Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species
Author
Johanson, Kjell Arne
Author
Oláh, János
text
Zootaxa
2012
3354
1
58
journal article
10.5281/zenodo.210736
61c2bd4f-8605-45f5-8d4a-89dc6f60914e
1175-5326
210736
Chimarra devoensis
,
new species
Figs. 26
,
139–144
The male genitalia of this species resemble those of
C. lavensis
and
C. vanuensis
,
particularly in the presence of a strongly dorsad-oriented dorsal branch of tergum X. In
C. devoensis
the dorsal process of tergum X is accompanied by a triangular, posterad oriented process between on the dorsal branches of tergum X. Segment IX has very long, anterolateral plates that are shorter in
C. lavensis
and
C. vanuensis
. In addition, the base of each gonopod is more strongly expanded ventrad compared to the other species. The species is easily distinguished from
C. vanuensis
by the absence of a large pale spot between Cu1 and end of Cu2 (arculus) in each forewing.
Male
. Head and thorax greyish brown, dorsal part of meso- and metathorax dark brown. Foreleg anterior claw as long as foreleg spur.
Wings
(
Fig. 26
). Forewings
5.1 mm
(n=1), brown. Forewings broad, ratio of length to breadth 3.2; R1 nearly straight before crossvein r; radial sector slightly produced anterad immediately before discoidal cell; discoidal cell originating at mid-length of wing, nearly
3x
longer than wide; median cell as long as discoidal cell; crossvein r originating from basis of fork I; fork I originating before crossvein s at distance equal to length of crossvein s; nygma located near base of fork II; fork
III 1
/6th as long as wing; fork V shorter than fork II; Cu2 ending in wing margin well separated from A.
Hind
wings
4.2 mm
(n=1), brown; ratio of length to breadth 3.0; margin weakly incurved where Cu1 and Cu2 fused with margin; fork I originating slightly before anterodistal corner of discoidal cell; fork III as long as discoidal cell and1/10th as long as wing; fork V slightly shorter than fork I; 1A+2Ab
3x
as long as 1A.
Male
genitalia
(
Figs. 139–144
). Segment IX about 1.5x taller than long in lateral view; anterodorsally produced into long, narrow, rounded, anterodorsad oriented plate; anteroventral plate large, triangular, pointing posterad in lateral view; anterodorsal margin widely and deeply concave, U-shaped; ventral margin irregularly convex, incised at vertical apodeme; each posterior margin forming uniformly convex plate originating immediately below cercus; segment IX ventrally with setae restricted to narrow band before posterior margin (
Fig. 141
). Dorsal margin of segment IX undulating in lateral view; anterior margin straight, membranous in dorsal view. In ventral view segment IX with almost parallel lateral margins, slightly narrower at transverse apodeme; anterior margin concave; posterior margin straight, without central projection. Tergum X nearly as long as segment IX, divided into pair of lateral branches at basis; in lateral view each lateral branch with large, straight, dorsad oriented dorsal branch, short, widely triangular posterad orienting process below dorsal branch, and long tube-shaped, nearly straight ventral branch with 2 apical sensillae. In dorsal and ventral view, ventral processes oriented posterad, slightly diverging, tapering along their length. Cerci small, wart-like, located at posterior margin of segment IX at mid-height of tergum X and covered by long setae. Gonopods longer than length of segment IX; divided into very broad basal third and narrow distal 2/3rds in lateral view; distal 2/3rds nearly parallel-sided; each gonopod unbranched in lateral view; with well-developed dorsomesal lobe and ventromesal lobe of mesal process in ventral view. Anterodorsal margin of each gonopod basally convex, distally straight; posteroventral margin concave, with strongly expanded setal bases; ventral margin of basis strongly expanded ventrad; distal apex without megasetae. In ventral view, gonopods slightly broader at base than rest of gonopods, with undulating margins; gonopods weakly curved mesad along their length. Mesal branch of each gonopod divided into well-developed ventromesal and dorsomesal lobe; ventromesal lobe strongly produced dorsad, darkly pigmented, covered by minute setae; dorsomesal lobe curved ventrad, with pointed apex. Phallic apparatus slightly longer than rest of genitalia, straight along its length; phallotheca, in lateral view nearly
4x
thicker than posterior part; in ventral view nearly
2x
thicker than posterior part; apicoventral spine absent; phallotremal sclerite forming simple, 3-rayed structure in lateral view; in ventral view forming single posterad directed arrow head situated on basis with 4 anterad pointing spines; 4 nearly black, short, variously directed endothecal spines present, about as long as diameter of narrowest part of phallotheca; endothecal spicules pointing posterodorsad, present in group below phallotremal sclerite.
Female
. Unknown.
Holotype
male: TAVEUNI:
Cakadrove Prov., Devo Peak, Malaise trap,
10–16.i.2003
, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc#37] [
FNIC
].
Etymology:
Devoensis
, after Devo Peak, the
type
locality of the species.
Distribution:
Taveuni.