158 lines
16 KiB
XML
158 lines
16 KiB
XML
<document id="4DD5C29D183315E97C818E8D1F4E9C91" ID-CLB-Dataset="31534" ID-DOI="http://dx.doi.org/10.3897/zookeys.715.20497" ID-GBIF-Dataset="138d6982-b157-4301-a3a0-72029f9044ba" ID-PMC="PMC5740420" ID-Pensoft-Pub="1313-2970-715-69" ID-PubMed="29302233" ID-ZooBank="763EF053AFAA41119CA96060F9E09E70" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-715-69" ModsDocOrigin="ZooKeys 715" ModsDocTitle="Revision of the genus Ptomaphagus Hellwig from eastern Asia (Coleoptera, Leiodidae, Cholevinae)" checkinTime="1510736981334" checkinUser="pensoft" docAuthor="Wang, Cheng-Bin, Perreau, Michel, Ruzicka, Jan & Nishikawa, Masaaki" docDate="2017" docId="926E6FD06F0423598DE9961C4A54490A" docLanguage="en" docName="ZooKeys 715: 69-92" docOrigin="ZooKeys 715" docSource="http://dx.doi.org/10.3897/zookeys.715.20497" docTitle="Ptomaphagus masumotoi Nishikawa 2011" docType="treatment" docVersion="6" lastPageNumber="79" masterDocId="7A4E6F14FFE5FFD08743FFBA6574A77E" masterDocTitle="Revision of the genus Ptomaphagus Hellwig from eastern Asia (Coleoptera, Leiodidae, Cholevinae)" masterLastPageNumber="92" masterPageNumber="69" pageNumber="75" updateTime="1701375221787" updateUser="ExternalLinkService">
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<mods:title id="46F500651B8CC43997EE148458C332AA">Revision of the genus Ptomaphagus Hellwig from eastern Asia (Coleoptera, Leiodidae, Cholevinae)</mods:title>
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<mods:namePart id="2E5C5F748FF6B76E482DBD1CCD6EFCCD">Wang, Cheng-Bin</mods:namePart>
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<mods:namePart id="4146A5E0FEF1F7F4E64F1F9106FE95AE">Perreau, Michel</mods:namePart>
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<mods:namePart id="9C83D77E060546387192DA094DFAFA90">Ruzicka, Jan</mods:namePart>
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<mods:namePart id="946E42601395D7139387EA68832D4796">Nishikawa, Masaaki</mods:namePart>
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<mods:title id="8F4D532D79C59D67860C35BE3727B982">ZooKeys</mods:title>
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<mods:date id="EC934B81C2481298C659F5692DB9EE9F">2017</mods:date>
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<mods:number id="14A5FA6F3E6B0CC77E47B0BD4ED3B87E">715</mods:number>
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<mods:url id="61833F9F99DBF80E98FC8A768870BBA0">http://dx.doi.org/10.3897/zookeys.715.20497</mods:url>
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<mods:classification id="4603395C8EB102ED6A8490A7D9E0A9C7">journal article</mods:classification>
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<mods:identifier id="E5EB78A32A16F10EA665C108B9ABD3F2" type="DOI">http://dx.doi.org/10.3897/zookeys.715.20497</mods:identifier>
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<treatment id="926E6FD06F0423598DE9961C4A54490A" ID-GBIF-Taxon="137306767" LSID="urn:lsid:plazi:treatment:926E6FD06F0423598DE9961C4A54490A" httpUri="http://treatment.plazi.org/id/926E6FD06F0423598DE9961C4A54490A" lastPageId="10" lastPageNumber="79" pageId="6" pageNumber="75">
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<subSubSection id="D1BE7AB31E15DA5172F0C323CF9E0E1D" pageId="6" pageNumber="75" type="nomenclature">
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<paragraph id="3A1344888C6F3036E8FAD0925C7EF67C" pageId="6" pageNumber="75">
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<taxonomicName id="7B13569AA85342D76D3294CCCF2E42D8" ID-CoL="4Q9F8" authority="Nishikawa, 2011" authorityName="Nishikawa" authorityYear="2011" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus masumotoi" order="Coleoptera" pageId="6" pageNumber="75" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="masumotoi">Ptomaphagus (s. str.) masumotoi Nishikawa, 2011</taxonomicName>
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Figs 1
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<normalizedToken id="35147973F66A8D81FE95DCAB28CCCD34" originalValue="C–D">C-D</normalizedToken>
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; 4
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<normalizedToken id="08F686AD1E8162A34123ACA876D5CC90" originalValue="A–J">A-J</normalizedToken>
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; 5
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<normalizedToken id="2520B6CC2DFD7D37E32BF9FB8768A540" originalValue="A–F">A-F</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection id="0BD1E36A7F3F006F3EB351FA6D907E0B" pageId="7" pageNumber="76" type="reference_group">
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<paragraph id="AFAFC8C9CE375FE41824EB944436ED56" pageId="7" pageNumber="76">
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<taxonomicName id="62ACABC571843F37466CA82D7446859A" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus masumotoi" order="Coleoptera" pageId="7" pageNumber="76" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="masumotoi">
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<pageBreakToken id="829A41857CF9FF092CF3A99AA705399A" pageId="7" pageNumber="76" start="start">Ptomaphagus</pageBreakToken>
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(s. str.) masumotoi
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</taxonomicName>
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Nishikawa, 2011: 97 (
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<taxonomicName id="357584BFE2E5621B3580C900DCD1A1C5" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus (Ptomaphagus)" order="Coleoptera" pageId="7" pageNumber="76" phylum="Arthropoda" rank="subGenus" subGenus="Ptomaphagus">Ptomaphagus (Ptomaphagus)</taxonomicName>
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; type locality: NW Thailand, Chiang Mai, Doi Inthanon [ca.
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<geoCoordinate id="74CA50B0292C28DCF68E446E84CF2F26" direction="north" orientation="latitude" precision="925" value="18.7">18°42'N</geoCoordinate>
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,
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<geoCoordinate id="4274A8A15E2AA0E9B61C906978E5D1CF" direction="east" orientation="longitude" precision="925" value="98.98333">098°59'E</geoCoordinate>
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], 1750 m; NSMT).
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</paragraph>
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</subSubSection>
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<subSubSection id="127208E44ED58CA842A9F23CC3DE419F" pageId="7" pageNumber="76" type="materials_examined">
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<paragraph id="85460612A67490BF97EDF612CF6EE992" pageId="7" pageNumber="76">Material examined.</paragraph>
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<paragraph id="91AA958C3AF3D9BEBFF651ED400069C8" pageId="7" pageNumber="76">
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Type material. Holotype: ♀, Doi Inthanon / 1750 m, Chiang / Mai, Thailand / 9-XI-1995 / K. MASUMOTO leg. // Holotype /
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<taxonomicName id="2CFA126BCF0355ED6413AA7E444D1F5D" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="7" pageNumber="76" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
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(
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<taxonomicName id="217ABCE16390BF239539D332C0C313FC" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus" order="Coleoptera" pageId="7" pageNumber="76" phylum="Arthropoda" rank="genus">Ptomaphagus</taxonomicName>
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) /
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<taxonomicName id="CE97AE2C7C988E7F86F615C5AE8F33E4" class="Insecta" family="Leiodidae" genus="Ptomaphagina" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Ptomaphagina masumotoi" order="Coleoptera" pageId="7" pageNumber="76" phylum="Arthropoda" rank="species" species="masumotoi">masumotoi</taxonomicName>
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M. Nishikawa, / 2011 / Design. M. Nishikawa, 2011 / # MNC 146Ch2P ♀ (NSMT). Paratype: 1♂, same data as holotype except: # MNC 147Ch2P ♂ (NSMT).
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</paragraph>
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</subSubSection>
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<subSubSection id="BAEB0FEA84A413A14CBC94A798EF31A8" lastPageId="10" lastPageNumber="79" pageId="7" pageNumber="76" type="redescription">
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<paragraph id="3FA8CEF7BF1A2A987992BDFCCC62B508" pageId="7" pageNumber="76">Redescription.</paragraph>
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<paragraph id="77FC0B47D002AAF8CB31D909419C6487" pageId="7" pageNumber="76">
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Male. EBL: 3.4 mm. Length of different body parts: HL: AL: PL: ELL = 0.6: -: 0.8: 2.0 mm; width: HW: EW: PW: ELW = 0.8: 0.1: 1.3: 1.5 mm. Proportion of antennomeres from base to tip in
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<normalizedToken id="1C9DC0C29DEEA1342E1B2FD8BB17EE35" originalValue="μm">μm</normalizedToken>
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(length
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<normalizedToken id="B7EB2D74C3F09A09E299363A9F2FD9BA" originalValue="×">x</normalizedToken>
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width): 121
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<normalizedToken id="8535F59E7590D166E3CA8AE312BE9EF0" originalValue="×">x</normalizedToken>
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57, 105
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<normalizedToken id="35EC9F2C7A2F68B6866FD446F8AD8884" originalValue="×">x</normalizedToken>
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62, 71
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<normalizedToken id="C6AA9F049893EA5320D65374E3EB5DA0" originalValue="×">x</normalizedToken>
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65, 48
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<normalizedToken id="AE6E04424CF4E767B89ED9F46AB31A02" originalValue="×">x</normalizedToken>
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70, 53
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<normalizedToken id="9228B1F374AE1784EB5028890DDF2A50" originalValue="×">x</normalizedToken>
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82, 35
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<normalizedToken id="39A3AC347C80CEF56ECDDC00E151A1A2" originalValue="×">x</normalizedToken>
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92, 94
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<normalizedToken id="C2B33DD4DE5DE0B8C68F4CD962741D39" originalValue="×">x</normalizedToken>
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111, 28
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<normalizedToken id="4FC7FB5FFBD841370ABA06F729CE4B6D" originalValue="×">x</normalizedToken>
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101, 83
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<normalizedToken id="AEC1E8A2A21CFEAEDE0BBE3E649F4AD2" originalValue="×">x</normalizedToken>
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112 (last two antennomeres missing).
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</paragraph>
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<paragraph id="7DC2429817033936CBD86D8784B97E8B" pageId="7" pageNumber="76">Habitus (Fig. 1C) elongated oval, regularly convex and sublustrous. Well pigmented: mostly brown; mouthparts, basal three antennomeres and apical half of ultimate antennomere, protarsi, and apex of meso- and metatarsi more or less paler. Dorsum continually clothed with fine, recumbent, yellowish pubescence. Insertions of pubescence on dorsal surfaces of pronotum, elytra and femora aligned along transverse striolations; interspace between two striolations glabrous.</paragraph>
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<paragraph id="A74DE4AC997DAA0E868DF9C9BAC333EE" lastPageId="8" lastPageNumber="77" pageId="7" pageNumber="76">
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Head transverse, HW/HL = 1.5. Clypeofrontal suture absent. Clypeus with anterior margin gently rounded. Compound eyes well developed, EW/HW = 0.1. Antennae
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<pageBreakToken id="E5B515BB44F676E4B271325537C9110C" pageId="8" pageNumber="77" start="start">(</pageBreakToken>
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Fig. 4A) slender; antennomere III shorter than II; VI with length/width = 0.4; X and XI of holotype missing.
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</paragraph>
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<paragraph id="947E345FE41838CCA5F07F590603100C" pageId="8" pageNumber="77">Pronotum (Fig. 4B) transverse, widest around hind angles, PW/PL = 1.6. Sides gently arched, simply narrowing from posterior to anterior; hind angles not projected backwards and bluntly rounded. Posterior margin widely protruded in middle part, emarginate near hind angles.</paragraph>
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<paragraph id="B789E7EA93483F4AAEC21D2CCB150959" pageId="9" pageNumber="78">
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<pageBreakToken id="E464BC39C1CA97C9ABBE84ACB8607753" pageId="9" pageNumber="78" start="start">Elytra</pageBreakToken>
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oval, widest at about basal 2/7, ELL/EW = 1.4. Sides weakly arched, gradually narrowing from widest part to apices; apices (Fig. 4G) narrowly rounded. Sutural striae present. Metathoracic wings fully developed.
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</paragraph>
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<paragraph id="53E68FF30C7E1AC1058400A67365C22D" pageId="9" pageNumber="78">Prolegs robust, with basal three protarsomeres (Fig. 4C) strongly expanded: TW/BTW = 1.0. Protibiae (Fig. 4E) strongly expanded towards apex. Profemora broad. Mesotibiae gently arcuate, mesotarsi simply linear. Metatibiae straight, but relatively short and thick.</paragraph>
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<paragraph id="D9225A467F34539462FA951E23428908" pageId="10" pageNumber="79">
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<pageBreakToken id="BAB10D0F0987581DB69A9D9B10409D3A" pageId="10" pageNumber="79" start="start">Abdominal</pageBreakToken>
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ventrite VIII (Fig. 4I) emarginate at posterior edge. Spiculum gastrale (Fig. 4J) of genital segment with about 1/5 of length protruding beyond anterior edge of epipleurite IX.
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</paragraph>
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<caption id="AB479877345FE4F8819BC75E8270A4CB" pageId="10" pageNumber="79">
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<paragraph id="5EA6550C64F4537769E94C8ECC67992E" pageId="10" pageNumber="79">
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Figure 4.
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<taxonomicName id="77E159453A743E507FEE955AAF059720" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus masumotoi" order="Coleoptera" pageId="10" pageNumber="79" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="masumotoi">Ptomaphagus (s. str.) masumotoi</taxonomicName>
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Nishikawa, 2011 (♂: paratype; ♀: holotype). A antenna ♂ (dorsal view) B pronotum ♂ (dorsal view) C protarsus ♂ (dorsal view) D protarsus ♀ (dorsal view) E protibia and profemur ♂ (ventral view) F protibia and profemur ♀ (ventral view) G elytral apex ♂ (dorsoapical view) H elytral apex ♀ (dorsoapical view) I ventrite VIII ♂ (ventral view) J genital segment ♂ (ventral view). Scale bars: 0.1 mm.
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</paragraph>
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</caption>
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<paragraph id="92E17E54E8FF3CA9478516EFBC633848" pageId="10" pageNumber="79">Aedeagus (Fig. 5A) slender, with median lobe gradually narrowing towards an oblong apex and terminated to a shortly rounded knob in dorsal view; opening of genital orifice situated on dorsal surface, deeply cut inwards on preapical left margin of median lobe. Ventral surface of the apex of the median lobe (Fig. 5C) inserted with five ventrally oriented setae on the left side and six ventrally oriented setae on the right side; parameres narrow, reaching about apical 1/7 of median lobe, each apex (Fig. 5D) with two long lateral setae and one similar apical seta. In lateral view (Fig. 5B), median lobe regularly bent ventrally but almost straight in apical half, and gradually tapering to a acuminate apex. Endophallus with stylus quite slender, a transverse nodule in middle region, a cheliform complex just below base of stylus, and a circular complex in the basal region.</paragraph>
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<paragraph id="96FC10A90F97D426325BAEF7869E3339" pageId="10" pageNumber="79">Female. Similar to male in general appearance (Fig. 1D), including elytral apices (Fig. 4H), but distinguished by the following characteristics: protarsi (Fig. 4D) simply linear; protibiae (Fig. 4F) slightly narrower; abdominal ventrite VIII (Fig. 5E) slightly protruded at median of posterior edge; genital segment and ovipositor as shown in Fig. 5F; spermatheca (Fig. 5F) sinuous or coiled in distal part, not coiled in proximal part.</paragraph>
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<caption id="16E00A3D32F213AA45219BD0F2E0E776" pageId="10" pageNumber="79">
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<paragraph id="29AFE9C3A7F4EF8997BF759954B94516" pageId="10" pageNumber="79">
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Figure 5.
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<taxonomicName id="EFBC8C633F3FAC2FACD239D5AB21A201" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus masumotoi" order="Coleoptera" pageId="10" pageNumber="79" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="masumotoi">Ptomaphagus (s. str.) masumotoi</taxonomicName>
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Nishikawa, 2011 (♂: paratype; ♀: holotype). A aedeagus (dorsal view) B aedeagus (lateral view) C aedeagal apex (ventral view) D paramere apex (lateral view) E ventrite VIII ♀ (ventral view) F spermatheca, genital segment and ovipositor (ventral view). Scale bars: 0.1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="EABE0192BA225D9966805464B6F3829F" pageId="10" pageNumber="79" type="distribution">
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<paragraph id="0FDC28BF464942CF384C690AC2F32454" pageId="10" pageNumber="79">Distribution.</paragraph>
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<paragraph id="F0A6212AEE492E5FBB717EB5B7FE6BC5" pageId="10" pageNumber="79">Thailand.</paragraph>
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</subSubSection>
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</treatment>
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</document> |