Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species Author Zhang, Guang Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology (Institute of Biodiversity), Yunnan University, Kunming, 650091 China & Guizhou Provincial Museum, Guiyang, 550081 China Author Gao, Jian-Jun 0000-0003-4636-2130 Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology (Institute of Biodiversity), Yunnan University, Kunming, 650091 China & gaojj @ ynu. edu. cn; https: // orcid. org / 0000 - 0003 - 4636 - 2130 gaojj@ynu.edu.cn Author Takano, Kohei Takenaka Graduate School of Environmental Earth Science, Hokkaido University, Sapporo 060 - 0810, Japan & Research Institute for Humanity and Nature, Kyoto 603 - 8047, Japan Author Yafuso, Masako 0000-0001-9737-7792 Uema 1 - 2 - 21, Naha, Okinawa 902 - 0073, Japan. yafusom @ juno. ocn. ne. jp; https: // orcid. org / 0000 - 0001 - 9737 - 7792 yafusom@juno.ocn.ne.jp Author Suwito, Awit 0000-0003-2600-2488 Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. sungaicola @ gmail. com; https: // orcid. org / 0000 - 0003 - 2600 - 2488 sungaicola@gmail.com Author Meleng, Paulus Ak 0000-0003-3777-0597 Research Development and Innovation Division, Forest Department Sarawak, KM 10, Jalan Datuk Amar Kalong Ningkan, 93250, Kuching, Sarawak, Malaysia. paulusm @ sarawak. gov. my; https: // orcid. org / 0000 - 0003 - 3777 - 0597 paulusm@sarawak.gov.my Author Toda, Masanori J. Hokkaido University Museum, Hokkaido University, Sapporo 060 - 0810, Japan text Zootaxa 2023 2023-05-05 5278 2 201 238 http://dx.doi.org/10.11646/zootaxa.5278.2.1 journal article 57601 10.11646/zootaxa.5278.2.1 2d4bcf32-8354-41cd-a2fd-05e47458c178 1175-5326 7905995 697AC989-141D-4A89-8F02-5A4644E303EA 4) Colocasiomyia oligochaeta Zhang & Toda , sp. nov. ( Figs 2–12D , 14D ) Colocasiomyia sp.2 aff. nepalensis : Sultana et al. , 2006: 694 ; Fartyal et al. , 2013: 768 . Diagnosis. Antennal arista approximately 1.8 times as long as first flagellomere ( Fig. 3D ). Distance between antennal scapes as wide as scape diameter ( Fig. 2D 2 ). Acrostichal setulae in 6 rows ( Fig. 6D ). All legs entirely grayish yellow to yellow except for grayish brown tarsomere Vs ( Fig. 14D ). Cercus slightly projected at apicoventral corner (Fig. 11-2D). Long, tube-like, membranous aedeagus ornamented with relatively sparse and weak spinules (Fig. 11-2D). Hypogynial valves with dorsal flaps fused into small, ligulate plate at distal end of basal portion ( Fig. 12D ). Description (♁ and ♀ ). Head. Antennal, first flagellomere approximately 1.7 times as long as pedicel ( Fig. 3D ). Gena grayish yellow ( Fig. 14D ). Supracervical setae 6–9 and postoculars 13–15 per side ( Fig. 4D ). Cibarial projections at anterolateral corners shorter than width of anterior margin; medial sensilla in parallel rows wider than sensilla campaniformia ( Fig. 5D 2 ). Prementum and neighboring lateral membrane with approximately 5 setae per side ( Fig. 5D ). Thorax. Scutum and scutellum glossy, blackish brown; pleura grayish brown ( Fig. 14D ). Legs. Fore tarsomere I as long as II+III ( Fig. 9D 1 ). Abdomen. Sternites grayish brown. Male terminalia. Cercus pubescent except for anterior to ventral margin, with approximately 32 setae. Female terminalia. Epigynium with 5–6 sensilla per side near lower posterior margin ( Fig. 12D 1 ). Measurements ( holotype /range in 5♁ and 5♀ , or less if noted, paratypes , in mm): BL = 1.51/♁ 1.30–1.62, ♀ 1.42–1.64 ( 4♀ ); ThL = 0.69/♁ 0.59–0.75, ♀ 0.59–0.71; WL = 1.35/♁ 1.20–1.44, ♀ 1.14–1.38; WW = 0.66/♁ 0.62–0.76, ♀ 0.54–0.72. Indices ( holotype /range in 5♁ and 5♀ , or less if noted, paratypes , in ratio): FW/HW = 0.56/0.56–0.62; ch/o = 0.34/0.34–0.39; rcorb = 0.44/0.47–0.60 (4♁ and 3♀ ); vb = 0.33/0.27–0.38; sctl = 0.62/0.61–0.74 (4♁ and 5♀ ); sctlp = 1.10/0.89–1.17; C = 1.44/1.28–1.47; 4c = 1.74/1.69–2.13; 4v = 2.57/2.60–3.15; 5x = 4.61/4.02–5.11; ac = 3.05/2.72–3.57; M = 1.15/1.10–1.34. Holotype . ♁ ( MZB ), “ Cibodas , West Java , Indonesia , 12.xii.2004 , K. T . Takano / ex inflorescence of Pinanga coronata ”. Paratypes . Indonesia : 5♁, 10♀ , same data as the holotype, 7 ♁, 20♀ , same data as the holotype except for 24.xii.2003 ( MZB , SEHU ); 10 ♁, 10♀ (#05180–99), same data as the holotype except for 24.xii.2003 ( KIZ ) . Distribution. Indonesia ( West Java ). Etymology. Referring to the lack of some setae from the head and thorax.