176 lines
20 KiB
XML
176 lines
20 KiB
XML
<document id="CA62C09B76B15CB358552111586499D0" ID-DOI="10.11646/phytotaxa.324.1.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="13697396" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725568149749" checkinUser="felipe" docAuthor="Elven, Reidar, Pedersen, A. Tiril M. & Bjorå, Charlotte S." docDate="2017" docId="038E0A32FFC3FFE64CECFA31FB69A979" docLanguage="en" docName="phytotaxa.324.1.4.pdf" docOrigin="Phytotaxa 324 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.324.1.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Carex saamica T. M. Pedersen & Elven 2017" docType="treatment" docVersion="2" lastPageNumber="70" masterDocId="FFB7724AFFC5FFE14C64FF85FFC4AB3A" masterDocTitle="The nomenclature of two hybrid taxa in Carex sect. Vesicariae (Cyperaceae) currently assigned as Carex rostrata var. borealis and Carex stenolepis" masterLastPageNumber="72" masterPageNumber="63" pageNumber="69" updateTime="1725571143640" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B247072C136CEAD60DB98F0F10A2C73A">The nomenclature of two hybrid taxa in Carex sect. Vesicariae (Cyperaceae) currently assigned as Carex rostrata var. borealis and Carex stenolepis</mods:title>
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<mods:namePart id="9A8E3E84235FD1D21A435A6AA9B55550">Elven, Reidar</mods:namePart>
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<mods:namePart id="80C811F1BE403D401C8ACD15FD48F53C">Pedersen, A. Tiril M.</mods:namePart>
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<mods:namePart id="DC1FD991702A7CFF662BE100E28F0555">Bjorå, Charlotte S.</mods:namePart>
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<mods:typeOfResource id="940FC8F979892E1DDF43583A9C73CC61">text</mods:typeOfResource>
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<mods:title id="0EBCE025BBE763D7C77FBE8C054EF286">Phytotaxa</mods:title>
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<mods:part id="AED3D84453BA1337F01301D9F5E4B971">
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<mods:date id="196C9AA98E8D92148AAF273EB23BD7A8">2017</mods:date>
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<mods:number id="208F33AF4880DA9EF47CBCCB1A8F9419">2017-10-06</mods:number>
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<mods:number id="1DC8AEC782722889733B6F9C464ED4F3">324</mods:number>
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<mods:url id="A95505E10562D01EEE9330A33C3BDBA3">http://dx.doi.org/10.11646/phytotaxa.324.1.4</mods:url>
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<treatment id="038E0A32FFC3FFE64CECFA31FB69A979" LSID="urn:lsid:plazi:treatment:038E0A32FFC3FFE64CECFA31FB69A979" httpUri="http://treatment.plazi.org/id/038E0A32FFC3FFE64CECFA31FB69A979" lastPageId="7" lastPageNumber="70" pageId="6" pageNumber="69">
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<subSubSection id="C33DE8AFFFC3FFE74CECFA31FD3EAEF5" box="[136,762,1460,1487]" pageId="6" pageNumber="69" type="nomenclature">
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<paragraph id="8B98BB24FFC3FFE74CECFA31FD3EAEF5" blockId="6.[136,762,1460,1487]" box="[136,762,1460,1487]" pageId="6" pageNumber="69">
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<heading id="D0D00C48FFC3FFE74CECFA31FD3EAEF5" box="[136,762,1460,1487]" fontSize="11" level="2" pageId="6" pageNumber="69" reason="3">
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<taxonomicName id="4C27C0A7FFC3FFE74CECFA31FD89AEF5" ID-CoL="8WS7K" authority="T. M. Pedersen & Elven" authorityName="T. M. Pedersen & Elven" authorityYear="2017" box="[136,589,1460,1487]" class="Liliopsida" family="Cyperaceae" genus="Carex" isHybrid="true" kingdom="Plantae" order="Poales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="saamica">
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<emphasis id="B9536736FFC3FFE74CECFA31FE80AEF4" bold="true" box="[136,324,1460,1487]" pageId="6" pageNumber="69">
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<emphasis id="B9536736FFC3FFE74CECFA31FF0AAEF4" bold="true" box="[136,206,1460,1486]" italics="true" pageId="6" pageNumber="69">Carex</emphasis>
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×
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<emphasis id="B9536736FFC3FFE74C81FA31FE80AEF4" bold="true" box="[229,324,1460,1486]" italics="true" pageId="6" pageNumber="69">saamica</emphasis>
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</emphasis>
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T.M.Pedersen & Elven
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</taxonomicName>
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, nomen novum
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33DE8AFFFC3FFE74CECFA5AFC3FAD2F" pageId="6" pageNumber="69" type="reference_group">
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<paragraph id="8B98BB24FFC3FFE74CECFA5AFBD6AECB" blockId="6.[136,1451,1500,1665]" box="[136,1042,1500,1521]" pageId="6" pageNumber="69">
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<emphasis id="B9536736FFC3FFE74CECFA5AFF52AECB" box="[136,150,1503,1521]" italics="true" pageId="6" pageNumber="69">≡</emphasis>
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<taxonomicName id="4C27C0A7FFC3FFE74CF2FA59FD77AECB" ID-CoL="5MW36" authority="Hartman 1820" authorityName="Hartman" authorityYear="1820" box="[150,691,1500,1521]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="variety" species="ampullacea" variety="borealis">
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<emphasis id="B9536736FFC3FFE74CF2FA59FEE6AECB" box="[150,290,1500,1521]" italics="true" pageId="6" pageNumber="69">C. ampullacea</emphasis>
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Goodenough var.
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<emphasis id="B9536736FFC3FFE74DB7FA59FDE5AECB" box="[467,545,1500,1521]" italics="true" pageId="6" pageNumber="69">borealis</emphasis>
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<bibRefCitation id="EFB6C6D5FFC3FFE74E43FA59FD77AECB" author="Hartman, C. J." box="[551,691,1500,1521]" pageId="6" pageNumber="69" refId="ref6128" refString="Hartman, C. J. (1820) Handbok i Skandinaviens flora. Ed. 1. Stockholm, Zacharias Haeggstroms Forlag." type="book" year="1820">Hartman 1820</bibRefCitation>
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</taxonomicName>
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, Handb. Skand. Fl.: 39. (nom. subs.)
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</paragraph>
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<paragraph id="8B98BB24FFC3FFE74CECF986FC3FAD2F" blockId="6.[136,1451,1500,1665]" box="[136,1019,1536,1557]" pageId="6" pageNumber="69">
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<emphasis id="B9536736FFC3FFE74CECF986FF52AD2F" box="[136,150,1539,1557]" italics="true" pageId="6" pageNumber="69">≡</emphasis>
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<taxonomicName id="4C27C0A7FFC3FFE74CF2F985FD35AD2F" ID-CoL="5MV58" authority="(Hartman) Kukenthal 1909" authorityName="Kukenthal" authorityYear="1909" baseAuthorityName="Hartman" baseAuthorityYear="1820" box="[150,753,1536,1557]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="variety" species="rostrata" variety="borealis">
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<emphasis id="B9536736FFC3FFE74CF2F985FEE5AD2F" box="[150,289,1536,1557]" italics="true" pageId="6" pageNumber="69">Carex rostrata</emphasis>
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Stokes var.
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<emphasis id="B9536736FFC3FFE74DF3F985FE21AD2F" box="[407,485,1536,1557]" italics="true" pageId="6" pageNumber="69">borealis</emphasis>
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(Hartman)
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<bibRefCitation id="EFB6C6D5FFC3FFE74E31F985FD35AD2F" author="Kukenthal, G." box="[597,753,1536,1557]" pageId="6" pageNumber="69" refId="ref6370" refString="Kukenthal, G. (1909) Cyperaceae - Caricoideae. In: Engler, A. (Ed.) Das Pflanzenreich. IV. Wilhelm Engelmann, Leipzig, pp. 20 (38)." type="journal volume" year="1909">Kükenthal 1909</bibRefCitation>
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</taxonomicName>
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, Pflanzenr. IV, 20(38): 723.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33DE8AFFFC3FFE64CECF9A1FB69A979" lastPageId="7" lastPageNumber="70" pageId="6" pageNumber="69" type="materials_examined">
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<paragraph id="8B98BB24FFC3FFE74CECF9A1FA4BAD03" blockId="6.[136,1451,1500,1665]" box="[136,1423,1572,1593]" pageId="6" pageNumber="69">
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<emphasis id="B9536736FFC3FFE74CECF9A1FF71AD03" box="[136,181,1572,1593]" italics="true" pageId="6" pageNumber="69">Type</emphasis>
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:—
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<collectingCountry id="F330FBB4FFC3FFE74CB5F9A1FEFEAD03" box="[209,314,1572,1593]" name="Sweden" pageId="6" pageNumber="69">SWEDEN</collectingCountry>
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,
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<collectingRegion id="49E375C6FFC3FFE74D20F9A1FE5FAD03" box="[324,411,1572,1593]" country="Sweden" name="Jamtland" pageId="6" pageNumber="69">Jämtland</collectingRegion>
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“Jaemtland subalpinis”. Herb. Hartman (
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<collectionCode id="ED3623E1FFC3FFE74F46F9A1FC92AD03" box="[802,854,1572,1593]" country="Sweden" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15542" lsid="urn:lsid:biocol.org:col:15542" name="Uppsala University, Museum of Evolution, Botany Section (Fytoteket)" pageId="6" pageNumber="69" type="Herbarium">UPS</collectionCode>
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!),
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<typeStatus id="549C0586FFC3FFE74F0FF9A1FC07AD03" box="[875,963,1572,1593]" pageId="6" pageNumber="69" type="lectotype">lectotype</typeStatus>
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(
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<figureCitation id="131CA7A1FFC3FFE74FB4F9A1FBCCAD03" box="[976,1032,1572,1593]" captionStart="FIGURE 1" captionStartId="3.[136,229,1984,2005]" captionTargetBox="[254,1334,187,1925]" captionTargetId="figure-14@3.[254,1334,187,1925]" captionTargetPageId="3" captionText="FIGURE 1. Lectotype of Carex rostrata var. borealis (Hartman) Kükenthal, from Sweden: Jämtland. From Uppsala Herbarium (UPS). Here referred to C. ×saamica T.M.Pedersen & Elven nom. nov. (= C. rostrata Stokes × C. rotundata Wahlenb.).A, habit; B, pistillate spike. © Museum of Evolution, Uppsala University (UPS)." figureDoi="http://doi.org/10.5281/zenodo.13697400" httpUri="https://zenodo.org/record/13697400/files/figure.png" pageId="6" pageNumber="69">Fig. 1</figureCitation>
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), designated here (or perhaps
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<typeStatus id="549C0586FFC3FFE7494FF9A1FA47AD03" box="[1323,1411,1572,1593]" pageId="6" pageNumber="69" type="holotype">holotype</typeStatus>
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).
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</paragraph>
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<paragraph id="8B98BB24FFC3FFE74CECF9CDFE8DADBB" blockId="6.[136,1451,1500,1665]" pageId="6" pageNumber="69">
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Etymology—The name
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<taxonomicName id="4C27C0A7FFC3FFE74D14F9CDFE0DAD67" authorityName="T. M. Pedersen & Elven" authorityYear="2017" box="[368,457,1608,1629]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="6" pageNumber="69" phylum="Tracheophyta" rank="species" species="saamica">“saamica</taxonomicName>
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” is chosen because the main range of this taxon seems to be within the Saami (Sápmi/Lapland) areas of Fennoscandia.
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</paragraph>
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<paragraph id="8B98BB24FFC3FFE64CECF937FD50AA8F" blockId="6.[136,1452,1714,2063]" lastBlockId="7.[136,1452,160,437]" lastPageId="7" lastPageNumber="70" pageId="6" pageNumber="69">
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Plants develop long-creeping rhizomes and forms open, extensive swards or clones, often of
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<quantity id="4CDF16C1FFC3FFE748F3F937FB28ADF1" box="[1175,1260,1714,1739]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="69" unit="m" value="1000.0">1000 m</quantity>
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² or more (Elven, Pedersen & Bjorå, pers. obs.). Withered leaf bases
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<taxonomicName id="4C27C0A7FFC3FFE74EB3F953FC9FADD5" box="[727,859,1750,1775]" form="dense" pageId="6" pageNumber="69" rank="form">form dense</taxonomicName>
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, dark brownish grey sheaths around base of culms. Leaves mainly basal, 1–2 blades arising from sheaths on the culms, erect, as long as or longer than culms, involute, smooth except for being scabrous on margins; abaxial surface green, adaxial surface pale green (but not visible unless the leaf is rolled out), both surfaces without pale spots or papillae. Culms 20–40(50) cm, erect, obtusely trigonous, smooth. Inflorescence (1)2–3 lateral pistillate spikes, the proximal one usually the largest, and 1–2(3) terminal staminate spikes, the distal one the largest. Lowermost bract
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<quantity id="4CDF16C1FFC3FFE74F4AF80FFC64AC99" box="[814,928,1930,1955]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" metricValueMax="1.8" metricValueMin="1.0" pageId="6" pageNumber="69" unit="cm" value="14.0" valueMax="18.0" valueMin="10.0">10–18 cm</quantity>
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, usually overtopping the inflorescence, narrow (
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<quantity id="4CDF16C1FFC3FFE74CF5F82BFF37ACFD" box="[145,243,1966,1991]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="6" pageNumber="69" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
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), moderately canaliculate; adaxial surface green, abaxial surface pale green, without pale spots or papillae. Distal bracts much shorter and narrower than the lowermost one. Pistillate spikes 20–35 ×
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<quantity id="4CDF16C1FFC3FFE748C7F857FAC1ACD1" box="[1187,1285,2002,2027]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="6" pageNumber="69" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">7–9 mm</quantity>
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, cylindrical or oblong, erect (even when long-pedunculate); proximal spike on peduncle
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<quantity id="4CDF16C1FFC3FFE74FBBF873FB99A335" box="[991,1117,2038,2063]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="6" pageNumber="69" unit="mm" value="20.0" valueMax="30.0" valueMin="10.0">10–30 mm</quantity>
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, more distal spikes on much shorter peduncles or subsessile. Pistillate scales lanceolate or ovate, 3.5–5.0 ×
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<quantity id="4CDF16C1FFC2FFE6486DFF25FB50AB83" box="[1033,1172,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="7" pageNumber="70" unit="mm" value="1.5" valueMax="1.8" valueMin="1.2">1.2–1.8 mm</quantity>
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, acute or obtuse, shorter and narrower than utricles, reddish brown with a pale brown or green midvein, darker brown towards the margins but with a narrow hyaline border and tip. Staminate spikes very narrow, the distal one 25–35 ×
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<quantity id="4CDF16C1FFC2FFE648C7FF6DFAEBAA3B" box="[1187,1327,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.3" metricValueMin="1.8" pageId="7" pageNumber="70" unit="mm" value="2.05" valueMax="2.3" valueMin="1.8">1.8–2.3 mm</quantity>
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. Staminate scales ovate, 3.0–4.5 ×
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||
<quantity id="4CDF16C1FFC2FFE64DF4FE89FDDEAA1F" box="[400,538,268,293]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="7" pageNumber="70" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8 mm</quantity>
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, obtuse, brown with a pale brown or green midvein, with a broad hyaline margin and tip. Stigmas 3. Utricles 4.0–5.5 × 2.0–3.0 mm, smooth, shiny, yellowish green during anthesis and turning pale brown during maturation, with indistinct veins, narrowing gradually into a short beak
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||
<quantity id="4CDF16C1FFC2FFE6485AFED1FB01AA57" box="[1086,1221,340,365]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="7" pageNumber="70" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0.3–0.5 mm</quantity>
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, distinctly bidentate. Utricles well-formed but usually (but not always) with undeveloped achenes. Stamens exerted and well developed; pollen variable, but usually looks well formed.
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</paragraph>
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<paragraph id="8B98BB24FFC2FFE64CECFE67FB69A979" blockId="7.[136,1452,482,579]" pageId="7" pageNumber="70">
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<taxonomicName id="4C27C0A7FFC2FFE64CECFE67FE35AAC7" authority="Lessing, Reise" authorityName="Lessing, Reise" authorityYear="1831" box="[136,497,482,509]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="stenolepis">
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||
<emphasis id="B9536736FFC2FFE64CECFE67FE83AAC7" bold="true" box="[136,327,482,509]" italics="true" pageId="7" pageNumber="70">Carex stenolepis</emphasis>
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||
<materialsCitation id="3B4FB179FFC2FFE64D2AFE61FE35AAC7" box="[334,497,484,509]" collectionCode="M, N, O" collectorName="den Loffoden" country="Norway" location="Nord-Trondelag" pageId="7" pageNumber="70" specimenCount="1" stateProvince="Nord-Trondelag" typeStatus="lectotype">Lessing, Reise</materialsCitation>
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||
</taxonomicName>
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durch Norwegen nach
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<collectorName id="26D2DEF2FFC2FFE64E92FE61FC4AAAC7" box="[758,910,484,509]" pageId="7" pageNumber="70">den Loffoden</collectorName>
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durch
|
||
<bibCitation id="5E0DB348FFC2FFE64FBFFE61FAB6AAC7" box="[987,1394,484,509]" page="301" pageId="7" pageNumber="70" score="0.0031996205649746983" volumeTitle="Lappland und Schweden" year="1831">Lappland und Schweden: 301. 1831</bibCitation>
|
||
.
|
||
<emphasis id="B9536736FFC2FFE64CECFD8FFF71A925" box="[136,181,522,543]" italics="true" pageId="7" pageNumber="70">Type</emphasis>
|
||
:—
|
||
<collectingCountry id="F330FBB4FFC2FFE64CABFD8FFEFDA925" box="[207,313,522,543]" name="Norway" pageId="7" pageNumber="70">NORWAY</collectingCountry>
|
||
,
|
||
<collectingRegion id="49E375C6FFC2FFE64D27FD8FFE25A925" box="[323,481,522,543]" country="Norway" name="Nord-Trondelag" pageId="7" pageNumber="70">Nord-Trøndelag</collectingRegion>
|
||
, Meråker: “Størdalen: Skurdalsporten,
|
||
<emphasis id="B9536736FFC2FFE64F3FFD8FFB90A925" box="[859,1108,522,543]" italics="true" pageId="7" pageNumber="70">
|
||
<taxonomicName id="4C27C0A7FFC2FFE64F3FFD8FFC37A925" authorityName="Linnaeus" authorityYear="1753" box="[859,1011,522,543]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="vesicaria">Carex vesicaria</taxonomicName>
|
||
–alpigena
|
||
</emphasis>
|
||
.” [1824],
|
||
<collectionCode id="ED3623E1FFC2FFE648DCFD8FFB14A925" box="[1208,1232,522,543]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="7" pageNumber="70" type="Herbarium">M</collectionCode>
|
||
.
|
||
<collectionCode id="ED3623E1FFC2FFE648B7FD8FFB23A925" box="[1235,1255,522,543]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="7" pageNumber="70" type="Herbarium">N</collectionCode>
|
||
. Blytt (
|
||
<collectionCode id="ED3623E1FFC2FFE64955FD8FFA83A925" box="[1329,1351,522,543]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="7" pageNumber="70" type="Herbarium">O</collectionCode>
|
||
!), left side plant,
|
||
<typeStatus id="549C0586FFC2FFE64C9DFDABFE9DA979" box="[249,345,558,579]" pageId="7" pageNumber="70" type="lectotype">Lectotype</typeStatus>
|
||
(designated here;
|
||
<figureCitation id="131CA7A1FFC2FFE64E6EFDABFD80A979" box="[522,580,558,579]" captionStart="FIGURE 2" captionStartId="4.[136,229,2029,2050]" captionTargetBox="[224,1364,190,2004]" captionTargetId="figure-15@4.[224,1364,190,2004]" captionTargetPageId="4" captionText="FIGURE 2. Lectoype of Carex stenolepis Lessing, designated here, from Norway: Nord-Trøndelag, Meråker. From Oslo Herbarium (O). The plants belong to a short grown form of C. rostrata. A, habit; B, pistillate spike; C, utricle." figureDoi="http://doi.org/10.5281/zenodo.13697402" httpUri="https://zenodo.org/record/13697402/files/figure.png" pageId="7" pageNumber="70">Fig. 2</figureCitation>
|
||
). The name is concluded to be a synonym of
|
||
<taxonomicName id="4C27C0A7FFC2FFE64F93FDABFB69A979" authority="Stokes." authorityName="Stokes." box="[1015,1197,558,579]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="7" pageNumber="70" phylum="Tracheophyta" rank="species" species="rostrata">
|
||
<emphasis id="B9536736FFC2FFE64F93FDABFB9BA979" box="[1015,1119,558,579]" italics="true" pageId="7" pageNumber="70">C. rostrata</emphasis>
|
||
Stokes.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |