Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae) Author Santos, Daubian https://orcid.org/0000-0003-1220-1267 Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil daubians@gmail.com Author Santos, Rodrigo dos Reis Pos-Graduacao em Entomologia, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Parana, Brazil Author Ribeiro, Guilherme Cunha Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil text Arthropod Systematics & amp; Phylogeny 2022 2022-09-12 80 439 494 http://dx.doi.org/10.3897/asp.80.e83035 journal article http://dx.doi.org/10.3897/asp.80.e83035 1864-8312-80-439 62FFB94CEBF441639F22881435EFC37C 3F4E21B524465471831D09738C2304A6 3.3.16. Amphineurus (Rhamphoneurus) morphyi sp. nov. Material examined. Holotype : ♂ , Chile, Chiloe Is., Dalcahue [ 42°22′S 73°39′W ], 1/ 4-IV-1968 , Pena (USNM)*. Paratypes : 1 ♂ , Chile , Osorno , Pucotrihue [ 40°32′S 73°42′W ], 60 m , 10-III-1958 , Pena (USNM) *; 1 ♂ , Chile, Chiloe Is., Dalcahue [ 42°22′S 73°39′W ], 10/ 12-II-1954 , Pena (USNM)*; 1 ♂ , Chile , Arauco , Nahuelbuta , Butamalal [ 37°49′S 73°14′W ], 1100-1400 m , 23/ 31-I-1954 , Pena (USNM) *. - Additional material : CHILE . 1 ♀ and 1 [sex unknown], Chiloe Is. , Chaiten [ 42°58′S 72°32′W ], 5/ 8-II-1954 , Pena (USNM); 1 [sex unknown], Chiloe Is. , Dalcahue [ 42°22′S 73°39′W ], 1/ 4-IV-1968 , Pena (USNM) . Etymology. The species is named after Charles Morphy D. dos Santos, a Brazilian entomologist who studied the taxonomy and biogeography of South American Diptera . Diagnosis. This species is distinguished by only a marking near R2, and basal deflection of R5 shorter than half-length of R2+3+4. Furthermore, the species is characterized by male terminalia with a triangular spur, tip of left mesal lobe bent, tip of right mesal lobe slightly curved and cercus thin and abruptly curved. Description. Wing length 5.63 mm, width 1.74 mm. - Coloration : General coloration yellowish brown. Rostrum and palpus dark brown. First segments of antenna yellow, remainding segments dark yellow. Head blackish brown. Eyes black. Thorax yellowish-brown, pleura yellow. Scutum with one reddish-brown stripe. Halter dark yellow with knob pale. Coxae and legs brownish yellow. Abdomen dark brown. - Head (dorsal view Fig. 21B , ventral view Fig. 21A ): Rostrum bifurcated with long appendices; first palpal segment shorter than terminal segment; scape thinner than pedicel; terminal flagellomere as long as penultimate. - Thorax (lateral view Fig. 21E , dorsal view Fig. 21F ): Anatergite shorter than katatergite. Halter with dilated knob. Wing (Fig. 21D ) almost clear of markings, except for marking highlighting R2; veins near fork of bM (Fig. 21C ) visible; basal deflection of R5 as long as r-m; M1+2 half-length of basal deflection of M1; R2+3+4 shorter than R2+3. - Female terminalia (Fig. 21H ): Female tergite IX wider and as long as tergite X; cercus curved and abruptly dorsad. - Male terminalia (Fig. 21G ): Male tergite IX slightly darkened along dorsal margin, with wide V-shaped median notch. Ventral branch of gonocoxite long. Lobe of gonostylus with rounded lobule; lobule longer than wide, longer than half-length of lobe of gonostylus, narrower than stem. Clasper of gonostylus with differently shaped branches: lateral branch straight; medial branch curved. Mesal lobes of gonocoxite asymmetrical; left mesal lobe with posterior projection anvil-shaped; right mesal lobe curved. Sheath of aedeagus straightened with conical projection, darkened only at tip. Figure 21. Amphineurus (Rhamphoneurus) morphyi sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh , sheath of aedeagus; anatg , anatergite; anepm , anepimeron; anepst , anepisternum; aprn , anteropronotum; cer , cercus; cgonst , clasper of gonostylus; cx , coxa; goncx , gonocoxite; hlt , halter; hyp val , hypogynial valve; k , knob of halter; kepm , katepimeron; kepst , katepisternum; ktg , katatergite; l ms , left mesal lobe of gonocoxite; lgonst ; lobe of gonostylus; mr , meron; ms lobe , mesal lobe of gonocoxite; mtanepst , metanepisternum; mtepm , metepimeron; mtg , mediotergite; mtkepst , metakatepisternum; mtn , metanotum; p , posterior basalare; patg , paratergite; pprn , postpronotum; presct , prescutum; r ms , right mesal lobe of gonocoxite; sct , scutum; sctl , scutellum; st , sternite; tg , tergite; t9 , male tergite IX. Remarks. Specimens of this new species were previously identified by C.P. Alexander as A. (R.) insanus Alexander. This species resembles A. (R.) amorimi sp. nov. but differs mainly in chaetotaxy of the thorax and gonocoxite, length of R2+3+4 and shape of mesal lobe.