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<mods:title id="6CB17F419D4FDB1503CC518963433067">Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species</mods:title>
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4) 
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Zhang &amp; Toda
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(
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Head parts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, right eye; 2, frontal view.Abbreviation: pc orb s, proclinate orbital seta. Scale bars: 0.1 mm." captionText-1="FIGURE 3. Right antennae (inner-lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Abbreviations: ar, arista; flgm 1, first flagellomere; ped, pedicel. Scale bars: 0.1 mm." captionText-2="FIGURE 4. Head parts (caudal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Abbreviations: dl a, dorsolateral tentorial apodeme; dm a, dorsomedial tentorial apodeme; i ocp s, inner occipital seta; o vt s, outer vertical seta; spc s, supracervical seta. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm." captionText-3="FIGURE 5. Mouthparts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, caudal view. Abbreviations: a s, anterior sensilla; cib, cibarium; clyp, clypeus; m s, medial sensilla; p s, posterior sensilla; plp, palpus; premnt, prementum; s c, sensilla campaniformia. Scale bars: 0.1 mm." captionText-4="FIGURE 6. Thoraxes (dorsal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E).Abbreviations: a dc s, anterior dorsocentral setae; pprn s, postpronotal seta; presct acr s, prescutellar acrostichal setae; trn sut, transverse suture. Scale bars: 0.5 mm." captionText-5="FIGURE 7. Thoraxes (lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Scale bars: 0.1 mm." captionText-6="FIGURE 8. Right wings of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, anterodistal portion (scale bars: 0.1 mm); 2, the whole (scale bars: 0.5 mm). Abbreviations: dm-m, discal-medial crossvein; M4, 4th branch of media." captionText-7="FIGURE 9. Legs of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, fore tibia (distalmost portion) and tarsus; 2, mid tibia (distal portion) and tarsus; 3, hind tibia (distal portion) and tarsus. Scale bars: 0.1 mm." captionText-8="FIGURE 10. Abdominal sternites of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, male; 2, female.Abbreviation: s, sternite. Scale bars: 0.1 mm." captionText-9="FIGURE 12. Female terminalia of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, ventral view. Abbreviations: epig, epigynium; epiprct, epiproct; hypg v, hypogynial valve; hyprct, hypoproct. Scale bars: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7906001" figureDoi-1="http://doi.org/10.5281/zenodo.7906005" figureDoi-2="http://doi.org/10.5281/zenodo.7906007" figureDoi-3="http://doi.org/10.5281/zenodo.7906011" figureDoi-4="http://doi.org/10.5281/zenodo.7906013" figureDoi-5="http://doi.org/10.5281/zenodo.7906015" figureDoi-6="http://doi.org/10.5281/zenodo.7906019" figureDoi-7="http://doi.org/10.5281/zenodo.7906021" figureDoi-8="http://doi.org/10.5281/zenodo.7906023" figureDoi-9="http://doi.org/10.5281/zenodo.7906029" httpUri-0="https://zenodo.org/record/7906001/files/figure.png" httpUri-1="https://zenodo.org/record/7906005/files/figure.png" httpUri-2="https://zenodo.org/record/7906007/files/figure.png" httpUri-3="https://zenodo.org/record/7906011/files/figure.png" httpUri-4="https://zenodo.org/record/7906013/files/figure.png" httpUri-5="https://zenodo.org/record/7906015/files/figure.png" httpUri-6="https://zenodo.org/record/7906019/files/figure.png" httpUri-7="https://zenodo.org/record/7906021/files/figure.png" httpUri-8="https://zenodo.org/record/7906023/files/figure.png" httpUri-9="https://zenodo.org/record/7906029/files/figure.png" pageId="25" pageNumber="226">Figs 2–12D</figureCitation>
, 
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<taxonomicName id="4C6F4D4CFFCFFFF9A5C6FC6B827FE6D0" ID-CoL="5ZMRT" authority=": Sultana et al., 2006: 694" authorityName="Sultana" authorityPageNumber="694" authorityYear="2006" box="[151,769,980,1005]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" isUncertain="true" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="226" phylum="Arthropoda" rank="species" species="undefined-2">
<emphasis id="B91BEADDFFCFFFF9A5C6FC6B8051E6D0" box="[151,303,980,1004]" italics="true" pageId="25" pageNumber="226">Colocasiomyia</emphasis>
sp.2 
<treatmentCitation id="0ACE10DEFFCFFFF9A434FC6B827FE6D0" author="Sultana, F. &amp; Hu, Y. G. &amp; Toda, M. J. &amp; Takenaka, K. &amp; Yafuso, M." box="[357,769,980,1004]" page="694" pageId="25" pageNumber="226" year="2006">
aff. 
<emphasis id="B91BEADDFFCFFFF9A4DFFC6B8087E6D0" box="[398,505,980,1004]" italics="true" pageId="25" pageNumber="226">nepalensis</emphasis>
: 
<bibRefCitation id="EFFE4B3EFFCFFFF9A757FC6B827FE6D0" author="Sultana, F. &amp; Hu, Y. G. &amp; Toda, M. J. &amp; Takenaka, K. &amp; Yafuso, M." box="[518,769,980,1004]" pageId="25" pageNumber="226" pagination="684 - 702" refId="ref21931" refString="Sultana, F., Hu, Y. G., Toda, M. J., Takenaka, K. &amp; Yafuso, M. (2006) Phylogeny and classification of Colocasiomyia (Diptera, Drosophilidae), and its evolution of pollination mutualism with aroid plants. Systematic Entomology, 31, 684 - 702. https: // doi. org / 10.1111 / j. 1365 - 3113.2006.00344. x" type="journal article" year="2006">
Sultana 
<emphasis id="B91BEADDFFCFFFF9A709FC6A83F2E6D0" box="[600,652,980,1004]" italics="true" pageId="25" pageNumber="226">et al.</emphasis>
, 2006: 694
</bibRefCitation>
</treatmentCitation>
</taxonomicName>
; 
<treatmentCitation id="0ACE10DEFFCFFFF9A65AFC6B857CE6D0" author="Fartyal, R. S. &amp; Gao, J. J. &amp; Toda, M. J. &amp; Hu, Y. G. &amp; Takano, K. T. &amp; Suwito, A. &amp; Katoh, T. &amp; Takigahira, T. &amp; Yin, J. T." box="[779,1026,980,1005]" page="768" pageId="25" pageNumber="226" year="2013">
<bibRefCitation id="EFFE4B3EFFCFFFF9A65AFC6B857CE6D0" author="Fartyal, R. S. &amp; Gao, J. J. &amp; Toda, M. J. &amp; Hu, Y. G. &amp; Takano, K. T. &amp; Suwito, A. &amp; Katoh, T. &amp; Takigahira, T. &amp; Yin, J. T." box="[779,1026,980,1005]" pageId="25" pageNumber="226" pagination="763 - 782" refId="ref20189" refString="Fartyal, R. S., Gao, J. J., Toda, M. J., Hu, Y. G., Takano, K. T., Suwito, A., Katoh, T., Takigahira, T. &amp; Yin, J. T. (2013) Colocasiomyia (Diptera: Drosophilidae) revised phylogenetically, with a new species group having peculiar lifecycles on monsteroid (Araceae) host plants. Systematic Entomology, 38, 763 - 782. https: // doi. org / 10.1111 / syen. 12027" type="journal article" year="2013">
Fartyal 
<emphasis id="B91BEADDFFCFFFF9A608FC6A82F2E6D0" box="[857,908,980,1004]" italics="true" pageId="25" pageNumber="226">et al.</emphasis>
, 2013: 768
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3756544FFCFFFF9A5C6FBB18430E185" pageId="25" pageNumber="226" type="diagnosis">
<paragraph id="8BD036CFFFCFFFF9A5C6FBB18430E185" blockId="25.[151,1437,1038,1931]" pageId="25" pageNumber="226">
<emphasis id="B91BEADDFFCFFFF9A5C6FBB1806CE114" bold="true" box="[151,274,1038,1064]" pageId="25" pageNumber="226">Diagnosis.</emphasis>
Antennal arista approximately 1.8 times as long as first flagellomere (
<figureCitation id="13542A4AFFCFFFF9A15AFBB0851DE115" box="[1035,1123,1039,1065]" captionStart="FIGURE 3" captionStartId="5.[151,250,1430,1454]" captionTargetBox="[151,1435,181,1406]" captionTargetId="figure-20@5.[151,1436,181,1406]" captionTargetPageId="5" captionText="FIGURE 3. Right antennae (inner-lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Abbreviations: ar, arista; flgm 1, first flagellomere; ped, pedicel. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906005" httpUri="https://zenodo.org/record/7906005/files/figure.png" pageId="25" pageNumber="226">Fig. 3D</figureCitation>
). Distance between antennal scapes as wide as scape diameter (
<figureCitation id="13542A4AFFCFFFF9A743FB8C830EE16F" box="[530,624,1075,1107]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1963]" captionTargetBox="[151,1433,634,1915]" captionTargetId="figure-203@4.[151,1436,633,1915]" captionTargetPageId="4" captionText="FIGURE 2. Head parts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, right eye; 2, frontal view.Abbreviation: pc orb s, proclinate orbital seta. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906001" httpUri="https://zenodo.org/record/7906001/files/figure.png" pageId="25" pageNumber="226">
Fig. 2D 
<subScript id="17EB348AFFCFFFF9A739FBFA830EE16F" attach="left" box="[616,624,1093,1107]" fontSize="6" pageId="25" pageNumber="226">2</subScript>
</figureCitation>
). Acrostichal setulae in 6 rows (
<figureCitation id="13542A4AFFCFFFF9A685FB8C8550E171" box="[980,1070,1075,1101]" captionStart="FIGURE 6" captionStartId="8.[151,250,1921,1945]" captionTargetBox="[151,1436,885,1898]" captionTargetId="figure-310@8.[151,1436,885,1898]" captionTargetPageId="8" captionText="FIGURE 6. Thoraxes (dorsal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E).Abbreviations: a dc s, anterior dorsocentral setae; pprn s, postpronotal seta; presct acr s, prescutellar acrostichal setae; trn sut, transverse suture. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7906013" httpUri="https://zenodo.org/record/7906013/files/figure.png" pageId="25" pageNumber="226">Fig. 6D</figureCitation>
). All legs entirely grayish yellow to yellow except for grayish brown tarsomere Vs (
<figureCitation id="13542A4AFFCFFFF9A79CFBE88248E14D" box="[717,822,1111,1137]" captionStart="FIGURE 14" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[222,1364,181,1919]" captionTargetId="figure-20@19.[222,1364,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Habitus (lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, male; 2, female. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906035" httpUri="https://zenodo.org/record/7906035/files/figure.png" pageId="25" pageNumber="226">Fig. 14D</figureCitation>
). Cercus slightly projected at apicoventral corner (Fig. 11-2D). Long, tube-like, membranous aedeagus ornamented with relatively sparse and weak spinules (Fig. 11-2D). Hypogynial valves with dorsal flaps fused into small, ligulate plate at distal end of basal portion (
<figureCitation id="13542A4AFFCFFFF9A188FB20843CE185" box="[1241,1346,1183,1209]" captionStart="FIGURE 12" captionStartId="16.[151,250,1932,1956]" captionTargetBox="[169,1419,420,1909]" captionTargetId="figure-108@16.[169,1419,417,1909]" captionTargetPageId="16" captionText="FIGURE 12. Female terminalia of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, ventral view. Abbreviations: epig, epigynium; epiprct, epiproct; hypg v, hypogynial valve; hyprct, hypoproct. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906029" httpUri="https://zenodo.org/record/7906029/files/figure.png" pageId="25" pageNumber="226">Fig. 12D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3756544FFCFFFF9A596FB7D8348E221" pageId="25" pageNumber="226" type="description">
<paragraph id="8BD036CFFFCFFFF9A596FB7D8045E051" blockId="25.[151,1437,1038,1931]" pageId="25" pageNumber="226">
<emphasis id="B91BEADDFFCFFFF9A596FB7D802FE1E0" bold="true" box="[199,337,1218,1244]" pageId="25" pageNumber="226">Description</emphasis>
(♁ and 
<specimenCount id="9D69FD46FFCFFFF9A4E4FB7C80B7E1E1" box="[437,457,1219,1245]" pageId="25" pageNumber="226" type="female">♀</specimenCount>
). Head. Antennal, first flagellomere approximately 1.7 times as long as pedicel (
<figureCitation id="13542A4AFFCFFFF9A03EFB7C81C3E03D" captionStart="FIGURE 3" captionStartId="5.[151,250,1430,1454]" captionTargetBox="[151,1435,181,1406]" captionTargetId="figure-20@5.[151,1436,181,1406]" captionTargetPageId="5" captionText="FIGURE 3. Right antennae (inner-lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Abbreviations: ar, arista; flgm 1, first flagellomere; ped, pedicel. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906005" httpUri="https://zenodo.org/record/7906005/files/figure.png" pageId="25" pageNumber="226">Fig. 3D</figureCitation>
). Gena grayish yellow (
<figureCitation id="13542A4AFFCFFFF9A49CFB588349E03D" box="[461,567,1255,1281]" captionStart="FIGURE 14" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[222,1364,181,1919]" captionTargetId="figure-20@19.[222,1364,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Habitus (lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, male; 2, female. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906035" httpUri="https://zenodo.org/record/7906035/files/figure.png" pageId="25" pageNumber="226">Fig. 14D</figureCitation>
). Supracervical setae 6–9 and postoculars 13–15 per side (
<figureCitation id="13542A4AFFCFFFF9A180FB588450E03D" box="[1233,1326,1255,1281]" captionStart="FIGURE 4" captionStartId="6.[151,250,1863,1887]" captionTargetBox="[162,1420,181,1839]" captionTargetId="figure-19@6.[157,1429,181,1839]" captionTargetPageId="6" captionText="FIGURE 4. Head parts (caudal view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Abbreviations: dl a, dorsolateral tentorial apodeme; dm a, dorsomedial tentorial apodeme; i ocp s, inner occipital seta; o vt s, outer vertical seta; spc s, supracervical seta. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906007" httpUri="https://zenodo.org/record/7906007/files/figure.png" pageId="25" pageNumber="226">Fig. 4D</figureCitation>
). Cibarial projections at anterolateral corners shorter than width of anterior margin; medial sensilla in parallel rows wider than sensilla campaniformia (
<figureCitation id="13542A4AFFCFFFF9A4FFFA90837BE075" box="[430,517,1326,1353]" captionStart="FIGURE 5" captionStartId="7.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1789]" captionTargetId="figure-20@7.[151,1436,181,1790]" captionTargetPageId="7" captionText="FIGURE 5. Mouthparts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, caudal view. Abbreviations: a s, anterior sensilla; cib, cibarium; clyp, clypeus; m s, medial sensilla; p s, posterior sensilla; plp, palpus; premnt, prementum; s c, sensilla campaniformia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906011" httpUri="https://zenodo.org/record/7906011/files/figure.png" pageId="25" pageNumber="226">Fig. 5D</figureCitation>
<figureCitation id="13542A4AFFCFFFF9A757FAFE8370E073" box="[518,526,1345,1359]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1963]" captionTargetBox="[151,1433,634,1915]" captionTargetId="figure-203@4.[151,1436,633,1915]" captionTargetPageId="4" captionText="FIGURE 2. Head parts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, right eye; 2, frontal view.Abbreviation: pc orb s, proclinate orbital seta. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906001" httpUri="https://zenodo.org/record/7906001/files/figure.png" pageId="25" pageNumber="226">
<subScript id="17EB348AFFCFFFF9A757FAFE8370E073" attach="right" box="[518,526,1345,1359]" fontSize="6" pageId="25" pageNumber="226">2</subScript>
</figureCitation>
). Prementum and neighboring lateral membrane with approximately 5 setae per side (
<figureCitation id="13542A4AFFCFFFF9A585FAEC8053E051" box="[212,301,1362,1389]" captionStart="FIGURE 5" captionStartId="7.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1789]" captionTargetId="figure-20@7.[151,1436,181,1790]" captionTargetPageId="7" captionText="FIGURE 5. Mouthparts of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, caudal view. Abbreviations: a s, anterior sensilla; cib, cibarium; clyp, clypeus; m s, medial sensilla; p s, posterior sensilla; plp, palpus; premnt, prementum; s c, sensilla campaniformia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906011" httpUri="https://zenodo.org/record/7906011/files/figure.png" pageId="25" pageNumber="226">Fig. 5D</figureCitation>
).
</paragraph>
<paragraph id="8BD036CFFFCFFFF9A596FAC885D9E0AD" blockId="25.[151,1437,1038,1931]" box="[199,1191,1399,1425]" pageId="25" pageNumber="226">
Thorax. Scutum and scutellum glossy, blackish brown; pleura grayish brown (
<figureCitation id="13542A4AFFCFFFF9A163FAC885E4E0AD" box="[1074,1178,1399,1425]" captionStart="FIGURE 14" captionStartId="19.[151,250,1943,1967]" captionTargetBox="[222,1364,181,1919]" captionTargetId="figure-20@19.[222,1364,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Habitus (lateral view) of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, male; 2, female. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906035" httpUri="https://zenodo.org/record/7906035/files/figure.png" pageId="25" pageNumber="226">Fig. 14D</figureCitation>
).
</paragraph>
<paragraph id="8BD036CFFFCFFFF9A596FA248380E089" blockId="25.[151,1437,1038,1931]" box="[199,766,1435,1467]" pageId="25" pageNumber="226">
Legs. Fore tarsomere I as long as II+III (
<figureCitation id="13542A4AFFCFFFF9A7DEFA248398E089" box="[655,742,1435,1461]" captionStart="FIGURE 9" captionStartId="11.[151,250,1926,1950]" captionTargetBox="[260,1328,181,1900]" captionTargetId="figure-20@11.[260,1328,181,1901]" captionTargetPageId="11" captionText="FIGURE 9. Legs of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, fore tibia (distalmost portion) and tarsus; 2, mid tibia (distal portion) and tarsus; 3, hind tibia (distal portion) and tarsus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906021" httpUri="https://zenodo.org/record/7906021/files/figure.png" pageId="25" pageNumber="226">Fig. 9D</figureCitation>
<figureCitation id="13542A4AFFCFFFF9A7B4FA128393E087" box="[741,749,1453,1467]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="25" pageNumber="226">
<subScript id="17EB348AFFCFFFF9A7B4FA128393E087" attach="left" box="[741,749,1453,1467]" fontSize="6" pageId="25" pageNumber="226">1</subScript>
</figureCitation>
).
</paragraph>
<paragraph id="8BD036CFFFCFFFF9A596FA00832BE0E5" blockId="25.[151,1437,1038,1931]" box="[199,597,1471,1497]" pageId="25" pageNumber="226">Abdomen. Sternites grayish brown.</paragraph>
<paragraph id="8BD036CFFFCFFFF9A596FA5C8445E0C1" blockId="25.[151,1437,1038,1931]" box="[199,1339,1507,1533]" pageId="25" pageNumber="226">Male terminalia. Cercus pubescent except for anterior to ventral margin, with approximately 32 setae.</paragraph>
<paragraph id="8BD036CFFFCFFFF9A596F9B88475E31D" blockId="25.[151,1437,1038,1931]" box="[199,1291,1542,1575]" pageId="25" pageNumber="226">
Female terminalia. Epigynium with 5–6 sensilla per side near lower posterior margin (
<figureCitation id="13542A4AFFCFFFF9A1DFF9B8858CE31D" box="[1166,1266,1543,1569]" captionStart="FIGURE 12" captionStartId="16.[151,250,1932,1956]" captionTargetBox="[169,1419,420,1909]" captionTargetId="figure-108@16.[169,1419,417,1909]" captionTargetPageId="16" captionText="FIGURE 12. Female terminalia of Colocasiomyia pinangae sp. nov. (A), C. besaris sp. nov. (B), C. luciphila sp. nov. (C), C. oligochaeta sp. nov. (D), and C. grimaldii sp. nov. (E). Subscript 1, lateral view; 2, ventral view. Abbreviations: epig, epigynium; epiprct, epiproct; hypg v, hypogynial valve; hyprct, hypoproct. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7906029" httpUri="https://zenodo.org/record/7906029/files/figure.png" pageId="25" pageNumber="226">Fig. 12D</figureCitation>
<figureCitation id="13542A4AFFCFFFF9A1A3F9A68584E31B" box="[1266,1274,1561,1575]" captionStart="FIGURE 1" captionStartId="3.[151,250,1943,1967]" captionTargetBox="[230,1356,181,1918]" captionTargetId="figure-20@3.[230,1357,181,1919]" captionTargetPageId="3" captionText="FIGURE 1. The unrooted neighbor-joining tree constructed for species delimitation of the Colocasiomyia zeylanica group based on DNA barcoding with the 658-bp fragment sequences of COI gene. Bootstrap percentages (≥50) are shown beside corresponding internal branches." figureDoi="http://doi.org/10.5281/zenodo.7905997" httpUri="https://zenodo.org/record/7905997/files/figure.png" pageId="25" pageNumber="226">
<subScript id="17EB348AFFCFFFF9A1A3F9A68584E31B" attach="left" box="[1266,1274,1561,1575]" fontSize="6" pageId="25" pageNumber="226">1</subScript>
</figureCitation>
).
</paragraph>
<paragraph id="8BD036CFFFCFFFF9A596F99480D7E3B1" blockId="25.[151,1437,1038,1931]" pageId="25" pageNumber="226">
Measurements (
<typeStatus id="54D4886DFFCFFFF9A42DF99480A1E379" box="[380,479,1579,1605]" pageId="25" pageNumber="226" type="holotype">holotype</typeStatus>
/range in 5♁ and 
<specimenCount id="9D69FD46FFCFFFF9A7FEF99583AAE379" box="[687,724,1578,1605]" pageId="25" pageNumber="226" type="female">5♀</specimenCount>
, or less if noted, 
<typeStatus id="54D4886DFFCFFFF9A6F5F994856CE379" box="[932,1042,1579,1605]" pageId="25" pageNumber="226" type="paratype">paratypes</typeStatus>
, in mm): BL = 1.51/♁ 1.30–1.62, 
<specimenCount id="9D69FD46FFCFFFF9A5C6F9F081D2E355" box="[151,172,1615,1641]" pageId="25" pageNumber="226" type="female">♀</specimenCount>
1.42–1.64 (
<specimenCount id="9D69FD46FFCFFFF9A469F9F08025E355" box="[312,347,1615,1641]" pageId="25" pageNumber="226" type="female">4♀</specimenCount>
); ThL = 0.69/♁ 0.59–0.75, 
<specimenCount id="9D69FD46FFCFFFF9A7CAF9F083CEE355" box="[667,688,1615,1641]" pageId="25" pageNumber="226" type="female">♀</specimenCount>
0.59–0.71; WL = 1.35/♁ 1.20–1.44, 
<specimenCount id="9D69FD46FFCFFFF9A10AF9F0850EE355" box="[1115,1136,1615,1641]" pageId="25" pageNumber="226" type="female">♀</specimenCount>
1.14–1.38; WW = 0.66/♁ 0.62–0.76, 
<specimenCount id="9D69FD46FFCFFFF9A444F9CC8054E3B1" box="[277,298,1651,1677]" pageId="25" pageNumber="226" type="female">♀</specimenCount>
0.54–0.72.
</paragraph>
<paragraph id="8BD036CFFFCFFFF9A596F9288348E221" blockId="25.[151,1437,1038,1931]" pageId="25" pageNumber="226">
Indices (
<typeStatus id="54D4886DFFCFFFF9A478F92880F2E38D" box="[297,396,1687,1713]" pageId="25" pageNumber="226" type="holotype">holotype</typeStatus>
/range in 5♁ and 
<specimenCount id="9D69FD46FFCFFFF9A709F9298303E38D" box="[600,637,1686,1713]" pageId="25" pageNumber="226" type="female">5♀</specimenCount>
, or less if noted, 
<typeStatus id="54D4886DFFCFFFF9A619F92882C8E38D" box="[840,950,1687,1713]" pageId="25" pageNumber="226" type="paratype">paratypes</typeStatus>
, in ratio): FW/HW = 0.56/0.56–0.62; ch/o = 0.34/0.34–0.39; rcorb = 0.44/0.47–0.60 (4♁ and 
<specimenCount id="9D69FD46FFCFFFF9A788F9048382E3E9" box="[729,764,1723,1749]" pageId="25" pageNumber="226" type="female">3♀</specimenCount>
); vb = 0.33/0.27–0.38; sctl = 0.62/0.61–0.74 (4♁ and 
<specimenCount id="9D69FD46FFCFFFF9A039F90584F5E3E9" box="[1384,1419,1722,1749]" pageId="25" pageNumber="226" type="female">5♀</specimenCount>
); sctlp = 1.10/0.89–1.17; C = 1.44/1.28–1.47; 4c = 1.74/1.69–2.13; 4v = 2.57/2.60–3.15; 5x = 4.61/4.02–5.11; ac = 3.05/2.72–3.57; M = 1.15/1.10–1.34.
</paragraph>
</subSubSection>
<subSubSection id="C3756544FFCFFFF9A596F8988428E2B5" pageId="25" pageNumber="226" type="materials_examined">
<paragraph id="8BD036CFFFCFFFF9A596F89884EEE27D" blockId="25.[151,1437,1038,1931]" box="[199,1424,1831,1859]" pageId="25" pageNumber="226">
<materialsCitation id="3B073C92FFCFFFF9A596F89884EEE27D" ID-GBIF-Occurrence="4101310314" box="[199,1424,1831,1859]" collectingDate="2004-12-12" collectionCode="MZB, T" country="Indonesia" location="Cibodas" pageId="25" pageNumber="226" specimenCount="1" stateProvince="Jawa Barat" typeStatus="holotype">
<emphasis id="B91BEADDFFCFFFF9A596F898804DE27D" bold="true" box="[199,307,1831,1857]" pageId="25" pageNumber="226">
<typeStatus id="54D4886DFFCFFFF9A596F8988053E27D" box="[199,301,1831,1857]" pageId="25" pageNumber="226" type="holotype">Holotype</typeStatus>
.
</emphasis>
♁ (
<collectionCode id="ED7EAE0AFFCFFFF9A409F89880EBE27D" box="[344,405,1831,1857]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="25" pageNumber="226" type="Museum">MZB</collectionCode>
), “ 
<location id="8EB06014FFCFFFF9A4FEF8988379E27D" LSID="urn:lsid:plazi:treatment:03C687D9FFCFFFFAA5C6FCC7857DE5E9:8EB06014FFCFFFF9A4FEF8988379E27D" box="[431,519,1831,1857]" country="Indonesia" name="Cibodas" pageId="25" pageNumber="226" stateProvince="Jawa Barat">Cibodas</location>
, 
<collectingRegion id="49ABF82DFFCFFFF9A740F8988306E27D" box="[529,632,1831,1857]" country="Indonesia" name="Jawa Barat" pageId="25" pageNumber="226">West Java</collectingRegion>
, 
<collectingCountry id="F378765FFFCFFFF9A7D3F8988399E27D" box="[642,743,1831,1857]" name="Indonesia" pageId="25" pageNumber="226">Indonesia</collectingCountry>
, 
<date id="FFD1100FFFCFFFF9A7A0F8988218E27D" box="[753,870,1831,1857]" pageId="25" pageNumber="226" value="2004-12-12">
<collectingDate id="EF95E9E7FFCFFFF9A7A0F8988218E27D" box="[753,870,1831,1857]" pageId="25" pageNumber="226" value="2004-12-12">12.xii.2004</collectingDate>
</date>
, K. 
<collectionCode id="ED7EAE0AFFCFFFF9A6D9F89882E7E27D" box="[904,921,1831,1857]" name="Tavera, Department of Geology and Geophysics" pageId="25" pageNumber="226">T</collectionCode>
. Takano / ex inflorescence of 
<taxonomicName id="4C6F4D4CFFCFFFF9A194F8988403E27C" box="[1221,1405,1831,1856]" class="Liliopsida" family="Arecaceae" genus="Pinanga" kingdom="Plantae" order="Arecales" pageId="25" pageNumber="226" phylum="Tracheophyta" rank="species" species="coronata">
<emphasis id="B91BEADDFFCFFFF9A194F8988403E27C" box="[1221,1405,1831,1856]" italics="true" pageId="25" pageNumber="226">Pinanga coronata</emphasis>
</taxonomicName>
”.
</materialsCitation>
</paragraph>
<paragraph id="8BD036CFFFCFFFF9A596F8F48428E2B5" blockId="25.[151,1437,1038,1931]" pageId="25" pageNumber="226">
<materialsCitation id="3B073C92FFCFFFF9A596F8F482EFE25B" ID-GBIF-Occurrence="4101310307" box="[199,913,1867,1895]" collectingDate="2004-12-12" collectionCode="MZB, T" country="Indonesia" location="Cibodas" pageId="25" pageNumber="226" specimenCount="10" specimenCount-female="10" stateProvince="Jawa Barat" typeStatus="paratype">
<emphasis id="B91BEADDFFCFFFF9A596F8F48038E259" bold="true" box="[199,326,1867,1893]" pageId="25" pageNumber="226">
<typeStatus id="54D4886DFFCFFFF9A596F8F4803FE259" box="[199,321,1867,1893]" pageId="25" pageNumber="226" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="F378765FFFCFFFF9A401F8F480BFE259" box="[336,449,1867,1893]" name="Indonesia" pageId="25" pageNumber="226">Indonesia</collectingCountry>
: 5♁, 
<specimenCount id="9D69FD46FFCFFFF9A753F8F4834BE259" box="[514,565,1867,1893]" pageId="25" pageNumber="226" type="female">10♀</specimenCount>
, same data as the holotype, 7
</materialsCitation>
♁, 
<materialsCitation id="3B073C92FFCFFFF9A6E2F8F4808BE2B7" ID-GBIF-Occurrence="4101310305" collectingDate="2003-12-24" collectionCode="MZB, SEHU" country="Indonesia" location="Cibodas" pageId="25" pageNumber="226" specimenCount="20" specimenCount-female="20" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="9D69FD46FFCFFFF9A6E2F8F48298E259" box="[947,998,1867,1893]" pageId="25" pageNumber="226" type="female">20♀</specimenCount>
, same data as the holotype except for 
<date id="FFD1100FFFCFFFF9A5C6F8D08069E2B5" box="[151,279,1903,1929]" pageId="25" pageNumber="226" value="2003-12-24">
<collectingDate id="EF95E9E7FFCFFFF9A5C6F8D08069E2B5" box="[151,279,1903,1929]" pageId="25" pageNumber="226" value="2003-12-24">24.xii.2003</collectingDate>
</date>
(
<collectionCode id="ED7EAE0AFFCFFFF9A477F8D08019E2B5" box="[294,359,1903,1929]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="25" pageNumber="226" type="Museum">MZB</collectionCode>
, 
<collectionCode id="ED7EAE0AFFCFFFF9A423F8D080BEE2B5" box="[370,448,1903,1929]" pageId="25" pageNumber="226">SEHU</collectionCode>
); 10
</materialsCitation>
♁, 
<materialsCitation id="3B073C92FFCFFFF9A743F8D0842CE2B5" ID-GBIF-Occurrence="4101310301" box="[530,1362,1902,1929]" collectingDate="2003-12-24" collectionCode="KIZ" country="Indonesia" location="Cibodas" pageId="25" pageNumber="226" specimenCount="10" specimenCount-female="10" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="9D69FD46FFCFFFF9A743F8D0833DE2B5" box="[530,579,1903,1929]" pageId="25" pageNumber="226" type="female">10♀</specimenCount>
(#05180–99), same data as the holotype except for 
<date id="FFD1100FFFCFFFF9A1D6F8D08479E2B5" box="[1159,1287,1903,1929]" pageId="25" pageNumber="226" value="2003-12-24">
<collectingDate id="EF95E9E7FFCFFFF9A1D6F8D08479E2B5" box="[1159,1287,1903,1929]" pageId="25" pageNumber="226" value="2003-12-24">24.xii.2003</collectingDate>
</date>
(
<collectionCode id="ED7EAE0AFFCFFFF9A047F8D08434E2B5" box="[1302,1354,1903,1929]" country="China" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="25" pageNumber="226">KIZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3756544FFCCFFFAA596FF29831BE58D" box="[199,613,150,177]" pageId="26" pageNumber="227" type="distribution">
<paragraph id="8BD036CFFFCCFFFAA596FF29831BE58D" blockId="26.[199,1027,150,213]" box="[199,613,150,177]" pageId="26" pageNumber="227">
<emphasis id="B91BEADDFFCCFFFAA596FF29801FE58C" bold="true" box="[199,353,150,176]" pageId="26" pageNumber="227">Distribution.</emphasis>
<collectingCountry id="F378765FFFCCFFFAA436FF2880AAE58D" box="[359,468,151,177]" name="Indonesia" pageId="26" pageNumber="227">Indonesia</collectingCountry>
(
<collectingRegion id="49ABF82DFFCCFFFAA4B2FF288327E58D" box="[483,601,151,177]" country="Indonesia" name="Jawa Barat" pageId="26" pageNumber="227">West Java</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3756544FFCCFFFAA596FF04857DE5E9" box="[199,1027,187,213]" pageId="26" pageNumber="227" type="etymology">
<paragraph id="8BD036CFFFCCFFFAA596FF04857DE5E9" blockId="26.[199,1027,150,213]" box="[199,1027,187,213]" pageId="26" pageNumber="227">
<emphasis id="B91BEADDFFCCFFFAA596FF048030E5E9" bold="true" box="[199,334,187,213]" pageId="26" pageNumber="227">Etymology.</emphasis>
Referring to the lack of some setae from the head and thorax.
</paragraph>
</subSubSection>
</treatment>
</document>