
<collectorName id="863453795A43FFDFFA90D1D518B5FC39" box="[1301,1432,998,1023]" pageId="24" pageNumber="225">A.E. Grube</collectorName>
,
<specimenCount id="3DC7FD265A43FFDFFF12D6391C0AFBE5" box="[151,295,1034,1059]" pageId="24" pageNumber="225" type="generic">2 specimens</specimenCount>
:
<collectionCode id="4DD0AE6A5A43FFDFFEB6D6391C5DFBE4" box="[307,368,1035,1058]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" pageId="24" pageNumber="225">ZMB</collectionCode>
F1941,
<typeStatus id="F47A880D5A43FFDFFE55D6391F7EFBE5" box="[464,595,1035,1059]" pageId="24" pageNumber="225" type="neotype">NEOTYPE</typeStatus>
</materialsCitation>
; ZMB 11529, 1 smaller specimen, broken in two pieces.
<materialsCitation id="9BA93CF25A43FFDFFB60D63919F5FBAD" ID-GBIF-Occurrence="1437751739" collectingDate="2014-06-22" collectionCode="APEM" collectorName="Folkestone Pomerania Survey" country="United Kingdom" elevation="33" latitude="51.457222" location="eastern English Channel" longLatPrecision="17" longitude="13.071944" municipality="Folkestone" pageId="24" pageNumber="225" specimenCount="3" typeStatus="neotype">
<collectingCountry id="53D6763F5A43FFDFFB60D6391842FBE5" box="[1253,1391,1035,1059]" name="United Kingdom" pageId="24" pageNumber="225">ENGLAND</collectingCountry>
, off
<collectingMunicipality id="CB1AACD55A43FFDFFF12D61C1C38FB81" box="[151,277,1070,1095]" pageId="24" pageNumber="225">Folkestone</collectingMunicipality>
,
<location id="2E1E60745A43FFDFFEA7D61D1F16FB81" LSID="urn:lsid:plazi:treatment:A36887B95A43FFDCFF12D0BD1C46F95D:2E1E60745A43FFDFFEA7D61D1F16FB81" box="[290,571,1070,1095]" country="United Kingdom" latitude="51.457222" longLatPrecision="17" longitude="13.071944" municipality="Folkestone" name="eastern English Channel" pageId="24" pageNumber="225">eastern English Channel</location>
,
<geoCoordinate id="4EF550685A43FFDFFDCCD61C1FDEFB80" box="[585,755,1070,1095]" direction="north" orientation="latitude" pageId="24" pageNumber="225" precision="15" value="51.457222">51° 27’ 26’’ N</geoCoordinate>
,
<geoCoordinate id="4EF550685A43FFDFFC84D61D1E84FB80" box="[769,937,1071,1095]" direction="east" orientation="longitude" pageId="24" pageNumber="225" precision="15" value="13.071944">13° 03’ 79’’ E</geoCoordinate>
, coarse sediment with
<taxonomicName id="ECC14D2C5A43FFDFFB37D61C18B1FB81" box="[1202,1436,1070,1095]" class="Polychaeta" family="Sabellariidae" genus="Sabellaria" kingdom="Animalia" order="Sabellida" pageId="24" pageNumber="225" phylum="Annelida" rank="species" species="spinulosa">
<emphasis id="19B5EABD5A43FFDFFB37D61C18B1FB81" box="[1202,1436,1070,1095]" italics="true" pageId="24" pageNumber="225">Sabellaria spinulosa</emphasis>
</taxonomicName>
reef,
<elevation id="A0ECD19C5A43FFDFFF55D6611C26FBAC" box="[208,267,1107,1131]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="24" pageNumber="225" unit="m" value="33.0">
<quantity id="EC399B4A5A43FFDFFF55D6611C26FBAC" box="[208,267,1107,1131]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="24" pageNumber="225" unit="m" value="33.0">33 m</quantity>
</elevation>
,
<collectingDate id="4F3BE9875A43FFDFFE92D6611C9DFBAD" box="[279,432,1107,1131]" pageId="24" pageNumber="225" value="2014-06-22">
<date id="5F7F106F5A43FFDFFE92D6611C9DFBAD" box="[279,432,1107,1131]" pageId="24" pageNumber="225" value="2014-06-22">22 June 2014</date>
</collectingDate>
,
<collectorName id="863453795A43FFDFFE3FD6601E26FBAD" box="[442,779,1106,1131]" pageId="24" pageNumber="225">Folkestone Pomerania Survey</collectorName>
,
<specimenCount id="3DC7FD265A43FFDFFC92D6611E8CFBAD" box="[791,929,1106,1131]" pageId="24" pageNumber="225" type="generic">3 specimens</specimenCount>
(collection of
<collectionCode id="4DD0AE6A5A43FFDFFBC1D66119BFFBAD" box="[1092,1170,1107,1131]" pageId="24" pageNumber="225">APEM</collectionCode>
Ltd.).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="63DB65245A43FFDEFF42D6441E88FF3E" lastPageId="25" lastPageNumber="226" pageId="24" pageNumber="225" type="description">
<paragraph id="2B7E36AF5A43FFDEFF42D6441E88FF3E" blockId="24.[151,1437,890,2031]" lastBlockId="25.[151,1437,151,248]" lastPageId="25" lastPageNumber="226" pageId="24" pageNumber="225">
<emphasis id="19B5EABD5A43FFDFFF42D6441C7CFB49" bold="true" box="[199,337,1142,1167]" pageId="24" pageNumber="225">Description</emphasis>
. Body of large size, elongated (
<figureCitation id="B3FA2A2A5A43FFDFFD65D6441E14FB49" box="[736,825,1142,1167]" captionStart="FIGURE 16" captionStartId="27.[151,250,1436,1458]" captionTargetBox="[235,1357,193,1408]" captionTargetId="figure@27.[229,1357,193,1414]" captionTargetPageId="27" captionText="FIGURE 16. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, complete specimen. B, midbody, dorsal view. C, posterior end, dorsal view. Syllis nigropunctata Haswell, 1886 (AM W. 29505). D, anterior end, dorsal view. E, prostomium and most anterior segments." httpUri="https://zenodo.org/record/343772/files/figure.png" pageId="24" pageNumber="225">Fig. 16</figureCitation>
A), complete,
<quantity id="EC399B4A5A43FFDFFC63D6451911FB48" box="[998,1084,1143,1167]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="24" pageNumber="225" unit="mm" value="33.0">33 mm</quantity>
long,
<quantity id="EC399B4A5A43FFDFFB14D64519F4FB48" box="[1169,1241,1143,1167]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="225" unit="mm" value="1.0">1 mm</quantity>
wide, with 133 chaetigers, tapered posteriorly (
<figureCitation id="B3FA2A2A5A43FFDFFD8BD6A81F4BFB75" box="[526,614,1178,1203]" captionStart="FIGURE 16" captionStartId="27.[151,250,1436,1458]" captionTargetBox="[235,1357,193,1408]" captionTargetId="figure@27.[229,1357,193,1414]" captionTargetPageId="27" captionText="FIGURE 16. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, complete specimen. B, midbody, dorsal view. C, posterior end, dorsal view. Syllis nigropunctata Haswell, 1886 (AM W. 29505). D, anterior end, dorsal view. E, prostomium and most anterior segments." httpUri="https://zenodo.org/record/343772/files/figure.png" pageId="24" pageNumber="225">Fig. 16</figureCitation>
A, C). Body opaque, dark, without colour pattern (
<figureCitation id="B3FA2A2A5A43FFDFFB53D6A8181FFB75" box="[1238,1330,1178,1203]" captionStart="FIGURE 14" captionStartId="25.[151,250,1894,1916]" captionTargetBox="[163,1429,317,1857]" captionTargetId="figure@25.[151,1436,301,1872]" captionTargetPageId="25" captionText="FIGURE 14. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, anterior end, dorsal view. B, anterior parapodium, lateral view. C, midbody parapodium, lateral view. D, posterior parapodium, lateral view. Scale bars: A: 0.4 mm. B – D: 0.2 mm." httpUri="https://zenodo.org/record/343770/files/figure.png" pageId="24" pageNumber="225">Figs 14</figureCitation>
A, 16A). Prostomium pentagonal; 4 eyes in trapezoidal arrangement. Palps robust, similar in length to prostomium (
<figureCitation id="B3FA2A2A5A43FFDFFAEAD68C1D99FB3D" captionStart="FIGURE 14" captionStartId="25.[151,250,1894,1916]" captionTargetBox="[163,1429,317,1857]" captionTargetId="figure@25.[151,1436,301,1872]" captionTargetPageId="25" captionText="FIGURE 14. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, anterior end, dorsal view. B, anterior parapodium, lateral view. C, midbody parapodium, lateral view. D, posterior parapodium, lateral view. Scale bars: A: 0.4 mm. B – D: 0.2 mm." httpUri="https://zenodo.org/record/343770/files/figure.png" pageId="24" pageNumber="225">Fig. 14</figureCitation>
A). Median antenna arising between posterior eyes, with 22 articles, slightly longer than combined length of prostomium and palps; lateral antennae shorter than median one, with 12–14 articles each. Peristomium shorter than subsequent segments (
<figureCitation id="B3FA2A2A5A43FFDFFE4AD7181F0EFA85" box="[463,547,1322,1347]" captionStart="FIGURE 14" captionStartId="25.[151,250,1894,1916]" captionTargetBox="[163,1429,317,1857]" captionTargetId="figure@25.[151,1436,301,1872]" captionTargetPageId="25" captionText="FIGURE 14. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, anterior end, dorsal view. B, anterior parapodium, lateral view. C, midbody parapodium, lateral view. D, posterior parapodium, lateral view. Scale bars: A: 0.4 mm. B – D: 0.2 mm." httpUri="https://zenodo.org/record/343770/files/figure.png" pageId="24" pageNumber="225">Fig. 14</figureCitation>
A). Dorsal tentacular cirri shorter than median antenna, with about 16 articles each; ventral tentacular cirri about half length of dorsal ones, with 12 articles each. Dorsal parapodial cirri similar or longer than body width on anterior segments, becoming similar to shorter than body width from midbody backwards, and more or less fusiform, especially from midbody backwards (
<figureCitation id="B3FA2A2A5A43FFDFFBA5D7A41954FA69" box="[1056,1145,1430,1455]" captionStart="FIGURE 14" captionStartId="25.[151,250,1894,1916]" captionTargetBox="[163,1429,317,1857]" captionTargetId="figure@25.[151,1436,301,1872]" captionTargetPageId="25" captionText="FIGURE 14. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, anterior end, dorsal view. B, anterior parapodium, lateral view. C, midbody parapodium, lateral view. D, posterior parapodium, lateral view. Scale bars: A: 0.4 mm. B – D: 0.2 mm." httpUri="https://zenodo.org/record/343770/files/figure.png" pageId="24" pageNumber="225">Figs 14</figureCitation>
A–C, 16A–C), with well defined cirrophores and articles; articles basally and distally smaller than medially; anteriormost dorsal parapodial cirri longer than dorsal tentacular cirri, with 23–26 articles the longer ones, 16–17 the shorter ones (
<figureCitation id="B3FA2A2A5A43FFDFFA8DD7EC1876FA31" box="[1288,1371,1502,1527]" captionStart="FIGURE 14" captionStartId="25.[151,250,1894,1916]" captionTargetBox="[163,1429,317,1857]" captionTargetId="figure@25.[151,1436,301,1872]" captionTargetPageId="25" captionText="FIGURE 14. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, anterior end, dorsal view. B, anterior parapodium, lateral view. C, midbody parapodium, lateral view. D, posterior parapodium, lateral view. Scale bars: A: 0.4 mm. B – D: 0.2 mm." httpUri="https://zenodo.org/record/343770/files/figure.png" pageId="24" pageNumber="225">Fig. 14</figureCitation>
A); at the proventricular level, all dorsal parapodial cirri become shorter and similar in length, with about 10–16 articles (
<figureCitation id="B3FA2A2A5A43FFDFFF1AD4141DDCF9F9" box="[159,241,1574,1599]" captionStart="FIGURE 14" captionStartId="25.[151,250,1894,1916]" captionTargetBox="[163,1429,317,1857]" captionTargetId="figure@25.[151,1436,301,1872]" captionTargetPageId="25" captionText="FIGURE 14. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, anterior end, dorsal view. B, anterior parapodium, lateral view. C, midbody parapodium, lateral view. D, posterior parapodium, lateral view. Scale bars: A: 0.4 mm. B – D: 0.2 mm." httpUri="https://zenodo.org/record/343770/files/figure.png" pageId="24" pageNumber="225">Fig. 14</figureCitation>
A, B), having 12-15 articles in midbody (
<figureCitation id="B3FA2A2A5A43FFDFFD4FD4141E30F9F9" box="[714,797,1574,1599]" captionStart="FIGURE 14" captionStartId="25.[151,250,1894,1916]" captionTargetBox="[163,1429,317,1857]" captionTargetId="figure@25.[151,1436,301,1872]" captionTargetPageId="25" captionText="FIGURE 14. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, anterior end, dorsal view. B, anterior parapodium, lateral view. C, midbody parapodium, lateral view. D, posterior parapodium, lateral view. Scale bars: A: 0.4 mm. B – D: 0.2 mm." httpUri="https://zenodo.org/record/343770/files/figure.png" pageId="24" pageNumber="225">Fig. 14</figureCitation>
C, B); in posteriormost segments, dorsal parapodial cirri with 10–14 articles each, being shorter and smaller than those of midbody, (
<figureCitation id="B3FA2A2A5A43FFDFFC71D478196BF9A5" box="[1012,1094,1610,1635]" captionStart="FIGURE 14" captionStartId="25.[151,250,1894,1916]" captionTargetBox="[163,1429,317,1857]" captionTargetId="figure@25.[151,1436,301,1872]" captionTargetPageId="25" captionText="FIGURE 14. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, anterior end, dorsal view. B, anterior parapodium, lateral view. C, midbody parapodium, lateral view. D, posterior parapodium, lateral view. Scale bars: A: 0.4 mm. B – D: 0.2 mm." httpUri="https://zenodo.org/record/343770/files/figure.png" pageId="24" pageNumber="225">Fig. 14</figureCitation>
D). Parapodia conical, distally bilobed on dorsal view (
<figureCitation id="B3FA2A2A5A43FFDFFE36D45C1F25F941" box="[435,520,1646,1671]" captionStart="FIGURE 14" captionStartId="25.[151,250,1894,1916]" captionTargetBox="[163,1429,317,1857]" captionTargetId="figure@25.[151,1436,301,1872]" captionTargetPageId="25" captionText="FIGURE 14. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, anterior end, dorsal view. B, anterior parapodium, lateral view. C, midbody parapodium, lateral view. D, posterior parapodium, lateral view. Scale bars: A: 0.4 mm. B – D: 0.2 mm." httpUri="https://zenodo.org/record/343770/files/figure.png" pageId="24" pageNumber="225">Fig. 14</figureCitation>
A). Ventral parapodial cirri digitiform, longer than parapodial lobes on anterior parapodia, becoming shorter from midbody onwards (
<figureCitation id="B3FA2A2A5A43FFDFFC8ED4A01E4DF96D" box="[779,864,1682,1707]" captionStart="FIGURE 14" captionStartId="25.[151,250,1894,1916]" captionTargetBox="[163,1429,317,1857]" captionTargetId="figure@25.[151,1436,301,1872]" captionTargetPageId="25" captionText="FIGURE 14. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, anterior end, dorsal view. B, anterior parapodium, lateral view. C, midbody parapodium, lateral view. D, posterior parapodium, lateral view. Scale bars: A: 0.4 mm. B – D: 0.2 mm." httpUri="https://zenodo.org/record/343770/files/figure.png" pageId="24" pageNumber="225">Fig. 14</figureCitation>
B–D). Chaetae compound heterogomph falcigers, with thick shafts and short, triangular, unidentate blades, or very indistinctly bidentate with a minute proximal tooth, sometimes with short spines on margin but usually smooth or almost smooth and of similar shape and size throughout body (
<figureCitation id="B3FA2A2A5A43FFDFFEE0D4CC1C95F8D1" box="[357,440,1790,1815]" captionStart="FIGURE 15" captionStartId="26.[151,250,1939,1961]" captionTargetBox="[301,1281,200,1885]" captionTargetId="figure@26.[291,1288,193,1894]" captionTargetPageId="26" captionText="FIGURE 15. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, compound chaetae, anterior parapodium. B, compound chaetae, midbody parapodium. C, compound chaetae, posterior parapodium. D, aciculae, anterior parapodium. E, aciculae, midbody parapodium. F, aciculae, posterior parapodium. Scale bars: A – F: 20 µm." httpUri="https://zenodo.org/record/343771/files/figure.png" pageId="24" pageNumber="225">Fig. 15</figureCitation>
A–C). Usually dorsalmost falciger of each parapodium with slightly thicker shaft than remaining ones, especially in midbody (
<figureCitation id="B3FA2A2A5A43FFDFFDDFD5101F81F8FD" box="[602,684,1826,1851]" captionStart="FIGURE 15" captionStartId="26.[151,250,1939,1961]" captionTargetBox="[301,1281,200,1885]" captionTargetId="figure@26.[291,1288,193,1894]" captionTargetPageId="26" captionText="FIGURE 15. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, compound chaetae, anterior parapodium. B, compound chaetae, midbody parapodium. C, compound chaetae, posterior parapodium. D, aciculae, anterior parapodium. E, aciculae, midbody parapodium. F, aciculae, posterior parapodium. Scale bars: A – F: 20 µm." httpUri="https://zenodo.org/record/343771/files/figure.png" pageId="24" pageNumber="225">Fig. 15</figureCitation>
C). Anterior parapodia each with 9 compound chaetae, with blades slightly elongated, 57–39 µm long (
<figureCitation id="B3FA2A2A5A43FFDFFDB6D5741FA8F899" box="[563,645,1862,1887]" captionStart="FIGURE 15" captionStartId="26.[151,250,1939,1961]" captionTargetBox="[301,1281,200,1885]" captionTargetId="figure@26.[291,1288,193,1894]" captionTargetPageId="26" captionText="FIGURE 15. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, compound chaetae, anterior parapodium. B, compound chaetae, midbody parapodium. C, compound chaetae, posterior parapodium. D, aciculae, anterior parapodium. E, aciculae, midbody parapodium. F, aciculae, posterior parapodium. Scale bars: A – F: 20 µm." httpUri="https://zenodo.org/record/343771/files/figure.png" pageId="24" pageNumber="225">Fig. 15</figureCitation>
A); midbody parapodia with 4–6 compound chaetae each, 43–34 µm long (
<figureCitation id="B3FA2A2A5A43FFDFFF59D5581C1CF845" box="[220,305,1898,1923]" captionStart="FIGURE 15" captionStartId="26.[151,250,1939,1961]" captionTargetBox="[301,1281,200,1885]" captionTargetId="figure@26.[291,1288,193,1894]" captionTargetPageId="26" captionText="FIGURE 15. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, compound chaetae, anterior parapodium. B, compound chaetae, midbody parapodium. C, compound chaetae, posterior parapodium. D, aciculae, anterior parapodium. E, aciculae, midbody parapodium. F, aciculae, posterior parapodium. Scale bars: A – F: 20 µm." httpUri="https://zenodo.org/record/343771/files/figure.png" pageId="24" pageNumber="225">Fig. 15</figureCitation>
B); posterior parapodia with 4–5 compound chaetae each, blades around 34 µm long (
<figureCitation id="B3FA2A2A5A43FFDFFAA0D5581857F845" box="[1317,1402,1898,1923]" captionStart="FIGURE 15" captionStartId="26.[151,250,1939,1961]" captionTargetBox="[301,1281,200,1885]" captionTargetId="figure@26.[291,1288,193,1894]" captionTargetPageId="26" captionText="FIGURE 15. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, compound chaetae, anterior parapodium. B, compound chaetae, midbody parapodium. C, compound chaetae, posterior parapodium. D, aciculae, anterior parapodium. E, aciculae, midbody parapodium. F, aciculae, posterior parapodium. Scale bars: A – F: 20 µm." httpUri="https://zenodo.org/record/343771/files/figure.png" pageId="24" pageNumber="225">Fig. 15</figureCitation>
C). Dorsal and ventral simple chaetae not seen. Anterior parapodia each with 3–4 small aciculae, distally blunt (
<figureCitation id="B3FA2A2A5A43FFDFFAEAD5BC1D99F80D" captionStart="FIGURE 15" captionStartId="26.[151,250,1939,1961]" captionTargetBox="[301,1281,200,1885]" captionTargetId="figure@26.[291,1288,193,1894]" captionTargetPageId="26" captionText="FIGURE 15. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, compound chaetae, anterior parapodium. B, compound chaetae, midbody parapodium. C, compound chaetae, posterior parapodium. D, aciculae, anterior parapodium. E, aciculae, midbody parapodium. F, aciculae, posterior parapodium. Scale bars: A – F: 20 µm." httpUri="https://zenodo.org/record/343771/files/figure.png" pageId="24" pageNumber="225">Fig. 15</figureCitation>
D); midbody parapodia with three aciculae, similar to those of anterior parapodia, but larger (
<figureCitation id="B3FA2A2A5A43FFDFFB7CD5801861F80D" box="[1273,1356,1970,1995]" captionStart="FIGURE 15" captionStartId="26.[151,250,1939,1961]" captionTargetBox="[301,1281,200,1885]" captionTargetId="figure@26.[291,1288,193,1894]" captionTargetPageId="26" captionText="FIGURE 15. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, compound chaetae, anterior parapodium. B, compound chaetae, midbody parapodium. C, compound chaetae, posterior parapodium. D, aciculae, anterior parapodium. E, aciculae, midbody parapodium. F, aciculae, posterior parapodium. Scale bars: A – F: 20 µm." httpUri="https://zenodo.org/record/343771/files/figure.png" pageId="24" pageNumber="225">Fig. 15</figureCitation>
E) two distally blunt and the rest slightly acuminate aciculae in each posterior parapodia (
<figureCitation id="B3FA2A2A5A43FFDFFBCED5E419B2F829" box="[1099,1183,2006,2031]" captionStart="FIGURE 15" captionStartId="26.[151,250,1939,1961]" captionTargetBox="[301,1281,200,1885]" captionTargetId="figure@26.[291,1288,193,1894]" captionTargetPageId="26" captionText="FIGURE 15. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, compound chaetae, anterior parapodium. B, compound chaetae, midbody parapodium. C, compound chaetae, posterior parapodium. D, aciculae, anterior parapodium. E, aciculae, midbody parapodium. F, aciculae, posterior parapodium. Scale bars: A – F: 20 µm." httpUri="https://zenodo.org/record/343771/files/figure.png" pageId="24" pageNumber="225">Fig. 15</figureCitation>
F). Pharynx extending through about 14 segments; pharyngeal tooth on anterior margin of pharynx. Proventricle through 13 segments; number of muscle cell rows not possible to count without dissection, since the body wall is quite opaque. Apparently, posterior part of body broken and regenerated (
<figureCitation id="B3FA2A2A5A42FFDEFCB4D2ED1EA9FF3E" box="[817,900,223,248]" captionStart="FIGURE 16" captionStartId="27.[151,250,1436,1458]" captionTargetBox="[235,1357,193,1408]" captionTargetId="figure@27.[229,1357,193,1414]" captionTargetPageId="27" captionText="FIGURE 16. Syllis monilaris Savigny in Lamarck, 1818 (AM W. 29492). A, complete specimen. B, midbody, dorsal view. C, posterior end, dorsal view. Syllis nigropunctata Haswell, 1886 (AM W. 29505). D, anterior end, dorsal view. E, prostomium and most anterior segments." httpUri="https://zenodo.org/record/343772/files/figure.png" pageId="25" pageNumber="226">Fig. 16</figureCitation>
C).
</paragraph>
</subSubSection>
<caption id="7FBE66275A42FFDEFF12D55418B1F85C" httpUri="https://zenodo.org/record/343770/files/figure.png" pageId="25" pageNumber="226" targetBox="[163,1429,317,1857]" targetPageId="25">
<paragraph id="2B7E36AF5A42FFDEFF12D55418B1F85C" blockId="25.[151,1436,1893,1946]" pageId="25" pageNumber="226">
<emphasis id="19B5EABD5A42FFDEFF12D5541C32F8BD" bold="true" box="[151,287,1894,1916]" pageId="25" pageNumber="226">FIGURE 14.</emphasis>
<taxonomicName id="ECC14D2C5A42FFDEFEA0D5571FF4F8BA" authority="Savigny in Lamarck, 1818" authorityName="Savigny in Lamarck" authorityYear="1818" box="[293,729,1893,1916]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="25" pageNumber="226" phylum="Annelida" rank="species" species="monilaris">
<emphasis id="19B5EABD5A42FFDEFEA0D5571C70F8BD" bold="true" box="[293,349,1893,1915]" italics="true" pageId="25" pageNumber="226">Syllis</emphasis>
<emphasis id="19B5EABD5A42FFDEFEE6D5571CEAF8BD" bold="true" box="[355,455,1893,1915]" italics="true" pageId="25" pageNumber="226">monilaris</emphasis>
Savigny in Lamarck, 1818
</taxonomicName>
(AM W.29492). A, anterior end, dorsal view. B, anterior parapodium, lateral view. C, midbody parapodium, lateral view. D, posterior parapodium, lateral view. Scale bars: A: 0.4mm. B–D: 0.2 mm.
</paragraph>
</caption>
<caption id="7FBE66275A41FFDDFF12D5A1191FF820" httpUri="https://zenodo.org/record/343771/files/figure.png" pageId="26" pageNumber="227" targetBox="[301,1281,200,1885]" targetPageId="26">
<paragraph id="2B7E36AF5A41FFDDFF12D5A1191FF820" blockId="26.[151,1435,1938,2022]" pageId="26" pageNumber="227">
<emphasis id="19B5EABD5A41FFDDFF12D5A11C0EF86E" bold="true" box="[151,291,1939,1961]" pageId="26" pageNumber="227">FIGURE 15.</emphasis>
<taxonomicName id="ECC14D2C5A41FFDDFEA8D5A01FD8F86F" authority="Savigny in Lamarck, 1818" authorityName="Savigny in Lamarck" authorityYear="1818" box="[301,757,1938,1961]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="26" pageNumber="227" phylum="Annelida" rank="species" species="monilaris">
<emphasis id="19B5EABD5A41FFDDFEA8D5A01CFEF86E" bold="true" box="[301,467,1938,1960]" italics="true" pageId="26" pageNumber="227">Syllis monilaris</emphasis>
Savigny in Lamarck, 1818
</taxonomicName>
(AM W.29492). A, compound chaetae, anterior parapodium. B, compound chaetae, midbody parapodium. C, compound chaetae, posterior parapodium. D, aciculae, anterior parapodium. E, aciculae, midbody parapodium. F, aciculae, posterior parapodium. Scale bars: A–F: 20 µm.
</paragraph>
</caption>
<caption id="7FBE66275A40FFDCFF12D7AE1C99FA29" httpUri="https://zenodo.org/record/343772/files/figure.png" pageId="27" pageNumber="228" targetBox="[235,1357,193,1408]" targetPageId="27">
<paragraph id="2B7E36AF5A40FFDCFF12D7AE1C99FA29" blockId="27.[151,1435,1435,1519]" pageId="27" pageNumber="228">
<emphasis id="19B5EABD5A40FFDCFF12D7AE1C0DFA77" bold="true" box="[151,288,1436,1458]" pageId="27" pageNumber="228">FIGURE 16.</emphasis>
<taxonomicName id="ECC14D2C5A40FFDCFEA2D7A91FF2FA74" authority="Savigny in Lamarck, 1818" authorityName="Savigny in Lamarck" authorityYear="1818" box="[295,735,1435,1458]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="27" pageNumber="228" phylum="Annelida" rank="species" species="monilaris">
<emphasis id="19B5EABD5A40FFDCFEA2D7A91CE7FA77" bold="true" box="[295,458,1435,1457]" italics="true" pageId="27" pageNumber="228">Syllis monilaris</emphasis>
Savigny
<emphasis id="19B5EABD5A40FFDCFDAED7AF1F12FA74" box="[555,575,1437,1458]" italics="true" pageId="27" pageNumber="228">in</emphasis>
Lamarck, 1818
</taxonomicName>
(AM W.29492). A, complete specimen. B, midbody, dorsal view. C, posterior end, dorsal view.
<taxonomicName id="ECC14D2C5A40FFDCFE35D7881E0CFA16" authority="Haswell, 1886" authorityName="Haswell" authorityYear="1886" box="[432,801,1466,1488]" class="Polychaeta" family="Syllidae" genus="Syllis" kingdom="Animalia" order="Phyllodocida" pageId="27" pageNumber="228" phylum="Annelida" rank="species" species="nigropunctata">
<emphasis id="19B5EABD5A40FFDCFE35D7881FA9FA16" bold="true" box="[432,644,1466,1488]" italics="true" pageId="27" pageNumber="228">Syllis nigropunctata</emphasis>
Haswell, 1886
</taxonomicName>
(AM W.29505). D, anterior end, dorsal view. E, prostomium and most anterior segments.
</paragraph>
</caption>
<subSubSection id="63DB65245A40FFDCFF42D4241C46F95D" pageId="27" pageNumber="228" type="discussion">
<paragraph id="2B7E36AF5A40FFDCFF42D4241C46F95D" blockId="27.[151,1436,1558,1691]" pageId="27" pageNumber="228">
<emphasis id="19B5EABD5A40FFDCFF42D4241C19F9E9" bold="true" box="[199,308,1558,1583]" pageId="27" pageNumber="228">Remarks</emphasis>
. This specimen agrees perfectly with the description of the neotype (
<bibRefCitation id="4F504B5E5A40FFDCFBBBD42418BDF9E9" author="Alvarez-Campos" box="[1086,1424,1558,1583]" pageId="27" pageNumber="228" refString="Alvarez-Campos, P., Gil, J. &amp; San Martin, G. (2015 b) Unveiling the Rosetta Stone of syllids: redescription and neotype designation of Syllis monilaris Savigny in Lamarck, 1818, type species of type genus of family Syllidae Grube, 1850 (Annelida). Zootaxa, 4040 (3), 317 - 330." type="journal article" year="2015" yearSuffix="b">
Álvarez-Campos
<emphasis id="19B5EABD5A40FFDCFA80D4251813F9E9" box="[1285,1342,1558,1583]" italics="true" pageId="27" pageNumber="228">et al.</emphasis>
2015b
</bibRefCitation>
), with small differences in size, number of articles of some cirri and other minor details.
<emphasis id="19B5EABD5A40FFDCFF42D46C1C4DF9B1" bold="true" box="[199,352,1630,1655]" pageId="27" pageNumber="228">Distribution.</emphasis>
Red Sea, Australia (Western Australia), English Channel; some previous dubious reports from the Mediterranean.
</paragraph>
</subSubSection>
</treatment>
</document>
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