251 lines
29 KiB
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251 lines
29 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/dez.65.21000" ID-GBIF-Dataset="03cce405-5887-4474-b040-35de18e8d020" ID-GBIF-Taxon="182223660" ID-Pensoft-Pub="1860-1324-1-1" ID-Pensoft-UUID="77D7AEC9294554E5BBDCABA37901022A" ID-Zenodo-Dep="1166397" ID-ZooBank="73DEE0F32BB04A21B4455E168FE50F54" LSID="urn:lsid:zoobank.org:act:03AF91A5-C31D-414B-852D-D6A40D71C718" ModsDocID="1860-1324-65-1-1" checkinTime="1621372489409" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2018" docId="92B4A6E2A7B1BD884D174D9FE634A14F" docLanguage="en" docName="DeutEntomolZeit 65(1): 1-63" docOrigin="Deutsche Entomologische Zeitschrift 65 (1)" docPubDate="2018-01-18" docSource="http://dx.doi.org/10.3897/dez.65.21000" docTitle="Eucyclothorax storeyi subsp. subgen. n." docType="treatment" docVersion="3" id="77D7AEC9294554E5BBDCABA37901022A" lastPageId="11" lastPageNumber="12" masterDocId="77D7AEC9294554E5BBDCABA37901022A" masterDocTitle="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel" masterLastPageNumber="63" masterPageNumber="1" pageId="9" pageNumber="10" updateTime="1643465225333" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liebherr, James K.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9831-884X</mods:nameIdentifier>
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<mods:affiliation>Cornell University Insect Collection, John H. and Anna B. Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
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<mods:nameIdentifier type="email">jkl5@cornell.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2018-01-18</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>65</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>63</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/dez.65.21000</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.65.21000</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-1324-1-1</mods:identifier>
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<mods:identifier type="ZooBank">73DEE0F32BB04A21B4455E168FE50F54</mods:identifier>
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<mods:identifier type="Pensoft-UUID">77D7AEC9294554E5BBDCABA37901022A</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1166397</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182223660" LSID="urn:lsid:zoobank.org:act:03AF91A5-C31D-414B-852D-D6A40D71C718" httpUri="http://treatment.plazi.org/id/92B4A6E2A7B1BD884D174D9FE634A14F" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
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<subSubSection pageId="9" pageNumber="10" type="nomenclature">
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<paragraph pageId="9" pageNumber="10">
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<taxonomicName LSID="http://zoobank.org/03AF91A5-C31D-414B-852D-D6A40D71C718" authority="Liebherr, 2018" authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Eucyclothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eucyclothorax subsp. subgen. n." order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="subSpecies" species="storeyi" subSpecies="subgen. n.">Eucyclothorax subgen. n.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="diagnosis">
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<paragraph pageId="9" pageNumber="10">Diagnosis.</paragraph>
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<paragraph lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
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Species of this subgenus can be diagnosed by the synapomorphic presence of punctures on the prosternum (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Thoracic structures of Mecyclothorax and Amblytelus spp. A-B. Pronotum, dorsal view: A, M. laterorectus; B, M. mouensis. C-F. Prosternum, ventral view: C, M. fleutiauxi; D, M. curtus; E, M. blackburni; F, M. moorei. G-J. Elytral apices, dorsal view: G, M. fleutiauxi; H, M. laterorectus; I, M. lissus; J, A. curtus. K-L. Elytral lateral margin showing eighth stria, lateral view: K, M. ambiguus; L, M. blackburni. Abbreviations include: aag, anteapical groove; acc, accessory elytral dorsal setae in interval 5; as, apical elytral seta; ca, carina laterad stria 7; ls, lateral elytral setae; ri, ridge mesad stria 8; sa, subapical elytral seta; sas, subapical sinuation of elytron; scs, supracarinal elytral setae; 8, medial portion of stria 8." figureDoi="10.3897/dez.65.21000.figure2" httpUri="https://binary.pensoft.net/fig/179958" pageId="9" pageNumber="10">2D-F</figureCitation>
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), and the presence of a distinctly punctate prosternal anteapical groove, or the presence of punctures so deep and large that they obscure any groove (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Thoracic structures of Mecyclothorax and Amblytelus spp. A-B. Pronotum, dorsal view: A, M. laterorectus; B, M. mouensis. C-F. Prosternum, ventral view: C, M. fleutiauxi; D, M. curtus; E, M. blackburni; F, M. moorei. G-J. Elytral apices, dorsal view: G, M. fleutiauxi; H, M. laterorectus; I, M. lissus; J, A. curtus. K-L. Elytral lateral margin showing eighth stria, lateral view: K, M. ambiguus; L, M. blackburni. Abbreviations include: aag, anteapical groove; acc, accessory elytral dorsal setae in interval 5; as, apical elytral seta; ca, carina laterad stria 7; ls, lateral elytral setae; ri, ridge mesad stria 8; sa, subapical elytral seta; sas, subapical sinuation of elytron; scs, supracarinal elytral setae; 8, medial portion of stria 8." figureDoi="10.3897/dez.65.21000.figure2" httpUri="https://binary.pensoft.net/fig/179958" pageId="9" pageNumber="10">2F</figureCitation>
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). The pronotal front angles narrow and are not protruded (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F9" captionText="Figure 8. Mecyclothorax spp., dorsal view, with subgeneric assignments: A, M. (Eucyclothorax, subgen. n.) Mecyclothorax blackburni (type species); B, M. (Qecyclothorax, subgen. n.) Mecyclothorax storeyi (type species); C, M. (Meonochilus, status n.) Meonochilus amplipennis (type species); D, M. (Mecyclothorax) montivagus (type species); E, M. (Mecyclothorax) lissus." figureDoi="10.3897/dez.65.21000.figure8" httpUri="https://binary.pensoft.net/fig/179964" pageId="9" pageNumber="10">8A</figureCitation>
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) in all species except
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<taxonomicName genus="M." lsidName="M. curtus" pageId="9" pageNumber="10" rank="species" species="curtus">
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<emphasis italics="true" pageId="9" pageNumber="10">M. curtus</emphasis>
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</taxonomicName>
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and
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<taxonomicName genus="M." lsidName="M." pageId="9" pageNumber="10" rank="genus">
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<emphasis italics="true" pageId="9" pageNumber="10">M.</emphasis>
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</taxonomicName>
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nsp. D, in which these angles are little protruded and more rounded (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Thoracic structures of Mecyclothorax and Amblytelus spp. A-B. Pronotum, dorsal view: A, M. laterorectus; B, M. mouensis. C-F. Prosternum, ventral view: C, M. fleutiauxi; D, M. curtus; E, M. blackburni; F, M. moorei. G-J. Elytral apices, dorsal view: G, M. fleutiauxi; H, M. laterorectus; I, M. lissus; J, A. curtus. K-L. Elytral lateral margin showing eighth stria, lateral view: K, M. ambiguus; L, M. blackburni. Abbreviations include: aag, anteapical groove; acc, accessory elytral dorsal setae in interval 5; as, apical elytral seta; ca, carina laterad stria 7; ls, lateral elytral setae; ri, ridge mesad stria 8; sa, subapical elytral seta; sas, subapical sinuation of elytron; scs, supracarinal elytral setae; 8, medial portion of stria 8." figureDoi="10.3897/dez.65.21000.figure2" httpUri="https://binary.pensoft.net/fig/179958" pageId="9" pageNumber="10">2D</figureCitation>
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). The paraglossae are generally
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<pageBreakToken pageId="10" pageNumber="11" start="start">elongate</pageBreakToken>
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(Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Head and mouthpart structures of Mecyclothorax spp. A-C. Head capsule, dorsal view: A, M. goweri; B, M. fleutiauxi; C, M. laterorectus. D-I, Mentum, labial palps and ligula: D, M. peryphoides, ventral view; E, M. storeyi, ventral view; F, M. amplipennis, dorsal view; G, M. montivagus, dorsal view; H, M. kanak, ventral view; I, M. fleutiauxi, ventral view. Abbreviations include: aso, anterior supraorbital seta; pso, posterior supraorbital seta." figureDoi="10.3897/dez.65.21000.figure1" httpUri="https://binary.pensoft.net/fig/179957" pageId="10" pageNumber="11">1D</figureCitation>
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), extended twice as far from the anterior margin of the ligula than the distance from their base to the ligular margin; this is reversed in
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<taxonomicName genus="M." lsidName="M. punctatus" pageId="10" pageNumber="11" rank="species" species="punctatus">
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<emphasis italics="true" pageId="10" pageNumber="11">M. punctatus</emphasis>
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</taxonomicName>
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which has shorter paraglossae. The elytral striae are distinctly punctate (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F9" captionText="Figure 8. Mecyclothorax spp., dorsal view, with subgeneric assignments: A, M. (Eucyclothorax, subgen. n.) Mecyclothorax blackburni (type species); B, M. (Qecyclothorax, subgen. n.) Mecyclothorax storeyi (type species); C, M. (Meonochilus, status n.) Meonochilus amplipennis (type species); D, M. (Mecyclothorax) montivagus (type species); E, M. (Mecyclothorax) lissus." figureDoi="10.3897/dez.65.21000.figure8" httpUri="https://binary.pensoft.net/fig/179964" pageId="10" pageNumber="11">8A</figureCitation>
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), distinguishing
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<taxonomicName authority="Liebherr, 2018" authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Eucyclothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eucyclothorax" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="11">Eucyclothorax</emphasis>
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</taxonomicName>
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spp. from species in
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<taxonomicName authority="Liebherr, 2018" authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Qecyclothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Qecyclothorax" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="11">Qecyclothorax</emphasis>
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</taxonomicName>
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, though not from other Australian members of subgenus
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<taxonomicName authority="Mecyclothorax" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax (Mecyclothorax)" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subGenus" subGenus="Mecyclothorax">Mecyclothorax Mecyclothorax</taxonomicName>
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such as
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<taxonomicName genus="M." lsidName="M. ambiguus" pageId="10" pageNumber="11" rank="species" species="ambiguus">
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<emphasis italics="true" pageId="10" pageNumber="11">M. ambiguus</emphasis>
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</taxonomicName>
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or
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<taxonomicName genus="M." lsidName="M. punctipennis" pageId="10" pageNumber="11" rank="species" species="punctipennis">
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<emphasis italics="true" pageId="10" pageNumber="11">M. punctipennis</emphasis>
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</taxonomicName>
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. Also, the eighth elytral stria is punctate at midlength in most species (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Thoracic structures of Mecyclothorax and Amblytelus spp. A-B. Pronotum, dorsal view: A, M. laterorectus; B, M. mouensis. C-F. Prosternum, ventral view: C, M. fleutiauxi; D, M. curtus; E, M. blackburni; F, M. moorei. G-J. Elytral apices, dorsal view: G, M. fleutiauxi; H, M. laterorectus; I, M. lissus; J, A. curtus. K-L. Elytral lateral margin showing eighth stria, lateral view: K, M. ambiguus; L, M. blackburni. Abbreviations include: aag, anteapical groove; acc, accessory elytral dorsal setae in interval 5; as, apical elytral seta; ca, carina laterad stria 7; ls, lateral elytral setae; ri, ridge mesad stria 8; sa, subapical elytral seta; sas, subapical sinuation of elytron; scs, supracarinal elytral setae; 8, medial portion of stria 8." figureDoi="10.3897/dez.65.21000.figure2" httpUri="https://binary.pensoft.net/fig/179958" pageId="10" pageNumber="11">2L</figureCitation>
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)-save
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<taxonomicName genus="M." lsidName="M. moorei" pageId="10" pageNumber="11" rank="species" species="moorei">
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<emphasis italics="true" pageId="10" pageNumber="11">M. moorei</emphasis>
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</taxonomicName>
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and
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<taxonomicName genus="M." lsidName="M. punctatus" pageId="10" pageNumber="11" rank="species" species="punctatus">
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<emphasis italics="true" pageId="10" pageNumber="11">M. punctatus</emphasis>
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</taxonomicName>
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-though this punctation is also seen in lesser development in some species of subgenus
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<taxonomicName authority="Mecyclothorax" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax (Mecyclothorax)" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subGenus" subGenus="Mecyclothorax">Mecyclothorax Mecyclothorax</taxonomicName>
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such as
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<taxonomicName genus="M." lsidName="M. punctipennis" pageId="10" pageNumber="11" rank="species" species="punctipennis">
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<emphasis italics="true" pageId="10" pageNumber="11">M. punctipennis</emphasis>
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</taxonomicName>
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. The surface of the basal abdominal ventrites is generally glossy without microsculpture, a synapomorphy, though this is reversed to a microsculptured surface in
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<taxonomicName genus="M." lsidName="M. lophoides" pageId="10" pageNumber="11" rank="species" species="lophoides">
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<emphasis italics="true" pageId="10" pageNumber="11">M. lophoides</emphasis>
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</taxonomicName>
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and
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<taxonomicName genus="M." lsidName="M." pageId="10" pageNumber="11" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="11">M.</emphasis>
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</taxonomicName>
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sp. n. D. In the male genitalia the right paramere is elongate, with only 2-7 setae present on the ventral surface (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paired left (above) and right (below) parameres of Mecyclothorax spp. (ectal view): A, M. blackburni; B, M. storeyi; C, M. amplipennis; D, M. montivagus; E, M. laterobustus. As the aedeagal assembly of M. storeyi males is inverted relative to other Mecyclothorax species (Moore 1984), the configuration of the upper, or right paramere of M. storeyi should be compared to the ventral, or left parameres of the other species, and vice versa." figureDoi="10.3897/dez.65.21000.figure5" httpUri="https://binary.pensoft.net/fig/179961" pageId="10" pageNumber="11">5A</figureCitation>
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), and the internal sac bears a dorsal plate as well as a sinuous flagellum (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Male aedeagal median lobe and parameres of Mecyclothorax spp., right lateral view, internal sac everted in C-G: A, M. blackburni; B-C, M. peryphoides; D, M. moorei; E, M. lewisensis; F, M. punctipennis; G, M. montivagus. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure4" httpUri="https://binary.pensoft.net/fig/179960" pageId="10" pageNumber="11">4C</figureCitation>
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). Finally, the female reproductive tract has the spermathecal duct entering at the juncture of the common oviduct with the bursa (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Female reproductive tract, gonocoxites and associated laterotergites, Amblytelus and Mecyclothorax spp: A, A. matthewsi; B, A. brevis; C, M. blackburni; D, M. storeyi; E, M. minutus; F, M. rectangulus. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure6" httpUri="https://binary.pensoft.net/fig/179962" pageId="10" pageNumber="11">6C</figureCitation>
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), a symplesiomorphy shared with
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<taxonomicName authority="Liebherr, 2018" authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Qecyclothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Qecyclothorax" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="11">Qecyclothorax</emphasis>
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</taxonomicName>
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spp. (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Female reproductive tract, gonocoxites and associated laterotergites, Amblytelus and Mecyclothorax spp: A, A. matthewsi; B, A. brevis; C, M. blackburni; D, M. storeyi; E, M. minutus; F, M. rectangulus. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure6" httpUri="https://binary.pensoft.net/fig/179962" pageId="10" pageNumber="11">6D</figureCitation>
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), some
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<taxonomicName authorityName="Erichson" authorityYear="1842" class="Insecta" family="Carabidae" genus="Amblytelus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Amblytelus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="10" pageNumber="11">Amblytelus</emphasis>
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</taxonomicName>
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spp. (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Female reproductive tract, gonocoxites and associated laterotergites, Amblytelus and Mecyclothorax spp: A, A. matthewsi; B, A. brevis; C, M. blackburni; D, M. storeyi; E, M. minutus; F, M. rectangulus. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure6" httpUri="https://binary.pensoft.net/fig/179962" pageId="10" pageNumber="11">6A-B</figureCitation>
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),
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<taxonomicName genus="M." lsidName="M. bellorum" pageId="10" pageNumber="11" rank="species" species="bellorum">
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<emphasis italics="true" pageId="10" pageNumber="11">M. bellorum</emphasis>
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</taxonomicName>
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of subgenus
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax (Meonochilus)" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subGenus" subGenus="Meonochilus">Mecyclothorax Meonochilus</taxonomicName>
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(
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.147.1898" author="Liebherr, JK" journalOrPublisher="from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae ZooKeys" pageId="51" pageNumber="52" pagination="277 - 335" refId="B57" refString="Liebherr, JK, 2011. Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoycea glacialis, gen. n. and sp. n. from the South Island, and revision of Meonochilus Liebherr and Marris from the North Island, New Zealand (Coleoptera, Carabidae ZooKeys 147: 277 - 335, DOI: https://doi.org/10.3897/zookeys.147.1898" title="Cladistic assessment of subtribal affinities within the tribe Moriomorphini with description of Rossjoycea glacialis, gen. n. and sp. n." url="https://doi.org/10.3897/zookeys.147.1898" volume="147" year="2011">Liebherr 2011</bibRefCitation>
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, fig. 12). A
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<pageBreakToken pageId="11" pageNumber="12" start="start">helminthoid</pageBreakToken>
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sclerite is also symplesiomorphically present (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Female reproductive tract, gonocoxites and associated laterotergites, Amblytelus and Mecyclothorax spp: A, A. matthewsi; B, A. brevis; C, M. blackburni; D, M. storeyi; E, M. minutus; F, M. rectangulus. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure6" httpUri="https://binary.pensoft.net/fig/179962" pageId="11" pageNumber="12">6C</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="11" pageNumber="12" type="member species">
|
||
<paragraph pageId="11" pageNumber="12">Member species.</paragraph>
|
||
<paragraph pageId="11" pageNumber="12">
|
||
All Australian
|
||
<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Mecyclothorax</emphasis>
|
||
</taxonomicName>
|
||
species have been examined, with the following species assignable to this subgenus represented in the analysis:
|
||
<taxonomicName genus="M." lsidName="M. blackburni" pageId="11" pageNumber="12" rank="species" species="blackburni">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">M. blackburni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. curtus" pageId="11" pageNumber="12" rank="species" species="curtus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">M. curtus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. eyrensis" pageId="11" pageNumber="12" rank="species" species="eyrensis">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">M. eyrensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. lophoides" pageId="11" pageNumber="12" rank="species" species="lophoides">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">M. lophoides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. moorei" pageId="11" pageNumber="12" rank="species" species="moorei">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">M. moorei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName genus="M." lsidName="M. peryphoides" pageId="11" pageNumber="12" rank="species" species="peryphoides">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">M. peryphoides</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName genus="M." lsidName="M. punctatus" pageId="11" pageNumber="12" rank="species" species="punctatus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">M. punctatus</emphasis>
|
||
</taxonomicName>
|
||
(Table
|
||
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Species-level taxa included in cladistic analysis. Mecyclothorax subgeneric classification based upon results of cladistic analysis. Type species indicated for Mecyclothorax generic-level taxa. Mecyclothorax andersoni represented by infraspecific aedeagal forms with either a short or long median lobe apex (Liebherr 2017 b). Mecyclothorax sp. n. D treated as non-valid terminal; species to be described subsequently." httpUri="http://table.plazi.org/id/170E9ACFC66D90BF4A660AFCD485EFB2" pageId="11" pageNumber="12" tableUuid="170E9ACFC66D90BF4A660AFCD485EFB2">1</tableCitation>
|
||
;
|
||
<bibRefCitation author="Moore, BP" journalOrPublisher="Journal of the Australian Entomological Society" pageId="52" pageNumber="53" refId="B80" refString="Moore, BP, Weir, TA, Pyke, JE, 1987. Coleoptera: Adephaga: Rhysodidae and Carabidae. In: Walton DW (Ed.) Zoological Catalogue of Australia Vol 4, Australian Government Printing Service, Canberra: 17-320." title="Coleoptera: Adephaga: Rhysodidae and Carabidae. In: Walton DW (Ed.) Zoological Catalogue of Australia Vol 4, Australian Government Printing Service, Canberra: 17 - 320." year="1987">Moore et al. 1987</bibRefCitation>
|
||
,
|
||
<bibRefCitation DOI="https://doi.org/10.3853/j.0067-1975.61.2009.1519" author="Baehr, M" journalOrPublisher="Records of the Australian Museum" pageId="50" pageNumber="51" pagination="89 - 92" refId="B12" refString="Baehr, M, 2009. A new species of the genus Mecyclothorax Sharp from New South Wales (Insecta: Carabidae: Psydrinae). Records of the Australian Museum 61: 89 - 92, DOI: https://doi.org/10.3853/j.0067-1975.61.2009.1519" title="A new species of the genus Mecyclothorax Sharp from New South Wales (Insecta: Carabidae: Psydrinae)." url="https://doi.org/10.3853/j.0067-1975.61.2009.1519" volume="61" year="2009">Baehr 2009</bibRefCitation>
|
||
).
|
||
<bibRefCitation author="Baehr, M" journalOrPublisher="Spixiana" pageId="50" pageNumber="51" pagination="93 - 97" refId="B14" refString="Baehr, M, 2016. A new subspecies of Mecyclothorax punctatus (Sloane) from south-western Australia. Spixiana 39: 93 - 97" title="A new subspecies of Mecyclothorax punctatus (Sloane) from south-western Australia." volume="39" year="2016">
|
||
<normalizedToken originalValue="Baehr’s">Baehr's</normalizedToken>
|
||
(2016)
|
||
</bibRefCitation>
|
||
southwestern Australian subspecies
|
||
<taxonomicName genus="M." lsidName="M. punctatus subsp. peckorum" pageId="11" pageNumber="12" rank="subSpecies" species="punctatus" subSpecies="peckorum">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">M. punctatus peckorum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation DOI="https://doi.org/10.1111/j.1440-6055.1984.tb01935.x" author="Moore, BP" journalOrPublisher="Journal of the Australian Entomological Society" pageId="52" pageNumber="53" pagination="161 - 166" refId="B77" refString="Moore, BP, 1984. Taxonomic notes on some Australasian Mecyclothorax Sharp (Coleoptera: Carabidae: Psydrinae) and descriptions of new species. Journal of the Australian Entomological Society 23: 161 - 166, DOI: https://doi.org/10.1111/j.1440-6055.1984.tb01935.x" title="Taxonomic notes on some Australasian Mecyclothorax Sharp (Coleoptera: Carabidae: Psydrinae) and descriptions of new species." url="https://doi.org/10.1111/j.1440-6055.1984.tb01935.x" volume="23" year="1984">Moore 1984</bibRefCitation>
|
||
, fig. 12) and
|
||
<taxonomicName genus="M." lsidName="M. cordicollis" pageId="11" pageNumber="12" rank="species" species="cordicollis">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">M. cordicollis</emphasis>
|
||
</taxonomicName>
|
||
(Sloane) complement this list. An undescribed species from Queensland-sp. n. D-will be described in a subsequent publication.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="11" pageNumber="12" type="etymology">
|
||
<paragraph pageId="11" pageNumber="12">Etymology.</paragraph>
|
||
<paragraph pageId="11" pageNumber="12">
|
||
To indicate membership of species of this subgenus in
|
||
<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Mecyclothorax</emphasis>
|
||
</taxonomicName>
|
||
, and to reflect the position of this taxon as the adelphotaxon to the remainder of
|
||
<taxonomicName authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Mecyclothorax</emphasis>
|
||
</taxonomicName>
|
||
sensu lato, the initial syllable
|
||
<normalizedToken originalValue="“eu”">"eu"</normalizedToken>
|
||
-the Ancient Greek prefix
|
||
<emphasis italics="true" pageId="11" pageNumber="12">
|
||
<normalizedToken originalValue="εύ">εύ</normalizedToken>
|
||
</emphasis>
|
||
, meaning
|
||
<normalizedToken originalValue="“proper”">"proper"</normalizedToken>
|
||
or
|
||
<normalizedToken originalValue="“true”">"true"</normalizedToken>
|
||
-is combined with the first name proposed for the genus,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cyclothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Cyclothorax" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Cyclothorax</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="MacLeay, W" journalOrPublisher="Transactions of the Entomological Society of New South Wales" pageId="52" pageNumber="53" pagination="79 - 205" refId="B74" refString="MacLeay, W, 1871. Notes on a collection of insects from Gayndah. Transactions of the Entomological Society of New South Wales 2 (2): 79 - 205" title="Notes on a collection of insects from Gayndah." volume="2" year="1871">MacLeay 1871</bibRefCitation>
|
||
), resulting in the subgenus name
|
||
<taxonomicName authority="Liebherr, 2018" authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Eucyclothorax" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eucyclothorax" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="11" pageNumber="12">Eucyclothorax</emphasis>
|
||
</taxonomicName>
|
||
. Based on the shared nominative masculine terminal syllable thorax, this epithet agrees in gender with the generic epithet.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |