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<mods:title id="28297445C20A39CF8244E11679EA604C">Phylogeny, taxonomy, and biogeography of the Sphaeriinae (Bivalvia: Sphaeriidae)</mods:title>
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<mods:affiliation id="07BBC6DBE39921A5A00D38294DEAFF49">Laboratory of Macroecology &amp; Biogeography of Invertebrates, Saint Petersburg State University, Universitetskaya Emb. 7 - 9, 199034 Saint Petersburg, Russian Federation</mods:affiliation>
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<mods:affiliation id="20BA81A59855DF6438CDF9832872C825">N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russian Federation</mods:affiliation>
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<mods:date id="2B3C39C0932DF1C331C19FB89E112FD3">2024</mods:date>
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<taxonomicName id="4C714D221828FFE53C16FCAAA5461A4C" ID-CoL="3C78F" authority="(Kuiper, 1975)" baseAuthorityName="Kuiper" baseAuthorityYear="1975" box="[304,601,793,820]" class="Bivalvia" family="Sphaeriidae" genus="Euglesa" kingdom="Animalia" order="Sphaeriida" pageId="18" pageNumber="323" phylum="Mollusca" rank="species" species="hinzi">
<emphasis id="B905EAB31828FFE53C16FCAAA5461A4C" bold="true" box="[304,601,793,820]" pageId="18" pageNumber="323">
<emphasis id="B905EAB31828FFE53C16FCAAA6AA1A4B" bold="true" box="[304,437,793,820]" italics="true" pageId="18" pageNumber="323">Euglesa hinzi</emphasis>
(Kuiper, 1975)
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(
<figureCitation id="134A2A241828FFE53D80FCF3A7F21A27" box="[166,237,832,856]" captionStart="Figure 9" captionStartId="19.[112,177,1463,1487]" captionTargetBox="[114,1458,145,1436]" captionTargetId="figure-216@19.[114,1458,144,1436]" captionTargetPageId="19" captionText="Figure 9. Shells of species, re-described in this research: (A) Euglesa khurbaensis (RMBH No. MSph-707, Russia, Primorsky Territory); (B) E. khurbaensis (RMBH No.MSph-707/2, Russia, Primorsky Territory); (C) E.lindholmi (RMBH No. MSph-370, Russia, Gydan Peninsula); (D) E. ljovushkini (RMBH No. MSph-335, Abkhazia); (E) E. ljovushkini (RMBH No. MSph-335/2, Abkhazia); (F) E. hinzi (RMBH No. MSph- 366, Russia, Gydan Peninsula); (G) E. subterranea (RMBH No. MSph-295, Georgia, Imereti); (H) E. nordenskioldi (Russia, Vaygach Island). Photos: O.V. Aksenova.Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13220108" httpUri="https://zenodo.org/record/13220108/files/figure.png" pageId="18" pageNumber="323">Fig. 9F</figureCitation>
)
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<taxonomicName id="4C714D221828FFE53DA7FCECA6841A08" authority="Kuiper, 1975" authorityName="Kuiper" authorityYear="1975" box="[129,411,863,887]" class="Bivalvia" family="Sphaeriidae" genus="Pisidium" kingdom="Animalia" order="Sphaeriida" pageId="18" pageNumber="323" phylum="Mollusca" rank="species" species="hinzi">
<emphasis id="B905EAB31828FFE53DA7FCECA6111A08" box="[129,270,863,887]" italics="true" pageId="18" pageNumber="323">Pisidium hinzi</emphasis>
Kuiper, 1975
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: text 27–31, figs 1–6, 28, 30.
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<emphasis id="B905EAB31828FFE53DA7FC2AA7EF1ACF" box="[129,240,921,944]" italics="true" pageId="18" pageNumber="323">Type series:</emphasis>
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:
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No. 192849, and
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<collectionCode id="ED60AE641828FFE53FF1FC2AA41A1ACF" box="[727,773,921,944]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="18" pageNumber="323" type="Museum">ZIN</collectionCode>
, nos 1,
<quantity id="4C899B441828FFE53DE4FC0BA7F71AAF" box="[194,232,952,976]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="18" pageNumber="323" unit="in" value="2.0">2 in</quantity>
the systematic catalogue.
</materialsCitation>
</paragraph>
<paragraph id="8BCE36A11828FFE53DBAFC6BA7E11D51" blockId="18.[129,778,921,1070]" pageId="18" pageNumber="323">
<materialsCitation id="3B193CFC1828FFE53DBAFC6BA7E51D51" collectorName="Varangerhalbiusel &amp; NO Norwegens'" country="Norway" elevation="147" location="Buetjorni" pageId="18" pageNumber="323" specimenCount="1" typeStatus="holotype">
<emphasis id="B905EAB31828FFE53DBAFC6BA63B1A90" box="[156,292,983,1007]" italics="true" pageId="18" pageNumber="323">Type locality:</emphasis>
<collectingCountry id="F36676311828FFE53C09FC6BA69D1A8F" box="[303,386,984,1008]" name="Norway" pageId="18" pageNumber="323">Norway</collectingCountry>
, ‘
<location id="8EAE607A1828FFE53CB2FC64A6E81A90" LSID="urn:lsid:plazi:treatment:03D887B71828FFE53C16FCAAA3781859:8EAE607A1828FFE53CB2FC64A6E81A90" box="[404,503,983,1007]" country="Norway" name="Buetjorni" pageId="18" pageNumber="323">Buetjörni</location>
(=Davggeluobal), 147 m NN, Flachsee (tiefste Stelle 9 m),
<collectorName id="268453771828FFE53CDCFC44A5AF1D70" box="[506,688,1015,1039]" pageId="18" pageNumber="323">Varangerhalbiusel</collectorName>
im
<collectorName id="268453771828FFE53FC7FC44A7E51D51" pageId="18" pageNumber="323">NO Norwegens’</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BCE36A11828FFE53DA7FBFCA5F31C7B" blockId="18.[129,778,1103,1284]" pageId="18" pageNumber="323">
<emphasis id="B905EAB31828FFE53DA7FBFCA6241D18" box="[129,315,1103,1127]" italics="true" pageId="18" pageNumber="323">Material examined:</emphasis>
<materialsCitation id="3B193CFC1828FFE53C6FFBFCA5F81C7B" collectingDate="2017-07" collectingDateMax="2019-08" collectingDateMin="2017-07" collectionCode="RMBH" country="Russia" county="River Basin" location="Yakutia" municipality="Yana River" pageId="18" pageNumber="323" specimenCount="27">
Russia:
<location id="8EAE607A1828FFE53CB1FBFCA6FD1D18" LSID="urn:lsid:plazi:treatment:03D887B71828FFE53C16FCAAA3781859:8EAE607A1828FFE53CB1FBFCA6FD1D18" box="[407,482,1103,1127]" country="Russia" county="River Basin" municipality="Yana River" name="Yakutia" pageId="18" pageNumber="323">Yakutia</location>
,
<location id="8EAE607A1828FFE53CCAFBFCA54E1D18" LSID="urn:lsid:plazi:treatment:03D887B71828FFE53C16FCAAA3781859:8EAE607A1828FFE53CCAFBFCA54E1D18" box="[492,593,1103,1127]" country="Russia" county="River Basin" municipality="Yana River" name="Ust-Kuiga" pageId="18" pageNumber="323">Ust-Kuiga</location>
,
<location id="8EAE607A1828FFE53F7DFBE3A4191D18" LSID="urn:lsid:plazi:treatment:03D887B71828FFE53C16FCAAA3781859:8EAE607A1828FFE53F7DFBE3A4191D18" box="[603,774,1103,1127]" country="Russia" county="River Basin" municipality="Yana River" name="Yana River Basin" pageId="18" pageNumber="323">
<collectingMunicipality id="6BAAACDB1828FFE53F7DFBE3A5D81D18" box="[603,711,1103,1127]" pageId="18" pageNumber="323">
Yana
<collectingCounty id="62AF4E2D1828FFE53FB7FBFCA5D81D18" box="[657,711,1103,1127]" pageId="18" pageNumber="323">River</collectingCounty>
</collectingMunicipality>
Basin
</location>
,
<date id="FFCF10611828FFE53DA7FBDCA7FB1DF8" box="[129,228,1135,1159]" pageId="18" pageNumber="323" value="2019-08">
<collectingDate id="EF8BE9891828FFE53DA7FBDCA7FB1DF8" box="[129,228,1135,1159]" pageId="18" pageNumber="323" value="2019-08">VIII.2019</collectingDate>
</date>
,
<collectorName id="268453771828FFE53DC8FBDCA6321DF8" box="[238,301,1135,1159]" pageId="18" pageNumber="323">Zubrii</collectorName>
leg.,
<specimenCount id="9D77FD281828FFE53C47FBDCA6F21DF8" box="[353,493,1135,1159]" count="1" pageId="18" pageNumber="323" type="generic">one specimen</specimenCount>
(lot no.
<collectionCode id="ED60AE641828FFE53F63FBDCA58F1DF9" box="[581,656,1135,1158]" pageId="18" pageNumber="323">RMBH</collectionCode>
MSph-572- 2); Gydan Peninsula
<date id="FFCF10611828FFE53C7CFB3DA6AA1DD9" box="[346,437,1166,1190]" pageId="18" pageNumber="323" value="2017-07">
<collectingDate id="EF8BE9891828FFE53C7CFB3DA6AA1DD9" box="[346,437,1166,1190]" pageId="18" pageNumber="323" value="2017-07">VII.2017</collectingDate>
</date>
,
<collectorName id="268453771828FFE53CE7FB3DA5031DD9" box="[449,540,1166,1190]" pageId="18" pageNumber="323">Sokolova</collectorName>
&amp;
<collectorName id="268453771828FFE53F66FB3DA5A61DD9" box="[576,697,1166,1190]" pageId="18" pageNumber="323">Shevchenko</collectorName>
leg.,
<specimenCount id="9D77FD281828FFE53FD6FB3DA7F61DBA" count="25" pageId="18" pageNumber="323" type="generic">25 specimens</specimenCount>
(lot nos
<collectionCode id="ED60AE641828FFE53C6CFB1DA68A1DBA" box="[330,405,1198,1221]" pageId="18" pageNumber="323">RMBH</collectionCode>
MSph-361, MSph-366, MSph-374), VashutkinylakesVIII.2016,
<collectorName id="268453771828FFE53CA9FB7EA6ED1D9A" box="[399,498,1229,1253]" pageId="18" pageNumber="323">Aksenova</collectorName>
,
<collectorName id="268453771828FFE53CDFFB7EA5501D9A" box="[505,591,1229,1253]" pageId="18" pageNumber="323">Aksenov</collectorName>
,
<collectorName id="268453771828FFE53F73FB7EA5B71D9A" box="[597,680,1229,1253]" pageId="18" pageNumber="323">Bespaliy</collectorName>
,
<collectorName id="268453771828FFE53F88FB7EA4161D9A" box="[686,777,1229,1253]" pageId="18" pageNumber="323">Sokolova</collectorName>
&amp;
<collectorName id="268453771828FFE53DBEFB5FA60E1C7B" box="[152,273,1260,1284]" pageId="18" pageNumber="323">Shevchenko</collectorName>
leg.,
<specimenCount id="9D77FD281828FFE53C67FB5FA6D51C7B" box="[321,458,1260,1284]" count="1" pageId="18" pageNumber="323" type="generic">one specimen</specimenCount>
(lot no.
<collectionCode id="ED60AE641828FFE53F3DFB5FA5771C7C" box="[539,616,1260,1283]" pageId="18" pageNumber="323">RMBH</collectionCode>
, MSph-161)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCE36A11828FFE53DA7FA95A5C51CA5" blockId="18.[129,778,1317,1498]" pageId="18" pageNumber="323">
<emphasis id="B905EAB31828FFE53DA7FA95A7E81C42" box="[129,247,1318,1341]" italics="true" pageId="18" pageNumber="323">Description:</emphasis>
The mean shell length ± SD is 2.4 ±
<quantity id="4C899B441828FFE53F97FA96A4161C42" box="[689,777,1317,1341]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="323" unit="mm" value="0.2">0.2 mm</quantity>
(
<emphasis id="B905EAB31828FFE53DAAFAF6A7801C23" box="[140,159,1349,1372]" italics="true" pageId="18" pageNumber="323">N</emphasis>
= 15), shell is rounded in shape (
<figureCitation id="134A2A241828FFE53F37FAF6A5441C22" box="[529,603,1349,1373]" captionStart="Figure 9" captionStartId="19.[112,177,1463,1487]" captionTargetBox="[114,1458,145,1436]" captionTargetId="figure-216@19.[114,1458,144,1436]" captionTargetPageId="19" captionText="Figure 9. Shells of species, re-described in this research: (A) Euglesa khurbaensis (RMBH No. MSph-707, Russia, Primorsky Territory); (B) E. khurbaensis (RMBH No.MSph-707/2, Russia, Primorsky Territory); (C) E.lindholmi (RMBH No. MSph-370, Russia, Gydan Peninsula); (D) E. ljovushkini (RMBH No. MSph-335, Abkhazia); (E) E. ljovushkini (RMBH No. MSph-335/2, Abkhazia); (F) E. hinzi (RMBH No. MSph- 366, Russia, Gydan Peninsula); (G) E. subterranea (RMBH No. MSph-295, Georgia, Imereti); (H) E. nordenskioldi (Russia, Vaygach Island). Photos: O.V. Aksenova.Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13220108" httpUri="https://zenodo.org/record/13220108/files/figure.png" pageId="18" pageNumber="323">Fig. 9F</figureCitation>
). The beaks are rounded, slightly prominent, offset to the posterior edge of the valves (
<figureCitation id="134A2A241828FFE53DEBFA30A63C1CE4" box="[205,291,1411,1435]" captionStart="Figure 10" captionStartId="20.[128,193,1856,1880]" captionTargetBox="[353,1249,143,1832]" captionTargetId="figure-9@20.[353,1249,143,1832]" captionTargetPageId="20" captionText="Figure 10. Morphology and anatomy of re-described species discussed in this research (teeth morphology and shell pores, anatomy of gills and siphons): (A) Euglesa hinzi (RMBH No. MSph-366, Russia, Gydan Peninsula); (B) E. khurbaensis (RMBH No. MSph-707/2, Russia, Primorsky Territory); (C) E.lindholmi (RMBH No.MSph-370, Russia, Gydan Peninsula); (D) E. ljovushkini (RMBH No. MSph-335, Abkhazia); (E) E. nordenskioldi (Russia, Vaygach Island); (F) E.subterranea (RMBH No. MSph-295, Georgia, Imereti). Photos: O.V. Aksenova. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13220112" httpUri="https://zenodo.org/record/13220112/files/figure.png" pageId="18" pageNumber="323">Fig. 10A</figureCitation>
). The cardinal tooth of the right valve is curved, with a branched posterior end. The cardinal teeth of the left valve are slightly curved, almost parallel to each other (
<figureCitation id="134A2A241828FFE53F52FA71A5D51CA5" box="[628,714,1474,1498]" captionStart="Figure 10" captionStartId="20.[128,193,1856,1880]" captionTargetBox="[353,1249,143,1832]" captionTargetId="figure-9@20.[353,1249,143,1832]" captionTargetPageId="20" captionText="Figure 10. Morphology and anatomy of re-described species discussed in this research (teeth morphology and shell pores, anatomy of gills and siphons): (A) Euglesa hinzi (RMBH No. MSph-366, Russia, Gydan Peninsula); (B) E. khurbaensis (RMBH No. MSph-707/2, Russia, Primorsky Territory); (C) E.lindholmi (RMBH No.MSph-370, Russia, Gydan Peninsula); (D) E. ljovushkini (RMBH No. MSph-335, Abkhazia); (E) E. nordenskioldi (Russia, Vaygach Island); (F) E.subterranea (RMBH No. MSph-295, Georgia, Imereti). Photos: O.V. Aksenova. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13220112" httpUri="https://zenodo.org/record/13220112/files/figure.png" pageId="18" pageNumber="323">Fig. 10A</figureCitation>
).
</paragraph>
<paragraph id="8BCE36A11828FFE53DA7FA48A52B1F6C" blockId="18.[129,564,1531,1555]" box="[129,564,1531,1555]" pageId="18" pageNumber="323">
<emphasis id="B905EAB31828FFE53DA7FA48A7E11F6C" box="[129,254,1531,1555]" italics="true" pageId="18" pageNumber="323">Distribution:</emphasis>
Northern Europe and Siberia.
</paragraph>
<paragraph id="8BCE36A11828FFE53DA7F986A3781859" blockId="18.[129,778,1589,2020]" lastBlockId="18.[825,1474,144,294]" pageId="18" pageNumber="323">
<emphasis id="B905EAB31828FFE53DA7F986A7EE1F33" box="[129,241,1589,1612]" italics="true" pageId="18" pageNumber="323">Comments:</emphasis>
We analysed the original descriptions and type specimens of some nominal species described from Eurasian localities where our speciments were found or from closely situated localities:
<taxonomicName id="4C714D221828FFE53C03F920A7DE1FB5" authority="(Starobogatov &amp; Streletzkaja, 1967)" baseAuthorityName="Starobogatov &amp; Streletzkaja" baseAuthorityYear="1967" class="Bivalvia" family="Sphaeriidae" genus="Euglesa" kingdom="Animalia" order="Sphaeriida" pageId="18" pageNumber="323" phylum="Mollusca" rank="species" species="jacutica">
<emphasis id="B905EAB31828FFE53C03F920A6DA1FD5" box="[293,453,1683,1707]" italics="true" pageId="18" pageNumber="323">Euglesa jacutica</emphasis>
(Starobogatov &amp; Streletzkaja, 1967)
</taxonomicName>
,
<taxonomicName id="4C714D221828FFE53DEAF901A5AA1FB5" authority="Dolgin &amp; Starobogatov, 1973" authorityName="Dolgin &amp; Starobogatov" authorityYear="1973" box="[204,693,1714,1738]" class="Bivalvia" family="Sphaeriidae" genus="Euglesa" kingdom="Animalia" order="Sphaeriida" pageId="18" pageNumber="323" phylum="Mollusca" rank="species" species="jamalensis">
<emphasis id="B905EAB31828FFE53DEAF901A69E1FB5" box="[204,385,1714,1738]" italics="true" pageId="18" pageNumber="323">Euglesa jamalensis</emphasis>
Dolgin &amp; Starobogatov, 1973
</taxonomicName>
,
<taxonomicName id="4C714D221828FFE53F99F901A5061F96" authority="Dolgin &amp; Korniushin, 1994" authorityName="Dolgin &amp; Korniushin" authorityYear="1994" class="Bivalvia" family="Sphaeriidae" genus="Euglesa" kingdom="Animalia" order="Sphaeriida" pageId="18" pageNumber="323" phylum="Mollusca" rank="species" species="johanseni">
<emphasis id="B905EAB31828FFE53F99F901A7C31F96" italics="true" pageId="18" pageNumber="323">Euglesa johanseni</emphasis>
Dolgin &amp; Korniushin, 1994
</taxonomicName>
,
<taxonomicName id="4C714D221828FFE53F0FF962A7A81E77" authority="Kuiper, 1975" authorityName="Kuiper" authorityYear="1975" class="Bivalvia" family="Sphaeriidae" genus="Euglesa" kingdom="Animalia" order="Sphaeriida" pageId="18" pageNumber="323" phylum="Mollusca" rank="species" species="hinzi">
<emphasis id="B905EAB31828FFE53F0FF962A5AD1F96" box="[553,690,1745,1769]" italics="true" pageId="18" pageNumber="323">Euglesa hinzi</emphasis>
Kuiper, 1975
</taxonomicName>
, and
<taxonomicName id="4C714D221828FFE53DCBF943A7AA1E57" authority="Starobogatov, Anistratenko &amp; Dolgin, 1990" authorityName="Starobogatov, Anistratenko &amp; Dolgin" authorityYear="1990" class="Bivalvia" family="Sphaeriidae" genus="Euglesa" kingdom="Animalia" order="Sphaeriida" pageId="18" pageNumber="323" phylum="Mollusca" rank="species" species="obensis">
<emphasis id="B905EAB31828FFE53DCBF943A69E1E77" box="[237,385,1776,1800]" italics="true" pageId="18" pageNumber="323">Euglesa obensis</emphasis>
Starobogatov, Anistratenko &amp; Dolgin, 1990
</taxonomicName>
(Supporting Information,
<figureCitation id="134A2A241828FFE53CF3F8A3A5001E57" box="[469,543,1808,1832]" captionStart="Figure 1" captionStartId="6.[129,191,1836,1860]" captionTargetBox="[186,1417,145,1801]" captionTargetId="figure-88@6.[185,1417,144,1808]" captionTargetPageId="6" captionText="Figure 1.Spatial distributionofthe numberofavailable 16S rRNA sequences (A), and species richness of thesubfamily Sphaeriinae (B), in theworld." figureDoi="http://doi.org/10.5281/zenodo.13220078" httpUri="https://zenodo.org/record/13220078/files/figure.png" pageId="18" pageNumber="323">Fig. S1</figureCitation>
). The molecular genetic data (16S rRNA sequences) obtained have no analogues in GenBank, with
<taxonomicName id="4C714D221828FFE53C73F8FDA6E51E19" baseAuthorityName="Clessin" baseAuthorityYear="1873" box="[341,506,1870,1894]" class="Bivalvia" family="Sphaeriidae" genus="Euglesa" kingdom="Animalia" order="Sphaeriida" pageId="18" pageNumber="323" phylum="Mollusca" rank="species" species="parvula">
<emphasis id="B905EAB31828FFE53C73F8FDA6E51E19" box="[341,506,1870,1894]" italics="true" pageId="18" pageNumber="323">Euglesa parvula</emphasis>
</taxonomicName>
(Clessin in Westerlund, 1873) being the closest ally. Morphologically, nominal species in this group have similar conchological features with the
<taxonomicName id="4C714D221828FFE53DA7F81EA7D01EBB" authorityName="Kuiper" authorityYear="1975" box="[129,207,1964,1988]" class="Bivalvia" family="Sphaeriidae" genus="Euglesa" kingdom="Animalia" order="Sphaeriida" pageId="18" pageNumber="323" phylum="Mollusca" rank="species" species="hinzi">
<emphasis id="B905EAB31828FFE53DA7F81EA7D01EBB" box="[129,207,1964,1988]" italics="true" pageId="18" pageNumber="323">E. hinzi</emphasis>
</taxonomicName>
group, i.e. the section
<taxonomicName id="4C714D221828FFE53C99F81EA7AA1E9B" authority="Dolgin &amp; Korniushin, 1993" authorityName="Dolgin &amp; Korniushin" authorityYear="1993" class="Bivalvia" family="Sphaeriidae" genus="Euglesa" kingdom="Animalia" order="Sphaeriida" pageId="18" pageNumber="323" phylum="Mollusca" rank="subGenus" subGenus="Hinziana">
<emphasis id="B905EAB31828FFE53C99F81EA5031EBB" box="[447,540,1965,1988]" italics="true" pageId="18" pageNumber="323">Hinziana</emphasis>
Dolgin &amp; Korniushin, 1993
</taxonomicName>
(
<bibRefCitation id="EFE04B501828FFE53DEDF87FA69F1E9B" author="Korniushin AV" box="[203,384,1996,2020]" pageId="18" pageNumber="323" refId="ref27522" refString="Korniushin AV. Bivalve Molluscs of the Superfamily Pisidioidea in the Palaearctic Region: Fauna, Systematics, Phylogeny. Kiev: Schmalhausen Institute of Zoology Press, 1996 [in Russian]." type="book" year="1996">Korniushin 1996</bibRefCitation>
). Thus, according to morphological features and molecular-genetic analysis, this species shoud be classified as
<taxonomicName id="4C714D221828FFE53E9FFF03A31919B8" authorityName="Kuiper" authorityYear="1975" box="[953,1030,175,199]" class="Bivalvia" family="Sphaeriidae" genus="Euglesa" kingdom="Animalia" order="Sphaeriida" pageId="18" pageNumber="323" phylum="Mollusca" rank="species" species="hinzi">
<emphasis id="B905EAB31828FFE53E9FFF03A31919B8" box="[953,1030,175,199]" italics="true" pageId="18" pageNumber="323">E. hinzi</emphasis>
</taxonomicName>
(
<figureCitation id="134A2A241828FFE5393EFF1CA37F19B8" box="[1048,1120,175,199]" captionStart="Figure 9" captionStartId="19.[112,177,1463,1487]" captionTargetBox="[114,1458,145,1436]" captionTargetId="figure-216@19.[114,1458,144,1436]" captionTargetPageId="19" captionText="Figure 9. Shells of species, re-described in this research: (A) Euglesa khurbaensis (RMBH No. MSph-707, Russia, Primorsky Territory); (B) E. khurbaensis (RMBH No.MSph-707/2, Russia, Primorsky Territory); (C) E.lindholmi (RMBH No. MSph-370, Russia, Gydan Peninsula); (D) E. ljovushkini (RMBH No. MSph-335, Abkhazia); (E) E. ljovushkini (RMBH No. MSph-335/2, Abkhazia); (F) E. hinzi (RMBH No. MSph- 366, Russia, Gydan Peninsula); (G) E. subterranea (RMBH No. MSph-295, Georgia, Imereti); (H) E. nordenskioldi (Russia, Vaygach Island). Photos: O.V. Aksenova.Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13220108" httpUri="https://zenodo.org/record/13220108/files/figure.png" pageId="18" pageNumber="323">Fig. 9F</figureCitation>
). The species status of
<taxonomicName id="4C714D221828FFE53870FF03A2A319B8" baseAuthorityName="Starobogatov &amp; Streletzkaja" baseAuthorityYear="1967" box="[1366,1468,176,199]" class="Bivalvia" family="Sphaeriidae" genus="Euglesa" kingdom="Animalia" order="Sphaeriida" pageId="18" pageNumber="323" phylum="Mollusca" rank="species" species="jacutica">
<emphasis id="B905EAB31828FFE53870FF03A2A319B8" box="[1366,1468,176,199]" italics="true" pageId="18" pageNumber="323">E. jacutica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C714D221828FFE53E1FFF7CA4A81999" authorityName="Dolgin &amp; Starobogatov" authorityYear="1973" box="[825,951,206,230]" class="Bivalvia" family="Sphaeriidae" genus="Euglesa" kingdom="Animalia" order="Sphaeriida" pageId="18" pageNumber="323" phylum="Mollusca" rank="species" species="jamalensis">
<emphasis id="B905EAB31828FFE53E1FFF7CA4A81999" box="[825,951,206,230]" italics="true" pageId="18" pageNumber="323">E. jamalensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C714D221828FFE53EE7FF7CA32A1999" authorityName="Dolgin &amp; Korniushin" authorityYear="1994" box="[961,1077,206,230]" class="Bivalvia" family="Sphaeriidae" genus="Euglesa" kingdom="Animalia" order="Sphaeriida" pageId="18" pageNumber="323" phylum="Mollusca" rank="species" species="johanseni">
<emphasis id="B905EAB31828FFE53EE7FF7CA32A1999" box="[961,1077,206,230]" italics="true" pageId="18" pageNumber="323">E. johanseni</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C714D221828FFE5394DFF7CA3D71999" authorityName="Starobogatov, Anistratenko &amp; Dolgin" authorityYear="1990" box="[1131,1224,206,230]" class="Bivalvia" family="Sphaeriidae" genus="Euglesa" kingdom="Animalia" order="Sphaeriida" pageId="18" pageNumber="323" phylum="Mollusca" rank="species" species="obensis">
<emphasis id="B905EAB31828FFE5394DFF7CA3D71999" box="[1131,1224,206,230]" italics="true" pageId="18" pageNumber="323">E. obensis</emphasis>
</taxonomicName>
needs to be clarified and cannot be solved here, since molecular studies of the
<typeStatus id="54CA88031828FFE5387DFF5DA2DD1879" box="[1371,1474,238,262]" pageId="18" pageNumber="323" type="topotype">topotypes</typeStatus>
of these species are necessary.
</paragraph>
</subSubSection>
</treatment>
</document>
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