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<document id="DCAAB3E8567451AC1247D900168A6749" ID-DOI="10.11646/zootaxa.5284.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7919001" ID-ZooBank="7334FB57-41A0-47CA-BCBF-FA108B13CEC1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1683703331630" checkinUser="plazi" docAuthor="Santos, Wildio Ikaro Da Graça, Dolibaina, Diego Rodrigo, Dias, Fernando Maia Silva, Mielke, Olaf Hermann Hendrik &amp; Casagrande, Mirna Martins" docDate="2023" docId="03FB5F61FFA5FFC8FF36FAD5099FFDA5" docLanguage="en" docName="zootaxa.5284.1.3.pdf" docOrigin="Zootaxa 5284 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5284.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Papilio amesis" docType="treatment" docVersion="1" lastPageNumber="98" masterDocId="FFC22719FFB1FFDDFFA1FFA90C03FFEE" masterDocTitle="A review of the South American metalmark genus Alesa Doubleday, 1847 (Lepidoptera: Riodinidae: Eurybiini) with the description of two new species" masterLastPageNumber="120" masterPageNumber="77" pageNumber="97" updateTime="1683836328005" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:titleInfo id="8CF623A140F18BDE664ACC6A4F5760F7">
<mods:title id="DCF2969A2F8E14C724AECEB17366E91D">A review of the South American metalmark genus Alesa Doubleday, 1847 (Lepidoptera: Riodinidae: Eurybiini) with the description of two new species</mods:title>
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<mods:namePart id="D7F1CFAA2E7C3BF733569AE42A004358">Santos, Wildio Ikaro Da Graça</mods:namePart>
<mods:nameIdentifier id="F3AE56740530B0828C6A0F8F4D82E66A" type="ORCID">0000-0003-3466-5820</mods:nameIdentifier>
<mods:affiliation id="9EFE74B195DA4859C34EB88EDBB4FBE4">Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, PO Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil &amp; wildio. ikaro @ gmail. com; https: // orcid. org / 0000 - 0003 - 3466 - 5820</mods:affiliation>
<mods:nameIdentifier id="C316DD169B5D4FF71CB33BF547ECD01E" type="email">wildio.ikaro@gmail.com</mods:nameIdentifier>
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<mods:namePart id="597C4F97347FE2A9BEEC0149E686CC68">Dolibaina, Diego Rodrigo</mods:namePart>
<mods:nameIdentifier id="07846CEA44B1236381EF74C4C2978934" type="ORCID">0000-0001-5258-3787</mods:nameIdentifier>
<mods:affiliation id="18775A1D198F6B2D196E156A91250F48">Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, PO Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil &amp; Reserva Serra Bonita, PO Box 001, 45.880 - 970, Camacan, Bahia, Brazil &amp; dirodrido @ gmail. com; https: // orcid. org / 0000 - 0001 - 5258 - 3787</mods:affiliation>
<mods:nameIdentifier id="8CABF1E48410E0E7C6F617E3800B251A" type="email">dirodrido@gmail.com</mods:nameIdentifier>
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<mods:role id="79F34AD6973C4FE6B22ECC0F7800643A">
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<mods:namePart id="D45053955ADC2E5CF6D95F72A540C6FE">Dias, Fernando Maia Silva</mods:namePart>
<mods:nameIdentifier id="C391EED98BB9C9B4D0237DADAE70E18E" type="ORCID">0000-0002-7198-7792</mods:nameIdentifier>
<mods:affiliation id="E868CC0AF3C93836740F8E89F52CF5C4">Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, PO Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil &amp; Departamento de Biologia Animal e Vegetal, Universidade Estadual de Londrina, PO Box 10011, 86.057 - 970, Londrina, Paraná, Brazil &amp; fernandodias @ uel. br; https: // orcid. org / 0000 - 0002 - 7198 - 7792</mods:affiliation>
<mods:nameIdentifier id="2F87602D9C465A1051DF154EA5C52C1C" type="email">fernandodias@uel.br</mods:nameIdentifier>
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<mods:namePart id="A18BA0B6393B34F9012B68F2A2EE5B03">Mielke, Olaf Hermann Hendrik</mods:namePart>
<mods:nameIdentifier id="B841AF3D45597F272920F43AE8CD139C" type="ORCID">0000-0003-3655-4606</mods:nameIdentifier>
<mods:affiliation id="17D5F459009FE7F1BFF467D093B33EC1">Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, PO Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil &amp; omhesp @ ufpr. br; https: // orcid. org / 0000 - 0003 - 3655 - 4606</mods:affiliation>
<mods:nameIdentifier id="B33BD56BD285785965F6C763D9106653" type="email">omhesp@ufpr.br</mods:nameIdentifier>
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<mods:namePart id="88E3092D42983428060648BEAC2F1CC5">Casagrande, Mirna Martins</mods:namePart>
<mods:nameIdentifier id="170F909E56BA182C222D1A88B46E5FF7" type="ORCID">0000-0002-6076-8463</mods:nameIdentifier>
<mods:affiliation id="593B017CAB0C026BC95E663D297A4E70">Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, PO Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil &amp; mibras @ ufpr. br; https: // orcid. org / 0000 - 0002 - 6076 - 8463</mods:affiliation>
<mods:nameIdentifier id="2FEF3E7A87353862BD1167D26E2A73FC" type="email">mibras@ufpr.br</mods:nameIdentifier>
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<mods:title id="685C906D8A0467FEF029F2107C03C34C">Zootaxa</mods:title>
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<mods:part id="2301DB3D398565C24FACA85F4F7B3506">
<mods:date id="1D2F5651AF287C002E013D93D70EE4E9">2023</mods:date>
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<mods:number id="61D9336F66F46350DA95F625E15DA4A4">2023-05-10</mods:number>
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<mods:number id="67A859FE103077E3298E6FC179771FE2">5284</mods:number>
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<mods:number id="B623D9039744219E8680951CFD8AAACB">1</mods:number>
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<mods:url id="8E4B76DA33B81298DC6A8F9EC60B5F57">http://dx.doi.org/10.11646/zootaxa.5284.1.3</mods:url>
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<mods:classification id="7295CD7387A3A7F1D463747AB59B1A32">journal article</mods:classification>
<mods:identifier id="7CB5A89E6C72BB5E7911794996337E8D" type="DOI">10.11646/zootaxa.5284.1.3</mods:identifier>
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<treatment id="03FB5F61FFA5FFC8FF36FAD5099FFDA5" ID-DOI="http://doi.org/10.5281/zenodo.7921239" ID-Zenodo-Dep="7921239" LSID="urn:lsid:plazi:treatment:03FB5F61FFA5FFC8FF36FAD5099FFDA5" httpUri="http://treatment.plazi.org/id/03FB5F61FFA5FFC8FF36FAD5099FFDA5" lastPageId="21" lastPageNumber="98" pageId="20" pageNumber="97">
<subSubSection id="C348BDFCFFA5FFC9FF36FAD50D5BFA78" box="[151,344,1404,1430]" pageId="20" pageNumber="97" type="nomenclature">
<paragraph id="8BEDEE77FFA5FFC9FF36FAD50D5BFA78" blockId="20.[151,344,1404,1430]" box="[151,344,1404,1430]" pageId="20" pageNumber="97">
<heading id="D0A5591BFFA5FFC9FF36FAD50D5BFA78" bold="true" box="[151,344,1404,1430]" fontSize="11" level="1" pageId="20" pageNumber="97" reason="1">
<emphasis id="B9263265FFA5FFC9FF36FAD50D5BFA78" bold="true" box="[151,344,1404,1430]" pageId="20" pageNumber="97">
“
<taxonomicName id="4C5295F4FFA5FFC9FF07FAD50CF4FA78" baseAuthorityName="Cramer" baseAuthorityYear="1777" box="[166,247,1404,1430]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="97" phylum="Arthropoda" rank="species" species="amesis">
<emphasis id="B9263265FFA5FFC9FF07FAD50CF4FA78" bold="true" box="[166,247,1404,1430]" italics="true" pageId="20" pageNumber="97">amesis</emphasis>
</taxonomicName>
group”
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C348BDFCFFA5FFC9FF36FA6D08B5F911" pageId="20" pageNumber="97" type="discussion">
<paragraph id="8BEDEE77FFA5FFC9FF36FA6D0D39F934" blockId="20.[151,1437,1476,1791]" pageId="20" pageNumber="97">
The “
<taxonomicName id="4C5295F4FFA5FFC9FF77FA6C0D21FA30" baseAuthorityName="Cramer" baseAuthorityYear="1777" box="[214,290,1477,1502]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="97" phylum="Arthropoda" rank="species" species="amesis">
<emphasis id="B9263265FFA5FFC9FF77FA6C0D21FA30" box="[214,290,1477,1502]" italics="true" pageId="20" pageNumber="97">amesis</emphasis>
</taxonomicName>
group” includes small-sized species and the two most common species of the genus (
<taxonomicName id="4C5295F4FFA5FFC9FB43FA6D094EFA30" baseAuthorityName="Cramer" baseAuthorityYear="1777" box="[1250,1357,1476,1502]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="97" phylum="Arthropoda" rank="species" species="amesis">
<emphasis id="B9263265FFA5FFC9FB43FA6D094EFA30" box="[1250,1357,1476,1502]" italics="true" pageId="20" pageNumber="97">A. amesis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5295F4FFA5FFC9FA25FA6D0CD8F9EC" authorityName="Bates" authorityYear="1867" class="Insecta" family="Riodinidae" genus="Alesa" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="97" phylum="Arthropoda" rank="species" species="lipara">
<emphasis id="B9263265FFA5FFC9FA25FA6D0CD8F9EC" italics="true" pageId="20" pageNumber="97">A. lipara</emphasis>
</taxonomicName>
). Males are characterized by the white tip of antennae; wings underside with a characteristic pattern; forewing with outer margin markedly projected in M
<subScript id="17D6EC32FFA5FFC9FDD1F9B70E7BF9C2" attach="left" box="[624,632,1566,1580]" fontSize="6" pageId="20" pageNumber="97">2</subScript>
–M
<subScript id="17D6EC32FFA5FFC9FD3EF9B70EA4F9C2" attach="left" box="[671,679,1566,1580]" fontSize="6" pageId="20" pageNumber="97">3</subScript>
, two distinct postdiscal spots in CuA
<subScript id="17D6EC32FFA5FFC9FB9FF9B70845F9C2" attach="left" box="[1086,1094,1566,1580]" fontSize="6" pageId="20" pageNumber="97">2</subScript>
-2A on the forewing underside; base of valva ventrally large; outer posterior process of valva about as long as inner posterior process; distal margin of the outer posterior process of valva with pointed internal margin; inner posterior process of valva wider than the outer posterior process. Females have wings upper side yellow-spotted; sterigma membranous medially; base of lamella postvaginalis with a small, rounded projection at each side; with two lateral signa on the middle of the corpus bursae.
</paragraph>
<paragraph id="8BEDEE77FFA5FFC9FF66F94D08B5F911" blockId="20.[151,1437,1476,1791]" box="[199,1206,1764,1791]" pageId="20" pageNumber="97">
<emphasis id="B9263265FFA5FFC9FF66F94D0D91F910" bold="true" box="[199,402,1764,1790]" pageId="20" pageNumber="97">Included species.</emphasis>
<taxonomicName id="4C5295F4FFA5FFC9FE39F94D0E07F910" baseAuthorityName="Cramer" baseAuthorityYear="1777" box="[408,516,1764,1790]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="97" phylum="Arthropoda" rank="species" species="amesis">
<emphasis id="B9263265FFA5FFC9FE39F94D0E07F910" box="[408,516,1764,1790]" italics="true" pageId="20" pageNumber="97">A. amesis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5295F4FFA5FFC9FDB3F94D0E83F910" authority="Santos &amp; Dolibaina &amp; Dias &amp; Mielke &amp; Casagrande, 2023" authorityName="Santos &amp; Dolibaina &amp; Dias &amp; Mielke &amp; Casagrande" authorityYear="2023" box="[530,640,1764,1790]" class="Insecta" family="Riodinidae" genus="Alesa" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="97" phylum="Arthropoda" rank="species" species="mariae" status="sp. nov.">
<emphasis id="B9263265FFA5FFC9FDB3F94D0E83F910" box="[530,640,1764,1790]" italics="true" pageId="20" pageNumber="97">A. mariae</emphasis>
</taxonomicName>
<emphasis id="B9263265FFA5FFC9FD26F94D0EE2F910" bold="true" box="[647,737,1764,1790]" pageId="20" pageNumber="97">
<taxonomicNameLabel id="A2158F1EFFA5FFC9FD26F94D0EE2F910" box="[647,737,1764,1790]" pageId="20" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C5295F4FFA5FFC9FD4EF94D0F51F910" authorityName="Bates" authorityYear="1867" box="[751,850,1764,1790]" class="Insecta" family="Riodinidae" genus="Alesa" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="97" phylum="Arthropoda" rank="species" species="lipara">
<emphasis id="B9263265FFA5FFC9FD4EF94D0F51F910" box="[751,850,1764,1790]" italics="true" pageId="20" pageNumber="97">A. lipara</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5295F4FFA5FFC9FCC1F94D0FEAF910" authorityName="Bates" authorityYear="1867" box="[864,1001,1764,1790]" class="Insecta" family="Riodinidae" genus="Alesa" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="97" phylum="Arthropoda" rank="species" species="hemiurga">
<emphasis id="B9263265FFA5FFC9FCC1F94D0FEAF910" box="[864,1001,1764,1790]" italics="true" pageId="20" pageNumber="97">A. hemiurga</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5295F4FFA5FFC9FB81F94D08ACF910" authorityName="Bates" authorityYear="1867" box="[1056,1199,1764,1790]" class="Insecta" family="Riodinidae" genus="Alesa" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="97" phylum="Arthropoda" rank="species" species="thelydrias">
<emphasis id="B9263265FFA5FFC9FB81F94D08ACF910" box="[1056,1199,1764,1790]" italics="true" pageId="20" pageNumber="97">A. thelydrias</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C348BDFCFFA5FFC8FF36F8F9099FFDA5" lastPageId="21" lastPageNumber="98" pageId="20" pageNumber="97" type="key">
<paragraph id="8BEDEE77FFA5FFC9FF36F8F90E83F884" blockId="20.[151,640,1872,1898]" box="[151,640,1872,1898]" pageId="20" pageNumber="97">
<heading id="D0A5591BFFA5FFC9FF36F8F90E83F884" bold="true" box="[151,640,1872,1898]" fontSize="11" level="1" pageId="20" pageNumber="97" reason="1">
<emphasis id="B9263265FFA5FFC9FF36F8F90E83F884" bold="true" box="[151,640,1872,1898]" pageId="20" pageNumber="97">
Key for males from the “
<taxonomicName id="4C5295F4FFA5FFC9FE6FF8F90E1CF884" baseAuthorityName="Cramer" baseAuthorityYear="1777" box="[462,543,1872,1898]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="97" phylum="Arthropoda" rank="species" species="amesis">
<emphasis id="B9263265FFA5FFC9FE6FF8F90E1CF884" bold="true" box="[462,543,1872,1898]" italics="true" pageId="20" pageNumber="97">amesis</emphasis>
</taxonomicName>
group”
</emphasis>
</heading>
</paragraph>
<keyStep id="30A6F6D2FFA5FFC9FF36F83C099FF829" pageId="20" pageNumber="97">
<paragraph id="8BEDEE77FFA5FFC9FF36F83C099FF844" blockId="20.[151,1436,1941,2022]" box="[151,1436,1941,1963]" pageId="20" pageNumber="97">
<keyLead id="30A34D42FFA5FFC9FF36F83C099FF844" box="[151,1436,1941,1963]" pageId="20" pageNumber="97">1 Forewing upper side with light silvery intervenal rays........................................................ 2</keyLead>
</paragraph>
<paragraph id="8BEDEE77FFA5FFC9FF36F81B099FF829" blockId="20.[151,1436,1941,2022]" box="[151,1436,1970,1992]" pageId="20" pageNumber="97">
<keyLead id="30A34D42FFA5FFC9FF36F81B099FF829" box="[151,1436,1970,1992]" pageId="20" pageNumber="97">1’ Forewing upper side without light silvery intervenal rays...................................................... 3</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BEDEE77FFA5FFC8FF36F879099FFF43" blockId="20.[151,1436,1941,2022]" lastBlockId="21.[151,1437,151,291]" lastPageId="21" lastPageNumber="98" pageId="20" pageNumber="97">
<keyLead id="30A34D42FFA5FFC8FF36F879099FFF43" lastPageId="21" lastPageNumber="98" pageId="20" pageNumber="97">
<keyStep id="30A6F6D2FFA5FFC8FF36F879099FFF43" lastPageId="21" lastPageNumber="98" pageId="20" pageNumber="97">
2 Wings upper side black without markings..............................................................
<taxonomicName id="4C5295F4FFA5FFC9FAFAF879099FF80B" baseAuthorityName="Cramer" baseAuthorityYear="1777" box="[1371,1436,2000,2021]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="20" pageNumber="97" phylum="Arthropoda" rank="species" species="amesis">
<emphasis id="B9263265FFA5FFC9FAFAF879099FF80B" box="[1371,1436,2000,2021]" italics="true" pageId="20" pageNumber="97">amesis</emphasis>
</taxonomicName>
2’ Wings upper side pale brown with reddish markings...............................................
<taxonomicName id="4C5295F4FFA4FFC8FAA7FF310949FF43" authority="Santos &amp; Dolibaina &amp; Dias &amp; Mielke &amp; Casagrande, 2023" authorityName="Santos &amp; Dolibaina &amp; Dias &amp; Mielke &amp; Casagrande" authorityYear="2023" box="[1286,1354,152,173]" class="Insecta" family="Riodinidae" genus="Alesa" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="species" species="mariae" status="sp. nov.">
<emphasis id="B9263265FFA4FFC8FAA7FF310949FF43" box="[1286,1354,152,173]" italics="true" pageId="21" pageNumber="98">mariae</emphasis>
</taxonomicName>
<emphasis id="B9263265FFA4FFC8FAF1FF31099FFF43" bold="true" box="[1360,1436,152,173]" pageId="21" pageNumber="98">
<taxonomicNameLabel id="A2158F1EFFA4FFC8FAF1FF31099FFF43" box="[1360,1436,152,173]" pageId="21" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyStep>
</keyLead>
</paragraph>
<keyStep id="30A6F6D2FFA4FFC8FF36FF1C099EFF09" pageId="21" pageNumber="98">
<paragraph id="8BEDEE77FFA4FFC8FF36FF1C099FFF24" blockId="21.[151,1437,151,291]" box="[151,1436,181,203]" pageId="21" pageNumber="98">
<keyLead id="30A34D42FFA4FFC8FF36FF1C099FFF24" box="[151,1436,181,203]" pageId="21" pageNumber="98">3 Wings upper side with spots or bands..................................................................... 4</keyLead>
</paragraph>
<paragraph id="8BEDEE77FFA4FFC8FF36FF7B099EFF09" blockId="21.[151,1437,151,291]" box="[151,1437,210,232]" pageId="21" pageNumber="98">
<keyLead id="30A34D42FFA4FFC8FF36FF7B099EFF09" box="[151,1437,210,232]" pageId="21" pageNumber="98">
3’ Wings upper side without spots or bands...............................................................
<taxonomicName id="4C5295F4FFA4FFC8FAC3FF7B099EFF09" authorityName="Bates" authorityYear="1867" box="[1378,1437,210,231]" class="Insecta" family="Riodinidae" genus="Alesa" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="species" species="lipara">
<emphasis id="B9263265FFA4FFC8FAC3FF7B099EFF09" box="[1378,1437,210,231]" italics="true" pageId="21" pageNumber="98">lipara</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A6F6D2FFA4FFC8FF36FF59099FFECC" pageId="21" pageNumber="98">
<paragraph id="8BEDEE77FFA4FFC8FF36FF59099FFEEA" blockId="21.[151,1437,151,291]" box="[151,1436,239,261]" pageId="21" pageNumber="98">
<keyLead id="30A34D42FFA4FFC8FF36FF59099FFEEA" box="[151,1436,239,261]" pageId="21" pageNumber="98">
4 Wings upper side with purplish sheen...............................................................
<taxonomicName id="4C5295F4FFA4FFC8FAE0FF46099FFEEA" authorityName="Bates" authorityYear="1867" box="[1345,1436,239,260]" class="Insecta" family="Riodinidae" genus="Alesa" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="species" species="hemiurga">
<emphasis id="B9263265FFA4FFC8FAE0FF46099FFEEA" box="[1345,1436,239,260]" italics="true" pageId="21" pageNumber="98">hemiurga</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEDEE77FFA4FFC8FF36FEA4099FFECC" blockId="21.[151,1437,151,291]" box="[151,1436,269,291]" pageId="21" pageNumber="98">
<keyLead id="30A34D42FFA4FFC8FF36FEA4099FFECC" box="[151,1436,269,291]" pageId="21" pageNumber="98">
4’ Wings upper side without purplish sheen............................................................
<taxonomicName id="4C5295F4FFA4FFC8FA9DFEA4099FFECC" authorityName="Bates" authorityYear="1867" box="[1340,1436,269,290]" class="Insecta" family="Riodinidae" genus="Alesa" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="species" species="thelydrias">
<emphasis id="B9263265FFA4FFC8FA9DFEA4099FFECC" box="[1340,1436,269,290]" italics="true" pageId="21" pageNumber="98">thelydrias</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BEDEE77FFA4FFC8FF36FEDC0E94FE61" blockId="21.[151,851,373,436]" box="[151,663,373,399]" pageId="21" pageNumber="98">
<heading id="D0A5591BFFA4FFC8FF36FEDC0E94FE61" bold="true" box="[151,663,373,399]" fontSize="11" level="1" pageId="21" pageNumber="98" reason="1">
<emphasis id="B9263265FFA4FFC8FF36FEDC0E94FE61" bold="true" box="[151,663,373,399]" pageId="21" pageNumber="98">
Key for females from the “
<taxonomicName id="4C5295F4FFA4FFC8FE44FEDC0E35FE61" baseAuthorityName="Cramer" baseAuthorityYear="1777" box="[485,566,373,399]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="species" species="amesis">
<emphasis id="B9263265FFA4FFC8FE44FEDC0E35FE61" bold="true" box="[485,566,373,399]" italics="true" pageId="21" pageNumber="98">amesis</emphasis>
</taxonomicName>
group”
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEDEE77FFA4FFC8FF36FE300F50FE5D" blockId="21.[151,851,373,436]" box="[151,851,409,436]" pageId="21" pageNumber="98">
(females of
<taxonomicName id="4C5295F4FFA4FFC8FEBCFE330D88FE5D" authority="Santos &amp; Dolibaina &amp; Dias &amp; Mielke &amp; Casagrande, 2023" authorityName="Santos &amp; Dolibaina &amp; Dias &amp; Mielke &amp; Casagrande" authorityYear="2023" box="[285,395,410,436]" class="Insecta" family="Riodinidae" genus="Alesa" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="species" species="mariae" status="sp. nov.">
<emphasis id="B9263265FFA4FFC8FEBCFE330D88FE5D" box="[285,395,410,436]" italics="true" pageId="21" pageNumber="98">A. mariae</emphasis>
</taxonomicName>
<emphasis id="B9263265FFA4FFC8FE33FE300DEFFE5D" bold="true" box="[402,492,409,435]" pageId="21" pageNumber="98">
<taxonomicNameLabel id="A2158F1EFFA4FFC8FE33FE300DEFFE5D" box="[402,492,409,435]" pageId="21" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C5295F4FFA4FFC8FD83FE330EB2FE5D" authorityName="Bates" authorityYear="1867" box="[546,689,409,436]" class="Insecta" family="Riodinidae" genus="Alesa" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="species" species="thelydrias">
<emphasis id="B9263265FFA4FFC8FD83FE330EB2FE5D" box="[546,689,409,436]" italics="true" pageId="21" pageNumber="98">A. thelydrias</emphasis>
</taxonomicName>
are unknown)
</paragraph>
<keyStep id="30A6F6D2FFA4FFC8FF36FE77099FFDFF" pageId="21" pageNumber="98">
<paragraph id="8BEDEE77FFA4FFC8FF36FE77099FFE1A" blockId="21.[151,1437,478,588]" box="[151,1436,478,500]" pageId="21" pageNumber="98">
<keyLead id="30A34D42FFA4FFC8FF36FE77099FFE1A" box="[151,1436,478,500]" pageId="21" pageNumber="98">
1 Dark blue iridescence at the center of the submarginal ocelli on the hindwing underside present...................
<taxonomicName id="4C5295F4FFA4FFC8FAFAFE76099FFE1A" baseAuthorityName="Cramer" baseAuthorityYear="1777" box="[1371,1436,479,500]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="species" species="amesis">
<emphasis id="B9263265FFA4FFC8FAFAFE76099FFE1A" box="[1371,1436,479,500]" italics="true" pageId="21" pageNumber="98">amesis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEDEE77FFA4FFC8FF36FE55099FFDFF" blockId="21.[151,1437,478,588]" box="[151,1436,507,529]" pageId="21" pageNumber="98">
<keyLead id="30A34D42FFA4FFC8FF36FE55099FFDFF" box="[151,1436,507,529]" pageId="21" pageNumber="98">1’ Dark blue iridescence at the center of the submarginal ocelli on the hindwing underside absent........................ 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A6F6D2FFA4FFC8FF36FDB0099FFDA5" pageId="21" pageNumber="98">
<paragraph id="8BEDEE77FFA4FFC8FF36FDB0099EFDC0" blockId="21.[151,1437,478,588]" box="[151,1437,537,559]" pageId="21" pageNumber="98">
<keyLead id="30A34D42FFA4FFC8FF36FDB0099EFDC0" box="[151,1437,537,559]" pageId="21" pageNumber="98">
2 Forewing length
<quantity id="4CAA4392FFA4FFC8FE27FDB00DFAFDC1" box="[390,505,537,559]" metricMagnitude="-2" metricUnit="m" metricValue="1.575" metricValueMax="1.65" metricValueMin="1.5" pageId="21" pageNumber="98" unit="mm" value="15.75" valueMax="16.5" valueMin="15.0">15–16,5mm</quantity>
........................................................................
<taxonomicName id="4C5295F4FFA4FFC8FAC3FDB0099EFDC0" authorityName="Bates" authorityYear="1867" box="[1378,1437,537,558]" class="Insecta" family="Riodinidae" genus="Alesa" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="species" species="lipara">
<emphasis id="B9263265FFA4FFC8FAC3FDB0099EFDC0" box="[1378,1437,537,558]" italics="true" pageId="21" pageNumber="98">lipara</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEDEE77FFA4FFC8FF36FD9F099FFDA5" blockId="21.[151,1437,478,588]" box="[151,1436,566,588]" pageId="21" pageNumber="98">
<keyLead id="30A34D42FFA4FFC8FF36FD9F099FFDA5" box="[151,1436,566,588]" pageId="21" pageNumber="98">
2’ Forewing length
<quantity id="4CAA4392FFA4FFC8FE27FD9F0DFAFDA2" box="[390,505,566,588]" metricMagnitude="-2" metricUnit="m" metricValue="1.475" metricValueMax="1.5" metricValueMin="1.45" pageId="21" pageNumber="98" unit="mm" value="14.75" valueMax="15.0" valueMin="14.5">14,5–15mm</quantity>
......................................................................
<taxonomicName id="4C5295F4FFA4FFC8FAE0FD9F099FFDA5" authorityName="Bates" authorityYear="1867" box="[1345,1436,566,587]" class="Insecta" family="Riodinidae" genus="Alesa" kingdom="Animalia" order="Lepidoptera" pageId="21" pageNumber="98" phylum="Arthropoda" rank="species" species="hemiurga">
<emphasis id="B9263265FFA4FFC8FAE0FD9F099FFDA5" box="[1345,1436,566,587]" italics="true" pageId="21" pageNumber="98">hemiurga</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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