185 lines
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185 lines
20 KiB
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<document ID-CLB-Dataset="26640" ID-DOI="http://dx.doi.org/10.3897/phytokeys.122.33032" ID-GBIF-Dataset="f497907c-ee54-41cd-b666-7292c766b9c3" ID-PMC="PMC6548746" ID-Pensoft-Pub="1314-2003-122-29" ID-Pensoft-UUID="FFB7FFA5CF25AD00FFBDFFEBFFFCFFDD" ID-PubMed="31182907" ID-Zenodo-Dep="3238737" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-2003-122-29" ModsDocOrigin="PhytoKeys 122" ModsDocTitle="Páramo Calamagrostis s.l. (Poaceae): An updated list and key to the species known or likely to occur in páramos of NW South America and southern Central America including two new species, one new variety and five new records for Colombia" checkinTime="1559110536903" checkinUser="pensoft" docAuthor="Sylvester, Steven P., Soreng, Robert J., Bravo-Pedraza, William J., Cuta-Alarcon, Lia E., Giraldo-Canas, Diego, Aguilar-Cano, Jose & Peterson, Paul M." docDate="2019" docId="3BE31AF4AA5AC5A372B2B37BC98A1985" docLanguage="en" docName="PhytoKeys 122: 29-78" docOrigin="PhytoKeys 122" docSource="http://dx.doi.org/10.3897/phytokeys.122.33032" docTitle="Calamagrostis planifolia Trin. ex Steud., Nomencl. Bot. (ed. 2) 1: 251. 1840. Deyeuxia planifolia Kunth, Nov. Gen. Sp. (quarto ed.) 1: 145. 1815 [1816]. Arundo planifolia (Kunth) Poir., Encycl. 4: 707. 1816" docType="treatment" docVersion="8" id="FFB7FFA5CF25AD00FFBDFFEBFFFCFFDD" lastPageNumber="29" masterDocId="FFB7FFA5CF25AD00FFBDFFEBFFFCFFDD" masterDocTitle="Paramo Calamagrostis s. l. (Poaceae): An updated list and key to the species known or likely to occur in paramos of NW South America and southern Central America including two new species, one new variety and five new records for Colombia" masterLastPageNumber="78" masterPageNumber="29" pageNumber="29" updateTime="1732696349232" updateUser="ExternalLinkService">
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<mods:title id="288C12A0332B3204D495AE5C2C1E10F4">Paramo Calamagrostis s. l. (Poaceae): An updated list and key to the species known or likely to occur in paramos of NW South America and southern Central America including two new species, one new variety and five new records for Colombia</mods:title>
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<mods:namePart id="1E1F4F35911C2BD928151B670BEAE9AB">Sylvester, Steven P.</mods:namePart>
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<mods:namePart id="C5F9D4CEE821F78D07C6D6354F41B5F2">Soreng, Robert J.</mods:namePart>
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<mods:namePart id="7535BCA2CC922E6145A26B3AF4414E25">Bravo-Pedraza, William J.</mods:namePart>
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<mods:namePart id="7E84D8724EBF00F7B175A750C8FA9F7C">Cuta-Alarcon, Lia E.</mods:namePart>
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<mods:namePart id="332DB8FB257A5010EF77535074964EE6">Giraldo-Canas, Diego</mods:namePart>
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<mods:namePart id="E903D56AE59E4E4EA3E820A45D8874EF">Aguilar-Cano, Jose</mods:namePart>
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<mods:namePart id="855F53CAC69D2AF873A546D1ECB2F2A1">Peterson, Paul M.</mods:namePart>
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<mods:typeOfResource id="DB129DB6E75A5A6E31FF52500AE8D4CD">text</mods:typeOfResource>
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<mods:title id="0D1BF27B740545BA59C85213AD8FDF68">PhytoKeys</mods:title>
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<mods:date id="733EFA0A420E81FA7E19F74A8A835C21">2019</mods:date>
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<mods:detail id="A3C98761DC0E51FFFD27A0AEFBF3D6B3" type="volume">
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<mods:number id="B98C929E6C106B67D23EA2E8CB618FEF">122</mods:number>
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<mods:start id="68C7528B3282AEBE1FE4B5BFC371BFD7">29</mods:start>
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<mods:end id="B1C1D71D235877A2C428A99245E60189">78</mods:end>
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<mods:url id="A2A22A9D8270F8358AB97BFA006F8BF0">http://dx.doi.org/10.3897/phytokeys.122.33032</mods:url>
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<mods:classification id="9F1E9AF162362DE5A151A1ACDEECEF41">journal article</mods:classification>
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<mods:identifier id="19C316AF10BE4F969A3AA87A75073328" type="DOI">http://dx.doi.org/10.3897/phytokeys.122.33032</mods:identifier>
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<mods:identifier id="8D26AF48647BD82CC093910C63B21989" type="Pensoft-Pub">1314-2003-122-29</mods:identifier>
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<treatment id="3BE31AF4AA5AC5A372B2B37BC98A1985" ID-GBIF-Taxon="157294003" LSID="urn:lsid:plazi:treatment:3BE31AF4AA5AC5A372B2B37BC98A1985" httpUri="http://treatment.plazi.org/id/3BE31AF4AA5AC5A372B2B37BC98A1985" lastPageNumber="29" pageId="0" pageNumber="29" scope_family="Poaceae">
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<subSubSection id="DAC606D4B49DB2DEF04E9947D8D688A9" pageId="0" pageNumber="29" type="nomenclature">
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<paragraph id="D5BFB8D10F0027645890A2D50D9FCAA2" pageId="0" pageNumber="29">
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<taxonomicName id="AE78511555C52FC8230AE7984E49212F" ID-CoL="PCSJ" LSID="9BE76333-271A-5B6B-91BC-F9CD2DE40409" authority="(Kunth) Trin. ex Steud., Nomencl. Bot. (ed. 2) 1: 251. 1840. Deyeuxia planifolia Kunth, Nov. Gen. Sp. (quarto ed.) 1: 145. 1815 [1816]. Arundo planifolia (Kunth) Poir., Encycl. 4: 707. 1816." class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis planifolia" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="planifolia">Calamagrostis planifolia (Kunth) Trin. ex Steud., Nomencl. Bot. (ed. 2) 1: 251. 1840. Deyeuxia planifolia Kunth, Nov. Gen. Sp. (quarto ed.) 1: 145. 1815[1816]. Arundo planifolia (Kunth) Poir., Encycl. 4: 707. 1816.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="5C8A22F9A6AEA784AFEE407158D9CBDF" pageId="0" pageNumber="29" type="reference_group">
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<paragraph id="D71955320194DD54F939F4546E8507FA" pageId="0" pageNumber="29">
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<emphasis id="CA41D9384D4F110EF8773D0FD71CFF72" bold="true" pageId="0" pageNumber="29">Type.</emphasis>
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PERU. In montanis Andinum Peruvianorum prope Guangamarca, 1250 hexap. [2286 m], 1833, Bonpland s.n. (holotype: P (P00729787 [image!]); isotype: BAA (BAA00001855 [image!]) fragm. ex P, P (P030106 [image!]) fragm., LE (LE-TRIN-1801.01!) fragm. ex Herb. Willd. 176).
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</paragraph>
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<paragraph id="56E036FCCCE04327839CD8EFED4AB839" pageId="0" pageNumber="29">
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=
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<taxonomicName id="51B173753E47071F3A05BF3A8EDCF1EE" authorityName="Pilg" authorityYear="1898" class="Liliopsida" family="Poaceae" genus="Deyeuxia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Deyeuxia pubescens" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="pubescens">Deyeuxia pubescens</taxonomicName>
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Pilg., Bot. Jahrb. Syst. 25(5): 712. 1898, syn. nov.
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<taxonomicName id="D62406F201BB494814EFD5ECE14D6A4B" authorityName="Pilg" authorityYear="1908" baseAuthorityName="Pilg" class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis pubescens" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="pubescens">Calamagrostis pubescens</taxonomicName>
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(Pilg.) Pilg., Bot. Jahrb. Syst. 42: 60. 1908, syn. nov. Type: COLOMBIA. [Crescit in monte ignivomo Pasto]
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<normalizedToken id="DABE9E8F97E2C2BDE55F57E3A96F23D9" originalValue="Volcán">Volcan</normalizedToken>
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de Pasto, 3400 m alt., 9 Dec. 1869,
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<normalizedToken id="48093CCF7142A84B8C5635699C85BE27" originalValue="Stübel">Stuebel</normalizedToken>
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389b (lectotype, designated by
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<bibRefCitation id="9B770E191F99A5F1D4B3AFEFB9449841" author="Vega, AS" journalOrPublisher="Botanica" pageId="0" pageNumber="29" pagination="31 - 35" refId="B44" refString="Vega, AS, Rugolo de Agrasar, ZE, 2013. Lectotypifications in taxa of the genera Calamagrostis, Deyeuxia, and Digitaria (Poaceae). Gayana. Botanica 70 (1): 31 - 35" title="Lectotypifications in taxa of the genera Calamagrostis, Deyeuxia, and Digitaria (Poaceae). Gayana." volume="70" year="2013">
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Vega and
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<normalizedToken id="3DCBEA70D37C9C321B9A38F6C432EA8E" originalValue="Rúgolo">Rugolo</normalizedToken>
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de Agrasar (2013
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</bibRefCitation>
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: 28): BAA (BAA00000811 [image!]) fragm. ex B; isolectotype: US (US00406355!) fragm. ex B).
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</paragraph>
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<paragraph id="369412A8247FCF3ACAA7298AAF5C9445" pageId="0" pageNumber="29">
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=
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<taxonomicName id="36F1E6FA426FA340E30084916DE3F209" authorityName="Hack" authorityYear="1902" class="Liliopsida" family="Poaceae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis pittieri" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="pittieri">Calamagrostis pittieri</taxonomicName>
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Hack., Oesterr. Bot. Z. 52(3): 108. 1902, syn. nov. Type: COSTA RICA. Cerro de Buena Vista, pres du sommit, [prope cacumen, Valle du General], 3100 m alt., 19 Jan. 1891, Pittier s.n. Pl. Costaric. Exs. 3359 (holotype: W (W19160029198 [image!]); isotypes: B, BR (BR0000006865702 [image!]), BAA (BAA00000766 [image!]) fragm. ex B).
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</paragraph>
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</subSubSection>
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<subSubSection id="6B3704033E88810DE79F75A3BA2BD9B8" pageId="0" pageNumber="29" type="comments">
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<paragraph id="B005E5C5E60C6BDFE613FCA164B94555" pageId="0" pageNumber="29">Comments.</paragraph>
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<paragraph id="84921732853A6E7DD567EEF170AAE876" pageId="0" pageNumber="29">
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<taxonomicName id="FCFC8F004F1F919078434A6BB8B2F46C" authorityName="Hack" authorityYear="1902" class="Liliopsida" family="Gramineae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis pittieri" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="pittieri">
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<emphasis id="46EA0C27B70AAADB12132FBFD9ACD715" italics="true" pageId="0" pageNumber="29">Calamagrostis pittieri</emphasis>
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</taxonomicName>
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, recorded for Costa Rica, Colombia and Venezuela (
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<bibRefCitation id="91AC63AC865F36B3A01FF6142A8C3265" author="Morales-Quiros, JF" editor="Hammel, BE" journalOrPublisher="Monographs in Systematic Botany from the Missouri Botanical Garden 93 (3)" pageId="0" pageNumber="29" pagination="598 - 821" refId="B21" refString="Morales-Quiros, JF, 2003. Poaceae. In: Hammel, BE, Grayum, MH, Herrera-Mora, C, Zamora-Villalobos, N, Eds., Manual of Plants of Costa Rica. Monographs in Systematic Botany from the Missouri Botanical Garden 93(3): 598 - 821" title="Poaceae." volumeTitle="Manual of Plants of Costa Rica." year="2003">
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Morales
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<normalizedToken id="259BB8DA02858969D615D23D4ECB4A77" originalValue="Quirós">Quiros</normalizedToken>
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2003
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</bibRefCitation>
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;
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<bibRefCitation id="91FC0F68785AC37B13952C5F2A4C8930" author="Hokche, O" journalOrPublisher="Contributions from the United States National Herbarium" pageId="0" pageNumber="29" refId="B13" refString="Hokche, O, Berry, PE, Huber, O, 2008. (2008) Nuevo Catalogo de la Flora Vascular de Venezuela. Fundacion Instituto Botanico de Venezuela, Caracas, 1-859." title="(2008) Nuevo Catalogo de la Flora Vascular de Venezuela. Fundacion Instituto Botanico de Venezuela, Caracas, 1 - 859." year="2008">Hokche et al. 2008</bibRefCitation>
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;
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<bibRefCitation id="9EF73802ADC8E6387F00294BFBBB191E" author="Giraldo-Canas, D" journalOrPublisher="Darwiniana" pageId="0" pageNumber="29" pagination="139 - 247" refId="B9" refString="Giraldo-Canas, D, 2011. Catalogo de la familia Poaceae en Colombia. Darwiniana 49: 139 - 247" title="Catalogo de la familia Poaceae en Colombia." volume="49" year="2011">
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<normalizedToken id="B33B7959017E812E24626B68EFBF0530" originalValue="Giraldo-Cañas">Giraldo-Canas</normalizedToken>
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2011
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</bibRefCitation>
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,
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<bibRefCitation id="87A7A0AB4BE29CF89F7E1018264B1DB2" author="Giraldo-Canas, D" journalOrPublisher="Darwiniana" pageId="0" pageNumber="29" refId="B10" refString="Giraldo-Canas, D, 2013. Las gramineas en Colombia: riqueza, distribucion, endemismo, invasion, migracion, usos y taxonomias populares. Biblioteca Jose Jeronimo Triana No. 26, 1-382." title="Las gramineas en Colombia: riqueza, distribucion, endemismo, invasion, migracion, usos y taxonomias populares. Biblioteca Jose Jeronimo Triana No. 26, 1 - 382." year="2013">2013</bibRefCitation>
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;
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<bibRefCitation id="DFDCE39C5422AA7EA23E337D9601D838" author="Giraldo-Canas, D" editor="Bernal, R" journalOrPublisher="2. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota D. C." pageId="0" pageNumber="29" pagination="2127 - 2205" refId="B11" refString="Giraldo-Canas, D, Londono, X, Clark, LG, 2016. Poaceae. In: Bernal, R, Gradstein, SR, Celis, M, Eds., Catalogo de plantas y liquenes de Colombia, vol. 2. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogota D.C.: 2127 - 2205" title="Poaceae." volumeTitle="Catalogo de plantas y liquenes de Colombia, vol." year="2016">
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<normalizedToken id="205FB27F1991A14BDB9A4990D81CF0ED" originalValue="Giraldo-Cañas">Giraldo-Canas</normalizedToken>
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et al. 2016
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</bibRefCitation>
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) is synonymised under
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<taxonomicName id="4ABAC474D32E34BD00A27D9DAA94C383" class="Liliopsida" family="Gramineae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis planifolia" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="planifolia">
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<emphasis id="7983981CAB28970CEACD6F2154F084C7" italics="true" pageId="0" pageNumber="29">Calamagrostis planifolia</emphasis>
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</taxonomicName>
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as no satisfactory characteristics were found to separate the two. Both have spikelets with florets bearing a pilose rachilla extension with hairs reaching from
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<normalizedToken id="15B8F4F70F625D1507F0C7351C2840FD" originalValue="¾">3/4</normalizedToken>
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to passing the apex of the lemma, an awn inserted in the upper half of the lemma, 2 anthers that usually are short, ca. 1-1.4 mm long and leaf blades that are usually flat (sometimes drying convolute) with variable indumentum.
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</paragraph>
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<paragraph id="A823D12BC5428B0AC3C69A588909D864" pageId="0" pageNumber="29">
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<taxonomicName id="1065D0C85CDB69D8B48F8052A8A26A5B" authorityName="Pilg" authorityYear="1908" baseAuthorityName="Pilg" class="Liliopsida" family="Gramineae" genus="Calamagrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Calamagrostis pubescens" order="Poales" pageId="0" pageNumber="29" phylum="Tracheophyta" rank="species" species="pubescens">
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<emphasis id="320F178F5E09FC56E1CB07A7DFA58F82" italics="true" pageId="0" pageNumber="29">Calamagrostis pubescens</emphasis>
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</taxonomicName>
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was considered an endemic species to Colombia (
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<bibRefCitation id="B3673662D532BC5B9F824ED867C19FDD" author="Giraldo-Canas, D" journalOrPublisher="Darwiniana" pageId="0" pageNumber="29" refId="B10" refString="Giraldo-Canas, D, 2013. Las gramineas en Colombia: riqueza, distribucion, endemismo, invasion, migracion, usos y taxonomias populares. Biblioteca Jose Jeronimo Triana No. 26, 1-382." title="Las gramineas en Colombia: riqueza, distribucion, endemismo, invasion, migracion, usos y taxonomias populares. Biblioteca Jose Jeronimo Triana No. 26, 1 - 382." year="2013">
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Giraldo-
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<normalizedToken id="C29E0C5ABED0719186BDB2327DA44E93" originalValue="Cañas">Canas</normalizedToken>
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2013
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</bibRefCitation>
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) and only known from the type specimen that was collected in southern Colombia from hills surrounding Pasto of departamento Narino. The B holotype was destroyed during World War II, with fragments only remaining at BAA and US. The BAA fragment consists of spikelets whose size and characteristics match
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<taxonomicName id="16D9E636CF246E779C1C010AE37D517F" class="Monocotyledoneae" family="Gramineae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. planifolia" order="Poales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="planifolia">
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<emphasis id="09101BD8AF83CDACC9CAE0E0AFBAB71D" italics="true" pageId="0" pageNumber="29">C. planifolia</emphasis>
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</taxonomicName>
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(Zulma
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<normalizedToken id="26D78D199C16B75A5E38DCCF013A9107" originalValue="Rúgolo">Rugolo</normalizedToken>
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de Agrasar, pers. comm.), and the description of vegetative characteristics in the protologue also matches
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<taxonomicName id="3DAEA26E7B5C0E8B8E47C37DA3A3BB57" class="Monocotyledoneae" family="Gramineae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. planifolia" order="Poales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="planifolia">
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<emphasis id="5E61424C47755F29720090700CFE41FF" italics="true" pageId="0" pageNumber="29">C. planifolia</emphasis>
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</taxonomicName>
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|
and so we consider
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|
<taxonomicName id="CF38F1402D707528BEA5C73BC890311D" class="Monocotyledoneae" family="Gramineae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. pubescens" order="Poales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="pubescens">
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|
<emphasis id="9E6CC53FB51DFE667000127A493762C2" italics="true" pageId="0" pageNumber="29">C. pubescens</emphasis>
|
|
</taxonomicName>
|
|
a synonym of
|
|
<taxonomicName id="447ABABE41AD31688BFE8F02BB2E54BC" class="Monocotyledoneae" family="Gramineae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. planifolia" order="Poales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="planifolia">
|
|
<emphasis id="4213FA47559F11C9095E3ECED751A991" italics="true" pageId="0" pageNumber="29">C. planifolia</emphasis>
|
|
</taxonomicName>
|
|
. Nevertheless, further study is needed for the
|
|
<taxonomicName id="0A4E61BADF540BA68636C63BE649E8D5" class="Monocotyledoneae" family="Gramineae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. planifolia" order="Poales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="planifolia">
|
|
<emphasis id="12F9A2BBA8F14C527590BA16BA929964" italics="true" pageId="0" pageNumber="29">C. planifolia</emphasis>
|
|
</taxonomicName>
|
|
complex in Colombia and there are certain crucial characters, such as number of stamens that need to be verified in these taxa. The
|
|
<taxonomicName id="91FD52A808F542F353DE17FF2DC9EB44" class="Monocotyledoneae" family="Gramineae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="C. pubescens" order="Poales" pageId="0" pageNumber="29" phylum="Angiospermae" rank="species" species="pubescens">
|
|
<emphasis id="27820490C2A962FE5811824BF2930A0E" italics="true" pageId="0" pageNumber="29">C. pubescens</emphasis>
|
|
</taxonomicName>
|
|
BAA fragment lacked anthers (Zulma
|
|
<normalizedToken id="4C1284AC1F5787E134DDDBFDC388127F" originalValue="Rúgolo">Rugolo</normalizedToken>
|
|
de Agrasar, pers. comm.) and
|
|
<bibRefCitation id="7A0587C7943A41260279D1F247DFE4BD" author="Pilger, RKF" journalOrPublisher="Botanische Jahrbuecher fuer Systematik, Pflanzengeschichte und Pflanzengeographie" pageId="0" pageNumber="29" pagination="709 - 721" refId="B26" refString="Pilger, RKF, 1898. Plantae Stuebelianae novae, Gramineae. Botanische Jahrbuecher fuer Systematik, Pflanzengeschichte und Pflanzengeographie 25 (5): 709 - 721" title="Plantae Stuebelianae novae, Gramineae." volume="25" year="1898">Pilger (1898</bibRefCitation>
|
|
: 712) did not describe them.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |