333 lines
29 KiB
XML
333 lines
29 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e83035" ID-Pensoft-Pub="1864-8312-80-439" ID-Pensoft-UUID="3F4E21B524465471831D09738C2304A6" ID-ZooBank="62FFB94CEBF441639F22881435EFC37C" ModsDocID="1864-8312-80-439" checkinTime="1663021027595" checkinUser="pensoft" docAuthor="Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha" docDate="2022" docId="229C83CB948C50B684BCA5FF071C86F1" docLanguage="en" docName="ArthropodSystPhyl 80: 439-494" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-09-12" docSource="http://dx.doi.org/10.3897/asp.80.e83035" docTitle="Amphineurus (Rhamphoneurus) morphyi Santos & Santos & Ribeiro 2022, sp. nov." docType="treatment" docUuid="3200E7A9-C1D2-4CE0-9A73-BDC7F670061B" docUuidSource="ZooBank" docVersion="1" id="3F4E21B524465471831D09738C2304A6" lastPageNumber="439" masterDocId="3F4E21B524465471831D09738C2304A6" masterDocTitle="Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)" masterLastPageNumber="494" masterPageNumber="439" pageNumber="439" updateTime="1663021027595" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Santos, Daubian</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1220-1267</mods:nameIdentifier>
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<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">daubians@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Santos, Rodrigo dos Reis</mods:namePart>
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<mods:affiliation>Pos-Graduacao em Entomologia, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Parana, Brazil</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ribeiro, Guilherme Cunha</mods:namePart>
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<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-09-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>80</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>439</mods:start>
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<mods:end>494</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.80.e83035</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e83035</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-80-439</mods:identifier>
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<mods:identifier type="ZooBank">62FFB94CEBF441639F22881435EFC37C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">3F4E21B524465471831D09738C2304A6</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:3200E7A9-C1D2-4CE0-9A73-BDC7F670061B" httpUri="http://treatment.plazi.org/id/229C83CB948C50B684BCA5FF071C86F1" lastPageNumber="439" pageId="0" pageNumber="439">
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<subSubSection pageId="0" pageNumber="439" type="nomenclature">
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<paragraph pageId="0" pageNumber="439">
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3.3.16.
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<taxonomicName LSID="http://zoobank.org/3200E7A9-C1D2-4CE0-9A73-BDC7F670061B" authority="Santos & Santos & Ribeiro, 2022" authorityName="Santos & Santos & Ribeiro" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) morphyi" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="morphyi" status="sp. nov." subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) morphyi</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="439">sp. nov.</taxonomicNameLabel>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="439" type="materials_examined">
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||
<paragraph pageId="0" pageNumber="439">Material examined.</paragraph>
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||
<paragraph pageId="0" pageNumber="439">
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||
<emphasis bold="true" pageId="0" pageNumber="439">
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||
<typeStatus>Holotype</typeStatus>
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||
</emphasis>
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||
:
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||
<specimenCount>♂</specimenCount>
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||
, Chile,
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||
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
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||
Is., Dalcahue [
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<geoCoordinate degrees="42" direction="south" minutes="22" orientation="latitude" precision="925" value="-42.366665">42°22′S</geoCoordinate>
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<geoCoordinate degrees="73" direction="west" minutes="39" orientation="longitude" precision="925" value="-73.65">73°39′W</geoCoordinate>
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], 1/
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<date value="1968-04-04">4-IV-1968</date>
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,
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<normalizedToken originalValue="Peña">Pena</normalizedToken>
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(USNM)*.
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<materialsCitation collectingDate="1958-03-10" county="Chile" elevation="60" latitude="-40.533333" location="Pucotrihue" longLatPrecision="1162" longitude="-73.7" municipality="Osorno" specimenCount="1" specimenCount-male="1" typeStatus="Paratypes">
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<emphasis bold="true" pageId="0" pageNumber="439">
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<typeStatus>Paratypes</typeStatus>
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</emphasis>
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:
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<specimenCount type="male">1 ♂</specimenCount>
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,
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<collectingCounty>Chile</collectingCounty>
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,
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<collectingMunicipality>Osorno</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:229C83CB948C50B684BCA5FF071C86F1:9A6A1B1CC6D4A019ACFF100337DC21D2" county="Chile" latitude="-40.533333" longLatPrecision="1162" longitude="-73.7" municipality="Osorno" name="Pucotrihue">Pucotrihue</location>
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[
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<geoCoordinate degrees="40" direction="south" minutes="32" orientation="latitude" precision="925" value="-40.533333">40°32′S</geoCoordinate>
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<geoCoordinate degrees="73" direction="west" minutes="42" orientation="longitude" precision="925" value="-73.7">73°42′W</geoCoordinate>
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],
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<quantity metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">
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<elevation metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">60 m</elevation>
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</quantity>
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,
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<collectingDate value="1958-03-10">10-III-1958</collectingDate>
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,
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<normalizedToken originalValue="Peña">Pena</normalizedToken>
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(USNM)
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</materialsCitation>
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*;
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<specimenCount type="male">1 ♂</specimenCount>
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, Chile,
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<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
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Is., Dalcahue [
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<geoCoordinate degrees="42" direction="south" minutes="22" orientation="latitude" precision="925" value="-42.366665">42°22′S</geoCoordinate>
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<geoCoordinate degrees="73" direction="west" minutes="39" orientation="longitude" precision="925" value="-73.65">73°39′W</geoCoordinate>
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], 10/
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<date value="1954-02-12">12-II-1954</date>
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,
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<normalizedToken originalValue="Peña">Pena</normalizedToken>
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(USNM)*;
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<materialsCitation collectingDate="1954-01-31" county="Chile" elevation="1250" latitude="-37.816666" location="Butamalal" longLatPrecision="1179" longitude="-73.23333" municipality="Nahuelbuta" specimenCount="1" specimenCount-male="1" stateProvince="Arauco" typeStatus="paratype">
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<specimenCount type="male">1 ♂</specimenCount>
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,
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<collectingCounty>Chile</collectingCounty>
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,
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<collectingRegion country="Chile" name="Araucania">Arauco</collectingRegion>
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,
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<collectingMunicipality>Nahuelbuta</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:229C83CB948C50B684BCA5FF071C86F1:491083A84BC6DE1A872DF51A1E8E0F71" county="Chile" latitude="-37.816666" longLatPrecision="1179" longitude="-73.23333" municipality="Nahuelbuta" name="Butamalal" stateProvince="Arauco">Butamalal</location>
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[
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<geoCoordinate degrees="37" direction="south" minutes="49" orientation="latitude" precision="925" value="-37.816666">37°49′S</geoCoordinate>
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<geoCoordinate degrees="73" direction="west" minutes="14" orientation="longitude" precision="925" value="-73.23333">73°14′W</geoCoordinate>
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],
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" unit="m" value="1250.0" valueMax="1400.0" valueMin="1100.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" unit="m" value="1250.0" valueMax="1400.0" valueMin="1100.0">1100-1400 m</elevation>
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</quantity>
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, 23/
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<collectingDate value="1954-01-31">31-I-1954</collectingDate>
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,
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<normalizedToken originalValue="Peña">Pena</normalizedToken>
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(USNM)
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</materialsCitation>
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*. -
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<emphasis bold="true" pageId="0" pageNumber="439">Additional material</emphasis>
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:
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<materialsCitation collectingDate="1954-02-08" collectingDateMax="1968-04-04" collectingDateMin="1954-02-08" collectorName="Chiloe Is., Chaiten, Pena, Dalcahue" country="CHILE" latitude="-42.366665" location="CHILE" longLatPrecision="1150" longitude="-73.65" specimenCount="1" specimenCount-female="1">
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<emphasis bold="true" pageId="0" pageNumber="439">
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<collectingCountry name="Chile">CHILE</collectingCountry>
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.
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</emphasis>
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<specimenCount type="female">1 ♀</specimenCount>
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and 1 [sex unknown],
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<collectorName>
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<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
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Is.
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</collectorName>
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,
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<collectorName>Chaiten</collectorName>
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[
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<geoCoordinate degrees="42" direction="south" minutes="58" orientation="latitude" precision="925" value="-42.966667">42°58′S</geoCoordinate>
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<geoCoordinate degrees="72" direction="west" minutes="32" orientation="longitude" precision="925" value="-72.53333">72°32′W</geoCoordinate>
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], 5/
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<collectingDate value="1954-02-08">8-II-1954</collectingDate>
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,
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<normalizedToken originalValue="Peña">
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<collectorName>Pena</collectorName>
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||
</normalizedToken>
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(USNM); 1 [sex unknown],
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<collectorName>
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<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
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Is.
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</collectorName>
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,
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<collectorName>Dalcahue</collectorName>
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[
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<geoCoordinate degrees="42" direction="south" minutes="22" orientation="latitude" precision="925" value="-42.366665">42°22′S</geoCoordinate>
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<geoCoordinate degrees="73" direction="west" minutes="39" orientation="longitude" precision="925" value="-73.65">73°39′W</geoCoordinate>
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], 1/
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<collectingDate value="1968-04-04">4-IV-1968</collectingDate>
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,
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<normalizedToken originalValue="Peña">Pena</normalizedToken>
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(USNM)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="439" type="etymology">
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<paragraph pageId="0" pageNumber="439">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="439">
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The species is named after Charles Morphy D. dos Santos, a Brazilian entomologist who studied the taxonomy and biogeography of South American
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<taxonomicName authorityName="Skuse" authorityYear="1890" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="439" rank="order">Diptera</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="439" type="diagnosis">
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<paragraph pageId="0" pageNumber="439">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="439">This species is distinguished by only a marking near R2, and basal deflection of R5 shorter than half-length of R2+3+4. Furthermore, the species is characterized by male terminalia with a triangular spur, tip of left mesal lobe bent, tip of right mesal lobe slightly curved and cercus thin and abruptly curved.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="439" type="description">
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<paragraph pageId="0" pageNumber="439">Description.</paragraph>
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<paragraph pageId="0" pageNumber="439">
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Wing length 5.63 mm, width 1.74 mm. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Coloration</emphasis>
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: General coloration yellowish brown. Rostrum and palpus dark brown. First segments of antenna yellow, remainding segments dark yellow. Head blackish brown. Eyes black. Thorax yellowish-brown, pleura yellow. Scutum with one reddish-brown stripe. Halter dark yellow with knob pale. Coxae and legs brownish yellow. Abdomen dark brown. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Head</emphasis>
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(dorsal view Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Amphineurus (Rhamphoneurus) morphyi sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure21" httpUri="https://binary.pensoft.net/fig/740574" pageId="0" pageNumber="439">21B</figureCitation>
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, ventral view Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Amphineurus (Rhamphoneurus) morphyi sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure21" httpUri="https://binary.pensoft.net/fig/740574" pageId="0" pageNumber="439">21A</figureCitation>
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): Rostrum bifurcated with long appendices; first palpal segment shorter than terminal segment; scape thinner than pedicel; terminal flagellomere as long as penultimate. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Thorax</emphasis>
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(lateral view Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Amphineurus (Rhamphoneurus) morphyi sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure21" httpUri="https://binary.pensoft.net/fig/740574" pageId="0" pageNumber="439">21E</figureCitation>
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, dorsal view Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Amphineurus (Rhamphoneurus) morphyi sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure21" httpUri="https://binary.pensoft.net/fig/740574" pageId="0" pageNumber="439">21F</figureCitation>
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): Anatergite shorter than katatergite. Halter with dilated knob. Wing (Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Amphineurus (Rhamphoneurus) morphyi sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure21" httpUri="https://binary.pensoft.net/fig/740574" pageId="0" pageNumber="439">21D</figureCitation>
|
||
) almost clear of markings, except for marking highlighting R2; veins near fork of bM (Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Amphineurus (Rhamphoneurus) morphyi sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure21" httpUri="https://binary.pensoft.net/fig/740574" pageId="0" pageNumber="439">21C</figureCitation>
|
||
) visible; basal deflection of R5 as long as r-m; M1+2 half-length of basal deflection of M1; R2+3+4 shorter than R2+3. -
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Female terminalia</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Amphineurus (Rhamphoneurus) morphyi sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure21" httpUri="https://binary.pensoft.net/fig/740574" pageId="0" pageNumber="439">21H</figureCitation>
|
||
): Female tergite IX wider and as long as tergite X; cercus curved and abruptly dorsad. -
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Male terminalia</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Amphineurus (Rhamphoneurus) morphyi sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure21" httpUri="https://binary.pensoft.net/fig/740574" pageId="0" pageNumber="439">21G</figureCitation>
|
||
): Male tergite IX slightly darkened along dorsal margin, with wide V-shaped median notch. Ventral branch of gonocoxite long. Lobe of gonostylus with rounded lobule; lobule longer than wide, longer than half-length of lobe of gonostylus, narrower than stem. Clasper of gonostylus with differently shaped branches: lateral branch straight; medial branch curved. Mesal lobes of gonocoxite asymmetrical; left mesal lobe with posterior projection anvil-shaped; right mesal lobe curved. Sheath of aedeagus straightened with conical projection, darkened only at tip.
|
||
</paragraph>
|
||
<caption doi="10.3897/asp.80.e83035.figure21" httpUri="https://binary.pensoft.net/fig/740574" pageId="0" pageNumber="439" start="Figure 21" startId="F21">
|
||
<paragraph pageId="0" pageNumber="439">
|
||
<emphasis bold="true" pageId="0" pageNumber="439">Figure 21.</emphasis>
|
||
<taxonomicName authorityName="Santos & Santos & Ribeiro" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) morphyi" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="morphyi" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) morphyi</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="439">A</emphasis>
|
||
head (ventral view);
|
||
<emphasis bold="true" pageId="0" pageNumber="439">B</emphasis>
|
||
head (dorsal view);
|
||
<emphasis bold="true" pageId="0" pageNumber="439">C</emphasis>
|
||
detail of wing;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">D</emphasis>
|
||
wing;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">E</emphasis>
|
||
thorax (lateral view);
|
||
<emphasis bold="true" pageId="0" pageNumber="439">F</emphasis>
|
||
thorax (dorsal view);
|
||
<emphasis bold="true" pageId="0" pageNumber="439">G</emphasis>
|
||
male tergite IX (dorsal view) and remainder of male terminalia (lateral view);
|
||
<emphasis bold="true" pageId="0" pageNumber="439">H</emphasis>
|
||
female terminalia (dorsal view). - Abbreviations:
|
||
<emphasis bold="true" pageId="0" pageNumber="439">aed sh</emphasis>
|
||
, sheath of aedeagus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">anatg</emphasis>
|
||
, anatergite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">anepm</emphasis>
|
||
, anepimeron;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">anepst</emphasis>
|
||
, anepisternum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">aprn</emphasis>
|
||
, anteropronotum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">cer</emphasis>
|
||
, cercus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">cgonst</emphasis>
|
||
, clasper of gonostylus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">cx</emphasis>
|
||
, coxa;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">goncx</emphasis>
|
||
, gonocoxite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">hlt</emphasis>
|
||
, halter;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">hyp val</emphasis>
|
||
, hypogynial valve;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">k</emphasis>
|
||
, knob of halter;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">kepm</emphasis>
|
||
, katepimeron;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">kepst</emphasis>
|
||
, katepisternum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">ktg</emphasis>
|
||
, katatergite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">l ms</emphasis>
|
||
, left mesal lobe of gonocoxite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">lgonst</emphasis>
|
||
; lobe of gonostylus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mr</emphasis>
|
||
, meron;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">ms lobe</emphasis>
|
||
, mesal lobe of gonocoxite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mtanepst</emphasis>
|
||
, metanepisternum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mtepm</emphasis>
|
||
, metepimeron;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mtg</emphasis>
|
||
, mediotergite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mtkepst</emphasis>
|
||
, metakatepisternum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mtn</emphasis>
|
||
, metanotum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">p</emphasis>
|
||
, posterior basalare;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">patg</emphasis>
|
||
, paratergite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">pprn</emphasis>
|
||
, postpronotum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">presct</emphasis>
|
||
, prescutum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">r ms</emphasis>
|
||
, right mesal lobe of gonocoxite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">sct</emphasis>
|
||
, scutum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">sctl</emphasis>
|
||
, scutellum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">st</emphasis>
|
||
, sternite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">tg</emphasis>
|
||
, tergite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">t9</emphasis>
|
||
, male tergite IX.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="439" type="remarks">
|
||
<paragraph pageId="0" pageNumber="439">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="439">
|
||
Specimens of this new species were previously identified by C.P. Alexander as
|
||
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) insanus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="insanus" subGenus="R.">A. (R.) insanus</taxonomicName>
|
||
Alexander. This species resembles
|
||
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) amorimi" order="Diptera" pageId="0" pageNumber="439" rank="species" species="amorimi" subGenus="R.">A. (R.) amorimi</taxonomicName>
|
||
sp. nov. but differs mainly in chaetotaxy of the thorax and gonocoxite, length of R2+3+4 and shape of mesal lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |