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<document id="2F97E3A1B983C40A727674C20379A80C" ID-DOI="10.11646/zootaxa.5481.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="12758944" ID-ZooBank="F86F4482-5F80-444F-A5DE-F1F1BAC04E41" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721216078619" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="A64A2B29256F163542FEFF4EFCB09B35" docLanguage="en" docName="zootaxa.5481.3.7.pdf" docOrigin="Zootaxa 5481 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5481.3.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hygrodromicus splendidus Zerche 1992" docType="treatment" docVersion="2" lastPageNumber="389" masterDocId="5A735351256B16304269FFD9FFC49D3C" masterDocTitle="On some species of the genus Hygrodromicus Tronquet with description of a new species from Pakistan (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="390" masterPageNumber="384" pageNumber="388" updateTime="1721216576459" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="570A5AF59AA522680C49C5F00F624A41">On some species of the genus Hygrodromicus Tronquet with description of a new species from Pakistan (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<treatment id="A64A2B29256F163542FEFF4EFCB09B35" LSID="urn:lsid:plazi:treatment:A64A2B29256F163542FEFF4EFCB09B35" httpUri="http://treatment.plazi.org/id/A64A2B29256F163542FEFF4EFCB09B35" lastPageId="5" lastPageNumber="389" pageId="4" pageNumber="388">
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<paragraph id="2E5C9A3F256F163442FEFF4EFD419D8D" blockId="4.[151,645,151,214]" box="[151,645,151,178]" pageId="4" pageNumber="388">
<heading id="75142D53256F163442FEFF4EFD419D8D" bold="true" box="[151,645,151,178]" fontSize="11" level="1" pageId="4" pageNumber="388" reason="1">
<taxonomicName id="E9E3E1BC256F163442FEFF4EFD419D8D" ID-CoL="3NBJ4" authority="Zerche, 1992" authorityName="Zerche" authorityYear="1992" box="[151,645,151,178]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="388" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="1C97462D256F163442FEFF4EFD419D8D" bold="true" box="[151,645,151,178]" pageId="4" pageNumber="388">
<emphasis id="1C97462D256F163442FEFF4EFE199D8E" bold="true" box="[151,477,151,178]" italics="true" pageId="4" pageNumber="388">Hygrodromicus splendidus</emphasis>
<bibRefCitation id="4A72E7CE256F1634438DFF4EFD419D8D" author="Zerche, L." box="[484,645,151,177]" pageId="4" pageNumber="388" pagination="105 - 142" refId="ref3633" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche, 1992</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<paragraph id="2E5C9A3F256F163442FEFF62FEE39DEA" blockId="4.[151,645,151,214]" box="[151,295,187,214]" pageId="4" pageNumber="388">
(
<figureCitation id="B6D886BA256F163442F6FF62FF2C9DE9" box="[159,232,187,213]" captionStart="FIGURES 12" captionStartId="2.[151,264,1207,1231]" captionTargetBox="[157,1430,281,1175]" captionTargetId="figure-60@2.[151,1436,270,1181]" captionTargetPageId="2" captionText="FIGURES 12. Habitus of Hygrodromicus: 1H. incrassatus (paratype), 2H. splendidus (Kazakhstan, Ketmen` Mts.). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12758946" httpUri="https://zenodo.org/record/12758946/files/figure.png" pageId="4" pageNumber="388">Figs 2</figureCitation>
,
<figureCitation id="B6D886BA256F1634429DFF62FEDB9DEA" box="[244,287,187,214]" captionStart-0="FIGURES 36" captionStart-1="FIGURE 7" captionStartId-0="3.[151,264,1842,1866]" captionStartId-1="4.[151,250,1743,1767]" captionTargetBox-0="[177,1417,713,1811]" captionTargetBox-1="[151,1435,664,1720]" captionTargetId-0="figure-228@3.[151,1436,705,1819]" captionTargetId-1="figure-160@4.[151,1435,664,1720]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 36. Aedeagus of Hygrodromicus (Figs 34: H. incrassatus (holotype); Figs. 56: H. splendidus (Kazakhstan: Ketmen` Mts.): 3, 5parameral view, 4, 6lateral view. Scale bars: 0.1 mm." captionText-1="FIGURE 7. Distribution of Hygrodromicus incrassatus (square) and H. splendidus (circles)." figureDoi-0="http://doi.org/10.5281/zenodo.12758950" figureDoi-1="http://doi.org/10.5281/zenodo.12758954" httpUri-0="https://zenodo.org/record/12758950/files/figure.png" httpUri-1="https://zenodo.org/record/12758954/files/figure.png" pageId="4" pageNumber="388">57</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="66F9C9B4256F163442FEFED9FDA19C2B" box="[151,613,255,280]" pageId="4" pageNumber="388" type="reference_group">
<paragraph id="2E5C9A3F256F163442FEFED9FDA19C2B" blockId="4.[151,613,255,280]" box="[151,613,255,280]" pageId="4" pageNumber="388">
<treatmentCitationGroup id="0EF3BD11256F163442FEFED9FDA19C2B" box="[151,613,255,280]" pageId="4" pageNumber="388">
<treatmentCitation id="AF42BC2E256F163442FEFED9FDA19C2B" author="Zerche, L." box="[151,613,255,280]" page="116" pageId="4" pageNumber="388" year="1992">
<taxonomicName id="E9E3E1BC256F163442FEFED9FDA19C2B" ID-CoL="3NBJ4" authority="Zerche, 1992: 116" authorityName="Zerche" authorityPageNumber="116" authorityYear="1992" box="[151,613,255,280]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="388" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="1C97462D256F163442FEFED9FE639C24" box="[151,423,256,280]" italics="true" pageId="4" pageNumber="388">Hygrodromicus splendidus</emphasis>
<bibRefCitation id="4A72E7CE256F163443C4FED9FDA19C2B" author="Zerche, L." box="[429,613,255,280]" pageId="4" pageNumber="388" pagination="105 - 142" refId="ref3633" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche, 1992: 116</bibRefCitation>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="66F9C9B4256F163442FEFE9CFA959CCC" pageId="4" pageNumber="388" type="materials_examined">
<paragraph id="2E5C9A3F256F163442FEFE9CFA959CCC" blockId="4.[151,1437,325,641]" pageId="4" pageNumber="388">
<emphasis id="1C97462D256F163442FEFE9CFEBB9C63" bold="true" box="[151,383,325,351]" pageId="4" pageNumber="388">Material examined.</emphasis>
<materialsCitation id="9E8B9062256F163443EAFE9CFBBD9C5C" box="[387,1145,325,352]" collectionCode="R" country="Kazakhstan" location="Dolaity" municipality="Ketmen Ridge" pageId="4" pageNumber="388" specimenCount="30" specimenCount-female="21" specimenCount-male="9" stateProvince="Almaty Oblysy">
<emphasis id="1C97462D256F163443EAFE9CFD7C9C63" bold="true" box="[387,696,325,351]" pageId="4" pageNumber="388">
<collectingCountry id="56F4DAAF256F163443EAFE9CFD899C63" box="[387,589,325,351]" name="Kazakhstan" pageId="4" pageNumber="388">KAZAKHSTAN</collectingCountry>
:
<collectingRegion id="EC2754DD256F1634403EFE9CFD709C63" box="[599,692,325,351]" country="Kazakhstan" name="Almaty Oblysy" pageId="4" pageNumber="388">ALMATY</collectingRegion>
:
</emphasis>
<specimenCount id="38E551B6256F163440D5FE9FFD3F9C5C" box="[700,763,326,352]" count="9" pageId="4" pageNumber="388" type="male">9 ♂♂</specimenCount>
,
<specimenCount id="38E551B6256F1634416FFE9FFC979C5C" box="[774,851,326,352]" count="21" pageId="4" pageNumber="388" type="female">21 ♀♀</specimenCount>
:
<collectingMunicipality id="CE380045256F16344134FE9FFBC59C5C" box="[861,1025,326,352]" pageId="4" pageNumber="388">Ketmen Ridge</collectingMunicipality>
,
<location id="2B3CCCE4256F16344663FE9FFB9A9C5C" LSID="urn:lsid:plazi:treatment:A64A2B29256F163542FEFF4EFCB09B35:2B3CCCE4256F16344663FE9FFB9A9C5C" box="[1034,1118,326,352]" country="Kazakhstan" municipality="Ketmen Ridge" name="Dolaity" pageId="4" pageNumber="388" stateProvince="Almaty Oblysy">Dolaity</location>
<collectionCode id="48F202FA256F1634460AFE9FFBBD9C5C" box="[1123,1145,326,352]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="388">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="9E8B9062256F163446E8FE9FFADB9C5C" box="[1153,1311,326,353]" collectingDate="1988-07-15" collectionCode="V" country="Kazakhstan" pageId="4" pageNumber="388" specimenCount="1" stateProvince="Almaty Oblysy">
<date id="5A5DBCFF256F163446E8FE9FFAC79C5D" box="[1153,1283,326,353]" pageId="4" pageNumber="388" value="1988-07-15">
<collectingDate id="4A194517256F163446E8FE9FFAC79C5D" box="[1153,1283,326,353]" pageId="4" pageNumber="388" value="1988-07-15">15.07.1988</collectingDate>
</date>
.
<collectionCode id="48F202FA256F16344762FE9FFADB9C5C" box="[1291,1311,326,352]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="388" type="Museum">V</collectionCode>
</materialsCitation>
.
<collectorName id="8316FFE9256F16344741FE9FFA599C5C" box="[1320,1437,326,352]" pageId="4" pageNumber="388">Kastcheev</collectorName>
leg. (cSh,
<collectionCode id="48F202FA256F16344360FEB3FE959CB8" box="[265,337,362,388]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="388">NMW</collectionCode>
,
<collectionCode id="48F202FA256F16344334FEB3FE549CB8" box="[349,400,362,388]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="388" type="Museum">ZIN</collectionCode>
);
<materialsCitation id="9E8B9062256F163443CDFEB3FC9C9CB8" box="[420,856,362,388]" collectingDate="1988-09-07" collectionCode="V" country="Kazakhstan" pageId="4" pageNumber="388" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Almaty Oblysy">
<specimenCount id="38E551B6256F163443CDFEB3FE149CB8" box="[420,464,362,388]" count="1" pageId="4" pageNumber="388" type="male">1 ♂</specimenCount>
,
<specimenCount id="38E551B6256F163443B5FEB3FDCD9CB8" box="[476,521,362,388]" count="1" pageId="4" pageNumber="388" type="female">1 ♀</specimenCount>
: same data,
<date id="5A5DBCFF256F163440FAFEB3FCD59CB8" box="[659,785,362,388]" pageId="4" pageNumber="388" value="1988-09-07">
<collectingDate id="4A194517256F163440FAFEB3FCD59CB8" box="[659,785,362,388]" pageId="4" pageNumber="388" value="1988-09-07">09.07.1988</collectingDate>
</date>
(cSh)
</materialsCitation>
;
<materialsCitation id="9E8B9062256F1634410DFEB3FB609CB8" box="[868,1188,362,388]" collectingDate="1988-09-07" collectionCode="R" country="Kazakhstan" location="Avat" pageId="4" pageNumber="388" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Almaty Oblysy">
<specimenCount id="38E551B6256F1634410DFEB3FC559CB8" box="[868,913,362,388]" count="1" pageId="4" pageNumber="388" type="male">1 ♂</specimenCount>
,
<specimenCount id="38E551B6256F163441F4FEB3FC0D9CB8" box="[925,969,362,388]" count="1" pageId="4" pageNumber="388" type="female">1 ♀</specimenCount>
: same data,
<location id="2B3CCCE4256F1634463AFEB3FB439CB8" LSID="urn:lsid:plazi:treatment:A64A2B29256F163542FEFF4EFCB09B35:2B3CCCE4256F1634463AFEB3FB439CB8" box="[1107,1159,362,388]" country="Kazakhstan" name="Avat" pageId="4" pageNumber="388" stateProvince="Almaty Oblysy">Avat</location>
<collectionCode id="48F202FA256F163446E7FEB3FB609CB8" box="[1166,1188,362,388]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="388">R</collectionCode>
</materialsCitation>
.
<date id="5A5DBCFF256F163446C6FEB3FAE99CB8" box="[1199,1325,362,388]" pageId="4" pageNumber="388" value="2009-07-14">14.07.2009</date>
(cSh);
<materialsCitation id="9E8B9062256F163447E9FEB3FD159C94" collectingDate="1988-09-07" collectionCode="R" country="Kazakhstan" location="Kulbastau" pageId="4" pageNumber="388" specimenCount="19" specimenCount-female="6" specimenCount-male="13" stateProvince="Almaty Oblysy">
<specimenCount id="38E551B6256F163447E9FEB3FF079C94" count="13" pageId="4" pageNumber="388" type="male">13 ♂♂</specimenCount>
,
<specimenCount id="38E551B6256F163442B9FE57FED59C94" box="[208,273,398,424]" count="6" pageId="4" pageNumber="388" type="female">6 ♀♀</specimenCount>
: same data,
<location id="2B3CCCE4256F163443F4FE57FDCB9C94" LSID="urn:lsid:plazi:treatment:A64A2B29256F163542FEFF4EFCB09B35:2B3CCCE4256F163443F4FE57FDCB9C94" box="[413,527,398,424]" country="Kazakhstan" name="Kulbastau" pageId="4" pageNumber="388" stateProvince="Almaty Oblysy">Kulbastau</location>
[=Kol`bastau]
<collectionCode id="48F202FA256F163440D2FE57FD159C94" box="[699,721,398,424]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="388">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="9E8B9062256F163440B4FE57FC249C94" box="[733,992,398,424]" collectingDate="1988-07-27" collectionCode="ZIN" country="Kazakhstan" pageId="4" pageNumber="388" specimenCount="1" stateProvince="Almaty Oblysy">
<date id="5A5DBCFF256F163440B4FE57FC9F9C94" box="[733,859,398,424]" pageId="4" pageNumber="388" value="1988-07-27">
<collectingDate id="4A194517256F163440B4FE57FC9F9C94" box="[733,859,398,424]" pageId="4" pageNumber="388" value="1988-07-27">27.07.1988</collectingDate>
</date>
(cSh,
<collectionCode id="48F202FA256F163441CDFE57FC139C94" box="[932,983,398,424]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="388" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9E8B9062256F16344185FE57FE9A9CF0" collectingDate="1991-05-18" collectionCode="ZIN" collectorName="Bolshoy Kyrgyzsay." country="Kazakhstan" pageId="4" pageNumber="388" specimenCount="3" specimenCount-female="3" stateProvince="Almaty Oblysy">
<specimenCount id="38E551B6256F16344185FE57FBE99C94" box="[1004,1069,398,424]" count="3" pageId="4" pageNumber="388" type="female">3 ♀♀</specimenCount>
: same data,
<collectorName id="8316FFE9256F163446D0FE57FA599C94" box="[1209,1437,398,424]" pageId="4" pageNumber="388">Bolshoy Kyrgyzsay.</collectorName>
<date id="5A5DBCFF256F163442FEFE6BFED29CF1" box="[151,278,434,461]" pageId="4" pageNumber="388" value="1991-05-18">
<collectingDate id="4A194517256F163442FEFE6BFED29CF1" box="[151,278,434,461]" pageId="4" pageNumber="388" value="1991-05-18">18.05.1991</collectingDate>
</date>
(cSh)
</materialsCitation>
;
<materialsCitation id="9E8B9062256F16344305FE6BFD369CF0" box="[364,754,434,460]" collectingDate="1988-10-06" collectionCode="ZIN" collectorName="Bolshoy Kyrgyzsay." country="Kazakhstan" pageId="4" pageNumber="388" specimenCount="1" specimenCount-female="1" stateProvince="Almaty Oblysy">
<specimenCount id="38E551B6256F16344305FE6BFE5F9CF0" box="[364,411,434,460]" count="1" pageId="4" pageNumber="388" type="female">1 ♀</specimenCount>
: same data,
<date id="5A5DBCFF256F16344042FE6BFD6D9CF0" box="[555,681,434,460]" pageId="4" pageNumber="388" value="1988-10-06">
<collectingDate id="4A194517256F16344042FE6BFD6D9CF0" box="[555,681,434,460]" pageId="4" pageNumber="388" value="1988-10-06">10.06.1988</collectingDate>
</date>
(cSh)
</materialsCitation>
;
<materialsCitation id="9E8B9062256F16344169FE68FB2E9CF0" box="[768,1258,433,460]" collectionCode="R" country="Kazakhstan" location="Koksay" municipality="Aksu-Dzhabagly" pageId="4" pageNumber="388" specimenCount="1" specimenCount-male="1" stateProvince="JAMBYL">
<emphasis id="1C97462D256F16344169FE68FCA09CF7" bold="true" box="[768,868,433,459]" pageId="4" pageNumber="388">
<collectingRegion id="EC2754DD256F16344169FE68FC9B9CF7" box="[768,863,433,459]" country="Kazakhstan" name="JAMBYL" pageId="4" pageNumber="388">JAMBYL</collectingRegion>
:
</emphasis>
<specimenCount id="38E551B6256F16344104FE6BFC589CF0" box="[877,924,434,460]" count="1" pageId="4" pageNumber="388" type="male">1 ♂</specimenCount>
:
<collectingMunicipality id="CE380045256F163441C0FE6BFBAC9CF0" box="[937,1128,434,460]" pageId="4" pageNumber="388">Aksu-Dzhabagly</collectingMunicipality>
,
<location id="2B3CCCE4256F1634461CFE6BFB0F9CF0" LSID="urn:lsid:plazi:treatment:A64A2B29256F163542FEFF4EFCB09B35:2B3CCCE4256F1634461CFE6BFB0F9CF0" box="[1141,1227,434,460]" country="Kazakhstan" municipality="Aksu-Dzhabagly" name="Koksay" pageId="4" pageNumber="388" stateProvince="JAMBYL">Koksay</location>
<collectionCode id="48F202FA256F163446BDFE6BFB2E9CF0" box="[1236,1258,434,460]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="388">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="9E8B9062256F1634469EFE6BFA5D9CF0" box="[1271,1433,434,460]" collectingDate="2001-02-07" collectionCode="V" country="Kazakhstan" pageId="4" pageNumber="388" specimenCount="1" stateProvince="JAMBYL">
<date id="5A5DBCFF256F1634469EFE6BFABC9CF0" box="[1271,1400,434,460]" pageId="4" pageNumber="388" value="2001-02-07">
<collectingDate id="4A194517256F1634469EFE6BFABC9CF0" box="[1271,1400,434,460]" pageId="4" pageNumber="388" value="2001-02-07">02.07.2001</collectingDate>
</date>
.
<collectionCode id="48F202FA256F163447ECFE6BFA5D9CF0" box="[1413,1433,434,460]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="388" type="Museum">V</collectionCode>
</materialsCitation>
. Kastcheev leg. (cSh);
<materialsCitation id="9E8B9062256F163443E7FE0CFA899CCC" box="[398,1357,469,499]" collectingDate="1994-09-06" collectorName="Kegety Gorge. &amp; S. Ovchinnikov" country="Kyrgyzstan" location="CHUY" pageId="4" pageNumber="388" specimenCount="1" specimenCount-female="1" stateProvince="CHUY">
<emphasis id="1C97462D256F163443E7FE0CFD6B9CCC" bold="true" box="[398,687,469,499]" pageId="4" pageNumber="388">
<collectingCountry id="56F4DAAF256F163443E7FE0CFD929CD3" box="[398,598,469,495]" name="Kyrgyzstan" pageId="4" pageNumber="388">KYRGYZSTAN</collectingCountry>
:
<collectingRegion id="EC2754DD256F1634400BFE0EFD619CCF" box="[610,677,471,499]" country="Kyrgyzstan" name="CHUY" pageId="4" pageNumber="388">CHÜY</collectingRegion>
:
</emphasis>
<specimenCount id="38E551B6256F163440DFFE0FFD269CCC" box="[694,738,470,496]" count="1" pageId="4" pageNumber="388" type="female">1 ♀</specimenCount>
:
<collectorName id="8316FFE9256F16344086FE0FFC509CCC" box="[751,916,470,496]" pageId="4" pageNumber="388">Kegety Gorge.</collectorName>
<date id="5A5DBCFF256F163441F3FE0FFBD89CCC" box="[922,1052,470,496]" pageId="4" pageNumber="388" value="1994-09-06">
<collectingDate id="4A194517256F163441F3FE0FFBD89CCC" box="[922,1052,470,496]" pageId="4" pageNumber="388" value="1994-09-06">09.06.1994</collectingDate>
</date>
.
<collectorName id="8316FFE9256F1634464FFE0FFB129CCC" box="[1062,1238,470,496]" pageId="4" pageNumber="388">S. Ovchinnikov</collectorName>
leg. (cSh)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="66F9C9B4256F163542AEFE20FA4199D5" lastPageId="5" lastPageNumber="389" pageId="4" pageNumber="388" type="description">
<paragraph id="2E5C9A3F256F163442AEFE20FDAF9FBC" blockId="4.[151,1437,325,641]" pageId="4" pageNumber="388">
<emphasis id="1C97462D256F163442AEFE20FEB19F2F" bold="true" box="[199,373,505,531]" pageId="4" pageNumber="388">Redescription.</emphasis>
Measurements (n = 50): HW: 0.941.01; HL: 0.740.75; OL: 0.250.26; LT: 0.170.18; AL (averaged): 3.46; PL: 0.880.94; PWmax: 1.331.41; PWmin: 1.231.25; ESL: 1.471.63; EW: 2.072.35; MTbL(averaged): 1.55; MTrL (averaged): 0.34 (MTrL 14: 0.19; MTrL 5: 0.15); AW: 2.132.36; AedL: 1.281.34; BL: 5.457.00 (
<typeStatus id="F158249D256F16344320FDBFFE759FBC" box="[329,433,614,640]" pageId="4" pageNumber="388" type="holotype">holotype</typeStatus>
:
<bibRefCitation id="4A72E7CE256F163443D2FDBFFDA39FBC" author="Zerche, L." box="[443,615,614,640]" pageId="4" pageNumber="388" pagination="105 - 142" refId="ref3633" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche (1992))</bibRefCitation>
.
</paragraph>
<caption id="7A9CCAB7256F163442FEF916FB879BDB" ID-DOI="http://doi.org/10.5281/zenodo.12758954" ID-Zenodo-Dep="12758954" box="[151,1091,1743,1767]" httpUri="https://zenodo.org/record/12758954/files/figure.png" pageId="4" pageNumber="388" startId="4.[151,250,1743,1767]" targetBox="[151,1435,664,1720]" targetPageId="4" targetType="figure">
<paragraph id="2E5C9A3F256F163442FEF916FB879BDB" blockId="4.[151,1091,1743,1767]" box="[151,1091,1743,1767]" pageId="4" pageNumber="388">
<emphasis id="1C97462D256F163442FEF916FED79BDB" bold="true" box="[151,275,1743,1767]" pageId="4" pageNumber="388">FIGURE 7.</emphasis>
Distribution of
<taxonomicName id="E9E3E1BC256F163443DFF916FD089BDB" authorityName="Shavrin" authorityYear="2024" box="[438,716,1743,1767]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="388" phylum="Arthropoda" rank="species" species="incrassatus">
<emphasis id="1C97462D256F163443DFF916FD089BDB" box="[438,716,1743,1767]" italics="true" pageId="4" pageNumber="388">Hygrodromicus incrassatus</emphasis>
</taxonomicName>
(square) and
<taxonomicName id="E9E3E1BC256F1634413EF909FC269BDB" authorityName="Zerche" authorityYear="1992" box="[855,994,1743,1767]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="388" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="1C97462D256F1634413EF909FC269BDB" box="[855,994,1743,1767]" italics="true" pageId="4" pageNumber="388">H. splendidus</emphasis>
</taxonomicName>
(circles).
</paragraph>
</caption>
<paragraph id="2E5C9A3F256F163442AEF8E0FA9A9ADF" blockId="4.[151,1437,1849,2020]" pageId="4" pageNumber="388">
Habitus as in
<figureCitation id="B6D886BA256F1634430FF8E0FE689A68" box="[358,428,1849,1876]" captionStart="FIGURES 12" captionStartId="2.[151,264,1207,1231]" captionTargetBox="[157,1430,281,1175]" captionTargetId="figure-60@2.[151,1436,270,1181]" captionTargetPageId="2" captionText="FIGURES 12. Habitus of Hygrodromicus: 1H. incrassatus (paratype), 2H. splendidus (Kazakhstan, Ketmen` Mts.). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12758946" httpUri="https://zenodo.org/record/12758946/files/figure.png" pageId="4" pageNumber="388">Fig. 2</figureCitation>
. Body reddish-brown to black, usually with distinctly paler elytra; mouthparts, antennae and legs brownish; apical segment of maxillary palpi and tarsi yellowish. Head with dense and fine isodiametric microculpture, coarser on infraorbital portions; neck with dense transverse or isodiametric sculpture; pronotum with dense and coarse isodiametric meshes, usually finer along middle and mediobasal third; scutellum with dense and fine transverse meshes; abdomen with dense and sometimes coarse isodametric or transverse microsculpture.
</paragraph>
<paragraph id="2E5C9A3F256E163542AEFF4EFDEC9C91" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">
Head 1.21.3 times as broad as long, with slightly elevated middle portion; medioapical depression deep and wide, narrowing posteriad; interocellar depression wide, relatively shallow or deep; anteocellar foveae narrow and long, strongly convergent diagonally toward level of anterior margins of eyes. Punctation irregular, fine, sometimes invisible in some specimens, denser and slightly larger between eyes; neck without visible punctation or with several punctures in middle. Ocelli small and indistinct (invisible in
<typeStatus id="F158249D256E16354154FEFEFC659C7D" box="[829,929,295,321]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
) or moderately large. Antenna reaching basal third or apical margins of elytra when reclined, with somewhat widened antennomeres; antennomere 3 slightly longer than 2, 45 slightly longer and usually indistinctly broader than 3, 610 slightly longer than 5 (sometimes antennomere 5 indistinctly shorter).
</paragraph>
<paragraph id="2E5C9A3F256E163542AEFE6EFECD9FB9" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">Pronotum markedly convex and distinctly transverse, 1.5 times as broad as long, 1.31.4 times as broad as head, from widest anterior portion strongly narrowed posteriad toward obtuse or subacute posterior angles; anterior angles widely rounded, sometimes slightly protruded anteriad; posterior margin straight or rounded, slightly narrower than posterior margin usually widely concave in middle. Punctation fine and dense, more distinct than that on head, finer in middle and sometimes with narrow impunctate median band. Scutellum without or with several fine punctures in middle.</paragraph>
<paragraph id="2E5C9A3F256E163542AEFD56FB9A9FF1" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">Elytra 1.11.3 times as broad as long. Punctation dense, distinctly larger and deeper than that on pronotum, with interstices between punctures in middle about one-two diameters of nearest punctures.</paragraph>
<paragraph id="2E5C9A3F256E163542AEFD0EFE7A9E05" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">
Abdomen slightly narrower (
<typeStatus id="F158249D256E16354064FD0EFDB69FCD" box="[525,626,727,753]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
) or broader than elytra, with two large and transverse tometose spots in the middle of abdominal tergite IV and two small spots in tergite V (some specimens witth indistinct spots); palisade fringe narrow or invisible.
</paragraph>
<paragraph id="2E5C9A3F256E163542AEFC9AFC1D992D" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">
Male.Apical margin of abdominal tergite VIII somewhat straight or rounded.Apical margin of abdominal sternite VIII sinuate. Aedeagus with small basal portion and moderately wide elongate median lobe, distinctly narrowed in apical third, slightly broadened in preapical portion and narrowed toward small rounded apex; parameres narrow, indistinctly broadened apically, usually not reaching apex of median lobe, with four short apical setae; internal sac relatively long, with two narrow sclerotized lobes in preapical portion and moderately long flagellum, spirally folded in basal portion (
<figureCitation id="B6D886BA256E163543CAFC2EFE2C992D" box="[419,488,1014,1041]" captionStart="FIGURES 36" captionStartId="3.[151,264,1842,1866]" captionTargetBox="[177,1417,713,1811]" captionTargetId="figure-228@3.[151,1436,705,1819]" captionTargetPageId="3" captionText="FIGURES 36. Aedeagus of Hygrodromicus (Figs 34: H. incrassatus (holotype); Figs. 56: H. splendidus (Kazakhstan: Ketmen` Mts.): 3, 5parameral view, 4, 6lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12758950" httpUri="https://zenodo.org/record/12758950/files/figure.png" pageId="5" pageNumber="389">Fig. 5</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B6D886BA256E163541F9FC2EFC11992D" box="[912,981,1015,1041]" captionStart="FIGURES 36" captionStartId="3.[151,264,1842,1866]" captionTargetBox="[177,1417,713,1811]" captionTargetId="figure-228@3.[151,1436,705,1819]" captionTargetPageId="3" captionText="FIGURES 36. Aedeagus of Hygrodromicus (Figs 34: H. incrassatus (holotype); Figs. 56: H. splendidus (Kazakhstan: Ketmen` Mts.): 3, 5parameral view, 4, 6lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12758950" httpUri="https://zenodo.org/record/12758950/files/figure.png" pageId="5" pageNumber="389">Fig. 6</figureCitation>
.
</paragraph>
<paragraph id="2E5C9A3F256E163542AEFBC2FF3F9965" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">Female. Apical margins of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII rounded.</paragraph>
<paragraph id="2E5C9A3F256E163542AEFBBBFA4199D5" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">
<emphasis id="1C97462D256E163542AEFBBBFE749941" bold="true" box="[199,432,1122,1149]" pageId="5" pageNumber="389">Comparative notes.</emphasis>
Based on the general shape of the aedeagus,
<taxonomicName id="E9E3E1BC256E163541C0FBBAFB879941" authorityName="Zerche" authorityYear="1992" box="[937,1091,1123,1149]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="1C97462D256E163541C0FBBAFC799940" box="[937,957,1123,1148]" italics="true" pageId="5" pageNumber="389">H</emphasis>
.
<emphasis id="1C97462D256E163541A2FBBAFB879941" box="[971,1091,1123,1149]" italics="true" pageId="5" pageNumber="389">splendidus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="E9E3E1BC256E163546BCFBBAFA529941" authorityName="Coiffait" authorityYear="1983" box="[1237,1430,1123,1149]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="cachemiricum">
<emphasis id="1C97462D256E163546BCFBBAFA529941" box="[1237,1430,1123,1149]" italics="true" pageId="5" pageNumber="389">H. cachemiricum</emphasis>
</taxonomicName>
,
<taxonomicName id="E9E3E1BC256E163542FEFB5EFEFF999C" authorityName="Shavrin" authorityYear="2024" box="[151,315,1159,1184]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="incrassatus" status="sp. n.">
<emphasis id="1C97462D256E163542FEFB5EFEFF999C" box="[151,315,1159,1184]" italics="true" pageId="5" pageNumber="389">H. incrassatus</emphasis>
</taxonomicName>
<emphasis id="1C97462D256E1635432DFB5EFE41999D" bold="true" box="[324,389,1159,1185]" pageId="5" pageNumber="389">
<taxonomicNameLabel id="07A4FB56256E1635432DFB5EFE41999D" box="[324,389,1159,1185]" pageId="5" pageNumber="389" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="E9E3E1BC256E163543FDFB5EFDFC999D" authorityName="Coiffait" authorityYear="1982" box="[404,568,1159,1185]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="1C97462D256E163543FDFB5EFDFC999D" box="[404,568,1159,1185]" italics="true" pageId="5" pageNumber="389">H. klapperichi</emphasis>
</taxonomicName>
,
<taxonomicName id="E9E3E1BC256E1635402EFB5EFD7C999C" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[583,696,1159,1184]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="1C97462D256E1635402EFB5EFD7C999C" box="[583,696,1159,1184]" italics="true" pageId="5" pageNumber="389">H. reitteri</emphasis>
</taxonomicName>
and
<taxonomicName id="E9E3E1BC256E1635409BFB5EFCA5999C" authorityName="Zerche" authorityYear="1992" box="[754,865,1159,1184]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="wrasei">
<emphasis id="1C97462D256E1635409BFB5EFCA5999C" box="[754,865,1159,1184]" italics="true" pageId="5" pageNumber="389">H. wrasei</emphasis>
</taxonomicName>
. From all these species it can be distinguished by the broader antennae, more convex pronotum, and details of the external and internal morphology of the aedeagus. Additionally, it differs from
<taxonomicName id="E9E3E1BC256E163543BDFB16FD5299D5" authorityName="Coiffait" authorityYear="1983" box="[468,662,1231,1257]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="cachemiricum">
<emphasis id="1C97462D256E163543BDFB16FD5299D5" box="[468,662,1231,1257]" italics="true" pageId="5" pageNumber="389">H. cachemiricum</emphasis>
</taxonomicName>
,
<taxonomicName id="E9E3E1BC256E163540CAFB16FC8299D5" authorityName="Coiffait" authorityYear="1982" box="[675,838,1231,1257]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="1C97462D256E163540CAFB16FC8299D5" box="[675,838,1231,1257]" italics="true" pageId="5" pageNumber="389">H. klapperichi</emphasis>
</taxonomicName>
and
<taxonomicName id="E9E3E1BC256E16354112FB16FC2F99D4" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[891,1003,1231,1256]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="1C97462D256E16354112FB16FC2F99D4" box="[891,1003,1231,1256]" italics="true" pageId="5" pageNumber="389">H. reitteri</emphasis>
</taxonomicName>
by the broader pronotum and elytra.
</paragraph>
</subSubSection>
<subSubSection id="66F9C9B4256E163542AEFB2BFEC39845" pageId="5" pageNumber="389" type="distribution">
<paragraph id="2E5C9A3F256E163542AEFB2BFCF6980D" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">
<emphasis id="1C97462D256E163542AEFB2BFEA59830" bold="true" box="[199,353,1266,1292]" pageId="5" pageNumber="389">Distribution.</emphasis>
<taxonomicName id="E9E3E1BC256E16354302FB2AFD5F9831" authorityName="Zerche" authorityYear="1992" box="[363,667,1267,1293]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="1C97462D256E16354302FB2AFD5F9831" box="[363,667,1267,1293]" italics="true" pageId="5" pageNumber="389">Hygrodromicus splendidus</emphasis>
</taxonomicName>
is known from several locality in the Middle Asia: south-western
<collectingCountry id="56F4DAAF256E163542FEFACEFED9980D" box="[151,285,1303,1329]" name="Kazakhstan" pageId="5" pageNumber="389">Kazakhstan</collectingCountry>
, north
<collectingCountry id="56F4DAAF256E16354303FACEFE2F980D" box="[362,491,1303,1329]" name="Kyrgyzstan" pageId="5" pageNumber="389">Kyrgyzstan</collectingCountry>
and north
<collectingCountry id="56F4DAAF256E1635400BFACEFD14980D" box="[610,720,1303,1329]" name="Tajikistan" pageId="5" pageNumber="389">Tajikistan</collectingCountry>
(
<figureCitation id="B6D886BA256E16354089FACEFCE0980D" box="[736,804,1303,1329]" captionStart="FIGURE 7" captionStartId="4.[151,250,1743,1767]" captionTargetBox="[151,1435,664,1720]" captionTargetId="figure-160@4.[151,1435,664,1720]" captionTargetPageId="4" captionText="FIGURE 7. Distribution of Hygrodromicus incrassatus (square) and H. splendidus (circles)." figureDoi="http://doi.org/10.5281/zenodo.12758954" httpUri="https://zenodo.org/record/12758954/files/figure.png" pageId="5" pageNumber="389">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="2E5C9A3F256E163542AEFAE3FEC39845" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">
<emphasis id="1C97462D256E163542AEFAE3FE8F9868" bold="true" box="[199,331,1338,1364]" pageId="5" pageNumber="389">Bionomics.</emphasis>
The
<typeStatus id="F158249D256E163543E6FAE2FE359869" box="[399,497,1339,1365]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
was collected at elevation
<quantity id="E91B37DA256E16354153FAE2FC1F9869" box="[826,987,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="389" unit="m" value="3500.0" valueMax="4000.0" valueMin="3000.0">30004000 m</quantity>
a.s.l. The detailed bionomical data is unknown.
</paragraph>
</subSubSection>
<subSubSection id="66F9C9B4256E163542AEFA5AFCB09B35" pageId="5" pageNumber="389" type="discussion">
<paragraph id="2E5C9A3F256E163542AEFA5AFCB09B35" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">
<emphasis id="1C97462D256E163542AEFA5AFEFF98A1" bold="true" box="[199,315,1411,1437]" pageId="5" pageNumber="389">Remarks.</emphasis>
<taxonomicName id="E9E3E1BC256E1635432FFA5AFDB298A1" authorityName="Zerche" authorityYear="1992" box="[326,630,1411,1437]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="1C97462D256E1635432FFA5AFDB298A1" box="[326,630,1411,1437]" italics="true" pageId="5" pageNumber="389">Hygrodromicus splendidus</emphasis>
</taxonomicName>
was originally described based on the
<typeStatus id="F158249D256E1635462DFA5AFB6298A1" box="[1092,1190,1411,1437]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
from
<collectingCountry id="56F4DAAF256E1635469BFA5AFAA498A1" box="[1266,1376,1411,1437]" name="Tajikistan" pageId="5" pageNumber="389">Tajikistan</collectingCountry>
(“… Pamir, Muksu Gebiet… Hochflâche bei Kishlak Ljarsch…”). The specimens studied by me are conspecific to the
<typeStatus id="F158249D256E163542FEFA12FF3898D9" box="[151,252,1483,1509]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
, but have some insignificant morphological differences. Thus, I decided to redescribed this species. It is here recorded from
<collectingCountry id="56F4DAAF256E1635431CFA36FE3C9B35" box="[373,504,1519,1545]" name="Kazakhstan" pageId="5" pageNumber="389">Kazakhstan</collectingCountry>
and
<collectingCountry id="56F4DAAF256E16354047FA36FD6B9B35" box="[558,687,1519,1545]" name="Kyrgyzstan" pageId="5" pageNumber="389">Kyrgyzstan</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>