163 lines
20 KiB
XML
163 lines
20 KiB
XML
<document id="E50001913FB94C7DBF7FF38800AFBF51" ID-DOI="10.5281/zenodo.210736" ID-GBIF-Dataset="61c2bd4f-8605-45f5-8d4a-89dc6f60914e" ID-ISSN="1175-5326" ID-Zenodo-Dep="210736" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460492877430" checkinUser="plazi" docAuthor="Johanson, Kjell Arne & Oláh, János" docDate="2012" docId="9F3E87DD5603FFF8E89AFE10FE40FEE6" docLanguage="en" docName="zt03354p058.pdf" docOrigin="Zootaxa 3354" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Chimarra devoensis Johanson & Oláh, 2012, new species" docType="treatment" docVersion="9" lastPageNumber="47" masterDocId="6307FFA5562EFFD6E80DFF84FF8BFFD9" masterDocTitle="Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species" masterLastPageNumber="58" masterPageNumber="1" pageNumber="46" updateTime="1698310454288" updateUser="plazi">
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<mods:title id="7F81FCCE8898A7A07EC63DD49A2525A1">Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species</mods:title>
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<mods:namePart id="8DEBD041059365B6569B6E90F1EDEB68">Johanson, Kjell Arne</mods:namePart>
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<mods:namePart id="23623272AF3133DC555E880F8C9A324B">Oláh, János</mods:namePart>
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<mods:typeOfResource id="A8697EED4A2A6212F762FDAB31472883">text</mods:typeOfResource>
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<mods:title id="4FACD9B65B8C5DFB48918A54D8B31D3A">Zootaxa</mods:title>
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<mods:part id="57691AED26C5CE92388B3C94411CB536">
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<mods:date id="74DEB57A265F9434A612BF628C80216B">2012</mods:date>
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<mods:identifier id="A63DFECF32D254E59FE00D361194C119" type="DOI">10.5281/zenodo.210736</mods:identifier>
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<mods:identifier id="DD9D955B532103BF01941B930DE92BD0" type="ISSN">1175-5326</mods:identifier>
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<treatment id="9F3E87DD5603FFF8E89AFE10FE40FEE6" ID-DOI="http://doi.org/10.5281/zenodo.5664510" ID-GBIF-Taxon="119520086" ID-Zenodo-Dep="5664510" LSID="urn:lsid:plazi:treatment:9F3E87DD5603FFF8E89AFE10FE40FEE6" httpUri="http://treatment.plazi.org/id/9F3E87DD5603FFF8E89AFE10FE40FEE6" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
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<subSubSection id="5F8D65405603FFFBE89AFE10FBFDFD13" pageId="45" pageNumber="46" type="nomenclature">
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<paragraph id="172836CB5603FFFBE89AFE10FDACFE74" blockId="45.[151,551,403,462]" box="[151,551,403,430]" pageId="45" pageNumber="46">
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<heading id="4C6081A75603FFFBE89AFE10FDACFE74" bold="true" box="[151,551,403,430]" fontSize="11" level="1" pageId="45" pageNumber="46" reason="1">
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<emphasis id="25E3EAD95603FFFBE89AFE10FDACFE74" bold="true" box="[151,551,403,430]" pageId="45" pageNumber="46">
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<taxonomicName id="D0974D485603FFFBE89AFE10FE03FE77" box="[151,392,404,430]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="devoensis" status="sp. nov.">
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<emphasis id="25E3EAD95603FFFBE89AFE10FE03FE77" bold="true" box="[151,392,404,430]" italics="true" pageId="45" pageNumber="46">Chimarra devoensis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="3ED057A25603FFFBE99AFE17FDACFE74" box="[407,551,403,429]" pageId="45" pageNumber="46" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="172836CB5603FFFBE89AFE32FEEAFE17" blockId="45.[151,551,403,462]" box="[151,353,438,462]" pageId="45" pageNumber="46">
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<figureCitation id="8FAC2A4E5603FFFBE89AFE32FF78FE17" box="[151,243,438,462]" captionStart="FIGURES 22 – 29" captionStartId="12.[151,264,1912,1935]" captionTargetBox="[224,1369,239,1869]" captionTargetId="figure@12.[198,1403,193,1901]" captionTargetPageId="12" captionText="FIGURES 22 – 29. Right wings, dorsal views. 22 — C. cartwrighti, new species, holotype; 23 — C. kimminsi, new species, holotype; 24 — C. kadavuensis, new species, holotype; 25 — C. lavensis, new species, holotype; 26 — C. devoensis, new species, holotype; 27 — C. helomyzida, new species, holotype; 28 — C. tipulida, new species, holotype; 29 — C. psychodida, new species, holotype." httpUri="https://zenodo.org/record/210744/files/figure.png" pageId="45" pageNumber="46">Figs. 26</figureCitation>
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,
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<figureCitation id="8FAC2A4E5603FFFBE8F2FE32FEEAFE17" box="[255,353,438,462]" captionStart="FIGURES 139 – 144" captionStartId="46.[151,264,1343,1366]" captionTargetBox="[177,1404,410,1269]" captionTargetId="figure@46.[151,1436,375,1321]" captionTargetPageId="46" captionText="FIGURES 139 – 144. Chimarra devoensis, new species, holotype. 139 — genitalia, left lateral view; 140 — genitalia, dorsal view; 141 — genitalia, ventral view; 142 — left gonopod, caudal view; 143 — phallic apparatus, left lateral view; 144 — phallic apparatus, ventral view." httpUri="https://zenodo.org/record/210767/files/figure.png" pageId="45" pageNumber="46">139–144</figureCitation>
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</paragraph>
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<paragraph id="172836CB5603FFFBE89AFE7AFBFDFD13" blockId="45.[151,1437,510,2014]" pageId="45" pageNumber="46">
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The male genitalia of this species resemble those of
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<taxonomicName id="D0974D485603FFFBEAEBFE7BFCEAFDCF" box="[742,865,510,534]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="lavensis">
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<emphasis id="25E3EAD95603FFFBEAEBFE7BFCEAFDCF" box="[742,865,510,534]" italics="true" pageId="45" pageNumber="46">C. lavensis</emphasis>
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</taxonomicName>
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and
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<emphasis id="25E3EAD95603FFFBEB94FE7BFBA4FDCF" box="[921,1071,511,534]" italics="true" pageId="45" pageNumber="46">
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<taxonomicName id="D0974D485603FFFBEB94FE7BFBA0FDCF" box="[921,1067,511,534]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="vanuensis">C. vanuensis</taxonomicName>
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,
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</emphasis>
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particularly in the presence of a strongly dorsad-oriented dorsal branch of tergum X. In
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<taxonomicName id="D0974D485603FFFBEAF2FDA7FC07FDE2" box="[767,908,547,571]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="devoensis">
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<emphasis id="25E3EAD95603FFFBEAF2FDA7FC07FDE2" box="[767,908,547,571]" italics="true" pageId="45" pageNumber="46">C. devoensis</emphasis>
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</taxonomicName>
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the dorsal process of tergum X is accompanied by a triangular, posterad oriented process between on the dorsal branches of tergum X. Segment IX has very long, anterolateral plates that are shorter in
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<taxonomicName id="D0974D485603FFFBEA4EFDEFFD35FD5A" box="[579,702,619,643]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="lavensis">
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<emphasis id="25E3EAD95603FFFBEA4EFDEFFD35FD5A" box="[579,702,619,643]" italics="true" pageId="45" pageNumber="46">C. lavensis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="D0974D485603FFFBEAFAFDEFFC0CFD5A" box="[759,903,619,643]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="vanuensis">
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<emphasis id="25E3EAD95603FFFBEAFAFDEFFC0CFD5A" box="[759,903,619,643]" italics="true" pageId="45" pageNumber="46">C. vanuensis</emphasis>
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</taxonomicName>
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. In addition, the base of each gonopod is more strongly expanded ventrad compared to the other species. The species is easily distinguished from
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<taxonomicName id="D0974D485603FFFBECE7FD0BFAF3FD7F" box="[1258,1400,655,678]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="vanuensis">
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<emphasis id="25E3EAD95603FFFBECE7FD0BFAF3FD7F" box="[1258,1400,655,678]" italics="true" pageId="45" pageNumber="46">C. vanuensis</emphasis>
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</taxonomicName>
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by the absence of a large pale spot between Cu1 and end of Cu2 (arculus) in each forewing.
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</paragraph>
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</subSubSection>
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<subSubSection id="5F8D65405603FFF8E8CAFD53FE29FF76" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
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<paragraph id="172836CB5603FFFBE8CAFD53FE12FCCB" blockId="45.[151,1437,510,2014]" pageId="45" pageNumber="46">
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<emphasis id="25E3EAD95603FFFBE8CAFD53FE8FFD36" bold="true" box="[199,260,727,751]" pageId="45" pageNumber="46">
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<collectingCountry id="6F80765B5603FFFBE8CAFD53FE8FFD36" box="[199,260,727,751]" name="Malta" pageId="45" pageNumber="46">Male</collectingCountry>
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</emphasis>
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. Head and thorax greyish brown, dorsal part of meso- and metathorax dark brown. Foreleg anterior claw as long as foreleg spur.
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</paragraph>
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<paragraph id="172836CB5603FFFBE8CAFC9AFE9CFB8F" blockId="45.[151,1437,510,2014]" pageId="45" pageNumber="46">
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<emphasis id="25E3EAD95603FFFBE8CAFC9AFE99FCEE" bold="true" box="[199,274,798,823]" pageId="45" pageNumber="46">Wings</emphasis>
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(
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<figureCitation id="8FAC2A4E5603FFFBE92EFC9AFEF3FCEF" box="[291,376,798,822]" captionStart="FIGURES 22 – 29" captionStartId="12.[151,264,1912,1935]" captionTargetBox="[224,1369,239,1869]" captionTargetId="figure@12.[198,1403,193,1901]" captionTargetPageId="12" captionText="FIGURES 22 – 29. Right wings, dorsal views. 22 — C. cartwrighti, new species, holotype; 23 — C. kimminsi, new species, holotype; 24 — C. kadavuensis, new species, holotype; 25 — C. lavensis, new species, holotype; 26 — C. devoensis, new species, holotype; 27 — C. helomyzida, new species, holotype; 28 — C. tipulida, new species, holotype; 29 — C. psychodida, new species, holotype." httpUri="https://zenodo.org/record/210744/files/figure.png" pageId="45" pageNumber="46">Fig. 26</figureCitation>
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). Forewings
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<quantity id="D06F9B2E5603FFFBEA07FC9AFDEAFCEF" box="[522,609,798,822]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="45" pageNumber="46" unit="mm" value="5.1">5.1 mm</quantity>
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(n=1), brown. Forewings broad, ratio of length to breadth 3.2; R1 nearly straight before crossvein r; radial sector slightly produced anterad immediately before discoidal cell; discoidal cell originating at mid-length of wing, nearly
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<date id="6329100B5603FFFBEA78FCE2FD1AFCA7" box="[629,657,870,894]" pageId="45" pageNumber="46">3x</date>
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longer than wide; median cell as long as discoidal cell; crossvein r originating from basis of fork I; fork I originating before crossvein s at distance equal to length of crossvein s; nygma located near base of fork II; fork
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<date id="6329100B5603FFFBEA6CFC2BFD05FC1F" box="[609,654,942,966]" pageId="45" pageNumber="46">III 1</date>
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/6th as long as wing; fork V shorter than fork II; Cu2 ending in wing margin well separated from A.
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<collectingCountry id="6F80765B5603FFFBEA0DFC56FDB3FC33" box="[512,568,978,1002]" name="India" pageId="45" pageNumber="46">Hind</collectingCountry>
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wings
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<quantity id="D06F9B2E5603FFFBEA83FC56FD6DFC33" box="[654,742,978,1002]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="45" pageNumber="46" unit="mm" value="4.2">4.2 mm</quantity>
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(n=1), brown; ratio of length to breadth 3.0; margin weakly incurved where Cu1 and Cu2 fused with margin; fork I originating slightly before anterodistal corner of discoidal cell; fork III as long as discoidal cell and1/10th as long as wing; fork V slightly shorter than fork I; 1A+2Ab
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<date id="6329100B5603FFFBED6CFB9EFAF6FBEB" box="[1377,1405,1050,1074]" pageId="45" pageNumber="46">3x</date>
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as long as 1A.
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</paragraph>
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<paragraph id="172836CB5603FFFBE8CAFBE7FA86F804" blockId="45.[151,1437,510,2014]" pageId="45" pageNumber="46">
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<emphasis id="25E3EAD95603FFFBE8CAFBE7FEFFFBA2" bold="true" box="[199,372,1122,1147]" pageId="45" pageNumber="46">
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<collectingCountry id="6F80765B5603FFFBE8CAFBE7FE8FFBA2" box="[199,260,1123,1147]" name="Malta" pageId="45" pageNumber="46">Male</collectingCountry>
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genitalia
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</emphasis>
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(
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<figureCitation id="8FAC2A4E5603FFFBE988FBE6FDA6FBA3" box="[389,557,1122,1146]" captionStart="FIGURES 139 – 144" captionStartId="46.[151,264,1343,1366]" captionTargetBox="[177,1404,410,1269]" captionTargetId="figure@46.[151,1436,375,1321]" captionTargetPageId="46" captionText="FIGURES 139 – 144. Chimarra devoensis, new species, holotype. 139 — genitalia, left lateral view; 140 — genitalia, dorsal view; 141 — genitalia, ventral view; 142 — left gonopod, caudal view; 143 — phallic apparatus, left lateral view; 144 — phallic apparatus, ventral view." httpUri="https://zenodo.org/record/210767/files/figure.png" pageId="45" pageNumber="46">Figs. 139–144</figureCitation>
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). Segment IX about 1.5x taller than long in lateral view; anterodorsally produced into long, narrow, rounded, anterodorsad oriented plate; anteroventral plate large, triangular, pointing posterad in lateral view; anterodorsal margin widely and deeply concave, U-shaped; ventral margin irregularly convex, incised at vertical apodeme; each posterior margin forming uniformly convex plate originating immediately below cercus; segment IX ventrally with setae restricted to narrow band before posterior margin (
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<figureCitation id="8FAC2A4E5603FFFBECD1FB69FAB6FADC" box="[1244,1341,1261,1285]" captionStart="FIGURES 139 – 144" captionStartId="46.[151,264,1343,1366]" captionTargetBox="[177,1404,410,1269]" captionTargetId="figure@46.[151,1436,375,1321]" captionTargetPageId="46" captionText="FIGURES 139 – 144. Chimarra devoensis, new species, holotype. 139 — genitalia, left lateral view; 140 — genitalia, dorsal view; 141 — genitalia, ventral view; 142 — left gonopod, caudal view; 143 — phallic apparatus, left lateral view; 144 — phallic apparatus, ventral view." httpUri="https://zenodo.org/record/210767/files/figure.png" pageId="45" pageNumber="46">Fig. 141</figureCitation>
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). Dorsal margin of segment IX undulating in lateral view; anterior margin straight, membranous in dorsal view. In ventral view segment IX with almost parallel lateral margins, slightly narrower at transverse apodeme; anterior margin concave; posterior margin straight, without central projection. Tergum X nearly as long as segment IX, divided into pair of lateral branches at basis; in lateral view each lateral branch with large, straight, dorsad oriented dorsal branch, short, widely triangular posterad orienting process below dorsal branch, and long tube-shaped, nearly straight ventral branch with 2 apical sensillae. In dorsal and ventral view, ventral processes oriented posterad, slightly diverging, tapering along their length. Cerci small, wart-like, located at posterior margin of segment IX at mid-height of tergum X and covered by long setae. Gonopods longer than length of segment IX; divided into very broad basal third and narrow distal 2/3rds in lateral view; distal 2/3rds nearly parallel-sided; each gonopod unbranched in lateral view; with well-developed dorsomesal lobe and ventromesal lobe of mesal process in ventral view. Anterodorsal margin of each gonopod basally convex, distally straight; posteroventral margin concave, with strongly expanded setal bases; ventral margin of basis strongly expanded ventrad; distal apex without megasetae. In ventral view, gonopods slightly broader at base than rest of gonopods, with undulating margins; gonopods weakly curved mesad along their length. Mesal branch of each gonopod divided into well-developed ventromesal and dorsomesal lobe; ventromesal lobe strongly produced dorsad, darkly pigmented, covered by minute setae; dorsomesal lobe curved ventrad, with pointed apex. Phallic apparatus slightly longer than rest of genitalia, straight along its length; phallotheca, in lateral view nearly
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<date id="6329100B5603FFFBEAD5F8BEFD7FF88B" box="[728,756,1850,1874]" pageId="45" pageNumber="46">4x</date>
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thicker than posterior part; in ventral view nearly
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<date id="6329100B5603FFFBED26F8BEFACCF88B" box="[1323,1351,1850,1874]" pageId="45" pageNumber="46">2x</date>
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thicker than posterior part; apicoventral spine absent; phallotremal sclerite forming simple, 3-rayed structure in lateral view; in ventral view forming single posterad directed arrow head situated on basis with 4 anterad pointing spines; 4 nearly black, short, variously directed endothecal spines present, about as long as diameter of narrowest part of phallotheca; endothecal spicules pointing posterodorsad, present in group below phallotremal sclerite.
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</paragraph>
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<paragraph id="172836CB5600FFF8E8CAFF1CFE29FF76" blockId="46.[151,1436,151,320]" box="[199,418,151,176]" pageId="46" pageNumber="47">
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<emphasis id="25E3EAD95600FFF8E8CAFF1CFE95FF69" bold="true" box="[199,286,152,176]" pageId="46" pageNumber="47">Female</emphasis>
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. Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="5F8D65405600FFF8E8CAFF38FDFDFF2E" pageId="46" pageNumber="47" type="materials_examined">
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||
<paragraph id="172836CB5600FFF8E8CAFF38FDFDFF2E" blockId="46.[151,1436,151,320]" pageId="46" pageNumber="47">
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<emphasis id="25E3EAD95600FFF8E8CAFF38FD9FFF0D" bold="true" box="[199,532,187,212]" pageId="46" pageNumber="47">
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<typeStatus id="C82C88695600FFF8E8CAFF38FEBFFF0D" box="[199,308,188,212]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
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male: TAVEUNI:
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</emphasis>
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Cakadrove Prov., Devo Peak, Malaise trap,
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<date id="6329100B5600FFF8EC26FF38FB4AFF0D" box="[1067,1217,188,212]" pageId="46" pageNumber="47" value="2003-01-10" valueMax="2003-01-16" valueMin="2003-01-10">10–16.i.2003</date>
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, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc#37] [
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<collectionCode id="7186AE0E5600FFF8EA2EFF5BFDE0FF2E" box="[547,619,223,247]" pageId="46" pageNumber="47">FNIC</collectionCode>
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].
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</paragraph>
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</subSubSection>
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<subSubSection id="5F8D65405600FFF8E8CAFE80FC7CFEC5" box="[199,1015,260,285]" pageId="46" pageNumber="47" type="etymology">
|
||
<paragraph id="172836CB5600FFF8E8CAFE80FC7CFEC5" blockId="46.[151,1436,151,320]" box="[199,1015,260,285]" pageId="46" pageNumber="47">
|
||
<emphasis id="25E3EAD95600FFF8E8CAFE80FEDAFEC5" bold="true" box="[199,337,260,284]" pageId="46" pageNumber="47">Etymology:</emphasis>
|
||
<taxonomicName id="D0974D485600FFF8E955FE81FE40FEC5" box="[344,459,261,284]" class="Insecta" family="Philopotamidae" genus="Chimarra" kingdom="Animalia" order="Trichoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="devoensis">
|
||
<emphasis id="25E3EAD95600FFF8E955FE81FE40FEC5" box="[344,459,261,284]" italics="true" pageId="46" pageNumber="47">Devoensis</emphasis>
|
||
</taxonomicName>
|
||
, after Devo Peak, the
|
||
<typeStatus id="C82C88695600FFF8EAC8FE81FD7EFEC4" box="[709,757,261,285]" pageId="46" pageNumber="47">type</typeStatus>
|
||
locality of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5F8D65405600FFF8E8CAFEA3FE40FEE6" box="[199,459,295,320]" pageId="46" pageNumber="47" type="distribution">
|
||
<paragraph id="172836CB5600FFF8E8CAFEA3FE40FEE6" blockId="46.[151,1436,151,320]" box="[199,459,295,320]" pageId="46" pageNumber="47">
|
||
<emphasis id="25E3EAD95600FFF8E8CAFEA3FEE9FE99" bold="true" box="[199,354,295,320]" pageId="46" pageNumber="47">Distribution:</emphasis>
|
||
Taveuni.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |