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<document id="119E79B249E83CBD4D9CFFAF6C298A37" ID-DOI="10.5281/zenodo.5311784" ID-ISSN="0374-1036" ID-Zenodo-Dep="5311784" ID-ZooBank="15CF0A9E-44F3-492B-88F7-A7922EF9F73A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630118666848" checkinUser="marcus" docAuthor="Angus, Robert B., Jia, Fenglong, Chen, Zhen-ning, Zhang, Ying, Vondráček, Dominik, Fikáček, Martin &amp; Sw, London" docDate="2016" docId="377A879F8D702844FE8058F17181FCB5" docLanguage="en" docName="ActaEntMusNatPra.2016.56.1.109-148.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 56 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Helophorus (Lihelophorus) ser Zaitzev 1908" docType="treatment" docVersion="3" lastPageNumber="127" masterDocId="CB43FFE78D792856FFE05B4D7368FFAC" masterDocTitle="Taxonomy, larval morphology and cytogenetics of Lihelophorus, the Tibetan endemic subgenus of Helophorus (Coleoptera: Hydrophiloidea)" masterLastPageNumber="148" masterPageNumber="109" pageNumber="118" updateTime="1699158837459" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E05D05938DD873E27B64FD774A04F4D9">Taxonomy, larval morphology and cytogenetics of Lihelophorus, the Tibetan endemic subgenus of Helophorus (Coleoptera: Hydrophiloidea)</mods:title>
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<mods:namePart id="6B0981C691E771B072C125FA737C0EBB">Jia, Fenglong</mods:namePart>
<mods:affiliation id="3E8B9E67750F757C5BAB045A73D01A50">Museum of Biology, Sun Yat-sen University, Guangzhou, China; e-mail: lssjϐl @ mail. sysu. edu. cn</mods:affiliation>
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<taxonomicName id="78D34D0A8D70285FFE8058F1703AFC7F" ID-CoL="3KHKG" authority="Zaitzev, 1908" authorityName="Zaitzev" authorityYear="1908" box="[352,850,956,979]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="ser" subGenus="Lihelophorus">
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<emphasis id="8DA7EA9B8D70285FFE8058F1728AFC7F" bold="true" box="[352,482,956,979]" italics="true" pageId="9" pageNumber="118">Helophorus</emphasis>
(
<emphasis id="8DA7EA9B8D70285FFE1258F171EEFC7F" bold="true" box="[498,646,956,979]" italics="true" pageId="9" pageNumber="118">Lihelophorus</emphasis>
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<bibRefCitation id="DB424B788D70285FFD5D58F1703AFC7F" author="ZAITZEV F. A." box="[701,850,956,979]" pageId="9" pageNumber="118" pagination="417 - 426" refId="ref20914" refString="ZAITZEV F. A. 1908: Beitrag zur Kenntniss der Waserkafer von Chinesisch-Centralasien. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St. Petersbourg 13: 417 - 426." type="journal article" year="1908">Zaitzev, 1908</bibRefCitation>
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(
<figureCitation id="27E82A0C8D70285FFE7758977149FC41" box="[407,545,986,1005]" captionStart="Fig" captionStartId="6.[151,182,1490,1509]" captionTargetBox="[152,1126,220,1478]" captionTargetId="figure-10@6.[151,1127,217,1486]" captionTargetPageId="6" captionText="Fig. 1. Adult morphology of Helophorus (Lihelophorus). ab H. yangae sp. nov. (a ventral view of the head; b detail of mentum and labial palps); cd maxillary galea and lacinia (c H. yangae sp. nov.; d H. ser Zaitzev, 1908); ef mandible (e H. yangae sp. nov.; f H. ser); g labial palpus of H. ser; hl hind legs in dorsal view (h H. yangae sp. nov.; i H. ser; j H. lamicola Zaitzev, 1908; k detail of apex of tibia and base of tarsus of H. lamicola; l detail of tarsomere 5 and claw of H. ser). Abbreviations: sh swimming hairs. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311786" httpUri="https://zenodo.org/record/5311786/files/figure.png" pageId="9" pageNumber="118">Figs 1d, fg, i, l</figureCitation>
;
<figureCitation id="27E82A0C8D70285FFDCB58977115FC41" box="[555,637,986,1005]" captionStart="Fig" captionStartId="7.[113,144,1463,1482]" captionTargetBox="[133,1067,220,1453]" captionTargetId="figure-13@7.[133,1068,219,1454]" captionTargetPageId="7" captionText="Fig. 2. Adult morphology of Helophorus (Lihelophorus). af details of left half of pronotum (a,c,e) and base of left elytron (b, d, f): ab H. yangae sp. nov.; cd H. ser Zaitzev, 1908; ef H. lamicola Zaitzev, 1908; arrow in (a) shows remnants of pronotal granulations, arrows in (b, d, f) show scutellary stria. gh prothoracic hypomeron and suprapleural area (g H. yangae sp. nov.; h H. ser). i detail of posterior margin of abdominal ventrite 5 of H. ser, showing the irregular denticulation. j ventral view of the thorax of H. ser. k dorsolateral portion of left elytron of H. lamicola. Abbreviations: ep epipleuron; li last elytral interval. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311790" httpUri="https://zenodo.org/record/5311790/files/figure.png" pageId="9" pageNumber="118">2cd, hj</figureCitation>
;
<figureCitation id="27E82A0C8D70285FFD66589771DBFC41" box="[646,691,986,1005]" captionStart="Fig" captionStartId="10.[151,182,1482,1501]" captionTargetBox="[159,1124,234,1349]" captionTargetId="figure-10@10.[151,1127,219,1365]" captionTargetPageId="10" captionText="Fig. 3. Habitus of the species of Helophorus (Lihelophorus). ac H. lamicola Zaitzev, 1908 (a Qinghai: Zhaling Hu (lectotype); bc Quinghai: Golo).dg H.ser Zaitzev, 1908 (de Qinghai:Golo; f Xizang, Nyainqentanglha Feng; g Gansu: Xiahe env.). hk H. yangae sp. nov. (h Xizang: 90 km W of Amdo; i Xizang: 25 km W of Amdo; j Qinghai: Golo; k India: Kar Tso (Orchymont coll.)). All specimens to scale." figureDoi="http://doi.org/10.5281/zenodo.5311792" httpUri="https://zenodo.org/record/5311792/files/figure.png" pageId="9" pageNumber="118">3dg</figureCitation>
;
<figureCitation id="27E82A0C8D70285FFD5C58977181FC41" box="[700,745,986,1005]" captionStart="Fig" captionStartId="11.[113,144,1530,1549]" captionTargetBox="[131,1080,229,1416]" captionTargetId="figure-13@11.[113,1091,219,1456]" captionTargetPageId="11" captionText="Fig.4. Details of the head and pronotum of the Helophorus (Lihelophorus) specimens.ac H. lamicola Zaitzev,1908 (a Qinghai: Zhaling Hu (lectotype); bc Qinghai: Golo). dh H. ser Zaitzev, 1908 (d Qinghai: valley of Alag Hu lake (lectotype); ef Qinghai: Golo; g Xizang, Nyainqentanglha Feng; h Gansu: Xiahe env.). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311794" httpUri="https://zenodo.org/record/5311794/files/figure.png" pageId="9" pageNumber="118">4dh</figureCitation>
;
<figureCitation id="27E82A0C8D70285FFD1258977072FC41" box="[754,794,986,1005]" captionStart="Fig" captionStartId="15.[113,144,1496,1515]" captionTargetBox="[149,1030,229,1439]" captionTargetId="figure-13@15.[139,1063,219,1474]" captionTargetPageId="15" captionText="Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. ae H. (L.) ser Zaitzev, 1908 (a lectotype, Qinghai: valley of Alag Hu lake; bc Xizang: valley SE of Dong La pass; d Qinghai: Golo, 20 km E of Maduo; e Qinghai: Zuimatan). fj H. (L.) lamicola Zaitzev, 1908 (f lectotype, Qinghai: Djarin Nor; gi Qinghai: Zuimatan; j paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). ko H. (L.) yangae sp. nov. (k holotype, Qinghai: Zuimatan; lm paratypes, same locality; n Qinghai: Golo, Yematan; o Xizang: Gongbujiangda)." figureDoi="http://doi.org/10.5281/zenodo.5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="9" pageNumber="118">6ae</figureCitation>
)
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<taxonomicName id="78D34D0A8D70285FFF9158B0714AFBBC" ID-CoL="3KHKG" authority="Zaitzev, 1908: 422" authorityName="Zaitzev" authorityPageNumber="422" authorityYear="1908" box="[113,546,1021,1040]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="ser" subGenus="Lihelophorus">
<emphasis id="8DA7EA9B8D70285FFF9158B073BEFBBC" box="[113,214,1021,1040]" italics="true" pageId="9" pageNumber="118">Helophorus</emphasis>
(
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<emphasis id="8DA7EA9B8D70285FFE8158B07213FBBC" box="[353,379,1021,1040]" italics="true" pageId="9" pageNumber="118">ser</emphasis>
<bibRefCitation id="DB424B788D70285FFE6058B0714AFBBC" author="ZAITZEV F. A." box="[384,546,1021,1040]" pageId="9" pageNumber="118" pagination="417 - 426" refId="ref20914" refString="ZAITZEV F. A. 1908: Beitrag zur Kenntniss der Waserkafer von Chinesisch-Centralasien. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St. Petersbourg 13: 417 - 426." type="journal article" year="1908">Zaitzev, 1908: 422</bibRefCitation>
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<emphasis id="8DA7EA9B8D70285FFF915F5A73BEFB86" box="[113,214,1047,1066]" italics="true" pageId="9" pageNumber="118">Helophorus</emphasis>
(
<emphasis id="8DA7EA9B8D70285FFF035F5A739EFB86" box="[227,246,1047,1066]" italics="true" pageId="9" pageNumber="118">L.</emphasis>
)
<emphasis id="8DA7EA9B8D70285FFEE25F5A7274FB86" box="[258,284,1047,1066]" italics="true" pageId="9" pageNumber="118">ser</emphasis>
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:
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<bibRefCitation id="DB424B788D70285FFEC75F5A72C9FB86" author="ANGUS R. B." box="[295,417,1047,1066]" pageId="9" pageNumber="118" pagination="185 - 206" refId="ref19760" refString="ANGUS R. B. 1995: Helophoridae: The Helophorus species of China, with notes on the species from neighbouring areas (Coleoptera). Pp 185 - 206. In: JACH M. A. &amp; JI L. (eds): Water Beetles of China. Volume 1. Zoologisch- - Botanische Gesellschaft &amp; Wiener Coleopterologenverein, Wien, 410 pp." type="book chapter" year="1995">
<collectingRegion id="7D17F86B8D70285FFEC75F5A7237FB84" box="[295,351,1047,1066]" country="United Kingdom" name="Angus" pageId="9" pageNumber="118">ANGUS</collectingRegion>
(1995)
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: 190
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(redescription,
<typeStatus id="6068882B8D70285FFDB25F5A71C8FB86" box="[594,672,1047,1066]" pageId="9" pageNumber="118" type="lectotype">lectotype</typeStatus>
designation);
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<bibRefCitation id="DB424B788D70285FFCFC5F5A70BAFB86" author="FIKACEK M. &amp; PROKIN A. &amp; ANGUS R. B. &amp; PONOMARENKO A. &amp; YUE Y. &amp; REN D. &amp; PROKOP J." box="[796,978,1047,1066]" pageId="9" pageNumber="118" pagination="420 - 447" refId="ref20446" refString="FIKACEK M., PROKIN A., ANGUS R. B., PONOMARENKO A., YUE Y., REN D. &amp; PROKOP J. 2012: Phylogeny and the fossil record of the Helophoridae reveal Jurassic origin of extant hydrophiloid lineages (Coleoptera: Polyphaga). Systematic Entomology 37: 420 - 447." type="journal article" year="2012">FIKÁČEK et al. (2012)</bibRefCitation>
: 437
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(phylogenetic analysis).
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</paragraph>
</subSubSection>
<subSubSection id="F7C965028D70285AFF915F117205FAFE" lastPageId="12" lastPageNumber="121" pageId="9" pageNumber="118" type="materials_examined">
<paragraph id="BF6C36898D70285FFF915F117027FBC3" blockId="9.[113,1090,1116,1589]" box="[113,847,1116,1135]" pageId="9" pageNumber="118">
<emphasis id="8DA7EA9B8D70285FFF915F117382FBC3" bold="true" box="[113,234,1116,1135]" pageId="9" pageNumber="118">
<typeStatus id="6068882B8D70285FFF915F1173F6FBC3" box="[113,158,1116,1135]" pageId="9" pageNumber="118">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="C7C476198D70285FFF0F5F11724CFBC3" box="[239,292,1116,1135]" name="China" pageId="9" pageNumber="118">China</collectingCountry>
,
<collectingRegion id="7D17F86B8D70285FFECD5F1172ABFBC3" box="[301,451,1116,1135]" country="China" name="Qinghai" pageId="9" pageNumber="118">Qinghai Province</collectingRegion>
, valley of Alag Hu lake,
<geoCoordinate id="DAE7504E8D70285FFD7A5F117187FBC3" box="[666,751,1116,1135]" degrees="35.573" direction="north" orientation="latitude" pageId="9" pageNumber="118" precision="55" value="35.573">35.573°N</geoCoordinate>
,
<geoCoordinate id="DAE7504E8D70285FFD185F117023FBC3" box="[760,843,1116,1135]" degrees="97.120" direction="east" orientation="longitude" pageId="9" pageNumber="118" precision="55" value="97.12">97.120°E</geoCoordinate>
.
</paragraph>
<paragraph id="BF6C36898D70285AFF915F3B7205FAFE" blockId="9.[113,1090,1116,1589]" lastBlockId="12.[151,1128,1156,1362]" lastPageId="12" lastPageNumber="121" pageId="9" pageNumber="118">
<emphasis id="8DA7EA9B8D70285FFF915F3B7224FB25" bold="true" box="[113,332,1142,1161]" pageId="9" pageNumber="118">Type material examined</emphasis>
(
<specimenCount id="A9D5FD008D70285FFEB95F3B72FEFB25" box="[345,406,1142,1161]" pageId="9" pageNumber="118" type="generic">1 spec.</specimenCount>
).
<materialsCitation id="0FBB3CD48D70285FFE435F3B7729FB13" collectingDate="1900-05-30" collectionCode="ZIN" collectorName="Ekcp. &amp; Eksp. Kozlova. &amp; Hel." determinerName="Zaitzev" location="Bydda" pageId="9" pageNumber="118" specimenCode="ANGUS 1995" specimenCount="1" typeStatus="lectotype">
<typeStatus id="6068882B8D70285FFE435F3B7294FB24" box="[419,508,1142,1161]" pageId="9" pageNumber="118" type="lectotype">LECTOTYPE</typeStatus>
(designated by
<bibRefCitation id="DB424B788D70285FFD605F3B7183FB25" author="ANGUS R. B." box="[640,747,1142,1161]" pageId="9" pageNumber="118" pagination="185 - 206" refId="ref19760" refString="ANGUS R. B. 1995: Helophoridae: The Helophorus species of China, with notes on the species from neighbouring areas (Coleoptera). Pp 185 - 206. In: JACH M. A. &amp; JI L. (eds): Water Beetles of China. Volume 1. Zoologisch- - Botanische Gesellschaft &amp; Wiener Coleopterologenverein, Wien, 410 pp." type="book chapter" year="1995">
<specimenCode id="EF759EF28D70285FFD605F3B7183FB25" box="[640,747,1142,1161]" collectionCode="ANGUS" pageId="9" pageNumber="118">ANGUS 1995</specimenCode>
</bibRefCitation>
): 1 J (
<collectionCode id="D9C2AE4C8D70285FFCC35F3B7023FB25" box="[803,843,1142,1161]" country="Russia" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="9" pageNumber="118" type="Museum">ZIN</collectionCode>
): LECTO- / TYPE [rounded label with violet margin] // Ю. cкл. xp. Буpxaн-
<location id="BA0C60528D70285FFDE65FDC7156FB08" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D70285FFDE65FDC7156FB08" box="[518,574,1169,1188]" name="Bydda" pageId="9" pageNumber="118">Бyддa</location>
: [= Yu. skl. khr. Burkhan-Budda:] / дoл. oЗ. Aлык- Нop. [dol. oz.Alyk-Nor]
<date id="CB6D10498D70285FFF0B5FE1722DFB13" box="[235,325,1196,1215]" pageId="9" pageNumber="118" value="1900-05-30">
<collectingDate id="DB29E9A18D70285FFF0B5FE1722DFB13" box="[235,325,1196,1215]" pageId="9" pageNumber="118" value="1900-05-30">30.V.1900</collectingDate>
</date>
. /
<collectorName id="1226535F8D70285FFEB65FE172E0FB13" box="[342,392,1196,1215]" pageId="9" pageNumber="118">Эkcп.</collectorName>
КoЗлoвa. [=
<collectorName id="1226535F8D70285FFE175FE17114FB13" box="[503,636,1196,1215]" pageId="9" pageNumber="118">Eksp. Kozlova.</collectorName>
] //
<collectorName id="1226535F8D70285FFD735FE171D1FB13" box="[659,697,1196,1215]" pageId="9" pageNumber="118">Hel.</collectorName>
(
<taxonomicName id="78D34D0A8D70285FFD215FE1705DFB13" authorityName="Zaitzev" authorityYear="1908" box="[705,821,1196,1215]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="147" phylum="Arthropoda" rank="subGenus" subGenus="Lihelophorus">Lihelophorus</taxonomicName>
) / ser
<taxonomicNameLabel id="969457E08D70285FFC865FE170FBFB13" box="[870,915,1196,1215]" pageId="9" pageNumber="118" rank="species">sp. n.</taxonomicNameLabel>
/ iv.08
<determinerName id="4AB0DA4D8D70285FFC2E5FE17766FB13" box="[974,1038,1196,1215]" pageId="9" pageNumber="118">Zaitzev</determinerName>
det. //
</materialsCitation>
<typeStatus id="6068882B8D70285FFF915F8B7381FB75" box="[113,233,1222,1241]" pageId="9" pageNumber="118" type="lectotype">LECTOTYPE</typeStatus>
<date id="CB6D10498D70285FFF0D5F8B7239FB75" box="[237,337,1222,1241]" pageId="9" pageNumber="118" value="1970-03">March 1970</date>
/
<taxonomicName id="78D34D0A8D70285FFEBE5F8B7178FB75" authority="Zaitz." authorityName="Zaitzev" authorityYear="1908" box="[350,528,1222,1241]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="ser">Helophorus ser Zaitz.</taxonomicName>
/ R. B.Angus det.// кoлл. Семенова- T.Ш. [koll. Semenova-T.Sh.]
<materialsCitation id="0FBB3CD48D70285FFF915FAC7157FAA3" collectingDate="1906-08" collectionCode="IRSNB" collectorName="Zugmayer &amp; Orchymont &amp; Knisch" country="China" location="Westtibet" municipality="Apo Zo" pageId="9" pageNumber="118" specimenCount="95" stateProvince="Xizang">
<emphasis id="8DA7EA9B8D70285FFF915FAC7212FB58" bold="true" box="[113,378,1249,1268]" pageId="9" pageNumber="118">Additional material examined</emphasis>
(
<specimenCount id="A9D5FD008D70285FFE655FAC72A5FB58" box="[389,461,1249,1268]" pageId="9" pageNumber="118" type="generic">95 spec.</specimenCount>
).
<emphasis id="8DA7EA9B8D70285FFE395FAC7118FB58" bold="true" box="[473,624,1249,1268]" pageId="9" pageNumber="118">
<collectingCountry id="C7C476198D70285FFE395FAC714AFB58" box="[473,546,1249,1268]" name="China" pageId="9" pageNumber="118">CHINA</collectingCountry>
:
<collectingRegion id="7D17F86B8D70285FFDCA5FAC7101FB5E" box="[554,617,1249,1268]" country="China" name="Xizang" pageId="9" pageNumber="118">XIZANG</collectingRegion>
:
</emphasis>
1 J (
<collectionCode id="D9C2AE4C8D70285FFD7D5FAC7188FB58" box="[669,736,1249,1268]" country="Belgium" name="Institut Royal des Sciences Naturelles de Belgique" pageId="9" pageNumber="118">IRSNB</collectionCode>
):
<collectingMunicipality id="5F08ACF38D70285FFD0B5FAC7044FB58" box="[747,812,1249,1268]" pageId="9" pageNumber="118">Apo Zo</collectingMunicipality>
,
<location id="BA0C60528D70285FFCD35FAC70ECFB58" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D70285FFCD35FAC70ECFB58" box="[819,900,1249,1268]" country="China" municipality="Apo Zo" name="Westtibet" pageId="9" pageNumber="118" stateProvince="Xizang">Westtibet</location>
,
<date id="CB6D10498D70285FFC6A5FAC708BFB58" box="[906,995,1249,1268]" pageId="9" pageNumber="118" value="1906-08">
<collectingDate id="DB29E9A18D70285FFC6A5FAC708BFB58" box="[906,995,1249,1268]" pageId="9" pageNumber="118" value="1906-08">Aug.1906</collectingDate>
</date>
,
<collectorName id="1226535F8D70285FFC0A5FAC7728FB58" box="[1002,1088,1249,1268]" pageId="9" pageNumber="118">Zugmayer</collectorName>
(identified as
<taxonomicName id="78D34D0A8D70285FFF065FB17223FAA3" authorityName="Zaitzev" authorityYear="1908" box="[230,331,1276,1295]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="118" phylum="Arthropoda" rank="species" species="lamicola">
<emphasis id="8DA7EA9B8D70285FFF065FB17223FAA3" box="[230,331,1276,1295]" italics="true" pageId="9" pageNumber="118">H. lamicola</emphasis>
</taxonomicName>
by
<collectorName id="1226535F8D70285FFE8E5FB172A5FAA3" box="[366,461,1276,1295]" pageId="9" pageNumber="118">Orchymont</collectorName>
and
<collectorName id="1226535F8D70285FFE1A5FB17151FAA3" box="[506,569,1276,1295]" pageId="9" pageNumber="118">Knisch</collectorName>
)
</materialsCitation>
;
<materialsCitation id="0FBB3CD48D70285FFDA95FB170DAFA85" collectingDate="2011-06-18" collectionCode="SE" collectorName="S. Tibet &amp; J. Schmidt" country="China" latitude="28.766666" location="Dongo La Pass" longLatPrecision="1230" longitude="87.95" pageId="9" pageNumber="118" specimenCount="1" stateProvince="Xizang">
1 unsexed spec. (BMNH):
<collectorName id="1226535F8D70285FFCD65FB17015FAA3" box="[822,893,1276,1295]" pageId="9" pageNumber="118">S. Tibet</collectorName>
, vall.
<collectionCode id="D9C2AE4C8D70285FFC525FB170A3FAA3" box="[946,971,1276,1295]" pageId="9" pageNumber="118">SE</collectionCode>
of
<location id="BA0C60528D70285FFC0A5FB173FEFA85" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D70285FFC0A5FB173FEFA85" country="China" latitude="28.766666" longLatPrecision="1230" longitude="87.95" name="Dongo La Pass" pageId="9" pageNumber="118" stateProvince="Xizang">Dongo La Pass</location>
[= valley
<locationDeviation id="AEDE2FAB8D70285FFF0F5E5B7219FA85" box="[239,369,1302,1321]" location="Dong La" pageId="9" pageNumber="118">
<collectionCode id="D9C2AE4C8D70285FFF0F5E5B7260FA85" box="[239,264,1302,1321]" pageId="9" pageNumber="118">SE</collectionCode>
of Dong La
</locationDeviation>
pass],
<geoCoordinate id="DAE7504E8D70285FFE4D5E5B729DFA85" box="[429,501,1302,1321]" degrees="28" direction="north" minutes="46" orientation="latitude" pageId="9" pageNumber="118" precision="925" value="28.766666">28°46ʹN</geoCoordinate>
,
<geoCoordinate id="DAE7504E8D70285FFE1F5E5B712CFA85" box="[511,580,1302,1321]" degrees="87" direction="east" minutes="57" orientation="longitude" pageId="9" pageNumber="118" precision="925" value="87.95">87°57ʹE</geoCoordinate>
, 49005300 m,
<date id="CB6D10498D70285FFD355E5B7047FA85" box="[725,815,1302,1321]" pageId="9" pageNumber="118" value="2011-06-18">
<collectingDate id="DB29E9A18D70285FFD355E5B7047FA85" box="[725,815,1302,1321]" pageId="9" pageNumber="118" value="2011-06-18">18.vi.2011</collectingDate>
</date>
,
<collectorName id="1226535F8D70285FFCD95E5B70FAFA85" box="[825,914,1302,1321]" pageId="9" pageNumber="118">J. Schmidt</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D70285FFC5F5E5B7198FAF3" collectingDate="2010-07-20" collectionCode="NMPC" collectorName="Schmidt" country="China" county="Tibet" elevation="5175" latitude="30.379723" location="Lha Tsu Vall" longLatPrecision="20" longitude="90.72278" municipality="Nyainqentanglha Feng" pageId="9" pageNumber="118" specimenCount="16" specimenCount-female="16" stateProvince="Xizang">
16 JJ
<specimenCount id="A9D5FD008D70285FFBE65E5B772AFA85" box="[1030,1090,1296,1321]" pageId="9" pageNumber="118" type="female">16 ♀♀</specimenCount>
(BMNH), 1 J 1 unsexed spec. (
<collectionCode id="D9C2AE4C8D70285FFE665E7C72AFFAE8" box="[390,455,1329,1348]" country="Czech Republic" name="National Museum Prague" pageId="9" pageNumber="118">NMPC</collectionCode>
): S.
<collectingCounty id="560D4E058D70285FFE155E7C7148FAE8" box="[501,544,1329,1348]" pageId="9" pageNumber="118">Tibet</collectingCounty>
E
<collectingMunicipality id="5F08ACF38D70285FFDDB5E7C7193FAE8" box="[571,763,1329,1348]" pageId="9" pageNumber="118">Nyainqentanglha Feng</collectingMunicipality>
[Mt.], E slope
<location id="BA0C60528D70285FFC605E7C7086FAE8" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D70285FFC605E7C7086FAE8" box="[896,1006,1329,1348]" country="China" county="Tibet" latitude="30.379723" longLatPrecision="20" longitude="90.72278" municipality="Nyainqentanglha Feng" name="Lha Tsu Vall" pageId="9" pageNumber="118" stateProvince="Xizang">Lha Tsu Vall</location>
[Lha Tsu valley],
<quantity id="782B9B6C8D70285FFF565E017243FAF3" box="[182,299,1356,1375]" metricMagnitude="3" metricUnit="m" metricValue="5.175" metricValueMax="5.35" metricValueMin="5.0" pageId="9" pageNumber="118" unit="m" value="5175.0" valueMax="5350.0" valueMin="5000.0">
<elevation id="34FED1BA8D70285FFF565E017243FAF3" box="[182,299,1356,1375]" metricMagnitude="3" metricUnit="m" metricValue="5.175" metricValueMax="5.35" metricValueMin="5.0" pageId="9" pageNumber="118" unit="m" value="5175.0" valueMax="5350.0" valueMin="5000.0">50005350 m</elevation>
</quantity>
,
<geoCoordinate id="DAE7504E8D70285FFED45E0172FFFAF3" box="[308,407,1356,1375]" degrees="30" direction="north" minutes="22" orientation="latitude" pageId="9" pageNumber="118" precision="15" seconds="47" value="30.379723">30°2247″N</geoCoordinate>
<geoCoordinate id="DAE7504E8D70285FFE7D5E01716EFAF3" box="[413,518,1356,1375]" degrees="90" direction="east" minutes="43" orientation="longitude" pageId="9" pageNumber="118" precision="15" seconds="22" value="90.72278">90°4322″E</geoCoordinate>
,
<date id="CB6D10498D70285FFDEF5E017118FAF3" box="[527,624,1356,1375]" pageId="9" pageNumber="118" value="2010-07-20">
<collectingDate id="DB29E9A18D70285FFDEF5E017118FAF3" box="[527,624,1356,1375]" pageId="9" pageNumber="118" value="2010-07-20">20.vii.2010</collectingDate>
</date>
, J.
<collectorName id="1226535F8D70285FFD6B5E0171B9FAF3" box="[651,721,1356,1375]" pageId="9" pageNumber="118">Schmidt</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D70285FFD1C5E017390FA65" collectingDate="2005-06-11" collectingDateMax="2015-06-28" collectingDateMin="2005-06-11" collectionCode="NMPC" collectorName="J. Schmidt &amp; J. Hajek &amp; D. Kral &amp; J. Ruzicka &amp; Gangca Dasi" country="China" elevation="5060" latitude="35.191666" location="Monda La" longLatPrecision="119" longitude="102.51" municipality="Pumo Tso" pageId="9" pageNumber="118" specimenCount="14" specimenCount-female="1" stateProvince="Gansu">
<specimenCount id="A9D5FD008D70285FFD1C5E017074FAF2" box="[764,796,1349,1375]" pageId="9" pageNumber="118" type="female">1 ♀</specimenCount>
(
<collectionCode id="D9C2AE4C8D70285FFCC75E017000FAF3" box="[807,872,1356,1375]" country="Czech Republic" name="National Museum Prague" pageId="9" pageNumber="118">NMPC</collectionCode>
):
<collectingMunicipality id="5F08ACF38D70285FFC965E0170A1FAF3" box="[886,969,1356,1375]" pageId="9" pageNumber="118">Pumo Tso</collectingMunicipality>
to
<location id="BA0C60528D70285FFC045E017755FAF3" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D70285FFC045E017755FAF3" box="[996,1085,1356,1375]" country="China" latitude="35.191666" longLatPrecision="119" longitude="102.51" municipality="Pumo Tso" name="Monda La" pageId="9" pageNumber="118" stateProvince="Gansu">Monda La</location>
,
<quantity id="782B9B6C8D70285FFF915E2B738EFAD5" box="[113,230,1382,1401]" metricMagnitude="3" metricUnit="m" metricValue="5.06" metricValueMax="5.12" metricValueMin="5.0" pageId="9" pageNumber="118" unit="m" value="5060.0" valueMax="5120.0" valueMin="5000.0">
<elevation id="34FED1BA8D70285FFF915E2B738EFAD5" box="[113,230,1382,1401]" metricMagnitude="3" metricUnit="m" metricValue="5.06" metricValueMax="5.12" metricValueMin="5.0" pageId="9" pageNumber="118" unit="m" value="5060.0" valueMax="5120.0" valueMin="5000.0">50005120 m</elevation>
</quantity>
,
<date id="CB6D10498D70285FFF0E5E2B7221FAD5" box="[238,329,1382,1401]" pageId="9" pageNumber="118" value="2015-06-28">
<collectingDate id="DB29E9A18D70285FFF0E5E2B7221FAD5" box="[238,329,1382,1401]" pageId="9" pageNumber="118" value="2015-06-28">28.vi.2015</collectingDate>
</date>
,
<geoCoordinate id="DAE7504E8D70285FFEB05E2B72FEFAD5" box="[336,406,1382,1401]" degrees="28" direction="north" minutes="29" orientation="latitude" pageId="9" pageNumber="118" precision="925" value="28.483334">28°29N</geoCoordinate>
<geoCoordinate id="DAE7504E8D70285FFE7A5E2B7288FAD5" box="[410,480,1382,1401]" degrees="90" direction="east" minutes="33" orientation="longitude" pageId="9" pageNumber="118" precision="925" value="90.55">90°33E</geoCoordinate>
,
<collectorName id="1226535F8D70285FFE075E2B7157FAD5" box="[487,575,1382,1401]" pageId="9" pageNumber="118">J. Schmidt</collectorName>
lgt. [DNA voucher MF1248].
<emphasis id="8DA7EA9B8D70285FFCDA5E2B7011FAD5" bold="true" box="[826,889,1382,1401]" pageId="9" pageNumber="118">
<collectingRegion id="7D17F86B8D70285FFCDA5E2B701BFAD4" box="[826,883,1382,1401]" country="China" name="Gansu" pageId="9" pageNumber="118">GANSU</collectingRegion>
:
</emphasis>
<specimenCount id="A9D5FD008D70285FFC9E5E2B70D0FAD5" box="[894,952,1382,1401]" pageId="9" pageNumber="118" type="generic">9 spec.</specimenCount>
(
<collectionCode id="D9C2AE4C8D70285FFC215E2B776AFAD5" box="[961,1026,1382,1401]" country="Czech Republic" name="National Museum Prague" pageId="9" pageNumber="118">NMPC</collectionCode>
):Xiahe (= Labrang) env.,
<geoCoordinate id="DAE7504E8D70285FFEE55ECC7232FA38" box="[261,346,1409,1428]" degrees="35" direction="north" minutes="11.5" orientation="latitude" pageId="9" pageNumber="118" precision="92" value="35.191666">35°11.5N</geoCoordinate>
<geoCoordinate id="DAE7504E8D70285FFEBE5ECC72D6FA38" box="[350,446,1409,1428]" degrees="102" direction="east" minutes="30.6" orientation="longitude" pageId="9" pageNumber="118" precision="92" value="102.51">102°30.6E</geoCoordinate>
,
<quantity id="782B9B6C8D70285FFE255ECC716EFA38" box="[453,518,1409,1428]" metricMagnitude="3" metricUnit="m" metricValue="2.94" pageId="9" pageNumber="118" unit="m" value="2940.0">
<elevation id="34FED1BA8D70285FFE255ECC716EFA38" box="[453,518,1409,1428]" metricMagnitude="3" metricUnit="m" metricValue="2.94" pageId="9" pageNumber="118" unit="m" value="2940.0">2940 m</elevation>
</quantity>
,
<date id="CB6D10498D70285FFDEE5ECC71E5FA38" box="[526,653,1409,1428]" pageId="9" pageNumber="118" value="2005-06-19" valueMax="2005-06-22" valueMin="2005-06-19">
<collectingDate id="DB29E9A18D70285FFDEE5ECC71E5FA38" box="[526,653,1409,1428]" pageId="9" pageNumber="118" value="2005-06-19" valueMax="2005-06-22" valueMin="2005-06-19">19.22.vi.2005</collectingDate>
</date>
,
<collectorName id="1226535F8D70285FFD755ECC71B3FA38" box="[661,731,1409,1428]" pageId="9" pageNumber="118">J. Hájek</collectorName>
,
<collectorName id="1226535F8D70285FFD025ECC7048FA38" box="[738,800,1409,1428]" pageId="9" pageNumber="118">D. Král</collectorName>
&amp;
<collectorName id="1226535F8D70285FFCDA5ECC70F8FA38" box="[826,912,1409,1428]" pageId="9" pageNumber="118">J. Růžička</collectorName>
lgt.
<emphasis id="8DA7EA9B8D70285FFC545ECC776BFA38" bold="true" box="[948,1027,1409,1428]" pageId="9" pageNumber="118">
<collectingRegion id="7D17F86B8D70285FFC545ECC7094FA3E" box="[948,1020,1409,1428]" country="China" name="Qinghai" pageId="9" pageNumber="118">QINGHAI</collectingRegion>
:
</emphasis>
<specimenCount id="A9D5FD008D70285FFBE75ECC772AFA38" box="[1031,1090,1409,1428]" pageId="9" pageNumber="118" type="generic">4 spec.</specimenCount>
(
<collectionCode id="D9C2AE4C8D70285FFF975ED173D0FA03" box="[119,184,1436,1455]" country="Czech Republic" name="National Museum Prague" pageId="9" pageNumber="118">NMPC</collectionCode>
):
<collectorName id="1226535F8D70285FFF255ED17258FA03" box="[197,304,1436,1455]" pageId="9" pageNumber="118">Gangca Dasi</collectorName>
(lamasery), 37°32.433.0N 100°05.306.0E,
<quantity id="782B9B6C8D70285FFD565ED17043FA03" box="[694,811,1436,1455]" metricMagnitude="3" metricUnit="m" metricValue="3.6725" metricValueMax="3.84" metricValueMin="3.505" pageId="9" pageNumber="118" unit="m" value="3672.5" valueMax="3840.0" valueMin="3505.0">
<elevation id="34FED1BA8D70285FFD565ED17043FA03" box="[694,811,1436,1455]" metricMagnitude="3" metricUnit="m" metricValue="3.6725" metricValueMax="3.84" metricValueMin="3.505" pageId="9" pageNumber="118" unit="m" value="3672.5" valueMax="3840.0" valueMin="3505.0">35053840 m</elevation>
</quantity>
,
<date id="CB6D10498D70285FFCD35ED170C5FA03" box="[819,941,1436,1455]" pageId="9" pageNumber="118" value="2005-06-11" valueMax="2005-06-12" valueMin="2005-06-11">
<collectingDate id="DB29E9A18D70285FFCD35ED170C5FA03" box="[819,941,1436,1455]" pageId="9" pageNumber="118" value="2005-06-11" valueMax="2005-06-12" valueMin="2005-06-11">1112.vi.2005</collectingDate>
</date>
,
<collectorName id="1226535F8D70285FFC555ED17093FA03" box="[949,1019,1436,1455]" pageId="9" pageNumber="118">J. Hájek</collectorName>
,
<collectorName id="1226535F8D70285FFBE25ED17728FA03" box="[1026,1088,1436,1455]" pageId="9" pageNumber="118">D. Král</collectorName>
&amp;
<collectorName id="1226535F8D70285FFF655EFB73B3FA65" box="[133,219,1462,1481]" pageId="9" pageNumber="118">J.Růžička</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D70285FFEE35EFB7211FA48" collectingDate="2005-07-13" collectingDateMax="2005-07-15" collectingDateMin="2005-07-13" collectionCode="NMPC" collectorName="J. Hajek &amp; D. Kral &amp; Ruzicka" country="China" elevation="3230" location="Haibu" pageId="9" pageNumber="118" specimenCount="5" stateProvince="Gansu">
1 J
<specimenCount id="A9D5FD008D70285FFEC45EFB7236FA65" box="[292,350,1462,1481]" pageId="9" pageNumber="118" type="generic">5 spec.</specimenCount>
(
<collectionCode id="D9C2AE4C8D70285FFE865EFB72CFFA65" box="[358,423,1462,1481]" country="Czech Republic" name="National Museum Prague" pageId="9" pageNumber="118">NMPC</collectionCode>
):
<location id="BA0C60528D70285FFE535EFB728EFA65" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D70285FFE535EFB728EFA65" box="[435,486,1462,1481]" country="China" name="Haibu" pageId="9" pageNumber="118" stateProvince="Gansu">Haibu</location>
env.,
<quantity id="782B9B6C8D70285FFDF45EFB71E0FA65" box="[532,648,1462,1481]" metricMagnitude="3" metricUnit="m" metricValue="3.2299999999999995" metricValueMax="3.27" metricValueMin="3.19" pageId="9" pageNumber="118" unit="m" value="3230.0" valueMax="3270.0" valueMin="3190.0">
<elevation id="34FED1BA8D70285FFDF45EFB71E0FA65" box="[532,648,1462,1481]" metricMagnitude="3" metricUnit="m" metricValue="3.2299999999999995" metricValueMax="3.27" metricValueMin="3.19" pageId="9" pageNumber="118" unit="m" value="3230.0" valueMax="3270.0" valueMin="3190.0">31903270 m</elevation>
</quantity>
,36°48.449.8N 100°45.449.7E,
<date id="CB6D10498D70285FFC4C5EFB7744FA65" box="[940,1068,1462,1481]" pageId="9" pageNumber="118" value="2005-07-13" valueMax="2005-07-15" valueMin="2005-07-13">
<collectingDate id="DB29E9A18D70285FFC4C5EFB7744FA65" box="[940,1068,1462,1481]" pageId="9" pageNumber="118" value="2005-07-13" valueMax="2005-07-15" valueMin="2005-07-13">1315.vii.2005</collectingDate>
</date>
,
<collectorName id="1226535F8D70285FFBD35EFB73CDFA48" pageId="9" pageNumber="118">J. Hájek</collectorName>
,
<collectorName id="1226535F8D70285FFF4D5E9C7383FA48" box="[173,235,1489,1508]" pageId="9" pageNumber="118">D. Král</collectorName>
&amp; J.
<collectorName id="1226535F8D70285FFEF65E9C7233FA48" box="[278,347,1489,1508]" pageId="9" pageNumber="118">Růžička</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D70285FFE645E9C7229FA53" collectingDate="2005-07-03" collectingDateMax="2005-07-05" collectingDateMin="2005-07-03" collectionCode="NMPC" collectorName="Hajek &amp; Kral &amp; Ruzicka" country="China" elevation="3240" latitude="37.168335" location="Hairag" longLatPrecision="117" longitude="100.41167" pageId="9" pageNumber="118" specimenCount="3" stateProvince="Gansu">
1 J
<specimenCount id="A9D5FD008D70285FFE475E9C728AFA48" box="[423,482,1489,1508]" pageId="9" pageNumber="118" type="generic">3 spec.</specimenCount>
(
<collectionCode id="D9C2AE4C8D70285FFE0B5E9C7144FA48" box="[491,556,1489,1508]" country="Czech Republic" name="National Museum Prague" pageId="9" pageNumber="118">NMPC</collectionCode>
):
<location id="BA0C60528D70285FFDD95E9C711CFA48" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D70285FFDD95E9C711CFA48" box="[569,628,1489,1508]" country="China" latitude="37.168335" longLatPrecision="117" longitude="100.41167" name="Hairag" pageId="9" pageNumber="118" stateProvince="Gansu">Hairag</location>
,
<quantity id="782B9B6C8D70285FFD9B5E9C71D3FA48" box="[635,699,1489,1508]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" pageId="9" pageNumber="118" unit="m" value="3240.0">
<elevation id="34FED1BA8D70285FFD9B5E9C71D3FA48" box="[635,699,1489,1508]" metricMagnitude="3" metricUnit="m" metricValue="3.2399999999999998" pageId="9" pageNumber="118" unit="m" value="3240.0">3240 m</elevation>
</quantity>
,
<geoCoordinate id="DAE7504E8D70285FFD245E9C7071FA48" box="[708,793,1489,1508]" degrees="37" direction="north" minutes="10.1" orientation="latitude" pageId="9" pageNumber="118" precision="92" value="37.168335">37°10.1N</geoCoordinate>
<geoCoordinate id="DAE7504E8D70285FFCFD5E9C7015FA48" box="[797,893,1489,1508]" degrees="100" direction="east" minutes="24.7" orientation="longitude" pageId="9" pageNumber="118" precision="92" value="100.41167">100°24.7E</geoCoordinate>
,
<date id="CB6D10498D70285FFC655E9C7098FA48" box="[901,1008,1489,1508]" pageId="9" pageNumber="118" value="2005-07-03" valueMax="2005-07-05" valueMin="2005-07-03">
<collectingDate id="DB29E9A18D70285FFC655E9C7098FA48" box="[901,1008,1489,1508]" pageId="9" pageNumber="118" value="2005-07-03" valueMax="2005-07-05" valueMin="2005-07-03">35.vii.2005</collectingDate>
</date>
, J.
<collectorName id="1226535F8D70285FFBE95E9C7755FA48" box="[1033,1085,1489,1508]" pageId="9" pageNumber="118">Hájek</collectorName>
, D.
<collectorName id="1226535F8D70285FFF6B5EA173D8FA53" box="[139,176,1516,1535]" pageId="9" pageNumber="118">Král</collectorName>
&amp; J.
<collectorName id="1226535F8D70285FFF3D5EA1724AFA53" box="[221,290,1516,1535]" pageId="9" pageNumber="118">Růžička</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D70285FFEAD5EA171D2F9B5" collectingDate="2013-06-08" collectorName="R. B. Angus &amp; F. L. Jia &amp; Zhang" country="China" elevation="4290" latitude="34.85472" location="Maduo" longLatPrecision="19" longitude="98.28833" municipality="Golo" pageId="9" pageNumber="118" specimenCount="1" stateProvince="Gansu">
1 J (BMNH):
<collectingMunicipality id="5F08ACF38D70285FFE295EA1729DFA53" box="[457,501,1516,1535]" pageId="9" pageNumber="118">Golo</collectingMunicipality>
,
<location id="BA0C60528D70285FFE1D5EA17153FA53" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D70285FFE1D5EA17153FA53" box="[509,571,1516,1535]" country="China" latitude="34.85472" longLatPrecision="19" longitude="98.28833" municipality="Golo" name="Maduo" pageId="9" pageNumber="118" stateProvince="Gansu">Maduo</location>
, roadside pools on river flats
<locationDeviation id="AEDE2FAB8D70285FFCCF5EA170BCFA53" box="[815,980,1516,1535]" location="Maduo." pageId="9" pageNumber="118">
<quantity id="782B9B6C8D70285FFCCF5EA1700CFA53" box="[815,868,1516,1535]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="118" unit="km" value="20.0">20 km</quantity>
E of Maduo.
</locationDeviation>
<geoCoordinate id="DAE7504E8D70285FFC385EA17755FA53" box="[984,1085,1516,1535]" degrees="34" direction="north" minutes="51" orientation="latitude" pageId="9" pageNumber="118" precision="15" seconds="17" value="34.85472">34°5117″N</geoCoordinate>
,
<geoCoordinate id="DAE7504E8D70285FFF915D4B73BCF9B5" box="[113,212,1542,1561]" degrees="98" direction="east" minutes="17" orientation="longitude" pageId="9" pageNumber="118" precision="15" seconds="18" value="98.28833">98°1718″E</geoCoordinate>
,
<quantity id="782B9B6C8D70285FFF3D5D4B7276F9B5" box="[221,286,1542,1561]" metricMagnitude="3" metricUnit="m" metricValue="4.29" pageId="9" pageNumber="118" unit="m" value="4290.0">
<elevation id="34FED1BA8D70285FFF3D5D4B7276F9B5" box="[221,286,1542,1561]" metricMagnitude="3" metricUnit="m" metricValue="4.29" pageId="9" pageNumber="118" unit="m" value="4290.0">4290 m</elevation>
</quantity>
,
<date id="CB6D10498D70285FFEC75D4B721FF9B5" box="[295,375,1542,1561]" pageId="9" pageNumber="118" value="2013-06-08">
<collectingDate id="DB29E9A18D70285FFEC75D4B721FF9B5" box="[295,375,1542,1561]" pageId="9" pageNumber="118" value="2013-06-08">8.vi.2013</collectingDate>
</date>
,
<collectorName id="1226535F8D70285FFE605D4B7281F9B5" box="[384,489,1542,1561]" pageId="9" pageNumber="118">R. B. Angus</collectorName>
,
<collectorName id="1226535F8D70285FFE115D4B715CF9B5" box="[497,564,1542,1561]" pageId="9" pageNumber="118">F. L. Jia</collectorName>
&amp; Y.
<collectorName id="1226535F8D70285FFD855D4B71F3F9B5" box="[613,667,1542,1561]" pageId="9" pageNumber="118">Zhang</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D70285FFD265D4B70B9F998" collectingDate="2013-06-07" collectorName="Angus &amp; Jia &amp; Zhang" country="China" county="Golo" elevation="4141" latitude="35.331112" location="Zuimatan" longLatPrecision="19" longitude="99.065834" municipality="Huamuxia" pageId="9" pageNumber="118" specimenCount="6" specimenCount-female="6" stateProvince="Gansu">
2 JJ
<specimenCount id="A9D5FD008D70285FFD1A5D4B7041F9B5" box="[762,809,1536,1561]" pageId="9" pageNumber="118" type="female">6 ♀♀</specimenCount>
(BMNH):
<collectingCounty id="560D4E058D70285FFC665D4B70DAF9B5" box="[902,946,1542,1561]" pageId="9" pageNumber="118">Golo</collectingCounty>
,
<collectingMunicipality id="5F08ACF38D70285FFC5B5D4B777CF9B5" box="[955,1044,1542,1561]" pageId="9" pageNumber="118">Huamuxia</collectingMunicipality>
,
<location id="BA0C60528D70285FFBFC5D4B73CEF998" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D70285FFBFC5D4B73CEF998" country="China" county="Golo" latitude="35.331112" longLatPrecision="19" longitude="99.065834" municipality="Huamuxia" name="Zuimatan" pageId="9" pageNumber="118" stateProvince="Gansu">Zuimatan</location>
, roadside pool,
<geoCoordinate id="DAE7504E8D70285FFEC85D6C72E5F998" box="[296,397,1569,1588]" degrees="35" direction="north" minutes="19" orientation="latitude" pageId="9" pageNumber="118" precision="15" seconds="52" value="35.331112">35°1952″N</geoCoordinate>
,
<geoCoordinate id="DAE7504E8D70285FFE745D6C7290F998" box="[404,504,1569,1588]" degrees="99" direction="east" minutes="03" orientation="longitude" pageId="9" pageNumber="118" precision="15" seconds="57" value="99.065834">99°0357″E</geoCoordinate>
,
<quantity id="782B9B6C8D70285FFE1F5D6C7157F998" box="[511,575,1569,1588]" metricMagnitude="3" metricUnit="m" metricValue="4.141" pageId="9" pageNumber="118" unit="m" value="4141.0">
<elevation id="34FED1BA8D70285FFE1F5D6C7157F998" box="[511,575,1569,1588]" metricMagnitude="3" metricUnit="m" metricValue="4.141" pageId="9" pageNumber="118" unit="m" value="4141.0">4141 m</elevation>
</quantity>
,
<date id="CB6D10498D70285FFDA75D6C71F0F998" box="[583,664,1569,1588]" pageId="9" pageNumber="118" value="2013-06-07">
<collectingDate id="DB29E9A18D70285FFDA75D6C71F0F998" box="[583,664,1569,1588]" pageId="9" pageNumber="118" value="2013-06-07">7.vi.2013</collectingDate>
</date>
, R. B.
<collectorName id="1226535F8D70285FFD2D5D6C706EF998" box="[717,774,1569,1588]" pageId="9" pageNumber="118">Angus</collectorName>
, F. L.
<collectorName id="1226535F8D70285FFCD75D6C7026F998" box="[823,846,1569,1588]" pageId="9" pageNumber="118">Jia</collectorName>
&amp; Y.
<collectorName id="1226535F8D70285FFC9D5D6C70DBF998" box="[893,947,1569,1588]" pageId="9" pageNumber="118">Zhang</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D70285AFC3C5D6C729CFB3B" collectionCode="SYSU" country="China" lastPageId="12" lastPageNumber="121" pageId="9" pageNumber="118" specimenCount="2" specimenCount-female="2" stateProvince="Gansu">
3 JJ
<specimenCount id="A9D5FD008D70285FFBEE5D6C7752F998" box="[1038,1082,1563,1588]" pageId="9" pageNumber="118" type="female">2♀♀</specimenCount>
, 6 unsexed spec. (
<collectionCode id="D9C2AE4C8D75285AFEC85FC9720AFB3B" box="[296,354,1156,1175]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="12" pageNumber="121" type="Herbarium">SYSU</collectionCode>
): same label data
</materialsCitation>
;
<materialsCitation id="0FBB3CD48D75285AFE1E5FC9773BFB3B" box="[510,1107,1156,1175]" collectionCode="NMPC" country="China" pageId="12" pageNumber="121" specimenCount="2" stateProvince="Gansu">
<specimenCount id="A9D5FD008D75285AFE1E5FC97152FB3B" box="[510,570,1156,1175]" pageId="12" pageNumber="121" type="generic">2 spec.</specimenCount>
(
<collectionCode id="D9C2AE4C8D75285AFDA45FC971EDFB3B" box="[580,645,1156,1175]" country="Czech Republic" name="National Museum Prague" pageId="12" pageNumber="121">NMPC</collectionCode>
): same label data [DNA vouchers MF658 and MF659]
</materialsCitation>
;
<materialsCitation id="0FBB3CD48D75285AFBBC5FC97741FB1E" collectingDate="1990-08-15" collectionCode="SYSU" collectorName="Xuezhong Zhang" country="China" elevation="5130" location="Kangzhagri" municipality="Kekexili" pageId="12" pageNumber="121" specimenCount="1" stateProvince="Gansu">
1 J, 2 unsexed spec. (
<collectionCode id="D9C2AE4C8D75285AFEA55FD27217FB1E" box="[325,383,1183,1202]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="12" pageNumber="121" type="Herbarium">SYSU</collectionCode>
):
<collectingMunicipality id="5F08ACF38D75285AFE6E5FD272BEFB1E" box="[398,470,1183,1202]" pageId="12" pageNumber="121">Kekexili</collectingMunicipality>
[= Kekesili],
<location id="BA0C60528D75285AFDAE5FD271DAFB1E" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D75285AFDAE5FD271DAFB1E" box="[590,690,1183,1202]" country="China" municipality="Kekexili" name="Kangzhagri" pageId="12" pageNumber="121" stateProvince="Gansu">Kangzhagri</location>
,
<quantity id="782B9B6C8D75285AFD5B5FD27195FB1E" box="[699,765,1183,1202]" metricMagnitude="3" metricUnit="m" metricValue="5.13" pageId="12" pageNumber="121" unit="m" value="5130.0">
<elevation id="34FED1BA8D75285AFD5B5FD27195FB1E" box="[699,765,1183,1202]" metricMagnitude="3" metricUnit="m" metricValue="5.13" pageId="12" pageNumber="121" unit="m" value="5130.0">5130 m</elevation>
</quantity>
,
<date id="CB6D10498D75285AFCE75FD27006FB1E" box="[775,878,1183,1202]" pageId="12" pageNumber="121" value="1990-08-15">
<collectingDate id="DB29E9A18D75285AFCE75FD27006FB1E" box="[775,878,1183,1202]" pageId="12" pageNumber="121" value="1990-08-15">15.viii.1990</collectingDate>
</date>
,
<collectorName id="1226535F8D75285AFC975FD27761FB1E" box="[887,1033,1183,1202]" pageId="12" pageNumber="121">Xuezhong Zhang</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D75285AFBD65FD2722EFB4B" collectingDate="1990-08-02" collectionCode="SYSU" collectorName="Kekexili &amp; Xinqingfeng &amp; Xuezhong Zhang" country="China" elevation="4900" location="Mt." pageId="12" pageNumber="121" specimenCount="1" specimenCount-female="1" stateProvince="Gansu">
2 JJ
<specimenCount id="A9D5FD008D75285AFF775FF473D0FB60" box="[151,184,1203,1228]" pageId="12" pageNumber="121" type="female">1 ♀</specimenCount>
(
<collectionCode id="D9C2AE4C8D75285AFF245FF47396FB60" box="[196,254,1209,1228]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="12" pageNumber="121" type="Herbarium">SYSU</collectionCode>
):
<collectorName id="1226535F8D75285AFEEE5FF47230FB60" box="[270,344,1209,1228]" pageId="12" pageNumber="121">Kekexili</collectorName>
,
<collectorName id="1226535F8D75285AFE815FF472A3FB60" box="[353,459,1209,1228]" pageId="12" pageNumber="121">Xinqingfeng</collectorName>
[= Syn Qing Feng, or Buka Daban Feng]
<location id="BA0C60528D75285AFCD65FF4703DFB60" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D75285AFCD65FF4703DFB60" box="[822,853,1209,1228]" country="China" name="Mt." pageId="12" pageNumber="121" stateProvince="Gansu">Mt.</location>
hot spring,
<quantity id="782B9B6C8D75285AFC5B5FF47095FB60" box="[955,1021,1209,1228]" metricMagnitude="3" metricUnit="m" metricValue="4.9" pageId="12" pageNumber="121" unit="m" value="4900.0">
<elevation id="34FED1BA8D75285AFC5B5FF47095FB60" box="[955,1021,1209,1228]" metricMagnitude="3" metricUnit="m" metricValue="4.9" pageId="12" pageNumber="121" unit="m" value="4900.0">4900 m</elevation>
</quantity>
,
<date id="CB6D10498D75285AFBE75FF4770BFB60" box="[1031,1123,1209,1228]" pageId="12" pageNumber="121" value="1990-08-02">
<collectingDate id="DB29E9A18D75285AFBE75FF4770BFB60" box="[1031,1123,1209,1228]" pageId="12" pageNumber="121" value="1990-08-02">2.viii.1990</collectingDate>
</date>
,
<collectorName id="1226535F8D75285AFF775F997240FB4B" box="[151,296,1236,1255]" pageId="12" pageNumber="121">Xuezhong Zhang</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D75285AFEB15F997708FB4B" box="[337,1120,1231,1256]" collectingDate="1990-08-01" collectionCode="SYSU" collectorName="Xuezhong Zhang" country="China" elevation="4950" location="Malanshan" municipality="Kekexili" pageId="12" pageNumber="121" specimenCount="1" stateProvince="Gansu">
1 J, 2 unsexed spec. (
<collectionCode id="D9C2AE4C8D75285AFDE65F997128FB4B" box="[518,576,1236,1255]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="12" pageNumber="121" type="Herbarium">SYSU</collectionCode>
):
<collectingMunicipality id="5F08ACF38D75285AFDAE5F9971F0FB4B" box="[590,664,1236,1255]" pageId="12" pageNumber="121">Kekexili</collectingMunicipality>
,
<location id="BA0C60528D75285AFD7F5F997195FB4B" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D75285AFD7F5F997195FB4B" box="[671,765,1236,1255]" country="China" municipality="Kekexili" name="Malanshan" pageId="12" pageNumber="121" stateProvince="Gansu">Malanshan</location>
,
<quantity id="782B9B6C8D75285AFCE55F99702DFB4B" box="[773,837,1236,1255]" metricMagnitude="3" metricUnit="m" metricValue="4.95" pageId="12" pageNumber="121" unit="m" value="4950.0">
<elevation id="34FED1BA8D75285AFCE55F99702DFB4B" box="[773,837,1236,1255]" metricMagnitude="3" metricUnit="m" metricValue="4.95" pageId="12" pageNumber="121" unit="m" value="4950.0">4950 m</elevation>
</quantity>
,
<date id="CB6D10498D75285AFCAD5F9970C1FB4B" box="[845,937,1236,1255]" pageId="12" pageNumber="121" value="1990-08-01">
<collectingDate id="DB29E9A18D75285AFCAD5F9970C1FB4B" box="[845,937,1236,1255]" pageId="12" pageNumber="121" value="1990-08-01">1.viii.1990</collectingDate>
</date>
,
<collectorName id="1226535F8D75285AFC515F997729FB4B" box="[945,1089,1236,1255]" pageId="12" pageNumber="121">Xuezhong Zhang</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D75285AFF775FA27020FAAE" box="[151,840,1257,1282]" collectingDate="1990-08-10" collectionCode="SYSU" collectorName="Xuezhong Zhang" country="China" elevation="4600" location="Kusaihu lake" municipality="Kekexili" pageId="12" pageNumber="121" specimenCount="1" stateProvince="Gansu">
1 J (
<collectionCode id="D9C2AE4C8D75285AFF245FA27396FAAE" box="[196,254,1263,1282]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="12" pageNumber="121" type="Herbarium">SYSU</collectionCode>
):
<collectingMunicipality id="5F08ACF38D75285AFEED5FA2723FFAAE" box="[269,343,1263,1282]" pageId="12" pageNumber="121">Kekexili</collectingMunicipality>
,
<location id="BA0C60528D75285AFE805FA272B9FAAE" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D75285AFE805FA272B9FAAE" box="[352,465,1263,1282]" country="China" municipality="Kekexili" name="Kusaihu lake" pageId="12" pageNumber="121" stateProvince="Gansu">Kusaihu lake</location>
,
<quantity id="782B9B6C8D75285AFE3A5FA27174FAAE" box="[474,540,1263,1282]" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="12" pageNumber="121" unit="m" value="4600.0">
<elevation id="34FED1BA8D75285AFE3A5FA27174FAAE" box="[474,540,1263,1282]" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="12" pageNumber="121" unit="m" value="4600.0">4600 m</elevation>
</quantity>
,
<date id="CB6D10498D75285AFDC65FA271E4FAAE" box="[550,652,1263,1282]" pageId="12" pageNumber="121" value="1990-08-10">
<collectingDate id="DB29E9A18D75285AFDC65FA271E4FAAE" box="[550,652,1263,1282]" pageId="12" pageNumber="121" value="1990-08-10">10.viii.1990</collectingDate>
</date>
,
<collectorName id="1226535F8D75285AFD765FA27040FAAE" box="[662,808,1263,1282]" pageId="12" pageNumber="121">Xuezhong Zhang</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D75285AFCB45FA27201FAFE" collectingDate="1932-08-10" collectingDateMax="2013-06-08" collectingDateMin="1932-08-10" collectionCode="SYSU, IRSNB" collectorName="R. B. Angus &amp; F. L. Jia &amp; Y. Zhang &amp; Indian Tibet, L &amp; Chushol &amp; Orchymont" country="India" county="Golo" elevation="4240" latitude="34.67972" location="Yematan" longLatPrecision="19" longitude="99.065834" municipality="Maduo" pageId="12" pageNumber="121" specimenCount="2" specimenCount-female="2" stateProvince="Jammu and Kashmir">
1 J
<specimenCount id="A9D5FD008D75285AFC9B5FA270F5FAAD" box="[891,925,1256,1282]" pageId="12" pageNumber="121" type="female">1 ♀</specimenCount>
(
<collectionCode id="D9C2AE4C8D75285AFC475FA27089FAAE" box="[935,993,1263,1282]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="12" pageNumber="121" type="Herbarium">SYSU</collectionCode>
):
<collectingCounty id="560D4E058D75285AFC115FA27775FAAE" box="[1009,1053,1263,1282]" pageId="12" pageNumber="121">Golo</collectingCounty>
,
<collectingMunicipality id="5F08ACF38D75285AFBC65FA2770CFAAE" box="[1062,1124,1263,1282]" pageId="12" pageNumber="121">Maduo</collectingMunicipality>
, roadside pools near
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,
<geoCoordinate id="DAE7504E8D75285AFE705E44729BFAB0" box="[400,499,1289,1308]" degrees="34" direction="north" minutes="40" orientation="latitude" pageId="12" pageNumber="121" precision="15" seconds="47" value="34.67972">34°4047″N</geoCoordinate>
<geoCoordinate id="DAE7504E8D75285AFE185E447133FAB0" box="[504,603,1289,1308]" degrees="99" direction="east" minutes="03" orientation="longitude" pageId="12" pageNumber="121" precision="15" seconds="57" value="99.065834">99°0357″E</geoCoordinate>
,
<elevation id="34FED1BA8D75285AFD835E4471A7FAB0" box="[611,719,1289,1308]" metricMagnitude="3" metricUnit="m" metricValue="4.24" pageId="12" pageNumber="121" unit="m" value="4240.0">
<quantity id="782B9B6C8D75285AFD835E4471CBFAB0" box="[611,675,1289,1308]" metricMagnitude="3" metricUnit="m" metricValue="4.24" pageId="12" pageNumber="121" unit="m" value="4240.0">4240 m</quantity>
a.s.l.
</elevation>
,
<date id="CB6D10498D75285AFD375E44704FFAB0" box="[727,807,1289,1308]" pageId="12" pageNumber="121" value="2013-06-08">
<collectingDate id="DB29E9A18D75285AFD375E44704FFAB0" box="[727,807,1289,1308]" pageId="12" pageNumber="121" value="2013-06-08">8.vi.2013</collectingDate>
</date>
,
<collectorName id="1226535F8D75285AFCD05E4470F0FAB0" box="[816,920,1289,1308]" pageId="12" pageNumber="121">R. B. Angus</collectorName>
,
<collectorName id="1226535F8D75285AFC7F5E447089FAB0" box="[927,993,1289,1308]" pageId="12" pageNumber="121">F. L. Jia</collectorName>
&amp;
<collectorName id="1226535F8D75285AFC1A5E44772FFAB0" box="[1018,1095,1289,1308]" pageId="12" pageNumber="121">Y. Zhang</collectorName>
lgt.
<emphasis id="8DA7EA9B8D75285AFF775E697274FA99" bold="true" box="[151,284,1316,1335]" pageId="12" pageNumber="121">
<collectingCountry id="C7C476198D75285AFF775E6973B0FA9B" box="[151,216,1316,1335]" name="India" pageId="12" pageNumber="121">INDIA</collectingCountry>
:
<collectingRegion id="7D17F86B8D75285AFF015E697274FA99" box="[225,284,1316,1335]" country="India" name="Jammu and Kashmir" pageId="12" pageNumber="121">JAMMU</collectingRegion>
</emphasis>
AND
<emphasis id="8DA7EA9B8D75285AFEA65E6972F1FA9B" bold="true" box="[326,409,1316,1335]" pageId="12" pageNumber="121">KASHMIR:</emphasis>
<specimenCount id="A9D5FD008D75285AFE7E5E6972D5FA9B" box="[414,445,1310,1335]" pageId="12" pageNumber="121" type="female">1 ♀</specimenCount>
(
<collectionCode id="D9C2AE4C8D75285AFE285E697163FA9B" box="[456,523,1316,1335]" country="Belgium" name="Institut Royal des Sciences Naturelles de Belgique" pageId="12" pageNumber="121">IRSNB</collectionCode>
):
<collectorName id="1226535F8D75285AFDF85E6971FCFA9B" box="[536,660,1316,1335]" pageId="12" pageNumber="121">Indian Tibet, L</collectorName>
73
<collectorName id="1226535F8D75285AFD4E5E69719EFA9B" box="[686,758,1316,1335]" pageId="12" pageNumber="121">Chushol</collectorName>
, pond below village,
<date id="CB6D10498D75285AFC475E697763FA9B" box="[935,1035,1316,1335]" pageId="12" pageNumber="121" value="1932-08-10">
<collectingDate id="DB29E9A18D75285AFC475E697763FA9B" box="[935,1035,1316,1335]" pageId="12" pageNumber="121" value="1932-08-10">10.viii.1932</collectingDate>
</date>
(identified as
<taxonomicName id="78D34D0A8D75285AFF4E5E727389FAFE" authorityName="Zaitzev" authorityYear="1908" box="[174,225,1343,1362]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="121" phylum="Arthropoda" rank="species" species="ser">
<emphasis id="8DA7EA9B8D75285AFF4E5E727389FAFE" box="[174,225,1343,1362]" italics="true" pageId="12" pageNumber="121">H. ser</emphasis>
</taxonomicName>
by
<collectorName id="1226535F8D75285AFEE05E72720BFAFE" box="[256,355,1343,1362]" pageId="12" pageNumber="121">Orchymont</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="EBAC66018D73285CFF775E8770E2F981" ID-DOI="http://doi.org/10.5281/zenodo.5311792" ID-Zenodo-Dep="5311792" httpUri="https://zenodo.org/record/5311792/files/figure.png" pageId="10" pageNumber="119" startId="10.[151,182,1482,1501]" targetBox="[159,1124,234,1349]" targetPageId="10">
<paragraph id="BF6C36898D73285CFF775E8770E2F981" blockId="10.[151,1127,1482,1581]" pageId="10" pageNumber="119">
Fig. 3. Habitus of the species of
<taxonomicName id="78D34D0A8D73285CFE495E8771FCFA71" authority="" authorityName="Zaitzev" authorityYear="1908" box="[425,660,1482,1501]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="subGenus" subGenus="Lihelophorus">
<emphasis id="8DA7EA9B8D73285CFE495E877166FA71" box="[425,526,1482,1501]" italics="true" pageId="10" pageNumber="119">Helophorus</emphasis>
(
<emphasis id="8DA7EA9B8D73285CFDF95E8771E3FA71" box="[537,651,1482,1501]" italics="true" pageId="10" pageNumber="119">Lihelophorus</emphasis>
)
</taxonomicName>
. ac
<taxonomicName id="78D34D0A8D73285CFD2E5E8770C3FA71" authority="Zaitzev, 1908" authorityName="Zaitzev" authorityYear="1908" box="[718,939,1482,1501]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="lamicola">
<emphasis id="8DA7EA9B8D73285CFD2E5E87705BFA71" box="[718,819,1482,1501]" italics="true" pageId="10" pageNumber="119">H. lamicola</emphasis>
<bibRefCitation id="DB424B788D73285CFCD75E8770C3FA71" author="ZAITZEV F. A." box="[823,939,1482,1501]" pageId="10" pageNumber="119" pagination="417 - 426" refId="ref20914" refString="ZAITZEV F. A. 1908: Beitrag zur Kenntniss der Waserkafer von Chinesisch-Centralasien. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St. Petersbourg 13: 417 - 426." type="journal article" year="1908">Zaitzev, 1908</bibRefCitation>
</taxonomicName>
(a Qinghai: Zhaling Hu (lectotype); bc Quinghai: Golo).dg
<taxonomicName id="78D34D0A8D73285CFDF05EA971D0FA5B" authority="Zaitzev, 1908" authorityName="Zaitzev" authorityYear="1908" box="[528,696,1508,1527]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="ser">
<emphasis id="8DA7EA9B8D73285CFDF05EA9712BFA5B" box="[528,579,1508,1527]" italics="true" pageId="10" pageNumber="119">H.ser</emphasis>
<bibRefCitation id="DB424B788D73285CFDA75EA971D0FA5B" author="ZAITZEV F. A." box="[583,696,1508,1527]" pageId="10" pageNumber="119" pagination="417 - 426" refId="ref20914" refString="ZAITZEV F. A. 1908: Beitrag zur Kenntniss der Waserkafer von Chinesisch-Centralasien. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St. Petersbourg 13: 417 - 426." type="journal article" year="1908">Zaitzev, 1908</bibRefCitation>
</taxonomicName>
(de Qinghai:Golo; f Xizang, Nyainqentanglha Feng; g Gansu: Xiahe env.). hk
<taxonomicName id="78D34D0A8D73285CFE355EB27146F9BE" authority="Angus &amp; Jia &amp; Chen &amp; Zhang &amp; Vondráček &amp; Fikáček &amp; Sw, 2016" authorityName="Angus &amp; Jia &amp; Chen &amp; Zhang &amp; Vondráček &amp; Fikáček &amp; Sw" authorityYear="2016" box="[469,558,1535,1554]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="119" phylum="Arthropoda" rank="species" species="yangae" status="sp. nov.">
<emphasis id="8DA7EA9B8D73285CFE355EB27146F9BE" box="[469,558,1535,1554]" italics="true" pageId="10" pageNumber="119">H. yangae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="969457E08D73285CFDD35EB2711FF9BE" box="[563,631,1535,1554]" pageId="10" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
(h Xizang: 90 km W of Amdo; i Xizang: 25 km W of Amdo; j Qinghai: Golo; k India: Kar Tso (Orchymont coll.)). All specimens to scale.
</paragraph>
</caption>
<caption id="EBAC66018D72285DFF915EB7772AF9EE" ID-DOI="http://doi.org/10.5281/zenodo.5311794" ID-Zenodo-Dep="5311794" httpUri="https://zenodo.org/record/5311794/files/figure.png" pageId="11" pageNumber="120" startId="11.[113,144,1530,1549]" targetBox="[131,1080,229,1416]" targetPageId="11">
<paragraph id="BF6C36898D72285DFF915EB7772AF9EE" blockId="11.[113,1090,1530,1602]" pageId="11" pageNumber="120">
Fig.4. Details of the head and pronotum of the
<taxonomicName id="78D34D0A8D72285DFE125EB771BFF9A1" authority="" authorityName="Zaitzev" authorityYear="1908" box="[498,727,1530,1549]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="subGenus" subGenus="Lihelophorus">
<emphasis id="8DA7EA9B8D72285DFE125EB7713DF9A1" box="[498,597,1530,1549]" italics="true" pageId="11" pageNumber="120">Helophorus</emphasis>
(
<emphasis id="8DA7EA9B8D72285DFD805EB771B8F9A1" box="[608,720,1530,1549]" italics="true" pageId="11" pageNumber="120">Lihelophorus</emphasis>
)
</taxonomicName>
specimens.ac
<taxonomicName id="78D34D0A8D72285DFC8A5EB77728F9A1" authority="Zaitzev, 1908" authorityName="Zaitzev" authorityYear="1908" box="[874,1088,1530,1549]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="lamicola">
<emphasis id="8DA7EA9B8D72285DFC8A5EB770A5F9A1" box="[874,973,1530,1549]" italics="true" pageId="11" pageNumber="120">H. lamicola</emphasis>
<bibRefCitation id="DB424B788D72285DFC305EB77728F9A1" author="ZAITZEV F. A." box="[976,1088,1530,1549]" pageId="11" pageNumber="120" pagination="417 - 426" refId="ref20914" refString="ZAITZEV F. A. 1908: Beitrag zur Kenntniss der Waserkafer von Chinesisch-Centralasien. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St. Petersbourg 13: 417 - 426." type="journal article" year="1908">Zaitzev,1908</bibRefCitation>
</taxonomicName>
(a Qinghai: Zhaling Hu (lectotype); bc Qinghai: Golo). dh
<taxonomicName id="78D34D0A8D72285DFD435D597027F98B" authority="Zaitzev, 1908" authorityName="Zaitzev" authorityYear="1908" box="[675,847,1556,1575]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="120" phylum="Arthropoda" rank="species" species="ser">
<emphasis id="8DA7EA9B8D72285DFD435D5971BFF98B" box="[675,727,1556,1575]" italics="true" pageId="11" pageNumber="120">H. ser</emphasis>
<bibRefCitation id="DB424B788D72285DFD3B5D597027F98B" author="ZAITZEV F. A." box="[731,847,1556,1575]" pageId="11" pageNumber="120" pagination="417 - 426" refId="ref20914" refString="ZAITZEV F. A. 1908: Beitrag zur Kenntniss der Waserkafer von Chinesisch-Centralasien. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St. Petersbourg 13: 417 - 426." type="journal article" year="1908">Zaitzev, 1908</bibRefCitation>
</taxonomicName>
(d Qinghai: valley of Alag Hu lake (lectotype); ef Qinghai: Golo; g Xizang, Nyainqentanglha Feng; h Gansu: Xiahe env.). Not to scale.
</paragraph>
</caption>
<caption id="EBAC66018D75285AFF775F6F770EFBFC" ID-DOI="http://doi.org/10.5281/zenodo.5311796" ID-Zenodo-Dep="5311796" httpUri="https://zenodo.org/record/5311796/files/figure.png" pageId="12" pageNumber="121" startId="12.[151,182,1058,1077]" targetBox="[164,1118,238,1008]" targetPageId="12">
<paragraph id="BF6C36898D75285AFF775F6F770EFBFC" blockId="12.[151,1127,1058,1104]" pageId="12" pageNumber="121">
Fig. 5. Details of the head and pronotum of the
<taxonomicName id="78D34D0A8D75285AFDCE5F6F7034FB99" authority="Angus &amp; Jia &amp; Chen &amp; Zhang &amp; Vondráček &amp; Fikáček &amp; Sw, 2016" authorityName="Angus &amp; Jia &amp; Chen &amp; Zhang &amp; Vondráček &amp; Fikáček &amp; Sw" authorityYear="2016" box="[558,860,1058,1077]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="121" phylum="Arthropoda" rank="species" species="yangae" status="sp. nov." subGenus="Lihelophorus">
<emphasis id="8DA7EA9B8D75285AFDCE5F6F71FBFB99" box="[558,659,1058,1077]" italics="true" pageId="12" pageNumber="121">Helophorus</emphasis>
(
<emphasis id="8DA7EA9B8D75285AFD7F5F6F7079FB99" box="[671,785,1058,1077]" italics="true" pageId="12" pageNumber="121">Lihelophorus</emphasis>
)
<emphasis id="8DA7EA9B8D75285AFCFE5F6F7034FB99" box="[798,860,1058,1077]" italics="true" pageId="12" pageNumber="121">yangae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="969457E08D75285AFC815F6F70CCFB99" box="[865,932,1058,1077]" pageId="12" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
(a Xizang: 90 km W of Amdo; b,d Qinghai: Golo; c Xizang: 25 km W of Amdo; e India: Kar Tso (Orchymont coll.). Not to scale.
</paragraph>
</caption>
<subSubSection id="F7C965028D75285BFF775E27700CFEBF" lastPageId="13" lastPageNumber="122" pageId="12" pageNumber="121" type="diagnosis">
<paragraph id="BF6C36898D75285BFF775E27700CFEBF" blockId="12.[151,1127,1386,1601]" lastBlockId="13.[113,1090,220,1587]" lastPageId="13" lastPageNumber="122" pageId="12" pageNumber="121">
<emphasis id="8DA7EA9B8D75285AFF775E2772F9FA2D" bold="true" box="[151,401,1386,1409]" pageId="12" pageNumber="121">Differential diagnosis.</emphasis>
Characterized by moderately wide pronotum with distinctly sinuate sides abruptly narrowed just before anterior corners, with five longitudinal grooves distinctly impressed (
<figureCitation id="27E82A0C8D75285AFEED5EE7721EFA6D" box="[269,374,1450,1473]" captionStart="Fig" captionStartId="7.[113,144,1463,1482]" captionTargetBox="[133,1067,220,1453]" captionTargetId="figure-13@7.[133,1068,219,1454]" captionTargetPageId="7" captionText="Fig. 2. Adult morphology of Helophorus (Lihelophorus). af details of left half of pronotum (a,c,e) and base of left elytron (b, d, f): ab H. yangae sp. nov.; cd H. ser Zaitzev, 1908; ef H. lamicola Zaitzev, 1908; arrow in (a) shows remnants of pronotal granulations, arrows in (b, d, f) show scutellary stria. gh prothoracic hypomeron and suprapleural area (g H. yangae sp. nov.; h H. ser). i detail of posterior margin of abdominal ventrite 5 of H. ser, showing the irregular denticulation. j ventral view of the thorax of H. ser. k dorsolateral portion of left elytron of H. lamicola. Abbreviations: ep epipleuron; li last elytral interval. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311790" httpUri="https://zenodo.org/record/5311790/files/figure.png" pageId="12" pageNumber="121">Figs 2cd</figureCitation>
,
<figureCitation id="27E82A0C8D75285AFE9F5EE772DCFA6D" box="[383,436,1450,1473]" captionStart="Fig" captionStartId="10.[151,182,1482,1501]" captionTargetBox="[159,1124,234,1349]" captionTargetId="figure-10@10.[151,1127,219,1365]" captionTargetPageId="10" captionText="Fig. 3. Habitus of the species of Helophorus (Lihelophorus). ac H. lamicola Zaitzev, 1908 (a Qinghai: Zhaling Hu (lectotype); bc Quinghai: Golo).dg H.ser Zaitzev, 1908 (de Qinghai:Golo; f Xizang, Nyainqentanglha Feng; g Gansu: Xiahe env.). hk H. yangae sp. nov. (h Xizang: 90 km W of Amdo; i Xizang: 25 km W of Amdo; j Qinghai: Golo; k India: Kar Tso (Orchymont coll.)). All specimens to scale." figureDoi="http://doi.org/10.5281/zenodo.5311792" httpUri="https://zenodo.org/record/5311792/files/figure.png" pageId="12" pageNumber="121">3ag</figureCitation>
,
<figureCitation id="27E82A0C8D75285AFE5D5EE7729DFA6D" box="[445,501,1450,1473]" captionStart="Fig" captionStartId="11.[113,144,1530,1549]" captionTargetBox="[131,1080,229,1416]" captionTargetId="figure-13@11.[113,1091,219,1456]" captionTargetPageId="11" captionText="Fig.4. Details of the head and pronotum of the Helophorus (Lihelophorus) specimens.ac H. lamicola Zaitzev,1908 (a Qinghai: Zhaling Hu (lectotype); bc Qinghai: Golo). dh H. ser Zaitzev, 1908 (d Qinghai: valley of Alag Hu lake (lectotype); ef Qinghai: Golo; g Xizang, Nyainqentanglha Feng; h Gansu: Xiahe env.). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311794" httpUri="https://zenodo.org/record/5311794/files/figure.png" pageId="12" pageNumber="121">4dh</figureCitation>
); in this it is more similar to
<taxonomicName id="78D34D0A8D75285AFCF35EE77016FA6D" authority="Angus &amp; Jia &amp; Chen &amp; Zhang &amp; Vondráček &amp; Fikáček &amp; Sw, 2016" authorityName="Angus &amp; Jia &amp; Chen &amp; Zhang &amp; Vondráček &amp; Fikáček &amp; Sw" authorityYear="2016" box="[787,894,1450,1473]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="121" phylum="Arthropoda" rank="species" species="yangae" status="sp. nov.">
<emphasis id="8DA7EA9B8D75285AFCF35EE77016FA6D" box="[787,894,1450,1473]" italics="true" pageId="12" pageNumber="121">H. yangae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="969457E08D75285AFC625EE770B9FA6D" box="[898,977,1450,1473]" pageId="12" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
(which differs by aedeagus with rounded parameral apices) than to
<taxonomicName id="78D34D0A8D75285AFD5D5E877051FA4D" authorityName="Zaitzev" authorityYear="1908" box="[701,825,1482,1505]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="121" phylum="Arthropoda" rank="species" species="lamicola">
<emphasis id="8DA7EA9B8D75285AFD5D5E877051FA4D" box="[701,825,1482,1505]" italics="true" pageId="12" pageNumber="121">H. lamicola</emphasis>
</taxonomicName>
. Elytral series, including the scutellary stria, are always very distinct (in contrast to
<taxonomicName id="78D34D0A8D75285AFD085EA7700FF9AD" authorityName="Zaitzev" authorityYear="1908" box="[744,871,1514,1537]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="121" phylum="Arthropoda" rank="species" species="lamicola">
<emphasis id="8DA7EA9B8D75285AFD085EA7700FF9AD" box="[744,871,1514,1537]" italics="true" pageId="12" pageNumber="121">H. lamicola</emphasis>
</taxonomicName>
). Meso- and metatarsal claws are long (as in
<taxonomicName id="78D34D0A8D75285AFE955D47729AF98D" authorityName="Zaitzev" authorityYear="1908" box="[373,498,1546,1569]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="121" phylum="Arthropoda" rank="species" species="lamicola">
<emphasis id="8DA7EA9B8D75285AFE955D47729AF98D" box="[373,498,1546,1569]" italics="true" pageId="12" pageNumber="121">H. lamicola</emphasis>
</taxonomicName>
, but unlike
<taxonomicName id="78D34D0A8D75285AFD8F5D4771B4F98D" authority="Angus &amp; Jia &amp; Chen &amp; Zhang &amp; Vondráček &amp; Fikáček &amp; Sw, 2016" authorityName="Angus &amp; Jia &amp; Chen &amp; Zhang &amp; Vondráček &amp; Fikáček &amp; Sw" authorityYear="2016" box="[623,732,1546,1569]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="121" phylum="Arthropoda" rank="species" species="yangae" status="sp. nov.">
<emphasis id="8DA7EA9B8D75285AFD8F5D4771B4F98D" box="[623,732,1546,1569]" italics="true" pageId="12" pageNumber="121">H. yangae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="969457E08D75285AFD025D477050F98D" box="[738,824,1546,1569]" pageId="12" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
). Metatarsomere 5 is usually longer (often much longer) than metatarsomeres 34 combined. The aedeagus is very similar to
<taxonomicName id="78D34D0A8D74285BFF6D5B917263FF5F" authorityName="Zaitzev" authorityYear="1908" box="[141,267,220,243]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="122" phylum="Arthropoda" rank="species" species="lamicola">
<emphasis id="8DA7EA9B8D74285BFF6D5B917263FF5F" box="[141,267,220,243]" italics="true" pageId="13" pageNumber="122">H. lamicola</emphasis>
</taxonomicName>
(based on pointed parameral apices) but the median lobe in narrower and its apical portion longer (
<figureCitation id="27E82A0C8D74285BFEBE5BB172A2FEBF" box="[350,458,252,275]" captionStart="Fig" captionStartId="15.[113,144,1496,1515]" captionTargetBox="[149,1030,229,1439]" captionTargetId="figure-13@15.[139,1063,219,1474]" captionTargetPageId="15" captionText="Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. ae H. (L.) ser Zaitzev, 1908 (a lectotype, Qinghai: valley of Alag Hu lake; bc Xizang: valley SE of Dong La pass; d Qinghai: Golo, 20 km E of Maduo; e Qinghai: Zuimatan). fj H. (L.) lamicola Zaitzev, 1908 (f lectotype, Qinghai: Djarin Nor; gi Qinghai: Zuimatan; j paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). ko H. (L.) yangae sp. nov. (k holotype, Qinghai: Zuimatan; lm paratypes, same locality; n Qinghai: Golo, Yematan; o Xizang: Gongbujiangda)." figureDoi="http://doi.org/10.5281/zenodo.5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="13" pageNumber="122">Figs 6ae</figureCitation>
). For detailed comparison see
<tableCitation id="F25103328D74285BFCEE5BB17008FEBF" box="[782,864,252,275]" captionStart="Table 2" captionStartId="16.[153,200,220,239]" captionText="Table 2. Comparison of adult morphology of Lihelophorus species." pageId="13" pageNumber="122">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F7C965028D74285BFF915A517037F99F" pageId="13" pageNumber="122" type="description">
<paragraph id="BF6C36898D74285BFF915A5170F1FE9F" blockId="13.[113,1090,220,1587]" box="[113,921,284,307]" pageId="13" pageNumber="122">
<emphasis id="8DA7EA9B8D74285BFF915A51727FFE9F" bold="true" box="[113,279,284,307]" pageId="13" pageNumber="122">Redescription.</emphasis>
Body length:
<quantity id="782B9B6C8D74285BFE4E5A517146FE9F" box="[430,558,284,307]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="5.7" metricValueMin="4.4" pageId="13" pageNumber="122" unit="mm" value="5.05" valueMax="5.7" valueMin="4.4">4.45.7 mm</quantity>
in males,
<quantity id="782B9B6C8D74285BFD7C5A517074FE9F" box="[668,796,284,307]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.1" metricValueMin="4.9" pageId="13" pageNumber="122" unit="mm" value="5.5" valueMax="6.1" valueMin="4.9">4.96.1 mm</quantity>
in females.
</paragraph>
<paragraph id="BF6C36898D74285BFF705A7170D5FE5F" blockId="13.[113,1090,220,1587]" pageId="13" pageNumber="122">
<emphasis id="8DA7EA9B8D74285BFF705A7173B9FEFF" box="[144,209,316,339]" italics="true" pageId="13" pageNumber="122">Head.</emphasis>
Dull bronze with green to purple reflections to completely black, closely and irregularly punctate, giving a rough effect in some portions, punctation consisting of two sizes of punctures. Y-groove shallow to moderately deep, with rather distinct stem wider than lateral arms, widened anteriorly. Maxillary palpi pale yellow, elongate, apical segment asymmetrical. Antennae yellow, 9-segmented. Maxillary galea in shape of simple pubescent lobe, without strong spines (
<figureCitation id="27E82A0C8D74285BFEEC5A917233FE5F" box="[268,347,476,499]" captionStart="Fig" captionStartId="6.[151,182,1490,1509]" captionTargetBox="[152,1126,220,1478]" captionTargetId="figure-10@6.[151,1127,217,1486]" captionTargetPageId="6" captionText="Fig. 1. Adult morphology of Helophorus (Lihelophorus). ab H. yangae sp. nov. (a ventral view of the head; b detail of mentum and labial palps); cd maxillary galea and lacinia (c H. yangae sp. nov.; d H. ser Zaitzev, 1908); ef mandible (e H. yangae sp. nov.; f H. ser); g labial palpus of H. ser; hl hind legs in dorsal view (h H. yangae sp. nov.; i H. ser; j H. lamicola Zaitzev, 1908; k detail of apex of tibia and base of tarsus of H. lamicola; l detail of tarsomere 5 and claw of H. ser). Abbreviations: sh swimming hairs. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311786" httpUri="https://zenodo.org/record/5311786/files/figure.png" pageId="13" pageNumber="122">Fig. 1d</figureCitation>
). Mandible rather short, with simple blunt apex (
<figureCitation id="27E82A0C8D74285BFC855A9170D8FE5F" box="[869,944,476,499]" captionStart="Fig" captionStartId="6.[151,182,1490,1509]" captionTargetBox="[152,1126,220,1478]" captionTargetId="figure-10@6.[151,1127,217,1486]" captionTargetPageId="6" captionText="Fig. 1. Adult morphology of Helophorus (Lihelophorus). ab H. yangae sp. nov. (a ventral view of the head; b detail of mentum and labial palps); cd maxillary galea and lacinia (c H. yangae sp. nov.; d H. ser Zaitzev, 1908); ef mandible (e H. yangae sp. nov.; f H. ser); g labial palpus of H. ser; hl hind legs in dorsal view (h H. yangae sp. nov.; i H. ser; j H. lamicola Zaitzev, 1908; k detail of apex of tibia and base of tarsus of H. lamicola; l detail of tarsomere 5 and claw of H. ser). Abbreviations: sh swimming hairs. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311786" httpUri="https://zenodo.org/record/5311786/files/figure.png" pageId="13" pageNumber="122">Fig. 1f</figureCitation>
).
</paragraph>
<paragraph id="BF6C36898D74285BFF705AB170E1FD1F" blockId="13.[113,1090,220,1587]" pageId="13" pageNumber="122">
<emphasis id="8DA7EA9B8D74285BFF705AB1739FFDBF" box="[144,247,508,531]" italics="true" pageId="13" pageNumber="122">Pronotum</emphasis>
. Yellow, wider than the head, moderately and evenly arched. Widest closely before anterior corners; sides strongly arcuate, abruptly narrowed anteriorly, straight to indistinctly sinuate posteriorly. Grooves distinct, usually deeply and sharply impressed; mid groove straight, narrow; submedian grooves slightly sinuate; submarginal grooves with their basal pits distinct, bent mesally in anterior fourth. Narrow raised lateral margin distinct. Intervals without granules but with large punctures and smaller ones between them.
</paragraph>
<paragraph id="BF6C36898D74285BFF7059F17279FC9F" blockId="13.[113,1090,220,1587]" pageId="13" pageNumber="122">
<emphasis id="8DA7EA9B8D74285BFF7059F173BAFD7F" box="[144,210,700,723]" italics="true" pageId="13" pageNumber="122">Elytra</emphasis>
. Yellow as pronotum, with or without apparent darker sutural Λ- mark and sublateral spots. Striae distinct, consisting of rather large punctures, scutellary stria always distinct. Interval punctures much smaller than the striae, with fine erect hairs. All intervals flat, including the 10
<superScript id="48A69BC18D74285BFF1E58567262FC85" attach="none" box="[254,266,795,809]" fontSize="6" pageId="13" pageNumber="122">th</superScript>
.
</paragraph>
<paragraph id="BF6C36898D74285BFF705871717BFC3F" blockId="13.[113,1090,220,1587]" pageId="13" pageNumber="122">
<emphasis id="8DA7EA9B8D74285BFF70587173AAFCFF" box="[144,194,828,851]" italics="true" pageId="13" pageNumber="122">Legs</emphasis>
. Long, with long swimming-hairs on meso- and metatibiae and meso- and metatarsi; metatarsomere 5 distinctly longer than metatarsomeres 34 combined; metatarsal claw ca. 0.5× as long as utimate metatarsomere.
</paragraph>
<paragraph id="BF6C36898D74285BFF7058D17178FBBF" blockId="13.[113,1090,220,1587]" pageId="13" pageNumber="122">
<emphasis id="8DA7EA9B8D74285BFF7058D1739FFC1F" box="[144,247,924,947]" italics="true" pageId="13" pageNumber="122">Aedeagus</emphasis>
. Parameres pointed, very weakly to strongly sinuate on outer margin. Phallobase ca. as long as or slightly shorther than parameres. Median lobe narrow, its distal portion long (in shape of more less equilateral triangle); base of median lobe (between the struts) arcuate, with median backwardly directed spur.
</paragraph>
<paragraph id="BF6C36898D74285BFF915F517037F99F" blockId="13.[113,1090,220,1587]" pageId="13" pageNumber="122">
<emphasis id="8DA7EA9B8D74285BFF915F51738EFB9F" bold="true" box="[113,230,1052,1075]" pageId="13" pageNumber="122">Variation.</emphasis>
The species varies considerably in size and body proportions; both smaller narrower (
<figureCitation id="27E82A0C8D74285BFF205F717266FBFF" box="[192,270,1084,1107]" captionStart="Fig" captionStartId="10.[151,182,1482,1501]" captionTargetBox="[159,1124,234,1349]" captionTargetId="figure-10@10.[151,1127,219,1365]" captionTargetPageId="10" captionText="Fig. 3. Habitus of the species of Helophorus (Lihelophorus). ac H. lamicola Zaitzev, 1908 (a Qinghai: Zhaling Hu (lectotype); bc Quinghai: Golo).dg H.ser Zaitzev, 1908 (de Qinghai:Golo; f Xizang, Nyainqentanglha Feng; g Gansu: Xiahe env.). hk H. yangae sp. nov. (h Xizang: 90 km W of Amdo; i Xizang: 25 km W of Amdo; j Qinghai: Golo; k India: Kar Tso (Orchymont coll.)). All specimens to scale." figureDoi="http://doi.org/10.5281/zenodo.5311792" httpUri="https://zenodo.org/record/5311792/files/figure.png" pageId="13" pageNumber="122">Fig. 3e</figureCitation>
) and larger wide (
<figureCitation id="27E82A0C8D74285BFE375F71712EFBFF" box="[471,582,1084,1107]" captionStart="Fig" captionStartId="10.[151,182,1482,1501]" captionTargetBox="[159,1124,234,1349]" captionTargetId="figure-10@10.[151,1127,219,1365]" captionTargetPageId="10" captionText="Fig. 3. Habitus of the species of Helophorus (Lihelophorus). ac H. lamicola Zaitzev, 1908 (a Qinghai: Zhaling Hu (lectotype); bc Quinghai: Golo).dg H.ser Zaitzev, 1908 (de Qinghai:Golo; f Xizang, Nyainqentanglha Feng; g Gansu: Xiahe env.). hk H. yangae sp. nov. (h Xizang: 90 km W of Amdo; i Xizang: 25 km W of Amdo; j Qinghai: Golo; k India: Kar Tso (Orchymont coll.)). All specimens to scale." figureDoi="http://doi.org/10.5281/zenodo.5311792" httpUri="https://zenodo.org/record/5311792/files/figure.png" pageId="13" pageNumber="122">Figs 3d, g</figureCitation>
) specimens are known. Coloration also varies slightly, especially as it concerns the colouration of the head (
<figureCitation id="27E82A0C8D74285BFCF55F1170EDFBDF" box="[789,901,1116,1139]" captionStart="Fig" captionStartId="11.[113,144,1530,1549]" captionTargetBox="[131,1080,229,1416]" captionTargetId="figure-13@11.[113,1091,219,1456]" captionTargetPageId="11" captionText="Fig.4. Details of the head and pronotum of the Helophorus (Lihelophorus) specimens.ac H. lamicola Zaitzev,1908 (a Qinghai: Zhaling Hu (lectotype); bc Qinghai: Golo). dh H. ser Zaitzev, 1908 (d Qinghai: valley of Alag Hu lake (lectotype); ef Qinghai: Golo; g Xizang, Nyainqentanglha Feng; h Gansu: Xiahe env.). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311794" httpUri="https://zenodo.org/record/5311794/files/figure.png" pageId="13" pageNumber="122">Figs 4dh</figureCitation>
) and the absence / presence of the dark patterns on elytra; elytra of the specimens from high altitudes of southern
<collectingRegion id="7D17F86B8D74285BFF355FD1724BFB1F" box="[213,291,1180,1203]" country="China" name="Xizang" pageId="13" pageNumber="122">Xizang</collectingRegion>
(collected by J. Schmidt) are often slightly darker than in other examined specimens (
<figureCitation id="27E82A0C8D74285BFF135FF1722AFB7F" box="[243,322,1212,1235]" captionStart="Fig" captionStartId="10.[151,182,1482,1501]" captionTargetBox="[159,1124,234,1349]" captionTargetId="figure-10@10.[151,1127,219,1365]" captionTargetPageId="10" captionText="Fig. 3. Habitus of the species of Helophorus (Lihelophorus). ac H. lamicola Zaitzev, 1908 (a Qinghai: Zhaling Hu (lectotype); bc Quinghai: Golo).dg H.ser Zaitzev, 1908 (de Qinghai:Golo; f Xizang, Nyainqentanglha Feng; g Gansu: Xiahe env.). hk H. yangae sp. nov. (h Xizang: 90 km W of Amdo; i Xizang: 25 km W of Amdo; j Qinghai: Golo; k India: Kar Tso (Orchymont coll.)). All specimens to scale." figureDoi="http://doi.org/10.5281/zenodo.5311792" httpUri="https://zenodo.org/record/5311792/files/figure.png" pageId="13" pageNumber="122">Fig. 3f</figureCitation>
). The pronotum varies slightly in the shape of the sides as well as in the depth and shape of the pronotal grooves (
<figureCitation id="27E82A0C8D74285BFD815F9171BBFB5F" box="[609,723,1244,1267]" captionStart="Fig" captionStartId="11.[113,144,1530,1549]" captionTargetBox="[131,1080,229,1416]" captionTargetId="figure-13@11.[113,1091,219,1456]" captionTargetPageId="11" captionText="Fig.4. Details of the head and pronotum of the Helophorus (Lihelophorus) specimens.ac H. lamicola Zaitzev,1908 (a Qinghai: Zhaling Hu (lectotype); bc Qinghai: Golo). dh H. ser Zaitzev, 1908 (d Qinghai: valley of Alag Hu lake (lectotype); ef Qinghai: Golo; g Xizang, Nyainqentanglha Feng; h Gansu: Xiahe env.). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311794" httpUri="https://zenodo.org/record/5311794/files/figure.png" pageId="13" pageNumber="122">Figs 4dh</figureCitation>
), with rare specimens with partly reduced shallow grooves (
<figureCitation id="27E82A0C8D74285BFE705FB172B5FABF" box="[400,477,1276,1299]" captionStart="Fig" captionStartId="11.[113,144,1530,1549]" captionTargetBox="[131,1080,229,1416]" captionTargetId="figure-13@11.[113,1091,219,1456]" captionTargetPageId="11" captionText="Fig.4. Details of the head and pronotum of the Helophorus (Lihelophorus) specimens.ac H. lamicola Zaitzev,1908 (a Qinghai: Zhaling Hu (lectotype); bc Qinghai: Golo). dh H. ser Zaitzev, 1908 (d Qinghai: valley of Alag Hu lake (lectotype); ef Qinghai: Golo; g Xizang, Nyainqentanglha Feng; h Gansu: Xiahe env.). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311794" httpUri="https://zenodo.org/record/5311794/files/figure.png" pageId="13" pageNumber="122">Fig. 4e</figureCitation>
) which may resemble
<taxonomicName id="78D34D0A8D74285BFD355FB1703EFABF" authorityName="Zaitzev" authorityYear="1908" box="[725,854,1276,1299]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="122" phylum="Arthropoda" rank="species" species="lamicola">
<emphasis id="8DA7EA9B8D74285BFD355FB1703EFABF" box="[725,854,1276,1299]" italics="true" pageId="13" pageNumber="122">H. lamicola</emphasis>
</taxonomicName>
. The aedeagus varies considerably in the shape of the apical portions of parameres: the variation is the same as in
<taxonomicName id="78D34D0A8D74285BFF4D5E717247FAFF" authorityName="Zaitzev" authorityYear="1908" box="[173,303,1340,1363]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="122" phylum="Arthropoda" rank="species" species="lamicola">
<emphasis id="8DA7EA9B8D74285BFF4D5E717247FAFF" box="[173,303,1340,1363]" italics="true" pageId="13" pageNumber="122">H. lamicola</emphasis>
</taxonomicName>
, see there for details. In addition to that, we observed the considerable variation in the length of the aedeagus (compare
<figureCitation id="27E82A0C8D74285BFD605E11703AFADF" box="[640,850,1372,1395]" captionStart="Fig" captionStartId="15.[113,144,1496,1515]" captionTargetBox="[149,1030,229,1439]" captionTargetId="figure-13@15.[139,1063,219,1474]" captionTargetPageId="15" captionText="Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. ae H. (L.) ser Zaitzev, 1908 (a lectotype, Qinghai: valley of Alag Hu lake; bc Xizang: valley SE of Dong La pass; d Qinghai: Golo, 20 km E of Maduo; e Qinghai: Zuimatan). fj H. (L.) lamicola Zaitzev, 1908 (f lectotype, Qinghai: Djarin Nor; gi Qinghai: Zuimatan; j paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). ko H. (L.) yangae sp. nov. (k holotype, Qinghai: Zuimatan; lm paratypes, same locality; n Qinghai: Golo, Yematan; o Xizang: Gongbujiangda)." figureDoi="http://doi.org/10.5281/zenodo.5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="13" pageNumber="122">Figs 6ac and 6de</figureCitation>
) which was not found in
<taxonomicName id="78D34D0A8D74285BFF705E31727BFA3F" authorityName="Zaitzev" authorityYear="1908" box="[144,275,1404,1427]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="122" phylum="Arthropoda" rank="species" species="lamicola">
<emphasis id="8DA7EA9B8D74285BFF705E31727BFA3F" box="[144,275,1404,1427]" italics="true" pageId="13" pageNumber="122">H. lamicola</emphasis>
</taxonomicName>
. Examined specimens with shorter aedeagus were mostly from southern
<collectingRegion id="7D17F86B8D74285BFF915ED173D7FA1F" box="[113,191,1436,1459]" country="China" name="Xizang" pageId="13" pageNumber="122">Xizang</collectingRegion>
(specimens collected by J. Schmidt,
<figureCitation id="27E82A0C8D74285BFD885ED171B2FA1F" box="[616,730,1436,1459]" captionStart="Fig" captionStartId="15.[113,144,1496,1515]" captionTargetBox="[149,1030,229,1439]" captionTargetId="figure-13@15.[139,1063,219,1474]" captionTargetPageId="15" captionText="Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. ae H. (L.) ser Zaitzev, 1908 (a lectotype, Qinghai: valley of Alag Hu lake; bc Xizang: valley SE of Dong La pass; d Qinghai: Golo, 20 km E of Maduo; e Qinghai: Zuimatan). fj H. (L.) lamicola Zaitzev, 1908 (f lectotype, Qinghai: Djarin Nor; gi Qinghai: Zuimatan; j paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). ko H. (L.) yangae sp. nov. (k holotype, Qinghai: Zuimatan; lm paratypes, same locality; n Qinghai: Golo, Yematan; o Xizang: Gongbujiangda)." figureDoi="http://doi.org/10.5281/zenodo.5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="13" pageNumber="122">Figs 6bc</figureCitation>
), but the
<typeStatus id="6068882B8D74285BFCA95ED170C5FA1F" box="[841,941,1436,1459]" pageId="13" pageNumber="122" type="lectotype">lectotype</typeStatus>
from central
<collectingRegion id="7D17F86B8D74285BFF915EF173A1FA7F" box="[113,201,1468,1491]" country="China" name="Qinghai" pageId="13" pageNumber="122">Qinghai</collectingRegion>
also has this kind of the aedeagus; on the other hand, most specimens examined from
<collectingRegion id="7D17F86B8D74285BFF4D5E91726DFA5F" box="[173,261,1500,1523]" country="China" name="Qinghai" pageId="13" pageNumber="122">Qinghai</collectingRegion>
and
<collectingRegion id="7D17F86B8D74285BFED95E917216FA5F" box="[313,382,1500,1523]" country="China" name="Gansu" pageId="13" pageNumber="122">Gansu</collectingRegion>
have longer aedeagus (
<figureCitation id="27E82A0C8D74285BFD9D5E917186FA5F" box="[637,750,1500,1523]" captionStart="Fig" captionStartId="15.[113,144,1496,1515]" captionTargetBox="[149,1030,229,1439]" captionTargetId="figure-13@15.[139,1063,219,1474]" captionTargetPageId="15" captionText="Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. ae H. (L.) ser Zaitzev, 1908 (a lectotype, Qinghai: valley of Alag Hu lake; bc Xizang: valley SE of Dong La pass; d Qinghai: Golo, 20 km E of Maduo; e Qinghai: Zuimatan). fj H. (L.) lamicola Zaitzev, 1908 (f lectotype, Qinghai: Djarin Nor; gi Qinghai: Zuimatan; j paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). ko H. (L.) yangae sp. nov. (k holotype, Qinghai: Zuimatan; lm paratypes, same locality; n Qinghai: Golo, Yematan; o Xizang: Gongbujiangda)." figureDoi="http://doi.org/10.5281/zenodo.5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="13" pageNumber="122">Figs 6de</figureCitation>
). The apparent paler coloration of the aedeagus of the
<typeStatus id="6068882B8D74285BFE8E5EB172BAF9BF" box="[366,466,1532,1555]" pageId="13" pageNumber="122" type="lectotype">lectotype</typeStatus>
and specimens collected by J. Schmidt (
<figureCitation id="27E82A0C8D74285BFC755EB1776AF9BF" box="[917,1026,1532,1555]" captionStart="Fig" captionStartId="15.[113,144,1496,1515]" captionTargetBox="[149,1030,229,1439]" captionTargetId="figure-13@15.[139,1063,219,1474]" captionTargetPageId="15" captionText="Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. ae H. (L.) ser Zaitzev, 1908 (a lectotype, Qinghai: valley of Alag Hu lake; bc Xizang: valley SE of Dong La pass; d Qinghai: Golo, 20 km E of Maduo; e Qinghai: Zuimatan). fj H. (L.) lamicola Zaitzev, 1908 (f lectotype, Qinghai: Djarin Nor; gi Qinghai: Zuimatan; j paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). ko H. (L.) yangae sp. nov. (k holotype, Qinghai: Zuimatan; lm paratypes, same locality; n Qinghai: Golo, Yematan; o Xizang: Gongbujiangda)." figureDoi="http://doi.org/10.5281/zenodo.5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="13" pageNumber="122">Figs 6ac</figureCitation>
) may be an artefact of long-term storage of the specimens in fixation fluid.
</paragraph>
</subSubSection>
<subSubSection id="F7C965028D772858FF775B91705EFE41" pageId="14" pageNumber="123" type="distribution">
<paragraph id="BF6C36898D772858FF775B91702CFE3F" blockId="14.[151,1128,220,403]" pageId="14" pageNumber="123">
<emphasis id="8DA7EA9B8D772858FF775B91724AFF5F" bold="true" box="[151,290,220,243]" pageId="14" pageNumber="123">Distribution</emphasis>
(Fig. 7a). Very widely distributed on the Tibetan Plateau, from Indian province Jammu and Kashmir to southern
<collectingRegion id="7D17F86B8D772858FE165BB17152FEBF" box="[502,570,252,275]" country="China" name="Gansu" pageId="14" pageNumber="123">Gansu</collectingRegion>
in
<collectingCountry id="C7C476198D772858FDBC5BB171F7FEBF" box="[604,671,252,275]" name="China" pageId="14" pageNumber="123">China</collectingCountry>
. It occurs at altitudes ranging from
<quantity id="782B9B6C8D772858FBF65BB1770FFEBF" box="[1046,1127,252,275]" metricMagnitude="3" metricUnit="m" metricValue="2.94" pageId="14" pageNumber="123" unit="m" value="2940.0">2940 m</quantity>
(
<collectingRegion id="7D17F86B8D772858FF7F5A517380FE9F" box="[159,232,284,307]" country="China" name="Gansu" pageId="14" pageNumber="123">Gansu</collectingRegion>
: Labrang) to ca.
<quantity id="782B9B6C8D772858FE755A51728CFE9F" box="[405,484,284,307]" metricMagnitude="3" metricUnit="m" metricValue="5.35" pageId="14" pageNumber="123" unit="m" value="5350.0">5350 m</quantity>
(
<collectingRegion id="7D17F86B8D772858FE115A51712BFE9F" box="[497,579,284,307]" country="China" name="Xizang" pageId="14" pageNumber="123">Xizang</collectingRegion>
: Nyainqentanglha Feng). As pointed out by Joachim Schmidt (pers. comm. to R. B. Angus) this makes it one of the highest-living of all
<taxonomicName id="78D34D0A8D772858FC0C5A71770CFEFF" box="[1004,1124,316,339]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="14" pageNumber="123" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
.
<emphasis id="8DA7EA9B8D772858FF775A1173BBFEDF" bold="true" box="[151,211,348,371]" pageId="14" pageNumber="123">Note.</emphasis>
The record of
<taxonomicName id="78D34D0A8D772858FE8E5A1172C7FEDF" authorityName="Zaitzev" authorityYear="1908" box="[366,431,348,371]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="123" phylum="Arthropoda" rank="species" species="ser">
<emphasis id="8DA7EA9B8D772858FE8E5A1172C7FEDF" box="[366,431,348,371]" italics="true" pageId="14" pageNumber="123">H. ser</emphasis>
</taxonomicName>
from Indian
<collectingRegion id="7D17F86B8D772858FDD75A117106FEDF" box="[567,622,348,371]" country="China" name="Xizang" pageId="14" pageNumber="123">Tibet</collectingRegion>
by
<bibRefCitation id="DB424B788D772858FD755A117008FEDF" author="ORCHYMONT A." box="[661,864,348,371]" pageId="14" pageNumber="123" pagination="1 - 16" refId="ref20836" refString="ORCHYMONT A. d' 1943: Les Palpicornia du Tibet (Coleoptera). Bulletin du Musee Royal d'Histoire Naturelle de Belgique 19 (57): 1 - 16." type="journal article" year="1943">ORCHYMONT (1943)</bibRefCitation>
was based on a misidentified specimen of
<taxonomicName id="78D34D0A8D772858FEBD5A3172A3FE3F" authorityName="Angus &amp; Jia &amp; Chen &amp; Zhang &amp; Vondráček &amp; Fikáček &amp; Sw" authorityYear="2016" box="[349,459,380,403]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="123" phylum="Arthropoda" rank="species" species="yangae">
<emphasis id="8DA7EA9B8D772858FEBD5A3172A3FE3F" box="[349,459,380,403]" italics="true" pageId="14" pageNumber="123">H. yangae</emphasis>
</taxonomicName>
(see under that species for details).
</paragraph>
<paragraph id="BF6C36898D772858FEED5AF1705EFE41" blockId="14.[269,1009,444,493]" pageId="14" pageNumber="123">
<emphasis id="8DA7EA9B8D772858FEED5AF1704AFE7F" bold="true" box="[269,802,444,467]" pageId="14" pageNumber="123">
<taxonomicName id="78D34D0A8D772858FEED5AF171FAFE7F" authorityName="Angus &amp; Jia &amp; Chen &amp; Zhang &amp; Vondráček &amp; Fikáček &amp; Sw" authorityYear="2016" box="[269,658,444,467]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="123" phylum="Arthropoda" rank="species" species="yangae" subGenus="Lihelophorus">
<emphasis id="8DA7EA9B8D772858FEED5AF172E7FE7F" bold="true" box="[269,399,444,467]" italics="true" pageId="14" pageNumber="123">Helophorus</emphasis>
(
<emphasis id="8DA7EA9B8D772858FE7F5AF1715BFE7F" bold="true" box="[415,563,444,467]" italics="true" pageId="14" pageNumber="123">Lihelophorus</emphasis>
)
<emphasis id="8DA7EA9B8D772858FDA35AF171FAFE7F" bold="true" box="[579,658,444,467]" italics="true" pageId="14" pageNumber="123">yangae</emphasis>
</taxonomicName>
<collectingRegion id="7D17F86B8D772858FD785AF1718AFE7F" box="[664,738,444,467]" country="United Kingdom" name="Angus" pageId="14" pageNumber="123">Angus</collectingRegion>
, Fiká
</emphasis>
č
<emphasis id="8DA7EA9B8D772858FCCE5AF17099FE7F" bold="true" box="[814,1009,444,467]" pageId="14" pageNumber="123">ek &amp; Jia, sp. nov.</emphasis>
(
<figureCitation id="27E82A0C8D772858FE2D5A977125FE41" box="[461,589,474,493]" captionStart="Fig" captionStartId="6.[151,182,1490,1509]" captionTargetBox="[152,1126,220,1478]" captionTargetId="figure-10@6.[151,1127,217,1486]" captionTargetPageId="6" captionText="Fig. 1. Adult morphology of Helophorus (Lihelophorus). ab H. yangae sp. nov. (a ventral view of the head; b detail of mentum and labial palps); cd maxillary galea and lacinia (c H. yangae sp. nov.; d H. ser Zaitzev, 1908); ef mandible (e H. yangae sp. nov.; f H. ser); g labial palpus of H. ser; hl hind legs in dorsal view (h H. yangae sp. nov.; i H. ser; j H. lamicola Zaitzev, 1908; k detail of apex of tibia and base of tarsus of H. lamicola; l detail of tarsomere 5 and claw of H. ser). Abbreviations: sh swimming hairs. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311786" httpUri="https://zenodo.org/record/5311786/files/figure.png" pageId="14" pageNumber="123">Figs 1ac, e, h</figureCitation>
;
<figureCitation id="27E82A0C8D772858FDB65A9771FFFE41" box="[598,663,474,493]" captionStart="Fig" captionStartId="7.[113,144,1463,1482]" captionTargetBox="[133,1067,220,1453]" captionTargetId="figure-13@7.[133,1068,219,1454]" captionTargetPageId="7" captionText="Fig. 2. Adult morphology of Helophorus (Lihelophorus). af details of left half of pronotum (a,c,e) and base of left elytron (b, d, f): ab H. yangae sp. nov.; cd H. ser Zaitzev, 1908; ef H. lamicola Zaitzev, 1908; arrow in (a) shows remnants of pronotal granulations, arrows in (b, d, f) show scutellary stria. gh prothoracic hypomeron and suprapleural area (g H. yangae sp. nov.; h H. ser). i detail of posterior margin of abdominal ventrite 5 of H. ser, showing the irregular denticulation. j ventral view of the thorax of H. ser. k dorsolateral portion of left elytron of H. lamicola. Abbreviations: ep epipleuron; li last elytral interval. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311790" httpUri="https://zenodo.org/record/5311790/files/figure.png" pageId="14" pageNumber="123">2ab, g</figureCitation>
;
<figureCitation id="27E82A0C8D772858FD405A9771A5FE41" box="[672,717,474,493]" captionStart="Fig" captionStartId="10.[151,182,1482,1501]" captionTargetBox="[159,1124,234,1349]" captionTargetId="figure-10@10.[151,1127,219,1365]" captionTargetPageId="10" captionText="Fig. 3. Habitus of the species of Helophorus (Lihelophorus). ac H. lamicola Zaitzev, 1908 (a Qinghai: Zhaling Hu (lectotype); bc Quinghai: Golo).dg H.ser Zaitzev, 1908 (de Qinghai:Golo; f Xizang, Nyainqentanglha Feng; g Gansu: Xiahe env.). hk H. yangae sp. nov. (h Xizang: 90 km W of Amdo; i Xizang: 25 km W of Amdo; j Qinghai: Golo; k India: Kar Tso (Orchymont coll.)). All specimens to scale." figureDoi="http://doi.org/10.5281/zenodo.5311792" httpUri="https://zenodo.org/record/5311792/files/figure.png" pageId="14" pageNumber="123">3hk</figureCitation>
;
<figureCitation id="27E82A0C8D772858FD365A977068FE41" box="[726,768,474,493]" captionStart="Fig" captionStartId="12.[151,182,1058,1077]" captionTargetBox="[164,1118,238,1008]" captionTargetId="figure-10@12.[151,1129,219,1044]" captionTargetPageId="12" captionText="Fig. 5. Details of the head and pronotum of the Helophorus (Lihelophorus) yangae sp. nov. (a Xizang: 90 km W of Amdo; b,d Qinghai: Golo; c Xizang: 25 km W of Amdo; e India: Kar Tso (Orchymont coll.). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311796" httpUri="https://zenodo.org/record/5311796/files/figure.png" pageId="14" pageNumber="123">5ae</figureCitation>
;
<figureCitation id="27E82A0C8D772858FCE95A977058FE41" box="[777,816,474,493]" captionStart="Fig" captionStartId="15.[113,144,1496,1515]" captionTargetBox="[149,1030,229,1439]" captionTargetId="figure-13@15.[139,1063,219,1474]" captionTargetPageId="15" captionText="Fig. 6.Aedeagophores of Helophorus (Lihelophorus) species. ae H. (L.) ser Zaitzev, 1908 (a lectotype, Qinghai: valley of Alag Hu lake; bc Xizang: valley SE of Dong La pass; d Qinghai: Golo, 20 km E of Maduo; e Qinghai: Zuimatan). fj H. (L.) lamicola Zaitzev, 1908 (f lectotype, Qinghai: Djarin Nor; gi Qinghai: Zuimatan; j paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). ko H. (L.) yangae sp. nov. (k holotype, Qinghai: Zuimatan; lm paratypes, same locality; n Qinghai: Golo, Yematan; o Xizang: Gongbujiangda)." figureDoi="http://doi.org/10.5281/zenodo.5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="14" pageNumber="123">6jn</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F7C965028D772858FF77594971E8FB4B" pageId="14" pageNumber="123" type="materials_examined">
<paragraph id="BF6C36898D772858FF77594970ECFDBB" blockId="14.[151,1128,516,1255]" box="[151,900,516,535]" pageId="14" pageNumber="123">
<emphasis id="8DA7EA9B8D772858FF7759497278FDBB" bold="true" box="[151,272,516,535]" pageId="14" pageNumber="123">
<typeStatus id="6068882B8D772858FF77594973ACFDBB" box="[151,196,516,535]" pageId="14" pageNumber="123">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="C7C476198D772858FEF559497222FDBB" box="[277,330,516,535]" name="China" pageId="14" pageNumber="123">China</collectingCountry>
,
<collectingRegion id="7D17F86B8D772858FEB359497284FDBB" box="[339,492,516,535]" country="China" name="Qinghai" pageId="14" pageNumber="123">Qinghai Province</collectingRegion>
. Zuimatan, Roadside pool,
<geoCoordinate id="DAE7504E8D772858FD3559497040FDBB" box="[725,808,516,535]" degrees="35.333" direction="north" orientation="latitude" pageId="14" pageNumber="123" precision="55" value="35.333">35.333°N</geoCoordinate>
<geoCoordinate id="DAE7504E8D772858FCCD594970E8FDBB" box="[813,896,516,535]" degrees="99.007" direction="east" orientation="longitude" pageId="14" pageNumber="123" precision="55" value="99.007">99.007°E</geoCoordinate>
.
</paragraph>
<paragraph id="BF6C36898D772858FF77595271E8FB4B" blockId="14.[151,1128,516,1255]" pageId="14" pageNumber="123">
<emphasis id="8DA7EA9B8D772858FF77595272E8FD9E" bold="true" box="[151,384,543,562]" pageId="14" pageNumber="123">
Type material (
<specimenCount id="A9D5FD008D772858FEC05952721EFD9E" box="[288,374,543,562]" pageId="14" pageNumber="123" type="generic">143 spec.</specimenCount>
):
</emphasis>
<materialsCitation id="0FBB3CD48D772858FE6559527041FDE0" collectingDate="2013-06-07" collectionCode="SYSU" collectorName="R. B. Angus &amp; F. L. Jia &amp; Zhang" county="Golo" elevation="4141" latitude="35.331112" location="Zuimatan" longLatPrecision="19" longitude="99.065834" municipality="Huamuxia" pageId="14" pageNumber="123" specimenCount="1" typeStatus="holotype">
<typeStatus id="6068882B8D772858FE65595272B0FD9C" box="[389,472,543,562]" pageId="14" pageNumber="123" type="holotype">HOLOTYPE</typeStatus>
: J (
<collectionCode id="D9C2AE4C8D772858FE1D59527150FD9E" box="[509,568,543,562]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="14" pageNumber="123" type="Herbarium">SYSU</collectionCode>
): CHINA QINGHAI
<collectingCounty id="560D4E058D772858FD1459527049FD9E" box="[756,801,543,562]" pageId="14" pageNumber="123">Golo</collectingCounty>
,
<collectingMunicipality id="5F08ACF38D772858FCC9595270ECFD9E" box="[809,900,543,562]" pageId="14" pageNumber="123">Huamuxia</collectingMunicipality>
,
<location id="BA0C60528D772858FC6B595270B7FD9E" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FC6B595270B7FD9E" box="[907,991,543,562]" county="Golo" latitude="35.331112" longLatPrecision="19" longitude="99.065834" municipality="Huamuxia" name="Zuimatan" pageId="14" pageNumber="123">Zuimatan</location>
,
<location id="BA0C60528D772858FC075952775EFD9E" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FC075952775EFD9E" box="[999,1078,543,562]" county="Golo" latitude="35.331112" longLatPrecision="19" longitude="99.065834" municipality="Huamuxia" name="Roadside" pageId="14" pageNumber="123">Roadside</location>
pool,
<geoCoordinate id="DAE7504E8D772858FF7759747395FDE0" box="[151,253,569,588]" degrees="35" direction="north" minutes="19" orientation="latitude" pageId="14" pageNumber="123" precision="15" seconds="52" value="35.331112">35°1952”N</geoCoordinate>
<geoCoordinate id="DAE7504E8D772858FEE05974720DFDE0" box="[256,357,569,588]" degrees="99" direction="east" minutes="03" orientation="longitude" pageId="14" pageNumber="123" precision="15" seconds="57" value="99.065834">99°0357”E</geoCoordinate>
,
<quantity id="782B9B6C8D772858FE8C597472C2FDE0" box="[364,426,569,588]" metricMagnitude="3" metricUnit="m" metricValue="4.141" pageId="14" pageNumber="123" unit="m" value="4141.0">
<elevation id="34FED1BA8D772858FE8C597472C2FDE0" box="[364,426,569,588]" metricMagnitude="3" metricUnit="m" metricValue="4.141" pageId="14" pageNumber="123" unit="m" value="4141.0">4141m</elevation>
</quantity>
,
<date id="CB6D10498D772858FE505974716BFDE0" box="[432,515,569,588]" pageId="14" pageNumber="123" value="2013-06-07">
<collectingDate id="DB29E9A18D772858FE505974716BFDE0" box="[432,515,569,588]" pageId="14" pageNumber="123" value="2013-06-07">7.vi.2013</collectingDate>
</date>
,
<collectorName id="1226535F8D772858FDE959747102FDE0" box="[521,618,569,588]" pageId="14" pageNumber="123">R.B.Angus</collectorName>
,
<collectorName id="1226535F8D772858FD91597471C1FDE0" box="[625,681,569,588]" pageId="14" pageNumber="123">F.L.Jia</collectorName>
&amp; Y.
<collectorName id="1226535F8D772858FD3359747062FDE0" box="[723,778,569,588]" pageId="14" pageNumber="123">Zhang</collectorName>
lgt
</materialsCitation>
.
<materialsCitation id="0FBB3CD48D772858FCD3597472C5FD2E" collectingDate="2013-06-12" collectingDateMax="2013-06-13" collectingDateMin="2013-06-12" collectionCode="SYSU" collectorName="R. B. Angus &amp; Angus" country="China" county="Golo" elevation="4141" latitude="35.331112" location="Zuimatan" longLatPrecision="19" longitude="99.065834" municipality="Huamuxia" pageId="14" pageNumber="123" specimenCount="2" specimenCount-female="2" stateProvince="Qinghai" typeStatus="paratype">
<typeStatus id="6068882B8D772858FCD3597470E2FDE7" box="[819,906,569,588]" pageId="14" pageNumber="123" type="paratype">PARATYPES</typeStatus>
:
<emphasis id="8DA7EA9B8D772858FC745974775DFDE0" bold="true" box="[916,1077,569,588]" pageId="14" pageNumber="123">
<collectingCountry id="C7C476198D772858FC74597470B6FDE0" box="[916,990,569,588]" name="China" pageId="14" pageNumber="123">CHINA</collectingCountry>
:
<collectingRegion id="7D17F86B8D772858FC0559747746FDE7" box="[997,1070,569,588]" country="China" name="Qinghai" pageId="14" pageNumber="123">QINGHAI</collectingRegion>
:
</emphasis>
2 JJ
<specimenCount id="A9D5FD008D772858FF77591973AFFDCB" box="[151,199,590,615]" pageId="14" pageNumber="123" type="female">2 ♀♀</specimenCount>
(BMNH): same data as the holotype [1 J: chromosome prep. 3,
<date id="CB6D10498D772858FD175919703CFDCB" box="[759,852,596,615]" pageId="14" pageNumber="123" value="2013-06-12">
<collectingDate id="DB29E9A18D772858FD175919703CFDCB" box="[759,852,596,615]" pageId="14" pageNumber="123" value="2013-06-12">12.vi.2013</collectingDate>
</date>
,
<collectorName id="1226535F8D772858FCBD591970ABFDCB" box="[861,963,596,615]" pageId="14" pageNumber="123">R.B. Angus</collectorName>
; 1 J: chromosome prep. 1,
<date id="CB6D10498D772858FF3C59227251FD2E" box="[220,313,623,642]" pageId="14" pageNumber="123" value="2013-06-13">
<collectingDate id="DB29E9A18D772858FF3C59227251FD2E" box="[220,313,623,642]" pageId="14" pageNumber="123" value="2013-06-13">13.vi.2013</collectingDate>
</date>
, R.B.
<collectorName id="1226535F8D772858FE8C592272C0FD2E" box="[364,424,623,642]" pageId="14" pageNumber="123">Angus</collectorName>
]
</materialsCitation>
;
<materialsCitation id="0FBB3CD48D772858FE565922705FFD2E" box="[438,823,616,642]" collectingDate="2013-06-07" collectionCode="SYSU" collectorName="R. B. Angus &amp; F. L. Jia &amp; Zhang" county="Golo" elevation="4141" latitude="35.331112" location="Zuimatan" longLatPrecision="19" longitude="99.065834" municipality="Huamuxia" pageId="14" pageNumber="123" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
2 JJ
<specimenCount id="A9D5FD008D772858FE0959227160FD2D" box="[489,520,616,642]" pageId="14" pageNumber="123" type="female">1 ♀</specimenCount>
(
<collectionCode id="D9C2AE4C8D772858FDF359227126FD2E" box="[531,590,623,642]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="14" pageNumber="123" type="Herbarium">SYSU</collectionCode>
): same data as the holotype
</materialsCitation>
;
<materialsCitation id="0FBB3CD48D772858FCA05922713AFD1B" collectingDate="2013-06-08" collectingDateMax="2013-06-10" collectingDateMin="2013-06-08" collectorName="R. B. Angus &amp; F. L. Jia &amp; Y. Zhang &amp; Angus" county="Golo" elevation="4240" latitude="34.67972" location="Yematan" longLatPrecision="19" longitude="99.065834" municipality="Maduo" pageId="14" pageNumber="123" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
1 J
<specimenCount id="A9D5FD008D772858FC84592270F9FD2D" box="[868,913,616,642]" pageId="14" pageNumber="123" type="female">2 ♀♀</specimenCount>
(BMNH):
<collectingCounty id="560D4E058D772858FC1059227775FD2E" box="[1008,1053,623,642]" pageId="14" pageNumber="123">Golo</collectingCounty>
,
<collectingMunicipality id="5F08ACF38D772858FBC55922770CFD2E" box="[1061,1124,623,642]" pageId="14" pageNumber="123">Maduo</collectingMunicipality>
, roadside pools near
<location id="BA0C60528D772858FEA459C472F9FD30" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FEA459C472F9FD30" box="[324,401,649,668]" county="Golo" latitude="34.67972" longLatPrecision="19" longitude="99.065834" municipality="Maduo" name="Yematan" pageId="14" pageNumber="123">Yematan</location>
,
<geoCoordinate id="DAE7504E8D772858FE7B59C47168FD30" box="[411,512,649,668]" degrees="34" direction="north" minutes="40" orientation="latitude" pageId="14" pageNumber="123" precision="15" seconds="47" value="34.67972">34°4047″N</geoCoordinate>
<geoCoordinate id="DAE7504E8D772858FDE659C47103FD30" box="[518,619,649,668]" degrees="99" direction="east" minutes="03" orientation="longitude" pageId="14" pageNumber="123" precision="15" seconds="57" value="99.065834">99°0357″E</geoCoordinate>
,
<quantity id="782B9B6C8D772858FD9559C471D1FD30" box="[629,697,649,668]" metricMagnitude="3" metricUnit="m" metricValue="4.24" pageId="14" pageNumber="123" unit="m" value="4240.0">
<elevation id="34FED1BA8D772858FD9559C471D1FD30" box="[629,697,649,668]" metricMagnitude="3" metricUnit="m" metricValue="4.24" pageId="14" pageNumber="123" unit="m" value="4240.0">4240 m</elevation>
</quantity>
,
<date id="CB6D10498D772858FD2259C4707DFD30" box="[706,789,649,668]" pageId="14" pageNumber="123" value="2013-06-08">
<collectingDate id="DB29E9A18D772858FD2259C4707DFD30" box="[706,789,649,668]" pageId="14" pageNumber="123" value="2013-06-08">8.vi.2013</collectingDate>
</date>
,
<collectorName id="1226535F8D772858FCFE59C470E2FD30" box="[798,906,649,668]" pageId="14" pageNumber="123">R. B. Angus</collectorName>
,
<collectorName id="1226535F8D772858FC7459C470B2FD30" box="[916,986,649,668]" pageId="14" pageNumber="123">F. L. Jia</collectorName>
&amp;
<collectorName id="1226535F8D772858FC1659C4772DFD30" box="[1014,1093,649,668]" pageId="14" pageNumber="123">Y. Zhang</collectorName>
lgt. [J: chromosome prep. 2,
<date id="CB6D10498D772858FE9759E972BCFD1B" box="[375,468,676,695]" pageId="14" pageNumber="123" value="2013-06-10">
<collectingDate id="DB29E9A18D772858FE9759E972BCFD1B" box="[375,468,676,695]" pageId="14" pageNumber="123" value="2013-06-10">10.vi.2013</collectingDate>
</date>
, R. B.
<collectorName id="1226535F8D772858FDF159E97125FD1B" box="[529,589,676,695]" pageId="14" pageNumber="123">Angus</collectorName>
]
</materialsCitation>
;
<materialsCitation id="0FBB3CD48D772858FDBC59E970CBFD1B" box="[604,931,670,696]" collectionCode="SYSU" pageId="14" pageNumber="123" specimenCount="9" specimenCount-female="9" typeStatus="paratype">
6 JJ
<specimenCount id="A9D5FD008D772858FD7359E971ACFD1B" box="[659,708,670,695]" pageId="14" pageNumber="123" type="female">9 ♀♀</specimenCount>
(
<collectionCode id="D9C2AE4C8D772858FD3059E97063FD1B" box="[720,779,676,695]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="14" pageNumber="123" type="Herbarium">SYSU</collectionCode>
): same label data
</materialsCitation>
;
<materialsCitation id="0FBB3CD48D772858FC4D59E9715BFD7E" collectionCode="NMPC" pageId="14" pageNumber="123" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="A9D5FD008D772858FC4D59E970B5FD1B" box="[941,989,670,695]" pageId="14" pageNumber="123" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="D9C2AE4C8D772858FC0A59E97744FD1B" box="[1002,1068,676,695]" country="Czech Republic" name="National Museum Prague" pageId="14" pageNumber="123">NMPC</collectionCode>
): same label data [DNA vouchers MF662 and MF664]
</materialsCitation>
;
<materialsCitation id="0FBB3CD48D772858FDDD59F2773AFD40" collectingDate="2013-06-08" collectionCode="SYSU" collectorName="R. B. Angus &amp; F. L. Jia &amp; Zhang" county="Golo" elevation="4240" latitude="34.67972" location="Heihexiang" longLatPrecision="19" longitude="99.065834" municipality="Maduo" pageId="14" pageNumber="123" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
3 JJ
<specimenCount id="A9D5FD008D772858FD9459F271CEFD7D" box="[628,678,696,722]" pageId="14" pageNumber="123" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="D9C2AE4C8D772858FD5159F27184FD7E" box="[689,748,703,722]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="14" pageNumber="123" type="Herbarium">SYSU</collectionCode>
):
<collectingCounty id="560D4E058D772858FD1C59F27041FD7E" box="[764,809,703,722]" pageId="14" pageNumber="123">Golo</collectingCounty>
,
<collectingMunicipality id="5F08ACF38D772858FCD359F2701AFD7E" box="[819,882,703,722]" pageId="14" pageNumber="123">Maduo</collectingMunicipality>
,
<location id="BA0C60528D772858FC9B59F27088FD7E" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FC9B59F27088FD7E" box="[891,992,703,722]" county="Golo" latitude="34.67972" longLatPrecision="19" longitude="99.065834" municipality="Maduo" name="Heihexiang" pageId="14" pageNumber="123">Heihexiang</location>
, roadside pools near
<location id="BA0C60528D772858FF225994723FFD40" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FF225994723FFD40" box="[194,343,729,748]" county="Golo" latitude="34.67972" longLatPrecision="19" longitude="99.065834" municipality="Maduo" name="Xingxinghai lake" pageId="14" pageNumber="123">Xingxinghai lake</location>
<geoCoordinate id="DAE7504E8D772858FEBC599472AAFD40" box="[348,450,729,748]" degrees="34" direction="north" minutes="40" orientation="latitude" pageId="14" pageNumber="123" precision="15" seconds="47" value="34.67972">34°4047″N</geoCoordinate>
<geoCoordinate id="DAE7504E8D772858FE2859947145FD40" box="[456,557,729,748]" degrees="99" direction="east" minutes="03" orientation="longitude" pageId="14" pageNumber="123" precision="15" seconds="57" value="99.065834">99°0357″E</geoCoordinate>
,
<elevation id="34FED1BA8D772858FDD7599471CFFD40" box="[567,679,729,748]" metricMagnitude="3" metricUnit="m" metricValue="4.24" pageId="14" pageNumber="123" unit="m" value="4240.0">
<quantity id="782B9B6C8D772858FDD759947110FD40" box="[567,632,729,748]" metricMagnitude="3" metricUnit="m" metricValue="4.24" pageId="14" pageNumber="123" unit="m" value="4240.0">4240 m</quantity>
a.s.l.
</elevation>
,
<date id="CB6D10498D772858FD515994706CFD40" box="[689,772,729,748]" pageId="14" pageNumber="123" value="2013-06-08">
<collectingDate id="DB29E9A18D772858FD515994706CFD40" box="[689,772,729,748]" pageId="14" pageNumber="123" value="2013-06-08">8.vi.2013</collectingDate>
</date>
,
<collectorName id="1226535F8D772858FCED59947011FD40" box="[781,889,729,748]" pageId="14" pageNumber="123">R. B. Angus</collectorName>
,
<collectorName id="1226535F8D772858FC63599470A1FD40" box="[899,969,729,748]" pageId="14" pageNumber="123">F. L. Jia</collectorName>
&amp; Y.
<collectorName id="1226535F8D772858FC1D5994775CFD40" box="[1021,1076,729,748]" pageId="14" pageNumber="123">Zhang</collectorName>
lgt
</materialsCitation>
;
<materialsCitation id="0FBB3CD48D772858FBBC5994715FFC8E" collectingDate="2013-06-08" collectorName="R. B. Angus &amp; F. L. Jia &amp; Zhang" elevation="4290" latitude="34.85472" location="Maduo" longLatPrecision="19" longitude="98.28833" municipality="Golo" pageId="14" pageNumber="123" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="A9D5FD008D772858FBBC599473B9FCAB" pageId="14" pageNumber="123" type="female">1 female</specimenCount>
(BMNH):
<collectingMunicipality id="5F08ACF38D772858FECF59B97234FCAB" box="[303,348,756,775]" pageId="14" pageNumber="123">Golo</collectingMunicipality>
,
<location id="BA0C60528D772858FE8459B972CBFCAB" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FE8459B972CBFCAB" box="[356,419,756,775]" latitude="34.85472" longLatPrecision="19" longitude="98.28833" municipality="Golo" name="Maduo" pageId="14" pageNumber="123">Maduo</location>
, roadside pools on river flats
<locationDeviation id="AEDE2FAB8D772858FD7B59B97029FCAB" box="[667,833,756,775]" location="Maduo." pageId="14" pageNumber="123">
<quantity id="782B9B6C8D772858FD7B59B971B8FCAB" box="[667,720,756,775]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="123" unit="km" value="20.0">20 km</quantity>
E of Maduo.
</locationDeviation>
<geoCoordinate id="DAE7504E8D772858FCA459B970C3FCAB" box="[836,939,756,775]" degrees="34" direction="north" minutes="51" orientation="latitude" pageId="14" pageNumber="123" precision="15" seconds="17" value="34.85472">34°5117″N</geoCoordinate>
,
<geoCoordinate id="DAE7504E8D772858FC5459B97771FCAB" box="[948,1049,756,775]" degrees="98" direction="east" minutes="17" orientation="longitude" pageId="14" pageNumber="123" precision="15" seconds="18" value="98.28833">98°1718″E</geoCoordinate>
,
<quantity id="782B9B6C8D772858FBC159B9770BFCAB" box="[1057,1123,756,775]" metricMagnitude="3" metricUnit="m" metricValue="4.29" pageId="14" pageNumber="123" unit="m" value="4290.0">
<elevation id="34FED1BA8D772858FBC159B9770BFCAB" box="[1057,1123,756,775]" metricMagnitude="3" metricUnit="m" metricValue="4.29" pageId="14" pageNumber="123" unit="m" value="4290.0">4290 m</elevation>
</quantity>
,
<date id="CB6D10498D772858FF7758427382FC8E" box="[151,234,783,802]" pageId="14" pageNumber="123" value="2013-06-08">
<collectingDate id="DB29E9A18D772858FF7758427382FC8E" box="[151,234,783,802]" pageId="14" pageNumber="123" value="2013-06-08">8.vi.2013</collectingDate>
</date>
,
<collectorName id="1226535F8D772858FF1358427237FC8E" box="[243,351,783,802]" pageId="14" pageNumber="123">R. B. Angus</collectorName>
,
<collectorName id="1226535F8D772858FE87584272C5FC8E" box="[359,429,783,802]" pageId="14" pageNumber="123">F. L. Jia</collectorName>
&amp; Y.
<collectorName id="1226535F8D772858FE005842717FFC8E" box="[480,535,783,802]" pageId="14" pageNumber="123">Zhang</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D772858FDA4584270E3FCFB" collectingDate="1986-07-20" collectingDateMax="2013-06-12" collectingDateMin="1986-07-20" collectionCode="SYSU" collectorName="F. L. Jia &amp; Y. Zhang &amp; R. B. Angus &amp; Rikaze &amp; Liang Ge-Qiu" elevation="3370" latitude="37.3" location="Gangca" longLatPrecision="1182" longitude="100.183334" municipality="Hu" pageId="14" pageNumber="123" specimenCount="1" specimenCount-female="1" stateProvince="Qinghai" typeStatus="paratype">
<specimenCount id="A9D5FD008D772858FDA45842710CFC8D" box="[580,612,776,802]" pageId="14" pageNumber="123" type="female">1 ♀</specimenCount>
(BMNH): N.
<collectingRegion id="7D17F86B8D772858FD3E5842704BFC8E" box="[734,803,783,802]" country="China" name="Qinghai" pageId="14" pageNumber="123">Qinghai</collectingRegion>
<collectingMunicipality id="5F08ACF38D772858FCC95842702DFC8E" box="[809,837,783,802]" pageId="14" pageNumber="123">Hu</collectingMunicipality>
,
<location id="BA0C60528D772858FCAE584270F9FC8E" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FCAE584270F9FC8E" box="[846,913,783,802]" latitude="37.3" longLatPrecision="1182" longitude="100.183334" municipality="Hu" name="Gangca" pageId="14" pageNumber="123" stateProvince="Qinghai">Gangca</location>
, roadside pool.
<geoCoordinate id="DAE7504E8D772858FBFA5842770BFC8E" box="[1050,1123,783,802]" degrees="37" direction="north" minutes="18" orientation="latitude" pageId="14" pageNumber="123" precision="925" value="37.3">37°18N</geoCoordinate>
,
<geoCoordinate id="DAE7504E8D772858FF7758647380FC90" box="[151,232,809,828]" degrees="100" direction="east" minutes="11" orientation="longitude" pageId="14" pageNumber="123" precision="925" value="100.183334">100°11E</geoCoordinate>
,
<quantity id="782B9B6C8D772858FF115864725CFC90" box="[241,308,809,828]" metricMagnitude="3" metricUnit="m" metricValue="3.37" pageId="14" pageNumber="123" unit="m" value="3370.0">
<elevation id="34FED1BA8D772858FF115864725CFC90" box="[241,308,809,828]" metricMagnitude="3" metricUnit="m" metricValue="3.37" pageId="14" pageNumber="123" unit="m" value="3370.0">3370 m</elevation>
</quantity>
,
<date id="CB6D10498D772858FEDD586472E7FC90" box="[317,399,809,828]" pageId="14" pageNumber="123" value="2013-06-05">
<collectingDate id="DB29E9A18D772858FEDD586472E7FC90" box="[317,399,809,828]" pageId="14" pageNumber="123" value="2013-06-05">5.vi.2013</collectingDate>
</date>
,
<collectorName id="1226535F8D772858FE78586472B5FC90" box="[408,477,809,828]" pageId="14" pageNumber="123">F. L. Jia</collectorName>
&amp;
<collectorName id="1226535F8D772858FE185864712EFC90" box="[504,582,809,828]" pageId="14" pageNumber="123">Y. Zhang</collectorName>
lgt. [chromosome prep. 2,
<date id="CB6D10498D772858FCCE586470E3FC90" box="[814,907,809,828]" pageId="14" pageNumber="123" value="2013-06-12">
<collectingDate id="DB29E9A18D772858FCCE586470E3FC90" box="[814,907,809,828]" pageId="14" pageNumber="123" value="2013-06-12">12.vi.2013</collectingDate>
</date>
,
<collectorName id="1226535F8D772858FC7458647769FC90" box="[916,1025,809,828]" pageId="14" pageNumber="123">R. B. Angus</collectorName>
].
<emphasis id="8DA7EA9B8D772858FBEF5864773EFC90" bold="true" box="[1039,1110,809,828]" pageId="14" pageNumber="123">
<collectingRegion id="7D17F86B8D772858FBEF58647727FC97" box="[1039,1103,809,828]" country="China" name="Xizang" pageId="14" pageNumber="123">XIZANG</collectingRegion>
:
</emphasis>
3 JJ 4 unsexed spec. (
<collectionCode id="D9C2AE4C8D772858FEA7580972EAFCFB" box="[327,386,836,855]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="14" pageNumber="123" type="Herbarium">SYSU</collectionCode>
):suburb of
<collectorName id="1226535F8D772858FE0258097177FCFB" box="[482,543,836,855]" pageId="14" pageNumber="123">Rikaze</collectorName>
,
<quantity id="782B9B6C8D772858FDC65809710EFCFB" box="[550,614,836,855]" metricMagnitude="3" metricUnit="m" metricValue="3.826" pageId="14" pageNumber="123" unit="m" value="3826.0">
<elevation id="34FED1BA8D772858FDC65809710EFCFB" box="[550,614,836,855]" metricMagnitude="3" metricUnit="m" metricValue="3.826" pageId="14" pageNumber="123" unit="m" value="3826.0">3826 m</elevation>
</quantity>
,
<date id="CB6D10498D772858FD8E58097199FCFB" box="[622,753,836,855]" pageId="14" pageNumber="123" value="1986-07-20" valueMax="1986-07-23" valueMin="1986-07-20">
<collectingDate id="DB29E9A18D772858FD8E58097199FCFB" box="[622,753,836,855]" pageId="14" pageNumber="123" value="1986-07-20" valueMax="1986-07-23" valueMin="1986-07-20">2023.vii.1986</collectingDate>
</date>
,
<collectorName id="1226535F8D772858FD1758097005FCFB" box="[759,877,836,855]" pageId="14" pageNumber="123">Liang Ge-Qiu</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D772858FC765809716BFCDE" collectingDate="1986-07-27" collectingDateMax="1986-07-28" collectingDateMin="1986-07-27" collectionCode="SYSU" collectorName="Liang Ge-Qiu" location="Jiangzi" pageId="14" pageNumber="123" specimenCount="1" stateProvince="Qinghai" typeStatus="paratype">
1 unsexed spec.(
<collectionCode id="D9C2AE4C8D772858FBC258097735FCFB" box="[1058,1117,836,855]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="14" pageNumber="123" type="Herbarium">SYSU</collectionCode>
):
<location id="BA0C60528D772858FF77581273BEFCDE" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FF77581273BEFCDE" box="[151,214,863,882]" name="Jiangzi" pageId="14" pageNumber="123" stateProvince="Qinghai">Jiangzi</location>
,
<date id="CB6D10498D772858FF3F5812720AFCDE" box="[223,354,863,882]" pageId="14" pageNumber="123" value="1986-07-27" valueMax="1986-07-28" valueMin="1986-07-27">
<collectingDate id="DB29E9A18D772858FF3F5812720AFCDE" box="[223,354,863,882]" pageId="14" pageNumber="123" value="1986-07-27" valueMax="1986-07-28" valueMin="1986-07-27">2728.vii.1986</collectingDate>
</date>
,
<collectorName id="1226535F8D772858FE8B5812728AFCDE" box="[363,482,863,882]" pageId="14" pageNumber="123">Liang Ge-Qiu</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D772858FDEF58127001FC20" collectingDate="2013-08-06" collectionCode="SYSU" collectorName="Jia Yue &amp; Jia Fenglong" elevation="4320" latitude="30.09533" location="Yangbajing town" longLatPrecision="1" longitude="90.53926" municipality="Dangxiong" pageId="14" pageNumber="123" specimenCount="3" specimenCount-female="3" stateProvince="Qinghai" typeStatus="paratype">
3 JJ
<specimenCount id="A9D5FD008D772858FDA45812711CFCDD" box="[580,628,856,882]" pageId="14" pageNumber="123" type="female">3 ♀♀</specimenCount>
, 19 unsexed spec. (
<collectionCode id="D9C2AE4C8D772858FCFC5812703FFCDE" box="[796,855,863,882]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="14" pageNumber="123" type="Herbarium">SYSU</collectionCode>
):
<collectingMunicipality id="5F08ACF38D772858FC86581270AFFCDE" box="[870,967,863,882]" pageId="14" pageNumber="123">Dangxiong</collectingMunicipality>
,
<location id="BA0C60528D772858FC2F5812770BFCDE" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FC2F5812770BFCDE" box="[975,1123,863,882]" latitude="30.09533" longLatPrecision="1" longitude="90.53926" municipality="Dangxiong" name="Yangbajing town" pageId="14" pageNumber="123" stateProvince="Qinghai">Yangbajing town</location>
,
<elevation id="34FED1BA8D772858FF775834726FFC20" box="[151,263,889,908]" metricMagnitude="3" metricUnit="m" metricValue="4.32" pageId="14" pageNumber="123" unit="m" value="4320.0">
<quantity id="782B9B6C8D772858FF77583473B0FC20" box="[151,216,889,908]" metricMagnitude="3" metricUnit="m" metricValue="4.32" pageId="14" pageNumber="123" unit="m" value="4320.0">4320 m</quantity>
a.s.l.
</elevation>
,
<date id="CB6D10498D772858FEF058347207FC20" box="[272,367,889,908]" pageId="14" pageNumber="123" value="2013-08-06">
<collectingDate id="DB29E9A18D772858FEF058347207FC20" box="[272,367,889,908]" pageId="14" pageNumber="123" value="2013-08-06">6.viii.2013</collectingDate>
</date>
,
<geoCoordinate id="DAE7504E8D772858FE9858347287FC20" box="[376,495,889,908]" degrees="30" direction="north" minutes="5" orientation="latitude" pageId="14" pageNumber="123" precision="1" seconds="43.18" value="30.09533">30°543.18″N</geoCoordinate>
,
<geoCoordinate id="DAE7504E8D772858FE1858347110FC20" box="[504,632,889,908]" degrees="90" direction="east" minutes="32" orientation="longitude" pageId="14" pageNumber="123" precision="1" seconds="21.33" value="90.53926">90°3221.33″E</geoCoordinate>
,
<collectorName id="1226535F8D772858FD61583471D6FC20" box="[641,702,889,908]" pageId="14" pageNumber="123">Jia Yue</collectorName>
&amp;
<collectorName id="1226535F8D772858FD3A58347021FC20" box="[730,841,889,908]" pageId="14" pageNumber="123">Jia Fenglong</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D772858FC955834720FFC6E" collectingDate="2013-08-02" collectionCode="SYSU" collectorName="Jia Yue &amp; Jia Fenglong" county="Zhanang County" elevation="3658" latitude="29.32309" location="Sangye Fery" longLatPrecision="1" longitude="91.509384" municipality="Nanshan" pageId="14" pageNumber="123" specimenCount="3" specimenCount-female="3" stateProvince="Qinghai" typeStatus="paratype">
2 JJ
<specimenCount id="A9D5FD008D772858FC4A583470B2FC20" box="[938,986,883,908]" pageId="14" pageNumber="123" type="female">3 ♀♀</specimenCount>
8 unsexed spec. (
<collectionCode id="D9C2AE4C8D772858FF7D58D973B0FC0B" box="[157,216,916,935]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="14" pageNumber="123" type="Herbarium">SYSU</collectionCode>
):
<collectingMunicipality id="5F08ACF38D772858FF0958D9725EFC0B" box="[233,310,916,935]" pageId="14" pageNumber="123">Nanshan</collectingMunicipality>
,
<collectingCounty id="560D4E058D772858FEA058D972B9FC0B" box="[320,465,916,935]" pageId="14" pageNumber="123">Zhanang County</collectingCounty>
,
<location id="BA0C60528D772858FE3B58D97121FC0B" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FE3B58D97121FC0B" box="[475,585,916,935]" county="Zhanang County" latitude="29.32309" longLatPrecision="1" longitude="91.509384" municipality="Nanshan" name="Sangye Fery" pageId="14" pageNumber="123" stateProvince="Qinghai">Sangye Fery</location>
,
<geoCoordinate id="DAE7504E8D772858FDB358D971BCFC0B" box="[595,724,916,935]" degrees="29" direction="north" minutes="19" orientation="latitude" pageId="14" pageNumber="123" precision="1" seconds="23.12" value="29.32309">29°1923.12″N</geoCoordinate>
,
<geoCoordinate id="DAE7504E8D772858FD3F58D97037FC0B" box="[735,863,916,935]" degrees="91" direction="east" minutes="30" orientation="longitude" pageId="14" pageNumber="123" precision="1" seconds="33.79" value="91.509384">91°3033.79″E</geoCoordinate>
,
<elevation id="34FED1BA8D772858FC8958D970B3FC0B" box="[873,987,916,935]" metricMagnitude="3" metricUnit="m" metricValue="3.658" pageId="14" pageNumber="123" unit="m" value="3658.0">
<quantity id="782B9B6C8D772858FC8958D970C4FC0B" box="[873,940,916,935]" metricMagnitude="3" metricUnit="m" metricValue="3.658" pageId="14" pageNumber="123" unit="m" value="3658.0">3658 m</quantity>
a.s.l.
</elevation>
,
<date id="CB6D10498D772858FC0658D9772DFC0B" box="[998,1093,916,935]" pageId="14" pageNumber="123" value="2013-08-02">
<collectingDate id="DB29E9A18D772858FC0658D9772DFC0B" box="[998,1093,916,935]" pageId="14" pageNumber="123" value="2013-08-02">2.viii.2013</collectingDate>
</date>
,
<collectorName id="1226535F8D772858FBAF58D973D0FC6E" pageId="14" pageNumber="123">Jia Yue</collectorName>
&amp;
<collectorName id="1226535F8D772858FF3658E2722EFC6E" box="[214,326,943,962]" pageId="14" pageNumber="123">Jia Fenglong</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D772858FE9558E27005FC70" collectingDate="2013-08-06" collectionCode="SYSU" collectorName="Jia Yue &amp; Jia Fenglong" county="Zhanang County" elevation="3575" latitude="29.323748" location="Sangye" longLatPrecision="1" longitude="91.50373" municipality="Nanshan" pageId="14" pageNumber="123" specimenCount="2" specimenCount-female="2" stateProvince="Qinghai" typeStatus="paratype">
2 JJ
<specimenCount id="A9D5FD008D772858FE4C58E272B6FC6D" box="[428,478,936,962]" pageId="14" pageNumber="123" type="female">2 ♀♀</specimenCount>
6 unsexed spec. (
<collectionCode id="D9C2AE4C8D772858FD9958E271DCFC6E" box="[633,692,943,962]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="14" pageNumber="123" type="Herbarium">SYSU</collectionCode>
):
<collectingMunicipality id="5F08ACF38D772858FD2558E2707AFC6E" box="[709,786,943,962]" pageId="14" pageNumber="123">Nanshan</collectingMunicipality>
,
<collectingCounty id="560D4E058D772858FCFB58E270C5FC6E" box="[795,941,943,962]" pageId="14" pageNumber="123">Zhanang County</collectingCounty>
, near
<location id="BA0C60528D772858FC0258E27749FC6E" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FC0258E27749FC6E" box="[994,1057,943,962]" county="Zhanang County" latitude="29.323748" longLatPrecision="1" longitude="91.50373" municipality="Nanshan" name="Sangye" pageId="14" pageNumber="123" stateProvince="Qinghai">Sangye</location>
temple,
<geoCoordinate id="DAE7504E8D772858FF7758847271FC70" box="[151,281,969,988]" degrees="29" direction="north" minutes="19" orientation="latitude" pageId="14" pageNumber="123" precision="1" seconds="25.49" value="29.323748">29°1925.49″N</geoCoordinate>
,
<geoCoordinate id="DAE7504E8D772858FEC2588472C9FC70" box="[290,417,969,988]" degrees="91" direction="east" minutes="30" orientation="longitude" pageId="14" pageNumber="123" precision="1" seconds="13.42" value="91.50373">91°3013.42″E</geoCoordinate>
,
<elevation id="34FED1BA8D772858FE4A58847170FC70" box="[426,536,969,988]" metricMagnitude="3" metricUnit="m" metricValue="3.575" pageId="14" pageNumber="123" unit="m" value="3575.0">
<quantity id="782B9B6C8D772858FE4A58847282FC70" box="[426,490,969,988]" metricMagnitude="3" metricUnit="m" metricValue="3.575" pageId="14" pageNumber="123" unit="m" value="3575.0">3575 m</quantity>
a.s.l.
</elevation>
,
<date id="CB6D10498D772858FDC1588471E8FC70" box="[545,640,969,988]" pageId="14" pageNumber="123" value="2013-08-06">
<collectingDate id="DB29E9A18D772858FDC1588471E8FC70" box="[545,640,969,988]" pageId="14" pageNumber="123" value="2013-08-06">6.viii.2013</collectingDate>
</date>
,
<collectorName id="1226535F8D772858FD68588471ADFC70" box="[648,709,969,988]" pageId="14" pageNumber="123">Jia Yue</collectorName>
&amp;
<collectorName id="1226535F8D772858FD0058847025FC70" box="[736,845,969,988]" pageId="14" pageNumber="123">Jia Fenglong</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D772858FC99588473B6FBBE" collectionCode="SYSU" collectorName="Jia Yue &amp; Jia Fenglong" county="Zhanang County" elevation="3620" latitude="29.32309" location="Sangye town" longLatPrecision="1" longitude="91.509384" municipality="Shannan" pageId="14" pageNumber="123" specimenCount="2" specimenCount-female="2" stateProvince="Qinghai" typeStatus="paratype">
2 JJ
<specimenCount id="A9D5FD008D772858FC4D588470B3FC70" box="[941,987,963,988]" pageId="14" pageNumber="123" type="female">2 ♀♀</specimenCount>
9 unsexed spec. (
<collectionCode id="D9C2AE4C8D772858FF7D58A973B0FC5B" box="[157,216,996,1015]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="14" pageNumber="123" type="Herbarium">SYSU</collectionCode>
):
<collectingMunicipality id="5F08ACF38D772858FF0658A9725AFC5B" box="[230,306,996,1015]" pageId="14" pageNumber="123">Shannan</collectingMunicipality>
,
<collectingCounty id="560D4E058D772858FED958A972AFFC5B" box="[313,455,996,1015]" pageId="14" pageNumber="123">Zhanang County</collectingCounty>
,
<location id="BA0C60528D772858FE2E58A97156FC5B" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FE2E58A97156FC5B" box="[462,574,996,1015]" county="Zhanang County" latitude="29.32309" longLatPrecision="1" longitude="91.509384" municipality="Shannan" name="Sangye town" pageId="14" pageNumber="123" stateProvince="Qinghai">Sangye town</location>
,
<geoCoordinate id="DAE7504E8D772858FDA558A971AFFC5B" box="[581,711,996,1015]" degrees="29" direction="north" minutes="19" orientation="latitude" pageId="14" pageNumber="123" precision="1" seconds="23.12" value="29.32309">29°1923.12″N</geoCoordinate>
,
<geoCoordinate id="DAE7504E8D772858FD2E58A97026FC5B" box="[718,846,996,1015]" degrees="91" direction="east" minutes="30" orientation="longitude" pageId="14" pageNumber="123" precision="1" seconds="33.79" value="91.509384">91°3033.79″E</geoCoordinate>
,
<elevation id="34FED1BA8D772858FCB558A970A9FC5B" box="[853,961,996,1015]" metricMagnitude="3" metricUnit="m" metricValue="3.62" pageId="14" pageNumber="123" unit="m" value="3620.0">
<quantity id="782B9B6C8D772858FCB558A970FDFC5B" box="[853,917,996,1015]" metricMagnitude="3" metricUnit="m" metricValue="3.62" pageId="14" pageNumber="123" unit="m" value="3620.0">3620 m</quantity>
a.s.l.
</elevation>
<collectorName id="1226535F8D772858FC2458A97768FC5B" box="[964,1024,996,1015]" pageId="14" pageNumber="123">Jia Yue</collectorName>
&amp;
<collectorName id="1226535F8D772858FBF958A973D5FBBE" pageId="14" pageNumber="123">Jia Fenglong</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D772858FF0A58B27260FB80" collectingDate="2013-08-05" collectionCode="SYSU" collectorName="Jia Yue &amp; Jia Fenglong" elevation="5030" latitude="28.96803" location="Langkazi" longLatPrecision="1" longitude="90.39797" municipality="Shannan" pageId="14" pageNumber="123" specimenCount="1" stateProvince="Qinghai" typeStatus="paratype">
1 J (
<collectionCode id="D9C2AE4C8D772858FEF558B27238FBBE" box="[277,336,1023,1042]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="14" pageNumber="123" type="Herbarium">SYSU</collectionCode>
):
<collectingMunicipality id="5F08ACF38D772858FEBF58B272C3FBBE" box="[351,427,1023,1042]" pageId="14" pageNumber="123">Shannan</collectingMunicipality>
,
<location id="BA0C60528D772858FE5358B2716DFBBE" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FE5358B2716DFBBE" box="[435,517,1023,1042]" latitude="28.96803" longLatPrecision="1" longitude="90.39797" municipality="Shannan" name="Langkazi" pageId="14" pageNumber="123" stateProvince="Qinghai">Langkazi</location>
,
<geoCoordinate id="DAE7504E8D772858FDED58B271ECFBBE" box="[525,644,1023,1042]" degrees="28" direction="north" minutes="58" orientation="latitude" pageId="14" pageNumber="123" precision="1" seconds="4.91" value="28.96803">28°584.91″N</geoCoordinate>
,
<geoCoordinate id="DAE7504E8D772858FD6E58B27065FBBE" box="[654,781,1023,1042]" degrees="90" direction="east" minutes="23" orientation="longitude" pageId="14" pageNumber="123" precision="1" seconds="52.72" value="90.39797">90°2352.72″E</geoCoordinate>
,
<elevation id="34FED1BA8D772858FCF658B270EDFBBE" box="[790,901,1023,1042]" metricMagnitude="3" metricUnit="m" metricValue="5.029999999999999" pageId="14" pageNumber="123" unit="m" value="5030.0">
<quantity id="782B9B6C8D772858FCF658B2703FFBBE" box="[790,855,1023,1042]" metricMagnitude="3" metricUnit="m" metricValue="5.029999999999999" pageId="14" pageNumber="123" unit="m" value="5030.0">5030 m</quantity>
a.s.l.
</elevation>
,
<date id="CB6D10498D772858FC6F58B27085FBBE" box="[911,1005,1023,1042]" pageId="14" pageNumber="123" value="2013-08-05">
<collectingDate id="DB29E9A18D772858FC6F58B27085FBBE" box="[911,1005,1023,1042]" pageId="14" pageNumber="123" value="2013-08-05">5.viii.2013</collectingDate>
</date>
,
<collectorName id="1226535F8D772858FC1658B2775BFBBE" box="[1014,1075,1023,1042]" pageId="14" pageNumber="123">Jia Yue</collectorName>
&amp;
<collectorName id="1226535F8D772858FBAF58B27380FB80" pageId="14" pageNumber="123">Jia Fenglong</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D772858FEF45F547130FBEB" collectingDate="2013-08-03" collectionCode="SYSU" collectorName="Jia Fenglong &amp; Jia Yue" elevation="4447" latitude="28.915985" location="Yangzhuoyongcuo lake" longLatPrecision="1" longitude="90.72477" pageId="14" pageNumber="123" specimenCount="2" specimenCount-female="2" stateProvince="Qinghai" typeStatus="paratype">
2 JJ
<specimenCount id="A9D5FD008D772858FEA85F54721FFB80" box="[328,375,1043,1068]" pageId="14" pageNumber="123" type="female">2 ♀♀</specimenCount>
, 7 unsexed spec. (
<collectionCode id="D9C2AE4C8D772858FDF45F547127FB80" box="[532,591,1049,1068]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="14" pageNumber="123" type="Herbarium">SYSU</collectionCode>
): near
<location id="BA0C60528D772858FD665F547039FB80" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FD665F547039FB80" box="[646,849,1049,1068]" latitude="28.915985" longLatPrecision="1" longitude="90.72477" name="Yangzhuoyongcuo lake" pageId="14" pageNumber="123" stateProvince="Qinghai">Yangzhuoyongcuo lake</location>
,
<geoCoordinate id="DAE7504E8D772858FCB95F5470B2FB80" box="[857,986,1049,1068]" degrees="28" direction="north" minutes="54" orientation="latitude" pageId="14" pageNumber="123" precision="1" seconds="57.55" value="28.915985">28°5457.55″N</geoCoordinate>
,
<geoCoordinate id="DAE7504E8D772858FC035F54770BFB80" box="[995,1123,1049,1068]" degrees="90" direction="east" minutes="43" orientation="longitude" pageId="14" pageNumber="123" precision="1" seconds="29.18" value="90.72477">90°4329.18″E</geoCoordinate>
,
<elevation id="34FED1BA8D772858FF775F79726DFBEB" box="[151,261,1076,1095]" metricMagnitude="3" metricUnit="m" metricValue="4.447" pageId="14" pageNumber="123" unit="m" value="4447.0">
<quantity id="782B9B6C8D772858FF775F7973B0FBEB" box="[151,216,1076,1095]" metricMagnitude="3" metricUnit="m" metricValue="4.447" pageId="14" pageNumber="123" unit="m" value="4447.0">4447 m</quantity>
a.s.l.
</elevation>
,
<date id="CB6D10498D772858FEEE5F797204FBEB" box="[270,364,1076,1095]" pageId="14" pageNumber="123" value="2013-08-03">
<collectingDate id="DB29E9A18D772858FEEE5F797204FBEB" box="[270,364,1076,1095]" pageId="14" pageNumber="123" value="2013-08-03">3.viii.2013</collectingDate>
</date>
,
<collectorName id="1226535F8D772858FE945F79728AFBEB" box="[372,482,1076,1095]" pageId="14" pageNumber="123">Jia Fenglong</collectorName>
&amp;
<collectorName id="1226535F8D772858FE1B5F797150FBEB" box="[507,568,1076,1095]" pageId="14" pageNumber="123">Jia Yue</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D772858FD835F797101FBCE" collectingDate="2013-08-02" collectionCode="SYSU" collectorName="Jia Fenglong &amp; Jia Yue" elevation="4155" latitude="29.88528" location="Gongbujiangda" longLatPrecision="1" longitude="93.24607" pageId="14" pageNumber="123" specimenCount="1" stateProvince="Qinghai" typeStatus="paratype">
1 J (
<collectionCode id="D9C2AE4C8D772858FD6D5F7971A0FBEB" box="[653,712,1076,1095]" country="Taiwan" lsid="urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="14" pageNumber="123" type="Herbarium">SYSU</collectionCode>
):
<location id="BA0C60528D772858FD365F797034FBEB" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FD365F797034FBEB" box="[726,860,1076,1095]" latitude="29.88528" longLatPrecision="1" longitude="93.24607" name="Gongbujiangda" pageId="14" pageNumber="123" stateProvince="Qinghai">Gongbujiangda</location>
,
<geoCoordinate id="DAE7504E8D772858FC845F7970B3FBEB" box="[868,987,1076,1095]" degrees="29" direction="north" minutes="53" orientation="latitude" pageId="14" pageNumber="123" precision="1" seconds="7.01" value="29.88528">29°537.01″N</geoCoordinate>
,
<geoCoordinate id="DAE7504E8D772858FC035F79770BFBEB" box="[995,1123,1076,1095]" degrees="93" direction="east" minutes="14" orientation="longitude" pageId="14" pageNumber="123" precision="1" seconds="45.88" value="93.24607">93°1445.88″E</geoCoordinate>
,
<elevation id="34FED1BA8D772858FF775F027261FBCE" box="[151,265,1103,1122]" metricMagnitude="3" metricUnit="m" metricValue="4.154999999999999" pageId="14" pageNumber="123" unit="m" value="4155.0">
<quantity id="782B9B6C8D772858FF775F0273B2FBCE" box="[151,218,1103,1122]" metricMagnitude="3" metricUnit="m" metricValue="4.154999999999999" pageId="14" pageNumber="123" unit="m" value="4155.0">4155 m</quantity>
a.s.l.
</elevation>
,
<date id="CB6D10498D772858FEF35F02721AFBCE" box="[275,370,1103,1122]" pageId="14" pageNumber="123" value="2013-08-02">
<collectingDate id="DB29E9A18D772858FEF35F02721AFBCE" box="[275,370,1103,1122]" pageId="14" pageNumber="123" value="2013-08-02">2.viii.2013</collectingDate>
</date>
,
<collectorName id="1226535F8D772858FE9C5F027283FBCE" box="[380,491,1103,1122]" pageId="14" pageNumber="123">Jia Fenglong</collectorName>
&amp;
<collectorName id="1226535F8D772858FDE95F02712FFBCE" box="[521,583,1103,1122]" pageId="14" pageNumber="123">Jia Yue</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="0FBB3CD48D772858FD975F027074FBD0" collectingDate="2013-08-02" collectionCode="SYSU" collectorName="Jia Yue" elevation="3872" latitude="29.153214" location="Jiamudui village" longLatPrecision="1" longitude="89.0113" municipality="Rikaze" pageId="14" pageNumber="123" specimenCount="2" specimenCount-female="2" stateProvince="Qinghai" typeStatus="paratype">
4 JJ
<specimenCount id="A9D5FD008D772858FD4E5F027188FBCD" box="[686,736,1096,1122]" pageId="14" pageNumber="123" type="female">2 ♀♀</specimenCount>
15 unsexed spec. (
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):
<collectingMunicipality id="5F08ACF38D772858FC315F027766FBCE" box="[977,1038,1103,1122]" pageId="14" pageNumber="123">Rikaze</collectingMunicipality>
,
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,
<geoCoordinate id="DAE7504E8D772858FF3D5F24723BFBD0" box="[221,339,1129,1148]" degrees="29" direction="north" minutes="9" orientation="latitude" pageId="14" pageNumber="123" precision="1" seconds="11.57" value="29.153214">29°911.57″N</geoCoordinate>
,
<geoCoordinate id="DAE7504E8D772858FEBD5F2472BAFBD0" box="[349,466,1129,1148]" degrees="89" direction="east" minutes="0" orientation="longitude" pageId="14" pageNumber="123" precision="1" seconds="40.67" value="89.0113">89°040.67″E</geoCoordinate>
,
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<quantity id="782B9B6C8D772858FE3C5F247176FBD0" box="[476,542,1129,1148]" metricMagnitude="3" metricUnit="m" metricValue="3.872" pageId="14" pageNumber="123" unit="m" value="3872.0">3872 m</quantity>
a.s.l.
</elevation>
,
<date id="CB6D10498D772858FDB75F2471DEFBD0" box="[599,694,1129,1148]" pageId="14" pageNumber="123" value="2013-08-02">
<collectingDate id="DB29E9A18D772858FDB75F2471DEFBD0" box="[599,694,1129,1148]" pageId="14" pageNumber="123" value="2013-08-02">2.viii.2013</collectingDate>
</date>
,
<collectorName id="1226535F8D772858FD5F5F247195FBD0" box="[703,765,1129,1148]" pageId="14" pageNumber="123">Jia Yue</collectorName>
lgt
</materialsCitation>
;
<materialsCitation id="0FBB3CD48D772858FCC65F2471D4FB3B" collectionCode="NHMW" collectorName="Cigetang &amp; X. Guo" elevation="4400" location="90 km W Amdo" pageId="14" pageNumber="123" specimenCount="3" specimenCount-female="3" stateProvince="Xizang" typeStatus="paratype">
1 J
<specimenCount id="A9D5FD008D772858FCAD5F247016FBD0" box="[845,894,1123,1148]" pageId="14" pageNumber="123" type="female">3 ♀♀</specimenCount>
(
<collectionCode id="D9C2AE4C8D772858FC695F2470BDFBD0" box="[905,981,1129,1148]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="123" type="Museum">NHMW</collectionCode>
): CHINA:
<collectingRegion id="7D17F86B8D772858FBDA5F24770EFBD0" box="[1082,1126,1129,1148]" country="China" name="Xizang" pageId="14" pageNumber="123">Tibet</collectingRegion>
9.VIII. /
<location id="BA0C60528D772858FF035FC97205FB3B" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FF035FC97205FB3B" box="[227,365,1156,1175]" name="90 km W Amdo" pageId="14" pageNumber="123" stateProvince="Xizang">
<quantity id="782B9B6C8D772858FF035FC97272FB3B" box="[227,282,1156,1175]" metricMagnitude="4" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="123" unit="km" value="90.0">90 km</quantity>
W Amdo
</location>
/
<collectorName id="1226535F8D772858FE605FC972A5FB3B" box="[384,461,1156,1175]" pageId="14" pageNumber="123">Cigetang</collectorName>
<quantity id="782B9B6C8D772858FE335FC97167FB3B" box="[467,527,1156,1175]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="14" pageNumber="123" unit="m" value="4400.0">
<elevation id="34FED1BA8D772858FE335FC97167FB3B" box="[467,527,1156,1175]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="14" pageNumber="123" unit="m" value="4400.0">4400m</elevation>
</quantity>
/ leg.
<collectorName id="1226535F8D772858FDA65FC971EEFB3B" box="[582,646,1156,1175]" pageId="14" pageNumber="123">X. Guo</collectorName>
1998
</materialsCitation>
;
<materialsCitation id="0FBB3CD48D772858FD2A5FC97114FB4B" collectionCode="NHMW, IRSNB" collectorName="X. Guo &amp; Indian Tibet &amp; Tzo-Kar &amp; Kar Tso &amp; Startsakpuk Tso &amp; Zaitzev'" country="India" determinerName="A. d'Orchymont" elevation="4400" location="Couna Lake" pageId="14" pageNumber="123" specimenCount="1" specimenCount-female="1" stateProvince="Jammu and Kashmir" typeStatus="paratype">
<specimenCount id="A9D5FD008D772858FD2A5FC97183FB3B" box="[714,747,1150,1175]" pageId="14" pageNumber="123" type="female">1 ♀</specimenCount>
(
<collectionCode id="D9C2AE4C8D772858FD165FC9702AFB3B" box="[758,834,1156,1175]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="123" type="Museum">NHMW</collectionCode>
): CHINA: Tibet 14.VIII. /
<locationDeviation id="AEDE2FAB8D772858FBD05FC97380FB1E" location="Amdo" pageId="14" pageNumber="123">
<quantity id="782B9B6C8D772858FBD05FC9770EFB3B" box="[1072,1126,1156,1175]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="123" unit="km" value="25.0">25 km</quantity>
W Amdo
</locationDeviation>
,
<quantity id="782B9B6C8D772858FF105FD27244FB1E" box="[240,300,1183,1202]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="14" pageNumber="123" unit="m" value="4400.0">
<elevation id="34FED1BA8D772858FF105FD27244FB1E" box="[240,300,1183,1202]" metricMagnitude="3" metricUnit="m" metricValue="4.4" pageId="14" pageNumber="123" unit="m" value="4400.0">4400m</elevation>
</quantity>
/
<location id="BA0C60528D772858FEDD5FD272CEFB1E" LSID="urn:lsid:plazi:treatment:377A879F8D702844FE8058F17181FCB5:BA0C60528D772858FEDD5FD272CEFB1E" box="[317,422,1183,1202]" country="India" name="Couna Lake" pageId="14" pageNumber="123" stateProvince="Jammu and Kashmir">Couna Lake</location>
brook / leg.
<collectorName id="1226535F8D772858FDC55FD2710DFB1E" box="[549,613,1183,1202]" pageId="14" pageNumber="123">X. Guo</collectorName>
1998.
<emphasis id="8DA7EA9B8D772858FD475FD27058FB1C" bold="true" box="[679,816,1183,1202]" pageId="14" pageNumber="123">
<collectingCountry id="C7C476198D772858FD475FD27181FB1E" box="[679,745,1183,1202]" name="India" pageId="14" pageNumber="123">INDIA</collectingCountry>
:
<collectingRegion id="7D17F86B8D772858FD135FD27058FB1C" box="[755,816,1183,1202]" country="India" name="Jammu and Kashmir" pageId="14" pageNumber="123">JAMMU</collectingRegion>
</emphasis>
AND
<emphasis id="8DA7EA9B8D772858FCBC5FD270D9FB1E" bold="true" box="[860,945,1183,1202]" pageId="14" pageNumber="123">KASHMIR:</emphasis>
1 J (
<collectionCode id="D9C2AE4C8D772858FC035FD2774FFB1E" box="[995,1063,1183,1202]" country="Belgium" name="Institut Royal des Sciences Naturelles de Belgique" pageId="14" pageNumber="123">IRSNB</collectionCode>
): male symbol //
<collectorName id="1226535F8D772858FF0C5FF4723BFB60" box="[236,339,1209,1228]" pageId="14" pageNumber="123">Indian Tibet</collectorName>
/
<collectorName id="1226535F8D772858FE835FF472C3FB60" box="[355,427,1209,1228]" pageId="14" pageNumber="123">Tzo-Kar</collectorName>
[=
<collectorName id="1226535F8D772858FE295FF47166FB60" box="[457,526,1209,1228]" pageId="14" pageNumber="123">Kar Tso</collectorName>
] Sta- / ktsak-Puk-tso [=
<collectorName id="1226535F8D772858FD3D5FF4700FFB60" box="[733,871,1209,1228]" pageId="14" pageNumber="123">Startsakpuk Tso</collectorName>
] / pool in swamp / 4-IX, 1932 //
<determinerName id="4AB0DA4D8D772858FF475F99725CFB4B" box="[167,308,1236,1255]" pageId="14" pageNumber="123">A. dOrchymont</determinerName>
det. / H. (
<taxonomicName id="78D34D0A8D772858FE695F997296FB4B" authorityName="Zaitzev" authorityYear="1908" box="[393,510,1236,1255]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="147" phylum="Arthropoda" rank="subGenus" subGenus="Lihelophorus">Lihelophorus</taxonomicName>
) / ser
<collectorName id="1226535F8D772858FDD25F997114FB4B" box="[562,636,1236,1255]" pageId="14" pageNumber="123">Zaitzev</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F7C965028D772844FF775E4D7240FE23" lastPageId="18" lastPageNumber="127" pageId="14" pageNumber="123" type="diagnosis">
<paragraph id="BF6C36898D772858FF775E4D70CCFA3B" blockId="14.[151,1127,1280,1591]" pageId="14" pageNumber="123">
<emphasis id="8DA7EA9B8D772858FF775E4D72F9FABB" bold="true" box="[151,401,1280,1303]" pageId="14" pageNumber="123">Differential diagnosis.</emphasis>
Distinguished from the other two
<taxonomicName id="78D34D0A8D772858FD1A5E4D70E0FABB" authorityName="Zaitzev" authorityYear="1908" box="[762,904,1280,1303]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="147" phylum="Arthropoda" rank="subGenus" subGenus="Lihelophorus">
<emphasis id="8DA7EA9B8D772858FD1A5E4D70E0FABB" box="[762,904,1280,1303]" italics="true" pageId="14" pageNumber="123">Lihelophorus</emphasis>
</taxonomicName>
species (
<taxonomicName id="78D34D0A8D772858FC095E4D770EFABB" authorityName="Zaitzev" authorityYear="1908" box="[1001,1126,1280,1303]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="123" phylum="Arthropoda" rank="species" species="lamicola">
<emphasis id="8DA7EA9B8D772858FC095E4D770EFABB" box="[1001,1126,1280,1303]" italics="true" pageId="14" pageNumber="123">H. lamicola</emphasis>
</taxonomicName>
and
<taxonomicName id="78D34D0A8D772858FF245E6D726CFA9B" authorityName="Zaitzev" authorityYear="1908" box="[196,260,1312,1335]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="123" phylum="Arthropoda" rank="species" species="ser">
<emphasis id="8DA7EA9B8D772858FF245E6D726CFA9B" box="[196,260,1312,1335]" italics="true" pageId="14" pageNumber="123">H. ser</emphasis>
</taxonomicName>
) by the rather wide evenly arched pronotum (width approximately equal to length of mesotarsus + claw) with rounded sides and narrow marginal grooves, rather short metatarsal claws and by the bluntly rounded apices of the parameres. For detailed comparison see
<tableCitation id="F25103328D772858FBF45E2D770BFADB" box="[1044,1123,1376,1399]" captionStart="Table 2" captionStartId="16.[153,200,220,239]" captionText="Table 2. Comparison of adult morphology of Lihelophorus species." pageId="14" pageNumber="123">Table 2</tableCitation>
.
<emphasis id="8DA7EA9B8D772858FF775ECD724AFA3B" bold="true" box="[151,290,1408,1431]" pageId="14" pageNumber="123">Description.</emphasis>
Body length:
<quantity id="782B9B6C8D772858FE5A5ECD7151FA3B" box="[442,569,1408,1431]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.1" metricValueMin="4.6" pageId="14" pageNumber="123" unit="mm" value="4.85" valueMax="5.1" valueMin="4.6">4.65.1 mm</quantity>
in males, 5.0
<quantity id="782B9B6C8D772858FD385ECD704FFA3B" box="[728,807,1408,1431]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="14" pageNumber="123" unit="mm" value="6.1">6.1 mm</quantity>
in females.
</paragraph>
<paragraph id="BF6C36898D772846FF555EED72C3FB1B" blockId="14.[151,1127,1280,1591]" lastBlockId="16.[151,1127,1152,1623]" lastPageId="16" lastPageNumber="125" pageId="14" pageNumber="123">
<emphasis id="8DA7EA9B8D772858FF555EED739FFA1B" box="[181,247,1440,1463]" italics="true" pageId="14" pageNumber="123">Head.</emphasis>
Greenish bronze to maroon bronze, surface with flattened granulation, the granules with large median pits and the grooves between the granules sometimes interrupted resulting in partial coalescence of the granules. Y-groove with the stem at least twice as wide as the arms, widened anteriorly, its floor rugulose. Maxillary palpi yellow, elongate, the apical segment asymmetrical, darkened towards the tip. Antennae with 9 antennomeres, yellow, the clubs a little darker. Maxillary galea angulate apically, with several stout setae (
<figureCitation id="27E82A0C8D692846FC7D5FCD7082FB3B" box="[925,1002,1152,1175]" captionStart="Fig" captionStartId="6.[151,182,1490,1509]" captionTargetBox="[152,1126,220,1478]" captionTargetId="figure-10@6.[151,1127,217,1486]" captionTargetPageId="6" captionText="Fig. 1. Adult morphology of Helophorus (Lihelophorus). ab H. yangae sp. nov. (a ventral view of the head; b detail of mentum and labial palps); cd maxillary galea and lacinia (c H. yangae sp. nov.; d H. ser Zaitzev, 1908); ef mandible (e H. yangae sp. nov.; f H. ser); g labial palpus of H. ser; hl hind legs in dorsal view (h H. yangae sp. nov.; i H. ser; j H. lamicola Zaitzev, 1908; k detail of apex of tibia and base of tarsus of H. lamicola; l detail of tarsomere 5 and claw of H. ser). Abbreviations: sh swimming hairs. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311786" httpUri="https://zenodo.org/record/5311786/files/figure.png" pageId="16" pageNumber="125">Fig. 1c</figureCitation>
). Mandible wide, apex bifid (
<figureCitation id="27E82A0C8D692846FEB15FED72F5FB1B" box="[337,413,1184,1207]" captionStart="Fig" captionStartId="6.[151,182,1490,1509]" captionTargetBox="[152,1126,220,1478]" captionTargetId="figure-10@6.[151,1127,217,1486]" captionTargetPageId="6" captionText="Fig. 1. Adult morphology of Helophorus (Lihelophorus). ab H. yangae sp. nov. (a ventral view of the head; b detail of mentum and labial palps); cd maxillary galea and lacinia (c H. yangae sp. nov.; d H. ser Zaitzev, 1908); ef mandible (e H. yangae sp. nov.; f H. ser); g labial palpus of H. ser; hl hind legs in dorsal view (h H. yangae sp. nov.; i H. ser; j H. lamicola Zaitzev, 1908; k detail of apex of tibia and base of tarsus of H. lamicola; l detail of tarsomere 5 and claw of H. ser). Abbreviations: sh swimming hairs. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311786" httpUri="https://zenodo.org/record/5311786/files/figure.png" pageId="16" pageNumber="125">Fig. 1e</figureCitation>
).
</paragraph>
<caption id="EBAC66018D762859FF915E95774BF9FA" ID-DOI="http://doi.org/10.5281/zenodo.5311798" ID-Zenodo-Dep="5311798" httpUri="https://zenodo.org/record/5311798/files/figure.png" pageId="15" pageNumber="124" startId="15.[113,144,1496,1515]" targetBox="[149,1030,229,1439]" targetPageId="15">
<paragraph id="BF6C36898D762859FF915E95774BF9FA" blockId="15.[113,1090,1496,1622]" pageId="15" pageNumber="124">
Fig. 6.Aedeagophores of
<taxonomicName id="78D34D0A8D762859FEA85E957158FA47" authority="" authorityName="Zaitzev" authorityYear="1908" box="[328,560,1496,1515]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="subGenus" subGenus="Lihelophorus">
<emphasis id="8DA7EA9B8D762859FEA85E9572C5FA47" box="[328,429,1496,1515]" italics="true" pageId="15" pageNumber="124">Helophorus</emphasis>
(
<emphasis id="8DA7EA9B8D762859FE585E957142FA47" box="[440,554,1496,1515]" italics="true" pageId="15" pageNumber="124">Lihelophorus</emphasis>
)
</taxonomicName>
species. ae
<taxonomicName id="78D34D0A8D762859FD4B5E957010FA47" authority="Zaitzev, 1908" authorityName="Zaitzev" authorityYear="1908" box="[683,888,1496,1515]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="species" species="ser" subGenus="Lihelophorus">
<emphasis id="8DA7EA9B8D762859FD4B5E9571A9FA47" box="[683,705,1496,1515]" italics="true" pageId="15" pageNumber="124">H.</emphasis>
(
<emphasis id="8DA7EA9B8D762859FD2B5E9571B5FA47" box="[715,733,1496,1515]" italics="true" pageId="15" pageNumber="124">L.</emphasis>
)
<emphasis id="8DA7EA9B8D762859FD075E957069FA47" box="[743,769,1496,1515]" italics="true" pageId="15" pageNumber="124">ser</emphasis>
<bibRefCitation id="DB424B788D762859FCE55E957010FA47" author="ZAITZEV F. A." box="[773,888,1496,1515]" pageId="15" pageNumber="124" pagination="417 - 426" refId="ref20914" refString="ZAITZEV F. A. 1908: Beitrag zur Kenntniss der Waserkafer von Chinesisch-Centralasien. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St. Petersbourg 13: 417 - 426." type="journal article" year="1908">Zaitzev, 1908</bibRefCitation>
</taxonomicName>
(a lectotype, Qinghai: valley of Alag Hu lake; bc Xizang: valley SE of Dong La pass; d Qinghai: Golo, 20 km E of Maduo; e Qinghai: Zuimatan). fj
<taxonomicName id="78D34D0A8D762859FED05D43715AF98D" authority="Zaitzev, 1908" authorityName="Zaitzev" authorityYear="1908" box="[304,562,1550,1569]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="species" species="lamicola" subGenus="Lihelophorus">
<emphasis id="8DA7EA9B8D762859FED05D43722FF98D" box="[304,327,1550,1569]" italics="true" pageId="15" pageNumber="124">H.</emphasis>
(
<emphasis id="8DA7EA9B8D762859FEB15D43720CF98D" box="[337,356,1550,1569]" italics="true" pageId="15" pageNumber="124">L.</emphasis>
)
<emphasis id="8DA7EA9B8D762859FE8F5D4372D2F98D" box="[367,442,1550,1569]" italics="true" pageId="15" pageNumber="124">lamicola</emphasis>
<bibRefCitation id="DB424B788D762859FE5E5D43715AF98D" author="ZAITZEV F. A." box="[446,562,1550,1569]" pageId="15" pageNumber="124" pagination="417 - 426" refId="ref20914" refString="ZAITZEV F. A. 1908: Beitrag zur Kenntniss der Waserkafer von Chinesisch-Centralasien. Annuaire du Musee Zoologique de l'Academie Imperiale des Sciences de St. Petersbourg 13: 417 - 426." type="journal article" year="1908">Zaitzev, 1908</bibRefCitation>
</taxonomicName>
(f lectotype, Qinghai: Djarin Nor; gi Qinghai: Zuimatan; j paralectotype, Xizang: S of Arka-tagh, 5073 m a.s.l., lgt. S. Hedin). ko
<taxonomicName id="78D34D0A8D762859FCE35D657017F997" authority="Angus &amp; Jia &amp; Chen &amp; Zhang &amp; Vondráček &amp; Fikáček &amp; Sw, 2016" authorityName="Angus &amp; Jia &amp; Chen &amp; Zhang &amp; Vondráček &amp; Fikáček &amp; Sw" authorityYear="2016" box="[771,895,1576,1595]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="124" phylum="Arthropoda" rank="species" species="yangae" status="sp. nov." subGenus="Lihelophorus">
<emphasis id="8DA7EA9B8D762859FCE35D657071F997" box="[771,793,1576,1595]" italics="true" pageId="15" pageNumber="124">H.</emphasis>
(
<emphasis id="8DA7EA9B8D762859FCC45D65705EF997" box="[804,822,1576,1595]" italics="true" pageId="15" pageNumber="124">L.</emphasis>
)
<emphasis id="8DA7EA9B8D762859FCA15D657017F997" box="[833,895,1576,1595]" italics="true" pageId="15" pageNumber="124">yangae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="969457E08D762859FC645D6570AEF997" box="[900,966,1576,1595]" pageId="15" pageNumber="124" rank="species">sp. nov.</taxonomicNameLabel>
(k holotype, Qinghai: Zuimatan; lm paratypes, same locality; n Qinghai: Golo, Yematan; o Xizang: Gongbujiangda).
</paragraph>
</caption>
<caption id="EBAC66018D692846FF795B9171BEFF43" box="[153,726,220,239]" pageId="16" pageNumber="125" startId="16.[153,200,220,239]">
<paragraph id="BF6C36898D692846FF795B9171BEFF43" blockId="16.[153,726,220,239]" box="[153,726,220,239]" pageId="16" pageNumber="125">
Table 2. Comparison of adult morphology of
<taxonomicName id="78D34D0A8D692846FDFA5B9171E4FF43" authorityName="Zaitzev" authorityYear="1908" box="[538,652,220,239]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="147" phylum="Arthropoda" rank="subGenus" subGenus="Lihelophorus">
<emphasis id="8DA7EA9B8D692846FDFA5B9171E4FF43" box="[538,652,220,239]" italics="true" pageId="16" pageNumber="125">Lihelophorus</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<paragraph id="BF6C36898D692846FF7B5A4870F7FBE2" pageId="16" pageNumber="125">
<table id="CDD3C4298D69D7A9FF7B5A487735FBE2" box="[155,1117,261,1102]" gridcols="4" gridrows="16" pageId="16" pageNumber="125">
<tr id="01E334CB8D69D7A9FF7B5A487735FEB4" box="[155,1117,261,280]" gridrow="0" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B5A48721AFEB4" box="[155,370,261,280]" gridcol="0" gridrow="0" pageId="16" pageNumber="125">
<emphasis id="8DA7EA9B8D692846FF7B5A487391FEB4" bold="true" box="[155,249,261,280]" pageId="16" pageNumber="125">Character</emphasis>
</th>
<th id="42325DB78D69D7A9FE615A487101FEB4" box="[385,617,261,280]" gridcol="1" gridrow="0" pageId="16" pageNumber="125">
<taxonomicName id="78D34D0A8D692846FE615A487280FEB4" authorityName="Zaitzev" authorityYear="1908" box="[385,488,261,280]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="125" phylum="Arthropoda" rank="species" species="lamicola">
<emphasis id="8DA7EA9B8D692846FE615A487280FEB4" bold="true" box="[385,488,261,280]" italics="true" pageId="16" pageNumber="125">H. lamicola</emphasis>
</taxonomicName>
</th>
<th id="42325DB78D69D7A9FD9A5A487009FEB4" box="[634,865,261,280]" gridcol="2" gridrow="0" pageId="16" pageNumber="125">
<taxonomicName id="78D34D0A8D692846FD9A5A4871D8FEB4" authorityName="Zaitzev" authorityYear="1908" box="[634,688,261,280]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="125" phylum="Arthropoda" rank="species" species="ser">
<emphasis id="8DA7EA9B8D692846FD9A5A4871D8FEB4" bold="true" box="[634,688,261,280]" italics="true" pageId="16" pageNumber="125">H. ser</emphasis>
</taxonomicName>
</th>
<th id="42325DB78D69D7A9FC945A487735FEB4" box="[884,1117,261,280]" gridcol="3" gridrow="0" pageId="16" pageNumber="125">
<emphasis id="8DA7EA9B8D692846FC945A487770FEB4" bold="true" box="[884,1048,261,280]" pageId="16" pageNumber="125">
<taxonomicName id="78D34D0A8D692846FC945A4870A6FEB4" authority="Angus &amp; Jia &amp; Chen &amp; Zhang &amp; Vondráček &amp; Fikáček &amp; Sw, 2016" authorityName="Angus &amp; Jia &amp; Chen &amp; Zhang &amp; Vondráček &amp; Fikáček &amp; Sw" authorityYear="2016" box="[884,974,261,280]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="125" phylum="Arthropoda" rank="species" species="yangae" status="sp. nov.">
<emphasis id="8DA7EA9B8D692846FC945A4870A6FEB4" bold="true" box="[884,974,261,280]" italics="true" pageId="16" pageNumber="125">H. yangae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="969457E08D692846FC335A487770FEB4" box="[979,1048,261,280]" pageId="16" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="01E334CB8D69D7A9FF7B5A6E7735FEC0" box="[155,1117,291,364]" gridrow="1" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B5A6E721AFEC0" box="[155,370,291,364]" gridcol="0" gridrow="1" pageId="16" pageNumber="125">Mandibular apex</th>
<td id="42325DB78D69D7A9FE615A6E7101FEC0" box="[385,617,291,364]" gridcol="1" gridrow="1" pageId="16" pageNumber="125">simple (as in Fig. 1f)</td>
<td id="42325DB78D69D7A9FD9A5A6E7009FEC0" box="[634,865,291,364]" gridcol="2" gridrow="1" pageId="16" pageNumber="125">simple (Fig. 1f)</td>
<td id="42325DB78D69D7A9FC945A6E7735FEC0" box="[884,1117,291,364]" gridcol="3" gridrow="1" pageId="16" pageNumber="125">bifid (Fig. 1e; rare specimens with simple apex exist)</td>
</tr>
<tr id="01E334CB8D69D7A9FF7B5A3A7735FE08" box="[155,1117,375,420]" gridrow="2" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B5A3A721AFE08" box="[155,370,375,420]" gridcol="0" gridrow="2" pageId="16" pageNumber="125">Lacinia</th>
<td id="42325DB78D69D7A9FE615A3A7101FE08" box="[385,617,375,420]" gridcol="1" gridrow="2" pageId="16" pageNumber="125">simple, with trichoid setae (as in Fig. 1d)</td>
<td id="42325DB78D69D7A9FD9A5A3A7009FE08" box="[634,865,375,420]" gridcol="2" gridrow="2" pageId="16" pageNumber="125">simple, with trichoid setae (Fig. 1d)</td>
<td id="42325DB78D69D7A9FC945A3A7735FE08" box="[884,1117,375,420]" gridcol="3" gridrow="2" pageId="16" pageNumber="125">bilobate, with stout setae distally (Fig. 1c)</td>
</tr>
<tr id="01E334CB8D69D7A9FF7B5AE27735FE6E" box="[155,1117,431,450]" gridrow="3" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B5AE2721AFE6E" box="[155,370,431,450]" gridcol="0" gridrow="3" pageId="16" pageNumber="125">Pronotum shape</th>
<td id="42325DB78D69D7A9FE615AE27101FE6E" box="[385,617,431,450]" gridcol="1" gridrow="3" pageId="16" pageNumber="125">flat</td>
<td id="42325DB78D69D7A9FD9A5AE27009FE6E" box="[634,865,431,450]" gridcol="2" gridrow="3" pageId="16" pageNumber="125">weakly arched</td>
<td id="42325DB78D69D7A9FC945AE27735FE6E" box="[884,1117,431,450]" gridcol="3" gridrow="3" pageId="16" pageNumber="125">highly and evenly arched</td>
</tr>
<tr id="01E334CB8D69D7A9FF7B5A807735FE57" box="[155,1117,461,507]" gridrow="4" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B5A80721AFE57" box="[155,370,461,507]" gridcol="0" gridrow="4" pageId="16" pageNumber="125">Pronotal grooves</th>
<td id="42325DB78D69D7A9FE615A807101FE57" box="[385,617,461,507]" gridcol="1" gridrow="4" pageId="16" pageNumber="125">largely reduced (Figs 2e, 4ac)</td>
<td id="42325DB78D69D7A9FD9A5A807009FE57" box="[634,865,461,507]" gridcol="2" gridrow="4" pageId="16" pageNumber="125">well developed (Figs 2c, 4dh)</td>
<td id="42325DB78D69D7A9FC945A807735FE57" box="[884,1117,461,507]" gridcol="3" gridrow="4" pageId="16" pageNumber="125">well developed (Figs 2a, 5)</td>
</tr>
<tr id="01E334CB8D69D7A9FF7B594B7735FDE2" box="[155,1117,518,590]" gridrow="5" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B594B721AFDE2" box="[155,370,518,590]" gridcol="0" gridrow="5" pageId="16" pageNumber="125">Lateral margins of pronotum</th>
<td id="42325DB78D69D7A9FE61594B7101FDE2" box="[385,617,518,590]" gridcol="1" gridrow="5" pageId="16" pageNumber="125">very weakly arcuate (Figs 4ac)</td>
<td id="42325DB78D69D7A9FD9A594B7009FDE2" box="[634,865,518,590]" gridcol="2" gridrow="5" pageId="16" pageNumber="125">strongly arcuate, strongly narrowed anteriorly (Figs 4dh)</td>
<td id="42325DB78D69D7A9FC94594B7735FDE2" box="[884,1117,518,590]" gridcol="3" gridrow="5" pageId="16" pageNumber="125">evenly rounded (Fig. 5)</td>
</tr>
<tr id="01E334CB8D69D7A9FF7B59147735FD2B" box="[155,1117,601,647]" gridrow="6" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B5914721AFD2B" box="[155,370,601,647]" gridcol="0" gridrow="6" pageId="16" pageNumber="125">Basal pronotal margin</th>
<td id="42325DB78D69D7A9FE6159147101FD2B" box="[385,617,601,647]" gridcol="1" gridrow="6" pageId="16" pageNumber="125">narrower than elytral bases combined (Figs 3ac)</td>
<td id="42325DB78D69D7A9FD9A59147009FD2B" box="[634,865,601,647]" gridcol="2" gridrow="6" pageId="16" pageNumber="125">narrower than elytral bases combined (Figs 3eg)</td>
<td id="42325DB78D69D7A9FC9459147735FD2B" box="[884,1117,601,647]" gridcol="3" gridrow="6" pageId="16" pageNumber="125">ca. as wide as elytral bases combined (Figs 3hk)</td>
</tr>
<tr id="01E334CB8D69D7A9FF7B59DF7735FD13" box="[155,1117,658,703]" gridrow="7" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B59DF721AFD13" box="[155,370,658,703]" gridcol="0" gridrow="7" pageId="16" pageNumber="125">Strial punctures of elytra</th>
<td id="42325DB78D69D7A9FE6159DF7101FD13" box="[385,617,658,703]" gridcol="1" gridrow="7" pageId="16" pageNumber="125">minute, scarcely impressed (Fig. 2b)</td>
<td id="42325DB78D69D7A9FD9A59DF7009FD13" box="[634,865,658,703]" gridcol="2" gridrow="7" pageId="16" pageNumber="125">coarse, well impressed (Fig. 2d)</td>
<td id="42325DB78D69D7A9FC9459DF7735FD13" box="[884,1117,658,703]" gridcol="3" gridrow="7" pageId="16" pageNumber="125">minute, scarcely impressed (Fig. 2f)</td>
</tr>
<tr id="01E334CB8D69D7A9FF7B59877735FD54" box="[155,1117,714,760]" gridrow="8" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B5987721AFD54" box="[155,370,714,760]" gridcol="0" gridrow="8" pageId="16" pageNumber="125">Interval punctures</th>
<td id="42325DB78D69D7A9FE6159877101FD54" box="[385,617,714,760]" gridcol="1" gridrow="8" pageId="16" pageNumber="125">ca. as large as strial punctures</td>
<td id="42325DB78D69D7A9FD9A59877009FD54" box="[634,865,714,760]" gridcol="2" gridrow="8" pageId="16" pageNumber="125">much smaller than strial punctres</td>
<td id="42325DB78D69D7A9FC9459877735FD54" box="[884,1117,714,760]" gridcol="3" gridrow="8" pageId="16" pageNumber="125">ca. large as strial punctures</td>
</tr>
<tr id="01E334CB8D69D7A9FF7B584E7735FC9C" box="[155,1117,771,816]" gridrow="9" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B584E721AFC9C" box="[155,370,771,816]" gridcol="0" gridrow="9" pageId="16" pageNumber="125">Punctures of scutellary stria</th>
<td id="42325DB78D69D7A9FE61584E7101FC9C" box="[385,617,771,816]" gridcol="1" gridrow="9" pageId="16" pageNumber="125">79, often become indis- tinct</td>
<td id="42325DB78D69D7A9FD9A584E7009FC9C" box="[634,865,771,816]" gridcol="2" gridrow="9" pageId="16" pageNumber="125">47, coarse and always distinct</td>
<td id="42325DB78D69D7A9FC94584E7735FC9C" box="[884,1117,771,816]" gridcol="3" gridrow="9" pageId="16" pageNumber="125">68, always distinct</td>
</tr>
<tr id="01E334CB8D69D7A9FF7B58767735FCE2" box="[155,1117,827,846]" gridrow="10" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B5876721AFCE2" box="[155,370,827,846]" gridcol="0" gridrow="10" pageId="16" pageNumber="125">Legs including claws</th>
<td id="42325DB78D69D7A9FE6158767101FCE2" box="[385,617,827,846]" gridcol="1" gridrow="10" pageId="16" pageNumber="125">conspicuously long</td>
<td id="42325DB78D69D7A9FD9A58767009FCE2" box="[634,865,827,846]" gridcol="2" gridrow="10" pageId="16" pageNumber="125">conspicuously long</td>
<td id="42325DB78D69D7A9FC9458767735FCE2" box="[884,1117,827,846]" gridcol="3" gridrow="10" pageId="16" pageNumber="125">less obviously long</td>
</tr>
<tr id="01E334CB8D69D7A9FF7B58147735FC0E" box="[155,1117,857,930]" gridrow="11" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B5814721AFC0E" box="[155,370,857,930]" gridcol="0" gridrow="11" pageId="16" pageNumber="125">Metarsomere 5</th>
<td id="42325DB78D69D7A9FE6158147101FC0E" box="[385,617,857,930]" gridcol="1" gridrow="11" pageId="16" pageNumber="125">ca. as long as metatar- someres 34 combined (sometimes longer)</td>
<td id="42325DB78D69D7A9FD9A58147009FC0E" box="[634,865,857,930]" gridcol="2" gridrow="11" pageId="16" pageNumber="125">much longer than meta- tarsomeres 34 combined (sometimes as long as)</td>
<td id="42325DB78D69D7A9FC9458147735FC0E" box="[884,1117,857,930]" gridcol="3" gridrow="11" pageId="16" pageNumber="125">slightly longer or as long as metatarsomeres 34 combined</td>
</tr>
<tr id="01E334CB8D69D7A9FF7B58E17735FC76" box="[155,1117,940,986]" gridrow="12" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B58E1721AFC76" box="[155,370,940,986]" gridcol="0" gridrow="12" pageId="16" pageNumber="125">Metatarsomere / tarsal claw</th>
<td id="42325DB78D69D7A9FE6158E17101FC76" box="[385,617,940,986]" gridcol="1" gridrow="12" pageId="16" pageNumber="125">1.5</td>
<td id="42325DB78D69D7A9FD9A58E17009FC76" box="[634,865,940,986]" gridcol="2" gridrow="12" pageId="16" pageNumber="125">1.7</td>
<td id="42325DB78D69D7A9FC9458E17735FC76" box="[884,1117,940,986]" gridcol="3" gridrow="12" pageId="16" pageNumber="125">1.9</td>
</tr>
<tr id="01E334CB8D69D7A9FF7B58A87735FC54" box="[155,1117,997,1016]" gridrow="13" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B58A8721AFC54" box="[155,370,997,1016]" gridcol="0" gridrow="13" pageId="16" pageNumber="125">Apex of parameres</th>
<td id="42325DB78D69D7A9FE6158A87101FC54" box="[385,617,997,1016]" gridcol="1" gridrow="13" pageId="16" pageNumber="125">pointed</td>
<td id="42325DB78D69D7A9FD9A58A87009FC54" box="[634,865,997,1016]" gridcol="2" gridrow="13" pageId="16" pageNumber="125">pointed</td>
<td id="42325DB78D69D7A9FC9458A87735FC54" box="[884,1117,997,1016]" gridcol="3" gridrow="13" pageId="16" pageNumber="125">blunt</td>
</tr>
<tr id="01E334CB8D69D7A9FF7B5F4E7735FB9D" box="[155,1117,1027,1073]" gridrow="14" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B5F4E721AFB9D" box="[155,370,1027,1073]" gridcol="0" gridrow="14" pageId="16" pageNumber="125">Apical portion of median lobe</th>
<td id="42325DB78D69D7A9FE615F4E7101FB9D" box="[385,617,1027,1073]" gridcol="1" gridrow="14" pageId="16" pageNumber="125">equilateral triangle</td>
<td id="42325DB78D69D7A9FD9A5F4E7009FB9D" box="[634,865,1027,1073]" gridcol="2" gridrow="14" pageId="16" pageNumber="125">isosceles triangle</td>
<td id="42325DB78D69D7A9FC945F4E7735FB9D" box="[884,1117,1027,1073]" gridcol="3" gridrow="14" pageId="16" pageNumber="125">isosceles to almost equilat- eral triangle</td>
</tr>
<tr id="01E334CB8D69D7A9FF7B5F767735FBE2" box="[155,1117,1083,1102]" gridrow="15" pageId="16" pageNumber="125">
<th id="42325DB78D69D7A9FF7B5F76721AFBE2" box="[155,370,1083,1102]" gridcol="0" gridrow="15" pageId="16" pageNumber="125">Phallobase</th>
<td id="42325DB78D69D7A9FE615F767101FBE2" box="[385,617,1083,1102]" gridcol="1" gridrow="15" pageId="16" pageNumber="125">short</td>
<td id="42325DB78D69D7A9FD9A5F767009FBE2" box="[634,865,1083,1102]" gridcol="2" gridrow="15" pageId="16" pageNumber="125">short to very long</td>
<td id="42325DB78D69D7A9FC945F767735FBE2" box="[884,1117,1083,1102]" gridcol="3" gridrow="15" pageId="16" pageNumber="125">short</td>
</tr>
</table>
</paragraph>
<paragraph id="BF6C36898D692846FF555F8D712EFADB" blockId="16.[151,1127,1152,1623]" pageId="16" pageNumber="125">
<emphasis id="8DA7EA9B8D692846FF555F8D7276FB7B" box="[181,286,1216,1239]" italics="true" pageId="16" pageNumber="125">Pronotum</emphasis>
. Yellow, grooves sometimes slightly darker. Moderately and very evenly arched, widest medially, the sides evenly curved. Grooves narrow and shallow, sometimes weaker over anterior fifth of the pronotum; submedian grooves weakly angled outwards medially, occasionally interrupted in the region of the angle; marginal grooves rather indistinct, not obviously widened medially. Intervals punctate, sometimes very finely so; surface between the punctures either smooth or rugulose.
</paragraph>
<paragraph id="BF6C36898D692846FF555ECD7257F9BB" blockId="16.[151,1127,1152,1623]" pageId="16" pageNumber="125">
<emphasis id="8DA7EA9B8D692846FF555ECD7390FA3B" box="[181,248,1408,1431]" italics="true" pageId="16" pageNumber="125">Elytra</emphasis>
. Yellow to dirty yellow with vague darker mottling, the brownish sutural Λ- mark absent or distinct. Striae present as rows of punctures, very weakly impressed below the level of the interstices. All interstices, including the outermost one, completely flat. Scutellary stria with 58 punctures. Elytra widest just behind middle, tapering to blunt apex, the sides gently rounded.
</paragraph>
<paragraph id="BF6C36898D692846FF555D6D770FF9FB" blockId="16.[151,1127,1152,1623]" pageId="16" pageNumber="125">
<emphasis id="8DA7EA9B8D692846FF555D6D7380F99B" box="[181,232,1568,1591]" italics="true" pageId="16" pageNumber="125">Legs</emphasis>
. Yellow with apical tarsal segment dark brown apically, and claws brown. Legs rather long, with long swimming-hairs on meso- and metatibiae and meso- and metatarsi;
</paragraph>
<paragraph id="BF6C36898D6B2844FF775B957129FEA3" blockId="18.[151,1127,216,719]" pageId="18" pageNumber="127">metatarsomere 5 distinctly ca. as long as metatarsomeres 34 combined; metatarsal claw ca. 0.4× as long as ultimate metatarsomere.</paragraph>
<paragraph id="BF6C36898D6B2844FF555A557240FE23" blockId="18.[151,1127,216,719]" pageId="18" pageNumber="127">
<emphasis id="8DA7EA9B8D6B2844FF555A557274FE83" box="[181,284,280,303]" italics="true" pageId="18" pageNumber="127">Aedeagus</emphasis>
. Parameres bluntly rounded apically, weakly sinuate on outer margin. Phallobase ca. as long as parameres. Median lobe wide, its distal portion short (in shape of more less isosceles triangle); base of median lobe (between the struts) straight, with median backwardly directed spur.
</paragraph>
</subSubSection>
<subSubSection id="F7C965028D6B2844FF775AD57737FDC3" pageId="18" pageNumber="127" type="description">
<paragraph id="BF6C36898D6B2844FF775AD57737FDC3" blockId="18.[151,1127,216,719]" pageId="18" pageNumber="127">
<emphasis id="8DA7EA9B8D6B2844FF775AD5726EFE03" bold="true" box="[151,262,408,431]" pageId="18" pageNumber="127">Variation.</emphasis>
The species varies slightly in size and body proportions, both smaller narrower (
<figureCitation id="27E82A0C8D6B2844FBDC5AD573C5FE63" captionStart="Fig" captionStartId="10.[151,182,1482,1501]" captionTargetBox="[159,1124,234,1349]" captionTargetId="figure-10@10.[151,1127,219,1365]" captionTargetPageId="10" captionText="Fig. 3. Habitus of the species of Helophorus (Lihelophorus). ac H. lamicola Zaitzev, 1908 (a Qinghai: Zhaling Hu (lectotype); bc Quinghai: Golo).dg H.ser Zaitzev, 1908 (de Qinghai:Golo; f Xizang, Nyainqentanglha Feng; g Gansu: Xiahe env.). hk H. yangae sp. nov. (h Xizang: 90 km W of Amdo; i Xizang: 25 km W of Amdo; j Qinghai: Golo; k India: Kar Tso (Orchymont coll.)). All specimens to scale." figureDoi="http://doi.org/10.5281/zenodo.5311792" httpUri="https://zenodo.org/record/5311792/files/figure.png" pageId="18" pageNumber="127">Fig. 3j</figureCitation>
) and larger wider (
<figureCitation id="27E82A0C8D6B2844FE945AF572D2FE63" box="[372,442,440,463]" captionStart="Fig" captionStartId="10.[151,182,1482,1501]" captionTargetBox="[159,1124,234,1349]" captionTargetId="figure-10@10.[151,1127,219,1365]" captionTargetPageId="10" captionText="Fig. 3. Habitus of the species of Helophorus (Lihelophorus). ac H. lamicola Zaitzev, 1908 (a Qinghai: Zhaling Hu (lectotype); bc Quinghai: Golo).dg H.ser Zaitzev, 1908 (de Qinghai:Golo; f Xizang, Nyainqentanglha Feng; g Gansu: Xiahe env.). hk H. yangae sp. nov. (h Xizang: 90 km W of Amdo; i Xizang: 25 km W of Amdo; j Qinghai: Golo; k India: Kar Tso (Orchymont coll.)). All specimens to scale." figureDoi="http://doi.org/10.5281/zenodo.5311792" httpUri="https://zenodo.org/record/5311792/files/figure.png" pageId="18" pageNumber="127">Fig. 3i</figureCitation>
) are known. Coloration varies slightly as well, both in coloration of the head (black or with weak greenish to purple sheen) and the elytra (from yellowish to brown, sutural Λ- mark present or absent). Pronotum varies slightly in the extent of the impression of the grooves as well as in their shape, in rare specimens the submedian grooves may be even Y-shaped anteriorly (
<figureCitation id="27E82A0C8D6B2844FDEE59757108FDE3" box="[526,608,568,591]" captionStart="Fig" captionStartId="12.[151,182,1058,1077]" captionTargetBox="[164,1118,238,1008]" captionTargetId="figure-10@12.[151,1129,219,1044]" captionTargetPageId="12" captionText="Fig. 5. Details of the head and pronotum of the Helophorus (Lihelophorus) yangae sp. nov. (a Xizang: 90 km W of Amdo; b,d Qinghai: Golo; c Xizang: 25 km W of Amdo; e India: Kar Tso (Orchymont coll.). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5311796" httpUri="https://zenodo.org/record/5311796/files/figure.png" pageId="18" pageNumber="127">Fig. 5d</figureCitation>
). Aedeagus varies slightly in the proportions of the median lobe, but otherwise its morphology is very constant in all specimens examined.
</paragraph>
</subSubSection>
<subSubSection id="F7C965028D6B2844FF7759357099FD23" box="[151,1009,632,655]" pageId="18" pageNumber="127" type="etymology">
<paragraph id="BF6C36898D6B2844FF7759357099FD23" blockId="18.[151,1127,216,719]" box="[151,1009,632,655]" pageId="18" pageNumber="127">
<emphasis id="8DA7EA9B8D6B2844FF7759357270FD23" bold="true" box="[151,280,632,655]" pageId="18" pageNumber="127">Etymology.</emphasis>
This species is named after Dr Xiaomei Yang, wife of Fenglong Jia.
</paragraph>
</subSubSection>
<subSubSection id="F7C965028D6B2844FF7759D57181FCB5" pageId="18" pageNumber="127" type="distribution">
<paragraph id="BF6C36898D6B2844FF7759D57105FD63" blockId="18.[151,1127,216,719]" pageId="18" pageNumber="127">
<emphasis id="8DA7EA9B8D6B2844FF7759D5724AFD03" bold="true" box="[151,290,664,687]" pageId="18" pageNumber="127">Distribution</emphasis>
(Fig. 7a). Widely distributed on the Tibetan Plateau, at altitudes ranging from
<quantity id="782B9B6C8D6B2844FF7759F57380FD63" box="[151,232,696,719]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="18" pageNumber="127" unit="m" value="3300.0">3300 m</quantity>
(Gangca) to ca.
<quantity id="782B9B6C8D6B2844FE7959F57282FD63" box="[409,490,696,719]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="127" unit="m" value="5000.0">5000 m</quantity>
(Langkazi).
</paragraph>
<paragraph id="BF6C36898D6B2844FDF559B27181FCB5" blockId="18.[533,745,767,793]" box="[533,745,767,793]" pageId="18" pageNumber="127">
<emphasis id="8DA7EA9B8D6B2844FDF559B27181FCB5" bold="true" box="[533,745,767,793]" pageId="18" pageNumber="127">Immature stages</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>