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<mods:title id="A7FA6A63A97F1C5F9DC056347ED960C6">A reassessment of the Neotropical genus Pseudonannolene Silvestri, 1895: cladistic analysis, biogeography, and taxonomic review (Spirostreptida: Pseudonannolenidae)</mods:title>
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<mods:namePart id="8D3D8E75E107C5541C40DE71B6A3DCD0">Iniesta, Luiz Felipe Moretti</mods:namePart>
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<mods:namePart id="B3607F474B6DB72D6B4728885EFA10F9">Bouzan, Rodrigo Salvador</mods:namePart>
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<mods:namePart id="3535AB48D7C8D186DD06D0CF110ECC72">Brescovit, Antonio Domingos</mods:namePart>
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<emphasis id="B925EABE1326B1744E87FE37FCD854C8" bold="true" box="[465,834,475,502]" italics="true" pageId="21" pageNumber="22">Pseudonannolene ambuatinga</emphasis>
Iniesta &amp; Ferreira, 2013
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<figureCitation id="136A2A291326B1744E33FE13FE765727" box="[357,492,511,537]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="160.[189,232,768,794]" captionStartId-1="161.[189,232,1668,1694]" captionTargetBox-0="[189,1398,285,725]" captionTargetBox-1="[189,1398,269,1626]" captionTargetId-0="figure-13@160.[189,1398,273,725]" captionTargetId-1="figure-11@161.[189,1398,269,1626]" captionTargetPageId-0="160" captionTargetPageId-1="161" captionText-0="Fig. 46. Pseudonannolene ambuatinga Iniesta &amp; Ferreira, 2013, paratype, ♀ (ISLA 2274), in lateral view. A. Anterior region. B. Posterior region. Scale bars: A = 0.5 mm; B = 1 mm." captionText-1="Fig. 47. Pseudonannolene ambuatinga Iniesta &amp; Ferreira, 2013, ♂ (IBSP 3442). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7891139" figureDoi-1="http://doi.org/10.5281/zenodo.7891141" httpUri-0="https://zenodo.org/record/7891139/files/figure.png" httpUri-1="https://zenodo.org/record/7891141/files/figure.png" pageId="21" pageNumber="22">Figs 46–47</figureCitation>
, 
<figureCitation id="136A2A291326B1744EA1FE12FDA15727" box="[503,571,510,537]" captionStart="Fig" captionStartId="268.[189,232,1741,1767]" captionTargetBox="[348,1239,265,1699]" captionTargetId="figure-13@268.[348,1239,265,1699]" captionTargetPageId="268" captionText="Fig. 165. Stereoscopic images of midbody legs of species of Pseudonannolene Silvestri, 1895. A. P. ambuatinga Iniesta &amp; Ferreira, 2013. B. P. anapophysis Fontanetti, 1996. C. P. alata sp. nov. D. P. bovei Silvestri, 1895. E. P. buhrnheimi Schubart, 1960. F. P. caatinga Iniesta &amp; Ferreira, 2014. G. P. callipyge Brölemann, 1902. H. P. curtipes Schubart, 1960. I. P. fontanettiae Iniesta &amp; Ferreira, 2014. J. P. halophila Schubart, 1949. K. P. imbirensis Fontanetti, 1996. L. P. inops Brölemann, 1929. M, P. leopoldoi Iniesta &amp; Ferreira, 2014. N. P. leucomelas Schubart, 1947. O. P. longicornis (Porat, 1888). P. P. lundi Iniesta &amp; Ferreira, 2015. Q. P. magna Udulutsch &amp; Pietrobon, 2003. R. P. maritima Schubart, 1949. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891411" httpUri="https://zenodo.org/record/7891411/files/figure.png" pageId="21" pageNumber="22">165A</figureCitation>
, 
<figureCitation id="136A2A291326B1744D11FE13FD135727" box="[583,649,511,537]" captionStart="Fig" captionStartId="270.[189,232,1892,1918]" captionTargetBox="[213,1374,265,1851]" captionTargetId="figure-14@270.[213,1374,265,1851]" captionTargetPageId="270" captionText="Fig. 167. Stereoscopic images of gnathochilarium and anterior sternum of species of Pseudonannolene Silvestri, 1895. A. P. albiventris Schubart, 1952. B. P. ambuatinga Iniesta &amp; Ferreira, 2013. C. P. anapophysis Fontanetti, 1996. D. P. buhrnheimi Schubart, 1960. E. P. caatinga Iniesta &amp; Ferreira, 2014. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891417" httpUri="https://zenodo.org/record/7891417/files/figure.png" pageId="21" pageNumber="22">167B</figureCitation>
, 
<figureCitation id="136A2A291326B1744DC3FE13FD425727" box="[661,728,511,537]" captionStart="Fig" captionStartId="280.[189,232,1787,1813]" captionTargetBox="[250,1337,265,1739]" captionTargetId="figure-14@280.[250,1337,265,1739]" captionTargetPageId="280" captionText="Fig. 177. Stereoscopic images of left vulva of species of Pseudonannolene Silvestri, 1895, in oral view. A. P. albiventris Schubart, 1952. B. P. alegrensis Silvestri, 1897. C. P. ambuatinga Iniesta &amp; Ferreira, 2013. D. P. anapophysis Fontanetti, 1996. E. P. buhrnheimi Schubart, 1960. F. P. caatinga Iniesta &amp; Ferreira, 2014. G. P. callipyge Brölemann, 1902. H. P. erikae Iniesta &amp; Ferreira, 2014. I. P. fontanettiae Iniesta &amp; Ferreira, 2014. J. P. halophila Schubart, 1949. K. P. imbirensis Fontanetti, 1996. L. P. inops Brölemann, 1929. M. P. leucocephalus Schubart, 1944. N. P. longicornis (Porat, 1888). O. P. lundi Iniesta &amp; Ferreira, 2015. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891439" httpUri="https://zenodo.org/record/7891439/files/figure.png" pageId="21" pageNumber="22">177C</figureCitation>
, 
<figureCitation id="136A2A291326B1744DB2FE13FC895727" box="[740,787,511,537]" captionStart="Fig" captionStartId="283.[189,232,1363,1389]" captionTargetBox="[192,1368,285,1297]" captionTargetId="figure-12@283.[189,1398,265,1328]" captionTargetPageId="283" captionText="Fig. 180. Distribution map of the species Pseudonannolene albiventris Schubart, 1952, P. alegrensis Silvestri, 1897, P. ambuatinga Iniesta &amp; Ferreira, 2013, P. anapophysis Fontanetti, 1996, and P. borelli Silvestri, 1895." figureDoi="http://doi.org/10.5281/zenodo.7891445" httpUri="https://zenodo.org/record/7891445/files/figure.png" pageId="21" pageNumber="22">180</figureCitation>
; Supp. file 4: 
<figureCitation id="136A2A291326B1744CE3FE13FBA85727" box="[949,1074,510,537]" captionStart="Fig" captionStartId="310.[242,285,1160,1186]" captionText="Fig. 215. Character 69 and its character states. Telopodites. A. Pseudonannolene ambuatinga Iniesta &amp; Ferreira, 2013 (IBSP 3442). B. P. alata sp. nov. (IBSP 7879). Scale bars = 0.2 mm." pageId="21" pageNumber="22">Figs 215A</figureCitation>
, 
<figureCitation id="136A2A291326B1744B69FE13FB195727" box="[1087,1155,511,537]" captionStart="Fig" captionStartId="311.[242,285,265,291]" captionText="Fig. 220. Characters 83–84 and their character states. Gonopods. A. Epinannolene sp. (ICN). B. Pseudonannolene typica Silvestri, 1895 (MCSN). C. Phallorthus colombianus Chamberlin, 1952 (FMNH). D. P. ambuatinga Iniesta &amp; Ferreira, 2013 (IBSP 3442). Scale bars: A = 0.5 mm; B = 0.1 mm; C–D = 0.2 mm." pageId="21" pageNumber="22">220D</figureCitation>
, 
<figureCitation id="136A2A291326B1744BD9FE13FB545727" box="[1167,1230,511,537]" captionStart="Fig" captionStartId="311.[242,285,572,598]" captionText="Fig. 222. Characters 86–90 and their character states. Internal branch. A. Phallorthus colombianus Chamberlin, 1952 (FMNH). B. Pseudonannolene ambuatinga Iniesta &amp; Ferreira, 2013 (IBSP 3442). C. Epinannolene exilio (Brölemann, 1904) (INPA). D. P. robsoni Iniesta &amp; Ferreira, 2014 (IBSP 3441). E. P. occidentalis Schubart, 1958 (IBSP 1998). F. P. typica Silvestri, 1895 (MCSN). SEM image of distal region of the internal branch, P. fontanettiae Iniesta &amp; Ferreira, 2014 (IBSP), in detail (distal projection highlighted in red). Scale bars = 0.2 mm." pageId="21" pageNumber="22">222B</figureCitation>
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<emphasis id="B925EABE1326B1744FEBFDD3FDBB5767" box="[189,545,575,601]" italics="true" pageId="21" pageNumber="22">Pseudonannolene ambuatinga</emphasis>
<bibRefCitation id="EFC04B5D1326B1744D7EFDD3FC155764" author="Iniesta L. F. &amp; Ferreira R. L." box="[552,911,575,602]" pageId="21" pageNumber="22" pagination="357 - 369" refId="ref103738" refString="Iniesta L. F. M &amp; Ferreira R. L. 2013 b. Two new species of Pseudonannolene Silvestri, 1895 from Brazilian limestone caves (Spirostreptida: Pseudonannolenidae): synotopy of a troglophilic and a troglobiotic species. Zootaxa 3702 (4): 357 - 369. https: // doi. org / 10.11646 / zootaxa. 3702.4.3" type="journal article" year="2013">Iniesta &amp; Ferreira, 2013b: 358</bibRefCitation>
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, figs 1–6.
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<emphasis id="B925EABE1326B1744FEBFD8EFD9B5742" box="[189,513,610,636]" italics="true" pageId="21" pageNumber="22">Pseudonannolene saguassu</emphasis>
<bibRefCitation id="EFC04B5D1326B1744D5EFD8EFCF55742" author="Iniesta L. F. &amp; Ferreira R. L." box="[520,879,610,636]" pageId="21" pageNumber="22" pagination="357 - 369" refId="ref103738" refString="Iniesta L. F. M &amp; Ferreira R. L. 2013 b. Two new species of Pseudonannolene Silvestri, 1895 from Brazilian limestone caves (Spirostreptida: Pseudonannolenidae): synotopy of a troglophilic and a troglobiotic species. Zootaxa 3702 (4): 357 - 369. https: // doi. org / 10.11646 / zootaxa. 3702.4.3" type="journal article" year="2013">Iniesta &amp; Ferreira, 2013b: 363</bibRefCitation>
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, figs 7–10. 
<emphasis id="B925EABE1326B1744CA3FD8EFBFD5742" bold="true" box="[1013,1127,610,636]" pageId="21" pageNumber="22">Syn. nov.</emphasis>
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<treatmentCitation id="0AF010BD1326B1744FEBFD69FCC957A1" LSID="urn:lsid:zoobank.org:act:D1CDA013-785D-4800-A746-87BF93FE48C8" author="Gallo J. S. &amp; Bichuette M. E." box="[189,851,645,671]" httpUri="http://treatment.plazi.org/id/ACC03CDBD3AC5FBFAAED7FAB7CDD12DB" page="37" pageId="21" pageNumber="22" year="2020">
<taxonomicName id="4C514D2F1326B1744FEBFD69FCC957A1" ID-CoL="0eefd911-79bd-466a-b097-02dda89aa3ab" authority="Gallo &amp; Bichuette, 2020: 37" authorityName="Gallo &amp; Bichuette" authorityPageNumber="37" authorityYear="2020" box="[189,851,645,671]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="canastra">
<emphasis id="B925EABE1326B1744FEBFD69FE6657A1" box="[189,508,645,671]" italics="true" pageId="21" pageNumber="22">Pseudonannolene canastra</emphasis>
<bibRefCitation id="EFC04B5D1326B1744D55FD69FCC957A1" author="Gallo J. S. &amp; Bichuette M. E." box="[515,851,645,671]" pageId="21" pageNumber="22" pagination="33 - 47" refId="ref102297" refString="Gallo J. S. &amp; Bichuette M. E. 2020. Pseudonannolene canastra sp. nov. (Diplopoda, Spirostreptida) - A new troglobitic millipede from the southwestern state of Minas Gerais, Brazil. Subterranean Biology 35: 33 - 47. https: // doi. org / 10.3897 / subtbiol. 35.51183" type="journal article" year="2020">Gallo &amp; Bichuette, 2020: 37</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 3–6. 
<emphasis id="B925EABE1326B1744C9DFD69FBA657A1" bold="true" box="[971,1084,645,671]" pageId="21" pageNumber="22">Syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AC1326B1744FEBFD2AFD49563D" blockId="21.[189,1398,710,806]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB4111821326B1744FEBFD2AFD49563D" pageId="21" pageNumber="22">
<taxonomicName id="4C514D2F1326B1744FEBFD2AFDB857DE" ID-CoL="5f971c04-42f3-4d35-a8b2-a9ea67eab5ff" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[189,546,710,736]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ambuatinga">
<emphasis id="B925EABE1326B1744FEBFD2AFDB857DE" box="[189,546,710,736]" italics="true" pageId="21" pageNumber="22">Pseudonannolene ambuatinga</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0AF010BD1326B1744D16FD2AFC0F57DE" author="Iniesta L. F. &amp; Ferreira R. L." box="[576,917,710,736]" page="363" pageId="21" pageNumber="22" year="2014">
<bibRefCitation id="EFC04B5D1326B1744D16FD2AFC0F57DE" author="Iniesta L. F. &amp; Ferreira R. L." box="[576,917,710,736]" pageId="21" pageNumber="22" pagination="361 - 397" refId="ref103858" refString="Iniesta L. F. M &amp; Ferreira R. L. 2014. New species of Pseudonannolene Silvestri, 1895 from Brazilian limestone caves with comments on the potential distribution of the genus in South America (Spirostreptida: Pseudonannolenidae). Zootaxa 3846 (3): 361 - 397. https: // doi. org / 10.11646 / zootaxa. 3846.3.3" type="journal article" year="2014">Iniesta &amp; Ferreira 2014: 363</bibRefCitation>
</treatmentCitation>
. — 
<bibRefCitation id="EFC04B5D1326B1744C90FD2AFA9F57DE" author="Karam-Gemael M. &amp; Izzo T. J. &amp; Chagas-Jr A." box="[966,1285,710,736]" pageId="21" pageNumber="22" pagination="255 - 269" refId="ref104442" refString="Karam-Gemael M., Izzo T. J. &amp; Chagas-Jr A. 2018. Why be red listed? Threatened Myriapoda species in Brazil with implications for their conservation. ZooKeys 741: 255 - 269. https: // doi. org / 10.3897 / zookeys. 741.21971" type="journal article" year="2018">
Karam-Gemael 
<emphasis id="B925EABE1326B1744BD5FD2BFB2557DE" box="[1155,1215,710,736]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
2018
</bibRefCitation>
: figs 2–3. — 
<treatmentCitation id="0AF010BD1326B1744E59FD04FDC3563D" author="Gallo J. S. &amp; Bichuette M. E." box="[271,601,744,771]" page="42" pageId="21" pageNumber="22" year="2019">
<bibRefCitation id="EFC04B5D1326B1744E59FD04FDC3563D" author="Gallo J. S. &amp; Bichuette M. E." box="[271,601,744,771]" pageId="21" pageNumber="22" pagination="41 - 55" refId="ref102230" refString="Gallo J. S. &amp; Bichuette M. E. 2019. O que mudou na distribuicao dos diplopodes Pseudonannolene Silvestri, 1895 nas cavernas do Brasil 18 anos apos a sinopse de Trajano e colaboradores (2000)?. Espeleo-Tema 29 (1): 41 - 55. https: // doi. org / 10.3897 / aca. 1. e 30225" type="journal article" year="2019">Gallo &amp; Bichuette 2019: 42</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="0AF010BD1326B1744D32FD05FD4A563D" author="Gallo J. S. &amp; Bichuette M. E." box="[612,720,745,771]" page="34" pageId="21" pageNumber="22" year="2020">
<bibRefCitation id="EFC04B5D1326B1744D32FD05FD4A563D" author="Gallo J. S. &amp; Bichuette M. E." box="[612,720,745,771]" pageId="21" pageNumber="22" pagination="33 - 47" refId="ref102297" refString="Gallo J. S. &amp; Bichuette M. E. 2020. Pseudonannolene canastra sp. nov. (Diplopoda, Spirostreptida) - A new troglobitic millipede from the southwestern state of Minas Gerais, Brazil. Subterranean Biology 35: 33 - 47. https: // doi. org / 10.3897 / subtbiol. 35.51183" type="journal article" year="2020">2020: 34</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEE36AC1326B1744FEBFCE7FCF1561B" blockId="21.[189,1398,710,806]" box="[189,875,779,806]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB4111821326B1744FEBFCE7FCF1561B" box="[189,875,779,806]" pageId="21" pageNumber="22">
<taxonomicName id="4C514D2F1326B1744FEBFCE7FD9B561B" ID-CoL="a9792bb3-b08c-4010-89c2-9b3a6730c895" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[189,513,779,805]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="saguassu">
<emphasis id="B925EABE1326B1744FEBFCE7FD9B561B" box="[189,513,779,805]" italics="true" pageId="21" pageNumber="22">Pseudonannolene saguassu</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0AF010BD1326B1744D48FCE7FCFD561B" author="Gallo J. S. &amp; Bichuette M. E." box="[542,871,779,805]" page="48" pageId="21" pageNumber="22" year="2019">
<bibRefCitation id="EFC04B5D1326B1744D48FCE7FCFD561B" author="Gallo J. S. &amp; Bichuette M. E." box="[542,871,779,805]" pageId="21" pageNumber="22" pagination="41 - 55" refId="ref102230" refString="Gallo J. S. &amp; Bichuette M. E. 2019. O que mudou na distribuicao dos diplopodes Pseudonannolene Silvestri, 1895 nas cavernas do Brasil 18 anos apos a sinopse de Trajano e colaboradores (2000)?. Espeleo-Tema 29 (1): 41 - 55. https: // doi. org / 10.3897 / aca. 1. e 30225" type="journal article" year="2019">Gallo &amp; Bichuette 2019: 48</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34B65271326B1744FEBFCA0FDAB5194" pageId="21" pageNumber="22" type="description">
<paragraph id="8BEE36AC1326B1744FEBFCA0FE665658" blockId="21.[189,508,844,870]" box="[189,508,844,870]" pageId="21" pageNumber="22">
<heading id="D0A681C01326B1744FEBFCA0FE665658" bold="true" box="[189,508,844,870]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B925EABE1326B1744FEBFCA0FE665658" bold="true" box="[189,508,844,870]" pageId="21" pageNumber="22">Justification of synonymy</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1326B1744FEBFC96FDAB5194" blockId="21.[189,1399,890,1194]" pageId="21" pageNumber="22">
Having studied the original descriptions, closely examined the 
<typeStatus id="54EA880E1326B1744C9DFC97FC6456AB" box="[971,1022,891,917]" pageId="21" pageNumber="22">type</typeStatus>
species of 
<taxonomicName id="4C514D2F1326B1744BC7FC97FAA556AA" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[1169,1343,890,916]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ambuatinga">
<emphasis id="B925EABE1326B1744BC7FC97FAA556AA" box="[1169,1343,890,916]" italics="true" pageId="21" pageNumber="22">P. ambuatinga</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C514D2F1326B1744FEBFC71FED35688" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[189,329,925,950]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="saguassu">
<emphasis id="B925EABE1326B1744FEBFC71FED35688" box="[189,329,925,950]" italics="true" pageId="21" pageNumber="22">P. saguassu</emphasis>
</taxonomicName>
and topotypes from caves in the Arcos-Pains-Doresópolis speleological unit, the species 
<taxonomicName id="4C514D2F1326B1744FEBFC2CFED356E7" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[189,329,960,985]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="saguassu">
<emphasis id="B925EABE1326B1744FEBFC2CFED356E7" box="[189,329,960,985]" italics="true" pageId="21" pageNumber="22">P. saguassu</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C514D2F1326B1744EDEFC2CFD9556E7" authorityName="Gallo &amp; Bichuette" authorityYear="2020" box="[392,527,960,985]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="canastra">
<emphasis id="B925EABE1326B1744EDEFC2CFD9556E7" box="[392,527,960,985]" italics="true" pageId="21" pageNumber="22">P. canastra</emphasis>
</taxonomicName>
are here treated as junior synonyms of 
<taxonomicName id="4C514D2F1326B1744CA0FC2CFB3B56E7" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[1014,1185,959,985]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ambuatinga">
<emphasis id="B925EABE1326B1744CA0FC2CFB3B56E7" box="[1014,1185,959,985]" italics="true" pageId="21" pageNumber="22">P. ambuatinga</emphasis>
</taxonomicName>
, according to the similarities in gonopod structure (telopodite and internal branch) and first leg-pair of males. Regarding the species 
<taxonomicName id="4C514D2F1326B1744E11FBE9FE505120" authorityName="Gallo &amp; Bichuette" authorityYear="2020" box="[327,458,1029,1054]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="canastra">
<emphasis id="B925EABE1326B1744E11FBE9FE505120" box="[327,458,1029,1054]" italics="true" pageId="21" pageNumber="22">P. canastra</emphasis>
</taxonomicName>
, the males described correspond to immatures due to the prefemoral process of first leg-pair being still incipient, short and with few and scattered setae (
<bibRefCitation id="EFC04B5D1326B1744B75FBCBFAE8517F" author="Gallo J. S. &amp; Bichuette M. E." box="[1059,1394,1063,1089]" pageId="21" pageNumber="22" pagination="33 - 47" refId="ref102297" refString="Gallo J. S. &amp; Bichuette M. E. 2020. Pseudonannolene canastra sp. nov. (Diplopoda, Spirostreptida) - A new troglobitic millipede from the southwestern state of Minas Gerais, Brazil. Subterranean Biology 35: 33 - 47. https: // doi. org / 10.3897 / subtbiol. 35.51183" type="journal article" year="2020">Gallo &amp; Bichuette 2020: 37</bibRefCitation>
, fig. 3d–e), and the gonopod not fully developed, mainly the internal branch and gonocoxa (
<bibRefCitation id="EFC04B5D1326B1744A44FBA6FE3D51B9" author="Gallo J. S. &amp; Bichuette M. E." pageId="21" pageNumber="22" pagination="33 - 47" refId="ref102297" refString="Gallo J. S. &amp; Bichuette M. E. 2020. Pseudonannolene canastra sp. nov. (Diplopoda, Spirostreptida) - A new troglobitic millipede from the southwestern state of Minas Gerais, Brazil. Subterranean Biology 35: 33 - 47. https: // doi. org / 10.3897 / subtbiol. 35.51183" type="journal article" year="2020">Gallo &amp; Bichuette 2020: 37</bibRefCitation>
, fig. 6a–d) (see previous sections for more details on gonopod morphology and ontogeny in 
<taxonomicName id="4C514D2F1326B1744E06FB63FDBB5197" authorityName="Silvestri" authorityYear="1895" box="[336,545,1167,1193]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B925EABE1326B1744E06FB63FDBB5197" box="[336,545,1167,1193]" italics="true" pageId="21" pageNumber="22">Pseudonannolene</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B65271326B1744FEBFB3CFD0550BF" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="8BEE36AC1326B1744FEBFB3CFEAC51D4" blockId="21.[189,310,1232,1258]" box="[189,310,1232,1258]" pageId="21" pageNumber="22">
<heading id="D0A681C01326B1744FEBFB3CFEAC51D4" bold="true" box="[189,310,1232,1258]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B925EABE1326B1744FEBFB3CFEAC51D4" bold="true" box="[189,310,1232,1258]" pageId="21" pageNumber="22">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1326B1744FEBFB12FD0550BF" blockId="21.[188,1400,1278,1409]" pageId="21" pageNumber="22">
Resembling 
<taxonomicName id="4C514D2F1326B1744E05FB13FE345026" authorityName="Iniesta &amp; Ferreira" authorityYear="2015" box="[339,430,1278,1304]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="lundi">
<emphasis id="B925EABE1326B1744E05FB13FE345026" box="[339,430,1278,1304]" italics="true" pageId="21" pageNumber="22">P. lundi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C514D2F1326B1744EB8FB13FDFD5026" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[494,615,1278,1304]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="spelaea">
<emphasis id="B925EABE1326B1744EB8FB13FDFD5026" box="[494,615,1278,1304]" italics="true" pageId="21" pageNumber="22">P. spelaea</emphasis>
</taxonomicName>
by having head, trunk, and legs depigmented (
<figureCitation id="136A2A291326B1744BC8FB12FB635027" box="[1182,1273,1278,1305]" captionStart="Fig" captionStartId="160.[189,232,768,794]" captionTargetBox="[189,1398,285,725]" captionTargetId="figure-13@160.[189,1398,273,725]" captionTargetPageId="160" captionText="Fig. 46. Pseudonannolene ambuatinga Iniesta &amp; Ferreira, 2013, paratype, ♀ (ISLA 2274), in lateral view. A. Anterior region. B. Posterior region. Scale bars: A = 0.5 mm; B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891139" httpUri="https://zenodo.org/record/7891139/files/figure.png" pageId="21" pageNumber="22">Fig. 46</figureCitation>
). Males of 
<taxonomicName id="4C514D2F1326B1744FE9FACEFEFD5005" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[191,359,1313,1339]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ambuatinga">
<emphasis id="B925EABE1326B1744FE9FACEFEFD5005" box="[191,359,1313,1339]" italics="true" pageId="21" pageNumber="22">P. ambuatinga</emphasis>
</taxonomicName>
differ from 
<taxonomicName id="4C514D2F1326B1744EA5FACEFDD15005" authorityName="Iniesta &amp; Ferreira" authorityYear="2015" box="[499,587,1313,1339]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="lundi">
<emphasis id="B925EABE1326B1744EA5FACEFDD15005" box="[499,587,1313,1339]" italics="true" pageId="21" pageNumber="22">P. lundi</emphasis>
</taxonomicName>
by having a subtriangular solenomere (
<figureCitation id="136A2A291326B1744B4CFACDFB1E5005" box="[1050,1156,1313,1339]" captionStart="Fig" captionStartId="161.[189,232,1668,1694]" captionTargetBox="[189,1398,269,1626]" captionTargetId="figure-11@161.[189,1398,269,1626]" captionTargetPageId="161" captionText="Fig. 47. Pseudonannolene ambuatinga Iniesta &amp; Ferreira, 2013, ♂ (IBSP 3442). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891141" httpUri="https://zenodo.org/record/7891141/files/figure.png" pageId="21" pageNumber="22">Fig. 47D</figureCitation>
) instead of a square-shaped square-shaped solenomere, and from 
<taxonomicName id="4C514D2F1326B1744D9CFAA8FCDA5063" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[714,832,1347,1373]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="spelaea">
<emphasis id="B925EABE1326B1744D9CFAA8FCDA5063" box="[714,832,1347,1373]" italics="true" pageId="21" pageNumber="22">P. spelaea</emphasis>
</taxonomicName>
by having seminal apophysis evident and by the number of ommatidia (ca 25) (
<figureCitation id="136A2A291326B1744D72FA8AFD0850BF" box="[548,658,1382,1409]" captionStart="Fig" captionStartId="160.[189,232,768,794]" captionTargetBox="[189,1398,285,725]" captionTargetId="figure-13@160.[189,1398,273,725]" captionTargetPageId="160" captionText="Fig. 46. Pseudonannolene ambuatinga Iniesta &amp; Ferreira, 2013, paratype, ♀ (ISLA 2274), in lateral view. A. Anterior region. B. Posterior region. Scale bars: A = 0.5 mm; B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891139" httpUri="https://zenodo.org/record/7891139/files/figure.png" pageId="21" pageNumber="22">Fig. 46A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B65271326B1744FEBFA4BFAB2532D" pageId="21" pageNumber="22" type="etymology">
<paragraph id="8BEE36AC1326B1744FEBFA4BFEDE50FF" blockId="21.[189,324,1447,1473]" box="[189,324,1447,1473]" pageId="21" pageNumber="22">
<heading id="D0A681C01326B1744FEBFA4BFEDE50FF" bold="true" box="[189,324,1447,1473]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B925EABE1326B1744FEBFA4BFEDE50FF" bold="true" box="[189,324,1447,1473]" pageId="21" pageNumber="22">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1326B1744FEBFA39FAB2532D" blockId="21.[189,1398,1493,1555]" pageId="21" pageNumber="22">
A combination of words of the Brazilian Indian language Tupi-Guarani, ‘ 
<emphasis id="B925EABE1326B1744B6BFA39FB1050D1" box="[1085,1162,1493,1519]" italics="true" pageId="21" pageNumber="22">ambus</emphasis>
’ = ‘millipede’, and ‘ 
<emphasis id="B925EABE1326B1744F91FA15FE99532C" box="[199,259,1529,1554]" italics="true" pageId="21" pageNumber="22">tinga</emphasis>
’ = ‘white’, referring to the body depigmentation of the species (
<bibRefCitation id="EFC04B5D1326B1744CA5FA14FA81532D" author="Iniesta L. F. &amp; Ferreira R. L." box="[1011,1307,1528,1555]" pageId="21" pageNumber="22" pagination="357 - 369" refId="ref103738" refString="Iniesta L. F. M &amp; Ferreira R. L. 2013 b. Two new species of Pseudonannolene Silvestri, 1895 from Brazilian limestone caves (Spirostreptida: Pseudonannolenidae): synotopy of a troglophilic and a troglobiotic species. Zootaxa 3702 (4): 357 - 369. https: // doi. org / 10.11646 / zootaxa. 3702.4.3" type="journal article" year="2013">Iniesta &amp; Ferreira 2013b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B65271326B1774FEBF9D4FBD25767" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="8BEE36AC1326B1744FEBF9D4FE30536C" blockId="21.[189,426,1592,1618]" box="[189,426,1592,1618]" pageId="21" pageNumber="22">
<heading id="D0A681C01326B1744FEBF9D4FE30536C" bold="true" box="[189,426,1592,1618]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B925EABE1326B1744FEBF9D4FE30536C" bold="true" box="[189,426,1592,1618]" pageId="21" pageNumber="22">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1326B1744FBCF98BFEFD53BF" blockId="21.[187,1398,1639,1735]" box="[234,359,1639,1665]" pageId="21" pageNumber="22">
<emphasis id="B925EABE1326B1744FBCF98BFEFD53BF" bold="true" box="[234,359,1639,1665]" pageId="21" pageNumber="22">
<typeStatus id="54EA880E1326B1744FBCF98BFEFD53BF" box="[234,359,1639,1665]" pageId="21" pageNumber="22" type="holotype">Holotypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BEE36AC1326B1744FEBF966FC3D53F8" blockId="21.[187,1398,1639,1735]" pageId="21" pageNumber="22">
<materialsCitation id="3B393CF11326B1744FEBF966FC3D53F8" ID-GBIF-Occurrence="4101310269" collectingDate="2009-01-28" collectionCode="R" collectorName="R. L. Ferreira" country="Brazil" latitude="-20.369646" location="cave Loca d'Agua de Baixo" longLatPrecision="1" longitude="-45.692917" municipality="Pains" pageId="21" pageNumber="22" specimenCode="ISLA 2267" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
<collectingCountry id="F346763C1326B1744FEBF966FEBC539A" box="[189,294,1674,1700]" name="Brazil" pageId="21" pageNumber="22">BRAZIL</collectingCountry>
• 
<specimenCount id="9D57FD251326B1744E6DF966FEC9539A" box="[315,339,1674,1700]" pageId="21" pageNumber="22" type="male">♂</specimenCount>
, 
<typeStatus id="54EA880E1326B1744E08F965FE5F539D" box="[350,453,1673,1699]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C514D2F1326B1744EBEF966FD15539D" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[488,655,1673,1699]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ambuatinga">
<emphasis id="B925EABE1326B1744EBEF966FD15539D" box="[488,655,1673,1699]" italics="true" pageId="21" pageNumber="22">P. ambuatinga</emphasis>
</taxonomicName>
; 
<collectingRegion id="4995F84E1326B1744DCBF965FCA2539D" box="[669,824,1673,1699]" country="Brazil" name="Minas Gerais" pageId="21" pageNumber="22">Minas Gerais</collectingRegion>
, 
<collectingMunicipality id="6B8AACD61326B1744C13F965FC1D539D" box="[837,903,1673,1699]" pageId="21" pageNumber="22">Pains</collectingMunicipality>
, 
<location id="8E8E60771326B1744CC7F966FB4F539D" LSID="urn:lsid:plazi:treatment:03F887BA1326B1764E87FE37FBE05726:8E8E60771326B1744CC7F966FB4F539D" box="[913,1237,1673,1702]" country="Brazil" latitude="-20.369646" longLatPrecision="1" longitude="-45.692917" municipality="Pains" name="cave Loca d'Agua de Baixo" pageId="21" pageNumber="22" stateProvince="Minas Gerais">cave Loca d’Água de Baixo</location>
; [
<geoCoordinate id="EE65506B1326B1744BB0F965FAE8539D" box="[1254,1394,1673,1699]" degrees="20.369647" direction="south" orientation="latitude" pageId="21" pageNumber="22" precision="1" value="-20.369646">-20.369647</geoCoordinate>
, 
<geoCoordinate id="EE65506B1326B1744FEDF947FED253F8" box="[187,328,1707,1734]" degrees="45.692915" direction="west" orientation="longitude" pageId="21" pageNumber="22" precision="1" value="-45.692917">-45.692915</geoCoordinate>
]; 
<date id="FFEF106C1326B1744E0DF940FE6953F8" box="[347,499,1708,1735]" pageId="21" pageNumber="22" value="2009-01-28">
<collectingDate id="EFABE9841326B1744E0DF940FE6953F8" box="[347,499,1708,1735]" pageId="21" pageNumber="22" value="2009-01-28">28 Jan. 2009</collectingDate>
</date>
; 
<collectorName id="26A4537A1326B1744EA9F941FD0253F8" box="[511,664,1708,1735]" pageId="21" pageNumber="22">R.L. Ferreira</collectorName>
<emphasis id="B925EABE1326B1744DF7F941FD4653F8" box="[673,732,1708,1734]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
leg.; 
<specimenCode id="DBF79ED71326B1744C48F940FC3853F8" box="[798,930,1708,1734]" pageId="21" pageNumber="22">ISLA 2267</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BEE36AC1326B1744FEBF901FD145214" blockId="21.[189,1398,1772,1834]" pageId="21" pageNumber="22">
<materialsCitation id="3B393CF11326B1744FEBF901FD145214" ID-GBIF-Occurrence="4101309711" collectingDate="2012-03-15" collectionCode="ISLA" collectorName="R. Zampaulo" country="Brazil" latitude="-20.384577" location="cave Eden" longLatPrecision="1" longitude="-45.666798" municipality="Pains" pageId="21" pageNumber="22" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
<collectingCountry id="F346763C1326B1744FEBF901FEBC5239" box="[189,294,1773,1799]" name="Brazil" pageId="21" pageNumber="22">BRAZIL</collectingCountry>
• 
<specimenCount id="9D57FD251326B1744E11F902FEC55236" box="[327,351,1774,1800]" pageId="21" pageNumber="22" type="male">♂</specimenCount>
, 
<typeStatus id="54EA880E1326B1744E39F901FE4C5239" box="[367,470,1773,1799]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C514D2F1326B1744D53F902FD085239" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[517,658,1774,1799]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="saguassu">
<emphasis id="B925EABE1326B1744D53F902FD085239" box="[517,658,1774,1799]" italics="true" pageId="21" pageNumber="22">P. saguassu</emphasis>
</taxonomicName>
; 
<collectingRegion id="4995F84E1326B1744DF3F901FCDC5239" box="[677,838,1773,1799]" country="Brazil" name="Minas Gerais" pageId="21" pageNumber="22">Minas Gerais</collectingRegion>
, 
<collectingMunicipality id="6B8AACD61326B1744C0EF901FC005239" box="[856,922,1773,1799]" pageId="21" pageNumber="22">Pains</collectingMunicipality>
, 
<location id="8E8E60771326B1744CFDF902FBB65234" LSID="urn:lsid:plazi:treatment:03F887BA1326B1764E87FE37FBE05726:8E8E60771326B1744CFDF902FBB65234" box="[939,1068,1774,1802]" country="Brazil" latitude="-20.384577" longLatPrecision="1" longitude="-45.666798" municipality="Pains" name="cave Eden" pageId="21" pageNumber="22" stateProvince="Minas Gerais">cave Éden</location>
; [
<geoCoordinate id="EE65506B1326B1744B14F901FB555236" box="[1090,1231,1773,1800]" degrees="20.384577" direction="south" orientation="latitude" pageId="21" pageNumber="22" precision="1" value="-20.384577">-20.384577</geoCoordinate>
, 
<geoCoordinate id="EE65506B1326B1744B88F900FAF15239" box="[1246,1387,1772,1799]" degrees="45.666798" direction="west" orientation="longitude" pageId="21" pageNumber="22" precision="1" value="-45.666798">-45.666798</geoCoordinate>
]; 
<date id="FFEF106C1326B1744FEBF8E3FEC45214" box="[189,350,1807,1834]" pageId="21" pageNumber="22" value="2012-03-15">
<collectingDate id="EFABE9841326B1744FEBF8E3FEC45214" box="[189,350,1807,1834]" pageId="21" pageNumber="22" value="2012-03-15">15 Mar. 2012</collectingDate>
</date>
; 
<collectorName id="26A4537A1326B1744E3CF8FCFD995214" box="[362,515,1808,1834]" pageId="21" pageNumber="22">R. Zampaulo</collectorName>
leg.; 
<collectionCode id="ED40AE691326B1744D13F8FCFD105214" box="[581,650,1808,1834]" pageId="21" pageNumber="22">ISLA</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BEE36AC1326B1744FBCF8BDFDC35255" blockId="21.[189,1399,1872,2003]" box="[234,601,1872,1901]" pageId="21" pageNumber="22">
<emphasis id="B925EABE1326B1744FBCF8BDFEFD5255" bold="true" box="[234,359,1873,1899]" pageId="21" pageNumber="22">
<typeStatus id="54EA880E1326B1744FBCF8BDFEFD5255" box="[234,359,1873,1899]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
</emphasis>
(total: 
<specimenCount id="9D57FD251326B1744EEBF8BDFE655253" box="[445,511,1873,1901]" pageId="21" pageNumber="22" type="male">
3 
<emphasis id="B925EABE1326B1744E85F8BDFE655253" bold="true" box="[467,511,1873,1901]" pageId="21" pageNumber="22">♂♂</emphasis>
</specimenCount>
, 
<specimenCount id="9D57FD251326B1744D5BF8BCFDD55253" box="[525,591,1872,1901]" pageId="21" pageNumber="22" type="female">
5 
<emphasis id="B925EABE1326B1744D75F8BDFDD55253" bold="true" box="[547,591,1873,1901]" pageId="21" pageNumber="22">♀♀</emphasis>
</specimenCount>
)
</paragraph>
<paragraph id="8BEE36AC1326B1774FEBF89FFE535490" blockId="21.[189,1399,1872,2003]" lastBlockId="22.[188,1399,265,430]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<collectingCountry id="F346763C1326B1744FEBF89FFEBC52B3" box="[189,294,1907,1933]" name="Brazil" pageId="21" pageNumber="22">BRAZIL</collectingCountry>
• 
<materialsCitation id="3B393CF11326B1744E6FF89FFB5052B3" ID-GBIF-Occurrence="4101309500" box="[313,1226,1907,1934]" collectingDate="2012-03-15" collectionCode="R" collectorName="Zampaulo" country="Iceland" latitude="-20.384577" location="cave Eden" longLatPrecision="1" longitude="-45.666798" municipality="Pains" pageId="21" pageNumber="22" specimenCode="ISLA 2272" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount id="9D57FD251326B1744E6FF89FFEFE52B0" box="[313,356,1907,1934]" pageId="21" pageNumber="22" type="male">1 ♂</specimenCount>
, 
<typeStatus id="54EA880E1326B1744E38F898FE4952B0" box="[366,467,1908,1934]" pageId="21" pageNumber="22" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C514D2F1326B1744EA2F898FD0052B3" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[500,666,1907,1933]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ambuatinga">
<emphasis id="B925EABE1326B1744EA2F898FD0052B3" box="[500,666,1907,1933]" italics="true" pageId="21" pageNumber="22">P. ambuatinga</emphasis>
</taxonomicName>
; same collection data as for holotype; 
<specimenCode id="DBF79ED71326B1744B1DF89FFB5052B3" box="[1099,1226,1907,1933]" pageId="21" pageNumber="22">ISLA 2272</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B393CF11326B1744B8BF89FFC46528E" ID-GBIF-Occurrence="4101309312" collectingDate="2012-03-15" collectionCode="R" collectorName="Zampaulo" country="Iceland" latitude="-20.384577" location="cave Eden" longLatPrecision="1" longitude="-45.666798" municipality="Pains" pageId="21" pageNumber="22" specimenCode="ISLA 2268" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount id="9D57FD251326B1744B8BF89FFA9C52B0" box="[1245,1286,1907,1934]" pageId="21" pageNumber="22" type="female">1 ♀</specimenCount>
, 
<typeStatus id="54EA880E1326B1744A47F898FAEC52B0" box="[1297,1398,1908,1934]" pageId="21" pageNumber="22" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C514D2F1326B1744F88F87BFE12528E" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[222,392,1942,1968]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ambuatinga">
<emphasis id="B925EABE1326B1744F88F87BFE12528E" box="[222,392,1942,1968]" italics="true" pageId="21" pageNumber="22">P. ambuatinga</emphasis>
</taxonomicName>
; same collection data as for holotype; 
<specimenCode id="DBF79ED71326B1744C0EF87AFC46528E" box="[856,988,1942,1968]" pageId="21" pageNumber="22">ISLA 2268</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B393CF11326B1744CA1F87AFD6952EC" ID-GBIF-Occurrence="4101309972" collectingDate="2012-03-15" collectionCode="R" collectorName="Zampaulo" country="Iceland" latitude="-20.384577" location="cave Eden" longLatPrecision="1" longitude="-45.666798" municipality="Pains" pageId="21" pageNumber="22" specimenCode="ISLA 2269" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount id="9D57FD251326B1744CA1F87AFBBF528E" box="[1015,1061,1942,1968]" pageId="21" pageNumber="22" type="female">1 ♀</specimenCount>
, 
<typeStatus id="54EA880E1326B1744B63F87AFB00528E" box="[1077,1178,1942,1968]" pageId="21" pageNumber="22" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C514D2F1326B1744B92F87BFAF4528E" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[1220,1390,1942,1968]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ambuatinga">
<emphasis id="B925EABE1326B1744B92F87BFAF4528E" box="[1220,1390,1942,1968]" italics="true" pageId="21" pageNumber="22">P. ambuatinga</emphasis>
</taxonomicName>
; same collection data as for holotype; 
<specimenCode id="DBF79ED71326B1744D27F855FD6952EC" box="[625,755,1976,2003]" pageId="21" pageNumber="22">ISLA 2269</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B393CF11326B1774C5DF855FE60541D" ID-GBIF-Occurrence="4101309890" collectingDate="2012-03-15" collectionCode="R" collectorName="Zampaulo" country="Iceland" lastPageId="22" lastPageNumber="23" latitude="-20.384577" location="cave Eden" longLatPrecision="1" longitude="-45.666798" municipality="Pains" pageId="21" pageNumber="22" specimenCode="ISLA 2270" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount id="9D57FD251326B1744C5DF855FCAC52ED" box="[779,822,1977,2003]" pageId="21" pageNumber="22" type="female">1 ♀</specimenCount>
, 
<typeStatus id="54EA880E1326B1744C13F855FC3052ED" box="[837,938,1977,2003]" pageId="21" pageNumber="22" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C514D2F1326B1744C86F855FBE252EC" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[976,1144,1976,2002]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ambuatinga">
<emphasis id="B925EABE1326B1744C86F855FBE252EC" box="[976,1144,1976,2002]" italics="true" pageId="21" pageNumber="22">P. ambuatinga</emphasis>
</taxonomicName>
; same collection data as for holotype; 
<specimenCode id="DBF79ED71325B1774E2FFEE5FE60541D" box="[377,506,265,291]" pageId="22" pageNumber="23">ISLA 2270</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B393CF11325B1774D59FEE5FF6D5478" ID-GBIF-Occurrence="4101310176" collectingDate="2012-03-15" collectionCode="R" collectorName="Zampaulo" country="Iceland" latitude="-20.384577" location="cave Eden" longLatPrecision="1" longitude="-45.666798" municipality="Pains" pageId="22" pageNumber="23" specimenCode="ISLA 2271" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="paratype">
<specimenCount id="9D57FD251325B1774D59FEE5FDA1541A" box="[527,571,265,292]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
, 
<typeStatus id="54EA880E1325B1774D1EFEE6FD37541A" box="[584,685,266,292]" pageId="22" pageNumber="23" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C514D2F1325B1774D84FEE6FCE3541D" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[722,889,265,291]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ambuatinga">
<emphasis id="B925EABE1325B1774D84FEE6FCE3541D" box="[722,889,265,291]" italics="true" pageId="22" pageNumber="23">P. ambuatinga</emphasis>
</taxonomicName>
; same collection data as for holotype; 
<specimenCode id="DBF79ED71325B1774A61FEE5FF6D5478" pageId="22" pageNumber="23">ISLA 2271</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B393CF11325B1774E58FEC0FC595456" ID-GBIF-Occurrence="4101309578" collectingDate="2012-03-15" collectionCode="R" collectorName="R. L. Ferreira &amp; P. Ratton &amp; M. Souza-Silva" country="Iceland" county="cave Eden" latitude="-20.384577" location="Minas Gerais" longLatPrecision="1" longitude="-45.666798" municipality="Pains" pageId="22" pageNumber="23" specimenCode="ISLA 2273" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D57FD251325B1774E58FEC0FEA15478" box="[270,315,300,326]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
, 
<typeStatus id="54EA880E1325B1774E10FEC0FE315478" box="[326,427,300,326]" pageId="22" pageNumber="23" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C514D2F1325B1774E99FEC0FDCD547B" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[463,599,300,325]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="saguassu">
<emphasis id="B925EABE1325B1774E99FEC0FDCD547B" box="[463,599,300,325]" italics="true" pageId="22" pageNumber="23">P. saguassu</emphasis>
</taxonomicName>
; 
<location id="8E8E60771325B1774D32FEC0FC9B5478" LSID="urn:lsid:plazi:treatment:03F887BA1326B1764E87FE37FBE05726:8E8E60771325B1774D32FEC0FC9B5478" box="[612,769,300,326]" country="Iceland" county="cave Eden" latitude="-20.384577" longLatPrecision="1" longitude="-45.666798" municipality="Pains" name="Minas Gerais" pageId="22" pageNumber="23">Minas Gerais</location>
, 
<collectingMunicipality id="6B8AACD61325B1774C58FEC0FCCA5478" box="[782,848,300,326]" pageId="22" pageNumber="23">Pains</collectingMunicipality>
, 
<collectingCounty id="628F4E201325B1774C0DFEC0FC4C5476" box="[859,982,300,328]" pageId="22" pageNumber="23">cave Éden</collectingCounty>
; [
<geoCoordinate id="EE65506B1325B1774CB1FEC7FBEE5478" box="[999,1140,299,326]" degrees="20.384577" direction="south" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="-20.384577">-20.384577</geoCoordinate>
, 
<geoCoordinate id="EE65506B1325B1774B28FEC7FA915478" box="[1150,1291,299,326]" degrees="45.666798" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="-45.666798">-45.666798</geoCoordinate>
]; 
<date id="FFEF106C1325B1774A4BFEC7FF605456" pageId="22" pageNumber="23" value="2012-03-15">
<collectingDate id="EFABE9841325B1774A4BFEC7FF605456" pageId="22" pageNumber="23" value="2012-03-15">15 Mar. 2012</collectingDate>
</date>
; 
<collectorName id="26A4537A1325B1774E53FEA3FE3A5456" box="[261,416,334,361]" pageId="22" pageNumber="23">R.L. Ferreira</collectorName>
, 
<collectorName id="26A4537A1325B1774EFCFEA3FD895457" box="[426,531,335,361]" pageId="22" pageNumber="23">P. Ratton</collectorName>
and 
<collectorName id="26A4537A1325B1774D1AFEA3FC985456" box="[588,770,334,361]" pageId="22" pageNumber="23">M. Souza-Silva</collectorName>
leg.; 
<specimenCode id="DBF79ED71325B1774C14FEA2FC595456" box="[834,963,334,360]" pageId="22" pageNumber="23">ISLA 2273</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B393CF11325B1774C8FFEA2FD2454B2" ID-GBIF-Occurrence="4101310075" collectingDate="2012-03-15" collectionCode="R" collectorName="L. Ferreira &amp; P. Ratton &amp; M. Souza-Silva" country="Iceland" county="cave Eden" latitude="-20.384577" location="Minas Gerais" longLatPrecision="1" longitude="-45.666798" municipality="Pains" pageId="22" pageNumber="23" specimenCode="ISLA 2275" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D57FD251325B1774C8FFEA2FB9F5457" box="[985,1029,334,361]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
, 
<typeStatus id="54EA880E1325B1774B4CFEA3FBE55457" box="[1050,1151,335,361]" pageId="22" pageNumber="23" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C514D2F1325B1774BF2FEA3FAB65456" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[1188,1324,335,360]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="saguassu">
<emphasis id="B925EABE1325B1774BF2FEA3FAB65456" box="[1188,1324,335,360]" italics="true" pageId="22" pageNumber="23">P. saguassu</emphasis>
</taxonomicName>
; same collection data as for preceding; 
<specimenCode id="DBF79ED71325B1774D6AFE9DFD2454B2" box="[572,702,369,396]" pageId="22" pageNumber="23">ISLA 2275</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B393CF11325B1774D81FE9DFE585490" ID-GBIF-Occurrence="4101309472" collectingDate="2012-03-15" collectionCode="R" collectorName="L. Ferreira &amp; P. Ratton &amp; M. Souza-Silva" country="Iceland" county="cave Eden" latitude="-20.384577" location="Minas Gerais" longLatPrecision="1" longitude="-45.666798" municipality="Pains" pageId="22" pageNumber="23" specimenCode="ISLA 2274" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D57FD251325B1774D81FE9DFC9954B2" box="[727,771,369,396]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
, 
<typeStatus id="54EA880E1325B1774C4FFE9EFCE454B2" box="[793,894,370,396]" pageId="22" pageNumber="23" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C514D2F1325B1774CF0FE9EFBB454B5" authorityName="Iniesta &amp; Ferreira" authorityYear="2013" box="[934,1070,370,395]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="saguassu">
<emphasis id="B925EABE1325B1774CF0FE9EFBB454B5" box="[934,1070,370,395]" italics="true" pageId="22" pageNumber="23">P. saguassu</emphasis>
</taxonomicName>
; same collection data as for preceding; 
<specimenCode id="DBF79ED71325B1774E16FE78FE585490" box="[320,450,404,430]" pageId="22" pageNumber="23">ISLA 2274</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36AC1325B1774FBCFE39FBD25767" blockId="22.[189,1398,469,601]" pageId="22" pageNumber="23">
<emphasis id="B925EABE1325B1774FBCFE39FE3C54D1" bold="true" box="[234,422,469,495]" pageId="22" pageNumber="23">Other material</emphasis>
(total: 
<specimenCount id="9D57FD251325B1774EADFE39FDBD54CC" box="[507,551,469,498]" pageId="22" pageNumber="23" type="male">
1 
<emphasis id="B925EABE1325B1774D47FE3AFDBD54CC" bold="true" box="[529,551,470,498]" pageId="22" pageNumber="23">♂</emphasis>
</specimenCount>
, 
<specimenCount id="9D57FD251325B1774D63FE39FDFB54CC" box="[565,609,469,498]" pageId="22" pageNumber="23" type="female">
1 
<emphasis id="B925EABE1325B1774D1DFE3AFDFB54CC" bold="true" box="[587,609,470,498]" pageId="22" pageNumber="23">♀</emphasis>
</specimenCount>
) 
<materialsCitation id="3B393CF11325B1774FEBFE14FAEA572C" ID-GBIF-Occurrence="4101309740" box="[189,1392,504,531]" country="Brazil" elevation="766" latitude="-20.28908" location="Minas Gerais" longLatPrecision="1" longitude="-45.540085" municipality="Arcos" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais">
<collectingCountry id="F346763C1325B1774FEBFE14FEBC572C" box="[189,294,504,530]" name="Brazil" pageId="22" pageNumber="23">BRAZIL</collectingCountry>
– 
<emphasis id="B925EABE1325B1774E1CFE14FE6F572C" bold="true" box="[330,501,504,530]" pageId="22" pageNumber="23">
<collectingRegion id="4995F84E1325B1774E1CFE14FE6F572C" box="[330,501,504,530]" country="Brazil" name="Minas Gerais" pageId="22" pageNumber="23">Minas Gerais</collectingRegion>
</emphasis>
• 
<specimenCount id="9D57FD251325B1774D46FE14FDDB572D" box="[528,577,504,531]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
; 
<collectingMunicipality id="6B8AACD61325B1774D18FE14FD0D572C" box="[590,663,504,530]" pageId="22" pageNumber="23">Arcos</collectingMunicipality>
,, 
<location id="8E8E60771325B1774DE0FE15FC0B572C" LSID="urn:lsid:plazi:treatment:03F887BA1326B1764E87FE37FBE05726:8E8E60771325B1774DE0FE15FC0B572C" box="[694,913,504,531]" country="Brazil" latitude="-20.28908" longLatPrecision="1" longitude="-45.540085" municipality="Arcos" name="cave Alinhamento" pageId="22" pageNumber="23" stateProvince="Minas Gerais">cave Alinhamento</location>
; [
<geoCoordinate id="EE65506B1325B1774CF3FE14FBA8572C" box="[933,1074,504,530]" degrees="20.289079" direction="south" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="-20.28908">-20.289079</geoCoordinate>
, 
<geoCoordinate id="EE65506B1325B1774B16FE14FB57572D" box="[1088,1229,504,531]" degrees="45.540084" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="-45.540085">-45.540084</geoCoordinate>
]; 
<elevation id="007CD19F1325B1774BB4FE14FAEA572C" box="[1250,1392,504,531]" metricMagnitude="2" metricUnit="m" metricValue="7.659999999999999" pageId="22" pageNumber="23" unit="m" value="766.0">
<quantity id="4CA99B491325B1774BB4FE14FAB4572D" box="[1250,1326,504,531]" metricMagnitude="2" metricUnit="m" metricValue="7.659999999999999" pageId="22" pageNumber="23" unit="m" value="766.0">766 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
; 
<materialsCitation id="3B393CF11325B1774FE9FDF0FD435708" ID-GBIF-Occurrence="4101309508" box="[191,729,539,566]" collectingDate="2002-06-01" collectionCode="R" collectorName="R. L. Ferreira" pageId="22" pageNumber="23" specimenCode="IBSP3442" specimenCount="1">
<date id="FFEF106C1325B1774FE9FDF0FEDC5708" box="[191,326,540,566]" pageId="22" pageNumber="23" value="2002-06-01">
<collectingDate id="EFABE9841325B1774FE9FDF0FEDC5708" box="[191,326,540,566]" pageId="22" pageNumber="23" value="2002-06-01">1 Jun. 2002</collectingDate>
</date>
; 
<collectorName id="26A4537A1325B1774E06FDF0FE7C570B" box="[336,486,539,566]" pageId="22" pageNumber="23">R.L. Ferreira</collectorName>
<emphasis id="B925EABE1325B1774EB0FDF0FD85570B" box="[486,543,539,565]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
leg.; 
<specimenCode id="DBF79ED71325B1774D09FDF0FD435708" box="[607,729,540,566]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="22" pageNumber="23">IBSP3442</specimenCode>
</materialsCitation>
• 
<materialsCitation id="3B393CF11325B1774DBBFDF0FBDE5767" ID-GBIF-Occurrence="4101309435" collectingDate="2008-01-25" collectionCode="IBSP" collectorName="E. O. Machado &amp; J. P. P. Barbosa" elevation="700" latitude="-20.286839" location="Iguatama" longLatPrecision="1" longitude="-45.79329" municipality="cave Arcaica" pageId="22" pageNumber="23" specimenCode="IBSP 3315" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D57FD251325B1774DBBFDF0FC805708" box="[749,794,540,566]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
; 
<location id="8E8E60771325B1774C70FDF0FC0C5708" LSID="urn:lsid:plazi:treatment:03F887BA1326B1764E87FE37FBE05726:8E8E60771325B1774C70FDF0FC0C5708" box="[806,918,540,566]" latitude="-20.286839" longLatPrecision="1" longitude="-45.79329" municipality="cave Arcaica" name="Iguatama" pageId="22" pageNumber="23">Iguatama</location>
, 
<collectingMunicipality id="6B8AACD61325B1774CF6FDF0FBA2570B" box="[928,1080,539,566]" pageId="22" pageNumber="23">cave Arcaica</collectingMunicipality>
; [
<geoCoordinate id="EE65506B1325B1774B1EFDF7FB4F570B" box="[1096,1237,539,565]" degrees="20.286839" direction="south" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="-20.286839">-20.286839</geoCoordinate>
, 
<geoCoordinate id="EE65506B1325B1774B88FDF7FAF15708" box="[1246,1387,539,566]" degrees="45.793289" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="-45.79329">-45.793289</geoCoordinate>
]; 
<elevation id="007CD19F1325B1774FEBFDD3FEDD5767" box="[189,327,575,601]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="22" pageNumber="23" unit="m" value="700.0">
<quantity id="4CA99B491325B1774FEBFDD3FE9D5767" box="[189,263,575,601]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="22" pageNumber="23" unit="m" value="700.0">700 m</quantity>
a.s.l.
</elevation>
; 
<date id="FFEF106C1325B1774E03FDD2FE775767" box="[341,493,574,601]" pageId="22" pageNumber="23" value="2008-01-25">
<collectingDate id="EFABE9841325B1774E03FDD2FE775767" box="[341,493,574,601]" pageId="22" pageNumber="23" value="2008-01-25">25 Jan. 2008</collectingDate>
</date>
; 
<collectorName id="26A4537A1325B1774EAFFDD3FD395767" box="[505,675,575,601]" pageId="22" pageNumber="23">E.O. Machado</collectorName>
and 
<collectorName id="26A4537A1325B1774D8AFDD3FCE55767" box="[732,895,575,601]" pageId="22" pageNumber="23">J.P.P. Barbosa</collectorName>
leg.; 
<specimenCode id="DBF79ED71325B1774C97FDD3FBDE5767" box="[961,1092,574,601]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="22" pageNumber="23">IBSP 3315</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B65271325B1774FEBFD6CFBB55201" pageId="22" pageNumber="23" type="description">
<paragraph id="8BEE36AC1325B1774FEBFD6CFE0957A4" blockId="22.[189,403,640,666]" box="[189,403,640,666]" pageId="22" pageNumber="23">
<heading id="D0A681C01325B1774FEBFD6CFE0957A4" bold="true" box="[189,403,640,666]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B925EABE1325B1774FEBFD6CFE0957A4" bold="true" box="[189,403,640,666]" pageId="22" pageNumber="23">Descriptive notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1325B1774FEBFD43FEA6562E" blockId="22.[189,1399,686,784]" pageId="22" pageNumber="23">
<smallCapsWord id="8D08A0701325B1774FEBFD43FEF457F9" baselines="706,706" box="[189,366,687,713]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="22" pageNumber="23">MEASUREMENTS</smallCapsWord>
. 58–60 body rings (1–2 apodous + telson). Males: body length 
<quantity id="4CA99B491325B1774B0DFD42FB6957F7" box="[1115,1267,686,713]" metricMagnitude="-2" metricUnit="m" metricValue="3.875" metricValueMax="4.25" metricValueMin="3.5" pageId="22" pageNumber="23" unit="mm" value="38.75" valueMax="42.5" valueMin="35.0">35–42.5 mm</quantity>
; maximum midbody diameter 
<quantity id="4CA99B491325B1774EFDFD3EFDDE57D2" box="[427,580,722,748]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="22" pageNumber="23" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.8–1.9 mm</quantity>
. Females: body length 
<quantity id="4CA99B491325B1774C22FD3EFC6457D2" box="[884,1022,722,748]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" metricValueMax="4.1" metricValueMin="3.6" pageId="22" pageNumber="23" unit="mm" value="38.5" valueMax="41.0" valueMin="36.0">36–41 mm</quantity>
; maximum midbody diameter 
<quantity id="4CA99B491325B1774FEBFD19FEAD562E" box="[189,311,757,784]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="3.1" metricValueMin="2.0" pageId="22" pageNumber="23" unit="mm" value="2.55" valueMax="3.1" valueMin="2.0">2–3.1 mm</quantity>
.
</paragraph>
<paragraph id="8BEE36AC1325B1774FEBFCDAFA84564C" blockId="22.[189,1398,822,883]" pageId="22" pageNumber="23">
<smallCapsWord id="8D08A0701325B1774FEBFCDAFE935670" baselines="841,841" box="[189,265,822,848]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Color" pageId="22" pageNumber="23">COLOR</smallCapsWord>
. Living specimens depigmented. Color when stored in 70% ethanol: uniform pale brownish whitish, slightly darker posteriorly on prozonites; head, collum, antennae, and legs light brown.
</paragraph>
<paragraph id="8BEE36AC1325B1774FEBFC75FC1656C6" blockId="22.[189,1399,921,1016]" pageId="22" pageNumber="23">
<smallCapsWord id="8D08A0701325B1774FEBFC75FF66568F" baselines="940,940" box="[189,252,921,947]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="22" pageNumber="23">HEAD</smallCapsWord>
. Antennae short (
<figureCitation id="136A2A291325B1774E86FC75FDA4568A" box="[464,574,921,948]" captionStart="Fig" captionStartId="160.[189,232,768,794]" captionTargetBox="[189,1398,285,725]" captionTargetId="figure-13@160.[189,1398,273,725]" captionTargetPageId="160" captionText="Fig. 46. Pseudonannolene ambuatinga Iniesta &amp; Ferreira, 2013, paratype, ♀ (ISLA 2274), in lateral view. A. Anterior region. B. Posterior region. Scale bars: A = 0.5 mm; B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891139" httpUri="https://zenodo.org/record/7891139/files/figure.png" pageId="22" pageNumber="23">Fig. 46A</figureCitation>
), just reaching back to end of ring 5 when extended dorsally; relative antennomere lengths 1&lt;2&lt;3&gt;4&gt;5≈6&gt;7. Mandibular cardo with ventral margin swollen. Ommatidial cluster well-developed, elliptical; ca 23 ommatidia in 4 rows.
</paragraph>
<paragraph id="8BEE36AC1325B1774FEBFBF3FED7519E" blockId="22.[189,1399,1054,1184]" pageId="22" pageNumber="23">
<smallCapsWord id="8D08A0701325B1774FEBFBF3FF645109" baselines="1074,1074" box="[189,254,1055,1081]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Body" pageId="22" pageNumber="23">BODY</smallCapsWord>
<smallCapsWord id="8D08A0701325B1774E5CFBC8FED25109" baselines="1074" box="[266,328,1060,1079]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="rings" pageId="22" pageNumber="23">RINGS</smallCapsWord>
. Collum with lateral lobes broadly rounded, with ca 10 striae (
<figureCitation id="136A2A291325B1774B0AFBF2FB545107" box="[1116,1230,1054,1081]" captionStart="Fig" captionStartId="160.[189,232,768,794]" captionTargetBox="[189,1398,285,725]" captionTargetId="figure-13@160.[189,1398,273,725]" captionTargetPageId="160" captionText="Fig. 46. Pseudonannolene ambuatinga Iniesta &amp; Ferreira, 2013, paratype, ♀ (ISLA 2274), in lateral view. A. Anterior region. B. Posterior region. Scale bars: A = 0.5 mm; B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7891139" httpUri="https://zenodo.org/record/7891139/files/figure.png" pageId="22" pageNumber="23">Fig. 46A</figureCitation>
). Very faintly constricted between prozonite and metazonite; prozonites smooth; metazonites laterally with transverse striae below ozopore. Anterior sterna in midbody rings subrectangular, without transverse striae (
<figureCitation id="136A2A291325B1774F90FB69FEDA519E" box="[198,320,1157,1184]" captionStart="Fig" captionStartId="270.[189,232,1892,1918]" captionTargetBox="[213,1374,265,1851]" captionTargetId="figure-14@270.[213,1374,265,1851]" captionTargetPageId="270" captionText="Fig. 167. Stereoscopic images of gnathochilarium and anterior sternum of species of Pseudonannolene Silvestri, 1895. A. P. albiventris Schubart, 1952. B. P. ambuatinga Iniesta &amp; Ferreira, 2013. C. P. anapophysis Fontanetti, 1996. D. P. buhrnheimi Schubart, 1960. E. P. caatinga Iniesta &amp; Ferreira, 2014. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891417" httpUri="https://zenodo.org/record/7891417/files/figure.png" pageId="22" pageNumber="23">Fig. 167B</figureCitation>
).
</paragraph>
<paragraph id="8BEE36AC1325B1774FEBFB2BFD4D5079" blockId="22.[188,1398,1222,1352]" pageId="22" pageNumber="23">
<smallCapsWord id="8D08A0701325B1774FEBFB2BFF6051E1" baselines="1242,1242" box="[189,250,1223,1249]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="First" pageId="22" pageNumber="23">FIRST</smallCapsWord>
<smallCapsWord id="8D08A0701325B1774E53FB20FEFE51E1" baselines="1242,1242,1242" box="[261,356,1223,1249]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg-pair" pageId="22" pageNumber="23">LEG-PAIR</smallCapsWord>
<smallCapsWord id="8D08A0701325B1774E39FB20FE1351E1" baselines="1242" box="[367,393,1228,1247]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="of" pageId="22" pageNumber="23">OF</smallCapsWord>
<smallCapsWord id="8D08A0701325B1774EC2FB20FE4051E1" baselines="1242" box="[404,474,1228,1247]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="males" pageId="22" pageNumber="23">MALES</smallCapsWord>
. Coxae (
<emphasis id="B925EABE1325B1774D1DFB2AFDFD51DE" bold="true" box="[587,615,1222,1248]" italics="true" pageId="22" pageNumber="23">cx</emphasis>
) elongated (as long as the sum of remaining podomere lengths), subtriangular, with the base slightly arched, densely setose (
<figureCitation id="136A2A291325B1774C2EFB04FC7F503D" box="[888,997,1256,1283]" captionStart="Fig" captionStartId="161.[189,232,1668,1694]" captionTargetBox="[189,1398,269,1626]" captionTargetId="figure-11@161.[189,1398,269,1626]" captionTargetPageId="161" captionText="Fig. 47. Pseudonannolene ambuatinga Iniesta &amp; Ferreira, 2013, ♂ (IBSP 3442). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891141" httpUri="https://zenodo.org/record/7891141/files/figure.png" pageId="22" pageNumber="23">Fig. 47A</figureCitation>
); prefemoral process (
<emphasis id="B925EABE1325B1774BBCFB04FA94503C" bold="true" box="[1258,1294,1256,1282]" italics="true" pageId="22" pageNumber="23">prf</emphasis>
) as wide as half of prefemur, subcylindrical, densely setose along in its entire extension (
<figureCitation id="136A2A291325B1774B2CFAE7FB72501B" box="[1146,1256,1291,1317]" captionStart="Fig" captionStartId="161.[189,232,1668,1694]" captionTargetBox="[189,1398,269,1626]" captionTargetId="figure-11@161.[189,1398,269,1626]" captionTargetPageId="161" captionText="Fig. 47. Pseudonannolene ambuatinga Iniesta &amp; Ferreira, 2013, ♂ (IBSP 3442). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891141" httpUri="https://zenodo.org/record/7891141/files/figure.png" pageId="22" pageNumber="23">Fig. 47B</figureCitation>
); remaining podomeres with setae along the mesal region.
</paragraph>
<paragraph id="8BEE36AC1325B1774FEBFA82FDB550F3" blockId="22.[189,1400,1390,1485]" pageId="22" pageNumber="23">
<smallCapsWord id="8D08A0701325B1774FEBFA82FE8E50B8" baselines="1409,1409" box="[189,276,1390,1416]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Second" pageId="22" pageNumber="23">SECOND</smallCapsWord>
<smallCapsWord id="8D08A0701325B1774E48FA9FFEE750B8" baselines="1409,1409,1409" box="[286,381,1390,1416]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="leg-pair" pageId="22" pageNumber="23">LEG-PAIR</smallCapsWord>
<smallCapsWord id="8D08A0701325B1774ED1FA9FFE3B50B8" baselines="1409" box="[391,417,1395,1414]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="of" pageId="22" pageNumber="23">OF</smallCapsWord>
<smallCapsWord id="8D08A0701325B1774EFCFA9FFE6A50B8" baselines="1409" box="[426,496,1395,1414]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="males" pageId="22" pageNumber="23">MALES</smallCapsWord>
. Coxa (
<emphasis id="B925EABE1325B1774D07FA82FDF750B6" bold="true" box="[593,621,1390,1416]" italics="true" pageId="22" pageNumber="23">cx</emphasis>
) large and subrectangular; penis (
<emphasis id="B925EABE1325B1774B52FA82FBB950B6" bold="true" box="[1028,1059,1390,1416]" italics="true" pageId="22" pageNumber="23">pn</emphasis>
) located at proximal region, rounded, not extended basally (
<figureCitation id="136A2A291325B1774D65FA7CFD3B5094" box="[563,673,1424,1450]" captionStart="Fig" captionStartId="161.[189,232,1668,1694]" captionTargetBox="[189,1398,269,1626]" captionTargetId="figure-11@161.[189,1398,269,1626]" captionTargetPageId="161" captionText="Fig. 47. Pseudonannolene ambuatinga Iniesta &amp; Ferreira, 2013, ♂ (IBSP 3442). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891141" httpUri="https://zenodo.org/record/7891141/files/figure.png" pageId="22" pageNumber="23">Fig. 47C</figureCitation>
); prefemur compressed dorsoventrally; remaining podomeres setose, with long setae mesally.
</paragraph>
<paragraph id="8BEE36AC1325B1774FEBFA1FFB3153E2" blockId="22.[189,1399,1523,1756]" pageId="22" pageNumber="23">
<smallCapsWord id="8D08A0701325B1774FEBFA1FFEAE5332" baselines="1542,1543" box="[189,308,1523,1549]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Gonopods" pageId="22" pageNumber="23">GONOPODS</smallCapsWord>
. Gonocoxa (
<emphasis id="B925EABE1325B1774E84FA18FE665330" bold="true" box="[466,508,1524,1550]" italics="true" pageId="22" pageNumber="23">gcx</emphasis>
) elongated, but less than twice the length of the telopodite, with the base slightly arched; slightly flattened antero-posteriorly (
<figureCitation id="136A2A291325B1774C1DF9FAFC44530E" box="[843,990,1558,1584]" captionStart="Fig" captionStartId="161.[189,232,1668,1694]" captionTargetBox="[189,1398,269,1626]" captionTargetId="figure-11@161.[189,1398,269,1626]" captionTargetPageId="161" captionText="Fig. 47. Pseudonannolene ambuatinga Iniesta &amp; Ferreira, 2013, ♂ (IBSP 3442). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891141" httpUri="https://zenodo.org/record/7891141/files/figure.png" pageId="22" pageNumber="23">Fig. 47D–F</figureCitation>
); with rows of papillae mesally. Seminal groove (
<emphasis id="B925EABE1325B1774EDFF9D4FE39536C" bold="true" box="[393,419,1592,1618]" italics="true" pageId="22" pageNumber="23">sg</emphasis>
) curved; arising medially on mesal cavity and terminating apically on the seminal apophysis (
<emphasis id="B925EABE1325B1774E13F9B7FEC5534B" bold="true" box="[325,351,1627,1653]" italics="true" pageId="22" pageNumber="23">sa</emphasis>
). Shoulder (
<emphasis id="B925EABE1325B1774EA3F9B6FD8B534B" bold="true" box="[501,529,1626,1653]" italics="true" pageId="22" pageNumber="23">sh</emphasis>
) inconspicuous. Telopodite (
<emphasis id="B925EABE1325B1774C3CF9B7FC1B534B" bold="true" box="[874,897,1627,1653]" italics="true" pageId="22" pageNumber="23">tp</emphasis>
) less than half as wide as 
<emphasis id="B925EABE1325B1774BE9F9B7FB73534B" bold="true" box="[1215,1257,1627,1653]" italics="true" pageId="22" pageNumber="23">gcx</emphasis>
(
<figureCitation id="136A2A291325B1774BACF9B6FAF3534B" box="[1274,1385,1626,1653]" captionStart="Fig" captionStartId="161.[189,232,1668,1694]" captionTargetBox="[189,1398,269,1626]" captionTargetId="figure-11@161.[189,1398,269,1626]" captionTargetPageId="161" captionText="Fig. 47. Pseudonannolene ambuatinga Iniesta &amp; Ferreira, 2013, ♂ (IBSP 3442). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891141" httpUri="https://zenodo.org/record/7891141/files/figure.png" pageId="22" pageNumber="23">Fig. 47D</figureCitation>
); solenomere (
<emphasis id="B925EABE1325B1774E03F990FEF353A9" bold="true" box="[341,361,1660,1687]" italics="true" pageId="22" pageNumber="23">sl</emphasis>
) with apicomesal process (
<emphasis id="B925EABE1325B1774DFEF991FD4653A9" bold="true" box="[680,732,1661,1687]" italics="true" pageId="22" pageNumber="23">amp</emphasis>
) subtriangular; ectal process absent; 
<emphasis id="B925EABE1325B1774BDBF991FB3D53A9" bold="true" box="[1165,1191,1661,1687]" italics="true" pageId="22" pageNumber="23">sa</emphasis>
located at medial portion, visible apically. Internal branch (
<emphasis id="B925EABE1325B1774DF4F973FD235384" bold="true" box="[674,697,1695,1722]" italics="true" pageId="22" pageNumber="23">ib</emphasis>
) shovel-shaped and rounded apically, with horizontal plate; setae restricted to the apical region of 
<emphasis id="B925EABE1325B1774D2AF92DFD0953E2" bold="true" box="[636,659,1729,1756]" italics="true" pageId="22" pageNumber="23">ib</emphasis>
exceeding seminal region of 
<emphasis id="B925EABE1325B1774CBBF92DFB9B53E2" bold="true" box="[1005,1025,1729,1756]" italics="true" pageId="22" pageNumber="23">sl</emphasis>
(
<figureCitation id="136A2A291325B1774B47F92EFB0453E2" box="[1041,1182,1730,1756]" captionStart="Fig" captionStartId="161.[189,232,1668,1694]" captionTargetBox="[189,1398,269,1626]" captionTargetId="figure-11@161.[189,1398,269,1626]" captionTargetPageId="161" captionText="Fig. 47. Pseudonannolene ambuatinga Iniesta &amp; Ferreira, 2013, ♂ (IBSP 3442). A. First leg-pair. B. Detail of prefemur. C. Second leg-pair. D. Detail of telopodites, in anal view. E. Gonopods, in anal view. F. Gonopods, in oral view. Abbreviations: see Material and methods. Scale bars: A, C, E–F = 0.5 mm; B, D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7891141" httpUri="https://zenodo.org/record/7891141/files/figure.png" pageId="22" pageNumber="23">Fig. 47D–F</figureCitation>
).
</paragraph>
<paragraph id="8BEE36AC1325B1774FEBF8EFFBB55201" blockId="22.[187,1398,1794,1855]" pageId="22" pageNumber="23">
<smallCapsWord id="8D08A0701325B1774FEBF8EFFE8E5225" baselines="1814,1814" box="[189,276,1795,1821]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Vulvae" pageId="22" pageNumber="23">VULVAE</smallCapsWord>
. As typical for the genus. Bursa subtriangular, glabrous (
<figureCitation id="136A2A291325B1774CF5F8EEFB815222" box="[931,1051,1794,1820]" captionStart="Fig" captionStartId="280.[189,232,1787,1813]" captionTargetBox="[250,1337,265,1739]" captionTargetId="figure-14@280.[250,1337,265,1739]" captionTargetPageId="280" captionText="Fig. 177. Stereoscopic images of left vulva of species of Pseudonannolene Silvestri, 1895, in oral view. A. P. albiventris Schubart, 1952. B. P. alegrensis Silvestri, 1897. C. P. ambuatinga Iniesta &amp; Ferreira, 2013. D. P. anapophysis Fontanetti, 1996. E. P. buhrnheimi Schubart, 1960. F. P. caatinga Iniesta &amp; Ferreira, 2014. G. P. callipyge Brölemann, 1902. H. P. erikae Iniesta &amp; Ferreira, 2014. I. P. fontanettiae Iniesta &amp; Ferreira, 2014. J. P. halophila Schubart, 1949. K. P. imbirensis Fontanetti, 1996. L. P. inops Brölemann, 1929. M. P. leucocephalus Schubart, 1944. N. P. longicornis (Porat, 1888). O. P. lundi Iniesta &amp; Ferreira, 2015. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.7891439" httpUri="https://zenodo.org/record/7891439/files/figure.png" pageId="22" pageNumber="23">Fig. 177C</figureCitation>
); internal valve subtriangular; operculum narrow, not curved medially; external valve wide, subtriangular.
</paragraph>
</subSubSection>
<subSubSection id="C34B65271325B1764FEBF889FD605456" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="distribution">
<paragraph id="8BEE36AC1325B1774FEBF889FECC5241" blockId="22.[189,342,1893,1919]" box="[189,342,1893,1919]" pageId="22" pageNumber="23">
<heading id="D0A681C01325B1774FEBF889FECC5241" bold="true" box="[189,342,1893,1919]" fontSize="11" level="3" pageId="22" pageNumber="23" reason="3">
<emphasis id="B925EABE1325B1774FEBF889FECC5241" bold="true" box="[189,342,1893,1919]" pageId="22" pageNumber="23">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1325B1764FEBF87FFD605456" blockId="22.[189,1399,1939,2000]" lastBlockId="23.[189,1399,265,360]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
A troglomorphic species known only from caves in the Karst region of Pains and surrounding municipalities (Arcos-Pains-Doresópolis speleological unit), state of 
<collectingRegion id="4995F84E1325B1774CA0F859FB0252F1" box="[1014,1176,1973,1999]" country="Brazil" name="Minas Gerais" pageId="22" pageNumber="23">Minas Gerais</collectingRegion>
, 
<collectingCountry id="F346763C1325B1774BF0F859FB7452F1" box="[1190,1262,1973,1999]" name="Brazil" pageId="22" pageNumber="23">Brazil</collectingCountry>
(
<figureCitation id="136A2A291325B1774A56F859FAF352EE" box="[1280,1385,1973,2000]" captionStart="Fig" captionStartId="283.[189,232,1363,1389]" captionTargetBox="[192,1368,285,1297]" captionTargetId="figure-12@283.[189,1398,265,1328]" captionTargetPageId="283" captionText="Fig. 180. Distribution map of the species Pseudonannolene albiventris Schubart, 1952, P. alegrensis Silvestri, 1897, P. ambuatinga Iniesta &amp; Ferreira, 2013, P. anapophysis Fontanetti, 1996, and P. borelli Silvestri, 1895." figureDoi="http://doi.org/10.5281/zenodo.7891445" httpUri="https://zenodo.org/record/7891445/files/figure.png" pageId="22" pageNumber="23">Fig. 180</figureCitation>
). This karst, which comprises the highest density of caves known for South America, harbors many other undescribed and described cave-dwelling species (
<bibRefCitation id="EFC04B5D1324B1764C06FEC0FBEF547B" author="Alvares E. S. S. &amp; Ferreira R. L." box="[848,1141,299,328]" pageId="23" pageNumber="24" pagination="41 - 43" refId="ref99054" refString="Alvares E. S. S. &amp; Ferreira R. L. 2002. Coarazuphium pains, a new species of troglobitic beetle from Brazil (Coleoptera: Carabidae: Zuphiini). Lundiana 3 (1): 41 - 43." type="journal article" year="2002">Álvares &amp; Ferreira 2002</bibRefCitation>
; 
<bibRefCitation id="EFC04B5D1324B1764BD7FEC7FAE8547B" author="Parizotto D. R. &amp; Pires A. C. &amp; Mise K. M. &amp; Ferreira R. L. &amp; Sessegolo G. C." box="[1153,1394,299,326]" pageId="23" pageNumber="24" pagination="401 - 409" refId="ref106238" refString="Parizotto D. R., Pires A. C., Mise K. M., Ferreira R. L. &amp; Sessegolo G. C. 2017. Troglobitic invertebrates: Improving the knowledge on the Brazilian subterranean biodiversity through an interactive multi-entry key. Zootaxa 4365 (4): 401 - 409. https: // doi. org / 10.11646 / zootaxa. 4365.4.1" type="journal article" year="2017">
Parizotto 
<emphasis id="B925EABE1324B1764BA4FEC0FABF547B" box="[1266,1317,299,325]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
. 2017
</bibRefCitation>
; 
<bibRefCitation id="EFC04B5D1324B1764FEBFEA2FE7C5456" author="Gallao J. E. &amp; Bichuette M. E." box="[189,486,334,360]" pageId="23" pageNumber="24" pagination="1 - 23" refId="ref102130" refString="Gallao J. E. &amp; Bichuette M. E. 2018. Brazilian obligatory subterranean fauna and threats to the hypogean environment. ZooKeys 746: 1 - 23. https: // doi. org / 10.3897 / zookeys. 746.15140" type="journal article" year="2018">Gallão &amp; Bichuette 2018</bibRefCitation>
; 
<bibRefCitation id="EFC04B5D1324B1764EA5FEA2FD745456" author="Pellegrini T. G. &amp; Ferreira R. L. &amp; Zampaulo R. A. &amp; Vieira L." box="[499,750,334,360]" pageId="23" pageNumber="24" pagination="287 - 304" refId="ref106308" refString="Pellegrini T. G., Ferreira R. L., Zampaulo R. A. &amp; Vieira L. 2020. Coarazuphium lundi (Carabidae: Zuphiini), a new Brazilian troglobitic beetle, with the designation of a neotype for C. pains Alvares &amp; Ferreira, 2002. Zootaxa 4878 (2): 287 - 304. https: // doi. org / 10.11646 / zootaxa. 4878.2.4" type="journal article" year="2020">
Pellegrini 
<emphasis id="B925EABE1324B1764D3AFEA3FD055456" box="[620,671,334,360]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
. 2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B65271324B1764FEBFE61FBE05726" pageId="23" pageNumber="24" type="discussion">
<paragraph id="8BEE36AC1324B1764FEBFE61FEDE5499" blockId="23.[189,324,397,423]" box="[189,324,397,423]" pageId="23" pageNumber="24">
<heading id="D0A681C01324B1764FEBFE61FEDE5499" bold="true" box="[189,324,397,423]" fontSize="11" level="3" pageId="23" pageNumber="24" reason="3">
<emphasis id="B925EABE1324B1764FEBFE61FEDE5499" bold="true" box="[189,324,397,423]" pageId="23" pageNumber="24">Comments</emphasis>
</heading>
</paragraph>
<paragraph id="8BEE36AC1324B1764FEBFE56FBE05726" blockId="23.[189,1399,442,536]" pageId="23" pageNumber="24">
Although the examination of the 
<typeStatus id="54EA880E1324B1764D0FFE57FD1654EB" box="[601,652,443,469]" pageId="23" pageNumber="24">type</typeStatus>
material of the junior synonym 
<taxonomicName id="4C514D2F1324B1764B77FE57FB3354EA" authorityName="Gallo &amp; Bichuette" authorityYear="2020" box="[1057,1193,443,468]" class="Diplopoda" family="Pseudonannolenidae" genus="Pseudonannolene" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="canastra">
<emphasis id="B925EABE1324B1764B77FE57FB3354EA" box="[1057,1193,443,468]" italics="true" pageId="23" pageNumber="24">P. canastra</emphasis>
</taxonomicName>
deposited at the Laboratório de Estudos Subterrâneos (LES/UFSCar) was not possible during this study, the original description and figures provided by 
<bibRefCitation id="EFC04B5D1324B1764D32FE12FC0A5726" author="Gallo J. S. &amp; Bichuette M. E." box="[612,912,510,536]" pageId="23" pageNumber="24" pagination="33 - 47" refId="ref102297" refString="Gallo J. S. &amp; Bichuette M. E. 2020. Pseudonannolene canastra sp. nov. (Diplopoda, Spirostreptida) - A new troglobitic millipede from the southwestern state of Minas Gerais, Brazil. Subterranean Biology 35: 33 - 47. https: // doi. org / 10.3897 / subtbiol. 35.51183" type="journal article" year="2020">Gallo &amp; Bichuette (2020)</bibRefCitation>
are highly detailed.
</paragraph>
</subSubSection>
</treatment>
</document>