A review of the South American metalmark genus Alesa Doubleday, 1847 (Lepidoptera: Riodinidae: Eurybiini) with the description of two new species Author Santos, Wildio Ikaro Da Graça 0000-0003-3466-5820 Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, PO Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil & wildio. ikaro @ gmail. com; https: // orcid. org / 0000 - 0003 - 3466 - 5820 wildio.ikaro@gmail.com Author Dolibaina, Diego Rodrigo 0000-0001-5258-3787 Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, PO Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil & Reserva Serra Bonita, PO Box 001, 45.880 - 970, Camacan, Bahia, Brazil & dirodrido @ gmail. com; https: // orcid. org / 0000 - 0001 - 5258 - 3787 dirodrido@gmail.com Author Dias, Fernando Maia Silva 0000-0002-7198-7792 Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, PO Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil & Departamento de Biologia Animal e Vegetal, Universidade Estadual de Londrina, PO Box 10011, 86.057 - 970, Londrina, Paraná, Brazil & fernandodias @ uel. br; https: // orcid. org / 0000 - 0002 - 7198 - 7792 fernandodias@uel.br Author Mielke, Olaf Hermann Hendrik 0000-0003-3655-4606 Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, PO Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil & omhesp @ ufpr. br; https: // orcid. org / 0000 - 0003 - 3655 - 4606 omhesp@ufpr.br Author Casagrande, Mirna Martins 0000-0002-6076-8463 Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, PO Box 19020, 81.531 - 980, Curitiba, Paraná, Brazil & mibras @ ufpr. br; https: // orcid. org / 0000 - 0002 - 6076 - 8463 mibras@ufpr.br text Zootaxa 2023 2023-05-10 5284 1 77 120 http://dx.doi.org/10.11646/zootaxa.5284.1.3 journal article 10.11646/zootaxa.5284.1.3 1175-5326 7919001 7334FB57-41A0-47CA-BCBF-FA108B13CEC1 “ amesis group” The “ amesis group” includes small-sized species and the two most common species of the genus ( A. amesis and A. lipara ). Males are characterized by the white tip of antennae; wings underside with a characteristic pattern; forewing with outer margin markedly projected in M 2 –M 3 , two distinct postdiscal spots in CuA 2 -2A on the forewing underside; base of valva ventrally large; outer posterior process of valva about as long as inner posterior process; distal margin of the outer posterior process of valva with pointed internal margin; inner posterior process of valva wider than the outer posterior process. Females have wings upper side yellow-spotted; sterigma membranous medially; base of lamella postvaginalis with a small, rounded projection at each side; with two lateral signa on the middle of the corpus bursae. Included species. A. amesis , A. mariae sp. nov. , A. lipara , A. hemiurga and A. thelydrias . Key for males from the “ amesis group” 1 Forewing upper side with light silvery intervenal rays........................................................ 2 1’ Forewing upper side without light silvery intervenal rays...................................................... 3 2 Wings upper side black without markings.............................................................. amesis 2’ Wings upper side pale brown with reddish markings............................................... mariae sp. nov. 3 Wings upper side with spots or bands..................................................................... 4 3’ Wings upper side without spots or bands............................................................... lipara 4 Wings upper side with purplish sheen............................................................... hemiurga 4’ Wings upper side without purplish sheen............................................................ thelydrias Key for females from the “ amesis group” (females of A. mariae sp. nov. and A. thelydrias are unknown) 1 Dark blue iridescence at the center of the submarginal ocelli on the hindwing underside present................... amesis 1’ Dark blue iridescence at the center of the submarginal ocelli on the hindwing underside absent........................ 2 2 Forewing length 15–16,5mm ........................................................................ lipara 2’ Forewing length 14,5–15mm ...................................................................... hemiurga