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<document id="9FD23803774666A14A86DFAE54E656D2" ID-DOI="10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4900311" IM.bibliography_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1622827801739" checkinUser="carolina" docAuthor="Morón, Miguel Angel &amp; Solís, Angel" docDate="2001" docId="03A887FE5D4C15242D0AD03CFC1BFCFD" docLanguage="en" docName="TheColeopBull.55.1.11-29.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (1)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Phyllophaga (Phyllophaga) talamancana Moron and Solis, new species" docType="treatment" docVersion="1" lastPageNumber="24" masterDocId="FF91FF865D4715292C3FD44FFF9EFFD8" masterDocTitle="Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)" masterLastPageNumber="29" masterPageNumber="11" pageNumber="22" updateTime="1730139982923" updateUser="juliana" zenodo-license-document="CLOSED">
<mods:mods id="559ACA7FA70FAFD42EAA8C8F87DF3A45" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8ACF0D118B15F00F9510DB63A9A12E35">
<mods:title id="2213881540FEC2B7EA8AD12F74BA8B19">Seven New Species Of Phyllophaga (S. Str.) Harris From Costa Rica (Coleoptera: Melolonthidae: Melolonthinae)</mods:title>
</mods:titleInfo>
<mods:name id="14F5994351B5E8B8B0AC4182A2F08F23" type="personal">
<mods:role id="9EC9F5E298BAC700E5174A0829C9197B">
<mods:roleTerm id="A284940A7BB9E5371F2849CE0277D281">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B2234BA5603B150C224280DC0707CB1">Morón, Miguel Angel</mods:namePart>
<mods:affiliation id="0D617EC914BD0CAD7D6998ADA7222E3A">Departamento de Entomología Instituto de Ecología, A. C. (sector SEP­CONACYT) Apartado Postal 63 Xalapa, Veracruz 91000, MÉXICO</mods:affiliation>
</mods:name>
<mods:name id="40E4D8EC01EE30005730EED4C48B7F59" type="personal">
<mods:role id="FDF4F75AA5B2432DA9DCF825D3944E06">
<mods:roleTerm id="6C875FC6763D9EC9CEF94DA34E8C9393">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7837FCBEC8810CF5AE5A81362723ED35">Solís, Angel</mods:namePart>
<mods:affiliation id="8C4737AA148D87C20AFA91610EA2E4AE">Departamento de Entomología Instituto Nacional de Biodiversidad (INBio) Apartado Postal 22 ­ 3100 Santo Domingo de Heredia, COSTA RICA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="11FFF6E92A06E35C328A169D166FBA7F">text</mods:typeOfResource>
<mods:relatedItem id="7ADE4BD762558A93A7E1B067B5F46F5C" type="host">
<mods:titleInfo id="877792B93757F4967BA851F9F0890ED6">
<mods:title id="0AFC44CDE6BDC2F349F831386AF7C963">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="A2CBC30DF363872B3ED04453BDAE4041">
<mods:date id="B871EE556745076916866D1A35D181DD">2001</mods:date>
<mods:detail id="AC0D7800B1E67A3CCAB1AB0C849999B6" type="pubDate">
<mods:number id="34037430E04299507F1AEF9A91D442EF">2001-03-01</mods:number>
</mods:detail>
<mods:detail id="D64DDD4BB4AE9B8C4DBA7A2EC36340F8" type="volume">
<mods:number id="DF05666CD6311F1EC3D5F3E8910C1E0E">55</mods:number>
</mods:detail>
<mods:detail id="5EDE30C9E83BCC463DB7DD2028FE937A" type="issue">
<mods:number id="8B9AB655DCD701F5D1A241243EE0ABC5">1</mods:number>
</mods:detail>
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<mods:start id="16A81D157141876F6161E6872EC8E7FC">11</mods:start>
<mods:end id="36CAEC7D57E9CDDC6087A9E3DD84BD01">29</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B08C9CEC838940B67D0D98726419F3C9">journal article</mods:classification>
<mods:identifier id="F0A11BAB9E768FFC80BE532E4FD89FA0" type="DOI">10.1649/0010-065x(2001)055[0011:snsops]2.0.co;2</mods:identifier>
<mods:identifier id="E9740E164D7EA7D9CFCFAF5FFFDE25BE" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="CBC02CFCCF2044FF7221DD63B6CE4F99" type="Zenodo-Dep">4900311</mods:identifier>
</mods:mods>
<treatment id="03A887FE5D4C15242D0AD03CFC1BFCFD" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD" httpUri="http://treatment.plazi.org/id/03A887FE5D4C15242D0AD03CFC1BFCFD" lastPageId="13" lastPageNumber="24" pageId="11" pageNumber="22">
<subSubSection id="C31B65635D4C15222D0AD03CFBA4FB5F" box="[309,1082,1138,1160]" pageId="11" pageNumber="22" type="nomenclature">
<paragraph id="8BBE36E85D4C15222D0AD03CFBA4FB5F" blockId="11.[309,1082,1138,1186]" box="[309,1082,1138,1160]" pageId="11" pageNumber="22">
<taxonomicName id="4C014D6B5D4C15222D0AD03CFC35FB50" authority="Moron and Solis" authorityName="Moron and Solis" box="[309,939,1139,1160]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="species" species="talamancana" status="sp. nov." subGenus="Phyllophaga">
<emphasis id="B975EAFA5D4C15222D0AD03CFE25FB50" box="[309,443,1139,1160]" italics="true" pageId="11" pageNumber="22">Phyllophaga</emphasis>
(
<emphasis id="B975EAFA5D4C15222DF3D03CFDCFFB50" box="[460,593,1139,1160]" italics="true" pageId="11" pageNumber="22">Phyllophaga</emphasis>
)
<emphasis id="B975EAFA5D4C15222E5CD03CFD75FB50" box="[611,747,1139,1160]" italics="true" pageId="11" pageNumber="22">talamancana</emphasis>
Morón and Solís
</taxonomicName>
,
<emphasis id="B975EAFA5D4C15222F88D03DFBA4FB5F" bold="true" box="[951,1082,1138,1159]" pageId="11" pageNumber="22">
<taxonomicNameLabel id="A24657815D4C15222F88D03DFBA4FB5F" box="[951,1082,1138,1159]" pageId="11" pageNumber="22" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31B65635D4C15252E46D0C2FD8EFAB0" lastPageId="12" lastPageNumber="23" pageId="11" pageNumber="22" type="description">
<paragraph id="8BBE36E85D4C15222E46D0C2FD69FB7A" blockId="11.[309,1082,1138,1186]" box="[633,759,1165,1186]" pageId="11" pageNumber="22">
<figureCitation id="133A2A6D5D4C15222E46D0C2FD69FB7A" box="[633,759,1165,1186]" captionStart="Figs" captionStartId="11.[303,346,988,1007]" captionTargetBox="[305,1093,273,963]" captionTargetId="figure-12@11.[297,1094,272,968]" captionTargetPageId="11" captionText="Figs. 1821. Phyllophaga talamancana. 18) protarsal claw, male; 19) genital capsule, lateral view; 20) paramera, distal view; 21) genital plates, female. Scale lines 5 1 mm, except Fig. 18 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900321" httpUri="https://zenodo.org/record/4900321/files/figure.png" pageId="11" pageNumber="22">Figs. 1821</figureCitation>
</paragraph>
<paragraph id="8BBE36E85D4C15252D10D0F8FCF6FC37" blockId="11.[279,1113,1207,1658]" lastBlockId="12.[307,1140,277,1007]" lastPageId="12" lastPageNumber="23" pageId="11" pageNumber="22">
<emphasis id="B975EAFA5D4C15222D10D0F8FE11FB12" bold="true" box="[303,399,1207,1226]" pageId="11" pageNumber="22">
<typeStatus id="54BA884A5D4C15222D10D0F8FE15FB12" box="[303,395,1207,1226]" pageId="11" pageNumber="22" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male. Clypeus, frons and pronotum shiny dark reddish brown; elytra light brown with iridescent luster, head and pronotum with abundant, erect, long setae; elytra with abundant, erect, long setae around scutellum and toward the apex, near suture, and vestiture of uniformly distributed short setae on rest of surface; mouthparts, sterna, pygidium and legs shiny reddish brown. Clypeus 4.2× wider than long, anterior border very scarcely sinuate, with poorly elevated margin, surface shallowly concave, with scarce irregularly distributed, deep, round shaped punctures, each with short setae. Frontoclypeal suture sinuate and deeply impressed. Frons 2.3× wider than long, convex, with reduced number of coarse wide punctures, with long slender setae at sides and on disk. Antenna 10­segmented, with 3­segmented club, lamellae of 8th to 10th segments 1.1× longer than length of preceeding 6 segments combined. Frons 4.0× wider than dorsal diameter of eye. Eye canthus long and wide, with 67 setae. Labrum bilobed, widely sinuate, with scattered slender setae. Mentum slightly convex, with 4 punctures in a transverse row, and long slender setae near the borders; anterior border briefly sinuate. Pronotum 1.4× wider than long and 2.7× wider than frons. Pronotal disk shiny, with deep, round punctures irregularly separated from one another by 17 diameters; lateral borders widely angled, lateral marginal bead crenulate, with regularly located, long, slender setae; basal bead indicated by punctures mainly toward the sides; anterior angles acute, clearly prominent; posterior angles obtuse, slighly prominent. Scutellum 1.5× wider than long, with some deep punctures each with short setae. Elytron 3.0× longer than wider, shiny, densely punctate; epipleural border very narrow, extended along the complete margin, provided with setae along first half, setae diminishing in size anteroposteriorly from long to minute; humeral callus rounded, prominent; apical callus rounded. Metathoracic wings completely developed. Propygidium shiny, densely punctate, setose. Pygidium moderately convex, shiny, shallowly punctate rugose with scattered erect setae; apical margin with 19 long, slender setae; basal margin effaced medially. Pterosternon with long, dense, yellowish setae. Visible abdominal sternites IIIV convex, smooth and glabrous near midline; sternite V convex, shiny, densely punctate with short setae at midline and sparse punctures with mixed long and short setae toward sides; anal plate narrowed, with a transverse shallow concavity and longitudinal furrow at midline, anterior and posterior borders slightly elevated, near the posterior border with 23 erect setae. Protibia slightly shorter than protarsus (0.9:1), with external border tridentate, basal tooth short, preapical spur acute, nearly straight, as long as 2nd protarsomere. Mesotibia with one oblique, sharp, setiferous carina on external side; upper apical spur straight, narrow, and 1.1× longer than lower spur. Metatibia 0.9× shorter than metatarsus, with one oblique, sharp, setiferous carina on external side; upper apical spur articulated, curved, with apex rounded, 1.1× longer than basal metatarsomere, and 1.4× longer than lower spur; lower apical spur articulated with tibial border, with rounded apex. Tarsomeres semicylindrical, not much elongate, with enlarged apex, some setae apically and ventrally one longitudinal carina with a line of short setae at each side. Protaromeres 1 4 with poorly developed subapical tubercles. Tarsal claws symmetrical, similar on all legs, each with large acute tooth at middle of ventral border and with finely serrate border toward the base (
<figureCitation id="133A2A6D5D4B15252E1AD706FDF0FC84" box="[549,622,841,860]" captionStart="Figs" captionStartId="11.[303,346,988,1007]" captionTargetBox="[305,1093,273,963]" captionTargetId="figure-12@11.[297,1094,272,968]" captionTargetPageId="11" captionText="Figs. 1821. Phyllophaga talamancana. 18) protarsal claw, male; 19) genital capsule, lateral view; 20) paramera, distal view; 21) genital plates, female. Scale lines 5 1 mm, except Fig. 18 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900321" httpUri="https://zenodo.org/record/4900321/files/figure.png" pageId="12" pageNumber="23">Fig. 18</figureCitation>
). Genital capsule with short parameres, dorsally and ventrally fused, ring­shaped, apex widely notched, ventrally with preapical minute, toothlike projections. Aedeagus long, with large preapical patches of spinules and strong sclerotized support with dorsal brush of long setae directed to the apex; basally with a pair of dorsal rounded, membranous sac­like structures (
<figureCitation id="133A2A6D5D4B15252F79D7E4FC24FC66" box="[838,954,939,958]" captionStart="Figs" captionStartId="11.[303,346,988,1007]" captionTargetBox="[305,1093,273,963]" captionTargetId="figure-12@11.[297,1094,272,968]" captionTargetPageId="11" captionText="Figs. 1821. Phyllophaga talamancana. 18) protarsal claw, male; 19) genital capsule, lateral view; 20) paramera, distal view; 21) genital plates, female. Scale lines 5 1 mm, except Fig. 18 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900321" httpUri="https://zenodo.org/record/4900321/files/figure.png" pageId="12" pageNumber="23">Figs. 1920</figureCitation>
). Tectum uniformly convex. Length of genital capsule from apex of parameres to border of basal piece:
<quantity id="4CF99B0D5D4B15252858D78CFEC4FC37" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="23" unit="mm" value="5.0">5 mm</quantity>
. Total body length:
<quantity id="4CF99B0D5D4B15252E25D793FDECFC37" box="[538,626,988,1007]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="12" pageNumber="23" unit="mm" value="19.1">19.1 mm</quantity>
. Humeral width: 8.0 mm.
</paragraph>
<paragraph id="8BBE36E85D4B15252D72D043FCA4FB23" blockId="12.[307,1140,1036,1658]" pageId="12" pageNumber="23">
<emphasis id="B975EAFA5D4B15252D72D043FE2CFBF9" bold="true" box="[333,434,1036,1057]" pageId="12" pageNumber="23">
<typeStatus id="54BA884A5D4B15252D72D043FE33FBF9" box="[333,429,1036,1057]" pageId="12" pageNumber="23" type="allotype">Allotype</typeStatus>
.
</emphasis>
Female. Similar to the male except as follows: visible abdominal sternites II to V less convex; anal plate 1.5× longer than male anal plate, convex, punctate, with 60 scattered slender setae. Metafemur 1.3× wider than femur of male. Metatibia at the apex 1.4× wider than male metatibia. Both apical spurs of metatibia articulated, wider than in the male. Pygidium less convex. Ventral genital plates moderately sclerotized, nearly symmetrical, ovate; dorsal plates narrowed, curved, with scattered setae at apex and distal border, apparently articulated baso­laterally with ventral plates (
<figureCitation id="133A2A6D5D4B15252FE4D084FBB4FB38" box="[987,1066,1227,1248]" captionStart="Figs" captionStartId="11.[303,346,988,1007]" captionTargetBox="[305,1093,273,963]" captionTargetId="figure-12@11.[297,1094,272,968]" captionTargetPageId="11" captionText="Figs. 1821. Phyllophaga talamancana. 18) protarsal claw, male; 19) genital capsule, lateral view; 20) paramera, distal view; 21) genital plates, female. Scale lines 5 1 mm, except Fig. 18 5 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4900321" httpUri="https://zenodo.org/record/4900321/files/figure.png" pageId="12" pageNumber="23">Fig. 21</figureCitation>
). Total body length:
<quantity id="4CF99B0D5D4B15252DFDD0A9FDBAFB23" box="[450,548,1254,1275]" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="12" pageNumber="23" unit="mm" value="19.3">19.3 mm</quantity>
. Humeral width:
<quantity id="4CF99B0D5D4B15252EDFD0A9FCA8FB23" box="[736,822,1254,1275]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="12" pageNumber="23" unit="mm" value="8.3">8.3 mm</quantity>
.
</paragraph>
<paragraph id="8BBE36E85D4B15252D72D14EFD8EFAB0" blockId="12.[307,1140,1036,1658]" pageId="12" pageNumber="23">
<emphasis id="B975EAFA5D4B15252D72D14EFDB3FACE" bold="true" box="[333,557,1281,1302]" pageId="12" pageNumber="23">
<typeStatus id="54BA884A5D4B15252D72D14EFE2DFACE" box="[333,435,1281,1302]" pageId="12" pageNumber="23" type="paratype">Paratype</typeStatus>
Variation.
</emphasis>
Male. Similar to
<typeStatus id="54BA884A5D4B15252ED4D14EFCD8FACE" box="[747,838,1281,1302]" pageId="12" pageNumber="23" type="holotype">holotype</typeStatus>
except in total body length:
<quantity id="4CF99B0D5D4B15252D0CD153FE0AFAE9" box="[307,404,1308,1329]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="12" pageNumber="23" unit="mm" value="18.2">18.2 mm</quantity>
, humeral width: 8.0 mm. Female. Similar to
<typeStatus id="54BA884A5D4B15252F41D153FC4CFAE9" box="[894,978,1308,1329]" pageId="12" pageNumber="23" type="allotype">allotype</typeStatus>
except in total body length:
<quantity id="4CF99B0D5D4B15252DF6D178FDB0FA94" box="[457,558,1335,1356]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="12" pageNumber="23" unit="mm" value="18.7">18.7 mm</quantity>
, humeral width: 8.0 mm, elytra of some specimens darker than
<typeStatus id="54BA884A5D4B15252D8BD11CFD92FAB0" box="[436,524,1363,1384]" pageId="12" pageNumber="23" type="allotype">allotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B65635D4B15242D72D121FDFAFEA2" lastPageId="13" lastPageNumber="24" pageId="12" pageNumber="23" type="materials_examined">
<paragraph id="8BBE36E85D4B15242D72D121FCAEFE9D" blockId="12.[307,1140,1036,1658]" lastBlockId="13.[279,1113,277,805]" lastPageId="13" lastPageNumber="24" pageId="12" pageNumber="23">
<emphasis id="B975EAFA5D4B15252D72D121FE44FA5B" bold="true" box="[333,474,1390,1411]" pageId="12" pageNumber="23">Type Series.</emphasis>
<location id="8EDE60335D4B15252DDCD121FDD3FA5B" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B15252DDCD121FDD3FA5B" box="[483,589,1390,1411]" name="Described" pageId="12" pageNumber="23">Described</location>
from
<specimenCount id="9D07FD615D4B15252EAAD121FD77FA5B" box="[661,745,1390,1411]" pageId="12" pageNumber="23" type="male">4 males</specimenCount>
and
<specimenCount id="9D07FD615D4B15252F1BD121FC0EFA5B" box="[804,912,1390,1411]" pageId="12" pageNumber="23" type="female">4 females</specimenCount>
.
<materialsCitation id="3B693CB55D4B15252FA2D121FC15FA62" collectingDate="1996-07-15" collectionCode="IN- BIO" collectorName="M. Moraga." country="Costa Rica" elevation="600" location="Cerro Pittier" municipality="Sendero" pageId="12" pageNumber="23" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="holotype">
<typeStatus id="54BA884A5D4B15252FA2D121FC60FA5B" box="[925,1022,1390,1411]" pageId="12" pageNumber="23" type="holotype">Holotype</typeStatus>
<specimenCount id="9D07FD615D4B15252836D121FBA2FA5B" box="[1033,1084,1390,1411]" pageId="12" pageNumber="23" type="male">male</specimenCount>
(
<collectionCode id="ED10AE2D5D4B15252872D121FEFBFA47" pageId="12" pageNumber="23">INBIO</collectionCode>
):
<collectingCountry id="F31676785D4B15252DBED1C5FDB8FA47" box="[385,550,1418,1439]" name="Costa Rica" pageId="12" pageNumber="23">COSTA RICA</collectingCountry>
:
<collectingRegion id="49C5F80A5D4B15252E0FD1C5FD3BFA47" box="[560,677,1418,1439]" country="Costa Rica" name="Puntarenas" pageId="12" pageNumber="23">Puntarenas</collectingRegion>
,
<collectingMunicipality id="6BDAAC925D4B15252E8ED1C5FC99FA47" box="[689,775,1418,1439]" pageId="12" pageNumber="23">Sendero</collectingMunicipality>
a
<location id="8EDE60335D4B15252F1DD1C5FC38FA47" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B15252F1DD1C5FC38FA47" box="[802,934,1418,1439]" country="Costa Rica" municipality="Sendero" name="Cerro Pittier" pageId="12" pageNumber="23" stateProvince="Puntarenas">Cerro Pittier</location>
,
<quantity id="4CF99B0D5D4B15252F8FD1C5FC6DFA47" box="[944,1011,1418,1439]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="23" unit="m" value="600.0">
<elevation id="002CD1DB5D4B15252F8FD1C5FC6DFA47" box="[944,1011,1418,1439]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="23" unit="m" value="600.0">600 m</elevation>
</quantity>
al Noroeste de la
<location id="8EDE60335D4B15252D50D1EAFD8DFA62" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B15252D50D1EAFD8DFA62" box="[367,531,1445,1466]" country="Costa Rica" municipality="Sendero" name="Estacion Pittier" pageId="12" pageNumber="23" stateProvince="Puntarenas">Estación Pittier</location>
, 1,750 m,
<date id="FFBF10285D4B15252EBAD1EAFD6FFA62" box="[645,753,1445,1466]" pageId="12" pageNumber="23" value="1996-07-15">
<collectingDate id="EFFBE9C05D4B15252EBAD1EAFD6FFA62" box="[645,753,1445,1466]" pageId="12" pageNumber="23" value="1996-07-15">15­VII­96</collectingDate>
</date>
,
<collectorName id="26F4533E5D4B15252EC1D1EAFC1CFA62" box="[766,898,1445,1466]" pageId="12" pageNumber="23">M. Moraga.</collectorName>
</materialsCitation>
<materialsCitation id="3B693CB55D4B15252FACD1EAFC28FA29" collectingDate="1994-10" collectionCode="IN- BIO" collectorName="Z. Fuentes" country="Costa Rica" elevation="1250" latitude="1.15" location="Sector Altamira" longLatPrecision="78" longitude="-1.35" municipality="Buenos Aires" pageId="12" pageNumber="23" specimenCount="1" stateProvince="Puntarenas" typeStatus="allotype">
<typeStatus id="54BA884A5D4B15252FACD1EAFC70FA62" box="[915,1006,1445,1466]" pageId="12" pageNumber="23" type="allotype">Allotype</typeStatus>
<specimenCount id="9D07FD615D4B15252FC8D1EAFBA0FA62" box="[1015,1086,1445,1466]" pageId="12" pageNumber="23" type="female">female</specimenCount>
(
<collectionCode id="ED10AE2D5D4B15252872D1EAFEFBFA0E" pageId="12" pageNumber="23">INBIO</collectionCode>
):
<collectingCountry id="F31676785D4B15252DBDD18EFDB6FA0E" box="[386,552,1473,1494]" name="Costa Rica" pageId="12" pageNumber="23">COSTA RICA</collectingCountry>
:
<collectingRegion id="49C5F80A5D4B15252E0CD18EFD36FA0E" box="[563,680,1473,1494]" country="Costa Rica" name="Puntarenas" pageId="12" pageNumber="23">Puntarenas</collectingRegion>
,
<collectingMunicipality id="6BDAAC925D4B15252E8BD18EFCD8FA0E" box="[692,838,1473,1494]" pageId="12" pageNumber="23">Buenos Aires</collectingMunicipality>
,
<location id="8EDE60335D4B15252F6DD18EFC63FA0E" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B15252F6DD18EFC63FA0E" box="[850,1021,1473,1494]" country="Costa Rica" latitude="1.15" longLatPrecision="78" longitude="-1.35" municipality="Buenos Aires" name="Sector Altamira" pageId="12" pageNumber="23" stateProvince="Puntarenas">Sector Altamira</location>
,
<locationDeviation id="9A0C2FCA5D4B15252836D18EFEF7FA29" pageId="12" pageNumber="23">1 km Suroeste</locationDeviation>
del
<location id="8EDE60335D4B15252DA8D193FDB6FA29" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B15252DA8D193FDB6FA29" box="[407,552,1500,1521]" country="Costa Rica" latitude="1.15" longLatPrecision="78" longitude="-1.35" municipality="Buenos Aires" name="Cerro Biolley" pageId="12" pageNumber="23" stateProvince="Puntarenas">Cerro Biolley</location>
,
<quantity id="4CF99B0D5D4B15252E0CD193FD48FA29" box="[563,726,1500,1521]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.35" metricValueMin="1.15" pageId="12" pageNumber="23" unit="m" value="1250.0" valueMax="1350.0" valueMin="1150.0">
<elevation id="002CD1DB5D4B15252E0CD193FD48FA29" box="[563,726,1500,1521]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.35" metricValueMin="1.15" pageId="12" pageNumber="23" unit="m" value="1250.0" valueMax="1350.0" valueMin="1150.0">1,1501,350 m</elevation>
</quantity>
,
<date id="FFBF10285D4B15252EDED193FCADFA29" box="[737,819,1500,1521]" pageId="12" pageNumber="23" value="1994-10">
<collectingDate id="EFFBE9C05D4B15252EDED193FCADFA29" box="[737,819,1500,1521]" pageId="12" pageNumber="23" value="1994-10">X­1994</collectingDate>
</date>
,
<collectorName id="26F4533E5D4B15252F01D193FC28FA29" box="[830,950,1500,1521]" pageId="12" pageNumber="23">Z. Fuentes</collectorName>
</materialsCitation>
.
<materialsCitation id="3B693CB55D4B15252FF5D193FEEBF9F0" collectingDate="1994-11" collectionCode="INBIO, MXAL" collectorName="M. Segura'" country="Costa Rica" county="Sendero Altamira" elevation="1375" latitude="1.15" location="Refugio Nacional de Fauna" longLatPrecision="78" longitude="-1.35" municipality="Estacion Pittier" pageId="12" pageNumber="23" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<typeStatus id="54BA884A5D4B15252FF5D193FBAAFA29" box="[970,1076,1500,1521]" pageId="12" pageNumber="23" type="paratype">Paratypes</typeStatus>
: same data as allotype except
<quantity id="4CF99B0D5D4B15252E7DD1B7FD70F9D5" box="[578,750,1528,1549]" metricMagnitude="3" metricUnit="m" metricValue="1.375" metricValueMax="1.45" metricValueMin="1.3" pageId="12" pageNumber="23" unit="m" value="1375.0" valueMax="1450.0" valueMin="1300.0">
<elevation id="002CD1DB5D4B15252E7DD1B7FD70F9D5" box="[578,750,1528,1549]" metricMagnitude="3" metricUnit="m" metricValue="1.375" metricValueMax="1.45" metricValueMin="1.3" pageId="12" pageNumber="23" unit="m" value="1375.0" valueMax="1450.0" valueMin="1300.0">1,3001,450 m</elevation>
</quantity>
,
<date id="FFBF10285D4B15252EC2D1B7FCA0F9D5" box="[765,830,1528,1549]" pageId="12" pageNumber="23" value="1994-11">
<collectingDate id="EFFBE9C05D4B15252EC2D1B7FCA0F9D5" box="[765,830,1528,1549]" pageId="12" pageNumber="23" value="1994-11">XI­94</collectingDate>
</date>
,
<collectorName id="26F4533E5D4B15252F72D1B7FC57F9D5" box="[845,969,1528,1549]" pageId="12" pageNumber="23">M. Segura</collectorName>
(
<specimenCount id="9D07FD615D4B15252FDCD1B7FBB0F9D5" box="[995,1070,1528,1549]" count="1" pageId="12" pageNumber="23" type="male">1 male</specimenCount>
,
<specimenCount id="9D07FD615D4B15252801D1B7FEF4F9F0" count="1" pageId="12" pageNumber="23" type="female">1 female</specimenCount>
);
</materialsCitation>
<materialsCitation id="3B693CB55D4B15252DBCD25CFC3CF99B" collectingDate="1995-10-04" collectionCode="INBIO, MXAL" collectorName="M. Moraga" country="Costa Rica" county="Sendero Altamira" elevation="1720" latitude="1.15" location="Estacion Pittier" longLatPrecision="78" longitude="-1.35" municipality="Estacion Pittier" pageId="12" pageNumber="23" specimenCode="INBIO CR 1002 468647" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<collectingRegion id="49C5F80A5D4B15252DBCD25CFE66F9F0" box="[387,504,1555,1576]" country="Costa Rica" name="Puntarenas" pageId="12" pageNumber="23">Puntarenas</collectingRegion>
,
<locationDeviation id="9A0C2FCA5D4B15252E35D25CFDAEF9F0" box="[522,560,1555,1576]" pageId="12" pageNumber="23">NO</locationDeviation>
<location id="8EDE60335D4B15252E02D25CFD78F9F0" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B15252E02D25CFD78F9F0" box="[573,742,1555,1576]" country="Costa Rica" county="Sendero Altamira" latitude="1.15" longLatPrecision="78" longitude="-1.35" municipality="Estacion Pittier" name="Estacion Pittier" pageId="12" pageNumber="23" stateProvince="Puntarenas">Estación Pittier</location>
,
<collectingCounty id="62DF4E645D4B15252EC8D25CFC27F9F0" box="[759,953,1555,1576]" pageId="12" pageNumber="23">Sendero Altamira</collectingCounty>
,
<date id="FFBF10285D4B15252FF5D25CFB86F9F0" box="[970,1048,1555,1576]" pageId="12" pageNumber="23" value="1995-10-04">
<collectingDate id="EFFBE9C05D4B15252FF5D25CFB86F9F0" box="[970,1048,1555,1576]" pageId="12" pageNumber="23" value="1995-10-04">4­X­95</collectingDate>
</date>
,
<quantity id="4CF99B0D5D4B15252816D25CFE11F99B" metricMagnitude="3" metricUnit="m" metricValue="1.72" metricValueMax="1.7399999999999998" metricValueMin="1.7" pageId="12" pageNumber="23" unit="m" value="1720.0" valueMax="1740.0" valueMin="1700.0">
<elevation id="002CD1DB5D4B15252816D25CFE11F99B" metricMagnitude="3" metricUnit="m" metricValue="1.72" metricValueMax="1.7399999999999998" metricValueMin="1.7" pageId="12" pageNumber="23" unit="m" value="1720.0" valueMax="1740.0" valueMin="1700.0">1,700 1,740 m</elevation>
</quantity>
,
<collectorName id="26F4533E5D4B15252DA2D261FD84F99B" box="[413,538,1582,1603]" pageId="12" pageNumber="23">M. Moraga</collectorName>
,
<specimenCode id="DBA79E935D4B15252E17D261FCABF99B" box="[552,821,1582,1603]" pageId="12" pageNumber="23">INBIO CR 1002 468647</specimenCode>
(
<specimenCount id="9D07FD615D4B15252F75D261FC09F99B" box="[842,919,1582,1603]" count="1" pageId="12" pageNumber="23" type="male">1 male</specimenCount>
);
</materialsCitation>
<materialsCitation id="3B693CB55D4B15252F92D261FE61F9A2" collectingDate="1995-01-18" collectionCode="INBIO, MXAL" collectorName="R. Villalobos" country="Costa Rica" county="Sendero Altamira" elevation="1670" latitude="1.15" location="PILA- ACLA" longLatPrecision="78" longitude="-1.35" municipality="Estacion Pittier" pageId="12" pageNumber="23" specimenCode="INBIO CR 1002 183335" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas" typeStatus="paratype">
<collectingRegion id="49C5F80A5D4B15252F92D261FBBCF99B" box="[941,1058,1582,1603]" country="Costa Rica" name="Puntarenas" pageId="12" pageNumber="23">Puntarenas</collectingRegion>
,
<location id="8EDE60335D4B1525280FD261FEE3F987" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B1525280FD261FEE3F987" country="Costa Rica" county="Sendero Altamira" latitude="1.15" longLatPrecision="78" longitude="-1.35" municipality="Estacion Pittier" name="PILA- ACLA" pageId="12" pageNumber="23" stateProvince="Puntarenas">PILA­ACLA</location>
,
<collectingMunicipality id="6BDAAC925D4B15252DB4D205FDAFF987" box="[395,561,1610,1631]" pageId="12" pageNumber="23">Estación Pittier</collectingMunicipality>
, 5/
<date id="FFBF10285D4B15252E6AD205FD3AF987" box="[597,676,1610,1631]" pageId="12" pageNumber="23" value="1995-01-18">
<collectingDate id="EFFBE9C05D4B15252E6AD205FD3AF987" box="[597,676,1610,1631]" pageId="12" pageNumber="23" value="1995-01-18">18­I­95</collectingDate>
</date>
,
<quantity id="4CF99B0D5D4B15252E8CD205FC91F987" box="[691,783,1610,1631]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="12" pageNumber="23" unit="m" value="1670.0">
<elevation id="002CD1DB5D4B15252E8CD205FC91F987" box="[691,783,1610,1631]" metricMagnitude="3" metricUnit="m" metricValue="1.67" pageId="12" pageNumber="23" unit="m" value="1670.0">1,670 m</elevation>
</quantity>
,
<collectorName id="26F4533E5D4B15252F21D205FC2EF987" box="[798,944,1610,1631]" pageId="12" pageNumber="23">R. Villalobos</collectorName>
,
<specimenCode id="DBA79E935D4B15252F81D205FE1FF9A2" pageId="12" pageNumber="23">INBIO CR 1002 183335</specimenCode>
(
<specimenCount id="9D07FD615D4B15252DABD22AFE6DF9A2" box="[404,499,1637,1658]" count="1" pageId="12" pageNumber="23" type="female">1 female</specimenCount>
);
</materialsCitation>
<materialsCitation id="3B693CB55D4B15242E38D22AFEC4FE9D" collectingDate="1992-04" collectionCode="INBIO, MXAL" collectorName="G. Mora'" country="Costa Rica" county="Sendero Altamira" elevation="1250" lastPageId="13" lastPageNumber="24" latitude="1.15" location="Refugio Nacional de Fauna Silvestre Tapanti´" longLatPrecision="78" longitude="-1.35" municipality="Quebrada Segunda" pageId="12" pageNumber="23" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cartago" typeStatus="paratype">
<collectingCountry id="F31676785D4B15252E2FD22AFD28F9A2" box="[528,694,1637,1658]" name="Costa Rica" pageId="12" pageNumber="23">COSTA RICA</collectingCountry>
:
<collectingRegion id="49C5F80A5D4B15252EFED22AFC88F9A2" box="[705,790,1637,1658]" country="Costa Rica" name="Cartago" pageId="12" pageNumber="23">Cartago</collectingRegion>
,
<location id="8EDE60335D4B15242F1DD22AFE2FFEF2" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4B15242F1DD22AFE2FFEF2" country="Costa Rica" county="Sendero Altamira" lastPageId="13" lastPageNumber="24" latitude="1.15" longLatPrecision="78" longitude="-1.35" municipality="Quebrada Segunda" name="Refugio Nacional de Fauna Sil- vestre Tapanti´" pageId="12" pageNumber="23" stateProvince="Cartago">Refugio Nacional de Fauna Silvestre Tapantı´</location>
,
<collectingMunicipality id="6BDAAC925D4A15242D81D55AFD14FEF2" box="[446,650,277,298]" pageId="13" pageNumber="24">Quebrada Segunda</collectingMunicipality>
,
<quantity id="4CF99B0D5D4A15242EA8D55AFD6DFEF2" box="[663,755,277,298]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="13" pageNumber="24" unit="m" value="1250.0">
<elevation id="002CD1DB5D4A15242EA8D55AFD6DFEF2" box="[663,755,277,298]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="13" pageNumber="24" unit="m" value="1250.0">1,250 m</elevation>
</quantity>
,
<date id="FFBF10285D4A15242F3FD55AFCA1FEF2" box="[768,831,277,298]" pageId="13" pageNumber="24" value="1992-04">
<collectingDate id="EFFBE9C05D4A15242F3FD55AFCA1FEF2" box="[768,831,277,298]" pageId="13" pageNumber="24" value="1992-04">IV­92</collectingDate>
</date>
,
<collectorName id="26F4533E5D4A15242F73D55AFC2DFEF2" box="[844,947,277,298]" pageId="13" pageNumber="24">G. Mora</collectorName>
(
<specimenCount id="9D07FD615D4A15242FF3D55AFB88FEF2" box="[972,1046,277,298]" count="1" pageId="13" pageNumber="24" type="male">1 male</specimenCount>
,
<specimenCount id="9D07FD615D4A1524281BD55AFED0FE9D" count="1" pageId="13" pageNumber="24" type="female">1 female</specimenCount>
).
</materialsCitation>
Paratypes deposited in
<collectionCode id="ED10AE2D5D4A15242E64D57FFD3AFE9D" box="[603,676,304,325]" pageId="13" pageNumber="24">INBIO</collectionCode>
and
<collectionCode id="ED10AE2D5D4A15242EE4D57FFCB5FE9D" box="[731,811,304,325]" pageId="13" pageNumber="24">MXAL</collectionCode>
.
</paragraph>
<paragraph id="8BBE36E85D4A15242D0DD505FDFAFEA2" blockId="13.[279,1113,277,805]" pageId="13" pageNumber="24">
<emphasis id="B975EAFA5D4A15242D0DD505FE48FE87" bold="true" box="[306,470,330,351]" pageId="13" pageNumber="24">Type Locality.</emphasis>
<location id="8EDE60335D4A15242DE1D504FD63FEB8" LSID="urn:lsid:plazi:treatment:03A887FE5D4C15242D0AD03CFC1BFCFD:8EDE60335D4A15242DE1D504FD63FEB8" box="[478,765,331,352]" country="Costa Rica" municipality="Cordillera de Talamanca" name="Falda Sur del Cerro Pittier" pageId="13" pageNumber="24">Falda Sur del Cerro Pittier</location>
,
<collectingMunicipality id="6BDAAC925D4A15242F36D504FB8EFEB8" box="[777,1040,331,352]" pageId="13" pageNumber="24">Cordillera de Talamanca</collectingMunicipality>
,
<collectingCountry id="F31676785D4A15242823D504FED9FEA2" name="Costa Rica" pageId="13" pageNumber="24">Costa Rica</collectingCountry>
(
<geoCoordinate id="EE35502F5D4A15242D67D52AFE56FEA2" box="[344,456,357,378]" pageId="13" pageNumber="24">9°01'450N</geoCoordinate>
;
<geoCoordinate id="EE35502F5D4A15242DEAD52AFDC6FEA2" box="[469,600,357,378]" pageId="13" pageNumber="24">82°57'550W</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B65635D4A15242D0DD5CFFCCAFDC2" pageId="13" pageNumber="24" type="biology_ecology">
<paragraph id="8BBE36E85D4A15242D0DD5CFFCCAFDC2" blockId="13.[279,1113,277,805]" pageId="13" pageNumber="24">
<emphasis id="B975EAFA5D4A15242D0DD5CFFE7DFE4D" bold="true" box="[306,483,384,405]" pageId="13" pageNumber="24">Biological Data.</emphasis>
Four males and
<specimenCount id="9D07FD615D4A15242EADD5CFFC80FE4D" box="[658,798,384,405]" pageId="13" pageNumber="24" type="female">three females</specimenCount>
of
<taxonomicName id="4C014D6B5D4A15242F7ED5CFFC78FE4D" box="[833,998,384,405]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="24" phylum="Arthropoda" rank="species" species="talamancana" status="sp. nov.">
<emphasis id="B975EAFA5D4A15242F7ED5CFFC78FE4D" box="[833,998,384,405]" italics="true" pageId="13" pageNumber="24">P. talamancana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24657815D4A15242FD2D5CFFB81FE4D" box="[1005,1055,384,405]" pageId="13" pageNumber="24" rank="species">n.sp.</taxonomicNameLabel>
were collected at lights in grasslands, near the forest, and
<specimenCount id="9D07FD615D4A15242F74D5D4FC5EFE68" box="[843,960,411,432]" pageId="13" pageNumber="24" type="female">one female</specimenCount>
was collected at lights in premontane rain forest, located between
<quantity id="4CF99B0D5D4A15242F01D5FAFC40FE12" box="[830,990,437,458]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.75" metricValueMin="1.25" pageId="13" pageNumber="24" unit="m" value="1500.0" valueMax="1750.0" valueMin="1250.0">
<geoCoordinate id="EE35502F5D4A15242F01D5FAFCE7FE12" box="[830,889,437,458]" degrees="1,250" direction="north" orientation="latitude" pageId="13" pageNumber="24" precision="55" value="1.25">1,250</geoCoordinate>
<geoCoordinate id="EE35502F5D4A15242F46D5FAFC5CFE12" box="[889,962,437,458]" degrees="1,750" direction="west" orientation="longitude" pageId="13" pageNumber="24" precision="55" value="-1.75">1,750</geoCoordinate>
m
</quantity>
of altitude. Phenology: January (1), April (2), July (1), October (2), November (2). Other species of
<taxonomicName id="4C014D6B5D4A15242DAFD5A4FD88FDD8" box="[400,534,491,512]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B975EAFA5D4A15242DAFD5A4FD88FDD8" box="[400,534,491,512]" italics="true" pageId="13" pageNumber="24">Phyllophaga</emphasis>
</taxonomicName>
flying at the same time and places were:
<emphasis id="B975EAFA5D4A15242FCCD5A4FEF4FDC3" italics="true" pageId="13" pageNumber="24">P. nevermannea</emphasis>
Saylor,
<emphasis id="B975EAFA5D4A15242DFAD649FDAAFDC3" box="[453,564,518,539]" italics="true" pageId="13" pageNumber="24">P. orosina</emphasis>
Moser,
<taxonomicName id="4C014D6B5D4A15242EB2D649FC8EFDC3" box="[653,784,518,539]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="24" phylum="Arthropoda" rank="species" species="tapantina" status="sp. nov.">
<emphasis id="B975EAFA5D4A15242EB2D649FC8EFDC3" box="[653,784,518,539]" italics="true" pageId="13" pageNumber="24">P. tapantina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24657815D4A15242F25D64AFCCAFDC2" box="[794,852,517,538]" pageId="13" pageNumber="24" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C31B65635D4A15242D0DD66FFC21FD28" pageId="13" pageNumber="24" type="discussion">
<paragraph id="8BBE36E85D4A15242D0DD66FFC21FD28" blockId="13.[279,1113,277,805]" pageId="13" pageNumber="24">
<emphasis id="B975EAFA5D4A15242D0DD66FFE00FDED" bold="true" box="[306,414,544,565]" pageId="13" pageNumber="24">Remarks.</emphasis>
<taxonomicName id="4C014D6B5D4A15242D94D66FFCF7FDED" box="[427,873,544,565]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="24" phylum="Arthropoda" rank="species" species="talamancana" status="sp. nov." subGenus="Phyllophaga">
<emphasis id="B975EAFA5D4A15242D94D66FFDAFFDED" box="[427,561,544,565]" italics="true" pageId="13" pageNumber="24">Phyllophaga</emphasis>
(
<emphasis id="B975EAFA5D4A15242E79D66FFD55FDED" box="[582,715,544,565]" italics="true" pageId="13" pageNumber="24">Phyllophaga</emphasis>
)
<emphasis id="B975EAFA5D4A15242EDED66FFCF7FDED" box="[737,873,544,565]" italics="true" pageId="13" pageNumber="24">talamancana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24657815D4A15242F49D66FFC2BFDED" box="[886,949,544,565]" pageId="13" pageNumber="24" rank="species">n. sp.</taxonomicNameLabel>
is not closely related with any known species group (
<emphasis id="B975EAFA5D4A15242E84D674FD6BFD88" box="[699,757,571,592]" italics="true" pageId="13" pageNumber="24">sensu</emphasis>
<bibRefCitation id="EF904B195D4A15242EC2D674FC1AFD88" author="Moron" box="[765,900,571,592]" firstAuthor="Moron" pageId="13" pageNumber="24" refId="ref11224" refString="Moron, M. A. 1986. El genero Phyllophaga en Mexico. Morfologia, distribucion y sistematica supraespecifica (Insecta: Coleoptera). Publ. 20, Instituto de Ecologia, Mexico, D. F. 341 pp." type="book" year="1986">Morón 1986</bibRefCitation>
). The general structure of the last sternites, shape of tarsal claws and the length of antennal club in the males suggest a vague relation with some species in the
<emphasis id="B975EAFA5D4A15242FF1D63FFBD6FD5D" box="[974,1096,624,645]" italics="true" pageId="13" pageNumber="24">schizorhina</emphasis>
group, and the sculpture of the head and pronotum resemble some species in the
<emphasis id="B975EAFA5D4A15242D6DD6E9FE5BFD63" box="[338,453,678,699]" italics="true" pageId="13" pageNumber="24">blanchardi</emphasis>
group. The shape and details of the male genital capsule, sculpture of the pronotum, pygidium and sternites, body luster and color will aid to separate this species from other Costarican
<emphasis id="B975EAFA5D4A15242F0DD694FC21FD28" box="[818,959,731,752]" italics="true" pageId="13" pageNumber="24">
<taxonomicName id="4C014D6B5D4A15242F0DD694FC25FD28" box="[818,955,731,752]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="24" phylum="Arthropoda" rank="genus">Phyllophaga</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31B65635D4A15242D0DD6BAFC1BFCFD" pageId="13" pageNumber="24" type="etymology">
<paragraph id="8BBE36E85D4A15242D0DD6BAFC1BFCFD" blockId="13.[279,1113,277,805]" pageId="13" pageNumber="24">
<emphasis id="B975EAFA5D4A15242D0DD6BAFE2FFCD2" bold="true" box="[306,433,757,778]" pageId="13" pageNumber="24">Etymology.</emphasis>
Derived from Cordillera de Talamanca that comprises the majority of the geographic range where this species lives.
</paragraph>
</subSubSection>
</treatment>
</document>