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<document id="565C8F36C3F73CF79A8B0D7886EED40C" ID-DOI="10.11646/zootaxa.5481.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="12758762" ID-ZooBank="798238C5-D310-4FA1-BF2D-6D365250655C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721215777745" checkinUser="plazi" docAuthor="Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Van Dam, Alex R., Helmick, Ericka E., Bartlett, Charles R., Martinez Aponte, Laura V., Ruiz, Alfredo Rodriguez, Bloch, Melody &amp; Bahder, Brian W." docDate="2024" docId="039B0E04FF88FFE6FF538490FB690EA4" docLanguage="en" docName="zootaxa.5481.3.3.pdf" docOrigin="Zootaxa 5481 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5481.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Colpoptera stonei Bahder &amp; Bartlett 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="350" masterDocId="FFA2767CFF8CFFEFFFC4810EFFF70C0D" masterDocTitle="A new species of planthopper in the genus Colpoptera (Hemiptera: Fulgoroidea: Nogodinidae) from the Caribbean coast of Costa Rica" masterLastPageNumber="352" masterPageNumber="341" pageNumber="345" updateTime="1721216188411" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6D9F5D060975CB0350E6485E28F6F05A">A new species of planthopper in the genus Colpoptera (Hemiptera: Fulgoroidea: Nogodinidae) from the Caribbean coast of Costa Rica</mods:title>
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<mods:namePart id="EFE298C48958208593A0DF0C78054361">Barrantes Barrantes, Edwin A.</mods:namePart>
<mods:affiliation id="EBF94CB85EBDA4DD2DA3C4F1CD6243D3">Universidad de Costa Rica - Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica</mods:affiliation>
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<mods:namePart id="5C2BED750E7673F12EFFF125A3F5F37D">Zumbado Echavarria, Marco A.</mods:namePart>
<mods:affiliation id="614C2984E6D76D529111BABA62089B19">Universidad de Costa Rica - Sede San Ramón, Departamento de Ciencias Naturales, de la Iglesia el Tremedal 400 mts al Oeste carretera hacia San Pedro, San Ramón, Alajuela, Costa Rica</mods:affiliation>
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<mods:affiliation id="1460FB5AD9904BD3D30D78488EBA3B3C">University of Florida, Department of Entomology and Nematology - Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA</mods:affiliation>
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<mods:affiliation id="13BDEC941AB5427ECAD4E05813A2339B">University of Delaware, Department of Entomology and Wildlife Ecology, 250 Townsend Hall, Newark, DE 19716 - 2160, USA</mods:affiliation>
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<mods:namePart id="0B70AFA5E7ABE3A76E0DA53E6DEC1168">Ruiz, Alfredo Rodriguez</mods:namePart>
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<mods:namePart id="FA423B78FC474E7E37DAD3BF44E44103">Bloch, Melody</mods:namePart>
<mods:affiliation id="CD65EDF3BA668A46FE9FA2ED7C71F26C">University of Florida, Department of Entomology and Nematology - Fort Lauderdale Research and Education Center; 3205 College Ave., Davie, FL 33314 - 7719, USA</mods:affiliation>
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<mods:namePart id="6FF95012C1C098BAF02F369D6D353E99">Bahder, Brian W.</mods:namePart>
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<heading id="D0C5087EFF88FFEBFF538490FD3209B4" bold="true" box="[151,709,1438,1465]" fontSize="11" level="1" pageId="4" pageNumber="345" reason="1">
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<taxonomicName id="4C32C491FF88FFEBFF538490FDAD09B4" authority="Bahder &amp; Bartlett" authorityName="Bahder &amp; Bartlett" authorityYear="2024" box="[151,602,1438,1465]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
<emphasis id="B9466300FF88FFEBFF538490FE9B09B5" bold="true" box="[151,364,1438,1465]" italics="true" pageId="4" pageNumber="345">Colpoptera stonei</emphasis>
Bahder &amp; Bartlett
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,
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<paragraph id="8B8DBF12FF88FFEBFF5384CDFED809D1" blockId="4.[151,709,1438,1501]" box="[151,303,1474,1501]" pageId="4" pageNumber="345">
(
<figureCitation id="1309A397FF88FFEBFF5B84CDFED009D1" box="[159,295,1474,1501]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStartId-0="6.[151,250,1341,1365]" captionStartId-1="6.[151,250,1923,1947]" captionStartId-2="7.[151,250,751,775]" captionStartId-3="7.[151,250,1454,1478]" captionStartId-4="8.[151,250,954,978]" captionTargetBox-0="[151,1436,181,1317]" captionTargetBox-1="[156,1434,1395,1893]" captionTargetBox-2="[151,1435,181,727]" captionTargetBox-3="[197,1376,799,1430]" captionTargetBox-4="[158,1423,181,930]" captionTargetId-0="figure-24@6.[151,1436,181,1317]" captionTargetId-1="figure-43@6.[151,1435,1389,1899]" captionTargetId-2="figure-17@7.[151,1435,181,727]" captionTargetId-3="figure-38@7.[197,1390,799,1430]" captionTargetId-4="figure-24@8.[158,1429,181,930]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 2. Adult habitus Colpoptera stonei sp. nov. (A) Male lateral view; (B) male dorsal view." captionText-1="FIGURE 3. Adult Colpoptera stonei sp. nov.; (A) Head, pronotum, mesonotum dorsal view; (B) head and pronotum lateral view;(C) head frontal view. Scale = 1 mm." captionText-2="FIGURE 4. Forewing venation of Colpoptera stonei sp. nov. Vein labeling nomenclature following Bourgoin et al. (2015)." captionText-3="FIGURE 5. Adult male Colpoptera stonei sp. nov., terminalia in left lateral view." captionText-4="FIGURE 6. Aedeagus of Colpoptera stonei sp. nov.: (A) right lateral view; (B) left lateral view; (C) dorsal view; (D) ventral view." figureDoi-0="http://doi.org/10.5281/zenodo.12758768" figureDoi-1="http://doi.org/10.5281/zenodo.12758770" figureDoi-2="http://doi.org/10.5281/zenodo.12758774" figureDoi-3="http://doi.org/10.5281/zenodo.12758776" figureDoi-4="http://doi.org/10.5281/zenodo.12758780" httpUri-0="https://zenodo.org/record/12758768/files/figure.png" httpUri-1="https://zenodo.org/record/12758770/files/figure.png" httpUri-2="https://zenodo.org/record/12758774/files/figure.png" httpUri-3="https://zenodo.org/record/12758776/files/figure.png" httpUri-4="https://zenodo.org/record/12758780/files/figure.png" pageId="4" pageNumber="345">Figures 26</figureCitation>
)
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<emphasis id="B9466300FF88FFEBFF538705FEC10A29" bold="true" box="[151,310,1546,1573]" pageId="4" pageNumber="345">Type locality.</emphasis>
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,
<collectingRegion id="49F671F0FF88FFEBFE0E8705FD750A28" box="[458,642,1547,1573]" country="Costa Rica" name="Limon" pageId="4" pageNumber="345">Limón Province</collectingRegion>
,
<collectingCountry id="F325FF82FF88FFEBFD498705FCF00A28" box="[653,775,1547,1573]" name="Costa Rica" pageId="4" pageNumber="345">Costa Rica</collectingCountry>
(
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<figureCitation id="1309A397FF88FFEBFBE38705FB9E0A28" box="[1063,1129,1547,1573]" captionStart="FIGURE 1" captionStartId="2.[151,250,1522,1546]" captionTargetBox="[151,1436,181,1496]" captionTargetId="figure-24@2.[151,1436,181,1498]" captionTargetPageId="2" captionText="FIGURE 1. Habitat and locality of Colpoptera stonei sp. nov.; red circle = Heredia locality, orange circle = Limón locality, habitat photo from Limón locality." figureDoi="http://doi.org/10.5281/zenodo.12758764" httpUri="https://zenodo.org/record/12758764/files/figure.png" pageId="4" pageNumber="345">Fig. 1</figureCitation>
)
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.
</paragraph>
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<subSubSection id="C328EC99FF88FFEBFF038721FED20AB8" pageId="4" pageNumber="345" type="etymology">
<paragraph id="8B8DBF12FF88FFEBFF038721FED20AB8" blockId="4.[151,1436,1546,2005]" pageId="4" pageNumber="345">
<emphasis id="B9466300FF88FFEBFF038721FEB90A44" bold="true" box="[199,334,1583,1609]" pageId="4" pageNumber="345">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C32C491FF88FFEBFDF88721FD7E0A45" authorityName="Bahder &amp; Bartlett" authorityYear="2024" box="[572,649,1583,1609]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="stonei">
<emphasis id="B9466300FF88FFEBFD828721FD7E0A45" box="[582,649,1583,1608]" italics="true" pageId="4" pageNumber="345">stonei</emphasis>
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is a patronym in honor of Dan and Eugene Stone, very close friends (almost older brothers) to the senior author. Their friendship, comedic relief, being fellow bandmates in Shutter and overall love and support was instrumental in his development and success in early academic years. Rock on Dan and Eugene!
</paragraph>
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<subSubSection id="C328EC99FF88FFEBFF0387B0FDDC0B64" pageId="4" pageNumber="345" type="diagnosis">
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<emphasis id="B9466300FF88FFEBFF0387B0FEB50AD5" bold="true" box="[199,322,1726,1752]" pageId="4" pageNumber="345">Diagnosis.</emphasis>
Pale species, body generally uniformly testaceous, slight fuscous mottling on pronotum and mesonotum, forewing transparent, slight fuscous patch at apex, face without markings.Aedeagus lightly sclerotized, nearly symmetrical, pair of strongly bifurcated dorsal processes of moderate length arising subapically on lateral margins with ventral bifurcations emerging mesad near midline (not on outer lateral margin) and pair of shorter, curved processes on ventral margin.
</paragraph>
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<paragraph id="8B8DBF12FF88FFEBFF03867CFB490BBC" blockId="4.[151,1436,1546,2005]" pageId="4" pageNumber="345">
<emphasis id="B9466300FF88FFEBFF03867CFEAE0B81" bold="true" box="[199,345,1906,1932]" pageId="4" pageNumber="345">Description.</emphasis>
Pale species, body generally uniformly testaceous, slight fuscous mottling on pronotum and mesonotum, forewing transparent, slight fuscous patch at apex, face without markings (
<figureCitation id="1309A397FF88FFEBFBA88699FB460BBC" box="[1132,1201,1943,1969]" captionStart="FIGURE 2" captionStartId="6.[151,250,1341,1365]" captionTargetBox="[151,1436,181,1317]" captionTargetId="figure-24@6.[151,1436,181,1317]" captionTargetPageId="6" captionText="FIGURE 2. Adult habitus Colpoptera stonei sp. nov. (A) Male lateral view; (B) male dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758768" httpUri="https://zenodo.org/record/12758768/files/figure.png" pageId="4" pageNumber="345">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8B8DBF12FF88FFEBFF0386B5FAD70BD8" blockId="4.[151,1436,1546,2005]" box="[199,1312,1978,2005]" pageId="4" pageNumber="345">
<emphasis id="B9466300FF88FFEBFF0386B5FEC10BD8" box="[199,310,1979,2005]" italics="true" pageId="4" pageNumber="345">Structure.</emphasis>
Body length with with wings; males (
<emphasis id="B9466300FF88FFEBFD2686B5FD070BD9" box="[738,752,1979,2004]" italics="true" pageId="4" pageNumber="345">n</emphasis>
=4)
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, females (
<emphasis id="B9466300FF88FFEBFBDE86B5FBCC0BD9" box="[1050,1083,1979,2004]" italics="true" pageId="4" pageNumber="345">n=</emphasis>
6) 4.44.6 (
<tableCitation id="C6B08AA9FF88FFEBFB7886B5FAE50BD8" box="[1212,1298,1979,2005]" captionStart="TABLE 3" captionStartId="5.[152,244,152,176]" captionText="TABLE 3. Biometric data for Colpoptera stonei sp. nov." pageId="4" pageNumber="345">Table 3</tableCitation>
).
</paragraph>
<caption id="DF4DEF9AFF89FFEAFF5C8196FD0B0CBC" box="[152,764,152,177]" pageId="5" pageNumber="346" startId="5.[152,244,152,176]" targetType="table">
<paragraph id="8B8DBF12FF89FFEAFF5C8196FD0B0CBC" blockId="5.[152,764,152,177]" box="[152,764,152,177]" pageId="5" pageNumber="346">
<emphasis id="B9466300FF89FFEAFF5C8196FEF80CBD" bold="true" box="[152,271,152,176]" pageId="5" pageNumber="346">TABLE 3.</emphasis>
Biometric data for
<taxonomicName id="4C32C491FF89FFEAFE248196FD570CBD" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[480,672,152,176]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
<emphasis id="B9466300FF89FFEAFE248196FD570CBD" box="[480,672,152,176]" italics="true" pageId="5" pageNumber="346">Colpoptera stonei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A275DE7BFF89FFEAFD628197FD0B0CBC" box="[678,764,153,177]" pageId="5" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8B8DBF12FF89FFEAFF6781CFFAA50F50" pageId="5" pageNumber="346">
<table id="F9324DB2FF890010FF6781CFFA9B0F50" box="[163,1388,193,861]" gridcols="5" gridrows="17" pageId="5" pageNumber="346">
<tr id="3502BD50FF890010FF6781CFFA9B0CD4" box="[163,1388,193,217]" gridrow="0" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF6781CFFE4C0CD4" box="[163,443,193,217]" gridcol="0" gridrow="0" pageId="5" pageNumber="346">Character</th>
<th id="76D3D42CFF890010FDAB81CFFC4A0CD4" box="[623,957,193,217]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="5" pageNumber="346">
Male (
<emphasis id="B9466300FF89FFEAFCD481CFFCD90CD5" box="[784,814,193,216]" italics="true" pageId="5" pageNumber="346">n=</emphasis>
4)
</th>
<th id="76D3D42CFF890010FBE981CFFA9B0CD4" box="[1069,1388,193,217]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="5" pageNumber="346">
Female (
<emphasis id="B9466300FF89FFEAFB1381CFFB130CD5" box="[1239,1252,193,216]" italics="true" pageId="5" pageNumber="346">n</emphasis>
=6)
</th>
</tr>
<tr id="3502BD50FF890010FF6781E7FA9B0D0C" box="[163,1388,233,257]" gridrow="1" pageId="5" pageNumber="346" rowspan-0="1">
<td id="76D3D42CFF890010FDAB81E7FD4C0D0C" box="[623,699,233,257]" gridcol="1" gridrow="1" pageId="5" pageNumber="346">Range</td>
<td id="76D3D42CFF890010FCF481E7FC4A0D0C" box="[816,957,233,257]" gridcol="2" gridrow="1" pageId="5" pageNumber="346">Average ± SE</td>
<td id="76D3D42CFF890010FBE981E7FB8E0D0C" box="[1069,1145,233,257]" gridcol="3" gridrow="1" pageId="5" pageNumber="346">Range</td>
<td id="76D3D42CFF890010FB2881E7FA9B0D0C" box="[1260,1388,233,257]" gridcol="4" gridrow="1" pageId="5" pageNumber="346">Average±SE</td>
</tr>
<tr id="3502BD50FF890010FF67801FFA9B0D24" box="[163,1388,273,297]" gridrow="2" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF67801FFE4C0D24" box="[163,443,273,297]" gridcol="0" gridrow="2" pageId="5" pageNumber="346">Body length, with wings</th>
<td id="76D3D42CFF890010FDAB801FFD4C0D24" box="[623,699,273,297]" gridcol="1" gridrow="2" pageId="5" pageNumber="346">4.24.3</td>
<td id="76D3D42CFF890010FCF4801FFC4A0D24" box="[816,957,273,297]" gridcol="2" gridrow="2" pageId="5" pageNumber="346">4.3±0.1</td>
<td id="76D3D42CFF890010FBE9801FFB8E0D24" box="[1069,1145,273,297]" gridcol="3" gridrow="2" pageId="5" pageNumber="346">4.44.6</td>
<td id="76D3D42CFF890010FB28801FFA9B0D24" box="[1260,1388,273,297]" gridcol="4" gridrow="2" pageId="5" pageNumber="346">4.5±0.1</td>
</tr>
<tr id="3502BD50FF890010FF678037FA9B0D5F" box="[163,1388,313,338]" gridrow="3" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF678037FE4C0D5F" box="[163,443,313,338]" gridcol="0" gridrow="3" pageId="5" pageNumber="346">Body length, no wings</th>
<td id="76D3D42CFF890010FDAB8037FD4C0D5F" box="[623,699,313,338]" gridcol="1" gridrow="3" pageId="5" pageNumber="346">2.52.5</td>
<td id="76D3D42CFF890010FCF48037FC4A0D5F" box="[816,957,313,338]" gridcol="2" gridrow="3" pageId="5" pageNumber="346">2.5±0.0</td>
<td id="76D3D42CFF890010FBE98037FB8E0D5F" box="[1069,1145,313,338]" gridcol="3" gridrow="3" pageId="5" pageNumber="346">2.82.9</td>
<td id="76D3D42CFF890010FB288037FA9B0D5F" box="[1260,1388,313,338]" gridcol="4" gridrow="3" pageId="5" pageNumber="346">2.9±0.1</td>
</tr>
<tr id="3502BD50FF890010FF67806CFA9B0D77" box="[163,1388,354,378]" gridrow="4" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF67806CFE4C0D77" box="[163,443,354,378]" gridcol="0" gridrow="4" pageId="5" pageNumber="346">Forewing length</th>
<td id="76D3D42CFF890010FDAB806CFD4C0D77" box="[623,699,354,378]" gridcol="1" gridrow="4" pageId="5" pageNumber="346">3.83.8</td>
<td id="76D3D42CFF890010FCF4806CFC4A0D77" box="[816,957,354,378]" gridcol="2" gridrow="4" pageId="5" pageNumber="346">3.8±0.0</td>
<td id="76D3D42CFF890010FBE9806CFB8E0D77" box="[1069,1145,354,378]" gridcol="3" gridrow="4" pageId="5" pageNumber="346">4.04.2</td>
<td id="76D3D42CFF890010FB28806CFA9B0D77" box="[1260,1388,354,378]" gridcol="4" gridrow="4" pageId="5" pageNumber="346">4.1±0.1</td>
</tr>
<tr id="3502BD50FF890010FF678084FA9B0DAF" box="[163,1388,394,418]" gridrow="5" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF678084FE4C0DAF" box="[163,443,394,418]" gridcol="0" gridrow="5" pageId="5" pageNumber="346">Vertex length</th>
<td id="76D3D42CFF890010FDAB8084FD4C0DAF" box="[623,699,394,418]" gridcol="1" gridrow="5" pageId="5" pageNumber="346">0.10.1</td>
<td id="76D3D42CFF890010FCF48084FC4A0DAF" box="[816,957,394,418]" gridcol="2" gridrow="5" pageId="5" pageNumber="346">0.1±0.0</td>
<td id="76D3D42CFF890010FBE98084FB8E0DAF" box="[1069,1145,394,418]" gridcol="3" gridrow="5" pageId="5" pageNumber="346">0.10.1</td>
<td id="76D3D42CFF890010FB288084FA9B0DAF" box="[1260,1388,394,418]" gridcol="4" gridrow="5" pageId="5" pageNumber="346">0.1±0.0</td>
</tr>
<tr id="3502BD50FF890010FF6780BCFA9B0DC7" box="[163,1388,434,458]" gridrow="6" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF6780BCFE4C0DC7" box="[163,443,434,458]" gridcol="0" gridrow="6" pageId="5" pageNumber="346">Vertex width, basal margin</th>
<td id="76D3D42CFF890010FDAB80BCFD4C0DC7" box="[623,699,434,458]" gridcol="1" gridrow="6" pageId="5" pageNumber="346">0.30.3</td>
<td id="76D3D42CFF890010FCF480BCFC4A0DC7" box="[816,957,434,458]" gridcol="2" gridrow="6" pageId="5" pageNumber="346">0.3±0.0</td>
<td id="76D3D42CFF890010FBE980BCFB8E0DC7" box="[1069,1145,434,458]" gridcol="3" gridrow="6" pageId="5" pageNumber="346">0.30.3</td>
<td id="76D3D42CFF890010FB2880BCFA9B0DC7" box="[1260,1388,434,458]" gridcol="4" gridrow="6" pageId="5" pageNumber="346">0.3±0.0</td>
</tr>
<tr id="3502BD50FF890010FF6780D4FA9B0DFE" box="[163,1388,474,499]" gridrow="7" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF6780D4FE4C0DFE" box="[163,443,474,499]" gridcol="0" gridrow="7" pageId="5" pageNumber="346">Vertex width, distal margin</th>
<td id="76D3D42CFF890010FDAB80D4FD4C0DFE" box="[623,699,474,499]" gridcol="1" gridrow="7" pageId="5" pageNumber="346">0.30.3</td>
<td id="76D3D42CFF890010FCF480D4FC4A0DFE" box="[816,957,474,499]" gridcol="2" gridrow="7" pageId="5" pageNumber="346">0.3±0.0</td>
<td id="76D3D42CFF890010FBE980D4FB8E0DFE" box="[1069,1145,474,499]" gridcol="3" gridrow="7" pageId="5" pageNumber="346">0.30.3</td>
<td id="76D3D42CFF890010FB2880D4FA9B0DFE" box="[1260,1388,474,499]" gridcol="4" gridrow="7" pageId="5" pageNumber="346">0.3±0.0</td>
</tr>
<tr id="3502BD50FF890010FF67830DFA9B0E16" box="[163,1388,515,539]" gridrow="8" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF67830DFE4C0E16" box="[163,443,515,539]" gridcol="0" gridrow="8" pageId="5" pageNumber="346">Pronotum length, midline</th>
<td id="76D3D42CFF890010FDAB830DFD4C0E16" box="[623,699,515,539]" gridcol="1" gridrow="8" pageId="5" pageNumber="346">0.20.2</td>
<td id="76D3D42CFF890010FCF4830DFC4A0E16" box="[816,957,515,539]" gridcol="2" gridrow="8" pageId="5" pageNumber="346">0.2±0.0</td>
<td id="76D3D42CFF890010FBE9830DFB8E0E16" box="[1069,1145,515,539]" gridcol="3" gridrow="8" pageId="5" pageNumber="346">0.20.2</td>
<td id="76D3D42CFF890010FB28830DFA9B0E16" box="[1260,1388,515,539]" gridcol="4" gridrow="8" pageId="5" pageNumber="346">0.2±0.0</td>
</tr>
<tr id="3502BD50FF890010FF678325FA9B0E4E" box="[163,1388,555,579]" gridrow="9" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF678325FE4C0E4E" box="[163,443,555,579]" gridcol="0" gridrow="9" pageId="5" pageNumber="346">Mesonotum length, midline</th>
<td id="76D3D42CFF890010FDAB8325FD4C0E4E" box="[623,699,555,579]" gridcol="1" gridrow="9" pageId="5" pageNumber="346">0.70.8</td>
<td id="76D3D42CFF890010FCF48325FC4A0E4E" box="[816,957,555,579]" gridcol="2" gridrow="9" pageId="5" pageNumber="346">0.8±0.0</td>
<td id="76D3D42CFF890010FBE98325FB8E0E4E" box="[1069,1145,555,579]" gridcol="3" gridrow="9" pageId="5" pageNumber="346">0.80.9</td>
<td id="76D3D42CFF890010FB288325FA9B0E4E" box="[1260,1388,555,579]" gridcol="4" gridrow="9" pageId="5" pageNumber="346">0.9±0.1</td>
</tr>
<tr id="3502BD50FF890010FF67835DFA9B0E66" box="[163,1388,595,619]" gridrow="10" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF67835DFE4C0E66" box="[163,443,595,619]" gridcol="0" gridrow="10" pageId="5" pageNumber="346">Mesonotum width</th>
<td id="76D3D42CFF890010FDAB835DFD4C0E66" box="[623,699,595,619]" gridcol="1" gridrow="10" pageId="5" pageNumber="346">0.80.8</td>
<td id="76D3D42CFF890010FCF4835DFC4A0E66" box="[816,957,595,619]" gridcol="2" gridrow="10" pageId="5" pageNumber="346">0.8±0.0</td>
<td id="76D3D42CFF890010FBE9835DFB8E0E66" box="[1069,1145,595,619]" gridcol="3" gridrow="10" pageId="5" pageNumber="346">0.80.8</td>
<td id="76D3D42CFF890010FB28835DFA9B0E66" box="[1260,1388,595,619]" gridcol="4" gridrow="10" pageId="5" pageNumber="346">0.8±0.0</td>
</tr>
<tr id="3502BD50FF890010FF678375FA9B0E9E" box="[163,1388,635,659]" gridrow="11" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF678375FE4C0E9E" box="[163,443,635,659]" gridcol="0" gridrow="11" pageId="5" pageNumber="346">Frons width, dorsal margin</th>
<td id="76D3D42CFF890010FDAB8375FD4C0E9E" box="[623,699,635,659]" gridcol="1" gridrow="11" pageId="5" pageNumber="346">0.40.5</td>
<td id="76D3D42CFF890010FCF48375FC4A0E9E" box="[816,957,635,659]" gridcol="2" gridrow="11" pageId="5" pageNumber="346">0.5±0.0</td>
<td id="76D3D42CFF890010FBE98375FB8E0E9E" box="[1069,1145,635,659]" gridcol="3" gridrow="11" pageId="5" pageNumber="346">0.40.5</td>
<td id="76D3D42CFF890010FB288375FA9B0E9E" box="[1260,1388,635,659]" gridcol="4" gridrow="11" pageId="5" pageNumber="346">0.5±0.0</td>
</tr>
<tr id="3502BD50FF890010FF6783AAFA9B0EB1" box="[163,1388,676,700]" gridrow="12" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF6783AAFE4C0EB1" box="[163,443,676,700]" gridcol="0" gridrow="12" pageId="5" pageNumber="346">Frons width, clypeal suture</th>
<td id="76D3D42CFF890010FDAB83AAFD4C0EB1" box="[623,699,676,700]" gridcol="1" gridrow="12" pageId="5" pageNumber="346">0.20.3</td>
<td id="76D3D42CFF890010FCF483AAFC4A0EB1" box="[816,957,676,700]" gridcol="2" gridrow="12" pageId="5" pageNumber="346">0.3±0.0</td>
<td id="76D3D42CFF890010FBE983AAFB8E0EB1" box="[1069,1145,676,700]" gridcol="3" gridrow="12" pageId="5" pageNumber="346">0.20.3</td>
<td id="76D3D42CFF890010FB2883AAFA9B0EB1" box="[1260,1388,676,700]" gridcol="4" gridrow="12" pageId="5" pageNumber="346">0.3±0.0</td>
</tr>
<tr id="3502BD50FF890010FF6783C2FA9B0EE9" box="[163,1388,716,740]" gridrow="13" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF6783C2FE4C0EE9" box="[163,443,716,740]" gridcol="0" gridrow="13" pageId="5" pageNumber="346">Frons width, widest</th>
<td id="76D3D42CFF890010FDAB83C2FD4C0EE9" box="[623,699,716,740]" gridcol="1" gridrow="13" pageId="5" pageNumber="346">0.50.5</td>
<td id="76D3D42CFF890010FCF483C2FC4A0EE9" box="[816,957,716,740]" gridcol="2" gridrow="13" pageId="5" pageNumber="346">0.5±0.0</td>
<td id="76D3D42CFF890010FBE983C2FB8E0EE9" box="[1069,1145,716,740]" gridcol="3" gridrow="13" pageId="5" pageNumber="346">0.50.5</td>
<td id="76D3D42CFF890010FB2883C2FA9B0EE9" box="[1260,1388,716,740]" gridcol="4" gridrow="13" pageId="5" pageNumber="346">0.5±0.0</td>
</tr>
<tr id="3502BD50FF890010FF6783FAFA9B0F01" box="[163,1388,756,780]" gridrow="14" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF6783FAFE4C0F01" box="[163,443,756,780]" gridcol="0" gridrow="14" pageId="5" pageNumber="346">Frons width, narrowest</th>
<td id="76D3D42CFF890010FDAB83FAFD4C0F01" box="[623,699,756,780]" gridcol="1" gridrow="14" pageId="5" pageNumber="346">0.40.5</td>
<td id="76D3D42CFF890010FCF483FAFC4A0F01" box="[816,957,756,780]" gridcol="2" gridrow="14" pageId="5" pageNumber="346">0.5±0.0</td>
<td id="76D3D42CFF890010FBE983FAFB8E0F01" box="[1069,1145,756,780]" gridcol="3" gridrow="14" pageId="5" pageNumber="346">0.40.5</td>
<td id="76D3D42CFF890010FB2883FAFA9B0F01" box="[1260,1388,756,780]" gridcol="4" gridrow="14" pageId="5" pageNumber="346">0.5±0.0</td>
</tr>
<tr id="3502BD50FF890010FF678212FA9B0F39" box="[163,1388,796,820]" gridrow="15" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF678212FE4C0F39" box="[163,443,796,820]" gridcol="0" gridrow="15" pageId="5" pageNumber="346">Frons length, midline</th>
<td id="76D3D42CFF890010FDAB8212FD4C0F39" box="[623,699,796,820]" gridcol="1" gridrow="15" pageId="5" pageNumber="346">0.60.6</td>
<td id="76D3D42CFF890010FCF48212FC4A0F39" box="[816,957,796,820]" gridcol="2" gridrow="15" pageId="5" pageNumber="346">0.6±0.0</td>
<td id="76D3D42CFF890010FBE98212FB8E0F39" box="[1069,1145,796,820]" gridcol="3" gridrow="15" pageId="5" pageNumber="346">0.60.6</td>
<td id="76D3D42CFF890010FB288212FA9B0F39" box="[1260,1388,796,820]" gridcol="4" gridrow="15" pageId="5" pageNumber="346">0.6±0.0</td>
</tr>
<tr id="3502BD50FF890010FF67824BFA9B0F50" box="[163,1388,837,861]" gridrow="16" pageId="5" pageNumber="346">
<th id="76D3D42CFF890010FF67824BFE4C0F50" box="[163,443,837,861]" gridcol="0" gridrow="16" pageId="5" pageNumber="346">Clypeus length</th>
<td id="76D3D42CFF890010FDAB824BFD4C0F50" box="[623,699,837,861]" gridcol="1" gridrow="16" pageId="5" pageNumber="346">0.20.2</td>
<td id="76D3D42CFF890010FCF4824BFC4A0F50" box="[816,957,837,861]" gridcol="2" gridrow="16" pageId="5" pageNumber="346">0.2±0.0</td>
<td id="76D3D42CFF890010FBE9824BFB8E0F50" box="[1069,1145,837,861]" gridcol="3" gridrow="16" pageId="5" pageNumber="346">0.30.3</td>
<td id="76D3D42CFF890010FB28824BFA9B0F50" box="[1260,1388,837,861]" gridcol="4" gridrow="16" pageId="5" pageNumber="346">0.3±0.0</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8DBF12FF89FFEAFF0382ADFD66087C" blockId="5.[151,1437,931,2001]" pageId="5" pageNumber="346">
<emphasis id="B9466300FF89FFEAFF0382ADFEF30FB0" box="[199,260,931,957]" italics="true" pageId="5" pageNumber="346">Head</emphasis>
. In dorsal view, vertex nearly rectangular, anterior margin linear, approximately level with anterior margin of eyes, posterior margin concave, median carina present, transverse carina at fastigium evident (
<figureCitation id="1309A397FF89FFEAFB0882C9FAD20FEC" box="[1228,1317,967,993]" captionStart="FIGURE 3" captionStartId="6.[151,250,1923,1947]" captionTargetBox="[156,1434,1395,1893]" captionTargetId="figure-43@6.[151,1435,1389,1899]" captionTargetPageId="6" captionText="FIGURE 3. Adult Colpoptera stonei sp. nov.; (A) Head, pronotum, mesonotum dorsal view; (B) head and pronotum lateral view;(C) head frontal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12758770" httpUri="https://zenodo.org/record/12758770/files/figure.png" pageId="5" pageNumber="346">Fig. 3A</figureCitation>
). In lateral view, generally rounded, slightly angled at fastigium, vertex slightly projected above eyes, frons curved dorsad, becoming linear near ventral margin of eyes (
<figureCitation id="1309A397FF89FFEAFD748501FCFB0824" box="[688,780,1039,1065]" captionStart="FIGURE 3" captionStartId="6.[151,250,1923,1947]" captionTargetBox="[156,1434,1395,1893]" captionTargetId="figure-43@6.[151,1435,1389,1899]" captionTargetPageId="6" captionText="FIGURE 3. Adult Colpoptera stonei sp. nov.; (A) Head, pronotum, mesonotum dorsal view; (B) head and pronotum lateral view;(C) head frontal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12758770" httpUri="https://zenodo.org/record/12758770/files/figure.png" pageId="5" pageNumber="346">Fig. 3B</figureCitation>
). In frontal view, transverse carina at fastigium evident, lateral margins expanding from dorsal margin, reaching widest point just below antennae then constricting strongly, frontoclypeal suture curved dorsad (
<figureCitation id="1309A397FF89FFEAFDE88559FD73087C" box="[556,644,1111,1137]" captionStart="FIGURE 3" captionStartId="6.[151,250,1923,1947]" captionTargetBox="[156,1434,1395,1893]" captionTargetId="figure-43@6.[151,1435,1389,1899]" captionTargetPageId="6" captionText="FIGURE 3. Adult Colpoptera stonei sp. nov.; (A) Head, pronotum, mesonotum dorsal view; (B) head and pronotum lateral view;(C) head frontal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12758770" httpUri="https://zenodo.org/record/12758770/files/figure.png" pageId="5" pageNumber="346">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="8B8DBF12FF89FFEAFF038575FC430960" blockId="5.[151,1437,931,2001]" pageId="5" pageNumber="346">
<emphasis id="B9466300FF89FFEAFF038575FEEF0898" box="[199,280,1147,1173]" italics="true" pageId="5" pageNumber="346">Thorax</emphasis>
. Pronotum in dorsal view widest at midpoint with anterior margin moderately convex, posterior margin moderately concave, tricarinate, carina at midline continuous from anterior to posterior margin, lateral carinae extending from anterior margin of pronotum, laterad, becoming obsolete ventrally (
<figureCitation id="1309A397FF89FFEAFB6285CDFAF608D0" box="[1190,1281,1219,1245]" captionStart="FIGURE 3" captionStartId="6.[151,250,1923,1947]" captionTargetBox="[156,1434,1395,1893]" captionTargetId="figure-43@6.[151,1435,1389,1899]" captionTargetPageId="6" captionText="FIGURE 3. Adult Colpoptera stonei sp. nov.; (A) Head, pronotum, mesonotum dorsal view; (B) head and pronotum lateral view;(C) head frontal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12758770" httpUri="https://zenodo.org/record/12758770/files/figure.png" pageId="5" pageNumber="346">Fig. 3A</figureCitation>
). Mesonotum approximately as wide as long at midpoint, tricarinate at anterior margin, two additional carinae arising at mid carina near anterior margin, extending diagonally to just beyond lateral carinae, slightly sinuate (
<figureCitation id="1309A397FF89FFEAFB088405FAD20928" box="[1228,1317,1291,1317]" captionStart="FIGURE 3" captionStartId="6.[151,250,1923,1947]" captionTargetBox="[156,1434,1395,1893]" captionTargetId="figure-43@6.[151,1435,1389,1899]" captionTargetPageId="6" captionText="FIGURE 3. Adult Colpoptera stonei sp. nov.; (A) Head, pronotum, mesonotum dorsal view; (B) head and pronotum lateral view;(C) head frontal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12758770" httpUri="https://zenodo.org/record/12758770/files/figure.png" pageId="5" pageNumber="346">Fig. 3A</figureCitation>
). In lateral view, greatly raised, angled where diagonal carinae arise at mid carina (
<figureCitation id="1309A397FF89FFEAFC6E8421FBF60944" box="[938,1025,1327,1353]" captionStart="FIGURE 3" captionStartId="6.[151,250,1923,1947]" captionTargetBox="[156,1434,1395,1893]" captionTargetId="figure-43@6.[151,1435,1389,1899]" captionTargetPageId="6" captionText="FIGURE 3. Adult Colpoptera stonei sp. nov.; (A) Head, pronotum, mesonotum dorsal view; (B) head and pronotum lateral view;(C) head frontal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12758770" httpUri="https://zenodo.org/record/12758770/files/figure.png" pageId="5" pageNumber="346">Fig. 3B</figureCitation>
). Hind tibiae with single lateral spine approximately 2/3 distance from femoral-tibial joint, spinulation 6-7-2.
</paragraph>
<paragraph id="8B8DBF12FF89FFEAFF038479FF0509B8" blockId="5.[151,1437,931,2001]" pageId="5" pageNumber="346">
Forewing reticulate, broadest basally, constricting at claval apex, expanding slightly, apex broadly rounded (
<figureCitation id="1309A397FF89FFEAFF5B8495FF1209B8" box="[159,229,1435,1461]" captionStart="FIGURE 4" captionStartId="7.[151,250,751,775]" captionTargetBox="[151,1435,181,727]" captionTargetId="figure-17@7.[151,1435,181,727]" captionTargetPageId="7" captionText="FIGURE 4. Forewing venation of Colpoptera stonei sp. nov. Vein labeling nomenclature following Bourgoin et al. (2015)." figureDoi="http://doi.org/10.5281/zenodo.12758774" httpUri="https://zenodo.org/record/12758774/files/figure.png" pageId="5" pageNumber="346">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8B8DBF12FF89FFEAFF0384B1FB940B68" blockId="5.[151,1437,931,2001]" pageId="5" pageNumber="346">
<emphasis id="B9466300FF89FFEAFF0384B1FE7709D4" box="[199,384,1471,1497]" italics="true" pageId="5" pageNumber="346">Male terminalia</emphasis>
. Pyfoger in lateral view nearly uniform width along entire length, anterior margin sinuate, slightly concave, posterior margin nearly linear, dorsal and ventral margin equal length, slightly sinuate, medioventral process lacking (
<figureCitation id="1309A397FF89FFEAFE908709FE6E0A2F" box="[340,409,1543,1570]" captionStart="FIGURE 5" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[197,1376,799,1430]" captionTargetId="figure-38@7.[197,1390,799,1430]" captionTargetPageId="7" captionText="FIGURE 5. Adult male Colpoptera stonei sp. nov., terminalia in left lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758776" httpUri="https://zenodo.org/record/12758776/files/figure.png" pageId="5" pageNumber="346">Fig. 5</figureCitation>
). Gonostyli in lateral view generally rounded with large dorsal process arising basally, with helical sclerotized ridges, apex blunt with posterior corner rounded, anterior corner hooked (
<figureCitation id="1309A397FF89FFEAFB0A8725FAE10A4B" box="[1230,1302,1579,1606]" captionStart="FIGURE 5" captionStartId="7.[151,250,1454,1478]" captionTargetBox="[197,1376,799,1430]" captionTargetId="figure-38@7.[197,1390,799,1430]" captionTargetPageId="7" captionText="FIGURE 5. Adult male Colpoptera stonei sp. nov., terminalia in left lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758776" httpUri="https://zenodo.org/record/12758776/files/figure.png" pageId="5" pageNumber="346">Fig. 5</figureCitation>
). Aedeagus nearly symmetrical with three pairs of processes (
<figureCitation id="1309A397FF89FFEAFD0D8741FCF90A64" box="[713,782,1615,1641]" captionStart="FIGURE 6" captionStartId="8.[151,250,954,978]" captionTargetBox="[158,1423,181,930]" captionTargetId="figure-24@8.[158,1429,181,930]" captionTargetPageId="8" captionText="FIGURE 6. Aedeagus of Colpoptera stonei sp. nov.: (A) right lateral view; (B) left lateral view; (C) dorsal view; (D) ventral view." figureDoi="http://doi.org/10.5281/zenodo.12758780" httpUri="https://zenodo.org/record/12758780/files/figure.png" pageId="5" pageNumber="346">Fig. 6</figureCitation>
); first pair (A1 &amp; A2) arising on subapical lateral margin, angled dorsad and cephalad, both processes straight, slightly more sclerotized than rest of aedeagus (
<figureCitation id="1309A397FF89FFEAFAD6877DFA790A80" box="[1298,1422,1651,1677]" captionStart="FIGURE 6" captionStartId="8.[151,250,954,978]" captionTargetBox="[158,1423,181,930]" captionTargetId="figure-24@8.[158,1429,181,930]" captionTargetPageId="8" captionText="FIGURE 6. Aedeagus of Colpoptera stonei sp. nov.: (A) right lateral view; (B) left lateral view; (C) dorsal view; (D) ventral view." figureDoi="http://doi.org/10.5281/zenodo.12758780" httpUri="https://zenodo.org/record/12758780/files/figure.png" pageId="5" pageNumber="346">Fig. 6A, B</figureCitation>
), second pair (A3 &amp; A4) arising on inner ventral margin just basad of A1 and A2, angled dorsad and cephalad, slightly curved mesad, apices barely crossing at midpoint, approximately twice the length of A1 and A2 (
<figureCitation id="1309A397FF89FFEAFB0787B5FAED0AD8" box="[1219,1306,1723,1749]" captionStart="FIGURE 6" captionStartId="8.[151,250,954,978]" captionTargetBox="[158,1423,181,930]" captionTargetId="figure-24@8.[158,1429,181,930]" captionTargetPageId="8" captionText="FIGURE 6. Aedeagus of Colpoptera stonei sp. nov.: (A) right lateral view; (B) left lateral view; (C) dorsal view; (D) ventral view." figureDoi="http://doi.org/10.5281/zenodo.12758780" httpUri="https://zenodo.org/record/12758780/files/figure.png" pageId="5" pageNumber="346">Fig. 6C</figureCitation>
). Third pair (A5 &amp; A6) arising on ventral margin midline, short, slightly curved distad from base, equally sclerotized relative to A1A4, approximately half length of A1 and A2 (
<figureCitation id="1309A397FF89FFEAFD3E860DFC8C0B10" box="[762,891,1795,1821]" captionStart="FIGURE 6" captionStartId="8.[151,250,954,978]" captionTargetBox="[158,1423,181,930]" captionTargetId="figure-24@8.[158,1429,181,930]" captionTargetPageId="8" captionText="FIGURE 6. Aedeagus of Colpoptera stonei sp. nov.: (A) right lateral view; (B) left lateral view; (C) dorsal view; (D) ventral view." figureDoi="http://doi.org/10.5281/zenodo.12758780" httpUri="https://zenodo.org/record/12758780/files/figure.png" pageId="5" pageNumber="346">Fig. 6C, D</figureCitation>
). Anal segment elongate, irregularly sinuate on dorsal and ventral margins, dorsal margin angled ventrad approximately ¼ from base, ventral margin with strong invagination approximately 2/3 from base, apex distad of invagination comma-shaped.
</paragraph>
</subSubSection>
<subSubSection id="C328EC99FF89FFEAFF038660FB840B84" box="[199,1139,1902,1929]" pageId="5" pageNumber="346" type="distribution">
<paragraph id="8B8DBF12FF89FFEAFF038660FB840B84" blockId="5.[151,1437,931,2001]" box="[199,1139,1902,1929]" pageId="5" pageNumber="346">
<emphasis id="B9466300FF89FFEAFF038660FE960B85" bold="true" box="[199,353,1902,1928]" pageId="5" pageNumber="346">Distribution.</emphasis>
Manzanillo,
<collectingRegion id="49F671F0FF89FFEAFE318661FD5A0B84" box="[501,685,1903,1929]" country="Costa Rica" name="Limon" pageId="5" pageNumber="346">Limón Province</collectingRegion>
,
<collectingCountry id="F325FF82FF89FFEAFD7C8661FCC10B84" box="[696,822,1903,1929]" name="Costa Rica" pageId="5" pageNumber="346">Costa Rica</collectingCountry>
; Almonds and Corals Hotel.
</paragraph>
</subSubSection>
<subSubSection id="C328EC99FF89FFE8FF03869DFE450B5F" lastPageId="7" lastPageNumber="348" pageId="5" pageNumber="346" type="materials_examined">
<paragraph id="8B8DBF12FF89FFEAFF03869DFCD10BDC" blockId="5.[151,1437,931,2001]" pageId="5" pageNumber="346">
<emphasis id="B9466300FF89FFEAFF03869DFE820BA1" bold="true" box="[199,373,1938,1965]" pageId="5" pageNumber="346">Type material.</emphasis>
<materialsCitation id="3B5AB54FFF89FFEAFEBF869DFCD10BDC" collectingDate="2018-06-16" collectionCode="FLREC" collectorName="Coll. &amp; B. W. Bahder" country="Costa Rica" location="Manzanillo" pageId="5" pageNumber="346" specimenCount="1" specimenCount-male="1" stateProvince="Limon" typeStatus="holotype">
<typeStatus id="548901B0FF89FFEAFEBF869DFE110BA0" box="[379,486,1939,1965]" pageId="5" pageNumber="346" type="holotype">Holotype</typeStatus>
, male “
<collectingCountry id="F325FF82FF89FFEAFDFC869DFD410BA0" box="[568,694,1939,1965]" name="Costa Rica" pageId="5" pageNumber="346">Costa Rica</collectingCountry>
,
<collectingRegion id="49F671F0FF89FFEAFD05869DFCFB0BA0" box="[705,780,1939,1965]" country="Costa Rica" name="Limon" pageId="5" pageNumber="346">Limón</collectingRegion>
Pr. / nr.
<location id="8EEDE9C9FF89FFEAFCAD869DFC1E0BA0" LSID="urn:lsid:plazi:treatment:039B0E04FF88FFE6FF538490FB690EA4:8EEDE9C9FF89FFEAFCAD869DFC1E0BA0" box="[873,1001,1939,1965]" country="Costa Rica" name="Manzanillo" pageId="5" pageNumber="346" stateProvince="Limon">Manzanillo</location>
/
<date id="FF8C99D2FF89FFEAFC3A869DFB8A0BA0" box="[1022,1149,1939,1965]" pageId="5" pageNumber="346" value="2018-06-16">
<collectingDate id="EFC8603AFF89FFEAFC3A869DFB8A0BA0" box="[1022,1149,1939,1965]" pageId="5" pageNumber="346" value="2018-06-16">16.VI.2018</collectingDate>
</date>
/ sweeping weeds /
<collectorName id="26C7DAC4FF89FFEAFA99869DFA6E0BA0" box="[1373,1433,1939,1965]" pageId="5" pageNumber="346">Coll.</collectorName>
:
<collectorName id="26C7DAC4FF89FFEAFF5386B9FED70BDC" box="[151,288,1975,2001]" pageId="5" pageNumber="346">B.W.Bahder</collectorName>
//
<typeStatus id="548901B0FF89FFEAFEFA86B9FE510BDC" box="[318,422,1975,2001]" pageId="5" pageNumber="346" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="4C32C491FF89FFEAFE7886B9FD710BDC" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[444,646,1975,2001]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="stonei">
<emphasis id="B9466300FF89FFEAFE7886B9FD710BDC" box="[444,646,1975,2001]" italics="true" pageId="5" pageNumber="346">Colpoptera stonei</emphasis>
</taxonomicName>
<specimenCount id="9D34749BFF89FFEAFD4986B9FD540BDC" box="[653,675,1975,2001]" count="1" pageId="5" pageNumber="346" type="male"></specimenCount>
” (
<collectionCode id="ED2327D7FF89FFEAFD7986B9FCEC0BDC" box="[701,795,1975,2001]" pageId="5" pageNumber="346">FLREC</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="DF4DEF9AFF8AFFE9FF538433FB890958" ID-DOI="http://doi.org/10.5281/zenodo.12758768" ID-Zenodo-Dep="12758768" box="[151,1150,1341,1365]" httpUri="https://zenodo.org/record/12758768/files/figure.png" pageId="6" pageNumber="347" startId="6.[151,250,1341,1365]" targetBox="[151,1436,181,1317]" targetPageId="6" targetType="figure">
<paragraph id="8B8DBF12FF8AFFE9FF538433FB890958" blockId="6.[151,1150,1341,1365]" box="[151,1150,1341,1365]" pageId="6" pageNumber="347">
<emphasis id="B9466300FF8AFFE9FF538433FEE40958" bold="true" box="[151,275,1341,1365]" pageId="6" pageNumber="347">FIGURE 2.</emphasis>
Adult habitus
<taxonomicName id="4C32C491FF8AFFE9FE6C8433FDA90958" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[424,606,1341,1365]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
<emphasis id="B9466300FF8AFFE9FE6C8433FDA90958" box="[424,606,1341,1365]" italics="true" pageId="6" pageNumber="347">Colpoptera stonei</emphasis>
</taxonomicName>
<emphasis id="B9466300FF8AFFE9FDA08433FD420958" bold="true" box="[612,693,1341,1365]" pageId="6" pageNumber="347">
<taxonomicNameLabel id="A275DE7BFF8AFFE9FDA08433FD420958" box="[612,693,1341,1365]" pageId="6" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) Male lateral view; (B) male dorsal view.
</paragraph>
</caption>
<caption id="DF4DEF9AFF8AFFE9FF53868DFDB00BB3" ID-DOI="http://doi.org/10.5281/zenodo.12758770" ID-Zenodo-Dep="12758770" httpUri="https://zenodo.org/record/12758770/files/figure.png" pageId="6" pageNumber="347" startId="6.[151,250,1923,1947]" targetBox="[156,1434,1395,1893]" targetPageId="6" targetType="figure">
<paragraph id="8B8DBF12FF8AFFE9FF53868DFDB00BB3" blockId="6.[151,1436,1923,1983]" pageId="6" pageNumber="347">
<emphasis id="B9466300FF8AFFE9FF53868DFEE20B96" bold="true" box="[151,277,1923,1947]" pageId="6" pageNumber="347">FIGURE 3.</emphasis>
Adult
<taxonomicName id="4C32C491FF8AFFE9FE99868DFDE10B96" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[349,534,1923,1947]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
<emphasis id="B9466300FF8AFFE9FE99868DFDE10B96" box="[349,534,1923,1947]" italics="true" pageId="6" pageNumber="347">Colpoptera stonei</emphasis>
</taxonomicName>
<emphasis id="B9466300FF8AFFE9FDD9868DFD870B96" bold="true" box="[541,624,1923,1947]" pageId="6" pageNumber="347">
<taxonomicNameLabel id="A275DE7BFF8AFFE9FDD9868DFD870B96" box="[541,624,1923,1947]" pageId="6" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; (A) Head, pronotum, mesonotum dorsal view; (B) head and pronotum lateral view;(C) head frontal view. Scale = 1 mm.
</paragraph>
</caption>
<caption id="DF4DEF9AFF8BFFE8FF5383E1FA8A0F0A" ID-DOI="http://doi.org/10.5281/zenodo.12758774" ID-Zenodo-Dep="12758774" box="[151,1405,751,775]" httpUri="https://zenodo.org/record/12758774/files/figure.png" pageId="7" pageNumber="348" startId="7.[151,250,751,775]" targetBox="[151,1435,181,727]" targetPageId="7" targetType="figure">
<paragraph id="8B8DBF12FF8BFFE8FF5383E1FA8A0F0A" blockId="7.[151,1405,751,775]" box="[151,1405,751,775]" pageId="7" pageNumber="348">
<emphasis id="B9466300FF8BFFE8FF5383E1FEE40F0A" bold="true" box="[151,275,751,775]" pageId="7" pageNumber="348">FIGURE 4.</emphasis>
Forewing venation of
<taxonomicName id="4C32C491FF8BFFE8FE3E83E1FD460F0A" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[506,689,751,775]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
<emphasis id="B9466300FF8BFFE8FE3E83E1FD460F0A" box="[506,689,751,775]" italics="true" pageId="7" pageNumber="348">Colpoptera stonei</emphasis>
</taxonomicName>
<emphasis id="B9466300FF8BFFE8FD7283E1FCF00F0A" bold="true" box="[694,775,751,775]" pageId="7" pageNumber="348">
<taxonomicNameLabel id="A275DE7BFF8BFFE8FD7283E1FCF00F0A" box="[694,775,751,775]" pageId="7" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Vein labeling nomenclature following
<bibRefCitation id="EFA3C2E3FF8BFFE8FB5083E1FA8F0F0A" author="Bourgoin, T. &amp; Wang, R. R. &amp; Ache, M. &amp; Hoch, H. &amp; Soulier-Perkins, A. &amp; Stroinski, A. &amp; Yap, S. &amp; Szwedo, J." box="[1172,1400,751,775]" pageId="7" pageNumber="348" pagination="63 - 77" refId="ref6059" refString="Bourgoin, T., Wang, R. R., Ache, M., Hoch, H., Soulier-Perkins, A., Stroinski, A., Yap, S. &amp; Szwedo, J. (2015) From micropterism to hyperpterism: recognition strategy and standardized homology-driven terminology of the forewing venation patterns in planthoppers (Hemiptera: Fulgoromorpha). Zoomorphology, 134 (1), 63 - 77. https: // doi. org / 10.1007 / s 00435 - 014 - 0243 - 6" type="journal article" year="2015">
Bourgoin
<emphasis id="B9466300FF8BFFE8FB3D83FEFAD10F0A" box="[1273,1318,751,775]" italics="true" pageId="7" pageNumber="348">et al</emphasis>
. (2015)
</bibRefCitation>
.
</paragraph>
</caption>
<caption id="DF4DEF9AFF8BFFE8FF5384A0FC2009CB" ID-DOI="http://doi.org/10.5281/zenodo.12758776" ID-Zenodo-Dep="12758776" box="[151,983,1454,1478]" httpUri="https://zenodo.org/record/12758776/files/figure.png" pageId="7" pageNumber="348" startId="7.[151,250,1454,1478]" targetBox="[197,1376,799,1430]" targetPageId="7" targetType="figure">
<paragraph id="8B8DBF12FF8BFFE8FF5384A0FC2009CB" blockId="7.[151,983,1454,1478]" box="[151,983,1454,1478]" pageId="7" pageNumber="348">
<emphasis id="B9466300FF8BFFE8FF5384A0FEE409CB" bold="true" box="[151,275,1454,1478]" pageId="7" pageNumber="348">FIGURE 5.</emphasis>
Adult male
<taxonomicName id="4C32C491FF8BFFE8FE5584A0FDBF09CB" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[401,584,1454,1478]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
<emphasis id="B9466300FF8BFFE8FE5584A0FDBF09CB" box="[401,584,1454,1478]" italics="true" pageId="7" pageNumber="348">Colpoptera stonei</emphasis>
</taxonomicName>
<emphasis id="B9466300FF8BFFE8FD8A84A0FD5209CB" bold="true" box="[590,677,1454,1478]" pageId="7" pageNumber="348">
<taxonomicNameLabel id="A275DE7BFF8BFFE8FD8A84A0FD5709CB" box="[590,672,1454,1478]" pageId="7" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
terminalia in left lateral view.
</paragraph>
</caption>
<paragraph id="8B8DBF12FF8BFFE8FF038717FE180A5B" blockId="7.[151,1437,1560,1874]" pageId="7" pageNumber="348">
<typeStatus id="548901B0FF8BFFE8FF038717FECF0A3E" box="[199,312,1561,1587]" pageId="7" pageNumber="348" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D34749BFF8BFFE8FE818716FE650A3F" box="[325,402,1560,1586]" count="2" pageId="7" pageNumber="348" type="male">2 male</specimenCount>
and
<specimenCount id="9D34749BFF8BFFE8FE088716FDCD0A3F" box="[460,570,1560,1586]" count="4" pageId="7" pageNumber="348" type="female">4 females</specimenCount>
same as
<materialsCitation id="3B5AB54FFF8BFFE8FD618716FE1C0A5B" collectingDate="2018-06-20" location="La Sleva Biological Station" pageId="7" pageNumber="348" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Heredia" typeStatus="holotype">
<typeStatus id="548901B0FF8BFFE8FD618716FCE70A3F" box="[677,784,1560,1586]" pageId="7" pageNumber="348" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D34749BFF8BFFE8FCD98716FC9D0A3F" box="[797,874,1560,1586]" count="1" pageId="7" pageNumber="348" type="male">1 male</specimenCount>
and
<specimenCount id="9D34749BFF8BFFE8FC618716FBE40A3F" box="[933,1043,1560,1586]" count="2" pageId="7" pageNumber="348" type="female">2 females</specimenCount>
from
<location id="8EEDE9C9FF8BFFE8FB9F8717FA6F0A3F" LSID="urn:lsid:plazi:treatment:039B0E04FF88FFE6FF538490FB690EA4:8EEDE9C9FF8BFFE8FB9F8717FA6F0A3F" box="[1115,1432,1560,1587]" name="La Sleva Biological Station" pageId="7" pageNumber="348" stateProvince="Heredia">La Sleva Biological Station</location>
,
<collectingRegion id="49F671F0FF8BFFE8FF538732FEAA0A5B" box="[151,349,1596,1622]" country="Costa Rica" name="Heredia" pageId="7" pageNumber="348">Heredia Province</collectingRegion>
,
<date id="FF8C99D2FF8BFFE8FEAC8732FE1C0A5B" box="[360,491,1596,1622]" pageId="7" pageNumber="348" value="2018-06-20">
<collectingDate id="EFC8603AFF8BFFE8FEAC8732FE1C0A5B" box="[360,491,1596,1622]" pageId="7" pageNumber="348" value="2018-06-20">20.VI.2018</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8DBF12FF8BFFE8FF03876EFC220A77" blockId="7.[151,1437,1560,1874]" box="[199,981,1632,1658]" pageId="7" pageNumber="348">
<emphasis id="B9466300FF8BFFE8FF03876EFE540A77" bold="true" box="[199,419,1632,1658]" pageId="7" pageNumber="348">Plant associations.</emphasis>
Unknown, collected sweeping herbaceous weeds.
</paragraph>
<paragraph id="8B8DBF12FF8BFFE8FF03878AFE450B5F" blockId="7.[151,1437,1560,1874]" pageId="7" pageNumber="348">
<emphasis id="B9466300FF8BFFE8FF03878AFE750A93" bold="true" box="[199,386,1668,1694]" pageId="7" pageNumber="348">Sequence Data.</emphasis>
For the COI gene, a 657 bp product was generated (GenBank Accession No. PP378885), for the 18S gene, a 1,265 bp product (GenBank Accession No. PP386391) was generated and for the 28S D9 D10 expansion region, a 829 bp product was generated (GenBank Accession No. PP386392). The phylogenies constructed demonstrate strong bootstrap support for the monophyly of
<taxonomicName id="4C32C491FF8BFFE8FC0E87FEFBBD0B07" authorityName="Burmeister" authorityYear="1835" box="[970,1098,1776,1802]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="B9466300FF8BFFE8FC0E87FEFBBD0B07" box="[970,1098,1776,1802]" italics="true" pageId="7" pageNumber="348">Colpoptera</emphasis>
</taxonomicName>
based on 18S, 28S, COI data and concatenated sequences of the three loci (100, 99 and 80, respectively) with
<taxonomicName id="4C32C491FF8BFFE8FBCD861AFB260B23" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[1033,1233,1812,1838]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
<emphasis id="B9466300FF8BFFE8FBCD861AFB260B23" box="[1033,1233,1812,1838]" italics="true" pageId="7" pageNumber="348">Colpoptera stonei</emphasis>
</taxonomicName>
<emphasis id="B9466300FF8BFFE8FB13861AFAD80B23" bold="true" box="[1239,1327,1812,1838]" pageId="7" pageNumber="348">
<taxonomicNameLabel id="A275DE7BFF8BFFE8FB13861AFAD80B23" box="[1239,1327,1812,1838]" pageId="7" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resolving within this clade (
<figureCitation id="1309A397FF8BFFE8FEA48636FE530B5F" box="[352,420,1848,1874]" captionStart="FIGURE 7" captionStartId="8.[151,250,1701,1725]" captionTargetBox="[157,1430,1042,1672]" captionTargetId="figure-50@8.[151,1436,1039,1678]" captionTargetPageId="8" captionText="FIGURE 7. Maximum Likelihood trees (1,000 replicates) exhibiting relationship of Colpoptera stonei sp. nov. to available nogodinid taxa: (A) 18S, (B) 28S, (C) COI and (D) consensus tree based on concatenated data for the three loci; scale bar = percent nucleotide difference." figureDoi="http://doi.org/10.5281/zenodo.12758782" httpUri="https://zenodo.org/record/12758782/files/figure.png" pageId="7" pageNumber="348">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF4DEF9AFF84FFE7FF5382B4FF390FFB" ID-DOI="http://doi.org/10.5281/zenodo.12758780" ID-Zenodo-Dep="12758780" httpUri="https://zenodo.org/record/12758780/files/figure.png" pageId="8" pageNumber="349" startId="8.[151,250,954,978]" targetBox="[158,1423,181,930]" targetPageId="8" targetType="figure">
<paragraph id="8B8DBF12FF84FFE7FF5382B4FF390FFB" blockId="8.[151,1436,954,1014]" pageId="8" pageNumber="349">
<emphasis id="B9466300FF84FFE7FF5382B4FEE30FDF" bold="true" box="[151,276,954,978]" pageId="8" pageNumber="349">FIGURE 6.</emphasis>
Aedeagus of
<taxonomicName id="4C32C491FF84FFE7FE6582B4FDAF0FDF" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[417,600,954,978]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="349" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
<emphasis id="B9466300FF84FFE7FE6582B4FDAF0FDF" box="[417,600,954,978]" italics="true" pageId="8" pageNumber="349">Colpoptera stonei</emphasis>
</taxonomicName>
<emphasis id="B9466300FF84FFE7FD9B82B5FD460FDE" bold="true" box="[607,689,955,979]" pageId="8" pageNumber="349">
<taxonomicNameLabel id="A275DE7BFF84FFE7FD9B82B5FD460FDE" box="[607,689,955,979]" pageId="8" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A) right lateral view; (B) left lateral view; (C) dorsal view; (D) ventral view.
</paragraph>
</caption>
<caption id="DF4DEF9AFF84FFE7FF5387ABFE310B08" ID-DOI="http://doi.org/10.5281/zenodo.12758782" ID-Zenodo-Dep="12758782" httpUri="https://zenodo.org/record/12758782/files/figure.png" pageId="8" pageNumber="349" startId="8.[151,250,1701,1725]" targetBox="[157,1430,1042,1672]" targetPageId="8" targetType="figure">
<paragraph id="8B8DBF12FF84FFE7FF5387ABFE310B08" blockId="8.[151,1436,1701,1797]" pageId="8" pageNumber="349">
<emphasis id="B9466300FF84FFE7FF5387ABFEE20AB0" bold="true" box="[151,277,1701,1725]" pageId="8" pageNumber="349">FIGURE 7.</emphasis>
Maximum Likelihood trees (1,000 replicates) exhibiting relationship of
<taxonomicName id="4C32C491FF84FFE7FBC387ABFB370AB0" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[1031,1216,1701,1725]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="349" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
<emphasis id="B9466300FF84FFE7FBC387ABFB370AB0" box="[1031,1216,1701,1725]" italics="true" pageId="8" pageNumber="349">Colpoptera stonei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A275DE7BFF84FFE7FB0C87ABFAEB0AB0" box="[1224,1308,1701,1725]" pageId="8" pageNumber="349" rank="species">
<emphasis id="B9466300FF84FFE7FB0C87ABFAE10AB0" bold="true" box="[1224,1302,1701,1725]" pageId="8" pageNumber="349">sp. nov</emphasis>
.
</taxonomicNameLabel>
to available nogodinid taxa: (A) 18S, (B) 28S, (C) COI and (D) consensus tree based on concatenated data for the three loci; scale bar = percent nucleotide difference.
</paragraph>
</caption>
<subSubSection id="C328EC99FF84FFE6FF038656FB690EA4" lastPageId="9" lastPageNumber="350" pageId="8" pageNumber="349" type="discussion">
<paragraph id="8B8DBF12FF84FFE6FF038656FD7A0E30" blockId="8.[151,1436,1879,2014]" lastBlockId="9.[151,1437,151,681]" lastPageId="9" lastPageNumber="350" pageId="8" pageNumber="349">
<emphasis id="B9466300FF84FFE7FF038656FECC0B7F" bold="true" box="[199,315,1880,1906]" pageId="8" pageNumber="349">Remarks.</emphasis>
The general habitus of
<taxonomicName id="4C32C491FF84FFE7FD918659FCD50B7C" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[597,802,1879,1905]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="349" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
<emphasis id="B9466300FF84FFE7FD918659FCD50B7C" box="[597,802,1879,1905]" italics="true" pageId="8" pageNumber="349">Colpoptera stonei</emphasis>
</taxonomicName>
<emphasis id="B9466300FF84FFE7FCE88656FC7E0B7F" bold="true" box="[812,905,1880,1906]" pageId="8" pageNumber="349">
<taxonomicNameLabel id="A275DE7BFF84FFE7FCE88656FC7E0B7F" box="[812,905,1880,1906]" pageId="8" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
appears similar to many described species of
<taxonomicName id="4C32C491FF84FFE7FF538675FEE00B98" authorityName="Burmeister" authorityYear="1835" box="[151,279,1915,1941]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="349" phylum="Arthropoda" rank="genus">
<emphasis id="B9466300FF84FFE7FF538675FEE00B98" box="[151,279,1915,1941]" italics="true" pageId="8" pageNumber="349">Colpoptera</emphasis>
</taxonomicName>
and both the form of the terminalia and geography appear to exclude the other described genera in the
<taxonomicName id="4C32C491FF84FFE7FF078691FEA60BB4" authorityName="Gnezdilov" authorityYear="2003" box="[195,337,1951,1977]" class="Insecta" family="Nogodinidae" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="tribe" tribe="Colpopterini">Colpopterini</taxonomicName>
(most genera are Antillean). The closest species in terms of coloration and general form of the aedeagus appears to be
<taxonomicName id="4C32C491FF84FFE7FE5B86CAFD5D0BD0" authority="Caldwell" authorityName="Caldwell" authorityYear="1945" box="[415,682,1987,2014]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="349" phylum="Arthropoda" rank="species" species="albavenosa">
<emphasis id="B9466300FF84FFE7FE5B86CAFDC80BD0" box="[415,575,1987,2014]" italics="true" pageId="8" pageNumber="349">C. albavenosa</emphasis>
Caldwell
</taxonomicName>
from
<collectingRegion id="49F671F0FF84FFE7FD2A86CAFC690BD3" box="[750,926,1987,2014]" country="Mexico" name="San Luis Potosi" pageId="8" pageNumber="349">San Luis Potosí</collectingRegion>
in northern
<collectingCountry id="F325FF82FF84FFE7FBE386CDFB8A0BD0" box="[1063,1149,1987,2013]" name="Mexico" pageId="8" pageNumber="349">Mexico</collectingCountry>
(
<bibRefCitation id="EFA3C2E3FF84FFE7FB4F86CDFAC40BD3" author="Caldwell, J. S." box="[1163,1331,1987,2014]" pageId="8" pageNumber="349" pagination="89 - 120" refId="ref6271" refString="Caldwell, J. S. (1945) Notes on Issidae from Mexico (Homoptera, Fulgoroidea). Annals of the Entomological Society of America, 38, 89 - 120. https: // doi. org / 10.1093 / aesa / 38.1.89" type="journal article" year="1945">Caldwell 1945</bibRefCitation>
, plate IV, fig. 2). However,
<taxonomicName id="4C32C491FF85FFE6FE9F8199FDD30CBD" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[347,548,151,177]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
<emphasis id="B9466300FF85FFE6FE9F8199FDD30CBD" box="[347,548,151,177]" italics="true" pageId="9" pageNumber="350">Colpoptera stonei</emphasis>
</taxonomicName>
<emphasis id="B9466300FF85FFE6FDEE8199FD740CBC" bold="true" box="[554,643,151,177]" pageId="9" pageNumber="350">
<taxonomicNameLabel id="A275DE7BFF85FFE6FDEE8199FD740CBC" box="[554,643,151,177]" pageId="9" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C32C491FF85FFE6FCD18199FC420CBC" authorityName="Caldwell" authorityYear="1945" box="[789,949,151,177]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="albavenosa">
<emphasis id="B9466300FF85FFE6FCD18199FC420CBC" box="[789,949,151,177]" italics="true" pageId="9" pageNumber="350">C. albavenosa</emphasis>
</taxonomicName>
in the anal segment where
<taxonomicName id="4C32C491FF85FFE6FB208199FA730CBC" authorityName="Caldwell" authorityYear="1945" box="[1252,1412,151,177]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="albavenosa">
<emphasis id="B9466300FF85FFE6FB208199FA730CBC" box="[1252,1412,151,177]" italics="true" pageId="9" pageNumber="350">C. albavenosa</emphasis>
</taxonomicName>
is noted as having a nearly straight segment whereas
<taxonomicName id="4C32C491FF85FFE6FD0A81B5FC600CD9" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[718,919,187,213]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
<emphasis id="B9466300FF85FFE6FD0A81B5FC600CD9" box="[718,919,187,213]" italics="true" pageId="9" pageNumber="350">Colpoptera stonei</emphasis>
</taxonomicName>
<emphasis id="B9466300FF85FFE6FC5A81B5FC000CD8" bold="true" box="[926,1015,187,213]" pageId="9" pageNumber="350">
<taxonomicNameLabel id="A275DE7BFF85FFE6FC5A81B5FC000CD8" box="[926,1015,187,213]" pageId="9" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has deep invaginations on the ventral margin.Also, the processes on the aedeagus of
<taxonomicName id="4C32C491FF85FFE6FD6581D1FCB60CF4" authorityName="Caldwell" authorityYear="1945" box="[673,833,223,249]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="albavenosa">
<emphasis id="B9466300FF85FFE6FD6581D1FCB60CF4" box="[673,833,223,249]" italics="true" pageId="9" pageNumber="350">C. albavenosa</emphasis>
</taxonomicName>
are noted as subequal in length whereas the difference in size is substantial in
<taxonomicName id="4C32C491FF85FFE6FE58800DFD920D11" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[412,613,259,285]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
<emphasis id="B9466300FF85FFE6FE58800DFD920D11" box="[412,613,259,285]" italics="true" pageId="9" pageNumber="350">Colpoptera stonei</emphasis>
</taxonomicName>
<emphasis id="B9466300FF85FFE6FDA8800DFD310D10" bold="true" box="[620,710,259,285]" pageId="9" pageNumber="350">
<taxonomicNameLabel id="A275DE7BFF85FFE6FDA8800DFD310D10" box="[620,710,259,285]" pageId="9" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(also
<bibRefCitation id="EFA3C2E3FF85FFE6FCCE800DFC4F0D10" author="Caldwell, J. S." box="[778,952,258,285]" pageId="9" pageNumber="350" pagination="89 - 120" refId="ref6271" refString="Caldwell, J. S. (1945) Notes on Issidae from Mexico (Homoptera, Fulgoroidea). Annals of the Entomological Society of America, 38, 89 - 120. https: // doi. org / 10.1093 / aesa / 38.1.89" type="journal article" year="1945">Caldwell, 1945</bibRefCitation>
, depicts three processes on each side of the aedeagus for
<taxonomicName id="4C32C491FF85FFE6FEE88029FE5B0D4C" authorityName="Caldwell" authorityYear="1945" box="[300,428,295,321]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="albavenosa">
<emphasis id="B9466300FF85FFE6FEE88029FE5B0D4C" box="[300,428,295,321]" italics="true" pageId="9" pageNumber="350">albavenosa</emphasis>
</taxonomicName>
rather than two per side for
<taxonomicName id="4C32C491FF85FFE6FD2C8029FCBB0D4D" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[744,844,295,321]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
<emphasis id="B9466300FF85FFE6FD2C8029FCBB0D4D" box="[744,844,295,321]" italics="true" pageId="9" pageNumber="350">C. stonei</emphasis>
</taxonomicName>
<emphasis id="B9466300FF85FFE6FC978029FC5B0D4C" bold="true" box="[851,940,295,321]" pageId="9" pageNumber="350">
<taxonomicNameLabel id="A275DE7BFF85FFE6FC978029FC5B0D4C" box="[851,940,295,321]" pageId="9" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). Finally, the gonostyli in
<taxonomicName id="4C32C491FF85FFE6FB178029FA830D4C" authorityName="Caldwell" authorityYear="1945" box="[1235,1396,295,321]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="albavenosa">
<emphasis id="B9466300FF85FFE6FB178029FA830D4C" box="[1235,1396,295,321]" italics="true" pageId="9" pageNumber="350">C. albavenosa</emphasis>
</taxonomicName>
are shown as elongated, hooked, and narrow where they are stout with helical ridges
<taxonomicName id="4C32C491FF85FFE6FBE48045FB740D69" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[1056,1155,331,357]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
<emphasis id="B9466300FF85FFE6FBE48045FB740D69" box="[1056,1155,331,357]" italics="true" pageId="9" pageNumber="350">C. stonei</emphasis>
</taxonomicName>
<emphasis id="B9466300FF85FFE6FB4D8045FB140D68" bold="true" box="[1161,1251,331,357]" pageId="9" pageNumber="350">
<taxonomicNameLabel id="A275DE7BFF85FFE6FB4D8045FB140D68" box="[1161,1251,331,357]" pageId="9" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Another species with similar form in the terminalia is
<taxonomicName id="4C32C491FF85FFE6FDF48061FCE10D84" authority="Caldwell." authorityName="Caldwell." box="[560,790,367,393]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="acutata">
<emphasis id="B9466300FF85FFE6FDF48061FD540D85" box="[560,675,367,393]" italics="true" pageId="9" pageNumber="350">C. acutata</emphasis>
Caldwell.
</taxonomicName>
The general armature of the aedeagus in general is similar, however, in
<taxonomicName id="4C32C491FF85FFE6FEE6809DFE720DA1" authorityName="Barrantes Barrantes &amp; Zumbado Echavarria &amp; Van Dam &amp; Helmick &amp; Bartlett &amp; Martinez Aponte &amp; Ruiz &amp; Bloch &amp; Bahder" authorityYear="2024" box="[290,389,403,429]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="stonei" status="sp. nov.">
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the ventral bifurcation of the processes arises mesad and in
<taxonomicName id="4C32C491FF85FFE6FB4F809DFB080DA1" box="[1163,1279,403,429]" class="Insecta" family="Nogodinidae" genus="Colpoptera" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="350" phylum="Arthropoda" rank="species" species="acutata">
<emphasis id="B9466300FF85FFE6FB4F809DFB080DA1" box="[1163,1279,403,429]" italics="true" pageId="9" pageNumber="350">C. acutata</emphasis>
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(based on the description) appear to arise on the lateral margins. Additionally,
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<emphasis id="B9466300FF85FFE6FCA680B9FC210DDD" box="[866,982,439,465]" italics="true" pageId="9" pageNumber="350">C. acutata</emphasis>
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is described as a red-brown species with white spots on the frons whereas
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<emphasis id="B9466300FF85FFE6FDB980D5FD200DF8" bold="true" box="[637,727,475,501]" pageId="9" pageNumber="350">
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is not red-brown and has an unmarked frons. Finally, the apex of the anal segment in
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<emphasis id="B9466300FF85FFE6FE5B80F1FDE10E15" box="[415,534,511,537]" italics="true" pageId="9" pageNumber="350">C. acutata</emphasis>
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appears narrowed and slightly hooked based on the description and it is much broaded and not hooked in
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<emphasis id="B9466300FF85FFE6FE0D832DFDDB0E31" box="[457,556,547,573]" italics="true" pageId="9" pageNumber="350">C. stonei</emphasis>
</taxonomicName>
<emphasis id="B9466300FF85FFE6FDF7832DFD7A0E30" bold="true" box="[563,653,547,573]" pageId="9" pageNumber="350">
<taxonomicNameLabel id="A275DE7BFF85FFE6FDF7832DFD7A0E30" box="[563,653,547,573]" pageId="9" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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While molecular data is useful for elucidating
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<emphasis id="B9466300FF85FFE6FD208349FC930E6C" box="[740,868,583,609]" italics="true" pageId="9" pageNumber="350">Colpoptera</emphasis>
</taxonomicName>
, very little data is available for other nogodinids taxa that would allow for generating a consensus tree among COI, 18S, and 28S. As new taxa are discovered and previously described taxa become available for analysis, this shortcoming can be addressed.
</paragraph>
</subSubSection>
</treatment>
</document>

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Genus
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Colpoptera
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species.
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Colpoptera sinuata
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(
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by subsequent designation of Distant 1910: 314)
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. (Abridged from
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). Body elongated, head about as broad as prothorax, eyes prominent. Frons elongate, with distinct median carina. Vertex transverse, anterior margin truncate, posterior margin concave. Pronotum narrow, anterior margin strongly convex, posterior margin concave. Mesonotum at midline longer than vertex + pronotum combined, bearing arcuate anterior transverse carinae and median carina. Forewings elongate and narrowed to rounded apex, with narrow hypocostal plate and many transverse veins apically, hindwings large and functional. Hind tibia with single lateral tooth distally. Male pygofer with nearly straight hind margin (elongate process absent). Aedeagus relatively simple, bilaterally symmetrical, with elongated, usually bifurcated, subapical lateral processes (“phallobase lobe” of
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); and with pair of shorter, simple. Gonostyli bearing broad capitulum, narrowing apically with broad lateral tooth.Anal tube elongated. Female sternite VII with rounded median process (
<bibRefCitation id="EFA3C2E3FF88FFEBFDB285E5FC590908" author="Gnezdilov, V. M. &amp; O'Brien, L. B." box="[630,942,1259,1285]" pageId="4" pageNumber="345" pagination="1 - 26" refId="ref6691" refString="Gnezdilov, V. M. &amp; O'Brien, L. B. (2008) New taxa and combinations in Neotropical Issidae (Hemiptera: Fulgoroidea). Insecta Mundi, 31, 1 - 26." type="journal article" year="2008">Gnezdilov &amp; OBrien 2008</bibRefCitation>
, fig. 59). Gonoplacs nearly triangular, fused medially; gonocoxa VIII with straight hind margin (
<bibRefCitation id="EFA3C2E3FF88FFEBFD218401FBD70924" author="Gnezdilov, V. M. &amp; O'Brien, L. B." box="[741,1056,1295,1321]" pageId="4" pageNumber="345" pagination="1 - 26" refId="ref6691" refString="Gnezdilov, V. M. &amp; O'Brien, L. B. (2008) New taxa and combinations in Neotropical Issidae (Hemiptera: Fulgoroidea). Insecta Mundi, 31, 1 - 26." type="journal article" year="2008">Gnezdilov &amp; OBrien 2008</bibRefCitation>
, figs. 6062). Anal tube long and narrow, distinctly longer than gonoplacs
<bibRefCitation id="EFA3C2E3FF88FFEBFDA4843DFC610940" author="Gnezdilov, V. M. &amp; O'Brien, L. B." box="[608,918,1331,1357]" pageId="4" pageNumber="345" pagination="1 - 26" refId="ref6691" refString="Gnezdilov, V. M. &amp; O'Brien, L. B. (2008) New taxa and combinations in Neotropical Issidae (Hemiptera: Fulgoroidea). Insecta Mundi, 31, 1 - 26." type="journal article" year="2008">Gnezdilov &amp; OBrien 2008</bibRefCitation>
, fig. 57).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,206 @@
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<mods:title id="C26210BD9EE5242060345130C58C36DA">Key to species of the leafhopper genus Fistulatus Zhang, Zhang &amp; Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species</mods:title>
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<taxonomicName id="4C7E4D05FFF1CF771ED9FE640F4B068E" authority="Hassan &amp; Xing" authorityName="Hassan &amp; Shah &amp; Xing" authorityYear="2024" box="[151,615,461,488]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov.">
<emphasis id="B90AEA94FFF1CF771ED9FE640C8E068E" bold="true" box="[151,418,461,488]" italics="true" pageId="2" pageNumber="365">Fistulatus motuoensis</emphasis>
Hassan &amp; Xing
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<paragraph id="8BC13686FFF1CF771ED9FE580C350562" blockId="2.[151,717,461,523]" box="[151,281,497,523]" pageId="2" pageNumber="365">
(
<figureCitation id="13452A03FFF1CF771ED1FE580C3C0562" box="[159,272,497,523]" captionStart-0="FIGURES 14" captionStart-1="FIGURES 513" captionStartId-0="2.[151,264,1905,1929]" captionStartId-1="3.[151,264,1846,1870]" captionTargetBox-0="[157,1429,805,1875]" captionTargetBox-1="[158,1429,188,1816]" captionTargetId-0="figure-522@2.[151,1435,799,1881]" captionTargetId-1="figure-17@3.[151,1435,181,1822]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 14. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male habitus: 1. Dorsal view; 2. Lateral view; 3. Face; 4. Head and thorax, dorsal view." captionText-1="FIGURES 513. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male. 5. Genitalia (after maceration), lateral view; 6. Male pygofer, lateral view; 7. Valve and subgenital plate, ventral view; 8. Style, lateral view; 9. Connective, dorsal view; 10.Aedeagus, dorsal view; 11. Aedeagus, ventral view; 12. Aedeagus, lateral view; 13. Aedeagus, anteroventral view focusing on the apex." figureDoi-0="http://doi.org/10.5281/zenodo.12758859" figureDoi-1="http://doi.org/10.5281/zenodo.12758861" httpUri-0="https://zenodo.org/record/12758859/files/figure.png" httpUri-1="https://zenodo.org/record/12758861/files/figure.png" pageId="2" pageNumber="365">Figs 113</figureCitation>
)
</paragraph>
<paragraph id="8BC13686FFF1CF771ED9FD9009F2046E" blockId="2.[151,1437,569,776]" pageId="2" pageNumber="365">
<emphasis id="B90AEA94FFF1CF771ED9FD900C05053A" bold="true" box="[151,297,569,595]" pageId="2" pageNumber="365">Description.</emphasis>
Body coloration light yellowish green (
<figureCitation id="13452A03FFF1CF771CADFD900E64053A" box="[739,840,569,595]" captionStart="FIGURES 14" captionStartId="2.[151,264,1905,1929]" captionTargetBox="[157,1429,805,1875]" captionTargetId="figure-522@2.[151,1435,799,1881]" captionTargetPageId="2" captionText="FIGURES 14. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male habitus: 1. Dorsal view; 2. Lateral view; 3. Face; 4. Head and thorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758859" httpUri="https://zenodo.org/record/12758859/files/figure.png" pageId="2" pageNumber="365">Figs 14</figureCitation>
). Head including eyes as broader as pronotum, crown light yellowish green (
<figureCitation id="13452A03FFF1CF771FDCFDF40CFF051E" box="[402,467,605,631]" captionStart="FIGURES 14" captionStartId="2.[151,264,1905,1929]" captionTargetBox="[157,1429,805,1875]" captionTargetId="figure-522@2.[151,1435,799,1881]" captionTargetPageId="2" captionText="FIGURES 14. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male habitus: 1. Dorsal view; 2. Lateral view; 3. Face; 4. Head and thorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758859" httpUri="https://zenodo.org/record/12758859/files/figure.png" pageId="2" pageNumber="365">Fig. 1</figureCitation>
), face light green, without markings. Antennae long, 1.5 x width of the head or longer, scape and pedicel brownish yellow, flagellum dark brown (
<figureCitation id="13452A03FFF1CF771D6DFD280E4A05F2" box="[803,870,641,667]" captionStart="FIGURES 14" captionStartId="2.[151,264,1905,1929]" captionTargetBox="[157,1429,805,1875]" captionTargetId="figure-522@2.[151,1435,799,1881]" captionTargetPageId="2" captionText="FIGURES 14. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male habitus: 1. Dorsal view; 2. Lateral view; 3. Face; 4. Head and thorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758859" httpUri="https://zenodo.org/record/12758859/files/figure.png" pageId="2" pageNumber="365">Fig. 3</figureCitation>
). Pronotum light yellowish green, posterior margin brownish yellow, with some dark brown irroration on anterior margin. Scutellum brownish yellow, with a pair median of longitudinal light green bands present or absent in some individuals (
<figureCitation id="13452A03FFF1CF771A6CFD600945058A" box="[1058,1129,713,739]" captionStart="FIGURES 14" captionStartId="2.[151,264,1905,1929]" captionTargetBox="[157,1429,805,1875]" captionTargetId="figure-522@2.[151,1435,799,1881]" captionTargetPageId="2" captionText="FIGURES 14. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male habitus: 1. Dorsal view; 2. Lateral view; 3. Face; 4. Head and thorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758859" httpUri="https://zenodo.org/record/12758859/files/figure.png" pageId="2" pageNumber="365">Fig. 4</figureCitation>
). Forewing light brownish, venation dark brown. Legs pale yellow, unmarked, except for darker bases of macrosetae (
<figureCitation id="13452A03FFF1CF771AC2FD4409FD046E" box="[1164,1233,749,775]" captionStart="FIGURES 14" captionStartId="2.[151,264,1905,1929]" captionTargetBox="[157,1429,805,1875]" captionTargetId="figure-522@2.[151,1435,799,1881]" captionTargetPageId="2" captionText="FIGURES 14. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male habitus: 1. Dorsal view; 2. Lateral view; 3. Face; 4. Head and thorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758859" httpUri="https://zenodo.org/record/12758859/files/figure.png" pageId="2" pageNumber="365">Fig. 2</figureCitation>
).
</paragraph>
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<emphasis id="B90AEA94FFF1CF771ED9F8D80C1600E0" bold="true" box="[151,314,1905,1929]" pageId="2" pageNumber="365">FIGURES 14.</emphasis>
<taxonomicName id="4C7E4D05FFF1CF771F0FF8D80FED00E0" authority="Hassan &amp; Xing" authorityName="Hassan &amp; Shah &amp; Xing" authorityYear="2024" box="[321,705,1905,1929]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov.">
<emphasis id="B90AEA94FFF1CF771F0FF8D80F3200E0" box="[321,542,1905,1929]" italics="true" pageId="2" pageNumber="365">Fistulatus motuoensis</emphasis>
Hassan &amp; Xing
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<taxonomicNameLabel id="A23957EFFFF1CF771C89F8D80E3400E0" box="[711,792,1905,1929]" pageId="2" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
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Male habitus: 1. Dorsal view; 2. Lateral view; 3. Face; 4. Head and thorax, dorsal view.
</paragraph>
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<paragraph id="8BC13686FFF0CF761ED9F89F08A900FF" blockId="3.[151,1436,1846,1942]" pageId="3" pageNumber="366">
<emphasis id="B90AEA94FFF0CF761ED9F89F0C650027" bold="true" box="[151,329,1846,1870]" pageId="3" pageNumber="366">FIGURES 513.</emphasis>
<taxonomicName id="4C7E4D05FFF0CF761F1FF89F0FF50027" authority="Hassan &amp; Xing" authorityName="Hassan &amp; Shah &amp; Xing" authorityYear="2024" box="[337,729,1846,1870]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="366" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov.">
<emphasis id="B90AEA94FFF0CF761F1FF89F0F1D0027" box="[337,561,1846,1870]" italics="true" pageId="3" pageNumber="366">Fistulatus motuoensis</emphasis>
Hassan &amp; Xing
</taxonomicName>
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<taxonomicNameLabel id="A23957EFFFF0CF761CAFF89F0E1F0027" box="[737,819,1846,1870]" pageId="3" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male. 5. Genitalia (after maceration), lateral view; 6. Male pygofer, lateral view; 7. Valve and subgenital plate, ventral view; 8. Style, lateral view; 9. Connective, dorsal view; 10. Aedeagus, dorsal view; 11. Aedeagus, ventral view; 12. Aedeagus, lateral view; 13. Aedeagus, anteroventral view focusing on the apex.
</paragraph>
</caption>
<paragraph id="8BC13686FFF7CF711E89FF3E0F0F06E0" blockId="4.[151,1437,150,645]" pageId="4" pageNumber="367">
Male genitalia (
<figureCitation id="13452A03FFF7CF711F36FF3E0CC007D8" box="[376,492,150,177]" captionStart="FIGURES 513" captionStartId="3.[151,264,1846,1870]" captionTargetBox="[158,1429,188,1816]" captionTargetId="figure-17@3.[151,1435,181,1822]" captionTargetPageId="3" captionText="FIGURES 513. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male. 5. Genitalia (after maceration), lateral view; 6. Male pygofer, lateral view; 7. Valve and subgenital plate, ventral view; 8. Style, lateral view; 9. Connective, dorsal view; 10.Aedeagus, dorsal view; 11. Aedeagus, ventral view; 12. Aedeagus, lateral view; 13. Aedeagus, anteroventral view focusing on the apex." figureDoi="http://doi.org/10.5281/zenodo.12758861" httpUri="https://zenodo.org/record/12758861/files/figure.png" pageId="4" pageNumber="367">Figs 513</figureCitation>
). Pygofer slightly wider than long, posteroventral margin of lobe with long, upward directed process, posterodorsally rounded with triangular apex, covered with numerous stout setae at posterolaterally (
<figureCitation id="13452A03FFF7CF711ED1FF760DCB0790" box="[159,231,223,249]" captionStart="FIGURES 513" captionStartId="3.[151,264,1846,1870]" captionTargetBox="[158,1429,188,1816]" captionTargetId="figure-17@3.[151,1435,181,1822]" captionTargetPageId="3" captionText="FIGURES 513. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male. 5. Genitalia (after maceration), lateral view; 6. Male pygofer, lateral view; 7. Valve and subgenital plate, ventral view; 8. Style, lateral view; 9. Connective, dorsal view; 10.Aedeagus, dorsal view; 11. Aedeagus, ventral view; 12. Aedeagus, lateral view; 13. Aedeagus, anteroventral view focusing on the apex." figureDoi="http://doi.org/10.5281/zenodo.12758861" httpUri="https://zenodo.org/record/12758861/files/figure.png" pageId="4" pageNumber="367">Fig. 6</figureCitation>
). Valve triangular. Subgenital plate triangular with narrow and elongated apex (
<figureCitation id="13452A03FFF7CF711ACCFF7609E50790" box="[1154,1225,223,249]" captionStart="FIGURES 513" captionStartId="3.[151,264,1846,1870]" captionTargetBox="[158,1429,188,1816]" captionTargetId="figure-17@3.[151,1435,181,1822]" captionTargetPageId="3" captionText="FIGURES 513. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male. 5. Genitalia (after maceration), lateral view; 6. Male pygofer, lateral view; 7. Valve and subgenital plate, ventral view; 8. Style, lateral view; 9. Connective, dorsal view; 10.Aedeagus, dorsal view; 11. Aedeagus, ventral view; 12. Aedeagus, lateral view; 13. Aedeagus, anteroventral view focusing on the apex." figureDoi="http://doi.org/10.5281/zenodo.12758861" httpUri="https://zenodo.org/record/12758861/files/figure.png" pageId="4" pageNumber="367">Fig. 7</figureCitation>
). Connective very short, Y-shaped, stem subequal to arms in length (
<figureCitation id="13452A03FFF7CF711C96FEAA0E320674" box="[728,798,259,285]" captionStart="FIGURES 513" captionStartId="3.[151,264,1846,1870]" captionTargetBox="[158,1429,188,1816]" captionTargetId="figure-17@3.[151,1435,181,1822]" captionTargetPageId="3" captionText="FIGURES 513. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male. 5. Genitalia (after maceration), lateral view; 6. Male pygofer, lateral view; 7. Valve and subgenital plate, ventral view; 8. Style, lateral view; 9. Connective, dorsal view; 10.Aedeagus, dorsal view; 11. Aedeagus, ventral view; 12. Aedeagus, lateral view; 13. Aedeagus, anteroventral view focusing on the apex." figureDoi="http://doi.org/10.5281/zenodo.12758861" httpUri="https://zenodo.org/record/12758861/files/figure.png" pageId="4" pageNumber="367">Fig. 9</figureCitation>
). Style moderately long, broad at base, curved laterally with apex rostriform (
<figureCitation id="13452A03FFF7CF711FDCFE8E0CF40628" box="[402,472,295,321]" captionStart="FIGURES 513" captionStartId="3.[151,264,1846,1870]" captionTargetBox="[158,1429,188,1816]" captionTargetId="figure-17@3.[151,1435,181,1822]" captionTargetPageId="3" captionText="FIGURES 513. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male. 5. Genitalia (after maceration), lateral view; 6. Male pygofer, lateral view; 7. Valve and subgenital plate, ventral view; 8. Style, lateral view; 9. Connective, dorsal view; 10.Aedeagus, dorsal view; 11. Aedeagus, ventral view; 12. Aedeagus, lateral view; 13. Aedeagus, anteroventral view focusing on the apex." figureDoi="http://doi.org/10.5281/zenodo.12758861" httpUri="https://zenodo.org/record/12758861/files/figure.png" pageId="4" pageNumber="367">Fig. 8</figureCitation>
). Aedeagus long and straight, with a pair of slender, retrorse subapical lateral spines arising from convex lateral flanges; shaft broadened at base and subapically in dorsal view, with apex fish-shaped in anteroventral view (
<figureCitation id="13452A03FFF7CF711FDAFEC60F3B06E0" box="[404,535,367,393]" captionStart="FIGURES 513" captionStartId="3.[151,264,1846,1870]" captionTargetBox="[158,1429,188,1816]" captionTargetId="figure-17@3.[151,1435,181,1822]" captionTargetPageId="3" captionText="FIGURES 513. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male. 5. Genitalia (after maceration), lateral view; 6. Male pygofer, lateral view; 7. Valve and subgenital plate, ventral view; 8. Style, lateral view; 9. Connective, dorsal view; 10.Aedeagus, dorsal view; 11. Aedeagus, ventral view; 12. Aedeagus, lateral view; 13. Aedeagus, anteroventral view focusing on the apex." figureDoi="http://doi.org/10.5281/zenodo.12758861" httpUri="https://zenodo.org/record/12758861/files/figure.png" pageId="4" pageNumber="367">Figs 1013</figureCitation>
).
</paragraph>
<paragraph id="8BC13686FFF7CF711E89FE3A0EC906C4" blockId="4.[151,1437,150,645]" box="[199,997,402,429]" pageId="4" pageNumber="367">
<emphasis id="B90AEA94FFF7CF711E89FE3A0C5F06C4" bold="true" box="[199,371,403,429]" pageId="4" pageNumber="367">Measurement.</emphasis>
Length (including tegmen):
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, 6.97.0 mm,
<specimenCount id="9D78FD0FFFF7CF711D22FE3A0EAD06C4" box="[876,897,403,429]" count="1" pageId="4" pageNumber="367" type="female"></specimenCount>
<quantity id="4C869B63FFF7CF711DC6FE3B0ECD06C4" box="[904,993,402,429]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="367" unit="mm" value="7.5">7.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF7CF711E89FE1E0E43069C" pageId="4" pageNumber="367" type="materials_examined">
<paragraph id="8BC13686FFF7CF711E89FE1E0E43069C" blockId="4.[151,1437,150,645]" pageId="4" pageNumber="367">
<emphasis id="B90AEA94FFF7CF711E89FE1E0C5506B9" bold="true" box="[199,377,438,465]" pageId="4" pageNumber="367">Type material.</emphasis>
<materialsCitation id="3B163CDBFFF7CF711FCBFE1E0E43069C" collectingDate="2020-08-08" collectionCode="GUGC" collectorName="Yongjin Sui" country="China" county="Motuo County" location="Autonomous Region" pageId="4" pageNumber="367" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="54C58824FFF7CF711FCBFE1E0CC106B8" box="[389,493,439,465]" pageId="4" pageNumber="367" type="holotype">Holotype</typeStatus>
<specimenCount id="9D78FD0FFFF7CF711FB6FE1E0F2306B8" box="[504,527,439,465]" count="1" pageId="4" pageNumber="367" type="male"></specimenCount>
,
<collectingCountry id="F3697616FFF7CF711C6EFE1E0F5106B8" box="[544,637,439,465]" name="China" pageId="4" pageNumber="367">CHINA</collectingCountry>
:
<collectingRegion id="49BAF864FFF7CF711CC5FE1E0FF006B8" box="[651,732,439,465]" country="China" name="Xizang" pageId="4" pageNumber="367">Xizang</collectingRegion>
<location id="8EA1605DFFF7CF711CA8FE1E0EFA06B8" LSID="urn:lsid:plazi:treatment:03D78790FFF1CF711ED9FE640E3505EC:8EA1605DFFF7CF711CA8FE1E0EFA06B8" box="[742,982,439,465]" country="China" county="Motuo County" name="Autonomous Region" pageId="4" pageNumber="367" stateProvince="Xizang">Autonomous Region</location>
,
<collectingCounty id="62A04E0AFFF7CF711DA8FE1E09BC06B8" box="[998,1168,439,465]" pageId="4" pageNumber="367">Motuo County</collectingCounty>
,
<date id="FFC01046FFF7CF711AD1FE1E087006B8" box="[1183,1372,439,465]" pageId="4" pageNumber="367" value="2020-08-08">
<collectingDate id="EF84E9AEFFF7CF711AD1FE1E087006B8" box="[1183,1372,439,465]" pageId="4" pageNumber="367" value="2020-08-08">08 August 2020</collectingDate>
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, coll.
<collectorName id="268B5350FFF7CF711ED9FE720C30069C" box="[151,284,475,501]" pageId="4" pageNumber="367">Yongjin Sui</collectorName>
(
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);
<typeStatus id="54C58824FFF7CF711FDAFE720F28069C" box="[404,516,475,501]" pageId="4" pageNumber="367" type="paratype">paratypes</typeStatus>
:
<specimenCount id="9D78FD0FFFF7CF711C40FE720F1F069C" box="[526,563,475,501]" count="1" pageId="4" pageNumber="367" type="male">1♂</specimenCount>
,
<specimenCount id="9D78FD0FFFF7CF711C71FE720F4E069C" box="[575,610,475,501]" count="1" pageId="4" pageNumber="367" type="female">1♀</specimenCount>
, same data as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF7CF711E89FE5609EF0570" box="[199,1219,511,537]" pageId="4" pageNumber="367" type="etymology">
<paragraph id="8BC13686FFF7CF711E89FE5609EF0570" blockId="4.[151,1437,150,645]" box="[199,1219,511,537]" pageId="4" pageNumber="367">
<emphasis id="B90AEA94FFF7CF711E89FE560C620570" bold="true" box="[199,334,511,537]" pageId="4" pageNumber="367">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C7E4D05FFF7CF711C77FE560FEC0571" authorityName="Hassan &amp; Shah &amp; Xing" authorityYear="2024" box="[569,704,511,537]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="motuoensis">
<emphasis id="B90AEA94FFF7CF711C0CFE560FEC0571" box="[578,704,511,536]" italics="true" pageId="4" pageNumber="367">motuoensis</emphasis>
</taxonomicName>
refers to the
<typeStatus id="54C58824FFF7CF711D15FE560EA70570" box="[859,907,511,537]" pageId="4" pageNumber="367">type</typeStatus>
locality of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF7CF711E89FD8A0E3505EC" pageId="4" pageNumber="367" type="discussion">
<paragraph id="8BC13686FFF7CF711E89FD8A0E3505EC" blockId="4.[151,1437,150,645]" pageId="4" pageNumber="367">
<emphasis id="B90AEA94FFF7CF711E89FD8A0C170554" bold="true" box="[199,315,547,573]" pageId="4" pageNumber="367">Remarks.</emphasis>
<taxonomicName id="4C7E4D05FFF7CF711F0DFD8A0FB80554" authority="Hassan &amp; Xing" authorityName="Hassan &amp; Shah &amp; Xing" authorityYear="2024" box="[323,660,547,573]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov.">
<emphasis id="B90AEA94FFF7CF711F0DFD8A0CF20555" box="[323,478,547,572]" italics="true" pageId="4" pageNumber="367">F. motuoensis</emphasis>
Hassan &amp; Xing
</taxonomicName>
<emphasis id="B90AEA94FFF7CF711CD2FD8A0FDB0554" bold="true" box="[668,759,547,573]" pageId="4" pageNumber="367">
<taxonomicNameLabel id="A23957EFFFF7CF711CD2FD8A0FDB0554" box="[668,759,547,573]" pageId="4" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is unique among its known congeners in having the ventral pygofer appendage relatively long and straight and by the aedeagal shaft with slender paired lateral prepical spines and the apex broadened and hammer-like in lateral view..
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,299 @@
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<mods:title id="CC173A326CC064DA69D4F912AB755FA2">Key to species of the leafhopper genus Fistulatus Zhang, Zhang &amp; Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species</mods:title>
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<paragraph id="8BC13686FFF4CF721ED9F9660FCA0180" blockId="7.[151,742,1743,1806]" box="[151,742,1743,1770]" pageId="7" pageNumber="370">
<heading id="D08981EAFFF4CF721ED9F9660FCA0180" bold="true" box="[151,742,1743,1770]" fontSize="11" level="1" pageId="7" pageNumber="370" reason="1">
<taxonomicName id="4C7E4D05FFF4CF721ED9F9660FCA0180" ID-CoL="6J6WR" authority="Zhang, Zhang &amp; Chen, 1997" authorityName="Zhang, Zhang &amp; Chen" authorityYear="1997" box="[151,742,1743,1770]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="sinensis">
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<emphasis id="B90AEA94FFF4CF721ED9F9660C550180" bold="true" box="[151,377,1743,1770]" italics="true" pageId="7" pageNumber="370">Fistulatus sinensis</emphasis>
<bibRefCitation id="EFEF4B77FFF4CF721FCEF9660FCA0180" author="Zhang, Y. L. &amp; Zhang, W. Z. &amp; Chen, B." box="[384,742,1743,1769]" pageId="7" pageNumber="370" pagination="235 - 245" refId="ref5993" refString="Zhang, Y. L., Zhang, W. Z. &amp; Chen, B. (1997) Selenocephaline leafhoppers (Homoptera: Cicadellidae) from Mt. Funiushan in Henan Province. Entomotaxonomia, 19, 235 - 245." type="journal article" year="1997">Zhang, Zhang &amp; Chen, 1997</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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(
<figureCitation id="13452A03FFF4CF721ED1F95A0C330067" box="[159,287,1779,1806]" captionStart="FIGURES 4059" captionStartId="7.[151,264,1421,1445]" captionTargetBox="[157,1430,187,1391]" captionTargetId="figure-17@7.[151,1435,181,1397]" captionTargetPageId="7" captionText="FIGURES 4059. Fistulatus species. 4044: F. monkoboensis (after Shang &amp; Zhang, 2003), 40: male pygofer, lateral view; 41: subgenital plate, valve, style and connective, ventral view; 42: aedeagus, dorsal view; 43: aedeagus, ventral view; 44: aedeagus, lateral view; 4549: F. rectilineus (after Shang et al., 2006), 45: male pygofer, lateral view; 46: style, dorsal view; 47: connective, dorsal view; 48: aedeagus, ventral view; 49: aedeagus, lateral view; 5054: F. sinensis (after Zhang et al., 1997), 50: male pygofer, lateral view; 51: style, dorsal view; 52: connective, dorsal view; 53: aedeagus, posteroventral view; 54: aedeagus, lateral view; 5559: F. quadrispinosus (after Lu &amp; Zhang, 2014), 55: male pygofer, lateral view; 56: subgenital plate, valve, style and connective, ventral view; 57: aedeagus, lateral view; 58: aedeagus, ventral view; 59: aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758871" httpUri="https://zenodo.org/record/12758871/files/figure.png" pageId="7" pageNumber="370">Figs 5054</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF4CF721ED9F8910F9B0039" box="[151,695,1848,1872]" pageId="7" pageNumber="370" type="reference_group">
<paragraph id="8BC13686FFF4CF721ED9F8910F9B0039" blockId="7.[151,695,1848,1872]" box="[151,695,1848,1872]" pageId="7" pageNumber="370">
<treatmentCitationGroup id="AB6E11A8FFF4CF721ED9F8910F9B0039" box="[151,695,1848,1872]" pageId="7" pageNumber="370">
<treatmentCitation id="0ADF1097FFF4CF721ED9F8910F9F0039" author="Zhang, Y. L. &amp; Zhang, W. Z. &amp; Chen, B." box="[151,691,1848,1872]" page="237" pageId="7" pageNumber="370" year="1997">
<taxonomicName id="4C7E4D05FFF4CF721ED9F8910F9F0039" ID-CoL="6J6WR" authority="Zhang, Zhang &amp; Chen, 1997: 237" authorityName="Zhang, Zhang &amp; Chen" authorityPageNumber="237" authorityYear="1997" box="[151,691,1848,1872]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B90AEA94FFF4CF721ED9F8910C7F0026" box="[151,339,1848,1872]" italics="true" pageId="7" pageNumber="370">Fistulatus sinensis</emphasis>
<bibRefCitation id="EFEF4B77FFF4CF721F17F8910F9F0039" author="Zhang, Y. L. &amp; Zhang, W. Z. &amp; Chen, B." box="[345,691,1848,1872]" pageId="7" pageNumber="370" pagination="235 - 245" refId="ref5993" refString="Zhang, Y. L., Zhang, W. Z. &amp; Chen, B. (1997) Selenocephaline leafhoppers (Homoptera: Cicadellidae) from Mt. Funiushan in Henan Province. Entomotaxonomia, 19, 235 - 245." type="journal article" year="1997">Zhang, Zhang &amp; Chen, 1997: 237</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF4CF7D1ED9F8D7080F0674" lastPageId="8" lastPageNumber="371" pageId="7" pageNumber="370" type="materials_examined">
<paragraph id="8BC13686FFF4CF721ED9F8D708B00089" blockId="7.[151,1436,1918,2017]" pageId="7" pageNumber="370">
<emphasis id="B90AEA94FFF4CF721ED9F8D70CAF00F1" bold="true" box="[151,387,1918,1944]" pageId="7" pageNumber="370">Material examined.</emphasis>
<materialsCitation id="3B163CDBFFF4CF721FC2F8D70C0300D5" collectingDate="2010-08-12" collectingDateMax="2010-08-13" collectingDateMin="2010-08-12" collectionCode="GUGC" collectorName="Zhihua Fan" country="China" location="Xiaowutai" municipality="Zhangjiakou City" pageId="7" pageNumber="370" specimenCount="1" specimenCount-male="1" stateProvince="Hebei">
<emphasis id="B90AEA94FFF4CF721FC2F8D70CDF00F1" bold="true" box="[396,499,1918,1944]" pageId="7" pageNumber="370">
<collectingCountry id="F3697616FFF4CF721FC2F8D70CC200F1" box="[396,494,1918,1944]" name="China" pageId="7" pageNumber="370">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="9D78FD0FFFF4CF721FB2F8D70F0D00F1" box="[508,545,1918,1944]" count="1" pageId="7" pageNumber="370" type="male">1♂</specimenCount>
,
<collectingRegion id="49BAF864FFF4CF721C7EF8D70F9600F1" box="[560,698,1918,1944]" country="China" name="Hebei" pageId="7" pageNumber="370">Hebei Prov.</collectingRegion>
,
<collectingMunicipality id="6BA5ACFCFFF4CF721C89F8D70EA300F1" box="[711,911,1918,1944]" pageId="7" pageNumber="370">Zhangjiakou City</collectingMunicipality>
,
<location id="8EA1605DFFF4CF721DD3F8D7093900F1" LSID="urn:lsid:plazi:treatment:03D78790FFF4CF7D1ED9F9660FD506C4:8EA1605DFFF4CF721DD3F8D7093900F1" box="[925,1045,1918,1944]" country="China" municipality="Zhangjiakou City" name="Xiaowutai" pageId="7" pageNumber="370" stateProvince="Hebei">Xiaowutai</location>
,
<date id="FFC01046FFF4CF721A6CF8D7082900F1" box="[1058,1285,1918,1944]" pageId="7" pageNumber="370" value="2010-08-12" valueMax="2010-08-13" valueMin="2010-08-12">
<collectingDate id="EF84E9AEFFF4CF721A6CF8D7082900F1" box="[1058,1285,1918,1944]" pageId="7" pageNumber="370" value="2010-08-12" valueMax="2010-08-13" valueMin="2010-08-12">1213 August 2010</collectingDate>
</date>
, coll.
<collectorName id="268B5350FFF4CF721B03F8D70DED00D5" pageId="7" pageNumber="370">Zhihua Fan</collectorName>
(
<collectionCode id="ED6FAE43FFF4CF721E9FF80B0C0B00D5" box="[209,295,1954,1980]" pageId="7" pageNumber="370">GUGC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B163CDBFFF4CF721F75F80B0C120089" collectingDate="2012-07-23" collectingDateMax="2012-07-26" collectingDateMin="2012-07-23" collectionCode="GUGC" collectorName="Qiongzhang Song" country="China" county="Yicheng County" location="Lishan" municipality="Linfen City" pageId="7" pageNumber="370" specimenCount="9" specimenCount-female="6" specimenCount-male="3" stateProvince="Shanxi">
<specimenCount id="9D78FD0FFFF4CF721F75F80B0C5F00D5" box="[315,371,1954,1980]" count="3" pageId="7" pageNumber="370" type="male">3♂♂</specimenCount>
<specimenCount id="9D78FD0FFFF4CF721F35F80B0C9800D5" box="[379,436,1954,1980]" count="6" pageId="7" pageNumber="370" type="female">6♀♀</specimenCount>
,
<collectingRegion id="49BAF864FFF4CF721F8CF80B0F7900D5" box="[450,597,1954,1980]" country="China" name="Shanxi" pageId="7" pageNumber="370">Shanxi Prov.</collectingRegion>
,
<collectingMunicipality id="6BA5ACFCFFF4CF721C2EF80B0FC800D5" box="[608,740,1954,1980]" pageId="7" pageNumber="370">Linfen City</collectingMunicipality>
,
<collectingCounty id="62A04E0AFFF4CF721CA1F80B0E8400D5" box="[751,936,1954,1980]" pageId="7" pageNumber="370">Yicheng County</collectingCounty>
,
<location id="8EA1605DFFF4CF721DFAF80B092F00D5" LSID="urn:lsid:plazi:treatment:03D78790FFF4CF7D1ED9F9660FD506C4:8EA1605DFFF4CF721DFAF80B092F00D5" box="[948,1027,1954,1980]" country="China" county="Yicheng County" municipality="Linfen City" name="Lishan" pageId="7" pageNumber="370" stateProvince="Shanxi">Lishan</location>
,
<date id="FFC01046FFF4CF721A41F80B09E300D5" box="[1039,1231,1954,1980]" pageId="7" pageNumber="370" value="2012-07-23" valueMax="2012-07-26" valueMin="2012-07-23">
<collectingDate id="EF84E9AEFFF4CF721A41F80B09E300D5" box="[1039,1231,1954,1980]" pageId="7" pageNumber="370" value="2012-07-23" valueMax="2012-07-26" valueMin="2012-07-23">2326 July 2012</collectingDate>
</date>
, coll.
<collectorName id="268B5350FFF4CF721B5DF80B0DFD0089" pageId="7" pageNumber="370">Qiongzhang Song</collectorName>
(
<collectionCode id="ED6FAE43FFF4CF721EAEF86F0C1A0089" box="[224,310,1990,2016]" pageId="7" pageNumber="370">GUGC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B163CDBFFF4CF721F07F86F08B40089" box="[329,1432,1990,2017]" collectingDate="2011-07-24" collectingDateMax="2011-07-25" collectingDateMin="2011-07-24" collectionCode="GUGC" collectorName="Hu Li" country="China" county="Yicheng County" location="Lishan" municipality="Linfen City" pageId="7" pageNumber="370" specimenCount="10" specimenCount-female="10" stateProvince="Shanxi">
<specimenCount id="9D78FD0FFFF4CF721F07F86F0CBD0089" box="[329,401,1990,2016]" count="10" pageId="7" pageNumber="370" type="female">10♀♀</specimenCount>
,
<collectingRegion id="49BAF864FFF4CF721FD0F86F0F1C0089" box="[414,560,1990,2016]" country="China" name="Shanxi" pageId="7" pageNumber="370">Shanxi Prov.</collectingRegion>
,
<collectingMunicipality id="6BA5ACFCFFF4CF721C75F86F0F920089" box="[571,702,1990,2016]" pageId="7" pageNumber="370">Linfen City</collectingMunicipality>
,
<collectingCounty id="62A04E0AFFF4CF721C87F86F0EAE0089" box="[713,898,1990,2016]" pageId="7" pageNumber="370">Yicheng County</collectingCounty>
,
<location id="8EA1605DFFF4CF721DC3F86F0EF00089" LSID="urn:lsid:plazi:treatment:03D78790FFF4CF7D1ED9F9660FD506C4:8EA1605DFFF4CF721DC3F86F0EF00089" box="[909,988,1990,2016]" country="China" county="Yicheng County" municipality="Linfen City" name="Lishan" pageId="7" pageNumber="370" stateProvince="Shanxi">Lishan</location>
,
<date id="FFC01046FFF4CF721DA9F86F09890089" box="[999,1189,1990,2017]" pageId="7" pageNumber="370" value="2011-07-24" valueMax="2011-07-25" valueMin="2011-07-24">
<collectingDate id="EF84E9AEFFF4CF721DA9F86F09890089" box="[999,1189,1990,2017]" pageId="7" pageNumber="370" value="2011-07-24" valueMax="2011-07-25" valueMin="2011-07-24">2425 July 2011</collectingDate>
</date>
, coll.
<collectorName id="268B5350FFF4CF721AA7F86F08070089" box="[1257,1323,1990,2016]" pageId="7" pageNumber="370">Hu Li</collectorName>
(
<collectionCode id="ED6FAE43FFF4CF721B74F86F08BC0089" box="[1338,1424,1990,2016]" pageId="7" pageNumber="370">GUGC</collectionCode>
)
</materialsCitation>
;
</paragraph>
<paragraph id="8BC13686FFFBCF7D1ED9FF3E080F0674" blockId="8.[151,1437,151,429]" pageId="8" pageNumber="371">
<materialsCitation id="3B163CDBFFFBCF7D1ED9FF3E0DD107BC" collectingDate="2012-07-12" collectingDateMax="2012-07-17" collectingDateMin="2012-07-12" collectionCode="GUGC" collectorName="Qiongzhang Song" county="Dushan County" location="Duliu Jiangyuan Wetland" pageId="8" pageNumber="371" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Guizhou">
<specimenCount id="9D78FD0FFFFBCF7D1ED9FF3E0DE307D8" box="[151,207,151,177]" count="6" pageId="8" pageNumber="371" type="male">6♂♂</specimenCount>
<specimenCount id="9D78FD0FFFFBCF7D1E97FF3E0C3E07D8" box="[217,274,151,177]" count="4" pageId="8" pageNumber="371" type="female">4♀♀</specimenCount>
,
<collectingRegion id="49BAF864FFFBCF7D1F6FFF3E0CE407D8" box="[289,456,151,177]" country="China" name="Guizhou" pageId="8" pageNumber="371">Guizhou Prov.</collectingRegion>
,
<collectingCounty id="62A04E0AFFFBCF7D1F9BFF3E0FA407D8" box="[469,648,151,177]" pageId="8" pageNumber="371">Dushan County</collectingCounty>
,
<location id="8EA1605DFFFBCF7D1CDBFF3E0E9507D8" LSID="urn:lsid:plazi:treatment:03D78790FFF4CF7D1ED9F9660FD506C4:8EA1605DFFFBCF7D1CDBFF3E0E9507D8" box="[661,953,151,177]" county="Dushan County" name="Duliu Jiangyuan Wetland" pageId="8" pageNumber="371" stateProvince="Guizhou">Duliu Jiangyuan Wetland</location>
,
<date id="FFC01046FFFBCF7D1D88FF3E09A507D8" box="[966,1161,151,177]" pageId="8" pageNumber="371" value="2012-07-12" valueMax="2012-07-17" valueMin="2012-07-12">
<collectingDate id="EF84E9AEFFFBCF7D1D88FF3E09A507D8" box="[966,1161,151,177]" pageId="8" pageNumber="371" value="2012-07-12" valueMax="2012-07-17" valueMin="2012-07-12">1217 July 2012</collectingDate>
</date>
, coll.
<collectorName id="268B5350FFFBCF7D1A9EFF3E08B107D8" box="[1232,1437,151,177]" pageId="8" pageNumber="371">Qiongzhang Song</collectorName>
(
<collectionCode id="ED6FAE43FFFBCF7D1ED1FF120DD907BC" box="[159,245,187,213]" pageId="8" pageNumber="371">GUGC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B163CDBFFFBCF7D1F44FF12084907BC" box="[266,1381,187,213]" collectingDate="2011-07-08" collectionCode="GUGC" collectorName="Hu Li" county="Xinglong County" location="Wulingshan" municipality="Chengde City" pageId="8" pageNumber="371" specimenCount="1" specimenCount-male="1" stateProvince="Hebei">
<specimenCount id="9D78FD0FFFFBCF7D1F44FF120C0207BC" box="[266,302,187,213]" count="1" pageId="8" pageNumber="371" type="male">1♂</specimenCount>
,
<collectingRegion id="49BAF864FFFBCF7D1F72FF120CE907BC" box="[316,453,187,213]" country="China" name="Hebei" pageId="8" pageNumber="371">Hebei Prov.</collectingRegion>
,
<collectingMunicipality id="6BA5ACFCFFFBCF7D1F9CFF120F4307BC" box="[466,623,187,213]" pageId="8" pageNumber="371">Chengde City</collectingMunicipality>
,
<collectingCounty id="62A04E0AFFFBCF7D1C32FF120E6F07BC" box="[636,835,187,213]" pageId="8" pageNumber="371">Xinglong County</collectingCounty>
,
<location id="8EA1605DFFFBCF7D1D01FF120EF407BC" LSID="urn:lsid:plazi:treatment:03D78790FFF4CF7D1ED9F9660FD506C4:8EA1605DFFFBCF7D1D01FF120EF407BC" box="[847,984,187,213]" county="Xinglong County" municipality="Chengde City" name="Wulingshan" pageId="8" pageNumber="371" stateProvince="Hebei">Wulingshan</location>
,
<date id="FFC01046FFFBCF7D1DABFF12094107BC" box="[997,1133,187,213]" pageId="8" pageNumber="371" value="2011-07-08">
<collectingDate id="EF84E9AEFFFBCF7D1DABFF12094107BC" box="[997,1133,187,213]" pageId="8" pageNumber="371" value="2011-07-08">8 July 2011</collectingDate>
</date>
, coll.
<collectorName id="268B5350FFFBCF7D1AFDFF1209DB07BC" box="[1203,1271,187,213]" pageId="8" pageNumber="371">Hu Li</collectorName>
(
<collectionCode id="ED6FAE43FFFBCF7D1B46FF12087207BC" box="[1288,1374,187,213]" pageId="8" pageNumber="371">GUGC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B163CDBFFFBCF7D1B3CFF1209C20790" collectingDate="2010-07-21" collectionCode="GUGC" collectorName="Hu Li &amp; Zhihua Fan" county="Liuba County" location="Yingpan Town" pageId="8" pageNumber="371" specimenCount="1" specimenCount-male="1" stateProvince="Shaanxi">
<specimenCount id="9D78FD0FFFFBCF7D1B3CFF1208BB07BC" box="[1394,1431,187,213]" count="1" pageId="8" pageNumber="371" type="male">1♂</specimenCount>
,
<collectingRegion id="49BAF864FFFBCF7D1ED9FF760C1B0790" box="[151,311,223,249]" country="China" name="Shaanxi" pageId="8" pageNumber="371">Shaanxi Prov.</collectingRegion>
,
<collectingCounty id="62A04E0AFFFBCF7D1F0AFF760CCE0790" box="[324,482,223,249]" pageId="8" pageNumber="371">Liuba County</collectingCounty>
,
<location id="8EA1605DFFFBCF7D1FA3FF760FBA0790" LSID="urn:lsid:plazi:treatment:03D78790FFF4CF7D1ED9F9660FD506C4:8EA1605DFFFBCF7D1FA3FF760FBA0790" box="[493,662,223,249]" county="Liuba County" name="Yingpan Town" pageId="8" pageNumber="371" stateProvince="Shaanxi">Yingpan Town</location>
,
<date id="FFC01046FFFBCF7D1CEDFF760E160790" box="[675,826,223,249]" pageId="8" pageNumber="371" value="2010-07-21">
<collectingDate id="EF84E9AEFFFBCF7D1CEDFF760E160790" box="[675,826,223,249]" pageId="8" pageNumber="371" value="2010-07-21">21 July 2010</collectingDate>
</date>
, coll.
<collectorName id="268B5350FFFBCF7D1DCEFF760EE80790" box="[896,964,223,249]" pageId="8" pageNumber="371">Hu Li</collectorName>
and
<collectorName id="268B5350FFFBCF7D1DB3FF7609530790" box="[1021,1151,223,249]" pageId="8" pageNumber="371">Zhihua Fan</collectorName>
(
<collectionCode id="ED6FAE43FFFBCF7D1ADEFF7609CA0790" box="[1168,1254,223,249]" pageId="8" pageNumber="371">GUGC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B163CDBFFFBCF7D1AB4FF7608330674" collectingDate="2012-07-12" collectionCode="GUGC" collectorName="Weicheng Yang &amp; Weibin Zheng" location="Ningshan Huoditang Forest" municipality="Pro." pageId="8" pageNumber="371" specimenCount="3" specimenCount-male="3" stateProvince="Shaanxi">
<specimenCount id="9D78FD0FFFFBCF7D1AB4FF7608190790" box="[1274,1333,223,249]" count="3" pageId="8" pageNumber="371" type="male">3♂♂</specimenCount>
,
<collectingRegion id="49BAF864FFFBCF7D1B0CFF7608B00790" box="[1346,1436,223,249]" country="China" name="Shaanxi" pageId="8" pageNumber="371">Shaanxi</collectingRegion>
<collectingMunicipality id="6BA5ACFCFFFBCF7D1ED9FEAA0DE40674" box="[151,200,259,285]" pageId="8" pageNumber="371">Pro.</collectingMunicipality>
,
<location id="8EA1605DFFFBCF7D1E9DFEAA0F250674" LSID="urn:lsid:plazi:treatment:03D78790FFF4CF7D1ED9F9660FD506C4:8EA1605DFFFBCF7D1E9DFEAA0F250674" box="[211,521,259,285]" municipality="Pro." name="Ningshan Huoditang Forest" pageId="8" pageNumber="371" stateProvince="Shaanxi">Ningshan Huoditang Forest</location>
Farm,
<date id="FFC01046FFFBCF7D1C17FEAA0FC00674" box="[601,748,259,285]" pageId="8" pageNumber="371" value="2012-07-12">
<collectingDate id="EF84E9AEFFFBCF7D1C17FEAA0FC00674" box="[601,748,259,285]" pageId="8" pageNumber="371" value="2012-07-12">12 July 2012</collectingDate>
</date>
, coll.
<collectorName id="268B5350FFFBCF7D1D61FEAA0EF20674" box="[815,990,259,285]" pageId="8" pageNumber="371">Weicheng Yang</collectorName>
and
<collectorName id="268B5350FFFBCF7D1A5AFEAA099F0674" box="[1044,1203,259,285]" pageId="8" pageNumber="371">Weibin Zheng</collectorName>
(
<collectionCode id="ED6FAE43FFFBCF7D1A8FFEAA083B0674" box="[1217,1303,259,285]" pageId="8" pageNumber="371">GUGC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFFBCF7D1E89FE8E09C4060C" pageId="8" pageNumber="371" type="discussion">
<paragraph id="8BC13686FFFBCF7D1E89FE8E09C4060C" blockId="8.[151,1437,151,429]" pageId="8" pageNumber="371">
<emphasis id="B90AEA94FFFBCF7D1E89FE8E0C170628" bold="true" box="[199,315,295,321]" pageId="8" pageNumber="371">Remarks.</emphasis>
This species can be distinguished from its congeners by the aedeagal shaft with a pair of spinelike lateral preapical processes in posterior view and the style with and apical extension in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFFBCF7D1E89FEC70FD506C4" pageId="8" pageNumber="371" type="distribution">
<paragraph id="8BC13686FFFBCF7D1E89FEC70FD506C4" blockId="8.[151,1437,151,429]" pageId="8" pageNumber="371">
<emphasis id="B90AEA94FFFBCF7D1E89FEC70C4F06E1" bold="true" box="[199,355,366,392]" pageId="8" pageNumber="371">Distribution:</emphasis>
<collectingCountry id="F3697616FFFBCF7D1F25FEC60C9E06E0" box="[363,434,367,393]" name="China" pageId="8" pageNumber="371">China</collectingCountry>
:
<collectingRegion id="49BAF864FFFBCF7D1FF0FEC60F3706E0" box="[446,539,367,393]" country="China" name="Shaanxi" pageId="8" pageNumber="371">Shaanxi</collectingRegion>
,
<collectingRegion id="49BAF864FFFBCF7D1C66FEC60F5806E0" box="[552,628,367,393]" country="China" name="Henan" pageId="8" pageNumber="371">Henan</collectingRegion>
,
<collectingRegion id="49BAF864FFFBCF7D1CCFFEC60FE506E0" box="[641,713,367,393]" country="China" name="Gansu" pageId="8" pageNumber="371">Gansu</collectingRegion>
(
<bibRefCitation id="EFEF4B77FFFBCF7D1C96FEC60E8B06E0" author="Zhang, Y. L. &amp; Zhang, W. Z. &amp; Chen, B." box="[728,935,367,393]" pageId="8" pageNumber="371" pagination="235 - 245" refId="ref5993" refString="Zhang, Y. L., Zhang, W. Z. &amp; Chen, B. (1997) Selenocephaline leafhoppers (Homoptera: Cicadellidae) from Mt. Funiushan in Henan Province. Entomotaxonomia, 19, 235 - 245." type="journal article" year="1997">
Zhang
<emphasis id="B90AEA94FFFBCF7D1D64FEC60E4806E0" box="[810,868,367,393]" italics="true" pageId="8" pageNumber="371">et al.</emphasis>
1997
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;
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Shang
<emphasis id="B90AEA94FFFBCF7D1A9BFEC6082306E0" box="[1237,1295,367,393]" italics="true" pageId="8" pageNumber="371">et al.</emphasis>
2006
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<bibRefCitation id="EFEF4B77FFFBCF7D1B11FEC60C0E06C4" author="Lu, L. &amp; Zhang, Y. L." pageId="8" pageNumber="371" pagination="247 - 250" refId="ref5506" refString="Lu, L. &amp; Zhang, Y. L. (2014) Taxonomy of the Oriental leafhopper genus Fistulatus (Hemiptera: Cicadellidae: Deltocephalinae), with description of a new species from China. Zootaxa, 3838 (2), 247 - 250. https: // doi. org / 10.11646 / zootaxa. 3838.2.10" type="journal article" year="2014">Lu &amp; Zhang 2014</bibRefCitation>
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Qu
<emphasis id="B90AEA94FFFBCF7D1F18FE3A0CA206C4" box="[342,398,403,429]" italics="true" pageId="8" pageNumber="371">et al.</emphasis>
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View file

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<emphasis id="B90AEA94FFF5CF731ED9F9AA0CB40177" bold="true" box="[151,408,1539,1566]" italics="true" pageId="6" pageNumber="369">Fistulatus rectilineus</emphasis>
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(
<figureCitation id="13452A03FFF5CF731ED1F98E0C330128" box="[159,287,1575,1601]" captionStart="FIGURES 4059" captionStartId="7.[151,264,1421,1445]" captionTargetBox="[157,1430,187,1391]" captionTargetId="figure-17@7.[151,1435,181,1397]" captionTargetPageId="7" captionText="FIGURES 4059. Fistulatus species. 4044: F. monkoboensis (after Shang &amp; Zhang, 2003), 40: male pygofer, lateral view; 41: subgenital plate, valve, style and connective, ventral view; 42: aedeagus, dorsal view; 43: aedeagus, ventral view; 44: aedeagus, lateral view; 4549: F. rectilineus (after Shang et al., 2006), 45: male pygofer, lateral view; 46: style, dorsal view; 47: connective, dorsal view; 48: aedeagus, ventral view; 49: aedeagus, lateral view; 5054: F. sinensis (after Zhang et al., 1997), 50: male pygofer, lateral view; 51: style, dorsal view; 52: connective, dorsal view; 53: aedeagus, posteroventral view; 54: aedeagus, lateral view; 5559: F. quadrispinosus (after Lu &amp; Zhang, 2014), 55: male pygofer, lateral view; 56: subgenital plate, valve, style and connective, ventral view; 57: aedeagus, lateral view; 58: aedeagus, ventral view; 59: aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758871" httpUri="https://zenodo.org/record/12758871/files/figure.png" pageId="6" pageNumber="369">Figs 4449</figureCitation>
)
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<paragraph id="8BC13686FFF5CF731ED9F9C50E0C01EA" blockId="6.[151,800,1643,1668]" box="[151,800,1643,1668]" pageId="6" pageNumber="369">
<treatmentCitationGroup id="AB6E11A8FFF5CF731ED9F9C50E0C01EA" box="[151,800,1643,1668]" pageId="6" pageNumber="369">
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<emphasis id="B90AEA94FFF5CF731ED9F9C50C4101ED" box="[151,365,1644,1668]" italics="true" pageId="6" pageNumber="369">Fistulatus rectilineus</emphasis>
Shang &amp; Zhang
</taxonomicName>
in
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Shang
<emphasis id="B90AEA94FFF5CF731C34F9C50F8101ED" box="[634,685,1644,1668]" italics="true" pageId="6" pageNumber="369">et al.</emphasis>
2006: 152
</bibRefCitation>
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<paragraph id="8BC13686FFF5CF731ED9F9180C130199" blockId="6.[151,1436,1713,1885]" pageId="6" pageNumber="369">
<emphasis id="B90AEA94FFF5CF731ED9F9180CAC01A2" bold="true" box="[151,384,1713,1739]" pageId="6" pageNumber="369">Material examined.</emphasis>
<materialsCitation id="3B163CDBFFF5CF731FC9F9180C170199" collectingDate="2012-07-31" collectionCode="GUGC" collectorName="Zhihua Fan &amp; Qiuxiao Tang" country="China" county="Kangding County" location="Kangding County" pageId="6" pageNumber="369" specimenCount="2" specimenCount-male="2" stateProvince="Sichuan">
<emphasis id="B90AEA94FFF5CF731FC9F9180CC201A2" bold="true" box="[391,494,1713,1739]" pageId="6" pageNumber="369">
<collectingCountry id="F3697616FFF5CF731FC9F9180CC501A2" box="[391,489,1713,1739]" name="China" pageId="6" pageNumber="369">CHINA</collectingCountry>
:
</emphasis>
<specimenCount id="9D78FD0FFFF5CF731FBAF91B0F0301A5" box="[500,559,1714,1740]" count="2" pageId="6" pageNumber="369" type="male">2♂♂</specimenCount>
,
<collectingRegion id="49BAF864FFF5CF731C74F91B0FF401A5" box="[570,728,1714,1740]" country="China" name="Sichuan" pageId="6" pageNumber="369">Sichuan Prov.</collectingRegion>
,
<collectingCounty id="62A04E0AFFF5CF731CADF91B0E8001A5" box="[739,940,1714,1740]" pageId="6" pageNumber="369">Kangding County</collectingCounty>
,
<date id="FFC01046FFF5CF731DF8F91B096601A5" box="[950,1098,1714,1740]" pageId="6" pageNumber="369" value="2012-07-31">
<collectingDate id="EF84E9AEFFF5CF731DF8F91B096601A5" box="[950,1098,1714,1740]" pageId="6" pageNumber="369" value="2012-07-31">31 July 2012</collectingDate>
</date>
, coll.
<collectorName id="268B5350FFF5CF731AC2F91B082001A5" box="[1164,1292,1714,1740]" pageId="6" pageNumber="369">Zhihua Fan</collectorName>
and
<collectorName id="268B5350FFF5CF731B0CF91B0DE30199" pageId="6" pageNumber="369">Qiuxiao Tang</collectorName>
(
<collectionCode id="ED6FAE43FFF5CF731E90F97F0C180199" box="[222,308,1750,1776]" pageId="6" pageNumber="369">GUGC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF5CF731E89F9530E350051" pageId="6" pageNumber="369" type="discussion">
<paragraph id="8BC13686FFF5CF731E89F9530E350051" blockId="6.[151,1436,1713,1885]" pageId="6" pageNumber="369">
<emphasis id="B90AEA94FFF5CF731E89F9530C17007D" bold="true" box="[199,315,1786,1812]" pageId="6" pageNumber="369">Remarks.</emphasis>
This species can be distinguished by the medially directed pygofer processes and the cylindrical, straight, and elongated aedeagal shaft, without processes.
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF5CF731E89F8E8087A0034" box="[199,1366,1857,1885]" pageId="6" pageNumber="369" type="distribution">
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<emphasis id="B90AEA94FFF5CF731E89F8E80C4F0032" bold="true" box="[199,355,1857,1883]" pageId="6" pageNumber="369">Distribution:</emphasis>
<collectingCountry id="F3697616FFF5CF731F24F8EB0C9D0035" box="[362,433,1858,1884]" name="China" pageId="6" pageNumber="369">China</collectingCountry>
:
<collectingRegion id="49BAF864FFF5CF731FF5F8EB0F390035" box="[443,533,1858,1884]" country="China" name="Sichuan" pageId="6" pageNumber="369">Sichuan</collectingRegion>
(
<bibRefCitation id="EFEF4B77FFF5CF731C6AF8EB0FC20035" author="Zhang, Y. L. &amp; Zhang, W. Z. &amp; Chen, B." box="[548,750,1858,1884]" pageId="6" pageNumber="369" pagination="235 - 245" refId="ref5993" refString="Zhang, Y. L., Zhang, W. Z. &amp; Chen, B. (1997) Selenocephaline leafhoppers (Homoptera: Cicadellidae) from Mt. Funiushan in Henan Province. Entomotaxonomia, 19, 235 - 245." type="journal article" year="1997">
Zhang
<emphasis id="B90AEA94FFF5CF731C3AF8EA0F810035" box="[628,685,1858,1884]" italics="true" pageId="6" pageNumber="369">et al.</emphasis>
1997
</bibRefCitation>
;
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Shang
<emphasis id="B90AEA94FFF5CF731D09F8EA0EAC0035" box="[839,896,1858,1884]" italics="true" pageId="6" pageNumber="369">et al.</emphasis>
2006
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Qu
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<emphasis id="B90AEA94FFF5CF731ED9FBD50CEA03FE" bold="true" box="[151,454,1148,1175]" italics="true" pageId="6" pageNumber="369">Fistulatus monkoboensis</emphasis>
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(
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<emphasis id="B90AEA94FFF5CF731ED9FB4C0CBF0394" box="[151,403,1253,1277]" italics="true" pageId="6" pageNumber="369">Fistulatus monkoboensis</emphasis>
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<emphasis id="B90AEA94FFF5CF731ED9FA820C27022C" bold="true" box="[151,267,1323,1349]" pageId="6" pageNumber="369">Remarks.</emphasis>
This species is similar to
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<emphasis id="B90AEA94FFF5CF731C72FA850F8A022C" box="[572,678,1323,1349]" italics="true" pageId="6" pageNumber="369">F. athena</emphasis>
Shang &amp; Zhang
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in the shape of the pygofer processes but can be distinguished by the absence of lateral processes on the connective and by the style with apical process strongly curved and not digitate in lateral view.
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<emphasis id="B90AEA94FFF5CF731E89FA3E0C4F02D8" bold="true" box="[199,355,1431,1457]" pageId="6" pageNumber="369">Distribution:</emphasis>
<collectingCountry id="F3697616FFF5CF731F24FA3E0CF902D8" box="[362,469,1431,1457]" name="Malaysia" pageId="6" pageNumber="369">Malaysia</collectingCountry>
:
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(
<bibRefCitation id="EFEF4B77FFF5CF731C7CFA3E0E0B02D8" author="Shang, S. Q. &amp; Zhang, Y. L." box="[562,807,1431,1457]" pageId="6" pageNumber="369" pagination="21 - 24" refId="ref5694" refString="Shang, S. Q. &amp; Zhang, Y. L. (2003) Two new species from Sabah (Kalimantan) of the genus Fistulatus (Homoptera: Cicadellidae: Selenocephalinae: Paraboloponini). Entomotaxonomia, 25, 21 - 24." type="journal article" year="2003">Shang &amp; Zhang 2003</bibRefCitation>
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Shang
<emphasis id="B90AEA94FFF5CF731DCEFA310E9402D8" box="[896,952,1431,1457]" italics="true" pageId="6" pageNumber="369">et al.</emphasis>
2006
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;
<bibRefCitation id="EFEF4B77FFF5CF731A48FA3109FF02D8" author="Lu, L. &amp; Zhang, Y. L." box="[1030,1235,1431,1458]" pageId="6" pageNumber="369" pagination="247 - 250" refId="ref5506" refString="Lu, L. &amp; Zhang, Y. L. (2014) Taxonomy of the Oriental leafhopper genus Fistulatus (Hemiptera: Cicadellidae: Deltocephalinae), with description of a new species from China. Zootaxa, 3838 (2), 247 - 250. https: // doi. org / 10.11646 / zootaxa. 3838.2.10" type="journal article" year="2014">Lu &amp; Zhang 2014</bibRefCitation>
;
<bibRefCitation id="EFEF4B77FFF5CF731A90FA3E08AF02DB" author="Qu, L. &amp; Webb, M. D. &amp; Dai, R. H." box="[1246,1411,1431,1458]" pageId="6" pageNumber="369" pagination="260 - 270" refId="ref5384" refString="Qu, L., Webb, M. D. &amp; Dai, R. H. (2015) A new genus and species of the leafhopper subtribe Paraboloponina from China (Hemiptera, Cicadellidae). Zootaxa, 3919 (2), 260 - 270. https: // doi. org / 10.11646 / zootaxa. 3919.2.2" type="journal article" year="2015">
Qu
<emphasis id="B90AEA94FFF5CF731B49FA31081302D8" box="[1287,1343,1431,1457]" italics="true" pageId="6" pageNumber="369">et al.</emphasis>
2015
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<mods:namePart id="B20A05184C7DD21E510A473AA08224DB">Shah, Bismillah</mods:namePart>
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<mods:namePart id="E5F41D07AD2237ADDDCF3BF935A7A9E5">Xing, Jichun</mods:namePart>
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<mods:affiliation id="1DC2A58768736ED25DB94BD2FE510AD1">Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China</mods:affiliation>
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<emphasis id="B90AEA94FFF5CF731ED9FDB40C550551" bold="true" box="[151,377,541,568]" italics="true" pageId="6" pageNumber="369">Fistulatus luteolus</emphasis>
<bibRefCitation id="EFEF4B77FFF5CF731FCEFDB40F7C0551" author="Cen, Y. W. &amp; Cai, P." box="[384,592,541,568]" pageId="6" pageNumber="369" pagination="116 - 120" refId="ref5050" refString="Cen, Y. W. &amp; Cai, P. (2002) Four new species of the subfamily Selenocephalinae from China (Homoptera: Cicadellidae). Acta Zootaxanomica Sinica, 27, 116 - 120." type="journal article" year="2002">Cen &amp; Cai, 2002</bibRefCitation>
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<paragraph id="8BC13686FFF5CF731ED9FDEB0C0B0534" blockId="6.[151,592,541,605]" box="[151,295,578,605]" pageId="6" pageNumber="369">
(
<figureCitation id="13452A03FFF5CF731ED1FDEB0C330534" box="[159,287,578,605]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="6" pageNumber="369">Figs 3539</figureCitation>
)
</paragraph>
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<taxonomicName id="4C7E4D05FFF5CF731ED9FD2F0F1605F7" ID-CoL="6J6X4" authority="Cen &amp; Cai, 2002: 119" authorityName="Cen &amp; Cai" authorityPageNumber="119" authorityYear="2002" box="[151,570,646,670]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="369" phylum="Arthropoda" rank="species" species="luteolus">
<emphasis id="B90AEA94FFF5CF731ED9FD2F0C7F05F7" box="[151,339,646,670]" italics="true" pageId="6" pageNumber="369">Fistulatus luteolus</emphasis>
<bibRefCitation id="EFEF4B77FFF5CF731F17FD2F0F1605F7" author="Cen, Y. W. &amp; Cai, P." box="[345,570,646,670]" pageId="6" pageNumber="369" pagination="116 - 120" refId="ref5050" refString="Cen, Y. W. &amp; Cai, P. (2002) Four new species of the subfamily Selenocephalinae from China (Homoptera: Cicadellidae). Acta Zootaxanomica Sinica, 27, 116 - 120." type="journal article" year="2002">Cen &amp; Cai, 2002: 119</bibRefCitation>
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.
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<subSubSection id="C364650DFFF5CF731ED9FD6508B604F2" pageId="6" pageNumber="369" type="materials_examined">
<paragraph id="8BC13686FFF5CF731ED9FD6508B604F2" blockId="6.[151,1437,716,1067]" pageId="6" pageNumber="369">
<emphasis id="B90AEA94FFF5CF731ED9FD650CAD058F" bold="true" box="[151,385,716,742]" pageId="6" pageNumber="369">Material examined.</emphasis>
<materialsCitation id="3B163CDBFFF5CF731FC9FD650C870462" collectingDate="2013-07-16" collectionCode="GUGC" collectorName="Dongliang Xing &amp; Zhimin Chang" country="China" location="Shenlongjia Guanmenshan" pageId="6" pageNumber="369" specimenCount="3" specimenCount-male="3" stateProvince="Hubei">
<emphasis id="B90AEA94FFF5CF731FC9FD650CC2058F" bold="true" box="[391,494,716,742]" pageId="6" pageNumber="369">
<collectingCountry id="F3697616FFF5CF731FC9FD650CC5058F" box="[391,489,716,742]" name="China" pageId="6" pageNumber="369">CHINA</collectingCountry>
:
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<specimenCount id="9D78FD0FFFF5CF731FBBFD640F03058E" box="[501,559,717,743]" count="3" pageId="6" pageNumber="369" type="male">3♂♂</specimenCount>
,
<collectingRegion id="49BAF864FFF5CF731C75FD650FE8058E" box="[571,708,716,743]" country="China" name="Hubei" pageId="6" pageNumber="369">Hubei Prov.</collectingRegion>
,
<location id="8EA1605DFFF5CF731C80FD650ED2058F" LSID="urn:lsid:plazi:treatment:03D78790FFF5CF731ED9FDB40E3D0343:8EA1605DFFF5CF731C80FD650ED2058F" box="[718,1022,716,742]" country="China" name="Shenlongjia Guanmenshan" pageId="6" pageNumber="369" stateProvince="Hubei">Shenlongjia Guanmenshan</location>
,
<date id="FFC01046FFF5CF731A47FD6509B1058E" box="[1033,1181,716,743]" pageId="6" pageNumber="369" value="2013-07-16">
<collectingDate id="EF84E9AEFFF5CF731A47FD6509B1058E" box="[1033,1181,716,743]" pageId="6" pageNumber="369" value="2013-07-16">16 July 2013</collectingDate>
</date>
, coll.
<collectorName id="268B5350FFF5CF731AAEFD6508B4058F" box="[1248,1432,716,742]" pageId="6" pageNumber="369">Dongliang Xing</collectorName>
,
<collectorName id="268B5350FFF5CF731ED9FD590C100463" box="[151,316,752,778]" pageId="6" pageNumber="369">Zhimin Chang</collectorName>
(
<collectionCode id="ED6FAE43FFF5CF731F00FD580C880462" box="[334,420,753,779]" pageId="6" pageNumber="369">GUGC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B163CDBFFF5CF731FF7FD580F360446" collectingDate="2013-08-05" collectionCode="GUGC" collectorName="Ling Qu &amp; Yunfei Wu" country="China" location="Badashan Tianpingshan Management Station" pageId="6" pageNumber="369" specimenCount="8" specimenCount-male="8" stateProvince="Hunan">
<specimenCount id="9D78FD0FFFF5CF731FF7FD580CDF0462" box="[441,499,753,779]" count="8" pageId="6" pageNumber="369" type="male">8♂♂</specimenCount>
,
<collectingRegion id="49BAF864FFF5CF731C4FFD580FBF0462" box="[513,659,753,779]" country="China" name="Hunan" pageId="6" pageNumber="369">Hunan Prov.</collectingRegion>
,
<location id="8EA1605DFFF5CF731CEEFD5909890463" LSID="urn:lsid:plazi:treatment:03D78790FFF5CF731ED9FDB40E3D0343:8EA1605DFFF5CF731CEEFD5909890463" box="[672,1189,752,779]" country="China" name="Badashan Tianpingshan Management Station" pageId="6" pageNumber="369" stateProvince="Hunan">Badashan Tianpingshan Management Station</location>
,
<date id="FFC01046FFF5CF731AFCFD5908720462" box="[1202,1374,752,779]" pageId="6" pageNumber="369" value="2013-08-05">
<collectingDate id="EF84E9AEFFF5CF731AFCFD5908720462" box="[1202,1374,752,779]" pageId="6" pageNumber="369" value="2013-08-05">5 August 2013</collectingDate>
</date>
, coll.
<collectorName id="268B5350FFF5CF731ED9FCBD0DDB0446" box="[151,247,788,815]" pageId="6" pageNumber="369">Ling Qu</collectorName>
and
<collectorName id="268B5350FFF5CF731F7FFCBD0C870446" box="[305,427,788,815]" pageId="6" pageNumber="369">Yunfei Wu</collectorName>
(
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)
</materialsCitation>
;
<materialsCitation id="3B163CDBFFF5CF731C66FCBC0F60043A" collectingDate="2012-08-12" collectingDateMax="2012-08-17" collectingDateMin="2012-08-12" collectionCode="GUGC" collectorName="Qiongzhang Song" country="China" county="Dushan County" location="Duliu Jiangyuan Wetland" pageId="6" pageNumber="369" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou">
<specimenCount id="9D78FD0FFFF5CF731C66FCBC0F600446" box="[552,588,789,815]" count="1" pageId="6" pageNumber="369" type="male">1♂</specimenCount>
,
<collectingRegion id="49BAF864FFF5CF731C15FCBD0E2E0446" box="[603,770,788,815]" country="China" name="Guizhou" pageId="6" pageNumber="369">Guizhou Prov.</collectingRegion>
,
<collectingCounty id="62A04E0AFFF5CF731D41FCBD0EED0446" box="[783,961,788,815]" pageId="6" pageNumber="369">Dushan County</collectingCounty>
,
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,
<date id="FFC01046FFF5CF731AB0FCBC0DFE043A" pageId="6" pageNumber="369" value="2012-08-12" valueMax="2012-08-17" valueMin="2012-08-12">
<collectingDate id="EF84E9AEFFF5CF731AB0FCBC0DFE043A" pageId="6" pageNumber="369" value="2012-08-12" valueMax="2012-08-17" valueMin="2012-08-12">1217 August 2012</collectingDate>
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, coll.
<collectorName id="268B5350FFF5CF731F58FC910CCC043A" box="[278,480,824,851]" pageId="6" pageNumber="369">Qiongzhang Song</collectorName>
(
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;
<materialsCitation id="3B163CDBFFF5CF731C16FC900F8E041E" collectingDate="2010-08-10" collectingDateMax="2010-08-12" collectingDateMin="2010-08-10" collectionCode="GUGC" collectorName="Zhimin Chang &amp; Yanli Zheng" country="China" location="Heihe National Forest" pageId="6" pageNumber="369" specimenCount="15" specimenCount-female="14" specimenCount-male="1" stateProvince="Shaanxi">
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<specimenCount id="9D78FD0FFFF5CF731C35FC900FEF043A" box="[635,707,825,851]" count="14" pageId="6" pageNumber="369" type="female">14♀♀</specimenCount>
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<collectingRegion id="49BAF864FFF5CF731C81FC910E42043A" box="[719,878,824,851]" country="China" name="Shaanxi" pageId="6" pageNumber="369">Shaanxi Prov.</collectingRegion>
,
<location id="8EA1605DFFF5CF731D37FC91095D043A" LSID="urn:lsid:plazi:treatment:03D78790FFF5CF731ED9FDB40E3D0343:8EA1605DFFF5CF731D37FC91095D043A" box="[889,1137,824,851]" country="China" name="Heihe National Forest" pageId="6" pageNumber="369" stateProvince="Shaanxi">Heihe National Forest</location>
Park,
<date id="FFC01046FFF5CF731AF7FC9008B4043A" box="[1209,1432,825,851]" pageId="6" pageNumber="369" value="2010-08-10" valueMax="2010-08-12" valueMin="2010-08-10">
<collectingDate id="EF84E9AEFFF5CF731AF7FC9008B4043A" box="[1209,1432,825,851]" pageId="6" pageNumber="369" value="2010-08-10" valueMax="2010-08-12" valueMin="2010-08-10">1012 August 2010</collectingDate>
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, coll.
<collectorName id="268B5350FFF5CF731E9EFCF50C5F041F" box="[208,371,860,886]" pageId="6" pageNumber="369">Zhimin Chang</collectorName>
and
<collectorName id="268B5350FFF5CF731FE4FCF50F1A041F" box="[426,566,860,886]" pageId="6" pageNumber="369">Yanli Zheng</collectorName>
(
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)
</materialsCitation>
;
<materialsCitation id="3B163CDBFFF5CF731CE0FCF40C1004F2" collectingDate="2010-08-14" collectionCode="GUGC" collectorName="Jichun Xing" country="China" location="Kuankuoshui" pageId="6" pageNumber="369" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Guizhou">
<specimenCount id="9D78FD0FFFF5CF731CE0FCF40FFA041E" box="[686,726,861,887]" count="1" pageId="6" pageNumber="369" type="male">1♂</specimenCount>
<specimenCount id="9D78FD0FFFF5CF731C98FCF40FD9041E" box="[726,757,861,887]" count="1" pageId="6" pageNumber="369" type="female">1♀</specimenCount>
,
<collectingRegion id="49BAF864FFF5CF731D4DFCF50E85041E" box="[771,937,860,887]" country="China" name="Guizhou" pageId="6" pageNumber="369">Guizhou Prov.</collectingRegion>
,
<location id="8EA1605DFFF5CF731DFAFCF50960041F" LSID="urn:lsid:plazi:treatment:03D78790FFF5CF731ED9FDB40E3D0343:8EA1605DFFF5CF731DFAFCF50960041F" box="[948,1100,860,886]" country="China" name="Kuankuoshui" pageId="6" pageNumber="369" stateProvince="Guizhou">Kuankuoshui</location>
,
<date id="FFC01046FFF5CF731A17FCF40823041E" box="[1113,1295,861,887]" pageId="6" pageNumber="369" value="2010-08-14">
<collectingDate id="EF84E9AEFFF5CF731A17FCF40823041E" box="[1113,1295,861,887]" pageId="6" pageNumber="369" value="2010-08-14">14 August 2010</collectingDate>
</date>
, coll.
<collectorName id="268B5350FFF5CF731B1DFCF50DE304F3" pageId="6" pageNumber="369">Jichun Xing</collectorName>
(
<collectionCode id="ED6FAE43FFF5CF731E90FC280C1804F2" box="[222,308,897,923]" pageId="6" pageNumber="369">GUGC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B163CDBFFF5CF731F09FC2808BA04F2" box="[327,1430,896,923]" collectingDate="2013-07-15" collectionCode="GUGC" collectorName="Ling Qu &amp; Yunfei Wu" country="China" location="Shenlongjia Guanmenshan" pageId="6" pageNumber="369" specimenCount="1" specimenCount-male="1" stateProvince="Hubei">
<specimenCount id="9D78FD0FFFF5CF731F09FC280C4704F2" box="[327,363,897,923]" count="1" pageId="6" pageNumber="369" type="male">1♂</specimenCount>
,
<collectingRegion id="49BAF864FFF5CF731F36FC290F2D04F2" box="[376,513,896,923]" country="China" name="Hubei" pageId="6" pageNumber="369">Hubei Prov.</collectingRegion>
,
<location id="8EA1605DFFF5CF731C45FC290E1704F3" LSID="urn:lsid:plazi:treatment:03D78790FFF5CF731ED9FDB40E3D0343:8EA1605DFFF5CF731C45FC290E1704F3" box="[523,827,896,922]" country="China" name="Shenlongjia Guanmenshan" pageId="6" pageNumber="369" stateProvince="Hubei">Shenlongjia Guanmenshan</location>
,
<date id="FFC01046FFF5CF731D08FC290EF604F2" box="[838,986,896,923]" pageId="6" pageNumber="369" value="2013-07-15">
<collectingDate id="EF84E9AEFFF5CF731D08FC290EF604F2" box="[838,986,896,923]" pageId="6" pageNumber="369" value="2013-07-15">15 July 2013</collectingDate>
</date>
, coll.
<collectorName id="268B5350FFF5CF731A53FC29095704F2" box="[1053,1147,896,923]" pageId="6" pageNumber="369">Ling Qu</collectorName>
and
<collectorName id="268B5350FFF5CF731AFFFC29080504F2" box="[1201,1321,896,923]" pageId="6" pageNumber="369">Yunfei Wu</collectorName>
(
<collectionCode id="ED6FAE43FFF5CF731B76FC2808A204F2" box="[1336,1422,897,923]" pageId="6" pageNumber="369">GUGC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF5CF731E89FC0C0E9F048B" pageId="6" pageNumber="369" type="discussion">
<paragraph id="8BC13686FFF5CF731E89FC0C0E9F048B" blockId="6.[151,1437,716,1067]" pageId="6" pageNumber="369">
<emphasis id="B90AEA94FFF5CF731E89FC0C0C1704D6" bold="true" box="[199,315,933,959]" pageId="6" pageNumber="369">Remarks.</emphasis>
This species is similar to
<taxonomicName id="4C7E4D05FFF5CF731C16FC0C0E8104D7" authority="Lu &amp; Zhang" authorityName="Lu &amp; Zhang" authorityYear="2014" box="[600,941,932,959]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="369" phylum="Arthropoda" rank="species" species="quadrispinosus">
<emphasis id="B90AEA94FFF5CF731C16FC0C0E3204D7" box="[600,798,932,958]" italics="true" pageId="6" pageNumber="369">F. quadrispinosus</emphasis>
Lu &amp; Zhang
</taxonomicName>
in having two pairs of pygofer processes but can be distinguished by the absence of processes on the aedeagal shaft.
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF5CF731E89FC450E3D0343" pageId="6" pageNumber="369" type="distribution">
<paragraph id="8BC13686FFF5CF731E89FC450E3D0343" blockId="6.[151,1437,716,1067]" pageId="6" pageNumber="369">
<emphasis id="B90AEA94FFF5CF731E89FC450C4F036F" bold="true" box="[199,355,1004,1030]" pageId="6" pageNumber="369">Distribution:</emphasis>
<collectingCountry id="F3697616FFF5CF731F25FC450C9E036F" box="[363,434,1004,1030]" name="China" pageId="6" pageNumber="369">China</collectingCountry>
:
<collectingRegion id="49BAF864FFF5CF731FF0FC450F09036F" box="[446,549,1004,1030]" country="China" name="Zhejiang" pageId="6" pageNumber="369">Zhejiang</collectingRegion>
,
<collectingRegion id="49BAF864FFF5CF731C7FFC440F56036E" box="[561,634,1005,1031]" country="China" name="Henan" pageId="6" pageNumber="369">Henan</collectingRegion>
(
<bibRefCitation id="EFEF4B77FFF5CF731CC4FC450E74036E" author="Zhang, Y. L. &amp; Zhang, W. Z. &amp; Chen, B." box="[650,856,1004,1031]" pageId="6" pageNumber="369" pagination="235 - 245" refId="ref5993" refString="Zhang, Y. L., Zhang, W. Z. &amp; Chen, B. (1997) Selenocephaline leafhoppers (Homoptera: Cicadellidae) from Mt. Funiushan in Henan Province. Entomotaxonomia, 19, 235 - 245." type="journal article" year="1997">
Zhang
<emphasis id="B90AEA94FFF5CF731C95FC440E39036F" box="[731,789,1004,1030]" italics="true" pageId="6" pageNumber="369">et al.</emphasis>
1997
</bibRefCitation>
;
<bibRefCitation id="EFEF4B77FFF5CF731D2BFC440907036E" author="Cen, Y. W. &amp; Cai, P." box="[869,1067,1004,1031]" pageId="6" pageNumber="369" pagination="116 - 120" refId="ref5050" refString="Cen, Y. W. &amp; Cai, P. (2002) Four new species of the subfamily Selenocephalinae from China (Homoptera: Cicadellidae). Acta Zootaxanomica Sinica, 27, 116 - 120." type="journal article" year="2002">Cen &amp; Cai, 2002</bibRefCitation>
;
<bibRefCitation id="EFEF4B77FFF5CF731A79FC45082F036F" author="Shang, S. Q. &amp; Shen L. &amp; Zhang, Y. L. &amp; Li, H. H." box="[1079,1283,1004,1030]" pageId="6" pageNumber="369" pagination="152 - 154" refId="ref5632" refString="Shang, S. Q., Shen L., Zhang, Y. L. &amp; Li, H. H. (2006) Taxonomy study on the leafhopper genus Fistulatus Zhang, with description of a new species from China (Hemiptera, Selenocephalinae, Paraboloponini). Acta Zootaxonomica Sinica, 31, 152 - 154." type="journal article" year="2006">
Shang
<emphasis id="B90AEA94FFF5CF731ACBFC440993036F" box="[1157,1215,1004,1030]" italics="true" pageId="6" pageNumber="369">et al.</emphasis>
2006
</bibRefCitation>
;
<bibRefCitation id="EFEF4B77FFF5CF731B41FC440DFF0342" author="Lu, L. &amp; Zhang, Y. L." pageId="6" pageNumber="369" pagination="247 - 250" refId="ref5506" refString="Lu, L. &amp; Zhang, Y. L. (2014) Taxonomy of the Oriental leafhopper genus Fistulatus (Hemiptera: Cicadellidae: Deltocephalinae), with description of a new species from China. Zootaxa, 3838 (2), 247 - 250. https: // doi. org / 10.11646 / zootaxa. 3838.2.10" type="journal article" year="2014">Lu &amp; Zhang 2014</bibRefCitation>
;
<bibRefCitation id="EFEF4B77FFF5CF731E90FBB80CAE0342" author="Qu, L. &amp; Webb, M. D. &amp; Dai, R. H." box="[222,386,1040,1067]" pageId="6" pageNumber="369" pagination="260 - 270" refId="ref5384" refString="Qu, L., Webb, M. D. &amp; Dai, R. H. (2015) A new genus and species of the leafhopper subtribe Paraboloponina from China (Hemiptera, Cicadellidae). Zootaxa, 3919 (2), 260 - 270. https: // doi. org / 10.11646 / zootaxa. 3919.2.2" type="journal article" year="2015">
Qu
<emphasis id="B90AEA94FFF5CF731F49FBB80C130343" box="[263,319,1040,1066]" italics="true" pageId="6" pageNumber="369">et al.</emphasis>
2015
</bibRefCitation>
),
<collectingRegion id="49BAF864FFF5CF731FDBFBB90CF00343" box="[405,476,1040,1066]" country="China" name="Hubei" pageId="6" pageNumber="369">Hubei</collectingRegion>
,
<collectingRegion id="49BAF864FFF5CF731FA6FBB80F190342" box="[488,565,1041,1067]" country="China" name="Hunan" pageId="6" pageNumber="369">Hunan</collectingRegion>
,
<collectingRegion id="49BAF864FFF5CF731C0FFBB90F880343" box="[577,676,1040,1066]" country="China" name="Guizhou" pageId="6" pageNumber="369">Guizhou</collectingRegion>
,
<collectingRegion id="49BAF864FFF5CF731CE1FBB90E210343" box="[687,781,1040,1066]" country="China" name="Shaanxi" pageId="6" pageNumber="369">Shaanxi</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,143 @@
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<mods:title id="7CA57CE5A3F0D0E5CF494EDDC9393877">Key to species of the leafhopper genus Fistulatus Zhang, Zhang &amp; Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species</mods:title>
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<mods:affiliation id="E2DA0704F47C1D37B75518F95CE31792">Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A &amp; F University, 666 Wusu Street, Linan, Hangzhou, Zhejiang 311300, P. R. China</mods:affiliation>
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<heading id="D08981EAFFF5CF731ED9FF3E0F9707D8" bold="true" box="[151,699,151,178]" fontSize="11" level="1" pageId="6" pageNumber="369" reason="1">
<taxonomicName id="4C7E4D05FFF5CF731ED9FF3E0F9707D8" ID-CoL="CCMWX" authority="Yang &amp; Dai, 2024" authorityName="Yang &amp; Dai" authorityYear="2024" box="[151,699,151,178]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="369" phylum="Arthropoda" rank="species" species="brevimarginalis">
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<emphasis id="B90AEA94FFF5CF731ED9FF3E0CF407DB" bold="true" box="[151,472,151,178]" italics="true" pageId="6" pageNumber="369">Fistulatus brevimarginalis</emphasis>
<bibRefCitation id="EFEF4B77FFF5CF731F91FF3E0F9707D8" author="Yang, Y. Q. &amp; Dai, R. H. &amp; You, S. Y." box="[479,699,151,177]" pageId="6" pageNumber="369" pagination="143 - 150" refId="ref5740" refString="Yang, Y. Q., Dai, R. H. &amp; You, S. Y. (2024) Key to species of the genus Fistulatus Zhang, 1997 (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of two new species from China. Zootaxa, Zootaxa, 5446 (1), 143 - 150. https: // doi. org / 10.11646 / zootaxa. 5446.1.9" type="journal article" year="2024">Yang &amp; Dai, 2024</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="C364650DFFF5CF731ED9FF120C0B07BC" box="[151,295,187,213]" pageId="6" pageNumber="369" type="description">
<paragraph id="8BC13686FFF5CF731ED9FF120C0B07BC" blockId="6.[151,699,151,213]" box="[151,295,187,213]" pageId="6" pageNumber="369">
(
<figureCitation id="13452A03FFF5CF731ED1FF120C3307BC" box="[159,287,187,213]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="6" pageNumber="369">Figs 3034</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF5CF731ED9FEA909890671" box="[151,1189,256,280]" pageId="6" pageNumber="369" type="reference_group">
<paragraph id="8BC13686FFF5CF731ED9FEA909890671" blockId="6.[151,1189,256,280]" box="[151,1189,256,280]" pageId="6" pageNumber="369">
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<emphasis id="B90AEA94FFF5CF731ED9FEA90C8F0671" box="[151,419,256,280]" italics="true" pageId="6" pageNumber="369">Fistulatus brevimarginalis</emphasis>
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<bibRefCitation id="EFEF4B77FFF5CF731FE7FEA90FBB0671" author="Yang, Y. Q. &amp; Dai, R. H. &amp; You, S. Y." box="[425,663,256,280]" pageId="6" pageNumber="369" pagination="143 - 150" refId="ref5740" refString="Yang, Y. Q., Dai, R. H. &amp; You, S. Y. (2024) Key to species of the genus Fistulatus Zhang, 1997 (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of two new species from China. Zootaxa, Zootaxa, 5446 (1), 143 - 150. https: // doi. org / 10.11646 / zootaxa. 5446.1.9" type="journal article" year="2024">Yang &amp; Dai, 2024: 117</bibRefCitation>
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.
<typeStatus id="54C58824FFF5CF731CEEFEA90FFE0671" box="[672,722,256,280]" pageId="6" pageNumber="369">Type</typeStatus>
locality:
<collectingCountry id="F3697616FFF5CF731D7CFEA90E430671" box="[818,879,256,280]" name="China" pageId="6" pageNumber="369">China</collectingCountry>
(
<collectingRegion id="49BAF864FFF5CF731D32FEA90EE30671" box="[892,975,256,280]" country="China" name="Yunnan" pageId="6" pageNumber="369">Yunnan</collectingRegion>
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<subSubSection id="C364650DFFF5CF731ED9FEEF0E7D06C1" pageId="6" pageNumber="369" type="discussion">
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<emphasis id="B90AEA94FFF5CF731ED9FEEF0C270609" bold="true" box="[151,267,326,352]" pageId="6" pageNumber="369">Remarks.</emphasis>
This species is similar to
<taxonomicName id="4C7E4D05FFF5CF731C78FEEE0EBB0609" authority="Yang &amp; Dai" authorityName="Yang &amp; Dai" authorityYear="2024" box="[566,919,326,352]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="369" phylum="Arthropoda" rank="species" species="brevimarginalis">
<emphasis id="B90AEA94FFF5CF731C78FEEE0E2F0609" box="[566,771,326,352]" italics="true" pageId="6" pageNumber="369">F. brevimarginalis</emphasis>
Yang &amp; Dai
</taxonomicName>
,
<taxonomicName id="4C7E4D05FFF5CF731DEDFEEE09B70609" authority="Cen &amp; Cai" authorityName="Cen &amp; Cai" authorityYear="2002" box="[931,1179,326,352]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="369" phylum="Arthropoda" rank="species" species="luteolus">
<emphasis id="B90AEA94FFF5CF731DEDFEEE09350609" box="[931,1049,326,352]" italics="true" pageId="6" pageNumber="369">F. luteolus</emphasis>
Cen &amp; Cai
</taxonomicName>
and
<taxonomicName id="4C7E4D05FFF5CF731A9AFEEE0C0D06ED" authority="Lu &amp; Zhang" authorityName="Lu &amp; Zhang" authorityYear="2014" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="369" phylum="Arthropoda" rank="species" species="quadrispinosus">
<emphasis id="B90AEA94FFF5CF731A9AFEEE08B00609" box="[1236,1436,326,352]" italics="true" pageId="6" pageNumber="369">F. quadrispinosus</emphasis>
Lu &amp; Zhang
</taxonomicName>
in having the pygofer with two pairs of processes but can be distinguished by the aedeagal shaft with a single dorsal preapical process and no paired apical processes.
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF5CF731E89FE180FC506A5" box="[199,745,433,460]" pageId="6" pageNumber="369" type="distribution">
<paragraph id="8BC13686FFF5CF731E89FE180FC506A5" blockId="6.[151,1436,326,460]" box="[199,745,433,460]" pageId="6" pageNumber="369">
<emphasis id="B90AEA94FFF5CF731E89FE180C4F06A2" bold="true" box="[199,355,433,459]" pageId="6" pageNumber="369">Distribution:</emphasis>
<collectingCountry id="F3697616FFF5CF731F24FE1B0C9D06A5" box="[362,433,434,460]" name="China" pageId="6" pageNumber="369">China</collectingCountry>
:
<collectingRegion id="49BAF864FFF5CF731FF4FE1B0F3C06A5" box="[442,528,434,460]" country="China" name="Yunnan" pageId="6" pageNumber="369">Yunnan</collectingRegion>
(
<bibRefCitation id="EFEF4B77FFF5CF731C50FE1B0FF106A5" author="Yang, Y. Q. &amp; Dai, R. H. &amp; You, S. Y." box="[542,733,434,460]" pageId="6" pageNumber="369" pagination="143 - 150" refId="ref5740" refString="Yang, Y. Q., Dai, R. H. &amp; You, S. Y. (2024) Key to species of the genus Fistulatus Zhang, 1997 (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of two new species from China. Zootaxa, Zootaxa, 5446 (1), 143 - 150. https: // doi. org / 10.11646 / zootaxa. 5446.1.9" type="journal article" year="2024">
Yang
<emphasis id="B90AEA94FFF5CF731C2FFE1A0FB506A5" box="[609,665,434,460]" italics="true" pageId="6" pageNumber="369">et al.</emphasis>
2024
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,155 @@
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<mods:title id="FB96DDDEE304A6EE54E21DBAA2B24305">Key to species of the leafhopper genus Fistulatus Zhang, Zhang &amp; Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species</mods:title>
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<mods:namePart id="E5D748AF7DFFE3A9027CABB3A5CC9D01">Hassan, Muhammad Asghar</mods:namePart>
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<mods:affiliation id="66D82C68CFA8AB787C321DA026A90DA3">Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China</mods:affiliation>
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<mods:namePart id="B5B1D200C263CEE1621A8D181821F25D">Shah, Bismillah</mods:namePart>
<mods:nameIdentifier id="BF1C7AD4A8DAE8B5865EEB7A2515CA84" type="ORCID">0000-0002-8407-8627</mods:nameIdentifier>
<mods:affiliation id="AD524B89FC0F92A975EA10840B18F32D">Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A &amp; F University, 666 Wusu Street, Linan, Hangzhou, Zhejiang 311300, P. R. China</mods:affiliation>
<mods:nameIdentifier id="17398DCB2338AEE54CF7F173C641D949" type="email">bismillahshah1990@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="2E5AAEFBF8797AB76380345B5AED8F4B">Xing, Jichun</mods:namePart>
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<mods:affiliation id="66C21FB2CEB4484866E19D428A7C0210">Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China</mods:affiliation>
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<treatment id="03D78790FFF7CF711ED9F93E0FD90098" ID-DOI="http://doi.org/10.5281/zenodo.12758877" ID-Zenodo-Dep="12758877" LSID="urn:lsid:plazi:treatment:03D78790FFF7CF711ED9F93E0FD90098" httpUri="http://treatment.plazi.org/id/03D78790FFF7CF711ED9F93E0FD90098" lastPageNumber="367" pageId="4" pageNumber="367">
<subSubSection id="C364650DFFF7CF711ED9F93E0FA301DB" box="[151,655,1687,1714]" pageId="4" pageNumber="367" type="nomenclature">
<paragraph id="8BC13686FFF7CF711ED9F93E0FA301DB" blockId="4.[151,655,1687,1751]" box="[151,655,1687,1714]" pageId="4" pageNumber="367">
<heading id="D08981EAFFF7CF711ED9F93E0FA301DB" bold="true" box="[151,655,1687,1714]" fontSize="11" level="1" pageId="4" pageNumber="367" reason="1">
<taxonomicName id="4C7E4D05FFF7CF711ED9F93E0FA301DB" ID-CoL="CCMX6" authority="Yang &amp; Dai, 2024" authorityName="Yang &amp; Dai" authorityYear="2024" box="[151,655,1687,1714]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="biconjugara">
<emphasis id="B90AEA94FFF7CF711ED9F93E0FA301DB" bold="true" box="[151,655,1687,1714]" pageId="4" pageNumber="367">
<emphasis id="B90AEA94FFF7CF711ED9F93E0C8001DB" bold="true" box="[151,428,1687,1714]" italics="true" pageId="4" pageNumber="367">Fistulatus biconjugara</emphasis>
<bibRefCitation id="EFEF4B77FFF7CF711FFDF9310FA301DB" author="Yang, Y. Q. &amp; Dai, R. H. &amp; You, S. Y." box="[435,655,1688,1714]" pageId="4" pageNumber="367" pagination="143 - 150" refId="ref5740" refString="Yang, Y. Q., Dai, R. H. &amp; You, S. Y. (2024) Key to species of the genus Fistulatus Zhang, 1997 (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of two new species from China. Zootaxa, Zootaxa, 5446 (1), 143 - 150. https: // doi. org / 10.11646 / zootaxa. 5446.1.9" type="journal article" year="2024">Yang &amp; Dai, 2024</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF7CF711ED9F9150C0B01BE" box="[151,295,1724,1751]" pageId="4" pageNumber="367" type="description">
<paragraph id="8BC13686FFF7CF711ED9F9150C0B01BE" blockId="4.[151,655,1687,1751]" box="[151,295,1724,1751]" pageId="4" pageNumber="367">
(
<figureCitation id="13452A03FFF7CF711ED1F9150C3301BE" box="[159,287,1724,1751]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="4" pageNumber="367">Figs 2529</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF7CF711ED9F8A909A80071" box="[151,1156,1792,1817]" pageId="4" pageNumber="367" type="reference_group">
<paragraph id="8BC13686FFF7CF711ED9F8A909A80071" blockId="4.[151,1156,1792,1817]" box="[151,1156,1792,1817]" pageId="4" pageNumber="367">
<treatmentCitationGroup id="AB6E11A8FFF7CF711ED9F8A909A80071" box="[151,1156,1792,1817]" pageId="4" pageNumber="367">
<treatmentCitation id="0ADF1097FFF7CF711ED9F8A90F5F0070" ID-DOI="http://doi.org/10.5281/zenodo.11084752" ID-Zenodo-Dep="11084752" author="Yang, Y. Q. &amp; Dai, R. H. &amp; You, S. Y." box="[151,627,1792,1817]" httpUri="http://treatment.plazi.org/id/137887D9FFC3C039FF3E04B920245482" page="117" pageId="4" pageNumber="367" year="2024">
<taxonomicName id="4C7E4D05FFF7CF711ED9F8A90F5F0070" ID-CoL="CCMX6" authority="Yang &amp; Dai, 2024: 117" authorityName="Yang &amp; Dai" authorityPageNumber="117" authorityYear="2024" box="[151,627,1792,1817]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="biconjugara">
<emphasis id="B90AEA94FFF7CF711ED9F8A90C530071" box="[151,383,1792,1816]" italics="true" pageId="4" pageNumber="367">Fistulatus biconjugara</emphasis>
<materialsCitation id="3B163CDBFFF7CF711FCBF8A80F5F0070" box="[389,627,1792,1817]" country="China" location="Fanjing Mountain" pageId="4" pageNumber="367" specimenCount="1" stateProvince="Guizhou">
<bibRefCitation id="EFEF4B77FFF7CF711FCBF8A80F5F0070" author="Yang, Y. Q. &amp; Dai, R. H. &amp; You, S. Y." box="[389,627,1792,1817]" pageId="4" pageNumber="367" pagination="143 - 150" refId="ref5740" refString="Yang, Y. Q., Dai, R. H. &amp; You, S. Y. (2024) Key to species of the genus Fistulatus Zhang, 1997 (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of two new species from China. Zootaxa, Zootaxa, 5446 (1), 143 - 150. https: // doi. org / 10.11646 / zootaxa. 5446.1.9" type="journal article" year="2024">Yang &amp; Dai, 2024: 117</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54C58824FFF7CF711C32F8A80F820070" box="[636,686,1793,1817]" pageId="4" pageNumber="367">Type</typeStatus>
locality:
<collectingCountry id="F3697616FFF7CF711D40F8A90E670071" box="[782,843,1792,1816]" name="China" pageId="4" pageNumber="367">China</collectingCountry>
(
<collectingRegion id="49BAF864FFF7CF711D16F8A90E990071" box="[856,949,1792,1816]" country="China" name="Guizhou" pageId="4" pageNumber="367">Guizhou</collectingRegion>
:
<location id="8EA1605DFFF7CF711DF0F8A909550071" LSID="urn:lsid:plazi:treatment:03D78790FFF7CF711ED9F93E0FD90098:8EA1605DFFF7CF711DF0F8A909550071" box="[958,1145,1792,1816]" country="China" name="Fanjing Mountain" pageId="4" pageNumber="367" stateProvince="Guizhou">Fanjing Mountain</location>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF7CF711ED9F8EE0F1700A4" pageId="4" pageNumber="367" type="discussion">
<paragraph id="8BC13686FFF7CF711ED9F8EE0F1700A4" blockId="4.[151,1436,1863,2033]" pageId="4" pageNumber="367">
<emphasis id="B90AEA94FFF7CF711ED9F8EE0C270008" bold="true" box="[151,267,1863,1889]" pageId="4" pageNumber="367">Remarks.</emphasis>
This species is similar to
<taxonomicName id="4C7E4D05FFF7CF711C78F8EE0EBB0008" authority="Yang &amp; Dai" authorityName="Yang &amp; Dai" authorityYear="2024" box="[566,919,1863,1889]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="brevimarginalis">
<emphasis id="B90AEA94FFF7CF711C78F8EE0E2F0008" box="[566,771,1863,1889]" italics="true" pageId="4" pageNumber="367">F. brevimarginalis</emphasis>
Yang &amp; Dai
</taxonomicName>
,
<taxonomicName id="4C7E4D05FFF7CF711DEDF8EE09B70008" authority="Cen &amp; Cai" authorityName="Cen &amp; Cai" authorityYear="2002" box="[931,1179,1863,1889]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="luteolus">
<emphasis id="B90AEA94FFF7CF711DEDF8EE09350008" box="[931,1049,1863,1889]" italics="true" pageId="4" pageNumber="367">F. luteolus</emphasis>
Cen &amp; Cai
</taxonomicName>
and
<taxonomicName id="4C7E4D05FFF7CF711A9AF8EE0C0F00EC" authority="Lu &amp; Zhang" authorityName="Lu &amp; Zhang" authorityYear="2014" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="quadrispinosus">
<emphasis id="B90AEA94FFF7CF711A9AF8EE08B00008" box="[1236,1436,1863,1889]" italics="true" pageId="4" pageNumber="367">F. quadrispinosus</emphasis>
Lu &amp; Zhang
</taxonomicName>
in having the pygofer with two pairs of processes but can be distinguished by the aedeagal shaft with only small distal processes (aedeagal shaft with two pairs of processes in
<taxonomicName id="4C7E4D05FFF7CF711D9BF82609B000C0" authorityName="Lu &amp; Zhang" authorityYear="2014" box="[981,1180,1935,1961]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="quadrispinosus">
<emphasis id="B90AEA94FFF7CF711D9BF82609B000C0" box="[981,1180,1935,1961]" italics="true" pageId="4" pageNumber="367">F. quadrispinosus</emphasis>
</taxonomicName>
and without processes in
<taxonomicName id="4C7E4D05FFF7CF711EFAF81A0CAC00A4" authorityName="Yang &amp; Dai" authorityYear="2024" box="[180,384,1971,1997]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="brevimarginalis">
<emphasis id="B90AEA94FFF7CF711EFAF81A0CAC00A4" box="[180,384,1971,1997]" italics="true" pageId="4" pageNumber="367">F. brevimarginalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7E4D05FFF7CF711FF9F81A0F0700A4" authorityName="Cen &amp; Cai" authorityYear="2002" box="[439,555,1971,1997]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="luteolus">
<emphasis id="B90AEA94FFF7CF711FF9F81A0F0700A4" box="[439,555,1971,1997]" italics="true" pageId="4" pageNumber="367">F. luteolus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF7CF711E89F87F0FD90098" box="[199,757,2006,2033]" pageId="4" pageNumber="367" type="distribution">
<paragraph id="8BC13686FFF7CF711E89F87F0FD90098" blockId="4.[151,1436,1863,2033]" box="[199,757,2006,2033]" pageId="4" pageNumber="367">
<emphasis id="B90AEA94FFF7CF711E89F87F0C4F0099" bold="true" box="[199,355,2006,2032]" pageId="4" pageNumber="367">Distribution:</emphasis>
<collectingCountry id="F3697616FFF7CF711F24F87E0C9D0098" box="[362,433,2007,2033]" name="China" pageId="4" pageNumber="367">China</collectingCountry>
:
<collectingRegion id="49BAF864FFF7CF711FF5F87E0F370098" box="[443,539,2007,2033]" country="China" name="Guizhou" pageId="4" pageNumber="367">Guizhou</collectingRegion>
(
<bibRefCitation id="EFEF4B77FFF7CF711C64F87E0FC50098" author="Yang, Y. Q. &amp; Dai, R. H. &amp; You, S. Y." box="[554,745,2007,2033]" pageId="4" pageNumber="367" pagination="143 - 150" refId="ref5740" refString="Yang, Y. Q., Dai, R. H. &amp; You, S. Y. (2024) Key to species of the genus Fistulatus Zhang, 1997 (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of two new species from China. Zootaxa, Zootaxa, 5446 (1), 143 - 150. https: // doi. org / 10.11646 / zootaxa. 5446.1.9" type="journal article" year="2024">
Yang
<emphasis id="B90AEA94FFF7CF711C23F87E0F890098" box="[621,677,2007,2033]" italics="true" pageId="4" pageNumber="367">et al.</emphasis>
2024
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,223 @@
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<mods:title id="E551E2277EECD656C8F88C026227E495">Key to species of the leafhopper genus Fistulatus Zhang, Zhang &amp; Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species</mods:title>
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<mods:namePart id="F7A85AA6C38F983D2F838B6B93DC2682">Shah, Bismillah</mods:namePart>
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<mods:affiliation id="A49F9A603AAD1F2466EE328ECCF3D803">Department of Forestry Protection, School of Forestry and Biotechnology, Zhejiang A &amp; F University, 666 Wusu Street, Linan, Hangzhou, Zhejiang 311300, P. R. China</mods:affiliation>
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<mods:affiliation id="8AB212BC38C49A926E38FD48680C6B4F">Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China</mods:affiliation>
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<taxonomicName id="4C7E4D05FFF7CF711ED9FB280F4103F2" ID-CoL="6J6WS" authority="Cen &amp; Cai, 2002" authorityName="Cen &amp; Cai" authorityYear="2002" box="[151,621,1153,1180]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="bidentatus">
<emphasis id="B90AEA94FFF7CF711ED9FB280F4103F2" bold="true" box="[151,621,1153,1180]" pageId="4" pageNumber="367">
<emphasis id="B90AEA94FFF7CF711ED9FB280CBB03F5" bold="true" box="[151,407,1153,1180]" italics="true" pageId="4" pageNumber="367">Fistulatus bidentatus</emphasis>
<bibRefCitation id="EFEF4B77FFF7CF711FD0FB280F4103F2" author="Cen, Y. W. &amp; Cai, P." box="[414,621,1153,1179]" pageId="4" pageNumber="367" pagination="116 - 120" refId="ref5050" refString="Cen, Y. W. &amp; Cai, P. (2002) Four new species of the subfamily Selenocephalinae from China (Homoptera: Cicadellidae). Acta Zootaxanomica Sinica, 27, 116 - 120." type="journal article" year="2002">Cen &amp; Cai, 2002</bibRefCitation>
</emphasis>
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<paragraph id="8BC13686FFF7CF711ED9FB0C0C0B03D6" blockId="4.[151,621,1153,1215]" box="[151,295,1189,1215]" pageId="4" pageNumber="367">
(
<figureCitation id="13452A03FFF7CF711ED1FB0C0C3303D6" box="[159,287,1189,1215]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="4" pageNumber="367">Figs 2024</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8BC13686FFF7CF711ED9FB430901026B" blockId="4.[151,1069,1258,1282]" box="[151,1069,1258,1282]" pageId="4" pageNumber="367">
<treatmentCitationGroup id="AB6E11A8FFF7CF711ED9FB430901026B" box="[151,1069,1258,1282]" pageId="4" pageNumber="367">
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<emphasis id="B90AEA94FFF7CF711ED9FB430C41026B" box="[151,365,1258,1282]" italics="true" pageId="4" pageNumber="367">Fistulatus bidentatus</emphasis>
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<bibRefCitation id="EFEF4B77FFF7CF711F3DFB430F78026B" author="Cen, Y. W. &amp; Cai, P." box="[371,596,1258,1282]" pageId="4" pageNumber="367" pagination="116 - 120" refId="ref5050" refString="Cen, Y. W. &amp; Cai, P. (2002) Four new species of the subfamily Selenocephalinae from China (Homoptera: Cicadellidae). Acta Zootaxanomica Sinica, 27, 116 - 120." type="journal article" year="2002">Cen &amp; Cai, 2002: 117</bibRefCitation>
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.
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locality:
<collectingCountry id="F3697616FFF7CF711CA1FB430E00026B" box="[751,812,1258,1282]" name="China" pageId="4" pageNumber="367">China</collectingCountry>
(
<collectingRegion id="49BAF864FFF7CF711D77FB430EB5026B" box="[825,921,1258,1282]" country="China" name="Zhejiang" pageId="4" pageNumber="367">Zhejiang</collectingRegion>
:
<location id="8EA1605DFFF7CF711DEFFB43090E026B" LSID="urn:lsid:plazi:treatment:03D78790FFF7CF711ED9FB280CD1012F:8EA1605DFFF7CF711DEFFB43090E026B" box="[929,1058,1258,1282]" country="China" name="Tianmushan" pageId="4" pageNumber="367" stateProvince="Zhejiang">Tianmushan</location>
).
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</paragraph>
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<subSubSection id="C364650DFFF7CF711ED9FA99092202FB" pageId="4" pageNumber="367" type="materials_examined">
<paragraph id="8BC13686FFF7CF711ED9FA99092202FB" blockId="4.[151,1437,1328,1607]" pageId="4" pageNumber="367">
<emphasis id="B90AEA94FFF7CF711ED9FA990C530223" bold="true" box="[151,383,1328,1354]" pageId="4" pageNumber="367">Material examined.</emphasis>
<materialsCitation id="3B163CDBFFF7CF711FCAFA9908B40223" box="[388,1432,1328,1355]" collectingDate="2012-05-15" collectionCode="GUGC" collectorName="Hu Li" country="China" location="Damingshan" municipality="Autonomous Region" pageId="4" pageNumber="367" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi">
<emphasis id="B90AEA94FFF7CF711FCAFA990CC70223" bold="true" box="[388,491,1328,1354]" pageId="4" pageNumber="367">
<collectingCountry id="F3697616FFF7CF711FCAFA990CCA0223" box="[388,486,1328,1354]" name="China" pageId="4" pageNumber="367">CHINA</collectingCountry>
:
</emphasis>
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,
<collectingRegion id="49BAF864FFF7CF711C6EFA990FAC0223" box="[544,640,1328,1354]" country="China" name="Guangxi" pageId="4" pageNumber="367">Guangxi</collectingRegion>
<collectingMunicipality id="6BA5ACFCFFF7CF711CCAFA990E5D0223" box="[644,881,1328,1354]" pageId="4" pageNumber="367">Autonomous Region</collectingMunicipality>
.,
<location id="8EA1605DFFF7CF711D30FA9909220223" LSID="urn:lsid:plazi:treatment:03D78790FFF7CF711ED9FB280CD1012F:8EA1605DFFF7CF711D30FA9909220223" box="[894,1038,1328,1354]" country="China" municipality="Autonomous Region" name="Damingshan" pageId="4" pageNumber="367" stateProvince="Guangxi">Damingshan</location>
,
<date id="FFC01046FFF7CF711A56FA9909810223" box="[1048,1197,1328,1355]" pageId="4" pageNumber="367" value="2012-05-15">
<collectingDate id="EF84E9AEFFF7CF711A56FA9909810223" box="[1048,1197,1328,1355]" pageId="4" pageNumber="367" value="2012-05-15">15 May 2012</collectingDate>
</date>
, coll.
<collectorName id="268B5350FFF7CF711AA3FA9808010223" box="[1261,1325,1328,1355]" pageId="4" pageNumber="367">Hu Li</collectorName>
(
<collectionCode id="ED6FAE43FFF7CF711B74FA9908BC0223" box="[1338,1424,1328,1354]" pageId="4" pageNumber="367">GUGC</collectionCode>
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</materialsCitation>
;
<materialsCitation id="3B163CDBFFF7CF711ED9FAFD09D60207" box="[151,1274,1364,1391]" collectingDate="2011-07-04" collectingDateMax="2011-07-05" collectingDateMin="2011-07-04" collectionCode="GUGC" collectorName="Jiankun Long &amp; Weibin Zheng" country="China" location="Leigongshan" pageId="4" pageNumber="367" specimenCount="3" specimenCount-male="3" stateProvince="Guizhou">
<specimenCount id="9D78FD0FFFF7CF711ED9FAFD0DFE0207" box="[151,210,1364,1390]" count="3" pageId="4" pageNumber="367" type="male">3♂♂</specimenCount>
,
<collectingRegion id="49BAF864FFF7CF711E90FAFD0CA90206" box="[222,389,1364,1391]" country="China" name="Guizhou" pageId="4" pageNumber="367">Guizhou Prov.</collectingRegion>
,
<location id="8EA1605DFFF7CF711FDFFAFD0F090207" LSID="urn:lsid:plazi:treatment:03D78790FFF7CF711ED9FB280CD1012F:8EA1605DFFF7CF711FDFFAFD0F090207" box="[401,549,1364,1390]" country="China" name="Leigongshan" pageId="4" pageNumber="367" stateProvince="Guizhou">Leigongshan</location>
,
<date id="FFC01046FFF7CF711C7EFAFD0FF80207" box="[560,724,1364,1391]" pageId="4" pageNumber="367" value="2011-07-04" valueMax="2011-07-05" valueMin="2011-07-04">
<collectingDate id="EF84E9AEFFF7CF711C7EFAFD0FF80207" box="[560,724,1364,1391]" pageId="4" pageNumber="367" value="2011-07-04" valueMax="2011-07-05" valueMin="2011-07-04">45 July 2011</collectingDate>
</date>
, coll.
<collectorName id="268B5350FFF7CF711D57FAFD0E980206" box="[793,948,1364,1391]" pageId="4" pageNumber="367">Jiankun Long</collectorName>
and
<collectorName id="268B5350FFF7CF711DA2FAFD09A10207" box="[1004,1165,1364,1390]" pageId="4" pageNumber="367">Weibin Zheng</collectorName>
(
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)
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;
<materialsCitation id="3B163CDBFFF7CF711B48FAFD092602FB" collectingDate="2009-07-20" collectionCode="GUGC" collectorName="Zehong Meng &amp; Yong Chen" country="China" location="Tianmushan" pageId="4" pageNumber="367" specimenCount="1" specimenCount-male="1" stateProvince="Zhejiang">
<specimenCount id="9D78FD0FFFF7CF711B48FAFD08070207" box="[1286,1323,1364,1390]" count="1" pageId="4" pageNumber="367" type="male">1♂</specimenCount>
,
<collectingRegion id="49BAF864FFF7CF711B77FAFD0DF902FA" country="China" name="Zhejiang" pageId="4" pageNumber="367">Zhejiang Prov.</collectingRegion>
,
<location id="8EA1605DFFF7CF711E91FAD10C4702FB" LSID="urn:lsid:plazi:treatment:03D78790FFF7CF711ED9FB280CD1012F:8EA1605DFFF7CF711E91FAD10C4702FB" box="[223,363,1400,1426]" country="China" name="Tianmushan" pageId="4" pageNumber="367" stateProvince="Zhejiang">Tianmushan</location>
,
<date id="FFC01046FFF7CF711F38FAD10F2602FB" box="[374,522,1400,1426]" pageId="4" pageNumber="367" value="2009-07-20">
<collectingDate id="EF84E9AEFFF7CF711F38FAD10F2602FB" box="[374,522,1400,1426]" pageId="4" pageNumber="367" value="2009-07-20">20 July 2009</collectingDate>
</date>
, coll.
<collectorName id="268B5350FFF7CF711C03FAD10FC602FA" box="[589,746,1400,1427]" pageId="4" pageNumber="367">Zehong Meng</collectorName>
and
<collectorName id="268B5350FFF7CF711D6EFAD00EB102FB" box="[800,925,1400,1427]" pageId="4" pageNumber="367">Yong Chen</collectorName>
(
<collectionCode id="ED6FAE43FFF7CF711DE3FAD1092F02FB" box="[941,1027,1400,1426]" pageId="4" pageNumber="367">GUGC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF7CF711E89FA350F190297" pageId="4" pageNumber="367" type="discussion">
<paragraph id="8BC13686FFF7CF711E89FA350F190297" blockId="4.[151,1437,1328,1607]" pageId="4" pageNumber="367">
<emphasis id="B90AEA94FFF7CF711E89FA350C1702DF" bold="true" box="[199,315,1436,1462]" pageId="4" pageNumber="367">Remarks.</emphasis>
This species resembles
<taxonomicName id="4C7E4D05FFF7CF711C12FA340E4D02DF" authority="Cen &amp; Cai" authorityName="Cen &amp; Cai" authorityYear="2002" box="[604,865,1436,1462]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="luteolus">
<emphasis id="B90AEA94FFF7CF711C12FA340FF902DF" box="[604,725,1436,1462]" italics="true" pageId="4" pageNumber="367">F. luteolus</emphasis>
Cen &amp; Cai
</taxonomicName>
and
<taxonomicName id="4C7E4D05FFF7CF711DEFFA34082002DF" authority="Lu &amp; Zhang" authorityName="Lu &amp; Zhang" authorityYear="2014" box="[929,1292,1436,1463]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="quadrispinosus">
<emphasis id="B90AEA94FFF7CF711DEFFA34094002DF" box="[929,1132,1436,1462]" italics="true" pageId="4" pageNumber="367">F. quadrispinosus</emphasis>
Lu &amp; Zhang
</taxonomicName>
in having a distinctive elongated aedeagal shaft with short processes near the midlength of the shaft but differs in having a pair of short distal processes on the shaft.
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF7CF711E89F9A10CD1012F" pageId="4" pageNumber="367" type="distribution">
<paragraph id="8BC13686FFF7CF711E89F9A10CD1012F" blockId="4.[151,1437,1328,1607]" pageId="4" pageNumber="367">
<emphasis id="B90AEA94FFF7CF711E89F9A10C4F014B" bold="true" box="[199,355,1544,1570]" pageId="4" pageNumber="367">Distribution:</emphasis>
<collectingCountry id="F3697616FFF7CF711F24F9A10C9D014B" box="[362,433,1544,1570]" name="China" pageId="4" pageNumber="367">China</collectingCountry>
:
<collectingRegion id="49BAF864FFF7CF711FF5F9A10F33014B" box="[443,543,1544,1570]" country="China" name="Zhejiang" pageId="4" pageNumber="367">Zhejiang</collectingRegion>
(
<bibRefCitation id="EFEF4B77FFF7CF711C63F9A10FDB014B" author="Zhang, Y. L. &amp; Zhang, W. Z. &amp; Chen, B." box="[557,759,1544,1570]" pageId="4" pageNumber="367" pagination="235 - 245" refId="ref5993" refString="Zhang, Y. L., Zhang, W. Z. &amp; Chen, B. (1997) Selenocephaline leafhoppers (Homoptera: Cicadellidae) from Mt. Funiushan in Henan Province. Entomotaxonomia, 19, 235 - 245." type="journal article" year="1997">
Zhang
<emphasis id="B90AEA94FFF7CF711C30F9A00F9A014B" box="[638,694,1544,1570]" italics="true" pageId="4" pageNumber="367">et al.</emphasis>
1997
</bibRefCitation>
;
<bibRefCitation id="EFEF4B77FFF7CF711D4DF9A10EE9014B" author="Cen, Y. W. &amp; Cai, P." box="[771,965,1544,1570]" pageId="4" pageNumber="367" pagination="116 - 120" refId="ref5050" refString="Cen, Y. W. &amp; Cai, P. (2002) Four new species of the subfamily Selenocephalinae from China (Homoptera: Cicadellidae). Acta Zootaxanomica Sinica, 27, 116 - 120." type="journal article" year="2002">Cen &amp; Cai, 2002</bibRefCitation>
;
<bibRefCitation id="EFEF4B77FFF7CF711D9EF9A109BB014B" author="Shang, S. Q. &amp; Shen L. &amp; Zhang, Y. L. &amp; Li, H. H." box="[976,1175,1544,1570]" pageId="4" pageNumber="367" pagination="152 - 154" refId="ref5632" refString="Shang, S. Q., Shen L., Zhang, Y. L. &amp; Li, H. H. (2006) Taxonomy study on the leafhopper genus Fistulatus Zhang, with description of a new species from China (Hemiptera, Selenocephalinae, Paraboloponini). Acta Zootaxonomica Sinica, 31, 152 - 154." type="journal article" year="2006">
Shang
<emphasis id="B90AEA94FFF7CF711A52F9A00979014B" box="[1052,1109,1544,1570]" italics="true" pageId="4" pageNumber="367">et al.</emphasis>
2006
</bibRefCitation>
;
<bibRefCitation id="EFEF4B77FFF7CF711AECF9A00843014B" author="Lu, L. &amp; Zhang, Y. L." box="[1186,1391,1544,1571]" pageId="4" pageNumber="367" pagination="247 - 250" refId="ref5506" refString="Lu, L. &amp; Zhang, Y. L. (2014) Taxonomy of the Oriental leafhopper genus Fistulatus (Hemiptera: Cicadellidae: Deltocephalinae), with description of a new species from China. Zootaxa, 3838 (2), 247 - 250. https: // doi. org / 10.11646 / zootaxa. 3838.2.10" type="journal article" year="2014">Lu &amp; Zhang 2014</bibRefCitation>
;
<bibRefCitation id="EFEF4B77FFF7CF711B34F9A10C3F012E" author="Qu, L. &amp; Webb, M. D. &amp; Dai, R. H." pageId="4" pageNumber="367" pagination="260 - 270" refId="ref5384" refString="Qu, L., Webb, M. D. &amp; Dai, R. H. (2015) A new genus and species of the leafhopper subtribe Paraboloponina from China (Hemiptera, Cicadellidae). Zootaxa, 3919 (2), 260 - 270. https: // doi. org / 10.11646 / zootaxa. 3919.2.2" type="journal article" year="2015">
Qu
<emphasis id="B90AEA94FFF7CF711ED9F9840DE3012F" box="[151,207,1580,1606]" italics="true" pageId="4" pageNumber="367">et al.</emphasis>
2015
</bibRefCitation>
);
<collectingRegion id="49BAF864FFF7CF711F68F9850CA5012F" box="[294,393,1580,1606]" country="China" name="Guangxi" pageId="4" pageNumber="367">Guangxi</collectingRegion>
,
<collectingRegion id="49BAF864FFF7CF711FDBF9850CD5012F" box="[405,505,1580,1606]" country="China" name="Guizhou" pageId="4" pageNumber="367">Guizhou</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,141 @@
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<mods:title id="8DE2997265F512F3AD11D9F48CE2B43D">Key to species of the leafhopper genus Fistulatus Zhang, Zhang &amp; Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species</mods:title>
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<heading id="D08981EAFFF7CF711ED9FD7F0FA80599" bold="true" box="[151,644,726,753]" fontSize="11" level="1" pageId="4" pageNumber="367" reason="1">
<taxonomicName id="4C7E4D05FFF7CF711ED9FD7F0FA80599" ID-CoL="6J795" authority="Shang &amp; Zhang, 2003" authorityName="Shang &amp; Zhang" authorityYear="2003" box="[151,644,726,753]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="athena">
<emphasis id="B90AEA94FFF7CF711ED9FD7F0FA80599" bold="true" box="[151,644,726,753]" pageId="4" pageNumber="367">
<emphasis id="B90AEA94FFF7CF711ED9FD7F0C400598" bold="true" box="[151,364,726,753]" italics="true" pageId="4" pageNumber="367">Fistulatus athena</emphasis>
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<paragraph id="8BC13686FFF7CF711ED9FD520C0B047C" blockId="4.[151,644,726,789]" box="[151,295,763,789]" pageId="4" pageNumber="367">
(
<figureCitation id="13452A03FFF7CF711ED1FD520C33047C" box="[159,287,763,789]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="4" pageNumber="367">Figs 1419</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF7CF711ED9FC960E80043E" box="[151,940,831,856]" pageId="4" pageNumber="367" type="reference_group">
<paragraph id="8BC13686FFF7CF711ED9FC960E80043E" blockId="4.[151,940,831,856]" box="[151,940,831,856]" pageId="4" pageNumber="367">
<treatmentCitationGroup id="AB6E11A8FFF7CF711ED9FC960E80043E" box="[151,940,831,856]" pageId="4" pageNumber="367">
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<taxonomicName id="4C7E4D05FFF7CF711ED9FC960F7B043E" ID-CoL="6J795" authority="Shang &amp; Zhang, 2003: 21" authorityName="Shang &amp; Zhang" authorityPageNumber="21" authorityYear="2003" box="[151,599,831,855]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="athena">
<emphasis id="B90AEA94FFF7CF711ED9FC960C64043E" box="[151,328,831,855]" italics="true" pageId="4" pageNumber="367">Fistulatus athena</emphasis>
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<bibRefCitation id="EFEF4B77FFF7CF711F00FC960F7B043E" author="Shang, S. Q. &amp; Zhang, Y. L." box="[334,599,831,855]" pageId="4" pageNumber="367" pagination="21 - 24" refId="ref5694" refString="Shang, S. Q. &amp; Zhang, Y. L. (2003) Two new species from Sabah (Kalimantan) of the genus Fistulatus (Homoptera: Cicadellidae: Selenocephalinae: Paraboloponini). Entomotaxonomia, 25, 21 - 24." type="journal article" year="2003">Shang &amp; Zhang, 2003: 21</bibRefCitation>
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<typeStatus id="54C58824FFF7CF711C2FFCE90FBF0431" box="[609,659,832,856]" pageId="4" pageNumber="367">Type</typeStatus>
locality:
<collectingCountry id="F3697616FFF7CF711CBDFC960E7C043E" box="[755,848,831,855]" name="Malaysia" pageId="4" pageNumber="367">Malaysia</collectingCountry>
(
<collectingRegion id="49BAF864FFF7CF711D13FC960E8C043E" box="[861,928,831,855]" country="Malaysia" name="Sabah" pageId="4" pageNumber="367">Sabah</collectingRegion>
).
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</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF7CF711ED9FC2C0C0C0362" pageId="4" pageNumber="367" type="discussion">
<paragraph id="8BC13686FFF7CF711ED9FC2C0C0C0362" blockId="4.[151,1436,901,1072]" pageId="4" pageNumber="367">
<emphasis id="B90AEA94FFF7CF711ED9FC2C0C2704F6" bold="true" box="[151,267,901,927]" pageId="4" pageNumber="367">Remarks.</emphasis>
This species is similar to
<taxonomicName id="4C7E4D05FFF7CF711C0CFC2F0EEF04F6" authority="Shang &amp; Zhang" authorityName="Shang &amp; Zhang" authorityYear="2003" box="[578,963,901,927]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="monkoboensis">
<emphasis id="B90AEA94FFF7CF711C0CFC2F0E2C04F6" box="[578,768,901,927]" italics="true" pageId="4" pageNumber="367">F. monkoboensis</emphasis>
Shang &amp; Zhang
</taxonomicName>
in having a distinctive shape of pygofer processes, which are directed ventrally and slightly backward in lateral view, but can be distinguished by the presence of lateral processes on connective and by the style with the apical process slightly curved and digitate in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C364650DFFF7CF711E89FBBC08A30359" box="[199,1423,1045,1072]" pageId="4" pageNumber="367" type="distribution">
<paragraph id="8BC13686FFF7CF711E89FBBC08A30359" blockId="4.[151,1436,901,1072]" box="[199,1423,1045,1072]" pageId="4" pageNumber="367">
<emphasis id="B90AEA94FFF7CF711E89FBBC0C4F0346" bold="true" box="[199,355,1045,1071]" pageId="4" pageNumber="367">Distribution:</emphasis>
<collectingCountry id="F3697616FFF7CF711F24FBBC0CF90346" box="[362,469,1045,1071]" name="Malaysia" pageId="4" pageNumber="367">Malaysia</collectingCountry>
:
<collectingRegion id="49BAF864FFF7CF711F91FBBC0F0F0346" box="[479,547,1045,1071]" country="Malaysia" name="Sabah" pageId="4" pageNumber="367">Sabah</collectingRegion>
(
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;
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Shang
<emphasis id="B90AEA94FFF7CF711DCEFBBF0E940346" box="[896,952,1045,1071]" italics="true" pageId="4" pageNumber="367">et al.</emphasis>
2006
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Qu
<emphasis id="B90AEA94FFF7CF711B49FBBF08130346" box="[1287,1343,1045,1071]" italics="true" pageId="4" pageNumber="367">et al.</emphasis>
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<emphasis id="B90AEA94FFFBCF7D1ED9FE570CFD0570" bold="true" box="[151,465,510,537]" italics="true" pageId="8" pageNumber="371">Fistulatus quadrispinosus</emphasis>
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(
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<emphasis id="B90AEA94FFFBCF7D1ED9FDCE0CB20516" box="[151,414,615,639]" italics="true" pageId="8" pageNumber="371">Fistulatus quadrispinosus</emphasis>
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<emphasis id="B90AEA94FFFBCF7D1ED9FD040C2705AE" bold="true" box="[151,267,685,711]" pageId="8" pageNumber="371">Remarks.</emphasis>
This species is similar to
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Cen &amp; Cai
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in having two pairs of processes on the pygofer but can be distinguished by a pair of narrow medial processes on the aedeagal shaft in ventral view.
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<emphasis id="B90AEA94FFFBCF7D1E89FD5C0C4F0466" bold="true" box="[199,355,757,783]" pageId="8" pageNumber="371">Distribution:</emphasis>
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:
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(
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Zhang
<emphasis id="B90AEA94FFFBCF7D1C30FD5F0F9A0466" box="[638,694,757,783]" italics="true" pageId="8" pageNumber="371">et al.</emphasis>
1997
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;
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Qu
<emphasis id="B90AEA94FFFBCF7D1A4BFD5F09110466" box="[1029,1085,757,783]" italics="true" pageId="8" pageNumber="371">et al.</emphasis>
2015
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Description (based on
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,
<figureCitation id="134F2A29932E904856A95D26478EFC35" box="[964,1002,871,897]" captionStart="FIGURE 3" captionStartId="7.[152,255,1309,1333]" captionTargetBox="[151,1436,181,1285]" captionTargetId="figure-19@7.[151,1436,181,1285]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Hippocrepis fuelleborni: A) Body in ventral view showing the distribution of the papillae. B) Detail of ventral papillae. C) Oral sucker and genital pore, ventral view. D) Details of tegument spines between oral sucker and genital pore, ventral view. E) Lateral view of the oral sucker region. F) Details of tegument spines in the anterior end of body, lateral view." pageId="5" pageNumber="230">3A</figureCitation>
<figureCitation id="134F2A29932E904856A95D26478EFC35" box="[964,1002,871,897]" captionStart="FIGURE 3" captionStartId="7.[152,255,1309,1333]" captionTargetBox="[151,1436,181,1285]" captionTargetId="figure-19@7.[151,1436,181,1285]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Hippocrepis fuelleborni: A) Body in ventral view showing the distribution of the papillae. B) Detail of ventral papillae. C) Oral sucker and genital pore, ventral view. D) Details of tegument spines between oral sucker and genital pore, ventral view. E) Lateral view of the oral sucker region. F) Details of tegument spines in the anterior end of body, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758989" httpUri="https://zenodo.org/record/12758989/files/figure.png" pageId="5" pageNumber="230">3A</figureCitation>
), 2.81 ± 0.44 (1.683.52) long by 0.77 ± 0.13 (0.451.04) wide. Lateral projections on each side of the oral sucker (
<figureCitation id="134F2A29932E904856825DCA4036FC11" box="[1007,1106,907,933]" captionStart="FIGURE 2" captionStartId="6.[152,255,1812,1836]" captionTargetBox="[164,1434,386,1780]" captionTargetId="figure-105@6.[151,1436,381,1788]" captionTargetPageId="6" captionText="FIGURE 2. Drawing of Hippocrepis fuelleborni: A) Complete adult specimen, ventral view. B) Distribution of papillae. C) Detail of cirrus sac, with internal seminal vesicle (isv) and cirrus (ci). D) Detail of the ovarian complex, showing ovary (ov), oviduct (ovd), ootype (oo), and vitelline duct (vd). E) Egg." pageId="5" pageNumber="230">Figs. 2B</figureCitation>
<figureCitation id="134F2A29932E904856825DCA4036FC11" box="[1007,1106,907,933]" captionStart="FIGURE 2" captionStartId="6.[152,255,1812,1836]" captionTargetBox="[164,1434,386,1780]" captionTargetId="figure-105@6.[151,1436,381,1788]" captionTargetPageId="6" captionText="FIGURE 2. Drawing of Hippocrepis fuelleborni: A) Complete adult specimen, ventral view. B) Distribution of papillae. C) Detail of cirrus sac, with internal seminal vesicle (isv) and cirrus (ci). D) Detail of the ovarian complex, showing ovary (ov), oviduct (ovd), ootype (oo), and vitelline duct (vd). E) Egg." figureDoi="http://doi.org/10.5281/zenodo.12758987" httpUri="https://zenodo.org/record/12758987/files/figure.png" pageId="5" pageNumber="230">Figs. 2B</figureCitation>
,
<figureCitation id="134F2A29932E904851305DCA40E5FC11" box="[1117,1153,907,933]" captionStart="FIGURE 3" captionStartId="7.[152,255,1309,1333]" captionTargetBox="[151,1436,181,1285]" captionTargetId="figure-19@7.[151,1436,181,1285]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Hippocrepis fuelleborni: A) Body in ventral view showing the distribution of the papillae. B) Detail of ventral papillae. C) Oral sucker and genital pore, ventral view. D) Details of tegument spines between oral sucker and genital pore, ventral view. E) Lateral view of the oral sucker region. F) Details of tegument spines in the anterior end of body, lateral view." pageId="5" pageNumber="230">3C</figureCitation>
<figureCitation id="134F2A29932E904851305DCA40E5FC11" box="[1117,1153,907,933]" captionStart="FIGURE 3" captionStartId="7.[152,255,1309,1333]" captionTargetBox="[151,1436,181,1285]" captionTargetId="figure-19@7.[151,1436,181,1285]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Hippocrepis fuelleborni: A) Body in ventral view showing the distribution of the papillae. B) Detail of ventral papillae. C) Oral sucker and genital pore, ventral view. D) Details of tegument spines between oral sucker and genital pore, ventral view. E) Lateral view of the oral sucker region. F) Details of tegument spines in the anterior end of body, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758989" httpUri="https://zenodo.org/record/12758989/files/figure.png" pageId="5" pageNumber="230">3C</figureCitation>
). Tegument covered with spines (
<figureCitation id="134F2A29932E904855815DEE4504FC7D" box="[236,352,943,969]" captionStart="FIGURE 3" captionStartId="7.[152,255,1309,1333]" captionTargetBox="[151,1436,181,1285]" captionTargetId="figure-19@7.[151,1436,181,1285]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Hippocrepis fuelleborni: A) Body in ventral view showing the distribution of the papillae. B) Detail of ventral papillae. C) Oral sucker and genital pore, ventral view. D) Details of tegument spines between oral sucker and genital pore, ventral view. E) Lateral view of the oral sucker region. F) Details of tegument spines in the anterior end of body, lateral view." pageId="5" pageNumber="230">Fig. 3EF</figureCitation>
<figureCitation id="134F2A29932E904855815DEE4504FC7D" box="[236,352,943,969]" captionStart="FIGURE 3" captionStartId="7.[152,255,1309,1333]" captionTargetBox="[151,1436,181,1285]" captionTargetId="figure-19@7.[151,1436,181,1285]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Hippocrepis fuelleborni: A) Body in ventral view showing the distribution of the papillae. B) Detail of ventral papillae. C) Oral sucker and genital pore, ventral view. D) Details of tegument spines between oral sucker and genital pore, ventral view. E) Lateral view of the oral sucker region. F) Details of tegument spines in the anterior end of body, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758989" httpUri="https://zenodo.org/record/12758989/files/figure.png" pageId="5" pageNumber="230">Fig. 3EF</figureCitation>
), smooth oral sucker edges, and covered with small spines from the oral sucker to the genital pore (
<figureCitation id="134F2A29932E904855F25D924578FC59" box="[159,284,979,1005]" captionStart="FIGURE 3" captionStartId="7.[152,255,1309,1333]" captionTargetBox="[151,1436,181,1285]" captionTargetId="figure-19@7.[151,1436,181,1285]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Hippocrepis fuelleborni: A) Body in ventral view showing the distribution of the papillae. B) Detail of ventral papillae. C) Oral sucker and genital pore, ventral view. D) Details of tegument spines between oral sucker and genital pore, ventral view. E) Lateral view of the oral sucker region. F) Details of tegument spines in the anterior end of body, lateral view." pageId="5" pageNumber="230">Fig. 3CD</figureCitation>
<figureCitation id="134F2A29932E904855F25D924578FC59" box="[159,284,979,1005]" captionStart="FIGURE 3" captionStartId="7.[152,255,1309,1333]" captionTargetBox="[151,1436,181,1285]" captionTargetId="figure-19@7.[151,1436,181,1285]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Hippocrepis fuelleborni: A) Body in ventral view showing the distribution of the papillae. B) Detail of ventral papillae. C) Oral sucker and genital pore, ventral view. D) Details of tegument spines between oral sucker and genital pore, ventral view. E) Lateral view of the oral sucker region. F) Details of tegument spines in the anterior end of body, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758989" httpUri="https://zenodo.org/record/12758989/files/figure.png" pageId="5" pageNumber="230">Fig. 3CD</figureCitation>
). Ventral surface concave and provided with numerous longitudinal rows of ventral papillae arranged irregularly, extending from the middle zone of the cirrus sac to posterior extremity (
<figureCitation id="134F2A29932E9048513B5DB640DEFBA5" box="[1110,1210,1015,1041]" captionStart="FIGURE 2" captionStartId="6.[152,255,1812,1836]" captionTargetBox="[164,1434,386,1780]" captionTargetId="figure-105@6.[151,1436,381,1788]" captionTargetPageId="6" captionText="FIGURE 2. Drawing of Hippocrepis fuelleborni: A) Complete adult specimen, ventral view. B) Distribution of papillae. C) Detail of cirrus sac, with internal seminal vesicle (isv) and cirrus (ci). D) Detail of the ovarian complex, showing ovary (ov), oviduct (ovd), ootype (oo), and vitelline duct (vd). E) Egg." pageId="5" pageNumber="230">Figs. 2B</figureCitation>
<figureCitation id="134F2A29932E9048513B5DB640DEFBA5" box="[1110,1210,1015,1041]" captionStart="FIGURE 2" captionStartId="6.[152,255,1812,1836]" captionTargetBox="[164,1434,386,1780]" captionTargetId="figure-105@6.[151,1436,381,1788]" captionTargetPageId="6" captionText="FIGURE 2. Drawing of Hippocrepis fuelleborni: A) Complete adult specimen, ventral view. B) Distribution of papillae. C) Detail of cirrus sac, with internal seminal vesicle (isv) and cirrus (ci). D) Detail of the ovarian complex, showing ovary (ov), oviduct (ovd), ootype (oo), and vitelline duct (vd). E) Egg." figureDoi="http://doi.org/10.5281/zenodo.12758987" httpUri="https://zenodo.org/record/12758987/files/figure.png" pageId="5" pageNumber="230">Figs. 2B</figureCitation>
,
<figureCitation id="134F2A29932E904851A55DB64169FBA5" box="[1224,1293,1015,1041]" captionStart="FIGURE 3" captionStartId="7.[152,255,1309,1333]" captionTargetBox="[151,1436,181,1285]" captionTargetId="figure-19@7.[151,1436,181,1285]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Hippocrepis fuelleborni: A) Body in ventral view showing the distribution of the papillae. B) Detail of ventral papillae. C) Oral sucker and genital pore, ventral view. D) Details of tegument spines between oral sucker and genital pore, ventral view. E) Lateral view of the oral sucker region. F) Details of tegument spines in the anterior end of body, lateral view." pageId="5" pageNumber="230">3AB</figureCitation>
<figureCitation id="134F2A29932E904851A55DB64169FBA5" box="[1224,1293,1015,1041]" captionStart="FIGURE 3" captionStartId="7.[152,255,1309,1333]" captionTargetBox="[151,1436,181,1285]" captionTargetId="figure-19@7.[151,1436,181,1285]" captionTargetPageId="7" captionText="FIGURE 3. SEM images of Hippocrepis fuelleborni: A) Body in ventral view showing the distribution of the papillae. B) Detail of ventral papillae. C) Oral sucker and genital pore, ventral view. D) Details of tegument spines between oral sucker and genital pore, ventral view. E) Lateral view of the oral sucker region. F) Details of tegument spines in the anterior end of body, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758989" httpUri="https://zenodo.org/record/12758989/files/figure.png" pageId="5" pageNumber="230">3AB</figureCitation>
,
<figureCitation id="134F2A29932E904850775DB64124FBA5" box="[1306,1344,1015,1041]" captionStart="FIGURE 4" captionStartId="9.[152,255,945,969]" captionTargetBox="[151,1436,181,921]" captionTargetId="figure-19@9.[151,1436,181,921]" captionTargetPageId="9" captionText="FIGURE 4. Histological section of Hippocrepis fuelleborni: A) Detail of ventral papillae. B) Sagittal section showing genital pore (gp) and cirrus (ci) with small spines. C) Posterior end of body: caeca (ca), ovary (ov), testis (te). D) Egg." pageId="5" pageNumber="230">4A</figureCitation>
<figureCitation id="134F2A29932E904850775DB64124FBA5" box="[1306,1344,1015,1041]" captionStart="FIGURE 4" captionStartId="9.[152,255,945,969]" captionTargetBox="[151,1436,181,921]" captionTargetId="figure-19@9.[151,1436,181,921]" captionTargetPageId="9" captionText="FIGURE 4. Histological section of Hippocrepis fuelleborni: A) Detail of ventral papillae. B) Sagittal section showing genital pore (gp) and cirrus (ci) with small spines. C) Posterior end of body: caeca (ca), ovary (ov), testis (te). D) Egg." figureDoi="http://doi.org/10.5281/zenodo.12758992" httpUri="https://zenodo.org/record/12758992/files/figure.png" pageId="5" pageNumber="230">4A</figureCitation>
), with a diameter of 51.97 ± 6.34 (43.0257.81) µm. Ventral sucker absent. Oral sucker subterminal, 0.28 ± 0.04 (0.210.34) long by 0.22 ± 0.003 (0.170.32) wide. Pharynx absent; oesophagus 0.05 ± 0.01 (0.040.07). Caeca with lateral diverticula in its entire extension, bordering vitelline follicles, fuse posteriorly to the ovary passing between testes, up to level of their posterior border (
<figureCitation id="134F2A29932E904857415AC646EBFB15" box="[556,655,1159,1185]" captionStart="FIGURE 2" captionStartId="6.[152,255,1812,1836]" captionTargetBox="[164,1434,386,1780]" captionTargetId="figure-105@6.[151,1436,381,1788]" captionTargetPageId="6" captionText="FIGURE 2. Drawing of Hippocrepis fuelleborni: A) Complete adult specimen, ventral view. B) Distribution of papillae. C) Detail of cirrus sac, with internal seminal vesicle (isv) and cirrus (ci). D) Detail of the ovarian complex, showing ovary (ov), oviduct (ovd), ootype (oo), and vitelline duct (vd). E) Egg." pageId="5" pageNumber="230">Figs. 2A</figureCitation>
<figureCitation id="134F2A29932E904857415AC646EBFB15" box="[556,655,1159,1185]" captionStart="FIGURE 2" captionStartId="6.[152,255,1812,1836]" captionTargetBox="[164,1434,386,1780]" captionTargetId="figure-105@6.[151,1436,381,1788]" captionTargetPageId="6" captionText="FIGURE 2. Drawing of Hippocrepis fuelleborni: A) Complete adult specimen, ventral view. B) Distribution of papillae. C) Detail of cirrus sac, with internal seminal vesicle (isv) and cirrus (ci). D) Detail of the ovarian complex, showing ovary (ov), oviduct (ovd), ootype (oo), and vitelline duct (vd). E) Egg." figureDoi="http://doi.org/10.5281/zenodo.12758987" httpUri="https://zenodo.org/record/12758987/files/figure.png" pageId="5" pageNumber="230">Figs. 2A</figureCitation>
,
<figureCitation id="134F2A29932E904857F65AC646DBFB15" box="[667,703,1159,1185]" captionStart="FIGURE 4" captionStartId="9.[152,255,945,969]" captionTargetBox="[151,1436,181,921]" captionTargetId="figure-19@9.[151,1436,181,921]" captionTargetPageId="9" captionText="FIGURE 4. Histological section of Hippocrepis fuelleborni: A) Detail of ventral papillae. B) Sagittal section showing genital pore (gp) and cirrus (ci) with small spines. C) Posterior end of body: caeca (ca), ovary (ov), testis (te). D) Egg." pageId="5" pageNumber="230">4C</figureCitation>
<figureCitation id="134F2A29932E904857F65AC646DBFB15" box="[667,703,1159,1185]" captionStart="FIGURE 4" captionStartId="9.[152,255,945,969]" captionTargetBox="[151,1436,181,921]" captionTargetId="figure-19@9.[151,1436,181,921]" captionTargetPageId="9" captionText="FIGURE 4. Histological section of Hippocrepis fuelleborni: A) Detail of ventral papillae. B) Sagittal section showing genital pore (gp) and cirrus (ci) with small spines. C) Posterior end of body: caeca (ca), ovary (ov), testis (te). D) Egg." figureDoi="http://doi.org/10.5281/zenodo.12758992" httpUri="https://zenodo.org/record/12758992/files/figure.png" pageId="5" pageNumber="230">4C</figureCitation>
). Testes irregularly lobed, oblique, posterior to caecal union, right testis 0.24 ± 0.05 (0.150.34) long by 0.19 ± 0.04 (0.130.27) wide, left testis 0.23 ± 0.05 (0.150.32) long by 0.19 ± 0.05 (0.130.28) wide. Cirrus sac median, elongate 0.68 ± 0.08 (0.550.88) long, containing cirrus and internal seminal vesicle (
<figureCitation id="134F2A29932E9048543E5AB245C8FAB9" box="[339,428,1267,1293]" captionStart="FIGURE 2" captionStartId="6.[152,255,1812,1836]" captionTargetBox="[164,1434,386,1780]" captionTargetId="figure-105@6.[151,1436,381,1788]" captionTargetPageId="6" captionText="FIGURE 2. Drawing of Hippocrepis fuelleborni: A) Complete adult specimen, ventral view. B) Distribution of papillae. C) Detail of cirrus sac, with internal seminal vesicle (isv) and cirrus (ci). D) Detail of the ovarian complex, showing ovary (ov), oviduct (ovd), ootype (oo), and vitelline duct (vd). E) Egg." pageId="5" pageNumber="230">Fig. 2C</figureCitation>
<figureCitation id="134F2A29932E9048543E5AB245C8FAB9" box="[339,428,1267,1293]" captionStart="FIGURE 2" captionStartId="6.[152,255,1812,1836]" captionTargetBox="[164,1434,386,1780]" captionTargetId="figure-105@6.[151,1436,381,1788]" captionTargetPageId="6" captionText="FIGURE 2. Drawing of Hippocrepis fuelleborni: A) Complete adult specimen, ventral view. B) Distribution of papillae. C) Detail of cirrus sac, with internal seminal vesicle (isv) and cirrus (ci). D) Detail of the ovarian complex, showing ovary (ov), oviduct (ovd), ootype (oo), and vitelline duct (vd). E) Egg." figureDoi="http://doi.org/10.5281/zenodo.12758987" httpUri="https://zenodo.org/record/12758987/files/figure.png" pageId="5" pageNumber="230">Fig. 2C</figureCitation>
). Cirrus not everted, with small spines (
<figureCitation id="134F2A29932E904856005AB247A2FAB9" box="[877,966,1267,1293]" captionStart="FIGURE 4" captionStartId="9.[152,255,945,969]" captionTargetBox="[151,1436,181,921]" captionTargetId="figure-19@9.[151,1436,181,921]" captionTargetPageId="9" captionText="FIGURE 4. Histological section of Hippocrepis fuelleborni: A) Detail of ventral papillae. B) Sagittal section showing genital pore (gp) and cirrus (ci) with small spines. C) Posterior end of body: caeca (ca), ovary (ov), testis (te). D) Egg." pageId="5" pageNumber="230">Fig. 4B</figureCitation>
<figureCitation id="134F2A29932E904856005AB247A2FAB9" box="[877,966,1267,1293]" captionStart="FIGURE 4" captionStartId="9.[152,255,945,969]" captionTargetBox="[151,1436,181,921]" captionTargetId="figure-19@9.[151,1436,181,921]" captionTargetPageId="9" captionText="FIGURE 4. Histological section of Hippocrepis fuelleborni: A) Detail of ventral papillae. B) Sagittal section showing genital pore (gp) and cirrus (ci) with small spines. C) Posterior end of body: caeca (ca), ovary (ov), testis (te). D) Egg." figureDoi="http://doi.org/10.5281/zenodo.12758992" httpUri="https://zenodo.org/record/12758992/files/figure.png" pageId="5" pageNumber="230">Fig. 4B</figureCitation>
). Genital pore median, posterior to caecal bifurcation. Ovary median, pretesticular, transversally elongated (
<figureCitation id="134F2A29932E904856145B5647B6FA85" box="[889,978,1303,1329]" captionStart="FIGURE 2" captionStartId="6.[152,255,1812,1836]" captionTargetBox="[164,1434,386,1780]" captionTargetId="figure-105@6.[151,1436,381,1788]" captionTargetPageId="6" captionText="FIGURE 2. Drawing of Hippocrepis fuelleborni: A) Complete adult specimen, ventral view. B) Distribution of papillae. C) Detail of cirrus sac, with internal seminal vesicle (isv) and cirrus (ci). D) Detail of the ovarian complex, showing ovary (ov), oviduct (ovd), ootype (oo), and vitelline duct (vd). E) Egg." pageId="5" pageNumber="230">Fig. 2D</figureCitation>
<figureCitation id="134F2A29932E904856145B5647B6FA85" box="[889,978,1303,1329]" captionStart="FIGURE 2" captionStartId="6.[152,255,1812,1836]" captionTargetBox="[164,1434,386,1780]" captionTargetId="figure-105@6.[151,1436,381,1788]" captionTargetPageId="6" captionText="FIGURE 2. Drawing of Hippocrepis fuelleborni: A) Complete adult specimen, ventral view. B) Distribution of papillae. C) Detail of cirrus sac, with internal seminal vesicle (isv) and cirrus (ci). D) Detail of the ovarian complex, showing ovary (ov), oviduct (ovd), ootype (oo), and vitelline duct (vd). E) Egg." figureDoi="http://doi.org/10.5281/zenodo.12758987" httpUri="https://zenodo.org/record/12758987/files/figure.png" pageId="5" pageNumber="230">Fig. 2D</figureCitation>
), 0.13 ± 0.03 (0.100.19) long by 0.24 ± 0.06 (0.100.36) wide. Vitelline follicles arranged in longitudinal series, extracaecal and pre-ovarian, ending close to testes, 0.60 ± 0.07 (0.500.70) long. Uterus transverse coils located in intercaecal space. Eggs oval, without polar filaments (
<figureCitation id="134F2A29932E904854625BC2450BFA29" box="[271,367,1411,1437]" captionStart="FIGURE 2" captionStartId="6.[152,255,1812,1836]" captionTargetBox="[164,1434,386,1780]" captionTargetId="figure-105@6.[151,1436,381,1788]" captionTargetPageId="6" captionText="FIGURE 2. Drawing of Hippocrepis fuelleborni: A) Complete adult specimen, ventral view. B) Distribution of papillae. C) Detail of cirrus sac, with internal seminal vesicle (isv) and cirrus (ci). D) Detail of the ovarian complex, showing ovary (ov), oviduct (ovd), ootype (oo), and vitelline duct (vd). E) Egg." pageId="5" pageNumber="230">Figs. 2E</figureCitation>
<figureCitation id="134F2A29932E904854625BC2450BFA29" box="[271,367,1411,1437]" captionStart="FIGURE 2" captionStartId="6.[152,255,1812,1836]" captionTargetBox="[164,1434,386,1780]" captionTargetId="figure-105@6.[151,1436,381,1788]" captionTargetPageId="6" captionText="FIGURE 2. Drawing of Hippocrepis fuelleborni: A) Complete adult specimen, ventral view. B) Distribution of papillae. C) Detail of cirrus sac, with internal seminal vesicle (isv) and cirrus (ci). D) Detail of the ovarian complex, showing ovary (ov), oviduct (ovd), ootype (oo), and vitelline duct (vd). E) Egg." figureDoi="http://doi.org/10.5281/zenodo.12758987" httpUri="https://zenodo.org/record/12758987/files/figure.png" pageId="5" pageNumber="230">Figs. 2E</figureCitation>
,
<figureCitation id="134F2A29932E904854165BC245C4FA29" box="[379,416,1411,1437]" captionStart="FIGURE 4" captionStartId="9.[152,255,945,969]" captionTargetBox="[151,1436,181,921]" captionTargetId="figure-19@9.[151,1436,181,921]" captionTargetPageId="9" captionText="FIGURE 4. Histological section of Hippocrepis fuelleborni: A) Detail of ventral papillae. B) Sagittal section showing genital pore (gp) and cirrus (ci) with small spines. C) Posterior end of body: caeca (ca), ovary (ov), testis (te). D) Egg." pageId="5" pageNumber="230">4D</figureCitation>
<figureCitation id="134F2A29932E904854165BC245C4FA29" box="[379,416,1411,1437]" captionStart="FIGURE 4" captionStartId="9.[152,255,945,969]" captionTargetBox="[151,1436,181,921]" captionTargetId="figure-19@9.[151,1436,181,921]" captionTargetPageId="9" captionText="FIGURE 4. Histological section of Hippocrepis fuelleborni: A) Detail of ventral papillae. B) Sagittal section showing genital pore (gp) and cirrus (ci) with small spines. C) Posterior end of body: caeca (ca), ovary (ov), testis (te). D) Egg." figureDoi="http://doi.org/10.5281/zenodo.12758992" httpUri="https://zenodo.org/record/12758992/files/figure.png" pageId="5" pageNumber="230">4D</figureCitation>
), 26.38 ± 11.94 (20.7069.15) µm long by 8.54 ± 1.09 (6.7210.25) µm wide.
</paragraph>
</subSubSection>

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<document id="EBA102375B841CBE206496B3C0312604" ID-DOI="10.11646/zootaxa.5481.3.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="12758924" ID-ZooBank="F885B41C-3D17-4CD2-8046-85C2D7EE4721" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721216146773" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="1A2C557E854F3E5B9EACFB88FE56F97B" docLanguage="en" docName="zootaxa.5481.3.8.pdf" docOrigin="Zootaxa 5481 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5481.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptomastax minutissima Jałoszyński, 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="394" masterDocId="E6152D06854E3E589E3BFFB7FFF1FF80" masterDocTitle="A new species of Leptomastax Pirazzoli in Israel (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="396" masterPageNumber="391" pageNumber="392" updateTime="1721216561249" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6746233A44AE19CA77E5C736B525383C">A new species of Leptomastax Pirazzoli in Israel (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:date id="9B971309AF8EEE2C884A63CDC1977192">2024</mods:date>
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<paragraph id="923AE468854F3E599EACFB88FDE6FBD9" blockId="1.[151,535,1087,1149]" box="[151,535,1087,1113]" pageId="1" pageNumber="392">
<heading id="C9725304854F3E599EACFB88FDE6FBD9" bold="true" box="[151,535,1087,1113]" fontSize="11" level="1" pageId="1" pageNumber="392" reason="1">
<emphasis id="A0F1387A854F3E599EACFB88FDE6FBD9" bold="true" box="[151,535,1087,1113]" pageId="1" pageNumber="392">
<taxonomicName id="55859FEB854F3E599EACFB88FE3CFBD9" authorityName="Jałoszyński" authorityYear="2024" box="[151,461,1087,1113]" class="Insecta" family="Staphylinidae" genus="Leptomastax" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="minutissima" status="sp. nov.">
<emphasis id="A0F1387A854F3E599EACFB88FE3CFBD9" bold="true" box="[151,461,1087,1113]" italics="true" pageId="1" pageNumber="392">Leptomastax minutissima</emphasis>
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<taxonomicNameLabel id="BBC28501854F3E599FEEFB88FDE6FBD9" box="[469,535,1087,1113]" pageId="1" pageNumber="392" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph id="923AE468854F3E599EACFBD4FEE8FBFD" blockId="1.[151,535,1087,1149]" box="[151,281,1123,1149]" pageId="1" pageNumber="392">
(
<figureCitation id="0ABEF8ED854F3E599EA4FBD4FEE1FBFD" box="[159,272,1123,1149]" captionStart-0="FIGURES 14" captionStart-1="FIGURES 57" captionStart-2="FIGURES 89" captionStart-3="FIGURES 1013" captionStartId-0="2.[151,264,1363,1387]" captionStartId-1="3.[151,264,1186,1210]" captionStartId-2="4.[151,264,1096,1120]" captionStartId-3="4.[151,264,2013,2037]" captionTargetBox-0="[151,1435,181,1340]" captionTargetBox-1="[151,1435,181,1162]" captionTargetBox-2="[158,1430,181,1072]" captionTargetBox-3="[159,1429,1187,1983]" captionTargetId-0="figure-21@2.[151,1435,181,1340]" captionTargetId-1="figure-15@3.[151,1435,181,1162]" captionTargetId-2="figure-21@4.[158,1430,181,1072]" captionTargetId-3="figure-54@4.[151,1436,1180,1990]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionText-0="FIGURES 14. Leptomastax minutissima sp. n., holotype male. Dorsal habitus (1); head, pronotum and elytral base in dorsal view (2); left antenna in dorsal view (3); elytra in dorsal view (4)." captionText-1="FIGURES 57. Leptomastax minutissima sp. n., holotype male. Head, prothorax and anterior region of mesothorax in ventral view (5); mouthparts in ventral view (6); left labial palp in ventral view (7)." captionText-2="FIGURES 89. Leptomastax minutissima sp. n., holotype male. Pterothorax and base of abdomen in ventral view (8); right mesocoxa and surrounding areas in ventral view (9)." captionText-3="FIGURES 1013. Leptomastax minutissima sp. n., holotype male. Aedeagus in ventral (10, 12) and lateral (11, 13) views." figureDoi-0="http://doi.org/10.5281/zenodo.12758926" figureDoi-1="http://doi.org/10.5281/zenodo.12758930" figureDoi-2="http://doi.org/10.5281/zenodo.12758932" figureDoi-3="http://doi.org/10.5281/zenodo.12758934" httpUri-0="https://zenodo.org/record/12758926/files/figure.png" httpUri-1="https://zenodo.org/record/12758930/files/figure.png" httpUri-2="https://zenodo.org/record/12758932/files/figure.png" httpUri-3="https://zenodo.org/record/12758934/files/figure.png" pageId="1" pageNumber="392">Figs 113</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DA9FB7E3854F3E599EACFB1CFE96FA8D" pageId="1" pageNumber="392" type="materials_examined">
<paragraph id="923AE468854F3E599EACFB1CFE96FA8D" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="392">
<emphasis id="A0F1387A854F3E599EACFB1CFEB6FB44" bold="true" box="[151,327,1194,1221]" pageId="1" pageNumber="392">Type material.</emphasis>
<materialsCitation id="22EDEE35854F3E599F74FB1CFE96FA8D" collectingDate="1982-04-20" collectionCode="MHNG" collectorName="N. Betzel &amp; P. Jaloszynski" country="Israel" location="ISRAEL" pageId="1" pageNumber="392" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="A0F1387A854F3E599F74FB1CFE34FB45" bold="true" box="[335,453,1195,1221]" pageId="1" pageNumber="392">
<typeStatus id="4D3E5ACA854F3E599F74FB1CFE31FB45" box="[335,448,1195,1221]" pageId="1" pageNumber="392" type="holotype">Holotype</typeStatus>
:
</emphasis>
(
<emphasis id="A0F1387A854F3E599FECFB1DFDA3FB44" bold="true" box="[471,594,1194,1220]" pageId="1" pageNumber="392">
<collectingCountry id="EA92A4F8854F3E599FECFB1DFDB4FB44" box="[471,581,1194,1220]" name="Israel" pageId="1" pageNumber="392">ISRAEL</collectingCountry>
):
</emphasis>
<specimenCount id="84832FE1854F3E599C60FB1CFD83FB45" box="[603,626,1195,1221]" count="1" pageId="1" pageNumber="392" type="male"></specimenCount>
, two labels: “ISRAEL: Galilea / Eilon,
<collectorName id="3F7081BE854F3E599A02FB1CFB54FB45" box="[1081,1189,1195,1221]" pageId="1" pageNumber="392">N. Betzel</collectorName>
/ valley,
<date id="E63BC2A8854F3E599B2BFB1CFA7AFB45" box="[1296,1419,1195,1221]" pageId="1" pageNumber="392" value="1982-04-20">
<collectingDate id="F67F3B40854F3E599B2BFB1CFA7AFB45" box="[1296,1419,1195,1221]" pageId="1" pageNumber="392" value="1982-04-20">20.IV.1982</collectingDate>
</date>
/ Löbl, Bes. nr. 13a” [white, printed], “
<taxonomicName id="55859FEB854F3E599C00FB78FD07FB69" authorityName="Pirazzoli" authorityYear="1855" box="[571,758,1231,1257]" class="Insecta" family="Staphylinidae" genus="Leptomastax" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="genus">
<emphasis id="A0F1387A854F3E599C00FB78FD07FB69" box="[571,758,1231,1257]" italics="true" pageId="1" pageNumber="392">LEPTOMASTAX</emphasis>
</taxonomicName>
/
<taxonomicName id="55859FEB854F3E599D37FB78FC62FB68" authorityName="Jałoszyński" authorityYear="2024" box="[780,915,1231,1256]" class="Insecta" family="Staphylinidae" genus="Leptomastax" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="392" phylum="Arthropoda" rank="species" species="minutissima">
<emphasis id="A0F1387A854F3E599D37FB78FC62FB68" box="[780,915,1231,1256]" italics="true" pageId="1" pageNumber="392">minutissima</emphasis>
</taxonomicName>
m. /
<collectorName id="3F7081BE854F3E599DF6FB78FB9DFB69" box="[973,1132,1231,1257]" pageId="1" pageNumber="392">P. Jałoszyński</collectorName>
, '24 /
<typeStatus id="4D3E5ACA854F3E599A95FB78FAA6FB69" box="[1198,1367,1231,1257]" pageId="1" pageNumber="392" type="holotype">HOLOTYPUS</typeStatus>
” [red, printed] (
<collectionCode id="F4947CAD854F3E599EC4FB44FEAAFA8D" box="[255,347,1267,1293]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="392" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA9FB7E3854F3E599EFCFAA1FDD4FA65" pageId="1" pageNumber="392" type="diagnosis">
<paragraph id="923AE468854F3E599EFCFAA1FDD4FA65" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="392">
<emphasis id="A0F1387A854F3E599EFCFAA1FEB3FAB0" bold="true" box="[199,322,1302,1328]" pageId="1" pageNumber="392">Diagnosis.</emphasis>
Body extremely small, below
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; eyes lacking; subocular and elytral macrosetae lacking; prosternum with two large lateral patches of modified setae; anterior region of mesoventral process much wider than its posterior intermesocoxal region; elytral rows of punctures irregular, subhumeral and marginal rows lacking; male with U-shaped impression with distinct lateral margins on posterior 1/3 of metaventrite; aedeagus with simple (i.e., unbranched) and nearly straight endophallus; each paramere with evenly rounded subapical and apical margin and with pointed apex directed mesally.
</paragraph>
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<subSubSection id="DA9FB7E3854F3E5A9EFCFA59FE6FF874" lastPageId="2" lastPageNumber="393" pageId="1" pageNumber="392" type="description">
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<emphasis id="A0F1387A854F3E599EFCFA59FEA8F988" bold="true" box="[199,345,1518,1544]" pageId="1" pageNumber="392">Description.</emphasis>
Body of male (
<figureCitation id="0ABEF8ED854F3E599C3CFA58FDB9F989" box="[519,584,1519,1545]" captionStart="FIGURES 14" captionStartId="2.[151,264,1363,1387]" captionTargetBox="[151,1435,181,1340]" captionTargetId="figure-21@2.[151,1435,181,1340]" captionTargetPageId="2" captionText="FIGURES 14. Leptomastax minutissima sp. n., holotype male. Dorsal habitus (1); head, pronotum and elytral base in dorsal view (2); left antenna in dorsal view (3); elytra in dorsal view (4)." figureDoi="http://doi.org/10.5281/zenodo.12758926" httpUri="https://zenodo.org/record/12758926/files/figure.png" pageId="1" pageNumber="392">Fig. 1</figureCitation>
) strongly elongate, strongly flattened, yellowish brown. BL
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.
</paragraph>
<paragraph id="923AE468854F3E599EFCF9A4FD02F961" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="392">
Head (
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) broadest near middle, eyeless, HL
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, HW
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; vertex and frons together slightly more than twice as wide as long; dorsal postmandibular impressions distinct; subocular (sensu Castellini (1994)) macrosetae absent, vertexal macrosetae (=
<emphasis id="A0F1387A854F3E599CBDF9ECFCABF9F5" box="[646,858,1627,1653]" italics="true" pageId="1" pageNumber="392">temporo-occipitale</emphasis>
of Castellini) present. Dorsum of head and genae (
<figureCitation id="0ABEF8ED854F3E599EA4F9C8FF12F919" box="[159,227,1662,1689]" captionStart="FIGURES 57" captionStartId="3.[151,264,1186,1210]" captionTargetBox="[151,1435,181,1162]" captionTargetId="figure-15@3.[151,1435,181,1162]" captionTargetPageId="3" captionText="FIGURES 57. Leptomastax minutissima sp. n., holotype male. Head, prothorax and anterior region of mesothorax in ventral view (5); mouthparts in ventral view (6); left labial palp in ventral view (7)." figureDoi="http://doi.org/10.5281/zenodo.12758930" httpUri="https://zenodo.org/record/12758930/files/figure.png" pageId="1" pageNumber="392">Fig. 5</figureCitation>
) impunctate. Antenna (
<figureCitation id="0ABEF8ED854F3E599FD6F9C8FDC0F919" box="[493,561,1663,1689]" captionStart="FIGURES 14" captionStartId="2.[151,264,1363,1387]" captionTargetBox="[151,1435,181,1340]" captionTargetId="figure-21@2.[151,1435,181,1340]" captionTargetPageId="2" captionText="FIGURES 14. Leptomastax minutissima sp. n., holotype male. Dorsal habitus (1); head, pronotum and elytral base in dorsal view (2); left antenna in dorsal view (3); elytra in dorsal view (4)." figureDoi="http://doi.org/10.5281/zenodo.12758926" httpUri="https://zenodo.org/record/12758926/files/figure.png" pageId="1" pageNumber="392">Fig. 3</figureCitation>
) slender, AnL
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, scape as long as antennomeres 25 together, pedicel 1.3 times as long as broad, antennomeres 310 each transverse (3 strongly, 10 weakly so), antennomere 11 nearly as long as 910 combined, 1.8 times as long as broad.
</paragraph>
<paragraph id="923AE468854F3E599EFCF95CFF12F8A9" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="392">Labial palps moderately long, palpomere 2 about twice as long as broad, palpomere 3 setiform, slightly longer than 2.</paragraph>
<paragraph id="923AE468854F3E599EFCF884FB8DF85D" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="392">
Pronotum (
<figureCitation id="0ABEF8ED854F3E599F7CF884FE7AF8CD" box="[327,395,1843,1869]" captionStart="FIGURES 14" captionStartId="2.[151,264,1363,1387]" captionTargetBox="[151,1435,181,1340]" captionTargetId="figure-21@2.[151,1435,181,1340]" captionTargetPageId="2" captionText="FIGURES 14. Leptomastax minutissima sp. n., holotype male. Dorsal habitus (1); head, pronotum and elytral base in dorsal view (2); left antenna in dorsal view (3); elytra in dorsal view (4)." figureDoi="http://doi.org/10.5281/zenodo.12758926" httpUri="https://zenodo.org/record/12758926/files/figure.png" pageId="1" pageNumber="392">Fig. 2</figureCitation>
) subtrapezoidal, broadest near anterior fourth; PL
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, PW
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. Anterior margin evenly and weakly rounded; lateral margins strongly rounded in anterior third and nearly straight in posterior half; posterior corners strongly obtuse-angled; posterior margin weakly and evenly rounded. Pronotal disc impunctate and asetose. Prosternum (
<figureCitation id="0ABEF8ED854F3E599FB3F828FE3AF839" box="[392,459,1950,1977]" captionStart="FIGURES 57" captionStartId="3.[151,264,1186,1210]" captionTargetBox="[151,1435,181,1162]" captionTargetId="figure-15@3.[151,1435,181,1162]" captionTargetPageId="3" captionText="FIGURES 57. Leptomastax minutissima sp. n., holotype male. Head, prothorax and anterior region of mesothorax in ventral view (5); mouthparts in ventral view (6); left labial palp in ventral view (7)." figureDoi="http://doi.org/10.5281/zenodo.12758930" httpUri="https://zenodo.org/record/12758930/files/figure.png" pageId="1" pageNumber="392">Fig. 5</figureCitation>
) with pair of large anterolateral patches of modified, strongly flattened and broadened scale-like setae and narrow row of similar setae along median portion of anterior margin.
</paragraph>
<caption id="C6FAB4E0854C3E5A9EACFAE4FCC0FA10" ID-DOI="http://doi.org/10.5281/zenodo.12758926" ID-Zenodo-Dep="12758926" httpUri="https://zenodo.org/record/12758926/files/figure.png" pageId="2" pageNumber="393" startId="2.[151,264,1363,1387]" targetBox="[151,1435,181,1340]" targetPageId="2" targetType="figure">
<paragraph id="923AE468854C3E5A9EACFAE4FCC0FA10" blockId="2.[151,1436,1363,1424]" pageId="2" pageNumber="393">
<emphasis id="A0F1387A854C3E5A9EACFAE4FECAFAEC" bold="true" box="[151,315,1363,1388]" pageId="2" pageNumber="393">FIGURES 14.</emphasis>
<taxonomicName id="55859FEB854C3E5A9F79FAE3FDB4FAEB" authorityName="Jałoszyński" authorityYear="2024" box="[322,581,1364,1387]" class="Insecta" family="Staphylinidae" genus="Leptomastax" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="393" phylum="Arthropoda" rank="species" species="minutissima" status="sp. nov.">
<emphasis id="A0F1387A854C3E5A9F79FAE3FDB4FAEB" box="[322,581,1364,1387]" italics="true" pageId="2" pageNumber="393">Leptomastax minutissima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBC28501854C3E5A9C77FAE3FD74FAEB" box="[588,645,1364,1387]" pageId="2" pageNumber="393" rank="species">sp. n.</taxonomicNameLabel>
, holotype male. Dorsal habitus (
<emphasis id="A0F1387A854C3E5A9DE9FAE3FC2EFAEC" bold="true" box="[978,991,1364,1388]" pageId="2" pageNumber="393">1</emphasis>
); head, pronotum and elytral base in dorsal view (
<emphasis id="A0F1387A854C3E5A9EECFACFFF15FA10" bold="true" box="[215,228,1400,1424]" pageId="2" pageNumber="393">2</emphasis>
); left antenna in dorsal view (
<emphasis id="A0F1387A854C3E5A9C2EFACFFDD3FA10" bold="true" box="[533,546,1400,1424]" pageId="2" pageNumber="393">3</emphasis>
); elytra in dorsal view (
<emphasis id="A0F1387A854C3E5A9D2DFACFFCD2FA10" bold="true" box="[790,803,1400,1424]" pageId="2" pageNumber="393">4</emphasis>
).
</paragraph>
</caption>
<paragraph id="923AE468854C3E5A9EFCFA55FAB3F9C4" blockId="2.[151,1437,1506,2036]" pageId="2" pageNumber="393">
Mesoventrite (
<figureCitation id="0ABEF8ED854C3E5A9F57FA55FE22FA7C" box="[364,467,1506,1532]" captionStart="FIGURES 89" captionStartId="4.[151,264,1096,1120]" captionTargetBox="[158,1430,181,1072]" captionTargetId="figure-21@4.[158,1430,181,1072]" captionTargetPageId="4" captionText="FIGURES 89. Leptomastax minutissima sp. n., holotype male. Pterothorax and base of abdomen in ventral view (8); right mesocoxa and surrounding areas in ventral view (9)." figureDoi="http://doi.org/10.5281/zenodo.12758932" httpUri="https://zenodo.org/record/12758932/files/figure.png" pageId="2" pageNumber="393">Figs 89</figureCitation>
) with scale-like setae on procoxal rests and elongate flattened setae on mesoventral process. Metaventrite (
<figureCitation id="0ABEF8ED854C3E5A9FAEF9B1FE06F9A0" box="[405,503,1542,1568]" captionStart="FIGURES 89" captionStartId="4.[151,264,1096,1120]" captionTargetBox="[158,1430,181,1072]" captionTargetId="figure-21@4.[158,1430,181,1072]" captionTargetPageId="4" captionText="FIGURES 89. Leptomastax minutissima sp. n., holotype male. Pterothorax and base of abdomen in ventral view (8); right mesocoxa and surrounding areas in ventral view (9)." figureDoi="http://doi.org/10.5281/zenodo.12758932" httpUri="https://zenodo.org/record/12758932/files/figure.png" pageId="2" pageNumber="393">Figs 89</figureCitation>
) with scale-like setae filling postmesocoxal foveae and with elongate flattened setae evenly distributed on remaining surface. Posterior third of metaventrite with median U-shaped impression.
</paragraph>
<paragraph id="923AE468854C3E5A9EFCF9F9FBE8F978" blockId="2.[151,1437,1506,2036]" pageId="2" pageNumber="393">
Elytra (
<figureCitation id="0ABEF8ED854C3E5A9F20F9F9FEAEF9E8" box="[283,351,1614,1640]" captionStart="FIGURES 14" captionStartId="2.[151,264,1363,1387]" captionTargetBox="[151,1435,181,1340]" captionTargetId="figure-21@2.[151,1435,181,1340]" captionTargetPageId="2" captionText="FIGURES 14. Leptomastax minutissima sp. n., holotype male. Dorsal habitus (1); head, pronotum and elytral base in dorsal view (2); left antenna in dorsal view (3); elytra in dorsal view (4)." figureDoi="http://doi.org/10.5281/zenodo.12758926" httpUri="https://zenodo.org/record/12758926/files/figure.png" pageId="2" pageNumber="393">Fig. 4</figureCitation>
) together oval and strongly elongate, broadest near middle; EL
<quantity id="557D498D854C3E5A9A10F9F9FB63F9E8" box="[1067,1170,1614,1640]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="2" pageNumber="393" unit="mm" value="0.68">0.68 mm</quantity>
, EW
<quantity id="557D498D854C3E5A9AEBF9F9FAC7F9E8" box="[1232,1334,1614,1641]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="393" unit="mm" value="0.35">0.35 mm</quantity>
, EI 1.93; humeral region strongly narrowing towards prothorax; lateral margins weakly rounded; apices separately rounded. Each elytron with sutural and humeral row of punctures, lacking subhumeral and marginal rows; punctures distinct but unevenly distributed, so that rows are somewhat irregular. Macrosetae (and setiferous punctures) in humeral and apical regions lacking. Elytra asetose and outside rows of punctures impunctate.
</paragraph>
<paragraph id="923AE468854C3E5A9EFCF8B5FB0AF89C" blockId="2.[151,1437,1506,2036]" box="[199,1275,1794,1820]" pageId="2" pageNumber="393">
Legs (
<figureCitation id="0ABEF8ED854C3E5A9F37F8B5FEA2F89C" box="[268,339,1794,1820]" captionStart="FIGURES 14" captionStartId="2.[151,264,1363,1387]" captionTargetBox="[151,1435,181,1340]" captionTargetId="figure-21@2.[151,1435,181,1340]" captionTargetPageId="2" captionText="FIGURES 14. Leptomastax minutissima sp. n., holotype male. Dorsal habitus (1); head, pronotum and elytral base in dorsal view (2); left antenna in dorsal view (3); elytra in dorsal view (4)." figureDoi="http://doi.org/10.5281/zenodo.12758926" httpUri="https://zenodo.org/record/12758926/files/figure.png" pageId="2" pageNumber="393">Figs 1</figureCitation>
,
<figureCitation id="0ABEF8ED854C3E5A9F5AF8B5FE81F89C" box="[353,368,1794,1820]" captionStart="FIGURES 89" captionStartId="4.[151,264,1096,1120]" captionTargetBox="[158,1430,181,1072]" captionTargetId="figure-21@4.[158,1430,181,1072]" captionTargetPageId="4" captionText="FIGURES 89. Leptomastax minutissima sp. n., holotype male. Pterothorax and base of abdomen in ventral view (8); right mesocoxa and surrounding areas in ventral view (9)." figureDoi="http://doi.org/10.5281/zenodo.12758932" httpUri="https://zenodo.org/record/12758932/files/figure.png" pageId="2" pageNumber="393">9</figureCitation>
) moderately long and slender, unmodified; each mesocoxa with two macrosetae.
</paragraph>
<paragraph id="923AE468854C3E5A9EFCF891FBACF8C0" blockId="2.[151,1437,1506,2036]" box="[199,1117,1830,1857]" pageId="2" pageNumber="393">
Abdomen (
<figureCitation id="0ABEF8ED854C3E5A9F7EF891FE76F8C0" box="[325,391,1830,1856]" captionStart="FIGURES 89" captionStartId="4.[151,264,1096,1120]" captionTargetBox="[158,1430,181,1072]" captionTargetId="figure-21@4.[158,1430,181,1072]" captionTargetPageId="4" captionText="FIGURES 89. Leptomastax minutissima sp. n., holotype male. Pterothorax and base of abdomen in ventral view (8); right mesocoxa and surrounding areas in ventral view (9)." figureDoi="http://doi.org/10.5281/zenodo.12758932" httpUri="https://zenodo.org/record/12758932/files/figure.png" pageId="2" pageNumber="393">Fig. 8</figureCitation>
) with regular setae almost evenly covering all exposed sternites.
</paragraph>
<paragraph id="923AE468854C3E5A9EFCF8FDFD78F850" blockId="2.[151,1437,1506,2036]" pageId="2" pageNumber="393">
Aedeagus (
<figureCitation id="0ABEF8ED854C3E5A9F72F8FDFE3FF8E4" box="[329,462,1866,1892]" captionStart="FIGURES 1013" captionStartId="4.[151,264,2013,2037]" captionTargetBox="[159,1429,1187,1983]" captionTargetId="figure-54@4.[151,1436,1180,1990]" captionTargetPageId="4" captionText="FIGURES 1013. Leptomastax minutissima sp. n., holotype male. Aedeagus in ventral (10, 12) and lateral (11, 13) views." figureDoi="http://doi.org/10.5281/zenodo.12758934" httpUri="https://zenodo.org/record/12758934/files/figure.png" pageId="2" pageNumber="393">Figs 1013</figureCitation>
) slender, AeL
<quantity id="557D498D854C3E5A9C45F8FDFD1BF8E5" box="[638,746,1866,1893]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="2" pageNumber="393" unit="mm" value="0.18">0.18 mm</quantity>
; median lobe in ventral view gradually narrowing towards subtriangular apex, in lateral view distal region nearly straight; sclerotized endophallus unipartite, tubular and unbranched, nearly straight; parameres with evenly rounded outer margins in subapical and apical regions, apex of each paramere pointed and directed mesally.
</paragraph>
<paragraph id="923AE468854C3E5A9EFCF86DFE6FF874" blockId="2.[151,1437,1506,2036]" box="[199,414,2010,2036]" pageId="2" pageNumber="393">Female. Unknown.</paragraph>
</subSubSection>
<caption id="C6FAB4E0854D3E5B9EACFB15FC67FB5F" ID-DOI="http://doi.org/10.5281/zenodo.12758930" ID-Zenodo-Dep="12758930" httpUri="https://zenodo.org/record/12758930/files/figure.png" pageId="3" pageNumber="394" startId="3.[151,264,1186,1210]" targetBox="[151,1435,181,1162]" targetPageId="3" targetType="figure">
<paragraph id="923AE468854D3E5B9EACFB15FC67FB5F" blockId="3.[151,1436,1186,1247]" pageId="3" pageNumber="394">
<emphasis id="A0F1387A854D3E5B9EACFB15FECAFB3B" bold="true" box="[151,315,1186,1211]" pageId="3" pageNumber="394">FIGURES 57.</emphasis>
<taxonomicName id="55859FEB854D3E5B9F79FB14FDB5FB3A" authorityName="Jałoszyński" authorityYear="2024" box="[322,580,1187,1210]" class="Insecta" family="Staphylinidae" genus="Leptomastax" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="minutissima" status="sp. nov.">
<emphasis id="A0F1387A854D3E5B9F79FB14FDB5FB3A" box="[322,580,1187,1210]" italics="true" pageId="3" pageNumber="394">Leptomastax minutissima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBC28501854D3E5B9C77FB14FD75FB3A" box="[588,644,1187,1210]" pageId="3" pageNumber="394" rank="species">sp. n.</taxonomicNameLabel>
, holotype male. Head, prothorax and anterior region of mesothorax in ventral view (
<emphasis id="A0F1387A854D3E5B9EECFB70FF15FB5F" bold="true" box="[215,228,1223,1247]" pageId="3" pageNumber="394">5</emphasis>
); mouthparts in ventral view (
<emphasis id="A0F1387A854D3E5B9C22FB71FDD7FB5E" bold="true" box="[537,550,1222,1246]" pageId="3" pageNumber="394">6</emphasis>
); left labial palp in ventral view (
<emphasis id="A0F1387A854D3E5B9D40FB70FC79FB5F" bold="true" box="[891,904,1223,1247]" pageId="3" pageNumber="394">7</emphasis>
).
</paragraph>
</caption>
<subSubSection id="DA9FB7E3854D3E5B9EFCFA87FDE8FACB" box="[199,537,1328,1355]" pageId="3" pageNumber="394" type="distribution">
<paragraph id="923AE468854D3E5B9EFCFA87FDE8FACB" blockId="3.[151,1437,1328,1787]" box="[199,537,1328,1355]" pageId="3" pageNumber="394">
<emphasis id="A0F1387A854D3E5B9EFCFA87FE90FACA" bold="true" box="[199,353,1328,1354]" pageId="3" pageNumber="394">Distribution.</emphasis>
Northern
<collectingCountry id="EA92A4F8854D3E5B9FEFFA86FDE7FACB" box="[468,534,1329,1355]" name="Israel" pageId="3" pageNumber="394">Israel</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DA9FB7E3854D3E5B9EFCFAE2FEDCFA13" pageId="3" pageNumber="394" type="etymology">
<paragraph id="923AE468854D3E5B9EFCFAE2FEDCFA13" blockId="3.[151,1437,1328,1787]" pageId="3" pageNumber="394">
<emphasis id="A0F1387A854D3E5B9EFCFAE2FEBFFAEF" bold="true" box="[199,334,1365,1391]" pageId="3" pageNumber="394">Etymology.</emphasis>
The specific adjective
<taxonomicName id="55859FEB854D3E5B9C52FAE2FD01FAEE" authorityName="Jałoszyński" authorityYear="2024" box="[617,752,1365,1390]" class="Insecta" family="Staphylinidae" genus="Leptomastax" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="minutissima">
<emphasis id="A0F1387A854D3E5B9C52FAE2FD01FAEE" box="[617,752,1365,1390]" italics="true" pageId="3" pageNumber="394">minutissima</emphasis>
</taxonomicName>
refers to the smallest body among all known species of
<taxonomicName id="55859FEB854D3E5B9EACFACEFED7FA12" authorityName="Pirazzoli" authorityYear="1855" box="[151,294,1401,1426]" class="Insecta" family="Staphylinidae" genus="Leptomastax" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="A0F1387A854D3E5B9EACFACEFED7FA12" box="[151,294,1401,1426]" italics="true" pageId="3" pageNumber="394">Leptomastax</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DA9FB7E3854D3E5B9EFCFA2AFE56F97B" pageId="3" pageNumber="394" type="discussion">
<paragraph id="923AE468854D3E5B9EFCFA2AFCE7F90F" blockId="3.[151,1437,1328,1787]" pageId="3" pageNumber="394">
<emphasis id="A0F1387A854D3E5B9EFCFA2AFECAFA37" bold="true" box="[199,315,1437,1463]" pageId="3" pageNumber="394">Remarks.</emphasis>
This is the smallest known species of
<taxonomicName id="55859FEB854D3E5B9CDAFA2AFC81FA36" authorityName="Pirazzoli" authorityYear="1855" box="[737,880,1437,1462]" class="Insecta" family="Staphylinidae" genus="Leptomastax" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="genus">
<emphasis id="A0F1387A854D3E5B9CDAFA2AFC81FA36" box="[737,880,1437,1462]" italics="true" pageId="3" pageNumber="394">Leptomastax</emphasis>
</taxonomicName>
; it can be easily identified by the slender, strongly elongate elytra, reduced and irregular elytral rows of punctures, lack of eyes, and aedeagal structures. The simple, unbranched endophallus in
<taxonomicName id="55859FEB854D3E5B9FF0FA52FD9EFA7E" authorityName="Jałoszyński" authorityYear="2024" box="[459,623,1509,1534]" class="Insecta" family="Staphylinidae" genus="Leptomastax" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="minutissima">
<emphasis id="A0F1387A854D3E5B9FF0FA52FD9EFA7E" box="[459,623,1509,1534]" italics="true" pageId="3" pageNumber="394">L. minutissima</emphasis>
</taxonomicName>
resembles that of
<taxonomicName id="55859FEB854D3E5B9D00FA52FBEEFA7F" authority="Meybohm" authorityName="Meybohm" authorityYear="2019" box="[827,1055,1509,1535]" class="Insecta" family="Staphylinidae" genus="Leptomastax" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="cretica">
<emphasis id="A0F1387A854D3E5B9D00FA52FC57FA7E" box="[827,934,1509,1534]" italics="true" pageId="3" pageNumber="394">L. cretica</emphasis>
Meybohm
</taxonomicName>
(in
<bibRefCitation id="F6149999854D3E5B9A71FA52FADBFA7F" author="Assing, V. &amp; Brachat, V. &amp; Meybohm, H." box="[1098,1322,1509,1535]" pageId="3" pageNumber="394" pagination="239 - 289" refId="ref2642" refString="Assing, V., Brachat, V. &amp; Meybohm, H. (2019) Monograph of the Staphylinidae of Crete (Greece). Part II. Descriptions of new species (Insecta: Coleoptera). Contributions to Entomology, 69 (2), 239 - 289. https: // doi. org / 10.21248 / contrib. entomol. 69.2.239 - 289" type="journal article" year="2019">
Assing
<emphasis id="A0F1387A854D3E5B9AA5FA52FB27FA7F" box="[1182,1238,1509,1535]" italics="true" pageId="3" pageNumber="394">et al.</emphasis>
(2019)
</bibRefCitation>
, fig. 149), but the latter species is much larger (BL
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) and has large eyes, each composed of “about 14 ommatidia” (
<bibRefCitation id="F6149999854D3E5B9EA4F99AFE80F9C7" author="Assing, V. &amp; Brachat, V. &amp; Meybohm, H." box="[159,369,1581,1607]" pageId="3" pageNumber="394" pagination="239 - 289" refId="ref2642" refString="Assing, V., Brachat, V. &amp; Meybohm, H. (2019) Monograph of the Staphylinidae of Crete (Greece). Part II. Descriptions of new species (Insecta: Coleoptera). Contributions to Entomology, 69 (2), 239 - 289. https: // doi. org / 10.21248 / contrib. entomol. 69.2.239 - 289" type="journal article" year="2019">
Assing
<emphasis id="A0F1387A854D3E5B9ECEF99AFEDCF9C7" box="[245,301,1581,1607]" italics="true" pageId="3" pageNumber="394">et al.</emphasis>
2019
</bibRefCitation>
). The apical region of median lobe and the endophallus in
<taxonomicName id="55859FEB854D3E5B9DC4F99AFB9BF9C6" authorityName="Meybohm" authorityYear="2019" box="[1023,1130,1581,1606]" class="Insecta" family="Staphylinidae" genus="Leptomastax" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="cretica">
<emphasis id="A0F1387A854D3E5B9DC4F99AFB9BF9C6" box="[1023,1130,1581,1606]" italics="true" pageId="3" pageNumber="394">L. cretica</emphasis>
</taxonomicName>
in lateral view are strongly curved (nearly straight in
<taxonomicName id="55859FEB854D3E5B9F84F9E6FD94F9EA" authorityName="Jałoszyński" authorityYear="2024" box="[447,613,1617,1642]" class="Insecta" family="Staphylinidae" genus="Leptomastax" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="minutissima">
<emphasis id="A0F1387A854D3E5B9F84F9E6FD94F9EA" box="[447,613,1617,1642]" italics="true" pageId="3" pageNumber="394">L. minutissima</emphasis>
</taxonomicName>
), and the parameral apices in ventral view in
<taxonomicName id="55859FEB854D3E5B9A4EF9E6FB10F9EA" authorityName="Meybohm" authorityYear="2019" box="[1141,1249,1617,1642]" class="Insecta" family="Staphylinidae" genus="Leptomastax" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="cretica">
<emphasis id="A0F1387A854D3E5B9A4EF9E6FB10F9EA" box="[1141,1249,1617,1642]" italics="true" pageId="3" pageNumber="394">L. cretica</emphasis>
</taxonomicName>
are rapidly bent mesally (evenly and gradually curved in
<taxonomicName id="55859FEB854D3E5B9C5BF9C2FCF4F90E" authorityName="Jałoszyński" authorityYear="2024" box="[608,773,1653,1678]" class="Insecta" family="Staphylinidae" genus="Leptomastax" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="394" phylum="Arthropoda" rank="species" species="minutissima">
<emphasis id="A0F1387A854D3E5B9C5BF9C2FCF4F90E" box="[608,773,1653,1678]" italics="true" pageId="3" pageNumber="394">L. minutissima</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="923AE468854D3E5B9EFCF92EFE56F97B" blockId="3.[151,1437,1328,1787]" pageId="3" pageNumber="394">
Notes containing detailed collecting data for the studied specimen have been lost; is is only known that most specimens taken during the Löbl-Besuchet 1982 fieldwork in Eilon come from sifted oak leaf litter (email from Ivan Löbl dated
<date id="E63BC2A8854D3E5B9F2CF956FE6DF97B" box="[279,412,1761,1787]" pageId="3" pageNumber="394" value="2024-06-18">18/06/2024</date>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="3D671FE6F665E3064399CDD720883DCB">Description of Neosilphitrombium rhipicephalum sp. nov. (Prostigmata: Neothrombiidae) ectoparasite on hard ticks from Iran</mods:title>
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Genus
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Diagnosis of larva (According to
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Saboori
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Neothrombiids with eyes 2 + 2; two dorsal scuta, AL setae bilobed and posterior or posteromedially to AM setae, sensilla posteromedially to PL setae, scutellum with two or more normal setae; coxal setal formula 3-4-2, all coxal setae bifid except lateral seta on coxa II (
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2b
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<emphasis id="1C97462D256A163142FEFE32FE439F3F" box="[151,391,491,515]" italics="true" pageId="1" pageNumber="385">Geodromicus perplexus</emphasis>
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</taxonomicName>
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.
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</paragraph>
<paragraph id="2E5C9A3F256A163142FEFDD3FD819F1E" blockId="1.[151,927,460,607]" box="[151,581,522,546]" pageId="1" pageNumber="385">
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<treatmentCitation id="AF42BC2E256A163142FEFDD3FD859F1E" author="Coiffait, H." box="[151,577,522,546]" page="271" pageId="1" pageNumber="385" year="1977">
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<emphasis id="1C97462D256A163142FEFDD3FEB39F1E" box="[151,375,522,546]" italics="true" pageId="1" pageNumber="385">Geodromicus palpalis</emphasis>
<bibRefCitation id="4A72E7CE256A16314314FDD3FD859F1E" author="Coiffait, H." box="[381,577,522,546]" pageId="1" pageNumber="385" pagination="243 - 275" refId="ref3399" refString="Coiffait, H. (1977) Staphilinides recoltes au Nepal par le Professeur Franz (2 e partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse, 112 (1976), 243 - 275." type="journal article" year="1977">Coiffait, 1977: 271</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="2E5C9A3F256A163142FEFDF0FDAE9F7D" blockId="1.[151,927,460,607]" box="[151,618,552,577]" pageId="1" pageNumber="385">
<treatmentCitationGroup id="0EF3BD11256A163142FEFDF0FDAE9F7D" box="[151,618,552,577]" pageId="1" pageNumber="385">
<treatmentCitation id="AF42BC2E256A163142FEFDF0FDDA9F7D" author="Coiffait, H." box="[151,542,552,577]" page="282" pageId="1" pageNumber="385" year="1983">
<taxonomicName id="E9E3E1BC256A163142FEFDF0FDDA9F7D" ID-CoL="3NBJM" authority="Coiffait, 1983: 282" authorityName="Coiffait" authorityPageNumber="282" authorityYear="1983" box="[151,542,552,577]" class="Insecta" family="Staphylinidae" genus="Hygrogeus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="ledouxi" status="syn. nov.">
<emphasis id="1C97462D256A163142FEFDF0FE939F7C" box="[151,343,552,576]" italics="true" pageId="1" pageNumber="385">Hygrogeus ledouxi</emphasis>
<bibRefCitation id="4A72E7CE256A16314334FDF1FDDA9F7D" author="Coiffait, H." box="[349,542,552,577]" pageId="1" pageNumber="385" pagination="273 - 289" refId="ref3438" refString="Coiffait, H. (1983) Nouveaux staphylinides du Cachemire (Coleoptera). Annales de la Societe entomologique de France, New Series, 19 (3), 273 - 289. https: // doi. org / 10.1080 / 21686351.1983.12278365" type="journal article" year="1983">Coiffait, 1983: 282</bibRefCitation>
</taxonomicName>
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<emphasis id="1C97462D256A1631404DFDF0FDAE9F7D" bold="true" box="[548,618,553,577]" pageId="1" pageNumber="385">
<taxonomicNameLabel id="07A4FB56256A1631404DFDF0FDAE9F7D" box="[548,618,553,577]" pageId="1" pageNumber="385">syn. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="2E5C9A3F256A163142FEFD9EFC5B9F63" blockId="1.[151,927,460,607]" box="[151,927,583,607]" pageId="1" pageNumber="385">
<treatmentCitationGroup id="0EF3BD11256A163142FEFD9EFC5B9F63" box="[151,927,583,607]" pageId="1" pageNumber="385">
<taxonomicName id="E9E3E1BC256A163142FEFD9EFE599F63" ID-CoL="3NBHT" baseAuthorityName="Cameron" baseAuthorityYear="1924" box="[151,413,583,607]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="1C97462D256A163142FEFD9EFE599F63" box="[151,413,583,607]" italics="true" pageId="1" pageNumber="385">Hygrodromicus coriaceus</emphasis>
</taxonomicName>
:
<treatmentCitation id="AF42BC2E256A163143C2FD9EFD9B9F63" author="Zerche, L." box="[427,607,583,607]" page="120" pageId="1" pageNumber="385" year="1992">
<bibRefCitation id="4A72E7CE256A163143C2FD9EFD9B9F63" author="Zerche, L." box="[427,607,583,607]" pageId="1" pageNumber="385" pagination="105 - 142" refId="ref3633" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche 1992: 120</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="AF42BC2E256A16314003FD9EFCE39F63" author="Shavrin, A. V." box="[618,807,583,607]" page="338" pageId="1" pageNumber="385" year="2021">
<bibRefCitation id="4A72E7CE256A16314003FD9EFCE39F63" author="Shavrin, A. V." box="[618,807,583,607]" pageId="1" pageNumber="385" pagination="335 - 348" refId="ref3483" refString="Shavrin, A. V. (2021) New taxonomic and faunistic data of the genus Hygrodromicus Tronquet, 1981 of the Himalayan Region (Coleoptera: Staphylinidaae: Omaliinae: Anthophagini). In: Hartmann, M., Barclay, M. &amp; Weipert, J. (Eds.), Biodiversitat und Naturausstattung im Himalaya VII. Verein der Freunde und Forderer des Naturkundmuseums Erfurt e. V., Erfurt, pp. 335 - 348." type="book chapter" year="2021">Shavrin 2021: 338</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="AF42BC2E256A1631415BFD9EFC5F9F63" ID-DOI="http://doi.org/10.5281/zenodo.12758936" ID-Zenodo-Dep="12758936" author="Shavrin, A. V." box="[818,923,583,607]" httpUri="http://treatment.plazi.org/id/A64A2B29256A163142FEFE5EFD669E2F" page="237" pageId="1" pageNumber="385" year="2024">
<bibRefCitation id="4A72E7CE256A1631415BFD9EFC5F9F63" author="Shavrin, A. V." box="[818,923,583,607]" pageId="1" pageNumber="385" pagination="236 - 246" refId="ref3567" refString="Shavrin, A. V. (2024) New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa, 5446 (2), 236 - 246. https: // doi. org / 10.11646 / zootaxa. 5446.2.5" type="journal article" year="2024">2024: 237</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="66F9C9B4256A163142FEFD54FD669E2F" pageId="1" pageNumber="385" type="discussion">
<paragraph id="2E5C9A3F256A163142FEFD54FD669E2F" blockId="1.[151,1437,653,787]" pageId="1" pageNumber="385">
<emphasis id="1C97462D256A163142FEFD54FECF9F9B" bold="true" box="[151,267,653,679]" pageId="1" pageNumber="385">Remarks.</emphasis>
<taxonomicName id="E9E3E1BC256A1631437AFD57FE239F9B" authorityName="Coiffait" authorityYear="1983" box="[275,487,653,679]" class="Insecta" family="Staphylinidae" genus="Hygrogeus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="1C97462D256A1631437AFD57FE239F9B" box="[275,487,653,679]" italics="true" pageId="1" pageNumber="385">Hygrogeus ledouxi</emphasis>
</taxonomicName>
based on the single
<typeStatus id="F158249D256A163140A6FD54FCF59F9B" box="[719,817,653,679]" pageId="1" pageNumber="385" type="holotype">holotype</typeStatus>
was originally described from “Cachemire, Pahalgam, Sheshnag…”. Based on the original description and the shape of the aedeagus (
<figureCitation id="B6D886BA256A16314679FD68FB749FF7" box="[1040,1200,689,715]" captionStart="FIGURES 12" captionStartId="2.[151,264,1207,1231]" captionTargetBox="[157,1430,281,1175]" captionTargetId="figure-60@2.[151,1436,270,1181]" captionTargetPageId="2" captionText="FIGURES 12. Habitus of Hygrodromicus: 1H. incrassatus (paratype), 2H. splendidus (Kazakhstan, Ketmen` Mts.). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12758946" httpUri="https://zenodo.org/record/12758946/files/figure.png" pageId="1" pageNumber="385">Figs 2I and 2J</figureCitation>
in
<bibRefCitation id="4A72E7CE256A163146BDFD68FAA99FF7" author="Coiffait, H." box="[1236,1389,689,715]" pageId="1" pageNumber="385" pagination="273 - 289" refId="ref3438" refString="Coiffait, H. (1983) Nouveaux staphylinides du Cachemire (Coleoptera). Annales de la Societe entomologique de France, New Series, 19 (3), 273 - 289. https: // doi. org / 10.1080 / 21686351.1983.12278365" type="journal article" year="1983">Coiffait 1983</bibRefCitation>
),
<taxonomicName id="E9E3E1BC256A163147E8FD6BFF2D9FD3" authorityName="Coiffait" authorityYear="1983" class="Insecta" family="Staphylinidae" genus="Hygrogeus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="1C97462D256A163147E8FD6BFF2D9FD3" italics="true" pageId="1" pageNumber="385">H. ledouxi</emphasis>
</taxonomicName>
is conspecific with the Himalayan
<taxonomicName id="E9E3E1BC256A1631401CFD0FFCC09FD3" baseAuthorityName="Cameron" baseAuthorityYear="1924" box="[629,772,726,751]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="coriaceus">
<emphasis id="1C97462D256A1631401CFD0FFCC09FD3" box="[629,772,726,751]" italics="true" pageId="1" pageNumber="385">H. coriaceus</emphasis>
</taxonomicName>
, also known from Kashmir (
<bibRefCitation id="4A72E7CE256A1631462FFD0CFB259FD3" author="Shavrin, A. V." box="[1094,1249,725,751]" pageId="1" pageNumber="385" pagination="335 - 348" refId="ref3483" refString="Shavrin, A. V. (2021) New taxonomic and faunistic data of the genus Hygrodromicus Tronquet, 1981 of the Himalayan Region (Coleoptera: Staphylinidaae: Omaliinae: Anthophagini). In: Hartmann, M., Barclay, M. &amp; Weipert, J. (Eds.), Biodiversitat und Naturausstattung im Himalaya VII. Verein der Freunde und Forderer des Naturkundmuseums Erfurt e. V., Erfurt, pp. 335 - 348." type="book chapter" year="2021">Shavrin 2021</bibRefCitation>
,
<bibRefCitation id="4A72E7CE256A16314685FD0CFAED9FD3" author="Shavrin, A. V." box="[1260,1321,725,751]" pageId="1" pageNumber="385" pagination="236 - 246" refId="ref3567" refString="Shavrin, A. V. (2024) New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa, 5446 (2), 236 - 246. https: // doi. org / 10.11646 / zootaxa. 5446.2.5" type="journal article" year="2024">2024</bibRefCitation>
). Thus,
<taxonomicName id="E9E3E1BC256A163147E8FD0FFF2D9E2F" authorityName="Coiffait" authorityYear="1983" class="Insecta" family="Staphylinidae" genus="Hygrogeus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="ledouxi">
<emphasis id="1C97462D256A163147E8FD0FFF2D9E2F" italics="true" pageId="1" pageNumber="385">H. ledouxi</emphasis>
</taxonomicName>
is synonymized with the latter species.
</paragraph>
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</treatment>
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View file

@ -0,0 +1,300 @@
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<emphasis id="1C97462D256A163142FEFC83FDEA9E49" bold="true" box="[151,558,858,885]" pageId="1" pageNumber="385">
<taxonomicName id="E9E3E1BC256A163142FEFC83FE219E49" authorityName="Shavrin" authorityYear="2024" box="[151,485,858,885]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="incrassatus" status="sp. nov.">
<emphasis id="1C97462D256A163142FEFC83FE219E49" bold="true" box="[151,485,858,885]" italics="true" pageId="1" pageNumber="385">Hygrodromicus incrassatus</emphasis>
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<taxonomicNameLabel id="07A4FB56256A16314385FC82FDEA9E49" box="[492,558,859,885]" pageId="1" pageNumber="385" rank="species">sp. n.</taxonomicNameLabel>
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(
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,
<figureCitation id="B6D886BA256A1631429DFCA6FEE49EA5" box="[244,288,895,921]" captionStart="FIGURES 36" captionStartId="3.[151,264,1842,1866]" captionTargetBox="[177,1417,713,1811]" captionTargetId="figure-228@3.[151,1436,705,1819]" captionTargetPageId="3" captionText="FIGURES 36. Aedeagus of Hygrodromicus (Figs 34: H. incrassatus (holotype); Figs. 56: H. splendidus (Kazakhstan: Ketmen` Mts.): 3, 5parameral view, 4, 6lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12758950" httpUri="https://zenodo.org/record/12758950/files/figure.png" pageId="1" pageNumber="385">34</figureCitation>
,
<figureCitation id="B6D886BA256A16314345FCA6FEFF9EA5" box="[300,315,895,921]" captionStart="FIGURE 7" captionStartId="4.[151,250,1743,1767]" captionTargetBox="[151,1435,664,1720]" captionTargetId="figure-160@4.[151,1435,664,1720]" captionTargetPageId="4" captionText="FIGURE 7. Distribution of Hygrodromicus incrassatus (square) and H. splendidus (circles)." figureDoi="http://doi.org/10.5281/zenodo.12758954" httpUri="https://zenodo.org/record/12758954/files/figure.png" pageId="1" pageNumber="385">7</figureCitation>
)
</paragraph>
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<emphasis id="1C97462D256A163142FEFC1EFE039EDC" bold="true" box="[151,455,966,993]" pageId="1" pageNumber="385">Type material examined.</emphasis>
<typeStatus id="F158249D256A163143BBFC1EFDFE9EDD" box="[466,570,967,993]" pageId="1" pageNumber="385" type="holotype">Holotype</typeStatus>
<specimenCount id="38E551B6256A1631402CFC1EFD9E9EDD" box="[581,602,967,993]" count="1" pageId="1" pageNumber="385" type="male"></specimenCount>
(a plastic plate with the aedeagus in Canada balsam was pinned under the card with the beetle; abdominal tergite VIII, sternite VIII, and the apical segment are glued on the same card under the specimen):
<collectingCountry id="56F4DAAF256A163143F9FBD6FDDB9915" box="[400,543,1039,1065]" name="Pakistan" pageId="1" pageNumber="385">PAKISTAN</collectingCountry>
: NW Frontier | province, Kaghan valley, nr. | Sighban,
<quantity id="E91B37DA256A163146F9FBD6FB2C9915" box="[1168,1256,1039,1065]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="385" unit="m" value="2200.0">2200 m</quantity>
, |
<date id="5A5DBCFF256A1631476AFBD6FA5C9915" box="[1283,1432,1039,1065]" pageId="1" pageNumber="385" value="2004-08-20">20.VIII.2004</date>
, leg.
<collectionCode id="48F202FA256A163142A0FBEAFF199971" box="[201,221,1075,1101]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="385" type="Museum">V</collectionCode>
. Gurko &lt;printed&gt;, Museum für Naturkunde | Berlin | Sammlung M. Schülke &lt;printed&gt;,
<typeStatus id="F158249D256A1631469DFBEAFA499971" box="[1268,1421,1075,1101]" pageId="1" pageNumber="385" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="E9E3E1BC256A163142FEFB8EFE81994D" authorityName="Tronquet" authorityYear="1981" box="[151,325,1111,1137]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="1C97462D256A163142FEFB8EFE81994D" box="[151,325,1111,1137]" italics="true" pageId="1" pageNumber="385">Hygrodromicus</emphasis>
</taxonomicName>
|
<taxonomicName id="E9E3E1BC256A16314331FB8EFE1C994C" authorityName="Shavrin" authorityYear="2024" box="[344,472,1111,1136]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="incrassatus" status="sp. nov.">
<emphasis id="1C97462D256A16314331FB8EFE1C994C" box="[344,472,1111,1136]" italics="true" pageId="1" pageNumber="385">incrassatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07A4FB56256A163143B6FB8EFDDF994D" box="[479,539,1111,1137]" pageId="1" pageNumber="385" rank="species">sp. n.</taxonomicNameLabel>
| Shavrin A.
<collectionCode id="48F202FA256A163140CCFB8EFD7F994D" box="[677,699,1111,1137]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="385" type="Museum">V</collectionCode>
. des. 2024 &lt;red, printed&gt; (cSch).
</paragraph>
<paragraph id="2E5C9A3F256A163142AEFBA2FACE9985" blockId="1.[151,1437,966,2037]" pageId="1" pageNumber="385">
<typeStatus id="F158249D256A163142AEFBA2FEFC99A9" box="[199,312,1147,1173]" pageId="1" pageNumber="385" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="38E551B6256A1631432DFBA2FE4099A9" box="[324,388,1147,1173]" count="2" pageId="1" pageNumber="385" type="female">2 ♀♀</specimenCount>
(
<figureCitation id="B6D886BA256A163143FDFBA2FE1C99A9" box="[404,472,1147,1173]" captionStart="FIGURES 12" captionStartId="2.[151,264,1207,1231]" captionTargetBox="[157,1430,281,1175]" captionTargetId="figure-60@2.[151,1436,270,1181]" captionTargetPageId="2" captionText="FIGURES 12. Habitus of Hygrodromicus: 1H. incrassatus (paratype), 2H. splendidus (Kazakhstan, Ketmen` Mts.). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12758946" httpUri="https://zenodo.org/record/12758946/files/figure.png" pageId="1" pageNumber="385">Fig. 1</figureCitation>
;
<specimenCount id="38E551B6256A1631438EFBA2FD4699A9" box="[487,642,1147,1173]" count="1" pageId="1" pageNumber="385" type="generic">one specimen</specimenCount>
without right fore leg): same data as the holotype, with additional red printed label:
<typeStatus id="F158249D256A1631435EFB46FE039985" box="[311,455,1183,1209]" pageId="1" pageNumber="385" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="E9E3E1BC256A163143B2FB46FD4D9985" authorityName="Tronquet" authorityYear="1981" box="[475,649,1183,1209]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="1C97462D256A163143B2FB46FD4D9985" box="[475,649,1183,1209]" italics="true" pageId="1" pageNumber="385">Hygrodromicus</emphasis>
</taxonomicName>
|
<taxonomicName id="E9E3E1BC256A163140F5FB46FCD89984" authorityName="Shavrin" authorityYear="2024" box="[668,796,1183,1208]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="385" phylum="Arthropoda" rank="species" species="incrassatus" status="sp. nov.">
<emphasis id="1C97462D256A163140F5FB46FCD89984" box="[668,796,1183,1208]" italics="true" pageId="1" pageNumber="385">incrassatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="07A4FB56256A1631414BFB46FC9A9985" box="[802,862,1183,1209]" pageId="1" pageNumber="385" rank="species">sp. n.</taxonomicNameLabel>
| Shavrin A.
<collectionCode id="48F202FA256A16314181FB46FC3A9985" box="[1000,1022,1183,1209]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="385" type="Museum">V</collectionCode>
. des. 2024 (cSch, cSh).
</paragraph>
</subSubSection>
<subSubSection id="66F9C9B4256A163342AEFB1BFEC19CED" lastPageId="3" lastPageNumber="387" pageId="1" pageNumber="385" type="description">
<paragraph id="2E5C9A3F256A163142AEFB1BFE0E9875" blockId="1.[151,1437,966,2037]" pageId="1" pageNumber="385">
<emphasis id="1C97462D256A163142AEFB1BFE9D99E0" bold="true" box="[199,345,1218,1244]" pageId="1" pageNumber="385">Description.</emphasis>
Measurements (n = 3): HW: 1.051.10; HL: 0.700.73; OL: 0.120.14; LT: 0.100.13; AL (
<typeStatus id="F158249D256A163142F6FB3EFECC983D" box="[159,264,1255,1281]" pageId="1" pageNumber="385" type="holotype">holotype</typeStatus>
): 4.15; PL: 0.921.05; PWmax: 1.261.45; PWmin: 1.031.20; ESL: 1.521.85; EW: 1.942.30; MTbL(
<typeStatus id="F158249D256A16314285FAD2FE969819" box="[236,338,1291,1317]" pageId="1" pageNumber="385" type="holotype">holotype</typeStatus>
): 1.55; MTrL (
<typeStatus id="F158249D256A1631439EFAD2FDA49819" box="[503,608,1291,1317]" pageId="1" pageNumber="385" type="holotype">holotype</typeStatus>
): 0.57 (MTrL 14: 0.32; MTrL 5: 0.25); AW: 2.242.40; AedL: 1.25; BL: 5.906.75 (
<typeStatus id="F158249D256A1631437FFAF6FEBA9875" box="[278,382,1327,1353]" pageId="1" pageNumber="385" type="holotype">holotype</typeStatus>
: 5.92).
</paragraph>
<paragraph id="2E5C9A3F256A163142AEFA8AFAB398C1" blockId="1.[151,1437,966,2037]" pageId="1" pageNumber="385">
Habitus as in
<figureCitation id="B6D886BA256A1631430CFA8AFE6C9851" box="[357,424,1363,1389]" captionStart="FIGURES 12" captionStartId="2.[151,264,1207,1231]" captionTargetBox="[157,1430,281,1175]" captionTargetId="figure-60@2.[151,1436,270,1181]" captionTargetPageId="2" captionText="FIGURES 12. Habitus of Hygrodromicus: 1H. incrassatus (paratype), 2H. splendidus (Kazakhstan, Ketmen` Mts.). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12758946" httpUri="https://zenodo.org/record/12758946/files/figure.png" pageId="1" pageNumber="385">Fig. 1</figureCitation>
. Body reddish-brown (elytra of
<typeStatus id="F158249D256A16314170FA8AFCBF9851" box="[793,891,1363,1389]" pageId="1" pageNumber="385" type="holotype">holotype</typeStatus>
distinctly paler); mouthparts, antennae and legs brownish; apical segment of maxillary palpi and tarsi yellowish. Head with regular, dense and fine isodiametric microreticulation, coarser in middle portion; neck with isodiametric sculpture, denser and coarser than that in middle portion of head; pronotum with dense isodiametric microsculpture similar to that in head, finer in middle; scutellum with dense transverse meshes in middle; abdomen with dense and moderately fine transverse microreticulation.
</paragraph>
<paragraph id="2E5C9A3F256A163142AEF9DEFE279A91" blockId="1.[151,1437,966,2037]" pageId="1" pageNumber="385">Head 1.5 times as broad as long, with markedly elevated portion between anterior margins of eyes and infraorbital portions; supra-antennal protuberances moderately wide and distinctly elevated; medioapical depression deep and wide, narrowing posteriad and indistinctly connected with moderately deep subrectangular interocellar depression; anteocellar foveae narrow and long, deep, convergent diagonally toward level of anterior margins of eyes; temples convex and slightly shorter than longitudinal length of eyes. Eyes moderately small, strongly convex. Punctation dense and moderately fine, sparser and finer in middle. Ocelli small, located distinctly below level of posterior margins of eyes, distance between ocelli slightly longer than distance between ocellus and posterior margin of eyes. Apical segment of maxillary palpi short, about three times shorter than preapical segment. Antenna long, almost reaching apical margins of elytra when reclined; basal antennomere distinctly more than three times as long as broad, antennomere 2 narrower and significantly shorter than basal antennomere, 3 distinctly longer than 2, 45 slightly longer than 3, 69 indistinctly longer and broader than 5, 10 slightly shorter than 9, apical antennomere about 1.3 times as long as 10.</paragraph>
<paragraph id="2E5C9A3F256A163242AEF86EFC809DC5" blockId="1.[151,1437,966,2037]" lastBlockId="2.[151,1436,151,249]" lastPageId="2" lastPageNumber="386" pageId="1" pageNumber="385">
Pronotum moderately convex, 1.3 times as broad as long, 1.21.3 times as broad as head, widest in anterior portion, strongly narrowed posteriad toward obtuse posterior angles, with indistinctly elevated longitudinal band in middle (
<typeStatus id="F158249D256916324298FF4EFE9D9D8D" box="[241,345,151,177]" pageId="2" pageNumber="386" type="holotype">holotype</typeStatus>
); posterior margin rounded, about as long as posterior margin slightly concave in middle.Punctation about as that in middle portion of head, but distinctly denser, sparser in median part, with narrow impunctate portion in middle. Scutellum with several fine and sparse punctation.
</paragraph>
<caption id="7A9CCAB72569163242FEFB6EFEDE99CF" ID-DOI="http://doi.org/10.5281/zenodo.12758946" ID-Zenodo-Dep="12758946" httpUri="https://zenodo.org/record/12758946/files/figure.png" pageId="2" pageNumber="386" startId="2.[151,264,1207,1231]" targetBox="[157,1430,281,1175]" targetPageId="2" targetType="figure">
<paragraph id="2E5C9A3F2569163242FEFB6EFEDE99CF" blockId="2.[151,1436,1207,1268]" pageId="2" pageNumber="386">
<emphasis id="1C97462D2569163242FEFB6EFEFF99F3" bold="true" box="[151,315,1207,1231]" pageId="2" pageNumber="386">FIGURES 12.</emphasis>
Habitus of
<taxonomicName id="E9E3E1BC2569163243DCFB6EFD9699F3" authorityName="Tronquet" authorityYear="1981" box="[437,594,1207,1231]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="genus">
<emphasis id="1C97462D2569163243DCFB6EFD9699F3" box="[437,594,1207,1231]" italics="true" pageId="2" pageNumber="386">Hygrodromicus</emphasis>
</taxonomicName>
: 1
<taxonomicName id="E9E3E1BC256916324010FB61FCC899F3" authorityName="Shavrin" authorityYear="2024" box="[633,780,1208,1231]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="incrassatus">
<emphasis id="1C97462D256916324010FB61FCC899F3" box="[633,780,1208,1231]" italics="true" pageId="2" pageNumber="386">H. incrassatus</emphasis>
</taxonomicName>
(paratype), 2
<taxonomicName id="E9E3E1BC2569163241CAFB61FBEA99F3" authorityName="Zerche" authorityYear="1992" box="[931,1070,1207,1231]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="1C97462D2569163241CAFB61FBEA99F3" box="[931,1070,1207,1231]" italics="true" pageId="2" pageNumber="386">H. splendidus</emphasis>
</taxonomicName>
(Kazakhstan, Ketmen` Mts.). Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph id="2E5C9A3F2569163242AEFA9FFA9498F0" blockId="2.[151,1437,1350,2024]" pageId="2" pageNumber="386">
Elytra flattened, 1.2 times as broad as long, 1.61.7 times as long as pronotum, strongly broadened posteriad, with straight or widely rounded apical margins. Punctation dense, distinctly larger and deeper than that on pronotum, denser and finer around scutellum, with interstices between punctures in middle about as long as diameter of one-two nearest puncture;
<typeStatus id="F158249D2569163243E6FA6BFE3E98F0" box="[399,506,1458,1484]" pageId="2" pageNumber="386" type="paratype">paratypes</typeStatus>
with finer and sparser punctation along suture. Hind wings fully developed.
</paragraph>
<paragraph id="2E5C9A3F2569163242AEFA0FFF0E9B04" blockId="2.[151,1437,1350,2024]" pageId="2" pageNumber="386">Abdomen distinctly broader than elytra, with two small and transverse tomentose spots in the middle of abdominal tergite V; apical margin of abdominal tergite VII with narrow palisade fringe. Punctation dense and fine.</paragraph>
<paragraph id="2E5C9A3F2569163242AEF99BFA949A2C" blockId="2.[151,1437,1350,2024]" pageId="2" pageNumber="386">
Male. Protarsomeres 14 wide. Apical portion of pronotum wide. Apical margin of abdominal tergite VIII somewhat straight. Apical margin of abdominal sternite VIII sinuate. Aedeagus with small narrow basal portion and elongate moderately wide median lobe, strongly narrowed in apical third, slightly broadened in preapical portion and narrowed toward rounded apex; parameres narrow, almost reaching apex of median lobe, each with four short apical setae; internal sac moderately long with two pairs of narrow elongate sclerotized structures and long narrow flagellum between them, spirally folded in basal portion (
<figureCitation id="B6D886BA256916324172F92FFC9B9A2C" box="[795,863,1782,1808]" captionStart="FIGURES 36" captionStartId="3.[151,264,1842,1866]" captionTargetBox="[177,1417,713,1811]" captionTargetId="figure-228@3.[151,1436,705,1819]" captionTargetPageId="3" captionText="FIGURES 36. Aedeagus of Hygrodromicus (Figs 34: H. incrassatus (holotype); Figs. 56: H. splendidus (Kazakhstan: Ketmen` Mts.): 3, 5parameral view, 4, 6lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12758950" httpUri="https://zenodo.org/record/12758950/files/figure.png" pageId="2" pageNumber="386">Fig. 3</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B6D886BA256916324761F92FFA889A2C" box="[1288,1356,1782,1808]" captionStart="FIGURES 36" captionStartId="3.[151,264,1842,1866]" captionTargetBox="[177,1417,713,1811]" captionTargetId="figure-228@3.[151,1436,705,1819]" captionTargetPageId="3" captionText="FIGURES 36. Aedeagus of Hygrodromicus (Figs 34: H. incrassatus (holotype); Figs. 56: H. splendidus (Kazakhstan: Ketmen` Mts.): 3, 5parameral view, 4, 6lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12758950" httpUri="https://zenodo.org/record/12758950/files/figure.png" pageId="2" pageNumber="386">Fig. 4</figureCitation>
.
</paragraph>
<paragraph id="2E5C9A3F2569163242AEF8C3FB039A64" blockId="2.[151,1437,1350,2024]" pageId="2" pageNumber="386">Female. Protarsomeres 14 moderately narrow. Apical portion of pronotum relatively narrow. Apical margins of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII somewhat rounded.</paragraph>
<paragraph id="2E5C9A3F2569163342AEF8B8FEC19CED" blockId="2.[151,1437,1350,2024]" lastBlockId="3.[151,1437,151,681]" lastPageId="3" lastPageNumber="387" pageId="2" pageNumber="386">
<emphasis id="1C97462D2569163242AEF8B8FE759A40" bold="true" box="[199,433,1889,1916]" pageId="2" pageNumber="386">Comparative notes.</emphasis>
Based on the general shape of the body and the aedeagus,
<taxonomicName id="E9E3E1BC25691632462CF8BBFB2C9A47" authorityName="Shavrin" authorityYear="2024" box="[1093,1256,1890,1915]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="incrassatus" status="sp. n.">
<emphasis id="1C97462D25691632462CF8BBFB2C9A47" box="[1093,1256,1890,1915]" italics="true" pageId="2" pageNumber="386">H. incrassatus</emphasis>
</taxonomicName>
<emphasis id="1C97462D256916324687F8BBFAEA9A40" bold="true" box="[1262,1326,1890,1916]" pageId="2" pageNumber="386">
<taxonomicNameLabel id="07A4FB56256916324687F8BBFAEA9A40" box="[1262,1326,1890,1916]" pageId="2" pageNumber="386" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="E9E3E1BC2569163242DDF85FFE049A9C" authority="Zerche, 1992" authorityName="Zerche" authorityYear="1992" box="[180,448,1926,1952]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="386" phylum="Arthropoda" rank="species" species="wrasei">
<emphasis id="1C97462D2569163242DDF85FFEE69AA3" box="[180,290,1926,1951]" italics="true" pageId="2" pageNumber="386">H. wrasei</emphasis>
<bibRefCitation id="4A72E7CE256916324340F85FFE049A9C" author="Zerche, L." box="[297,448,1926,1952]" pageId="2" pageNumber="386" pagination="105 - 142" refId="ref3633" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche, 1992</bibRefCitation>
</taxonomicName>
, known from
<collectingCountry id="56F4DAAF256916324034F85FFD0F9A9C" box="[605,715,1926,1952]" name="Tajikistan" pageId="2" pageNumber="386">Tajikistan</collectingCountry>
(
<bibRefCitation id="4A72E7CE2569163240B3F85FFCA89A9C" author="Zerche, L." box="[730,876,1926,1952]" pageId="2" pageNumber="386" pagination="105 - 142" refId="ref3633" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche 1992</bibRefCitation>
,
<bibRefCitation id="4A72E7CE25691632411EF85FFBD79A9C" author="Shavrin, A. V." box="[887,1043,1926,1952]" pageId="2" pageNumber="386" pagination="236 - 246" refId="ref3567" refString="Shavrin, A. V. (2024) New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa, 5446 (2), 236 - 246. https: // doi. org / 10.11646 / zootaxa. 5446.2.5" type="journal article" year="2024">Shavrin 2024</bibRefCitation>
). It can be distinguished from it by the larger body, the presence of ocelli, longer antennomeres 610, more transverse pronotum (especially in male) with more narrowed laterobasal margins, longer median lobe of the aedeagus with different shape of the apical portion, and the presence of the flagellum (missing in
<taxonomicName id="E9E3E1BC256816334096FF4EFCA99D8C" authorityName="Zerche" authorityYear="1992" box="[767,877,151,177]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="387" phylum="Arthropoda" rank="species" species="wrasei">
<emphasis id="1C97462D256816334096FF4EFCD79D8C" box="[767,787,151,176]" italics="true" pageId="3" pageNumber="387">H</emphasis>
.
<emphasis id="1C97462D25681633414BFF4EFCA99D8C" box="[802,877,151,176]" italics="true" pageId="3" pageNumber="387">wrasei</emphasis>
</taxonomicName>
). Based on the general shape of the aedeagus,
<taxonomicName id="E9E3E1BC2568163347E8FF4EFED39DE8" authorityName="Shavrin" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="387" phylum="Arthropoda" rank="species" species="incrassatus" status="sp. n.">
<emphasis id="1C97462D2568163347E8FF4EFED39DE8" italics="true" pageId="3" pageNumber="387">H. incrassatus</emphasis>
</taxonomicName>
<emphasis id="1C97462D256816334377FF62FE999DE9" bold="true" box="[286,349,187,213]" pageId="3" pageNumber="387">
<taxonomicNameLabel id="07A4FB56256816334377FF62FE999DE9" box="[286,349,187,213]" pageId="3" pageNumber="387" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is also similar to
<taxonomicName id="E9E3E1BC25681633404AFF62FC439DE9" authority="Coiffait, 1983" authorityName="Coiffait" authorityYear="1983" box="[547,903,187,213]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="387" phylum="Arthropoda" rank="species" species="cachemiricum">
<emphasis id="1C97462D25681633404AFF62FD209DE9" box="[547,740,187,213]" italics="true" pageId="3" pageNumber="387">H. cachemiricum</emphasis>
<bibRefCitation id="4A72E7CE256816334083FF62FC439DE9" author="Coiffait, H." box="[746,903,187,213]" pageId="3" pageNumber="387" pagination="273 - 289" refId="ref3438" refString="Coiffait, H. (1983) Nouveaux staphylinides du Cachemire (Coleoptera). Annales de la Societe entomologique de France, New Series, 19 (3), 273 - 289. https: // doi. org / 10.1080 / 21686351.1983.12278365" type="journal article" year="1983">Coiffait, 1983</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="56F4DAAF2568163341FFFF62FC119DE9" box="[918,981,187,213]" name="India" pageId="3" pageNumber="387">India</collectingCountry>
: Kashmir),
<taxonomicName id="E9E3E1BC25681633463EFF62FA589DE9" authority="Coiffait, 1982" authorityName="Coiffait" authorityYear="1982" box="[1111,1436,187,213]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="387" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="1C97462D25681633463EFF62FB3D9DE9" box="[1111,1273,187,213]" italics="true" pageId="3" pageNumber="387">H. klapperichi</emphasis>
Coiffait, 1982
</taxonomicName>
(
<collectingCountry id="56F4DAAF2568163342F6FF06FEEA9DC5" box="[159,302,223,249]" name="Afghanistan" pageId="3" pageNumber="387">Afghanistan</collectingCountry>
), and Middle Asian
<taxonomicName id="E9E3E1BC256816334071FF06FCB59DC5" authority="(Scheerpeltz, 1933)" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[536,881,223,249]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="387" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="1C97462D256816334071FF06FD4E9DC4" box="[536,650,223,248]" italics="true" pageId="3" pageNumber="387">H. reitteri</emphasis>
(Scheerpeltz, 1933)
</taxonomicName>
and
<taxonomicName id="E9E3E1BC2568163341C5FF06FB8C9DC5" authorityName="Zerche" authorityYear="1992" box="[940,1096,223,249]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="387" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="1C97462D2568163341C5FF06FB8C9DC5" box="[940,1096,223,249]" italics="true" pageId="3" pageNumber="387">H. splendidus</emphasis>
</taxonomicName>
. It can be distinguished from these species by the following morphological features: from
<taxonomicName id="E9E3E1BC256816334125FEDAFBC09C21" authorityName="Coiffait" authorityYear="1983" box="[844,1028,259,285]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="387" phylum="Arthropoda" rank="species" species="cachemiricus">
<emphasis id="1C97462D256816334125FEDAFBC09C21" box="[844,1028,259,285]" italics="true" pageId="3" pageNumber="387">H. cachemiricus</emphasis>
</taxonomicName>
by the broader pronotum, narrower and longer median lobe, with shorter apical portion; from
<taxonomicName id="E9E3E1BC256816334159FEFEFC109C7D" authorityName="Coiffait" authorityYear="1982" box="[816,980,295,321]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="387" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="1C97462D256816334159FEFEFC109C7D" box="[816,980,295,321]" italics="true" pageId="3" pageNumber="387">H. klapperichi</emphasis>
</taxonomicName>
and
<taxonomicName id="E9E3E1BC256816334664FEFEFBBA9C7C" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[1037,1150,295,320]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="387" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="1C97462D256816334664FEFEFBBA9C7C" box="[1037,1150,295,320]" italics="true" pageId="3" pageNumber="387">H. reitteri</emphasis>
</taxonomicName>
by the distinctly broader pronotum and elytra, shorter median lobe with different shape of the apical portion; from
<taxonomicName id="E9E3E1BC25681633461FFE92FACB9C59" authorityName="Zerche" authorityYear="1992" box="[1142,1295,331,357]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="387" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="1C97462D25681633461FFE92FACB9C59" box="[1142,1295,331,357]" italics="true" pageId="3" pageNumber="387">H. splendidus</emphasis>
</taxonomicName>
by narrower antennae and the less convex pronotum, the longer parameres and different shape of the apical portion of the median lobe. From all these species
<taxonomicName id="E9E3E1BC2568163343BCFE4AFDBC9C90" authorityName="Shavrin" authorityYear="2024" box="[469,632,403,428]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="387" phylum="Arthropoda" rank="species" species="incrassatus" status="sp. n.">
<emphasis id="1C97462D2568163343BCFE4AFDBC9C90" box="[469,632,403,428]" italics="true" pageId="3" pageNumber="387">H. incrassatus</emphasis>
</taxonomicName>
<emphasis id="1C97462D256816334017FE4AFD7A9C91" bold="true" box="[638,702,403,429]" pageId="3" pageNumber="387">
<taxonomicNameLabel id="07A4FB56256816334017FE4AFD7A9C91" box="[638,702,403,429]" pageId="3" pageNumber="387" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished by the details of the internal structure of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="66F9C9B42568163342AEFE03FEA39F5D" pageId="3" pageNumber="387" type="distribution">
<paragraph id="2E5C9A3F2568163342AEFE03FF719F25" blockId="3.[151,1437,151,681]" pageId="3" pageNumber="387">
<emphasis id="1C97462D2568163342AEFE03FEA59CC8" bold="true" box="[199,353,474,500]" pageId="3" pageNumber="387">Distribution.</emphasis>
<taxonomicName id="E9E3E1BC25681633430EFE02FD5F9CC8" authorityName="Shavrin" authorityYear="2024" box="[359,667,475,501]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="387" phylum="Arthropoda" rank="species" species="incrassatus" status="sp. nov.">
<emphasis id="1C97462D25681633430EFE02FD5F9CC8" box="[359,667,475,501]" italics="true" pageId="3" pageNumber="387">Hygrodromicus incrassatus</emphasis>
</taxonomicName>
<emphasis id="1C97462D2568163340C8FE02FD249CC9" bold="true" box="[673,736,475,501]" pageId="3" pageNumber="387">
<taxonomicNameLabel id="07A4FB562568163340C8FE02FD249CC9" box="[673,736,475,501]" pageId="3" pageNumber="387" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="F158249D256816334186FE02FBDB9CC9" box="[1007,1055,475,501]" pageId="3" pageNumber="387">type</typeStatus>
locality in northern
<collectingCountry id="56F4DAAF256816334768FE02FAA49CC9" box="[1281,1376,475,501]" name="Pakistan" pageId="3" pageNumber="387">Pakistan</collectingCountry>
(
<figureCitation id="B6D886BA256816334706FE02FF639F25" captionStart="FIGURE 7" captionStartId="4.[151,250,1743,1767]" captionTargetBox="[151,1435,664,1720]" captionTargetId="figure-160@4.[151,1435,664,1720]" captionTargetPageId="4" captionText="FIGURE 7. Distribution of Hygrodromicus incrassatus (square) and H. splendidus (circles)." figureDoi="http://doi.org/10.5281/zenodo.12758954" httpUri="https://zenodo.org/record/12758954/files/figure.png" pageId="3" pageNumber="387">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="2E5C9A3F2568163342AEFDFBFEA39F5D" blockId="3.[151,1437,151,681]" pageId="3" pageNumber="387">
<emphasis id="1C97462D2568163342AEFDFBFE8F9F00" bold="true" box="[199,331,546,572]" pageId="3" pageNumber="387">Bionomics.</emphasis>
The
<typeStatus id="F158249D2568163343EEFDFAFE739F01" box="[391,439,547,573]" pageId="3" pageNumber="387">type</typeStatus>
specimens were collected at an elevation of
<quantity id="E91B37DA2568163341DEFDFAFBCA9F01" box="[951,1038,547,573]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="3" pageNumber="387" unit="m" value="2200.0">2200 m</quantity>
a.s.l. The detailed bionomical data remains unknown.
</paragraph>
</subSubSection>
<subSubSection id="66F9C9B42568163342AEFDB2FED99F95" pageId="3" pageNumber="387" type="etymology">
<paragraph id="2E5C9A3F2568163342AEFDB2FED99F95" blockId="3.[151,1437,151,681]" pageId="3" pageNumber="387">
<emphasis id="1C97462D2568163342AEFDB2FE8A9FB9" bold="true" box="[199,334,619,645]" pageId="3" pageNumber="387">Etymology.</emphasis>
The specific epithet is the Latin
<taxonomicName id="E9E3E1BC2568163340AAFDB2FC879FB8" authorityName="Shavrin" authorityYear="2024" box="[707,835,619,644]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="387" phylum="Arthropoda" rank="species" species="incrassatus">
<emphasis id="1C97462D2568163340AAFDB2FC879FB8" box="[707,835,619,644]" italics="true" pageId="3" pageNumber="387">incrassatus</emphasis>
</taxonomicName>
, -
<emphasis id="1C97462D256816334132FDB2FCAD9FB8" box="[859,873,619,644]" italics="true" pageId="3" pageNumber="387">a</emphasis>
, -
<emphasis id="1C97462D2568163341E8FDB2FC679FB8" box="[897,931,619,644]" italics="true" pageId="3" pageNumber="387">um</emphasis>
(thickened). It alludes to the wide pronotum of the male.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,351 @@
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<paragraph id="2E5C9A3F256F163442FEFF4EFD419D8D" blockId="4.[151,645,151,214]" box="[151,645,151,178]" pageId="4" pageNumber="388">
<heading id="75142D53256F163442FEFF4EFD419D8D" bold="true" box="[151,645,151,178]" fontSize="11" level="1" pageId="4" pageNumber="388" reason="1">
<taxonomicName id="E9E3E1BC256F163442FEFF4EFD419D8D" ID-CoL="3NBJ4" authority="Zerche, 1992" authorityName="Zerche" authorityYear="1992" box="[151,645,151,178]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="388" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="1C97462D256F163442FEFF4EFD419D8D" bold="true" box="[151,645,151,178]" pageId="4" pageNumber="388">
<emphasis id="1C97462D256F163442FEFF4EFE199D8E" bold="true" box="[151,477,151,178]" italics="true" pageId="4" pageNumber="388">Hygrodromicus splendidus</emphasis>
<bibRefCitation id="4A72E7CE256F1634438DFF4EFD419D8D" author="Zerche, L." box="[484,645,151,177]" pageId="4" pageNumber="388" pagination="105 - 142" refId="ref3633" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche, 1992</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="66F9C9B4256F163442FEFF62FEE39DEA" box="[151,295,187,214]" pageId="4" pageNumber="388" type="description">
<paragraph id="2E5C9A3F256F163442FEFF62FEE39DEA" blockId="4.[151,645,151,214]" box="[151,295,187,214]" pageId="4" pageNumber="388">
(
<figureCitation id="B6D886BA256F163442F6FF62FF2C9DE9" box="[159,232,187,213]" captionStart="FIGURES 12" captionStartId="2.[151,264,1207,1231]" captionTargetBox="[157,1430,281,1175]" captionTargetId="figure-60@2.[151,1436,270,1181]" captionTargetPageId="2" captionText="FIGURES 12. Habitus of Hygrodromicus: 1H. incrassatus (paratype), 2H. splendidus (Kazakhstan, Ketmen` Mts.). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12758946" httpUri="https://zenodo.org/record/12758946/files/figure.png" pageId="4" pageNumber="388">Figs 2</figureCitation>
,
<figureCitation id="B6D886BA256F1634429DFF62FEDB9DEA" box="[244,287,187,214]" captionStart-0="FIGURES 36" captionStart-1="FIGURE 7" captionStartId-0="3.[151,264,1842,1866]" captionStartId-1="4.[151,250,1743,1767]" captionTargetBox-0="[177,1417,713,1811]" captionTargetBox-1="[151,1435,664,1720]" captionTargetId-0="figure-228@3.[151,1436,705,1819]" captionTargetId-1="figure-160@4.[151,1435,664,1720]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 36. Aedeagus of Hygrodromicus (Figs 34: H. incrassatus (holotype); Figs. 56: H. splendidus (Kazakhstan: Ketmen` Mts.): 3, 5parameral view, 4, 6lateral view. Scale bars: 0.1 mm." captionText-1="FIGURE 7. Distribution of Hygrodromicus incrassatus (square) and H. splendidus (circles)." figureDoi-0="http://doi.org/10.5281/zenodo.12758950" figureDoi-1="http://doi.org/10.5281/zenodo.12758954" httpUri-0="https://zenodo.org/record/12758950/files/figure.png" httpUri-1="https://zenodo.org/record/12758954/files/figure.png" pageId="4" pageNumber="388">57</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="66F9C9B4256F163442FEFED9FDA19C2B" box="[151,613,255,280]" pageId="4" pageNumber="388" type="reference_group">
<paragraph id="2E5C9A3F256F163442FEFED9FDA19C2B" blockId="4.[151,613,255,280]" box="[151,613,255,280]" pageId="4" pageNumber="388">
<treatmentCitationGroup id="0EF3BD11256F163442FEFED9FDA19C2B" box="[151,613,255,280]" pageId="4" pageNumber="388">
<treatmentCitation id="AF42BC2E256F163442FEFED9FDA19C2B" author="Zerche, L." box="[151,613,255,280]" page="116" pageId="4" pageNumber="388" year="1992">
<taxonomicName id="E9E3E1BC256F163442FEFED9FDA19C2B" ID-CoL="3NBJ4" authority="Zerche, 1992: 116" authorityName="Zerche" authorityPageNumber="116" authorityYear="1992" box="[151,613,255,280]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="388" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="1C97462D256F163442FEFED9FE639C24" box="[151,423,256,280]" italics="true" pageId="4" pageNumber="388">Hygrodromicus splendidus</emphasis>
<bibRefCitation id="4A72E7CE256F163443C4FED9FDA19C2B" author="Zerche, L." box="[429,613,255,280]" pageId="4" pageNumber="388" pagination="105 - 142" refId="ref3633" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche, 1992: 116</bibRefCitation>
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</paragraph>
</subSubSection>
<subSubSection id="66F9C9B4256F163442FEFE9CFA959CCC" pageId="4" pageNumber="388" type="materials_examined">
<paragraph id="2E5C9A3F256F163442FEFE9CFA959CCC" blockId="4.[151,1437,325,641]" pageId="4" pageNumber="388">
<emphasis id="1C97462D256F163442FEFE9CFEBB9C63" bold="true" box="[151,383,325,351]" pageId="4" pageNumber="388">Material examined.</emphasis>
<materialsCitation id="9E8B9062256F163443EAFE9CFBBD9C5C" box="[387,1145,325,352]" collectionCode="R" country="Kazakhstan" location="Dolaity" municipality="Ketmen Ridge" pageId="4" pageNumber="388" specimenCount="30" specimenCount-female="21" specimenCount-male="9" stateProvince="Almaty Oblysy">
<emphasis id="1C97462D256F163443EAFE9CFD7C9C63" bold="true" box="[387,696,325,351]" pageId="4" pageNumber="388">
<collectingCountry id="56F4DAAF256F163443EAFE9CFD899C63" box="[387,589,325,351]" name="Kazakhstan" pageId="4" pageNumber="388">KAZAKHSTAN</collectingCountry>
:
<collectingRegion id="EC2754DD256F1634403EFE9CFD709C63" box="[599,692,325,351]" country="Kazakhstan" name="Almaty Oblysy" pageId="4" pageNumber="388">ALMATY</collectingRegion>
:
</emphasis>
<specimenCount id="38E551B6256F163440D5FE9FFD3F9C5C" box="[700,763,326,352]" count="9" pageId="4" pageNumber="388" type="male">9 ♂♂</specimenCount>
,
<specimenCount id="38E551B6256F1634416FFE9FFC979C5C" box="[774,851,326,352]" count="21" pageId="4" pageNumber="388" type="female">21 ♀♀</specimenCount>
:
<collectingMunicipality id="CE380045256F16344134FE9FFBC59C5C" box="[861,1025,326,352]" pageId="4" pageNumber="388">Ketmen Ridge</collectingMunicipality>
,
<location id="2B3CCCE4256F16344663FE9FFB9A9C5C" LSID="urn:lsid:plazi:treatment:A64A2B29256F163542FEFF4EFCB09B35:2B3CCCE4256F16344663FE9FFB9A9C5C" box="[1034,1118,326,352]" country="Kazakhstan" municipality="Ketmen Ridge" name="Dolaity" pageId="4" pageNumber="388" stateProvince="Almaty Oblysy">Dolaity</location>
<collectionCode id="48F202FA256F1634460AFE9FFBBD9C5C" box="[1123,1145,326,352]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="388">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="9E8B9062256F163446E8FE9FFADB9C5C" box="[1153,1311,326,353]" collectingDate="1988-07-15" collectionCode="V" country="Kazakhstan" pageId="4" pageNumber="388" specimenCount="1" stateProvince="Almaty Oblysy">
<date id="5A5DBCFF256F163446E8FE9FFAC79C5D" box="[1153,1283,326,353]" pageId="4" pageNumber="388" value="1988-07-15">
<collectingDate id="4A194517256F163446E8FE9FFAC79C5D" box="[1153,1283,326,353]" pageId="4" pageNumber="388" value="1988-07-15">15.07.1988</collectingDate>
</date>
.
<collectionCode id="48F202FA256F16344762FE9FFADB9C5C" box="[1291,1311,326,352]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="388" type="Museum">V</collectionCode>
</materialsCitation>
.
<collectorName id="8316FFE9256F16344741FE9FFA599C5C" box="[1320,1437,326,352]" pageId="4" pageNumber="388">Kastcheev</collectorName>
leg. (cSh,
<collectionCode id="48F202FA256F16344360FEB3FE959CB8" box="[265,337,362,388]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="388">NMW</collectionCode>
,
<collectionCode id="48F202FA256F16344334FEB3FE549CB8" box="[349,400,362,388]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="388" type="Museum">ZIN</collectionCode>
);
<materialsCitation id="9E8B9062256F163443CDFEB3FC9C9CB8" box="[420,856,362,388]" collectingDate="1988-09-07" collectionCode="V" country="Kazakhstan" pageId="4" pageNumber="388" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Almaty Oblysy">
<specimenCount id="38E551B6256F163443CDFEB3FE149CB8" box="[420,464,362,388]" count="1" pageId="4" pageNumber="388" type="male">1 ♂</specimenCount>
,
<specimenCount id="38E551B6256F163443B5FEB3FDCD9CB8" box="[476,521,362,388]" count="1" pageId="4" pageNumber="388" type="female">1 ♀</specimenCount>
: same data,
<date id="5A5DBCFF256F163440FAFEB3FCD59CB8" box="[659,785,362,388]" pageId="4" pageNumber="388" value="1988-09-07">
<collectingDate id="4A194517256F163440FAFEB3FCD59CB8" box="[659,785,362,388]" pageId="4" pageNumber="388" value="1988-09-07">09.07.1988</collectingDate>
</date>
(cSh)
</materialsCitation>
;
<materialsCitation id="9E8B9062256F1634410DFEB3FB609CB8" box="[868,1188,362,388]" collectingDate="1988-09-07" collectionCode="R" country="Kazakhstan" location="Avat" pageId="4" pageNumber="388" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Almaty Oblysy">
<specimenCount id="38E551B6256F1634410DFEB3FC559CB8" box="[868,913,362,388]" count="1" pageId="4" pageNumber="388" type="male">1 ♂</specimenCount>
,
<specimenCount id="38E551B6256F163441F4FEB3FC0D9CB8" box="[925,969,362,388]" count="1" pageId="4" pageNumber="388" type="female">1 ♀</specimenCount>
: same data,
<location id="2B3CCCE4256F1634463AFEB3FB439CB8" LSID="urn:lsid:plazi:treatment:A64A2B29256F163542FEFF4EFCB09B35:2B3CCCE4256F1634463AFEB3FB439CB8" box="[1107,1159,362,388]" country="Kazakhstan" name="Avat" pageId="4" pageNumber="388" stateProvince="Almaty Oblysy">Avat</location>
<collectionCode id="48F202FA256F163446E7FEB3FB609CB8" box="[1166,1188,362,388]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="388">R</collectionCode>
</materialsCitation>
.
<date id="5A5DBCFF256F163446C6FEB3FAE99CB8" box="[1199,1325,362,388]" pageId="4" pageNumber="388" value="2009-07-14">14.07.2009</date>
(cSh);
<materialsCitation id="9E8B9062256F163447E9FEB3FD159C94" collectingDate="1988-09-07" collectionCode="R" country="Kazakhstan" location="Kulbastau" pageId="4" pageNumber="388" specimenCount="19" specimenCount-female="6" specimenCount-male="13" stateProvince="Almaty Oblysy">
<specimenCount id="38E551B6256F163447E9FEB3FF079C94" count="13" pageId="4" pageNumber="388" type="male">13 ♂♂</specimenCount>
,
<specimenCount id="38E551B6256F163442B9FE57FED59C94" box="[208,273,398,424]" count="6" pageId="4" pageNumber="388" type="female">6 ♀♀</specimenCount>
: same data,
<location id="2B3CCCE4256F163443F4FE57FDCB9C94" LSID="urn:lsid:plazi:treatment:A64A2B29256F163542FEFF4EFCB09B35:2B3CCCE4256F163443F4FE57FDCB9C94" box="[413,527,398,424]" country="Kazakhstan" name="Kulbastau" pageId="4" pageNumber="388" stateProvince="Almaty Oblysy">Kulbastau</location>
[=Kol`bastau]
<collectionCode id="48F202FA256F163440D2FE57FD159C94" box="[699,721,398,424]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="388">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="9E8B9062256F163440B4FE57FC249C94" box="[733,992,398,424]" collectingDate="1988-07-27" collectionCode="ZIN" country="Kazakhstan" pageId="4" pageNumber="388" specimenCount="1" stateProvince="Almaty Oblysy">
<date id="5A5DBCFF256F163440B4FE57FC9F9C94" box="[733,859,398,424]" pageId="4" pageNumber="388" value="1988-07-27">
<collectingDate id="4A194517256F163440B4FE57FC9F9C94" box="[733,859,398,424]" pageId="4" pageNumber="388" value="1988-07-27">27.07.1988</collectingDate>
</date>
(cSh,
<collectionCode id="48F202FA256F163441CDFE57FC139C94" box="[932,983,398,424]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="388" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="9E8B9062256F16344185FE57FE9A9CF0" collectingDate="1991-05-18" collectionCode="ZIN" collectorName="Bolshoy Kyrgyzsay." country="Kazakhstan" pageId="4" pageNumber="388" specimenCount="3" specimenCount-female="3" stateProvince="Almaty Oblysy">
<specimenCount id="38E551B6256F16344185FE57FBE99C94" box="[1004,1069,398,424]" count="3" pageId="4" pageNumber="388" type="female">3 ♀♀</specimenCount>
: same data,
<collectorName id="8316FFE9256F163446D0FE57FA599C94" box="[1209,1437,398,424]" pageId="4" pageNumber="388">Bolshoy Kyrgyzsay.</collectorName>
<date id="5A5DBCFF256F163442FEFE6BFED29CF1" box="[151,278,434,461]" pageId="4" pageNumber="388" value="1991-05-18">
<collectingDate id="4A194517256F163442FEFE6BFED29CF1" box="[151,278,434,461]" pageId="4" pageNumber="388" value="1991-05-18">18.05.1991</collectingDate>
</date>
(cSh)
</materialsCitation>
;
<materialsCitation id="9E8B9062256F16344305FE6BFD369CF0" box="[364,754,434,460]" collectingDate="1988-10-06" collectionCode="ZIN" collectorName="Bolshoy Kyrgyzsay." country="Kazakhstan" pageId="4" pageNumber="388" specimenCount="1" specimenCount-female="1" stateProvince="Almaty Oblysy">
<specimenCount id="38E551B6256F16344305FE6BFE5F9CF0" box="[364,411,434,460]" count="1" pageId="4" pageNumber="388" type="female">1 ♀</specimenCount>
: same data,
<date id="5A5DBCFF256F16344042FE6BFD6D9CF0" box="[555,681,434,460]" pageId="4" pageNumber="388" value="1988-10-06">
<collectingDate id="4A194517256F16344042FE6BFD6D9CF0" box="[555,681,434,460]" pageId="4" pageNumber="388" value="1988-10-06">10.06.1988</collectingDate>
</date>
(cSh)
</materialsCitation>
;
<materialsCitation id="9E8B9062256F16344169FE68FB2E9CF0" box="[768,1258,433,460]" collectionCode="R" country="Kazakhstan" location="Koksay" municipality="Aksu-Dzhabagly" pageId="4" pageNumber="388" specimenCount="1" specimenCount-male="1" stateProvince="JAMBYL">
<emphasis id="1C97462D256F16344169FE68FCA09CF7" bold="true" box="[768,868,433,459]" pageId="4" pageNumber="388">
<collectingRegion id="EC2754DD256F16344169FE68FC9B9CF7" box="[768,863,433,459]" country="Kazakhstan" name="JAMBYL" pageId="4" pageNumber="388">JAMBYL</collectingRegion>
:
</emphasis>
<specimenCount id="38E551B6256F16344104FE6BFC589CF0" box="[877,924,434,460]" count="1" pageId="4" pageNumber="388" type="male">1 ♂</specimenCount>
:
<collectingMunicipality id="CE380045256F163441C0FE6BFBAC9CF0" box="[937,1128,434,460]" pageId="4" pageNumber="388">Aksu-Dzhabagly</collectingMunicipality>
,
<location id="2B3CCCE4256F1634461CFE6BFB0F9CF0" LSID="urn:lsid:plazi:treatment:A64A2B29256F163542FEFF4EFCB09B35:2B3CCCE4256F1634461CFE6BFB0F9CF0" box="[1141,1227,434,460]" country="Kazakhstan" municipality="Aksu-Dzhabagly" name="Koksay" pageId="4" pageNumber="388" stateProvince="JAMBYL">Koksay</location>
<collectionCode id="48F202FA256F163446BDFE6BFB2E9CF0" box="[1236,1258,434,460]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="388">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="9E8B9062256F1634469EFE6BFA5D9CF0" box="[1271,1433,434,460]" collectingDate="2001-02-07" collectionCode="V" country="Kazakhstan" pageId="4" pageNumber="388" specimenCount="1" stateProvince="JAMBYL">
<date id="5A5DBCFF256F1634469EFE6BFABC9CF0" box="[1271,1400,434,460]" pageId="4" pageNumber="388" value="2001-02-07">
<collectingDate id="4A194517256F1634469EFE6BFABC9CF0" box="[1271,1400,434,460]" pageId="4" pageNumber="388" value="2001-02-07">02.07.2001</collectingDate>
</date>
.
<collectionCode id="48F202FA256F163447ECFE6BFA5D9CF0" box="[1413,1433,434,460]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="388" type="Museum">V</collectionCode>
</materialsCitation>
. Kastcheev leg. (cSh);
<materialsCitation id="9E8B9062256F163443E7FE0CFA899CCC" box="[398,1357,469,499]" collectingDate="1994-09-06" collectorName="Kegety Gorge. &amp; S. Ovchinnikov" country="Kyrgyzstan" location="CHUY" pageId="4" pageNumber="388" specimenCount="1" specimenCount-female="1" stateProvince="CHUY">
<emphasis id="1C97462D256F163443E7FE0CFD6B9CCC" bold="true" box="[398,687,469,499]" pageId="4" pageNumber="388">
<collectingCountry id="56F4DAAF256F163443E7FE0CFD929CD3" box="[398,598,469,495]" name="Kyrgyzstan" pageId="4" pageNumber="388">KYRGYZSTAN</collectingCountry>
:
<collectingRegion id="EC2754DD256F1634400BFE0EFD619CCF" box="[610,677,471,499]" country="Kyrgyzstan" name="CHUY" pageId="4" pageNumber="388">CHÜY</collectingRegion>
:
</emphasis>
<specimenCount id="38E551B6256F163440DFFE0FFD269CCC" box="[694,738,470,496]" count="1" pageId="4" pageNumber="388" type="female">1 ♀</specimenCount>
:
<collectorName id="8316FFE9256F16344086FE0FFC509CCC" box="[751,916,470,496]" pageId="4" pageNumber="388">Kegety Gorge.</collectorName>
<date id="5A5DBCFF256F163441F3FE0FFBD89CCC" box="[922,1052,470,496]" pageId="4" pageNumber="388" value="1994-09-06">
<collectingDate id="4A194517256F163441F3FE0FFBD89CCC" box="[922,1052,470,496]" pageId="4" pageNumber="388" value="1994-09-06">09.06.1994</collectingDate>
</date>
.
<collectorName id="8316FFE9256F1634464FFE0FFB129CCC" box="[1062,1238,470,496]" pageId="4" pageNumber="388">S. Ovchinnikov</collectorName>
leg. (cSh)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="66F9C9B4256F163542AEFE20FA4199D5" lastPageId="5" lastPageNumber="389" pageId="4" pageNumber="388" type="description">
<paragraph id="2E5C9A3F256F163442AEFE20FDAF9FBC" blockId="4.[151,1437,325,641]" pageId="4" pageNumber="388">
<emphasis id="1C97462D256F163442AEFE20FEB19F2F" bold="true" box="[199,373,505,531]" pageId="4" pageNumber="388">Redescription.</emphasis>
Measurements (n = 50): HW: 0.941.01; HL: 0.740.75; OL: 0.250.26; LT: 0.170.18; AL (averaged): 3.46; PL: 0.880.94; PWmax: 1.331.41; PWmin: 1.231.25; ESL: 1.471.63; EW: 2.072.35; MTbL(averaged): 1.55; MTrL (averaged): 0.34 (MTrL 14: 0.19; MTrL 5: 0.15); AW: 2.132.36; AedL: 1.281.34; BL: 5.457.00 (
<typeStatus id="F158249D256F16344320FDBFFE759FBC" box="[329,433,614,640]" pageId="4" pageNumber="388" type="holotype">holotype</typeStatus>
:
<bibRefCitation id="4A72E7CE256F163443D2FDBFFDA39FBC" author="Zerche, L." box="[443,615,614,640]" pageId="4" pageNumber="388" pagination="105 - 142" refId="ref3633" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche (1992))</bibRefCitation>
.
</paragraph>
<caption id="7A9CCAB7256F163442FEF916FB879BDB" ID-DOI="http://doi.org/10.5281/zenodo.12758954" ID-Zenodo-Dep="12758954" box="[151,1091,1743,1767]" httpUri="https://zenodo.org/record/12758954/files/figure.png" pageId="4" pageNumber="388" startId="4.[151,250,1743,1767]" targetBox="[151,1435,664,1720]" targetPageId="4" targetType="figure">
<paragraph id="2E5C9A3F256F163442FEF916FB879BDB" blockId="4.[151,1091,1743,1767]" box="[151,1091,1743,1767]" pageId="4" pageNumber="388">
<emphasis id="1C97462D256F163442FEF916FED79BDB" bold="true" box="[151,275,1743,1767]" pageId="4" pageNumber="388">FIGURE 7.</emphasis>
Distribution of
<taxonomicName id="E9E3E1BC256F163443DFF916FD089BDB" authorityName="Shavrin" authorityYear="2024" box="[438,716,1743,1767]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="388" phylum="Arthropoda" rank="species" species="incrassatus">
<emphasis id="1C97462D256F163443DFF916FD089BDB" box="[438,716,1743,1767]" italics="true" pageId="4" pageNumber="388">Hygrodromicus incrassatus</emphasis>
</taxonomicName>
(square) and
<taxonomicName id="E9E3E1BC256F1634413EF909FC269BDB" authorityName="Zerche" authorityYear="1992" box="[855,994,1743,1767]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="388" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="1C97462D256F1634413EF909FC269BDB" box="[855,994,1743,1767]" italics="true" pageId="4" pageNumber="388">H. splendidus</emphasis>
</taxonomicName>
(circles).
</paragraph>
</caption>
<paragraph id="2E5C9A3F256F163442AEF8E0FA9A9ADF" blockId="4.[151,1437,1849,2020]" pageId="4" pageNumber="388">
Habitus as in
<figureCitation id="B6D886BA256F1634430FF8E0FE689A68" box="[358,428,1849,1876]" captionStart="FIGURES 12" captionStartId="2.[151,264,1207,1231]" captionTargetBox="[157,1430,281,1175]" captionTargetId="figure-60@2.[151,1436,270,1181]" captionTargetPageId="2" captionText="FIGURES 12. Habitus of Hygrodromicus: 1H. incrassatus (paratype), 2H. splendidus (Kazakhstan, Ketmen` Mts.). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12758946" httpUri="https://zenodo.org/record/12758946/files/figure.png" pageId="4" pageNumber="388">Fig. 2</figureCitation>
. Body reddish-brown to black, usually with distinctly paler elytra; mouthparts, antennae and legs brownish; apical segment of maxillary palpi and tarsi yellowish. Head with dense and fine isodiametric microculpture, coarser on infraorbital portions; neck with dense transverse or isodiametric sculpture; pronotum with dense and coarse isodiametric meshes, usually finer along middle and mediobasal third; scutellum with dense and fine transverse meshes; abdomen with dense and sometimes coarse isodametric or transverse microsculpture.
</paragraph>
<paragraph id="2E5C9A3F256E163542AEFF4EFDEC9C91" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">
Head 1.21.3 times as broad as long, with slightly elevated middle portion; medioapical depression deep and wide, narrowing posteriad; interocellar depression wide, relatively shallow or deep; anteocellar foveae narrow and long, strongly convergent diagonally toward level of anterior margins of eyes. Punctation irregular, fine, sometimes invisible in some specimens, denser and slightly larger between eyes; neck without visible punctation or with several punctures in middle. Ocelli small and indistinct (invisible in
<typeStatus id="F158249D256E16354154FEFEFC659C7D" box="[829,929,295,321]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
) or moderately large. Antenna reaching basal third or apical margins of elytra when reclined, with somewhat widened antennomeres; antennomere 3 slightly longer than 2, 45 slightly longer and usually indistinctly broader than 3, 610 slightly longer than 5 (sometimes antennomere 5 indistinctly shorter).
</paragraph>
<paragraph id="2E5C9A3F256E163542AEFE6EFECD9FB9" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">Pronotum markedly convex and distinctly transverse, 1.5 times as broad as long, 1.31.4 times as broad as head, from widest anterior portion strongly narrowed posteriad toward obtuse or subacute posterior angles; anterior angles widely rounded, sometimes slightly protruded anteriad; posterior margin straight or rounded, slightly narrower than posterior margin usually widely concave in middle. Punctation fine and dense, more distinct than that on head, finer in middle and sometimes with narrow impunctate median band. Scutellum without or with several fine punctures in middle.</paragraph>
<paragraph id="2E5C9A3F256E163542AEFD56FB9A9FF1" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">Elytra 1.11.3 times as broad as long. Punctation dense, distinctly larger and deeper than that on pronotum, with interstices between punctures in middle about one-two diameters of nearest punctures.</paragraph>
<paragraph id="2E5C9A3F256E163542AEFD0EFE7A9E05" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">
Abdomen slightly narrower (
<typeStatus id="F158249D256E16354064FD0EFDB69FCD" box="[525,626,727,753]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
) or broader than elytra, with two large and transverse tometose spots in the middle of abdominal tergite IV and two small spots in tergite V (some specimens witth indistinct spots); palisade fringe narrow or invisible.
</paragraph>
<paragraph id="2E5C9A3F256E163542AEFC9AFC1D992D" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">
Male.Apical margin of abdominal tergite VIII somewhat straight or rounded.Apical margin of abdominal sternite VIII sinuate. Aedeagus with small basal portion and moderately wide elongate median lobe, distinctly narrowed in apical third, slightly broadened in preapical portion and narrowed toward small rounded apex; parameres narrow, indistinctly broadened apically, usually not reaching apex of median lobe, with four short apical setae; internal sac relatively long, with two narrow sclerotized lobes in preapical portion and moderately long flagellum, spirally folded in basal portion (
<figureCitation id="B6D886BA256E163543CAFC2EFE2C992D" box="[419,488,1014,1041]" captionStart="FIGURES 36" captionStartId="3.[151,264,1842,1866]" captionTargetBox="[177,1417,713,1811]" captionTargetId="figure-228@3.[151,1436,705,1819]" captionTargetPageId="3" captionText="FIGURES 36. Aedeagus of Hygrodromicus (Figs 34: H. incrassatus (holotype); Figs. 56: H. splendidus (Kazakhstan: Ketmen` Mts.): 3, 5parameral view, 4, 6lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12758950" httpUri="https://zenodo.org/record/12758950/files/figure.png" pageId="5" pageNumber="389">Fig. 5</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B6D886BA256E163541F9FC2EFC11992D" box="[912,981,1015,1041]" captionStart="FIGURES 36" captionStartId="3.[151,264,1842,1866]" captionTargetBox="[177,1417,713,1811]" captionTargetId="figure-228@3.[151,1436,705,1819]" captionTargetPageId="3" captionText="FIGURES 36. Aedeagus of Hygrodromicus (Figs 34: H. incrassatus (holotype); Figs. 56: H. splendidus (Kazakhstan: Ketmen` Mts.): 3, 5parameral view, 4, 6lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12758950" httpUri="https://zenodo.org/record/12758950/files/figure.png" pageId="5" pageNumber="389">Fig. 6</figureCitation>
.
</paragraph>
<paragraph id="2E5C9A3F256E163542AEFBC2FF3F9965" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">Female. Apical margins of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII rounded.</paragraph>
<paragraph id="2E5C9A3F256E163542AEFBBBFA4199D5" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">
<emphasis id="1C97462D256E163542AEFBBBFE749941" bold="true" box="[199,432,1122,1149]" pageId="5" pageNumber="389">Comparative notes.</emphasis>
Based on the general shape of the aedeagus,
<taxonomicName id="E9E3E1BC256E163541C0FBBAFB879941" authorityName="Zerche" authorityYear="1992" box="[937,1091,1123,1149]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="1C97462D256E163541C0FBBAFC799940" box="[937,957,1123,1148]" italics="true" pageId="5" pageNumber="389">H</emphasis>
.
<emphasis id="1C97462D256E163541A2FBBAFB879941" box="[971,1091,1123,1149]" italics="true" pageId="5" pageNumber="389">splendidus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="E9E3E1BC256E163546BCFBBAFA529941" authorityName="Coiffait" authorityYear="1983" box="[1237,1430,1123,1149]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="cachemiricum">
<emphasis id="1C97462D256E163546BCFBBAFA529941" box="[1237,1430,1123,1149]" italics="true" pageId="5" pageNumber="389">H. cachemiricum</emphasis>
</taxonomicName>
,
<taxonomicName id="E9E3E1BC256E163542FEFB5EFEFF999C" authorityName="Shavrin" authorityYear="2024" box="[151,315,1159,1184]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="incrassatus" status="sp. n.">
<emphasis id="1C97462D256E163542FEFB5EFEFF999C" box="[151,315,1159,1184]" italics="true" pageId="5" pageNumber="389">H. incrassatus</emphasis>
</taxonomicName>
<emphasis id="1C97462D256E1635432DFB5EFE41999D" bold="true" box="[324,389,1159,1185]" pageId="5" pageNumber="389">
<taxonomicNameLabel id="07A4FB56256E1635432DFB5EFE41999D" box="[324,389,1159,1185]" pageId="5" pageNumber="389" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="E9E3E1BC256E163543FDFB5EFDFC999D" authorityName="Coiffait" authorityYear="1982" box="[404,568,1159,1185]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="1C97462D256E163543FDFB5EFDFC999D" box="[404,568,1159,1185]" italics="true" pageId="5" pageNumber="389">H. klapperichi</emphasis>
</taxonomicName>
,
<taxonomicName id="E9E3E1BC256E1635402EFB5EFD7C999C" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[583,696,1159,1184]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="1C97462D256E1635402EFB5EFD7C999C" box="[583,696,1159,1184]" italics="true" pageId="5" pageNumber="389">H. reitteri</emphasis>
</taxonomicName>
and
<taxonomicName id="E9E3E1BC256E1635409BFB5EFCA5999C" authorityName="Zerche" authorityYear="1992" box="[754,865,1159,1184]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="wrasei">
<emphasis id="1C97462D256E1635409BFB5EFCA5999C" box="[754,865,1159,1184]" italics="true" pageId="5" pageNumber="389">H. wrasei</emphasis>
</taxonomicName>
. From all these species it can be distinguished by the broader antennae, more convex pronotum, and details of the external and internal morphology of the aedeagus. Additionally, it differs from
<taxonomicName id="E9E3E1BC256E163543BDFB16FD5299D5" authorityName="Coiffait" authorityYear="1983" box="[468,662,1231,1257]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="cachemiricum">
<emphasis id="1C97462D256E163543BDFB16FD5299D5" box="[468,662,1231,1257]" italics="true" pageId="5" pageNumber="389">H. cachemiricum</emphasis>
</taxonomicName>
,
<taxonomicName id="E9E3E1BC256E163540CAFB16FC8299D5" authorityName="Coiffait" authorityYear="1982" box="[675,838,1231,1257]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="klapperichi">
<emphasis id="1C97462D256E163540CAFB16FC8299D5" box="[675,838,1231,1257]" italics="true" pageId="5" pageNumber="389">H. klapperichi</emphasis>
</taxonomicName>
and
<taxonomicName id="E9E3E1BC256E16354112FB16FC2F99D4" baseAuthorityName="Scheerpeltz" baseAuthorityYear="1933" box="[891,1003,1231,1256]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="1C97462D256E16354112FB16FC2F99D4" box="[891,1003,1231,1256]" italics="true" pageId="5" pageNumber="389">H. reitteri</emphasis>
</taxonomicName>
by the broader pronotum and elytra.
</paragraph>
</subSubSection>
<subSubSection id="66F9C9B4256E163542AEFB2BFEC39845" pageId="5" pageNumber="389" type="distribution">
<paragraph id="2E5C9A3F256E163542AEFB2BFCF6980D" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">
<emphasis id="1C97462D256E163542AEFB2BFEA59830" bold="true" box="[199,353,1266,1292]" pageId="5" pageNumber="389">Distribution.</emphasis>
<taxonomicName id="E9E3E1BC256E16354302FB2AFD5F9831" authorityName="Zerche" authorityYear="1992" box="[363,667,1267,1293]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="1C97462D256E16354302FB2AFD5F9831" box="[363,667,1267,1293]" italics="true" pageId="5" pageNumber="389">Hygrodromicus splendidus</emphasis>
</taxonomicName>
is known from several locality in the Middle Asia: south-western
<collectingCountry id="56F4DAAF256E163542FEFACEFED9980D" box="[151,285,1303,1329]" name="Kazakhstan" pageId="5" pageNumber="389">Kazakhstan</collectingCountry>
, north
<collectingCountry id="56F4DAAF256E16354303FACEFE2F980D" box="[362,491,1303,1329]" name="Kyrgyzstan" pageId="5" pageNumber="389">Kyrgyzstan</collectingCountry>
and north
<collectingCountry id="56F4DAAF256E1635400BFACEFD14980D" box="[610,720,1303,1329]" name="Tajikistan" pageId="5" pageNumber="389">Tajikistan</collectingCountry>
(
<figureCitation id="B6D886BA256E16354089FACEFCE0980D" box="[736,804,1303,1329]" captionStart="FIGURE 7" captionStartId="4.[151,250,1743,1767]" captionTargetBox="[151,1435,664,1720]" captionTargetId="figure-160@4.[151,1435,664,1720]" captionTargetPageId="4" captionText="FIGURE 7. Distribution of Hygrodromicus incrassatus (square) and H. splendidus (circles)." figureDoi="http://doi.org/10.5281/zenodo.12758954" httpUri="https://zenodo.org/record/12758954/files/figure.png" pageId="5" pageNumber="389">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="2E5C9A3F256E163542AEFAE3FEC39845" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">
<emphasis id="1C97462D256E163542AEFAE3FE8F9868" bold="true" box="[199,331,1338,1364]" pageId="5" pageNumber="389">Bionomics.</emphasis>
The
<typeStatus id="F158249D256E163543E6FAE2FE359869" box="[399,497,1339,1365]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
was collected at elevation
<quantity id="E91B37DA256E16354153FAE2FC1F9869" box="[826,987,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="389" unit="m" value="3500.0" valueMax="4000.0" valueMin="3000.0">30004000 m</quantity>
a.s.l. The detailed bionomical data is unknown.
</paragraph>
</subSubSection>
<subSubSection id="66F9C9B4256E163542AEFA5AFCB09B35" pageId="5" pageNumber="389" type="discussion">
<paragraph id="2E5C9A3F256E163542AEFA5AFCB09B35" blockId="5.[151,1437,151,1545]" pageId="5" pageNumber="389">
<emphasis id="1C97462D256E163542AEFA5AFEFF98A1" bold="true" box="[199,315,1411,1437]" pageId="5" pageNumber="389">Remarks.</emphasis>
<taxonomicName id="E9E3E1BC256E1635432FFA5AFDB298A1" authorityName="Zerche" authorityYear="1992" box="[326,630,1411,1437]" class="Insecta" family="Staphylinidae" genus="Hygrodromicus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="389" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="1C97462D256E1635432FFA5AFDB298A1" box="[326,630,1411,1437]" italics="true" pageId="5" pageNumber="389">Hygrodromicus splendidus</emphasis>
</taxonomicName>
was originally described based on the
<typeStatus id="F158249D256E1635462DFA5AFB6298A1" box="[1092,1190,1411,1437]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
from
<collectingCountry id="56F4DAAF256E1635469BFA5AFAA498A1" box="[1266,1376,1411,1437]" name="Tajikistan" pageId="5" pageNumber="389">Tajikistan</collectingCountry>
(“… Pamir, Muksu Gebiet… Hochflâche bei Kishlak Ljarsch…”). The specimens studied by me are conspecific to the
<typeStatus id="F158249D256E163542FEFA12FF3898D9" box="[151,252,1483,1509]" pageId="5" pageNumber="389" type="holotype">holotype</typeStatus>
, but have some insignificant morphological differences. Thus, I decided to redescribed this species. It is here recorded from
<collectingCountry id="56F4DAAF256E1635431CFA36FE3C9B35" box="[373,504,1519,1545]" name="Kazakhstan" pageId="5" pageNumber="389">Kazakhstan</collectingCountry>
and
<collectingCountry id="56F4DAAF256E16354047FA36FD6B9B35" box="[558,687,1519,1545]" name="Kyrgyzstan" pageId="5" pageNumber="389">Kyrgyzstan</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,691 @@
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<mods:title id="4232CB2883C154ACFF5993510D8BDF10">Two new species of the genus Sycanus Amyot &amp; Serville (Insecta: Hemiptera: Reduviidae: Harpactorinae) from Vietnam</mods:title>
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<mods:namePart id="E37D866CE7308DE5A5780A5FBC76A448">Truong, Xuan Lam</mods:namePart>
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<mods:affiliation id="43B662ADB46DD3D65E52E38A2638A70E">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Hanoi, Vietnam, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:affiliation id="6FC7B88553E53A9EF5715BC41BB52CEF">Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam &amp; Vinh University, 182 Le Duan, Vinh, Nghe An, Vietnam</mods:affiliation>
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<mods:namePart id="C055AF8CD1234039EF3DCE043B2DC668">Nguyen, Dai Dac</mods:namePart>
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<mods:namePart id="972F3EF63D4B21C4D75AEFD483AADBB1">Ha, Ngoc Linh</mods:namePart>
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<taxonomicName id="F7D218ABE803FFBDFF55EF7FFD49896B" authority="Truong &amp; Ha" authorityName="Truong &amp; Giang &amp; Nguyen &amp; Chau &amp; Ha" authorityYear="2024" box="[151,648,587,614]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="thuathienhuensis" status="sp. nov.">
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Truong &amp; Ha
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,
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</subSubSection>
<subSubSection id="78C830A3E803FFBDFF55EF5BFE828987" box="[151,323,623,649]" pageId="12" pageNumber="313" type="description">
<paragraph id="306D6328E803FFBDFF55EF5BFE828987" blockId="12.[151,754,587,649]" box="[151,323,623,649]" pageId="12" pageNumber="313">
(
<figureCitation id="A8E97FADE803FFBDFF5DEF5BFEC28987" box="[159,259,623,649]" captionStart="FIGURE 4" captionStartId="9.[151,250,1203,1227]" captionTargetBox="[163,1422,193,1166]" captionTargetId="figure-18@9.[151,1436,181,1180]" captionTargetPageId="9" captionText="FIGURE 4. Body in dorsal view of two new morphospecies, α and β. A, Sycanus thuathienhuensis Truong &amp; Ha, sp. nov. (= α), paratype, ♂, TXL2024-009; B, S. taynguyenensis Truong &amp; Ha, sp. nov. (= β), holotype, ♂, NDD2019-285." figureDoi="http://doi.org/10.5281/zenodo.12758817" httpUri="https://zenodo.org/record/12758817/files/figure.png" pageId="12" pageNumber="313">Figs. 4A</figureCitation>
,
<figureCitation id="A8E97FADE803FFBDFECDEF5BFEDE8987" box="[271,287,623,649]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="12" pageNumber="313">6</figureCitation>
,
<figureCitation id="A8E97FADE803FFBDFEE9EF5BFEFB8987" box="[299,314,623,649]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="12" pageNumber="313">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="78C830A3E803FFBDFF55EF83FC358FC4" pageId="12" pageNumber="313" type="materials_examined">
<paragraph id="306D6328E803FFBDFF55EF83FD1E88C0" blockId="12.[151,1437,695,2018]" pageId="12" pageNumber="313">
<emphasis id="02A6BF3AE803FFBDFF55EF83FE8D89DF" bold="true" box="[151,332,695,721]" pageId="12" pageNumber="313">Type material.</emphasis>
<materialsCitation id="80BA6975E803FFBDFE9BEF83FDE589F8" collectingDate="2024-02-02" collectionCode="IEBR" collectorName="XL Truong" country="Vietnam" location="Bach Ma National Park" pageId="12" pageNumber="313" specimenCount="1" specimenCount-male="1" stateProvince="Thua Thien Hue Province" typeStatus="holotype">
<emphasis id="02A6BF3AE803FFBDFE9BEF83FE0C89DF" bold="true" box="[345,461,695,721]" pageId="12" pageNumber="313">
<typeStatus id="EF69DD8AE803FFBDFE9BEF83FE0989DF" box="[345,456,695,721]" pageId="12" pageNumber="313" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="26D4A8A1E803FFBDFE19EF8CFE3289DC" box="[475,499,696,722]" count="1" pageId="12" pageNumber="313" type="male"></specimenCount>
; TXL2024-012;
<collectingCountry id="48C523B8E803FFBDFD7EEF83FCDF89DF" box="[700,798,695,721]" name="Vietnam" pageId="12" pageNumber="313">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE803FFBDFCECEF83FBA689DF" box="[814,1127,695,722]" inferredBySuffix="true" pageId="12" pageNumber="313">Thua Thien Hue Province</collectingRegion>
,
<location id="350D35F3E803FFBDFBBBEF83FA5989DF" LSID="urn:lsid:plazi:treatment:B87BD23EE803FFA0FF55EF7FFDCB881B:350D35F3E803FFBDFBBBEF83FA5989DF" box="[1145,1432,695,722]" country="Vietnam" name="Bach Ma National Park" pageId="12" pageNumber="313" stateProvince="Thua Thien Hue Province">Bach Ma National Park</location>
;
<date id="446C45E8E803FFBDFF55EFEFFECC89FB" box="[151,269,731,757]" pageId="12" pageNumber="313" value="2024-02-02">
<collectingDate id="5428BC00E803FFBDFF55EFEFFECC89FB" box="[151,269,731,757]" pageId="12" pageNumber="313" value="2024-02-02">02.ii.2024</collectingDate>
</date>
;
<collectorName id="9D2706FEE803FFBDFED9EFE8FE5A89F8" box="[283,411,732,758]" pageId="12" pageNumber="313">XL Truong</collectorName>
leg.;
<collectionCode id="56C3FBEDE803FFBDFE23EFE8FDE589F8" box="[481,548,732,758]" pageId="12" pageNumber="313">IEBR</collectionCode>
</materialsCitation>
.
<materialsCitation id="80BA6975E803FFBDFDF1EFEFFD0D8814" collectingDate="2018-05-05" collectionCode="IEBR" collectorName="XL Truong" country="Vietnam" location="Kon Chu Rang Nature Reserve" pageId="12" pageNumber="313" specimenCount="1" specimenCount-male="1" stateProvince="Gia Lai" typeStatus="paratype">
<emphasis id="02A6BF3AE803FFBDFDF1EFEFFD7089FB" bold="true" box="[563,689,731,757]" pageId="12" pageNumber="313">
<typeStatus id="EF69DD8AE803FFBDFDF1EFEFFD6C89FB" box="[563,685,731,757]" pageId="12" pageNumber="313" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="26D4A8A1E803FFBDFD7EEFEFFD2089FB" box="[700,737,731,757]" count="1" pageId="12" pageNumber="313" type="male">1♂</specimenCount>
; TXL2018-004;
<collectingCountry id="48C523B8E803FFBDFC67EFEFFBC689FB" box="[933,1031,731,757]" name="Vietnam" pageId="12" pageNumber="313">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE803FFBDFBD7EFEFFB1F89FB" box="[1045,1246,731,757]" country="Vietnam" name="Gia Lai" pageId="12" pageNumber="313">Gia Lai Province</collectingRegion>
,
<location id="350D35F3E803FFBDFB2FEFE8FE898814" LSID="urn:lsid:plazi:treatment:B87BD23EE803FFA0FF55EF7FFDCB881B:350D35F3E803FFBDFB2FEFE8FE898814" country="Vietnam" name="Kon Chu Rang Nature Reserve" pageId="12" pageNumber="313" stateProvince="Gia Lai">Kon Chu Rang Nature Reserve</location>
;
<date id="446C45E8E803FFBDFE90EFCBFE058814" box="[338,452,767,794]" pageId="12" pageNumber="313" value="2018-05-05">
<collectingDate id="5428BC00E803FFBDFE90EFCBFE058814" box="[338,452,767,794]" pageId="12" pageNumber="313" value="2018-05-05">05.v.2018</collectingDate>
</date>
;
<collectorName id="9D2706FEE803FFBDFE0DEE34FD8B8814" box="[463,586,768,794]" pageId="12" pageNumber="313">XL Truong</collectorName>
leg.;
<collectionCode id="56C3FBEDE803FFBDFD4BEE34FD0D8814" box="[649,716,768,794]" pageId="12" pageNumber="313">IEBR</collectionCode>
</materialsCitation>
.
<materialsCitation id="80BA6975E803FFBDFD14EFCBFDB28830" collectingDate="2022-05-01" collectionCode="IEBR" collectorName="DD Nguyen" country="Vietnam" location="Tay Giang Forest" pageId="12" pageNumber="313" specimenCount="1" specimenCount-male="1" stateProvince="Quang Nam" typeStatus="paratype">
<specimenCount id="26D4A8A1E803FFBDFD14EFCBFD3B8817" box="[726,762,767,793]" count="1" pageId="12" pageNumber="313" type="male">1♂</specimenCount>
; NDD2022-009;
<collectingCountry id="48C523B8E803FFBDFC7FEFCBFBDE8817" box="[957,1055,767,793]" name="Vietnam" pageId="12" pageNumber="313">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE803FFBDFBEBEFCBFADC8817" box="[1065,1309,767,794]" country="Vietnam" name="Quang Nam" pageId="12" pageNumber="313">Quang Nam Province</collectingRegion>
,
<location id="350D35F3E803FFBDFAEAEE34FF208830" LSID="urn:lsid:plazi:treatment:B87BD23EE803FFA0FF55EF7FFDCB881B:350D35F3E803FFBDFAEAEE34FF208830" country="Vietnam" name="Tay Giang Forest" pageId="12" pageNumber="313" stateProvince="Quang Nam">Tay Giang Forest</location>
;
<date id="446C45E8E803FFBDFF2EEE17FE9C8833" box="[236,349,803,829]" pageId="12" pageNumber="313" value="2022-05-01">
<collectingDate id="5428BC00E803FFBDFF2EEE17FE9C8833" box="[236,349,803,829]" pageId="12" pageNumber="313" value="2022-05-01">01.v.2022</collectingDate>
</date>
;
<collectorName id="9D2706FEE803FFBDFEABEE10FE308830" box="[361,497,804,830]" pageId="12" pageNumber="313">DD Nguyen</collectorName>
leg.;
<collectionCode id="56C3FBEDE803FFBDFDF2EE10FDB28830" box="[560,627,804,830]" pageId="12" pageNumber="313">IEBR</collectionCode>
</materialsCitation>
.
<materialsCitation id="80BA6975E803FFBDFDBCEE17FDCC886C" collectingDate="2022-05-01" collectionCode="IEBR" collectorName="DD Nguyen" country="Vietnam" location="Tay Giang Forest" pageId="12" pageNumber="313" specimenCount="1" specimenCount-female="1" stateProvince="Quang Nam" typeStatus="paratype">
<specimenCount id="26D4A8A1E803FFBDFDBCEE17FD638833" box="[638,674,803,829]" count="1" pageId="12" pageNumber="313" type="female">1♀</specimenCount>
; NDD2022-063;
<collectingCountry id="48C523B8E803FFBDFCA4EE17FC098833" box="[870,968,803,829]" name="Vietnam" pageId="12" pageNumber="313">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE803FFBDFC10EE17FB068833" box="[978,1223,803,830]" country="Vietnam" name="Quang Nam" pageId="12" pageNumber="313">Quang Nam Province</collectingRegion>
,
<location id="350D35F3E803FFBDFB10EE10FA598830" LSID="urn:lsid:plazi:treatment:B87BD23EE803FFA0FF55EF7FFDCB881B:350D35F3E803FFBDFB10EE10FA598830" box="[1234,1432,803,830]" country="Vietnam" name="Tay Giang Forest" pageId="12" pageNumber="313" stateProvince="Quang Nam">Tay Giang Forest</location>
;
<date id="446C45E8E803FFBDFF55EE73FF3B886F" box="[151,250,839,865]" pageId="12" pageNumber="313" value="2022-05-01">
<collectingDate id="5428BC00E803FFBDFF55EE73FF3B886F" box="[151,250,839,865]" pageId="12" pageNumber="313" value="2022-05-01">1.v.2022</collectingDate>
</date>
;
<collectorName id="9D2706FEE803FFBDFEC7EE7CFE4C886C" box="[261,397,840,866]" pageId="12" pageNumber="313">DD Nguyen</collectorName>
leg.;
<collectionCode id="56C3FBEDE803FFBDFE08EE7CFDCC886C" box="[458,525,840,866]" pageId="12" pageNumber="313">IEBR</collectionCode>
</materialsCitation>
.
<materialsCitation id="80BA6975E803FFBDFDD5EE73FDDC8888" collectingDate="2024-02-02" collectionCode="IEBR" collectorName="XL Truong" country="Vietnam" location="Bach Ma National Park" pageId="12" pageNumber="313" specimenCount="1" specimenCount-female="1" stateProvince="Thua Thien Hue Province" typeStatus="paratype">
<specimenCount id="26D4A8A1E803FFBDFDD5EE73FDFA886F" box="[535,571,839,865]" count="1" pageId="12" pageNumber="313" type="female">1♀</specimenCount>
; TXL2024-059;
<collectingCountry id="48C523B8E803FFBDFD35EE73FC98886F" box="[759,857,839,865]" name="Vietnam" pageId="12" pageNumber="313">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE803FFBDFCA1EE73FB45886F" box="[867,1156,839,866]" inferredBySuffix="true" pageId="12" pageNumber="313">Thua Thien Hue Province</collectingRegion>
,
<location id="350D35F3E803FFBDFB4DEE73FA59886F" LSID="urn:lsid:plazi:treatment:B87BD23EE803FFA0FF55EF7FFDCB881B:350D35F3E803FFBDFB4DEE73FA59886F" box="[1167,1432,839,866]" country="Vietnam" name="Bach Ma National Park" pageId="12" pageNumber="313" stateProvince="Thua Thien Hue Province">Bach Ma National Park</location>
;
<date id="446C45E8E803FFBDFF55EE5FFECC888B" box="[151,269,875,901]" pageId="12" pageNumber="313" value="2024-02-02">
<collectingDate id="5428BC00E803FFBDFF55EE5FFECC888B" box="[151,269,875,901]" pageId="12" pageNumber="313" value="2024-02-02">02.ii.2024</collectingDate>
</date>
;
<collectorName id="9D2706FEE803FFBDFEDBEE58FE568888" box="[281,407,876,902]" pageId="12" pageNumber="313">XL Truong</collectorName>
leg.;
<collectionCode id="56C3FBEDE803FFBDFE18EE58FDDC8888" box="[474,541,876,902]" pageId="12" pageNumber="313">IEBR</collectionCode>
</materialsCitation>
.
<materialsCitation id="80BA6975E803FFBDFDEBEE5FFD1A88A4" collectingDate="2024-02-04" collectionCode="IEBR" collectorName="PT Giang" country="Vietnam" location="Bach Ma National Park" pageId="12" pageNumber="313" specimenCount="2" specimenCount-female="2" stateProvince="Thua Thien Hue Province" typeStatus="paratype">
<specimenCount id="26D4A8A1E803FFBDFDEBEE5FFD8F888B" box="[553,590,875,901]" count="2" pageId="12" pageNumber="313" type="female">2♀</specimenCount>
; PTG2024-060; PTG2024-061;
<collectingCountry id="48C523B8E803FFBDFC7CEE5FFBE1888B" box="[958,1056,875,901]" name="Vietnam" pageId="12" pageNumber="313">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE803FFBDFBEEEE5FFA94888B" box="[1068,1365,875,902]" inferredBySuffix="true" pageId="12" pageNumber="313">Thua Thien Hue Province</collectingRegion>
,
<location id="350D35F3E803FFBDFAA1EE5FFEA388A7" LSID="urn:lsid:plazi:treatment:B87BD23EE803FFA0FF55EF7FFDCB881B:350D35F3E803FFBDFAA1EE5FFEA388A7" country="Vietnam" name="Bach Ma National Park" pageId="12" pageNumber="313" stateProvince="Thua Thien Hue Province">Bach Ma National Park</location>
;
<date id="446C45E8E803FFBDFEAFEEBBFE2388A7" box="[365,482,911,937]" pageId="12" pageNumber="313" value="2024-02-04">
<collectingDate id="5428BC00E803FFBDFEAFEEBBFE2388A7" box="[365,482,911,937]" pageId="12" pageNumber="313" value="2024-02-04">04.ii.2024</collectingDate>
</date>
;
<collectorName id="9D2706FEE803FFBDFE2CEEA4FD9888A7" box="[494,601,911,938]" pageId="12" pageNumber="313">PT Giang</collectorName>
leg.;
<collectionCode id="56C3FBEDE803FFBDFD5AEEA4FD1A88A4" box="[664,731,912,938]" pageId="12" pageNumber="313">IEBR</collectionCode>
</materialsCitation>
.
<materialsCitation id="80BA6975E803FFBDFD24EEBBFD1A88C0" collectingDate="2024-02-05" collectionCode="IEBR" collectorName="PT Giang" country="Vietnam" location="Bach Ma National Park" pageId="12" pageNumber="313" specimenCount="1" specimenCount-female="1" stateProvince="Thua Thien Hue Province" typeStatus="paratype">
<specimenCount id="26D4A8A1E803FFBDFD24EEBBFCCB88A7" box="[742,778,911,937]" count="1" pageId="12" pageNumber="313" type="female">1♀</specimenCount>
; PTG2024-062;
<collectingCountry id="48C523B8E803FFBDFC05EEBBFBE888A7" box="[967,1065,911,937]" name="Vietnam" pageId="12" pageNumber="313">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE803FFBDFBF1EEBBFA9688A7" box="[1075,1367,911,938]" inferredBySuffix="true" pageId="12" pageNumber="313">Thua Thien Hue Province</collectingRegion>
,
<location id="350D35F3E803FFBDFAA1EEBBFEA388C3" LSID="urn:lsid:plazi:treatment:B87BD23EE803FFA0FF55EF7FFDCB881B:350D35F3E803FFBDFAA1EEBBFEA388C3" country="Vietnam" name="Bach Ma National Park" pageId="12" pageNumber="313" stateProvince="Thua Thien Hue Province">Bach Ma National Park</location>
;
<date id="446C45E8E803FFBDFEAFEE87FE2388C0" box="[365,482,947,974]" pageId="12" pageNumber="313" value="2024-02-05">
<collectingDate id="5428BC00E803FFBDFEAFEE87FE2388C0" box="[365,482,947,974]" pageId="12" pageNumber="313" value="2024-02-05">05.ii.2024</collectingDate>
</date>
;
<collectorName id="9D2706FEE803FFBDFE2FEE80FD9888C3" box="[493,601,947,974]" pageId="12" pageNumber="313">PT Giang</collectorName>
leg.;
<collectionCode id="56C3FBEDE803FFBDFD5AEE80FD1A88C0" box="[664,731,948,974]" pageId="12" pageNumber="313">IEBR</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="306D6328E803FFBDFF05EEE3FC358FC4" blockId="12.[151,1437,695,2018]" pageId="12" pageNumber="313">
<emphasis id="02A6BF3AE803FFBDFF05EEE3FE6688FF" bold="true" box="[199,423,983,1009]" pageId="12" pageNumber="313">Non-type material.</emphasis>
<materialsCitation id="80BA6975E803FFBDFE6EEEE3FE558F18" collectingDate="2016-04-26" collectionCode="IEBR" collectorName="XL Truong" country="Vietnam" location="Kon Chu Rang Nature Reserve" pageId="12" pageNumber="313" specimenCount="1" specimenCount-male="1" stateProvince="Gia Lai">
<specimenCount id="26D4A8A1E803FFBDFE6EEEE3FE1088FF" box="[428,465,983,1009]" count="1" pageId="12" pageNumber="313" type="male">1♂</specimenCount>
; TXL2016-514;
<collectingCountry id="48C523B8E803FFBDFD48EEE3FD2D88FF" box="[650,748,983,1009]" name="Vietnam" pageId="12" pageNumber="313">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE803FFBDFD37EEE3FC7288FF" box="[757,947,983,1009]" country="Vietnam" name="Gia Lai" pageId="12" pageNumber="313">Gia Lai Province</collectingRegion>
,
<location id="350D35F3E803FFBDFC7EEEECFAD488FC" LSID="urn:lsid:plazi:treatment:B87BD23EE803FFA0FF55EF7FFDCB881B:350D35F3E803FFBDFC7EEEECFAD488FC" box="[956,1301,983,1010]" country="Vietnam" name="Kon Chu Rang Nature Reserve" pageId="12" pageNumber="313" stateProvince="Gia Lai">Kon Chu Rang Nature Reserve</location>
;
<date id="446C45E8E803FFBDFADDEEE3FA5988FF" box="[1311,1432,983,1009]" pageId="12" pageNumber="313" value="2016-04-26">
<collectingDate id="5428BC00E803FFBDFADDEEE3FA5988FF" box="[1311,1432,983,1009]" pageId="12" pageNumber="313" value="2016-04-26">26.iv.2016</collectingDate>
</date>
;
<collectorName id="9D2706FEE803FFBDFF55EEC8FED38F18" box="[151,274,1020,1046]" pageId="12" pageNumber="313">XL Truong</collectorName>
leg.;
<collectionCode id="56C3FBEDE803FFBDFE93EEC8FE558F18" box="[337,404,1020,1046]" pageId="12" pageNumber="313">IEBR</collectionCode>
</materialsCitation>
.
<materialsCitation id="80BA6975E803FFBDFE5CEECFFE528F34" collectingDate="2016-04-27" collectionCode="IEBR" collectorName="XL Truong" country="Vietnam" location="Kon Chu Rang Nature Reserve" pageId="12" pageNumber="313" specimenCount="1" specimenCount-female="1" stateProvince="Gia Lai">
<specimenCount id="26D4A8A1E803FFBDFE5CEECFFE038F1B" box="[414,450,1019,1045]" count="1" pageId="12" pageNumber="313" type="female">1♀</specimenCount>
; TXL2016-521;
<collectingCountry id="48C523B8E803FFBDFDBCEECFFD218F1B" box="[638,736,1019,1045]" name="Vietnam" pageId="12" pageNumber="313">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE803FFBDFD28EECFFC6A8F1B" box="[746,939,1019,1045]" country="Vietnam" name="Gia Lai" pageId="12" pageNumber="313">Gia Lai Province</collectingRegion>
,
<location id="350D35F3E803FFBDFC74EEC8FAD58F18" LSID="urn:lsid:plazi:treatment:B87BD23EE803FFA0FF55EF7FFDCB881B:350D35F3E803FFBDFC74EEC8FAD58F18" box="[950,1300,1019,1046]" country="Vietnam" name="Kon Chu Rang Nature Reserve" pageId="12" pageNumber="313" stateProvince="Gia Lai">Kon Chu Rang Nature Reserve</location>
;
<date id="446C45E8E803FFBDFADCEECFFA598F1B" box="[1310,1432,1019,1045]" pageId="12" pageNumber="313" value="2016-04-27">
<collectingDate id="5428BC00E803FFBDFADCEECFFA598F1B" box="[1310,1432,1019,1045]" pageId="12" pageNumber="313" value="2016-04-27">27.iv.2016</collectingDate>
</date>
;
<collectorName id="9D2706FEE803FFBDFF55E914FED38F34" box="[151,274,1056,1082]" pageId="12" pageNumber="313">XL Truong</collectorName>
leg.;
<collectionCode id="56C3FBEDE803FFBDFE92E914FE528F34" box="[336,403,1056,1082]" pageId="12" pageNumber="313">IEBR</collectionCode>
</materialsCitation>
.
<materialsCitation id="80BA6975E803FFBDFE5CE92BFEAA8F50" collectingDate="2016-05-04" collectionCode="IEBR" collectorName="XL Truong" country="Vietnam" location="Chu Yang Sin National Park" pageId="12" pageNumber="313" specimenCount="1" specimenCount-female="1" stateProvince="Dak Lak">
<specimenCount id="26D4A8A1E803FFBDFE5CE92BFE038F37" box="[414,450,1055,1081]" count="1" pageId="12" pageNumber="313" type="female">1♀</specimenCount>
; TXL2016-604;
<collectingCountry id="48C523B8E803FFBDFDBCE92BFD218F37" box="[638,736,1055,1081]" name="Vietnam" pageId="12" pageNumber="313">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE803FFBDFD28E92BFC768F37" box="[746,951,1055,1081]" country="Vietnam" name="Dak Lak" pageId="12" pageNumber="313">Dak Lak Province</collectingRegion>
,
<location id="350D35F3E803FFBDFC00E92BFB3E8F37" LSID="urn:lsid:plazi:treatment:B87BD23EE803FFA0FF55EF7FFDCB881B:350D35F3E803FFBDFC00E92BFB3E8F37" box="[962,1279,1055,1082]" country="Vietnam" name="Chu Yang Sin National Park" pageId="12" pageNumber="313" stateProvince="Dak Lak">Chu Yang Sin National Park</location>
;
<date id="446C45E8E803FFBDFAC8E92BFAAC8F37" box="[1290,1389,1055,1081]" pageId="12" pageNumber="313" value="2016-05-04">
<collectingDate id="5428BC00E803FFBDFAC8E92BFAAC8F37" box="[1290,1389,1055,1081]" pageId="12" pageNumber="313" value="2016-05-04">4.v.2016</collectingDate>
</date>
;
<collectorName id="9D2706FEE803FFBDFABAE914FF298F50" pageId="12" pageNumber="313">XL Truong</collectorName>
leg.;
<collectionCode id="56C3FBEDE803FFBDFEEAE970FEAA8F50" box="[296,363,1092,1118]" pageId="12" pageNumber="313">IEBR</collectionCode>
</materialsCitation>
.
<materialsCitation id="80BA6975E803FFBDFEB4E977FC508F8C" collectingDate="2018-05-05" collectionCode="IEBR" collectorName="XL Truong" country="Vietnam" location="Kon Chu Rang Nature Reserve" pageId="12" pageNumber="313" specimenCount="4" specimenCount-female="4" stateProvince="Gia Lai">
<specimenCount id="26D4A8A1E803FFBDFEB4E977FE5A8F53" box="[374,411,1091,1117]" count="4" pageId="12" pageNumber="313" type="female">4♀</specimenCount>
; TXL2018-001; TXL2018-002; TXL2018-003; TXL2018-005;
<collectingCountry id="48C523B8E803FFBDFBABE977FB0A8F53" box="[1129,1227,1091,1117]" name="Vietnam" pageId="12" pageNumber="313">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE803FFBDFB14E977FA598F53" box="[1238,1432,1091,1117]" country="Vietnam" name="Gia Lai" pageId="12" pageNumber="313">Gia Lai Province</collectingRegion>
,
<location id="350D35F3E803FFBDFF55E95CFDC18F8C" LSID="urn:lsid:plazi:treatment:B87BD23EE803FFA0FF55EF7FFDCB881B:350D35F3E803FFBDFF55E95CFDC18F8C" box="[151,512,1127,1154]" country="Vietnam" name="Kon Chu Rang Nature Reserve" pageId="12" pageNumber="313" stateProvince="Gia Lai">Kon Chu Rang Nature Reserve</location>
;
<date id="446C45E8E803FFBDFDCFE953FDBE8F8C" box="[525,639,1127,1154]" pageId="12" pageNumber="313" value="2018-05-05">
<collectingDate id="5428BC00E803FFBDFDCFE953FDBE8F8C" box="[525,639,1127,1154]" pageId="12" pageNumber="313" value="2018-05-05">05.v.2018</collectingDate>
</date>
;
<collectorName id="9D2706FEE803FFBDFD4FE95CFCCA8F8C" box="[653,779,1128,1154]" pageId="12" pageNumber="313">XL Truong</collectorName>
leg.;
<collectionCode id="56C3FBEDE803FFBDFC8CE95CFC508F8C" box="[846,913,1128,1154]" pageId="12" pageNumber="313">IEBR</collectionCode>
</materialsCitation>
.
<materialsCitation id="80BA6975E803FFBDFC5CE953FB808FA8" collectingDate="2018-05-08" collectionCode="IEBR" collectorName="XL Truong" country="Vietnam" location="Chu Yang Sin National Park" pageId="12" pageNumber="313" specimenCount="2" specimenCount-female="2" stateProvince="Dak Lak">
<specimenCount id="26D4A8A1E803FFBDFC5CE953FC028F8F" box="[926,963,1127,1153]" count="2" pageId="12" pageNumber="313" type="female">2♀</specimenCount>
; TXL2018-043; TXL2018-044;
<collectingCountry id="48C523B8E803FFBDFAF4E953FA598F8F" box="[1334,1432,1127,1153]" name="Vietnam" pageId="12" pageNumber="313">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE803FFBDFF55E9BFFEA68FAB" box="[151,359,1163,1189]" country="Vietnam" name="Dak Lak" pageId="12" pageNumber="313">Dak Lak Province</collectingRegion>
,
<location id="350D35F3E803FFBDFEB1E9BFFD768FAB" LSID="urn:lsid:plazi:treatment:B87BD23EE803FFA0FF55EF7FFDCB881B:350D35F3E803FFBDFEB1E9BFFD768FAB" box="[371,695,1163,1190]" country="Vietnam" name="Chu Yang Sin National Park" pageId="12" pageNumber="313" stateProvince="Dak Lak">Chu Yang Sin National Park</location>
;
<date id="446C45E8E803FFBDFD01E9BFFCF58FAB" box="[707,820,1163,1189]" pageId="12" pageNumber="313" value="2018-05-08">
<collectingDate id="5428BC00E803FFBDFD01E9BFFCF58FAB" box="[707,820,1163,1189]" pageId="12" pageNumber="313" value="2018-05-08">08.v.2018</collectingDate>
</date>
;
<collectorName id="9D2706FEE803FFBDFC83E9B8FC7C8FA8" box="[833,957,1164,1190]" pageId="12" pageNumber="313">XL Truong</collectorName>
leg.;
<collectionCode id="56C3FBEDE803FFBDFC3CE9B8FB808FA8" box="[1022,1089,1164,1190]" pageId="12" pageNumber="313">IEBR</collectionCode>
</materialsCitation>
.
<materialsCitation id="80BA6975E803FFBDFB8FE9BFFC2E8FC4" collectingDate="2022-05-02" collectionCode="IEBR" collectorName="DD Nguyen" country="Vietnam" location="Tay Giang Forest" pageId="12" pageNumber="313" specimenCount="1" specimenCount-female="1" stateProvince="Quang Nam">
<specimenCount id="26D4A8A1E803FFBDFB8FE9BFFBB08FAB" box="[1101,1137,1163,1189]" count="1" pageId="12" pageNumber="313" type="female">1♀</specimenCount>
; NDD2022-024;
<collectingCountry id="48C523B8E803FFBDFAF4E9BFFA598FAB" box="[1334,1432,1163,1189]" name="Vietnam" pageId="12" pageNumber="313">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE803FFBDFF55E99BFE4D8FC7" box="[151,396,1199,1226]" country="Vietnam" name="Quang Nam" pageId="12" pageNumber="313">Quang Nam Province</collectingRegion>
,
<location id="350D35F3E803FFBDFE55E984FD9C8FC4" LSID="urn:lsid:plazi:treatment:B87BD23EE803FFA0FF55EF7FFDCB881B:350D35F3E803FFBDFE55E984FD9C8FC4" box="[407,605,1199,1226]" country="Vietnam" name="Tay Giang Forest" pageId="12" pageNumber="313" stateProvince="Quang Nam">Tay Giang Forest</location>
;
<date id="446C45E8E803FFBDFDAAE99BFD1B8FC7" box="[616,730,1199,1225]" pageId="12" pageNumber="313" value="2022-05-02">
<collectingDate id="5428BC00E803FFBDFDAAE99BFD1B8FC7" box="[616,730,1199,1225]" pageId="12" pageNumber="313" value="2022-05-02">02.v.2022</collectingDate>
</date>
;
<collectorName id="9D2706FEE803FFBDFD27E984FCAC8FC4" box="[741,877,1200,1226]" pageId="12" pageNumber="313">DD Nguyen</collectorName>
leg.;
<collectionCode id="56C3FBEDE803FFBDFC6EE984FC2E8FC4" box="[940,1007,1200,1226]" pageId="12" pageNumber="313">IEBR</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="78C830A3E803FFBEFF05E9E7FDA98C7F" lastPageId="15" lastPageNumber="316" pageId="12" pageNumber="313" type="diagnosis">
<paragraph id="306D6328E803FFBDFF05E9E7FADE8EAF" blockId="12.[151,1437,695,2018]" pageId="12" pageNumber="313">
<emphasis id="02A6BF3AE803FFBDFF05E9E7FE838FE3" bold="true" box="[199,322,1235,1261]" pageId="12" pageNumber="313">Diagnosis.</emphasis>
Body ochreous; anterior pronotal lobe black with some rows of short bent cream-yellow short and slender setae (
<figureCitation id="A8E97FADE803FFBDFEFFE9C3FE578E1F" box="[317,406,1271,1297]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="12" pageNumber="313">Fig. 6F</figureCitation>
); posterior pronotal lobe ochreous, rugulose, punctured, covered with short erect slender setae (
<figureCitation id="A8E97FADE803FFBDFF1DE82FFEF58E3B" box="[223,308,1307,1333]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="12" pageNumber="313">Fig. 6F</figureCitation>
); scutellum black with yellow latero-posterior margin (
<figureCitation id="A8E97FADE803FFBDFC62E82FFC378E3B" box="[928,1014,1307,1333]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="12" pageNumber="313">Fig. 6F</figureCitation>
); scutellar spine sub-vertical, short or almost tubercle-liked (
<quantity id="F72ACECDE803FFBDFE57E80BFDFE8E57" box="[405,575,1343,1370]" metricMagnitude="-4" metricUnit="m" metricValue="9.85" metricValueMax="15.3" metricValueMin="4.4" pageId="12" pageNumber="313" unit="mm" value="0.985" valueMax="1.53" valueMin="0.44">0.441.53 mm</quantity>
), not bifid at apex, yellow or brownish-yellow with black base (
<figureCitation id="A8E97FADE803FFBDFAD4E80BFA4E8E57" box="[1302,1423,1343,1369]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="12" pageNumber="313">Fig. 6B, F</figureCitation>
); coria yellow or ochreous with transection with clavus dark brown, clavus dark brown (
<figureCitation id="A8E97FADE803FFBDFBB6E857FB118E73" box="[1140,1232,1379,1405]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="12" pageNumber="313">Fig. 6G</figureCitation>
); male pygophore brownish yellow with ventrally centrally largely suffused with dark brown or blackish brown (
<figureCitation id="A8E97FADE803FFBDFB7BE8B3FAD08EAF" box="[1209,1297,1415,1441]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="12" pageNumber="313">Fig. 6C</figureCitation>
).
</paragraph>
<paragraph id="306D6328E803FFBDFF05E89FFA568C07" blockId="12.[151,1437,695,2018]" pageId="12" pageNumber="313">
This species is very similar to
<taxonomicName id="F7D218ABE803FFBDFDE8E89FFD248ECB" authorityName="Hsiao" authorityYear="1979" box="[554,741,1451,1477]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="croceus">
<emphasis id="02A6BF3AE803FFBDFDE8E89FFD248ECB" box="[554,741,1451,1477]" italics="true" pageId="12" pageNumber="313">Sycanus croceus</emphasis>
</taxonomicName>
in general appearance, especially in body colour, pronotum, and thoracic sterna. But the new species can be distinguished from
<taxonomicName id="F7D218ABE803FFBDFC58E8FBFBCF8EE7" authorityName="Hsiao" authorityYear="1979" box="[922,1038,1487,1513]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="croceus">
<emphasis id="02A6BF3AE803FFBDFC58E8FBFBCF8EE7" box="[922,1038,1487,1513]" italics="true" pageId="12" pageNumber="313">S. croceus</emphasis>
</taxonomicName>
by a combination of the following characteristics: a brownish yellow stripe running between postero-upper corner of compound eye and ocellus of same side (
<figureCitation id="A8E97FADE803FFBDFEDBEB23FEB08D3F" box="[281,369,1559,1585]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="12" pageNumber="313">Fig. 6E</figureCitation>
) (in
<taxonomicName id="F7D218ABE803FFBDFE68EB23FDDE8D3F" authorityName="Hsiao" authorityYear="1979" box="[426,543,1559,1585]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="croceus">
<emphasis id="02A6BF3AE803FFBDFE68EB23FDDE8D3F" box="[426,543,1559,1585]" italics="true" pageId="12" pageNumber="313">S. croceus</emphasis>
</taxonomicName>
without any yellow stripe), abdominal sternites with small longitudinal black markings segmentally laterally (
<figureCitation id="A8E97FADE803FFBDFE3CEB0FFD928D5B" box="[510,595,1595,1621]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="12" pageNumber="313">Fig. 6C</figureCitation>
) (in
<taxonomicName id="F7D218ABE803FFBDFD45EB0FFD398D5B" authorityName="Hsiao" authorityYear="1979" box="[647,760,1595,1621]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="croceus">
<emphasis id="02A6BF3AE803FFBDFD45EB0FFD398D5B" box="[647,760,1595,1621]" italics="true" pageId="12" pageNumber="313">S. croceus</emphasis>
</taxonomicName>
with small round spots segmentally laterally), large marking of laterotergite V reach antero-lateral margin of laterotergite V (in
<taxonomicName id="F7D218ABE803FFBDFCB2EB6BFC208D77" authorityName="Hsiao" authorityYear="1979" box="[880,993,1631,1657]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="croceus">
<emphasis id="02A6BF3AE803FFBDFCB2EB6BFC208D77" box="[880,993,1631,1657]" italics="true" pageId="12" pageNumber="313">S. croceus</emphasis>
</taxonomicName>
not extend to the antero-lateral margin), scutellar spine short or almost tubercle-liked, not bifid at apex, yellow or brownish-yellow with black base (
<figureCitation id="A8E97FADE803FFBDFAA6EBB7FF168DCF" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="12" pageNumber="313">Figs. 6B, F</figureCitation>
) (in
<taxonomicName id="F7D218ABE803FFBDFECEEB93FEBF8DCF" authorityName="Hsiao" authorityYear="1979" box="[268,382,1703,1729]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="croceus">
<emphasis id="02A6BF3AE803FFBDFECEEB93FEBF8DCF" box="[268,382,1703,1729]" italics="true" pageId="12" pageNumber="313">S. croceus</emphasis>
</taxonomicName>
long, apex bi-forked, wholly yellow or orange), distal dorsal lobe of endosoma (ddl) with two very large and long horn-shaped prickles centrally, surrounded by large prickles (
<figureCitation id="A8E97FADE803FFBDFBE1EBFFFB538DEB" box="[1059,1170,1739,1765]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="12" pageNumber="313">Fig. 7IK</figureCitation>
) (in
<taxonomicName id="F7D218ABE803FFBDFB05EBFFFAF88DEB" authorityName="Hsiao" authorityYear="1979" box="[1223,1337,1739,1765]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="croceus">
<emphasis id="02A6BF3AE803FFBDFB05EBFFFAF88DEB" box="[1223,1337,1739,1765]" italics="true" pageId="12" pageNumber="313">S. croceus</emphasis>
</taxonomicName>
ddl with two large horned processes, apical part laterally with three pairs of larger spines and eight pairs of smaller spines).
</paragraph>
<paragraph id="306D6328E803FFBDFF05EA20FAC38CEF" blockId="12.[151,1437,695,2018]" pageId="12" pageNumber="313">
Moreover, the new species is also similar to
<taxonomicName id="F7D218ABE803FFBDFD06EA27FC4B8C23" authorityName="Hsiao" authorityYear="1979" box="[708,906,1811,1837]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="02A6BF3AE803FFBDFD06EA27FC4B8C23" box="[708,906,1811,1837]" italics="true" pageId="12" pageNumber="313">Sycanus insularis</emphasis>
</taxonomicName>
in general morphology. But the former can be distinguished from the latter by the following features: abdominal sternites with small longitudinal black markings segmentally laterally (
<figureCitation id="A8E97FADE803FFBDFE4DEA6FFE248C7B" box="[399,485,1883,1909]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="12" pageNumber="313">Fig. 6C</figureCitation>
) (in
<taxonomicName id="F7D218ABE803FFBDFDDBEA6FFD578C7B" authorityName="Hsiao" authorityYear="1979" box="[537,662,1883,1909]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="02A6BF3AE803FFBDFDDBEA6FFD578C7B" box="[537,662,1883,1909]" italics="true" pageId="12" pageNumber="313">S. insularis</emphasis>
</taxonomicName>
with small round spots segmentally laterally), scutellar spine short or almost tubercle-liked, not bifid at apex, yellow or brownish-yellow with black base (
<figureCitation id="A8E97FADE803FFBDFBFEEA4BFB6F8C97" box="[1084,1198,1919,1945]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="12" pageNumber="313">Fig. 6B, F</figureCitation>
) (in
<taxonomicName id="F7D218ABE803FFBDFB23EA4BFA9F8C97" authorityName="Hsiao" authorityYear="1979" box="[1249,1374,1919,1945]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="02A6BF3AE803FFBDFB23EA4BFA9F8C97" box="[1249,1374,1919,1945]" italics="true" pageId="12" pageNumber="313">S. insularis</emphasis>
</taxonomicName>
long, apex bifid, black or yellow), coria yellow or ochreous with transection with clavus dark brown, clavus dark brown brown (
<figureCitation id="A8E97FADE803FFBDFF2CEAF3FE878CEF" box="[238,326,1991,2017]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="12" pageNumber="313">Fig. 6G</figureCitation>
) (in
<taxonomicName id="F7D218ABE803FFBDFEB9EAF3FE388CEF" authorityName="Hsiao" authorityYear="1979" box="[379,505,1991,2017]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="02A6BF3AE803FFBDFEB9EAF3FE388CEF" box="[379,505,1991,2017]" italics="true" pageId="12" pageNumber="313">S. insularis</emphasis>
</taxonomicName>
anterior half black or dark brown, remaining pale yellow or luteous).
</paragraph>
<caption id="64AD33A0E802FFBCFF55EA70FDDF8CAA" ID-DOI="http://doi.org/10.5281/zenodo.12758821" ID-Zenodo-Dep="12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="13" pageNumber="314" startId="13.[151,250,1860,1884]" targetBox="[151,1435,191,1836]" targetPageId="13" targetType="figure">
<paragraph id="306D6328E802FFBCFF55EA70FDDF8CAA" blockId="13.[151,1437,1860,1956]" pageId="13" pageNumber="314">
<emphasis id="02A6BF3AE802FFBCFF55EA70FED28C52" bold="true" box="[151,275,1860,1884]" pageId="13" pageNumber="314">FIGURE 6.</emphasis>
<taxonomicName id="F7D218ABE802FFBCFED9EA71FD718C52" authority="Truong &amp; Ha" authorityName="Truong &amp; Giang &amp; Nguyen &amp; Chau &amp; Ha" authorityYear="2024" box="[283,688,1860,1885]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="thuathienhuensis" status="sp. nov.">
<emphasis id="02A6BF3AE802FFBCFED9EA71FDE18C52" box="[283,544,1860,1884]" italics="true" pageId="13" pageNumber="314">Sycanus thuathienhuensis</emphasis>
Truong &amp; Ha
</taxonomicName>
,
<emphasis id="02A6BF3AE802FFBCFD7FEA71FCCF8C53" bold="true" box="[701,782,1861,1885]" pageId="13" pageNumber="314">
<taxonomicNameLabel id="19950241E802FFBCFD7FEA71FCCF8C53" box="[701,782,1861,1885]" pageId="13" pageNumber="314" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="02A6BF3AE802FFBCFCDEEA70FBF58C53" box="[796,1076,1860,1887]" italics="true" pageId="13" pageNumber="314">holotype, ♂, TXL2024-012.</emphasis>
A,
<emphasis id="02A6BF3AE802FFBCFB98EA70FAED8C52" box="[1114,1324,1860,1884]" italics="true" pageId="13" pageNumber="314">body in dorsal view;</emphasis>
B,
<emphasis id="02A6BF3AE802FFBCFA92EA70FED98C8E" italics="true" pageId="13" pageNumber="314">body in lateral view;</emphasis>
C, body
<emphasis id="02A6BF3AE802FFBCFEB3EA5DFCC38C8E" box="[369,770,1896,1920]" italics="true" pageId="13" pageNumber="314">in ventral view; D, head in lateral view;</emphasis>
E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing.
</paragraph>
</caption>
<paragraph id="306D6328E801FFBFFF05EDA3FB4F8A87" blockId="14.[151,1436,151,393]" pageId="14" pageNumber="315">
Furthermore, the new species has a very close relationship to
<taxonomicName id="F7D218ABE801FFBFFCB0EDA3FBCF8BBF" authorityName="Hsiao" authorityYear="1979" box="[882,1038,151,177]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="02A6BF3AE801FFBFFCB0EDA3FBCF8BBF" box="[882,1038,151,177]" italics="true" pageId="14" pageNumber="315">Sycanus rufus</emphasis>
</taxonomicName>
in molecular phylogenetic analyses with a minimum interspecific divergence of 2.0% in p-distance and
<quantity id="F72ACECDE801FFBFFC55ED8FFC278BDB" box="[919,998,187,213]" metricMagnitude="-4" metricUnit="m" metricValue="5.08" pageId="14" pageNumber="315" unit="in" value="0.02">0.02 in</quantity>
K
<specimenCount id="26D4A8A1E801FFBFFBC0ED8FFBE58BDB" box="[1026,1060,187,213]" count="2" pageId="14" pageNumber="315" type="generic">2P.</specimenCount>
However, the two species can be delimited by the external morphology, i.e., body ochreous (in
<taxonomicName id="F7D218ABE801FFBFFC92EDEBFC668BF7" authorityName="Hsiao" authorityYear="1979" box="[848,935,223,249]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="02A6BF3AE801FFBFFC92EDEBFC668BF7" box="[848,935,223,249]" italics="true" pageId="14" pageNumber="315">S. rufus</emphasis>
</taxonomicName>
body red), posterior pronotal lobe ochreous, rugulose, punctured, covered with short erect slender setae (
<figureCitation id="A8E97FADE801FFBFFC94EC37FC6C8A13" box="[854,941,259,285]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6F</figureCitation>
) (in
<taxonomicName id="F7D218ABE801FFBFFC28EC37FB858A13" authorityName="Hsiao" authorityYear="1979" box="[1002,1092,259,285]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="02A6BF3AE801FFBFFC28EC37FB858A13" box="[1002,1092,259,285]" italics="true" pageId="14" pageNumber="315">S. rufus</emphasis>
</taxonomicName>
red, rugulose, not punctured, clothes with procumbent short yellow setae), distal dorsal lobe of endosoma (ddl) with two very large and long horn-shaped prickles centrally, surrounded by large prickles (
<figureCitation id="A8E97FADE801FFBFFC86EC7FFC7F8A6B" box="[836,958,331,357]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="14" pageNumber="315">Figs. 7IK</figureCitation>
) (in
<taxonomicName id="F7D218ABE801FFBFFC31EC7FFB888A6B" authorityName="Hsiao" authorityYear="1979" box="[1011,1097,331,357]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="rufus">
<emphasis id="02A6BF3AE801FFBFFC31EC7FFB888A6B" box="[1011,1097,331,357]" italics="true" pageId="14" pageNumber="315">S. rufus</emphasis>
</taxonomicName>
ddl with two long and slender prickles, and laterally with two pairs of larger spines and about 10 pairs of smaller spines).
</paragraph>
<paragraph id="306D6328E801FFBFFF55ECEFFEA98AFA" blockId="14.[151,360,474,501]" box="[151,360,474,501]" pageId="14" pageNumber="315">
<emphasis id="02A6BF3AE801FFBFFF55ECEFFEA98AFA" bold="true" box="[151,360,474,501]" pageId="14" pageNumber="315">Male description.</emphasis>
</paragraph>
<paragraph id="306D6328E801FFBFFF55EF16FC788FCB" blockId="14.[151,1437,546,2013]" pageId="14" pageNumber="315">
<emphasis id="02A6BF3AE801FFBFFF55EF16FEDE8932" bold="true" box="[151,287,546,572]" pageId="14" pageNumber="315">Coloration.</emphasis>
Body ochreous. Head dorsum, neck, clypeus, maxillary plate, antenniferous tubercles, and posterior half of head venter black; labrum and anterior margin of gena dark brown; gena and anterior half of head venter luteous or yellow; a brownish yellow stripe running between postero-upper corner of compound eye and ocellus of same side; first visible labial segment blackish brown at basal half, paler as dark brown toward tip; base of second visible labial segment dark brown; remaining of labium brown, paler toward tip (
<figureCitation id="A8E97FADE801FFBFFBF3EF87FB6C89C3" box="[1073,1197,691,717]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6D, E</figureCitation>
). Scape, pedicel, and first flagellomere blackish brown; second flagellomere dark brown, paler toward tip. Collar, anterolateral angle, anterior margin of anterior pronotal lobe, acetabulum, prosternum, and stridulatory sulcus yellow (
<figureCitation id="A8E97FADE801FFBFFB10EFCFFAE7881B" box="[1234,1318,763,789]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6F</figureCitation>
); posterior pronotal lobe yellow or ochreous (
<figureCitation id="A8E97FADE801FFBFFDD5EE2BFDAD8837" box="[535,620,799,825]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6F</figureCitation>
); scutellum black with yellow latero-posterior margin (
<figureCitation id="A8E97FADE801FFBFFB16EE2BFAE88837" box="[1236,1321,799,825]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6F</figureCitation>
); scutellar spine yellow or brownish-yellow with black base or wholly black (
<figureCitation id="A8E97FADE801FFBFFC67EE77FC3F8853" box="[933,1022,835,861]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6F</figureCitation>
); meso- and metapleura, meso- and metasterna, black or blackish brown; coxae yellow; trochanters blackish brown; femora, tibiae, and tarsi black or blackish brown. Coria yellow or ochreous with transection with clavus dark brown; clavus dark brown; membrane pale bronzy brown, semi-hyaline (
<figureCitation id="A8E97FADE801FFBFFDDCEE9BFDB888C7" box="[542,633,943,969]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6G</figureCitation>
). Hind wings faintly semi-hyaline (
<figureCitation id="A8E97FADE801FFBFFBCCEE9BFBAB88C7" box="[1038,1130,943,969]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6H</figureCitation>
). Abdominal mediotergites blackish brown or black with irregular brown or yellow suffusions; laterotergites yellow or orangish yellow with lateral half of red, reddish yellow, or orange (
<figureCitation id="A8E97FADE801FFBFFD51EEC3FD2A8F1F" box="[659,747,1015,1041]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6B</figureCitation>
); laterotegites IIIVII segmentally large black marking; large marking of laterotergites VI and V reach antero-lateral margin of latetotergites; large marking of laterotergites III, VI, and VII far from reaching the lateral margin of laterotergites; abdominal sternites yellow or luteous with broad transversal black strip in anterior margin of each segment and discontinuous medial longitudinal black strip and small longitudinal black markings segmentally laterally (
<figureCitation id="A8E97FADE801FFBFFC89E9B3FC648FAF" box="[843,933,1159,1185]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6C</figureCitation>
). Pygophore brownish yellow with ventrally centrally largely suffused with dark brown or blackish brown (
<figureCitation id="A8E97FADE801FFBFFC91E99FFC6D8FCB" box="[851,940,1195,1221]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6C</figureCitation>
).
</paragraph>
<paragraph id="306D6328E801FFBEFF05E9FAFE48896F" blockId="14.[151,1437,546,2013]" lastBlockId="15.[151,1437,151,1077]" lastPageId="15" lastPageNumber="316" pageId="14" pageNumber="315">
<emphasis id="02A6BF3AE801FFBFFF05E9FAFE808FE6" bold="true" box="[199,321,1230,1256]" pageId="14" pageNumber="315">Structure.</emphasis>
Body large-sized (
<quantity id="F72ACECDE801FFBFFDEEE9FBFD398FE7" box="[556,760,1231,1257]" metricMagnitude="-2" metricUnit="m" metricValue="1.8239999999999998" metricValueMax="1.908" metricValueMin="1.7399999999999998" pageId="14" pageNumber="315" unit="mm" value="18.24" valueMax="19.08" valueMin="17.4">17.4019.08 mm</quantity>
), elongated and posteriorly widened (
<figureCitation id="A8E97FADE801FFBFFB01E9FBFA878FE7" box="[1219,1350,1231,1257]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6AC</figureCitation>
). Head tubular, slender, elongated, head and neck together much longer than pronotum (
<figureCitation id="A8E97FADE801FFBFFBFAE9C7FB768E03" box="[1080,1207,1267,1293]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6A, B</figureCitation>
); anteocular area of head elongate-conical; anteclypeus cylindrical and prominent; postocular area of head weakly globose, distinctly wider than anteocular area, shorter than anteocular area, constricted behind compound eyes, with a wide and deep interocular sulcus; neck long (
<figureCitation id="A8E97FADE801FFBFFE3BE86BFDBB8E77" box="[505,634,1375,1401]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6D, E</figureCitation>
). Compound eyes protruding laterally, nearly globose, with posterior margin sub-straight, oblique with respect to ventral margin of head; lateral ocelli produced, slightly elevated behind interocular sulcus, separated from each other; interspace between lateral ocelli wider than distance between compound eye and lateral ocellus (
<figureCitation id="A8E97FADE801FFBFFDE1E8FFFD5E8EEB" box="[547,671,1483,1509]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6D, E</figureCitation>
). First visible labial segment slightly thicker and much shorter than second segment, longer than anteocular area of head, not extending beyond level of middle of compound eye when labium laid backward (
<figureCitation id="A8E97FADE801FFBFFE63EB27FE3F8D23" box="[417,510,1555,1581]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6D</figureCitation>
); proportional average length of first to third visible labial segments 2.4:3.7:0.6. Scape about 1.6 times as long as head, about 2.5 times as long as pedicel, almost as long as second flagellomere; first flagellomere slightly shorter than pedicel and about 1/3 times as long as second flagellomere; proportional average length of scape, pedicel, first and second flagellomeres 7.0:2.9:2.3:6.8. Collar thick in dorsal view, with anterolateral angle roundly produced anteriorly; anterior pronotal lobe small, hemisphered and bulged, smooth, deeply depressed at base; posterior pronotal lobe rugulose and punctured, shallowly depressed on disc, with slightly swollen anteromedial elevation (never sulcate or concave); humerus bluntly triangular, with round apex; posterior margin of pronotum weakly concave, but slightly convex centrally; posterior angles round, slightly exceeded to posterior margin of pronotum (
<figureCitation id="A8E97FADE801FFBFFE3CEA07FD978C43" box="[510,598,1843,1869]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6F</figureCitation>
). Scutellum triangular, somewhat triangularly depressed basally, apically produced with a scutellar spine, and sloping downward, posterior apex round (
<figureCitation id="A8E97FADE801FFBFFBC3EA63FB978C7F" box="[1025,1110,1879,1905]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6F</figureCitation>
); scutellar spine sub-vertical, short or almost tubercle-liked (
<quantity id="F72ACECDE801FFBFFE38EA4FFD648C9B" box="[506,677,1915,1941]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="12.1" metricValueMin="4.4" pageId="14" pageNumber="315" unit="mm" value="0.825" valueMax="1.21" valueMin="0.44">0.441.21 mm</quantity>
), not bifid at apex (
<figureCitation id="A8E97FADE801FFBFFC48EA4FFC248C9B" box="[906,997,1915,1941]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6B</figureCitation>
). Femora elongated, slender but stout, moderate subnodulose apically; tibiae slender and elongated. Hemelytra surpassing beyond apex of abdomen when fully closed, 0.7 times as long as body length (
<figureCitation id="A8E97FADE801FFBFFD02EAF7FCDC8CD3" box="[704,797,1987,2013]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="14" pageNumber="315">Fig. 6A</figureCitation>
); discal cell nearly diamond-shaped, longer than width (
<figureCitation id="A8E97FADE800FFBEFF5DEDA3FF3A8BBF" box="[159,251,151,177]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="15" pageNumber="316">Fig. 6G</figureCitation>
); Sc 0.8 times as long as hemelytron length, 1.4 times as long as R + M (
<figureCitation id="A8E97FADE800FFBEFB80EDA3FB5C8BBF" box="[1090,1181,151,177]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="15" pageNumber="316">Fig. 6G</figureCitation>
). Hind wing about 3.0 times as long as maximum width (
<figureCitation id="A8E97FADE800FFBEFDE0ED8FFDBF8BDB" box="[546,638,187,213]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="15" pageNumber="316">Fig. 6H</figureCitation>
). Laterotergites much dilated and ascending with segmental incisures, posto-lateral margin of each laterotegite slightly produced than antero-lateral margin of following laterotegite (
<figureCitation id="A8E97FADE800FFBEFAADEDEBFF1F8A13" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="15" pageNumber="316">Fig. 6AC</figureCitation>
). Pygophore ovoid (
<figureCitation id="A8E97FADE800FFBEFE7FEC37FDF68A13" box="[445,567,259,285]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="15" pageNumber="316">Fig. 7BD</figureCitation>
); median process of pygophore (mpp) posteriorly produced, 0.52 times as long as wide in dorsal view, with apical margin convex, distinctly produced a round apex, apicolateral corner sub-round, distinctly produced posterolaterad (
<figureCitation id="A8E97FADE800FFBEFDEAEC7FFD638A6B" box="[552,674,331,357]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="15" pageNumber="316">Fig. 7BE</figureCitation>
); paramere long, slender, clavate, somewhat incurved and larger in apical part, with round apex (
<figureCitation id="A8E97FADE800FFBEFE27EC5BFD9C8A87" box="[485,605,367,393]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="15" pageNumber="316">Fig. 7F, G</figureCitation>
). Aedeagus in dorsal view ovoid, dorsally sclerotized, and in lateral view long and narrow (
<figureCitation id="A8E97FADE800FFBEFEA3ECA7FE7A8AA3" box="[353,443,403,429]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="15" pageNumber="316">Fig. 7H</figureCitation>
); articulatory apparatus (aa) in ventral view with basal plate arms relatively slender and jointly forming a V-shape, and in lateral view arched strongly (
<figureCitation id="A8E97FADE800FFBEFC88EC83FC768ADF" box="[842,951,439,465]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="15" pageNumber="316">Fig. 7H, I</figureCitation>
); dorsal phallothecal sclerite (dps) in lateral view with posteromedian weakly produced (
<figureCitation id="A8E97FADE800FFBEFD4BECEFFD198AFB" box="[649,728,475,501]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="15" pageNumber="316">Fig. 7I</figureCitation>
); spoon-like sclerites (sps) largely anteriorly produced, hyaline and glabrous (
<figureCitation id="A8E97FADE800FFBEFEF4ECCBFE648917" box="[310,421,511,537]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="15" pageNumber="316">Fig. 7H, I</figureCitation>
); membranous sac-like lobes almost disappeared (
<figureCitation id="A8E97FADE800FFBEFC1BECCBFBF38917" box="[985,1074,511,537]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="15" pageNumber="316">Fig. 7H</figureCitation>
); distal dorsal lobe of endosoma (ddl) round and weakly bulged, with two very large and long horn-shaped prickles centrally, surrounded by large prickles (
<figureCitation id="A8E97FADE800FFBEFF3DEF73FEBC896F" box="[255,381,583,609]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="15" pageNumber="316">Fig. 7HK</figureCitation>
).
</paragraph>
<paragraph id="306D6328E800FFBEFF05EF5EFBB18F3B" blockId="15.[151,1437,151,1077]" pageId="15" pageNumber="316">
<emphasis id="02A6BF3AE800FFBEFF05EF5EFEFB898A" bold="true" box="[199,314,618,644]" pageId="15" pageNumber="316">Vestiture.</emphasis>
Body clothed with pale, shining griseous, slender setae. Head covered with very short bent slender setae, interleaved with short erect slender setae; labium almost glabrous except base of first visible segment with a few tiny slender setae (
<figureCitation id="A8E97FADE800FFBEFE51EF87FDCB89C3" box="[403,522,691,717]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="15" pageNumber="316">Fig. 6D, E</figureCitation>
). Scape and pedicel covered with short slender sub-erect setae; remaining antennae covered with short vertical setae, denser toward tip. Collar, anterolateral angles with tiny bent setae abundantly; anterior pronotal lobe with some rows of tiny bent setae, somewhat interleave with slender erect setae; posterior pronotal lobe covered with short slender setae and posteriorly cover with slender setae; scutellum densely covered with long, slender, vertical, erect black setae (
<figureCitation id="A8E97FADE800FFBEFD52EE77FD248853" box="[656,741,835,861]" captionStart="FIGURE 6" captionStartId="13.[151,250,1860,1884]" captionTargetBox="[151,1435,191,1836]" captionTargetId="figure-18@13.[151,1435,181,1836]" captionTargetPageId="13" captionText="FIGURE 6. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., holotype, ♂, TXL2024-012. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra; F, right hind wing." figureDoi="http://doi.org/10.5281/zenodo.12758821" httpUri="https://zenodo.org/record/12758821/files/figure.png" pageId="15" pageNumber="316">Fig. 6F</figureCitation>
). Coxae and trochanters covered with short bent setae; femora, tibiae, and tarsi densely covered with long, slender, erect setae. Coria and clavus densely covered with short bent setae. Abdominal mediotergites sparsely covered with slender erect setae; lateral margins of laterotergites densely covered with short erect setae; abdominal sternites and laterotergites covered with short oblique cream-yellow setae, somewhat interleaved with long slender erect setae. Male genitalia pygophore ventrally covered with short slender bent setae and posteriorly densely covered with short and long slender setae (
<figureCitation id="A8E97FADE800FFBEFC33EEC3FBA88F1F" box="[1009,1129,1015,1041]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="15" pageNumber="316">Fig. 7BE</figureCitation>
); paramere almost glabrous except posterior apex densely covered with short and long thick erect setae (
<figureCitation id="A8E97FADE800FFBEFC2CE92FFBA28F3B" box="[1006,1123,1051,1077]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="15" pageNumber="316">Fig. 7F, G</figureCitation>
).
</paragraph>
<paragraph id="306D6328E800FFBEFF55E957FE438F72" blockId="15.[151,784,1122,1185]" box="[151,386,1122,1149]" pageId="15" pageNumber="316">
<emphasis id="02A6BF3AE800FFBEFF55E957FE438F72" bold="true" box="[151,386,1122,1149]" pageId="15" pageNumber="316">Female description.</emphasis>
</paragraph>
<paragraph id="306D6328E800FFBEFF55E9B3FCD18FAF" blockId="15.[151,784,1122,1185]" box="[151,784,1159,1185]" pageId="15" pageNumber="316">General external morphology similar to that of the male.</paragraph>
<paragraph id="306D6328E800FFBEFF55E9FAFE598E5B" blockId="15.[151,1437,1230,1905]" pageId="15" pageNumber="316">
<emphasis id="02A6BF3AE800FFBEFF55E9FAFEDE8FE6" bold="true" box="[151,287,1230,1256]" pageId="15" pageNumber="316">Coloration.</emphasis>
Almost similar to male but differ in the following characters. Abdominal sternite VII yellow with dark brown suffusion near posterior margin, and blackish brown or back posterior margin; Gonocoxa VIII yellow with inner margin black; abdominal tergite IX and gonapophysis VIII black; abdominal laterotergite VIII orange or sanguineous (
<figureCitation id="A8E97FADE800FFBEFEF3E80FFE4A8E5B" box="[305,395,1339,1365]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="15" pageNumber="316">Fig. 7A</figureCitation>
).
</paragraph>
<paragraph id="306D6328E800FFBEFF05E86AFEAE8DEF" blockId="15.[151,1437,1230,1905]" pageId="15" pageNumber="316">
<emphasis id="02A6BF3AE800FFBEFF05E86AFE808E76" bold="true" box="[199,321,1374,1400]" pageId="15" pageNumber="316">Structure.</emphasis>
Almost the same as male but larger than male and differ in the following characters. Body large-sized (
<quantity id="F72ACECDE800FFBEFF26E8B7FE6F8E93" box="[228,430,1411,1437]" metricMagnitude="-2" metricUnit="m" metricValue="2.3525" metricValueMax="2.487" metricValueMin="2.218" pageId="15" pageNumber="316" unit="mm" value="23.525" valueMax="24.87" valueMin="22.18">22.1824.87 mm</quantity>
), elongated, and somewhat robust. Head shorter than or almost as long as pronotum. Proportional average length of first to third visible labial segments 2.6:4.2:0.7. Scape about 1.9 times as long as head, about 2.4 times as long as pedicel, slightly longer than second flagellomere; first flagellomere slightly shorter than pedicel and about 1/3 times as long as second flagellomere; proportional average length of scape, pedicel, first and second flagellomeres 7.3:3.0:2.5:6.7. Abdominal laterotergite VIII with thin posterior margin, not exceed the posterior margin of abdominal tergite IX; abdominal sternite VII forming a semi-circular or wide sub-pentagonal median concavity, with posteromedian margin gently U-shaped, with inner posterolateral margin slightly concave; gonocoxa VIII subtriangular, produced mesad and forming a pointed apex at apical inner corner, and with inner margin weakly incurved in posterior 2/3; gonapophysis VIII subtriangular, posteriorly produced; abdominal tergite IX small (
<figureCitation id="A8E97FADE800FFBEFECAEBF3FEA38DEF" box="[264,354,1735,1761]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="15" pageNumber="316">Fig. 7A</figureCitation>
).
</paragraph>
<paragraph id="306D6328E800FFBEFF05EBDEFDA98C7F" blockId="15.[151,1437,1230,1905]" pageId="15" pageNumber="316">
<emphasis id="02A6BF3AE800FFBEFF05EBDEFEFB8C0A" bold="true" box="[199,314,1770,1796]" pageId="15" pageNumber="316">Vestiture.</emphasis>
Almost the same as male except for the following characters.Anterior 1/3 of gonocoxa VIII almost no setae; posterior 2/3 of gonocoxa VIII covered with short and long bent black setae; posterior margin of gonapophysis VIII densely covered with long bent black setae; abdominal laterotergite VIII and abdominal tergite IX covered with short and long bent black setae (
<figureCitation id="A8E97FADE800FFBEFDC3EA63FD9A8C7F" box="[513,603,1879,1905]" captionStart="FIGURE 7" captionStartId="16.[151,250,1874,1898]" captionTargetBox="[317,1270,181,1850]" captionTargetId="figure-24@16.[317,1270,181,1850]" captionTargetPageId="16" captionText="FIGURE 7. Sycanus thuathienhuensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia, holotype, ♂, TXL2024-012. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="15" pageNumber="316">Fig. 7A</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="64AD33A0E81FFFA1FF55EA66FB878CD8" ID-DOI="http://doi.org/10.5281/zenodo.12758823" ID-Zenodo-Dep="12758823" httpUri="https://zenodo.org/record/12758823/files/figure.png" pageId="16" pageNumber="317" startId="16.[151,250,1874,1898]" targetBox="[317,1270,181,1850]" targetPageId="16" targetType="figure">
<paragraph id="306D6328E81FFFA1FF55EA66FB878CD8" blockId="16.[151,1436,1874,2006]" pageId="16" pageNumber="317">
<emphasis id="02A6BF3AE81FFFA1FF55EA66FED28C64" bold="true" box="[151,275,1874,1898]" pageId="16" pageNumber="317">FIGURE 7.</emphasis>
<taxonomicName id="F7D218ABE81FFFA1FEDBEA66FD6B8C64" authority="Truong &amp; Ha" authorityName="Truong &amp; Giang &amp; Nguyen &amp; Chau &amp; Ha" authorityYear="2024" box="[281,682,1874,1899]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="thuathienhuensis" status="sp. nov.">
<emphasis id="02A6BF3AE81FFFA1FEDBEA66FDDC8C64" box="[281,541,1874,1898]" italics="true" pageId="16" pageNumber="317">Sycanus thuathienhuensis</emphasis>
Truong &amp; Ha
</taxonomicName>
,
<emphasis id="02A6BF3AE81FFFA1FD74EA66FCC78C64" bold="true" box="[694,774,1874,1898]" pageId="16" pageNumber="317">
<taxonomicNameLabel id="19950241E81FFFA1FD74EA66FCC78C64" box="[694,774,1874,1898]" pageId="16" pageNumber="317" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A, female genitalia in ventral view, paratype, ♀, NDD2022-063; BK, male genitalia,
<emphasis id="02A6BF3AE81FFFA1FEB1EA42FD4C8C80" box="[371,653,1910,1936]" italics="true" pageId="16" pageNumber="317">holotype, ♂, TXL2024-012.</emphasis>
BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl).
</paragraph>
</caption>
<subSubSection id="78C830A3E81EFFA0FF05EDA3FB8D896F" pageId="17" pageNumber="318" type="description">
<paragraph id="306D6328E81EFFA0FF05EDA3FB8D896F" blockId="17.[151,1437,150,789]" pageId="17" pageNumber="318">
<emphasis id="02A6BF3AE81EFFA0FF05EDA3FEBF8BBF" bold="true" box="[199,382,151,177]" pageId="17" pageNumber="318">Measurements.</emphasis>
All dimensions are given in mm.
<emphasis id="02A6BF3AE81EFFA0FCCBEDA3FC718BBA" bold="true" box="[777,944,151,180]" pageId="17" pageNumber="318">
<typeStatus id="EF69DD8AE81EFFA0FCCBEDA3FCB78BBF" box="[777,886,151,177]" pageId="17" pageNumber="318" type="holotype">Holotype</typeStatus>
(
<specimenCount id="26D4A8A1E81EFFA0FC4BEDACFC5E8BBA" box="[905,927,152,180]" count="1" pageId="17" pageNumber="318" type="male"></specimenCount>
):
</emphasis>
BL 18.89; HL 3.75; AoL 1.73; AoW 0.87; PoL 1.12; PoW 1.27; NL 1.17; OE 1.78; IE 0.70; ED 1.09; OD 0.22; OCD 0.36; COD 0.20; R1L 2.50; R2L 3.88; R3L 0.61; A1L 7.79; A2L 2.88; A3L 2.30; A4L 7.06; PnL 3.62; PnW 4.26; APL 1.31; PPL 2.30; SPL 0.46; HeL 13.84; HeW 4.58; Sc 10.23; R+M 7.38; HWL 9.63; HWW 3.17; AFL 7.09; ATL 8.29; MFL 6.34; MTL 7.89; PFL 8.41; PTL 11.08.
<emphasis id="02A6BF3AE81EFFA0FEA1EC13FDCA8A4A" bold="true" box="[355,523,295,324]" pageId="17" pageNumber="318">
<typeStatus id="EF69DD8AE81EFFA0FEA1EC13FE118A4F" box="[355,464,295,321]" pageId="17" pageNumber="318" type="paratype">Paratype</typeStatus>
(
<specimenCount id="26D4A8A1E81EFFA0FE26EC1CFE3B8A4A" box="[484,506,296,324]" count="1" pageId="17" pageNumber="318" type="male"></specimenCount>
):
</emphasis>
BL 19.08; HL 3.61; AoL 1.62; AoW 0.88; PoL 1.10; PoW 1.23; NL 1.14; OE 1.69; IE 0.69; ED 1.03; OD 0.19; OCD 0.46; COD 0.20; R1L 2.32; R2L 3.73; R3L 0.60; A1L 6.82; A2L 3.08; A3L 2.15; A4L 6.60; PnL 3.52; PnW 4.22; APL 1.20; PPL 2.31; SPL 1.21; HeL 13.49; HeW 4.04; Sc 9.95; R+M 6.00; AFL 7.72; ATL 8.19; MFL 6.25; MTL 7.63; PFL 8.10; PTL 10.44.
<emphasis id="02A6BF3AE81EFFA0FC45ECA7FBEC8ABE" bold="true" box="[903,1069,403,432]" pageId="17" pageNumber="318">
<typeStatus id="EF69DD8AE81EFFA0FC45ECA7FC358AA3" box="[903,1012,403,429]" pageId="17" pageNumber="318" type="paratype">Paratype</typeStatus>
(
<specimenCount id="26D4A8A1E81EFFA0FBC4ECA0FBDD8ABE" box="[1030,1052,404,432]" count="1" pageId="17" pageNumber="318" type="female"></specimenCount>
):
</emphasis>
BL 22.1824.87; HL 3.814.23; AoL 1.852.04; AoW 0.941.04; PoL 0.951.23; PoW 1.251.46; NL 1.351.42; OE 1.731.87; IE 0.760.84; ED 1.041.10; OD 0.210.28; OCD 0.450.58; COD 0.180.21; R1L 2.502.72; R2L 4.034.36; R3L 0.630.81; A1L 6.837.82; A2L 3.043.08; A3L 2.262.66; A4L 6.477.03; PnL 3.714.74; PnW 5.145.63; APL 1.031.45; PPL 2.693.27; SPL 0.961.53; HeL 16.0518.06; HeW 4.675.13; Sc 11.6912.98; R+M 7.938.83; AFL 7.338.08; ATL 8.589.00; MFL 6.196.71; MTL 7.898.40; PFL 8.119.22; PTL 11.3712.05.
</paragraph>
</subSubSection>
<subSubSection id="78C830A3E81EFFA0FF05EF5EFE8889A7" pageId="17" pageNumber="318" type="distribution">
<paragraph id="306D6328E81EFFA0FF05EF5EFE8889A7" blockId="17.[151,1437,150,789]" pageId="17" pageNumber="318">
<emphasis id="02A6BF3AE81EFFA0FF05EF5EFEA0898A" bold="true" box="[199,353,618,644]" pageId="17" pageNumber="318">Distribution.</emphasis>
<collectingCountry id="48C523B8E81EFFA0FEA5EF5FFE08898B" box="[359,457,619,645]" name="Vietnam" pageId="17" pageNumber="318">Vietnam</collectingCountry>
, Middle and Southern
<collectingCountry id="48C523B8E81EFFA0FD06EF5FFCE2898B" box="[708,803,619,645]" name="Vietnam" pageId="17" pageNumber="318">Vietnam</collectingCountry>
(Thua Thien Hue,
<collectingRegion id="F216ADCAE81EFFA0FC37EF5FFBBE898B" box="[1013,1151,619,645]" country="Vietnam" name="Quang Nam" pageId="17" pageNumber="318">Quang Nam</collectingRegion>
,
<collectingRegion id="F216ADCAE81EFFA0FB48EF5FFAC3898B" box="[1162,1282,619,645]" country="Vietnam" name="Binh Dinh" pageId="17" pageNumber="318">Binh Dinh</collectingRegion>
,
<collectingRegion id="F216ADCAE81EFFA0FACEEF5FFAA2898B" box="[1292,1379,619,645]" country="Vietnam" name="Gia Lai" pageId="17" pageNumber="318">Gia Lai</collectingRegion>
,
<collectingRegion id="F216ADCAE81EFFA0FAAFEF5FFF0289A7" country="Vietnam" name="Dak Lak" pageId="17" pageNumber="318">Dak Lak</collectingRegion>
Provinces).
</paragraph>
</subSubSection>
<subSubSection id="78C830A3E81EFFA0FF05EF87FC6C89C3" box="[199,941,690,717]" pageId="17" pageNumber="318" type="materials_examined">
<paragraph id="306D6328E81EFFA0FF05EF87FC6C89C3" blockId="17.[151,1437,150,789]" box="[199,941,690,717]" pageId="17" pageNumber="318">
<materialsCitation id="80BA6975E81EFFA0FF05EF87FC6889C3" box="[199,937,690,717]" country="Vietnam" location="Thua Thien Hue" municipality="Bach Ma National Park" pageId="17" pageNumber="318" specimenCount="1" typeStatus="holotype">
<emphasis id="02A6BF3AE81EFFA0FF05EF87FEA789C2" bold="true" box="[199,358,690,717]" pageId="17" pageNumber="318">
<typeStatus id="EF69DD8AE81EFFA0FF05EF87FEC389C3" box="[199,258,691,717]" pageId="17" pageNumber="318">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="48C523B8E81EFFA0FEAEEF87FE0F89C3" box="[364,462,691,717]" name="Vietnam" pageId="17" pageNumber="318">Vietnam</collectingCountry>
,
<location id="350D35F3E81EFFA0FE1BEF87FD5389C3" LSID="urn:lsid:plazi:treatment:B87BD23EE803FFA0FF55EF7FFDCB881B:350D35F3E81EFFA0FE1BEF87FD5389C3" box="[473,658,691,717]" country="Vietnam" municipality="Bach Ma National Park" name="Thua Thien Hue" pageId="17" pageNumber="318">Thua Thien Hue</location>
,
<collectingMunicipality id="D009F952E81EFFA0FD5CEF87FC6889C3" box="[670,937,691,717]" pageId="17" pageNumber="318">Bach Ma National Park</collectingMunicipality>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="78C830A3E81EFFA0FF05EFE3FDCB881B" pageId="17" pageNumber="318" type="etymology">
<paragraph id="306D6328E81EFFA0FF05EFE3FDCB881B" blockId="17.[151,1437,150,789]" pageId="17" pageNumber="318">
<emphasis id="02A6BF3AE81EFFA0FF05EFE3FE8F89FF" bold="true" box="[199,334,727,753]" pageId="17" pageNumber="318">Etymology.</emphasis>
The specific epithet refers to its occurrence in Thua Thien Hue, the North Central Coast region, the Central of
<collectingCountry id="48C523B8E81EFFA0FECCEFCFFEB1881B" box="[270,368,763,789]" name="Vietnam" pageId="17" pageNumber="318">Vietnam</collectingCountry>
; an adjective.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,492 @@
<document id="124C40535C955479EB239DD32668E542" ID-DOI="10.11646/zootaxa.5481.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="12758808" ID-ZooBank="0803FD80-03E1-4751-BC89-0BC1D44CA26C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721215784887" checkinUser="plazi" docAuthor="Truong, Xuan Lam, Giang, Phan Thi, Nguyen, Dai Dac, Chau, Tran Pham Minh &amp; Ha, Ngoc Linh" docDate="2024" docId="B87BD23EE81EFFA4FF55EE6BFE528DB3" docLanguage="en" docName="zootaxa.5481.3.1.pdf" docOrigin="Zootaxa 5481 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sycanus taynguyenensis Truong &amp; Giang &amp; Nguyen &amp; Chau &amp; Ha 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="322" masterDocId="4442AA46E80FFFB1FFC2ED34FFC18B0E" masterDocTitle="Two new species of the genus Sycanus Amyot &amp; Serville (Insecta: Hemiptera: Reduviidae: Harpactorinae) from Vietnam" masterLastPageNumber="325" masterPageNumber="301" pageNumber="318" updateTime="1721216262946" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="BE85D39491DFCCE3BF7A232E2A7C4B84" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EEB6FCA2D2A12F5E496A69AE4A90BAD4">
<mods:title id="7A17747D475BB5F4B00772B316FF8687">Two new species of the genus Sycanus Amyot &amp; Serville (Insecta: Hemiptera: Reduviidae: Harpactorinae) from Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="A928321DA866FEEC31F77D11D76DD798" type="personal">
<mods:role id="D6CDBABFCE2E6C594ADFD8C7226E4491">
<mods:roleTerm id="1A6A8893902D4044B095D97E14F88974">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0D2F99D003BEE60BDC90C17DF9C3F0F7">Truong, Xuan Lam</mods:namePart>
<mods:nameIdentifier id="0325A3A97C0F84F2401F3C1445181F8A" type="ORCID">0000-0002-1758-903X</mods:nameIdentifier>
<mods:affiliation id="DEB5B59FC4C8CAF7B45C23B225ACB801">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Hanoi, Vietnam, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="FBAE531BF149DA7430F9DD2A8F92C0E9" type="email">txlam.iebr@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="F1ACC5596DB91EF61D9D84361EA03048">Giang, Phan Thi</mods:namePart>
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<mods:affiliation id="86541D068BBA3CC369CB57BEED8E01E4">Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam &amp; Vinh University, 182 Le Duan, Vinh, Nghe An, Vietnam</mods:affiliation>
<mods:nameIdentifier id="2E73B80B1669CADD87F44483AE30B4B1" type="email">tragiangdhv8668@gmail.com</mods:nameIdentifier>
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<mods:namePart id="61969D485B2BAC20F4263E99B62D43BE">Nguyen, Dai Dac</mods:namePart>
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<mods:affiliation id="026D09FDAF9FD340D47B22A08FCC0F4E">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Hanoi, Vietnam, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="BAA0D6638D4912276C9B53C6785FD6CF" type="email">daindiebr@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D98C0BF7816FCCADFDD18727B255C9F1">Chau, Tran Pham Minh</mods:namePart>
<mods:nameIdentifier id="5D8C68308AE568B5D23632211A4B051E" type="ORCID">0009-0009-6264-0592</mods:nameIdentifier>
<mods:affiliation id="4D08836FC03FECDDF0495181FAC7EE58">Faculty of Biology, Vietnam National University - VNU University of Science, 334 Nguyen Trai, Thanh Xuan, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="4F6D0F70AFEE5928472A87E815D30CFF" type="email">tranphamminhchau_t66@hus.edu.vn</mods:nameIdentifier>
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<mods:namePart id="2E3314478CC2BF7F9FA49998F9294138">Ha, Ngoc Linh</mods:namePart>
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<mods:affiliation id="36D75FDCE0EE6F90EC47ACF2F64581F9">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Hanoi, Vietnam, 18 Hoang Quoc Viet Road, Nghia Do, Cau Giay, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="6F60CD34BCE81B3376D99BD3BAA637E2" type="email">linh.hangoc02@gmail.com</mods:nameIdentifier>
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<mods:date id="EAB2024CE9F21C508D42E1DBFC8E77D2">2024</mods:date>
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<heading id="6B25D444E81EFFA0FF55EE6BFD1B8877" bold="true" box="[151,730,862,889]" fontSize="11" level="1" pageId="17" pageNumber="318" reason="1">
<emphasis id="02A6BF3AE81EFFA0FF55EE6BFD1B8877" bold="true" box="[151,730,862,889]" pageId="17" pageNumber="318">
<taxonomicName id="F7D218ABE81EFFA0FF55EE6BFDAE8877" authority="Truong &amp; Ha" authorityName="Truong &amp; Giang &amp; Nguyen &amp; Chau &amp; Ha" authorityYear="2024" box="[151,623,862,889]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="taynguyenensis" status="sp. nov.">
<emphasis id="02A6BF3AE81EFFA0FF55EE6BFE7F8877" bold="true" box="[151,446,863,889]" italics="true" pageId="17" pageNumber="318">Sycanus taynguyenensis</emphasis>
Truong &amp; Ha
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,
<taxonomicNameLabel id="19950241E81EFFA0FDBEEE6BFD1B8877" box="[636,730,863,889]" pageId="17" pageNumber="318" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="78C830A3E81EFFA0FF55EEB7FE808893" box="[151,321,899,925]" pageId="17" pageNumber="318" type="description">
<paragraph id="306D6328E81EFFA0FF55EEB7FE808893" blockId="17.[151,730,862,925]" box="[151,321,899,925]" pageId="17" pageNumber="318">
(
<figureCitation id="A8E97FADE81EFFA0FF5DEEB7FEC08893" box="[159,257,899,925]" captionStart="FIGURE 4" captionStartId="9.[151,250,1203,1227]" captionTargetBox="[163,1422,193,1166]" captionTargetId="figure-18@9.[151,1436,181,1180]" captionTargetPageId="9" captionText="FIGURE 4. Body in dorsal view of two new morphospecies, α and β. A, Sycanus thuathienhuensis Truong &amp; Ha, sp. nov. (= α), paratype, ♂, TXL2024-009; B, S. taynguyenensis Truong &amp; Ha, sp. nov. (= β), holotype, ♂, NDD2019-285." figureDoi="http://doi.org/10.5281/zenodo.12758817" httpUri="https://zenodo.org/record/12758817/files/figure.png" pageId="17" pageNumber="318">Figs. 4B</figureCitation>
,
<figureCitation id="A8E97FADE81EFFA0FECFEEB7FEDD8893" box="[269,284,899,925]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="17" pageNumber="318">8</figureCitation>
,
<figureCitation id="A8E97FADE81EFFA0FEEBEEB7FEF98893" box="[297,312,899,925]" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="17" pageNumber="318">9</figureCitation>
)
</paragraph>
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<subSubSection id="78C830A3E81EFFA0FF55EEFFFCC88F97" pageId="17" pageNumber="318" type="materials_examined">
<paragraph id="306D6328E81EFFA0FF55EEFFFDED8F23" blockId="17.[151,1437,970,2041]" pageId="17" pageNumber="318">
<emphasis id="02A6BF3AE81EFFA0FF55EEFFFE8588EA" bold="true" box="[151,324,970,997]" pageId="17" pageNumber="318">Type material.</emphasis>
<materialsCitation id="80BA6975E81EFFA0FE89EEFFFEB58F07" collectingDate="2019-09-19" collectionCode="IEBR" collectorName="DD Nguyen" country="Vietnam" location="Ea So Natural Reserve" pageId="17" pageNumber="318" specimenCount="1" specimenCount-male="1" stateProvince="Dak Lak" typeStatus="holotype">
<emphasis id="02A6BF3AE81EFFA0FE89EEFFFE7E88EB" bold="true" box="[331,447,971,997]" pageId="17" pageNumber="318">
<typeStatus id="EF69DD8AE81EFFA0FE89EEFFFE7B88EB" box="[331,442,971,997]" pageId="17" pageNumber="318" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="26D4A8A1E81EFFA0FE07EEFFFE1C88EB" box="[453,477,971,997]" count="1" pageId="17" pageNumber="318" type="male"></specimenCount>
; NDD2019-285;
<collectingCountry id="48C523B8E81EFFA0FD5CEEFFFCC188EB" box="[670,768,971,997]" name="Vietnam" pageId="17" pageNumber="318">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE81EFFA0FCC8EEFFFC1988EB" box="[778,984,971,997]" country="Vietnam" name="Dak Lak" pageId="17" pageNumber="318">Dak Lak Province</collectingRegion>
,
<location id="350D35F3E81EFFA0FC21EEFFFB2288EB" LSID="urn:lsid:plazi:treatment:B87BD23EE81EFFA4FF55EE6BFE528DB3:350D35F3E81EFFA0FC21EEFFFB2288EB" box="[995,1251,971,997]" country="Vietnam" name="Ea So Natural Reserve" pageId="17" pageNumber="318" stateProvince="Dak Lak">Ea So Natural Reserve</location>
;
<date id="446C45E8E81EFFA0FB2CEEFFFAA888EB" box="[1262,1385,971,997]" pageId="17" pageNumber="318" value="2019-09-19">
<collectingDate id="5428BC00E81EFFA0FB2CEEFFFAA888EB" box="[1262,1385,971,997]" pageId="17" pageNumber="318" value="2019-09-19">19.ix.2019</collectingDate>
</date>
;
<collectorName id="9D2706FEE81EFFA0FAB6EEFFFF318F07" pageId="17" pageNumber="318">DD Nguyen</collectorName>
leg.;
<collectionCode id="56C3FBEDE81EFFA0FEF3EEDBFEB58F07" box="[305,372,1007,1033]" pageId="17" pageNumber="318">IEBR</collectionCode>
</materialsCitation>
.
<materialsCitation id="80BA6975E81EFFA0FEBDEEDBFDE98F23" collectingDate="2018-09-26" collectionCode="IEBR" collectorName="DD Nguyen" country="Vietnam" location="Kon Chu Rang Natural Reserve" pageId="17" pageNumber="318" specimenCount="1" specimenCount-female="1" stateProvince="Gia Lai" typeStatus="paratype">
<emphasis id="02A6BF3AE81EFFA0FEBDEEDBFE3F8F07" bold="true" box="[383,510,1007,1033]" pageId="17" pageNumber="318">
<typeStatus id="EF69DD8AE81EFFA0FEBDEEDBFE388F07" box="[383,505,1007,1033]" pageId="17" pageNumber="318" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="26D4A8A1E81EFFA0FDC4EEDBFDEB8F07" box="[518,554,1007,1033]" count="1" pageId="17" pageNumber="318" type="female">1♀</specimenCount>
; NDD2018-395;
<collectingCountry id="48C523B8E81EFFA0FD2DEEDBFC908F07" box="[751,849,1007,1033]" name="Vietnam" pageId="17" pageNumber="318">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE81EFFA0FC9FEEDBFBE08F07" box="[861,1057,1007,1033]" country="Vietnam" name="Gia Lai" pageId="17" pageNumber="318">Gia Lai Province</collectingRegion>
,
<location id="350D35F3E81EFFA0FBEFEEDBFA598F07" LSID="urn:lsid:plazi:treatment:B87BD23EE81EFFA4FF55EE6BFE528DB3:350D35F3E81EFFA0FBEFEEDBFA598F07" box="[1069,1432,1007,1033]" country="Vietnam" name="Kon Chu Rang Natural Reserve" pageId="17" pageNumber="318" stateProvince="Gia Lai">Kon Chu Rang Natural Reserve</location>
;
<date id="446C45E8E81EFFA0FF55E927FED38F23" box="[151,274,1043,1069]" pageId="17" pageNumber="318" value="2018-09-26">
<collectingDate id="5428BC00E81EFFA0FF55E927FED38F23" box="[151,274,1043,1069]" pageId="17" pageNumber="318" value="2018-09-26">26.ix.2018</collectingDate>
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;
<collectorName id="9D2706FEE81EFFA0FEDCE927FE678F23" box="[286,422,1043,1069]" pageId="17" pageNumber="318">DD Nguyen</collectorName>
leg.;
<collectionCode id="56C3FBEDE81EFFA0FE27E927FDE98F23" box="[485,552,1043,1069]" pageId="17" pageNumber="318">IEBR</collectionCode>
</materialsCitation>
.
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<emphasis id="02A6BF3AE81EFFA0FF05E903FE6F8F5E" bold="true" box="[199,430,1078,1105]" pageId="17" pageNumber="318">Non-type material.</emphasis>
<materialsCitation id="80BA6975E81EFFA0FE79E903FC8F8F7B" collectingDate="2019-03-30" collectionCode="IEBR" collectorName="Nguyen Thi Quynh Hoa" country="Vietnam" location="Dray Sap" municipality="Krong Ana" pageId="17" pageNumber="318" specimenCount="1" specimenCount-male="1" stateProvince="Dak Lak">
<specimenCount id="26D4A8A1E81EFFA0FE79E903FE218F5F" box="[443,480,1079,1105]" count="1" pageId="17" pageNumber="318" type="male">1♂</specimenCount>
; Qhoa-M10-01;
<collectingCountry id="48C523B8E81EFFA0FD64E903FCC98F5F" box="[678,776,1079,1105]" name="Vietnam" pageId="17" pageNumber="318">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE81EFFA0FCDBE903FC338F5F" box="[793,1010,1079,1105]" country="Vietnam" name="Dak Lak" pageId="17" pageNumber="318">Dak Lak Province</collectingRegion>
,
<collectingMunicipality id="D009F952E81EFFA0FBC1E903FB468F5F" box="[1027,1159,1079,1105]" pageId="17" pageNumber="318">Krong Ana</collectingMunicipality>
,
<location id="350D35F3E81EFFA0FB5AE903FACB8F5F" LSID="urn:lsid:plazi:treatment:B87BD23EE81EFFA4FF55EE6BFE528DB3:350D35F3E81EFFA0FB5AE903FACB8F5F" box="[1176,1290,1079,1105]" country="Vietnam" municipality="Krong Ana" name="Dray Sap" pageId="17" pageNumber="318" stateProvince="Dak Lak">Dray Sap</location>
;
<date id="446C45E8E81EFFA0FAD9E903FA598F5F" box="[1307,1432,1079,1105]" pageId="17" pageNumber="318" value="2019-03-30">
<collectingDate id="5428BC00E81EFFA0FAD9E903FA598F5F" box="[1307,1432,1079,1105]" pageId="17" pageNumber="318" value="2019-03-30">30.iii.2019</collectingDate>
</date>
;
<collectorName id="9D2706FEE81EFFA0FF55E96FFE688F7B" box="[151,425,1115,1141]" pageId="17" pageNumber="318">Nguyen Thi Quynh Hoa</collectorName>
(Tay Nguyen University) leg.;
<collectionCode id="56C3FBEDE81EFFA0FCC9E96FFC8F8F7B" box="[779,846,1115,1141]" pageId="17" pageNumber="318">IEBR</collectionCode>
</materialsCitation>
.
<materialsCitation id="80BA6975E81EFFA0FC98E96FFCC58F97" collectingDate="2023-09-15" collectionCode="IEBR" collectorName="XL Truong" country="Vietnam" location="Ea So Natural Reserve" pageId="17" pageNumber="318" specimenCount="1" specimenCount-male="1" stateProvince="Dak Lak">
<specimenCount id="26D4A8A1E81EFFA0FC98E96FFCBE8F7B" box="[858,895,1115,1141]" count="1" pageId="17" pageNumber="318" type="male">1♂</specimenCount>
; TXL-M10-02;
<collectingCountry id="48C523B8E81EFFA0FBF7E96FFB568F7B" box="[1077,1175,1115,1141]" name="Vietnam" pageId="17" pageNumber="318">Vietnam</collectingCountry>
,
<collectingRegion id="F216ADCAE81EFFA0FB61E96FFAB38F7B" box="[1187,1394,1115,1141]" country="Vietnam" name="Dak Lak" pageId="17" pageNumber="318">Dak Lak Province</collectingRegion>
,
<location id="350D35F3E81EFFA0FABDE96FFEB48F97" LSID="urn:lsid:plazi:treatment:B87BD23EE81EFFA4FF55EE6BFE528DB3:350D35F3E81EFFA0FABDE96FFEB48F97" country="Vietnam" name="Ea So Natural Reserve" pageId="17" pageNumber="318" stateProvince="Dak Lak">Ea So Natural Reserve</location>
;
<date id="446C45E8E81EFFA0FE42E94AFE3B8F97" box="[384,506,1150,1177]" pageId="17" pageNumber="318" value="2023-09-15">
<collectingDate id="5428BC00E81EFFA0FE42E94AFE3B8F97" box="[384,506,1150,1177]" pageId="17" pageNumber="318" value="2023-09-15">15.ix.2023</collectingDate>
</date>
;
<collectorName id="9D2706FEE81EFFA0FDC4E94BFD438F97" box="[518,642,1151,1177]" pageId="17" pageNumber="318">XL Truong</collectorName>
leg.;
<collectionCode id="56C3FBEDE81EFFA0FD03E94BFCC58F97" box="[705,772,1151,1177]" pageId="17" pageNumber="318">IEBR</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="78C830A3E81EFFA4FF05E996FD2C8E03" lastPageId="21" lastPageNumber="322" pageId="17" pageNumber="318" type="diagnosis">
<paragraph id="306D6328E81EFFA0FF05E996FB718E9B" blockId="17.[151,1437,970,2041]" pageId="17" pageNumber="318">
<emphasis id="02A6BF3AE81EFFA0FF05E996FE838FB2" bold="true" box="[199,322,1186,1212]" pageId="17" pageNumber="318">Diagnosis.</emphasis>
Body large-sized and pale orange brown; head long, almost black with a brownish yellow suffusion next to outer side of ocellus (
<figureCitation id="A8E97FADE81EFFA0FE28E9F3FDA68FEF" box="[490,615,1223,1249]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="17" pageNumber="318">Fig. 8D, E</figureCitation>
); base of scape blackish brown, scape interleaved brownish yellow and blackish brown, pedicel dark brown with a brown suffusion medially, first and second flagellomeres brown; anterior pronotal lobe orange brown, somewhat with irregular dark brown suffusion and posterior pronotal lobe rugulose, pale orange brown or reddish brown (
<figureCitation id="A8E97FADE81EFFA0FDFAE807FD4C8E43" box="[568,653,1331,1357]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="17" pageNumber="318">Fig. 8F</figureCitation>
); scutellum black with orange brown centrally and scutellar spine sub-vertical, short, bifid at apex, and wholly orange brown or reddish brown (
<figureCitation id="A8E97FADE81EFFA0FC14E863FB868E7F" box="[982,1095,1367,1393]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="17" pageNumber="318">Fig 8B, F</figureCitation>
); basal 2/3 of coria and whole clavus black or dark brown; remaining of coria white or orange; clavus dark brown (
<figureCitation id="A8E97FADE81EFFA0FB8BE84FFB628E9B" box="[1097,1187,1403,1429]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="17" pageNumber="318">Fig. 8G</figureCitation>
).
</paragraph>
<paragraph id="306D6328E81EFFA0FF05E8ABFD6C8DD7" blockId="17.[151,1437,970,2041]" pageId="17" pageNumber="318">
The general appearance and male genitalia of this species are highly similar to that of the
<taxonomicName id="F7D218ABE81EFFA0FB66E8ABFF3E8ED3" authority=", However" authorityName="However" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="sichuanensis">
<emphasis id="02A6BF3AE81EFFA0FB66E8ABFA5D8EB7" box="[1188,1436,1439,1465]" italics="true" pageId="17" pageNumber="318">Sycanus sichuanensis,</emphasis>
However
</taxonomicName>
, this new species can be distinguished from the latter by the following features: pronotum with anterior pronotal lobe orange brown, somewhat with irregular dark brown suffusion and posterior pronotal lobe pale orange brown or reddish brown (
<figureCitation id="A8E97FADE81EFFA0FE7BEB3FFDCF8D2B" box="[441,526,1547,1573]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="17" pageNumber="318">Fig. 8F</figureCitation>
) (in
<taxonomicName id="F7D218ABE81EFFA0FD87EB3FFD308D2B" box="[581,753,1547,1573]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="sichuanensis">
<emphasis id="02A6BF3AE81EFFA0FD87EB3FFD308D2B" box="[581,753,1547,1573]" italics="true" pageId="17" pageNumber="318">S. sichuanensis</emphasis>
</taxonomicName>
pronotum wholly black); laterotergites much dilated, posto-lateral margin of each laterotegite elevated, slightly produced than antero-lateral margin of following laterotegite (
<figureCitation id="A8E97FADE81EFFA0FF5DEB67FEDB8D63" box="[159,282,1619,1645]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="17" pageNumber="318">Fig. 8AC</figureCitation>
) (in
<taxonomicName id="F7D218ABE81EFFA0FE93EB67FE3F8D63" box="[337,510,1619,1645]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="sichuanensis">
<emphasis id="02A6BF3AE81EFFA0FE93EB67FE3F8D63" box="[337,510,1619,1645]" italics="true" pageId="17" pageNumber="318">S. sichuanensis</emphasis>
</taxonomicName>
laterotergites distinctly roundly laterally dilated, middle part of each laterotergite elevated); basal 2/3 of coria and whole clavus black or dark brown and remaining of coria white or orange (
<figureCitation id="A8E97FADE81EFFA0FAFFEB43FA558D9F" box="[1341,1428,1655,1681]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="17" pageNumber="318">Fig. 8G</figureCitation>
) (in
<taxonomicName id="F7D218ABE81EFFA0FF7DEBAFFEAD8DBB" box="[191,364,1691,1717]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="sichuanensis">
<emphasis id="02A6BF3AE81EFFA0FF7DEBAFFEAD8DBB" box="[191,364,1691,1717]" italics="true" pageId="17" pageNumber="318">S. sichuanensis</emphasis>
</taxonomicName>
coria yellowish to milk-white with basal part and apical angle black); coxae orange brown or reddish brown (in
<taxonomicName id="F7D218ABE81EFFA0FEA7EB8BFDD18DD7" box="[357,528,1727,1753]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="sichuanensis">
<emphasis id="02A6BF3AE81EFFA0FEA7EB8BFDD18DD7" box="[357,528,1727,1753]" italics="true" pageId="17" pageNumber="318">S. sichuanensis</emphasis>
</taxonomicName>
coxae black).
</paragraph>
<paragraph id="306D6328E81EFFA3FF05EBD7FBC98A4F" blockId="17.[151,1437,970,2041]" lastBlockId="18.[151,1437,151,321]" lastPageId="18" lastPageNumber="319" pageId="17" pageNumber="318">
Moreover, the new species and
<taxonomicName id="F7D218ABE81EFFA0FDECEBD7FD148DF2" authorityName="Hsiao" authorityYear="1979" box="[558,725,1763,1789]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="02A6BF3AE81EFFA0FDECEBD7FD148DF2" box="[558,725,1763,1789]" italics="true" pageId="17" pageNumber="318">Sycanus minor</emphasis>
</taxonomicName>
have high similarity in general appearance, i.e., male genitalia structure and general body coloration, and a very close relationship in molecular phylogenetic analyses (minimum interspecific p-distance of 1.8 %). However, the new species is distinct from
<taxonomicName id="F7D218ABE81EFFA0FBCEEA1FFBAC8C4A" authorityName="Hsiao" authorityYear="1979" box="[1036,1133,1835,1861]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="02A6BF3AE81EFFA0FBCEEA1FFBAC8C4A" box="[1036,1133,1835,1861]" italics="true" pageId="17" pageNumber="318">S. minor</emphasis>
</taxonomicName>
by the following features: pronotum with anterior pronotal lobe orange brown, somewhat with irregular dark brown suffusion and posterior pronotal lobe pale orange brown or reddish brown (
<figureCitation id="A8E97FADE81EFFA0FD19EA47FCEF8C83" box="[731,814,1907,1933]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="17" pageNumber="318">Fig. 8F</figureCitation>
) (in
<taxonomicName id="F7D218ABE81EFFA0FCA6EA47FC028C82" authorityName="Hsiao" authorityYear="1979" box="[868,963,1907,1933]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="02A6BF3AE81EFFA0FCA6EA47FC028C82" box="[868,963,1907,1933]" italics="true" pageId="17" pageNumber="318">S. minor</emphasis>
</taxonomicName>
pronotum black with lateral and posterior margins orange or anterior, lateral and posterior margins orange); abdominal sternites dark brown with discontinuous medial longitudinal black strip and small longitudinal black markings segmentally laterally (
<figureCitation id="A8E97FADE81EFFA0FB6CEA8FFAC58CDB" box="[1198,1284,1979,2005]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="17" pageNumber="318">Fig. 8C</figureCitation>
) (in
<taxonomicName id="F7D218ABE81EFFA0FAFEEA8FFA5D8CDA" authorityName="Hsiao" authorityYear="1979" box="[1340,1436,1979,2005]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="02A6BF3AE81EFFA0FAFEEA8FFA5D8CDA" box="[1340,1436,1979,2005]" italics="true" pageId="17" pageNumber="318">S. minor</emphasis>
</taxonomicName>
yellowish to red with median longitudinal stripe, segmental transversal stripe black); base of scape blackish brown, scape interleaved brownish yellow and blackish brown, pedicel dark brown with a brown suffusion medially, first and second flagellomeres brown (in
<taxonomicName id="F7D218ABE81DFFA3FDB3ED8FFD138BDA" authorityName="Hsiao" authorityYear="1979" box="[625,722,187,213]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="02A6BF3AE81DFFA3FDB3ED8FFD138BDA" box="[625,722,187,213]" italics="true" pageId="18" pageNumber="319">S. minor</emphasis>
</taxonomicName>
black), distal dorsal lobe of endosoma (ddl) centrally covered longitudinally with tiny prickles, surrounded with four large and long horn-shaped prickles centrally, and a few smaller prickles (
<figureCitation id="A8E97FADE81DFFA3FE9FEC37FE188A13" box="[349,473,259,285]" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="18" pageNumber="319">Fig. 9HK</figureCitation>
) (in
<taxonomicName id="F7D218ABE81DFFA3FDD0EC37FDB58A12" authorityName="Hsiao" authorityYear="1979" box="[530,628,259,285]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="02A6BF3AE81DFFA3FDD0EC37FDB58A12" box="[530,628,259,285]" italics="true" pageId="18" pageNumber="319">S. minor</emphasis>
</taxonomicName>
ddl with two feebly sclerotized stripe-shaped sclerites, apically with a feebly sclerotized horned process and laterally with five pairs of small spines).
</paragraph>
<caption id="64AD33A0E81DFFA3FF55EAA2FE9B8CF8" ID-DOI="http://doi.org/10.5281/zenodo.12758829" ID-Zenodo-Dep="12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="18" pageNumber="319" startId="18.[151,250,1942,1966]" targetBox="[183,1403,355,1919]" targetPageId="18" targetType="figure">
<paragraph id="306D6328E81DFFA3FF55EAA2FE9B8CF8" blockId="18.[151,1437,1942,2038]" pageId="18" pageNumber="319">
<emphasis id="02A6BF3AE81DFFA3FF55EAA2FED58CA0" bold="true" box="[151,276,1942,1966]" pageId="18" pageNumber="319">FIGURE 8.</emphasis>
<emphasis id="02A6BF3AE81DFFA3FED9EAA3FD698CA0" box="[283,680,1943,1966]" italics="true" pageId="18" pageNumber="319">
<taxonomicName id="F7D218ABE81DFFA3FED9EAA3FD658CA0" authority="Truong &amp; Ha" authorityName="Truong &amp; Giang &amp; Nguyen &amp; Chau &amp; Ha" authorityYear="2024" box="[283,676,1943,1966]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="taynguyenensis" status="sp. nov.">Sycanus taynguyenensis Truong &amp; Ha</taxonomicName>
,
</emphasis>
<emphasis id="02A6BF3AE81DFFA3FD6DEAA2FCC08CA0" bold="true" box="[687,769,1942,1966]" pageId="18" pageNumber="319">
<taxonomicNameLabel id="19950241E81DFFA3FD6DEAA2FCC08CA0" box="[687,769,1942,1966]" pageId="18" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="02A6BF3AE81DFFA3FCCDEAA2FBF38CA0" box="[783,1074,1942,1969]" italics="true" pageId="18" pageNumber="319">holotype, ♂, NDD2019-285.</emphasis>
A,
<emphasis id="02A6BF3AE81DFFA3FB9AEAA2FAEA8CA0" box="[1112,1323,1942,1966]" italics="true" pageId="18" pageNumber="319">body in dorsal view;</emphasis>
B,
<emphasis id="02A6BF3AE81DFFA3FA92EAA2FED98CDC" italics="true" pageId="18" pageNumber="319">body in lateral view;</emphasis>
C, body
<emphasis id="02A6BF3AE81DFFA3FEB3EA8FFCC38CDC" box="[369,770,1978,2002]" italics="true" pageId="18" pageNumber="319">in ventral view; D, head in lateral view;</emphasis>
E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra.
</paragraph>
</caption>
<paragraph id="306D6328E81CFFA2FF55EDA2FEA98BBE" blockId="19.[151,360,150,176]" box="[151,360,150,176]" pageId="19" pageNumber="320">
<emphasis id="02A6BF3AE81CFFA2FF55EDA2FEA98BBE" bold="true" box="[151,360,150,176]" pageId="19" pageNumber="320">Male description.</emphasis>
</paragraph>
<paragraph id="306D6328E81CFFA2FF55EDEAFC2F8836" blockId="19.[151,1437,222,2012]" pageId="19" pageNumber="320">
<emphasis id="02A6BF3AE81CFFA2FF55EDEAFEEE8BF6" bold="true" box="[151,303,222,248]" pageId="19" pageNumber="320">Colouration.</emphasis>
Body pale orange brown. Head dorsum, neck, clypeus, antenniferous tubercles, labrum, and head venter black; maxillary plate, gena black or blackish brown; a brownish yellow suffusion next to outer side of ocellus; first visible labial segment blackish brown; base of second visible labial segment dark brown; remaining of labium brown, paler as pale brown toward tip (
<figureCitation id="A8E97FADE81CFFA2FD5AEC7EFCCF8A6A" box="[664,782,330,356]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8D, E</figureCitation>
). Base of scape blackish brown; scape alternating brownish yellow and blackish brown; pedicel dark brown with a brown suffusion medially; first and second flagellomeres brown. Collar pale orange brown or pale reddish brown (
<figureCitation id="A8E97FADE81CFFA2FCE4ECA6FCBC8AA2" box="[806,893,402,428]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8F</figureCitation>
); anterolateral angle, anterior pronotal lobe and prosternum orange brown, somewhat with irregular dark brown suffusion (
<figureCitation id="A8E97FADE81CFFA2FC37EC82FB8C8ADE" box="[1013,1101,438,464]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8F</figureCitation>
); anterior acetabulum orange brown; stridulatory sulcus luteous; posterior pronotal lobe pale orange brown or reddish brown (
<figureCitation id="A8E97FADE81CFFA2FB04ECEEFADA8AFA" box="[1222,1307,474,500]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8F</figureCitation>
); scutellum black with orange brown centrally (
<figureCitation id="A8E97FADE81CFFA2FDFBECCAFD538916" box="[569,658,510,536]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8F</figureCitation>
); scutellar spine wholly orange brown or reddish brown (
<figureCitation id="A8E97FADE81CFFA2FAF7ECCAFA4F8916" box="[1333,1422,510,536]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8F</figureCitation>
); meso- and metapleura, meso- and metasterna, dark brown with irregular blackish brown sufusions; coxae orange brown or reddish brown; trochanters and basal half of femora dark brown; apical half of femora, tibiae, and tarsi blackish brown. Basal 2/3 of coria and whole clavus black or dark brown; remaining of coria white or orange; clavus dark brown; membrane pale bronzy brown, semi-hyaline (
<figureCitation id="A8E97FADE81CFFA2FCF2EFBAFC4D89A6" box="[816,908,654,680]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8G</figureCitation>
). Hind wings faintly semi-hyaline. Abdominal mediotergites dark brown with irregular brown suffusions; laterotergites sanguineous or reddish orange (
<figureCitation id="A8E97FADE81CFFA2FAFFEF86FF6D89FE" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8A, B</figureCitation>
); each laterotegites IIIVII with large black markings in anterior and posterior margins ((
<figureCitation id="A8E97FADE81CFFA2FB77EFE2FACF89FE" box="[1205,1294,726,752]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8B</figureCitation>
); abdominal sternites dark brown with discontinuous medial longitudinal black strip and small longitudinal black markings segmentally laterally (
<figureCitation id="A8E97FADE81CFFA2FE53EE2AFE288836" box="[401,489,798,824]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8C</figureCitation>
). Pygophore orange or sanguineous (
<figureCitation id="A8E97FADE81CFFA2FC4BEE2AFC208836" box="[905,993,798,824]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8C</figureCitation>
).
</paragraph>
<paragraph id="306D6328E81CFFA2FF05EE76FDCE8852" blockId="19.[151,1437,222,2012]" box="[199,527,834,860]" pageId="19" pageNumber="320">
<emphasis id="02A6BF3AE81CFFA2FF05EE76FF398852" bold="true" box="[199,248,834,860]" pageId="19" pageNumber="320">Var.</emphasis>
abdomen beneath black.
</paragraph>
<paragraph id="306D6328E81CFFA5FF05EE52FDD28BDB" blockId="19.[151,1437,222,2012]" lastBlockId="20.[151,1436,151,213]" lastPageId="20" lastPageNumber="321" pageId="19" pageNumber="320">
<emphasis id="02A6BF3AE81CFFA2FF05EE52FE80888F" bold="true" box="[199,321,870,897]" pageId="19" pageNumber="320">Structure.</emphasis>
Body large-sized (
<quantity id="F72ACECDE81CFFA2FDD4EE52FD4C888E" box="[534,653,870,897]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="19" pageNumber="320" unit="mm" value="15.5">15.50 mm</quantity>
), elongated and posteriorly widened. Head tubular, slender, elongated (
<figureCitation id="A8E97FADE81CFFA2FF5DEEBEFEDA88AA" box="[159,283,906,932]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8D, E</figureCitation>
); head and neck together much longer than pronotum (
<figureCitation id="A8E97FADE81CFFA2FC4DEEBEFBCA88AA" box="[911,1035,906,932]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8A, B</figureCitation>
); anteocular area of head elongate-conical; anteclypeus cylindrical and prominent; postocular area of head weakly globose, distinctly wider than anteocular area, shorter than anteocular area, constricted behind compound eyes, with a wide and deep interocular sulcus; neck long (
<figureCitation id="A8E97FADE81CFFA2FEABEEC2FE238F1E" box="[361,482,1014,1040]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8D, E</figureCitation>
). Compound eyes protruding laterally, nearly globose, oblique with respect to ventral margin of head; lateral ocelli produced, slightly elevated behind interocular sulcus, separated from each other; interspace between lateral ocelli wider than distance between compound eye and lateral ocellus (
<figureCitation id="A8E97FADE81CFFA2FB18E90AFA948F56" box="[1242,1365,1086,1112]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8D, E</figureCitation>
). First visible labial segment slightly thicker and much shorter than second segment, longer than anteocular area of head, not extending beyond level of middle of compound eye when labium laid backward (
<figureCitation id="A8E97FADE81CFFA2FB97E9B2FB718FAE" box="[1109,1200,1158,1184]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8D</figureCitation>
); proportional length of first to third visible labial segments 2.2:3.7:0.7. Scape about 1.7 times as long as head, about 2.4 times as long as pedicel, longer than second flagellomere; first flagellomere about half as long as pedicel and about 1/4 times as long as second flagellomere; proportional length of scape, pedicel, first and second flagellomeres 5.5:2.3:1.3:4.8. Collar thin, almost invisible in dorsal view, with anterolateral angle roundly produced anteriorly; anterior pronotal lobe small, hemisphered and bulged, smooth, deeply depressed at base; posterior pronotal lobe rugulose, shallowly depressed on disc, with slightly swollen anteromedial elevation (never sulcate or concave); humerus bluntly triangular, with round apex; posterior margin of pronotum substraight; posterior angles round, weakly exceeded to posterior margin of pronotum (
<figureCitation id="A8E97FADE81CFFA2FDD3E892FDA68ECE" box="[529,615,1446,1472]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8F</figureCitation>
). Scutellum triangular, somewhat triangularly depressed basally, apically produced with a scutellar spine, and sloping downward, posterior apex round (
<figureCitation id="A8E97FADE81CFFA2FBC3E8FEFB948EEA" box="[1025,1109,1482,1508]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8F</figureCitation>
); scutellar spine sub-vertical, short (
<quantity id="F72ACECDE81CFFA2FF22E8DAFE8A8D06" box="[224,331,1518,1545]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="19" pageNumber="320" unit="mm" value="1.45">1.45 mm</quantity>
), bifid at apex (
<figureCitation id="A8E97FADE81CFFA2FDC2E8DAFDB88D06" box="[512,633,1518,1544]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8B, F</figureCitation>
). Femora elongated, slender but stout, moderate subnodulose apically; tibiae slender and elongated. Hemelytra surpassing beyond apex of abdomen when fully closed, 0.7 times as long as body length (
<figureCitation id="A8E97FADE81CFFA2FE8FEB02FE698D5E" box="[333,424,1590,1616]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8A</figureCitation>
); discal cell nearly diamond-shaped, longer than width (
<figureCitation id="A8E97FADE81CFFA2FBE5EB02FB438D5E" box="[1063,1154,1590,1616]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8G</figureCitation>
); Sc 0.7 times as long as hemelytron length, 1.4 times as long as R + M (
<figureCitation id="A8E97FADE81CFFA2FD04EB6EFCE28D7A" box="[710,803,1626,1652]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8G</figureCitation>
). Hind wing damaged. Laterotergites much dilated and ascending with segmental incisures, posto-lateral margin of each laterotegite slightly produced than antero-lateral margin of following laterotegite (
<figureCitation id="A8E97FADE81CFFA2FDD8EB96FDB48DB2" box="[538,629,1698,1724]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="19" pageNumber="320">Fig. 8B</figureCitation>
). Pygophore ovoid (
<figureCitation id="A8E97FADE81CFFA2FCA1EB96FC238DB2" box="[867,994,1698,1724]" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="19" pageNumber="320">Fig. 9BD</figureCitation>
); median process of pygophore (mpp) posteriorly produced, 0.39 times as long as wide in dorsal view, with apical margin slightly convex, distinctly produced a round apex, apicolateral corner sub-round, distinctly produced posterolaterad (
<figureCitation id="A8E97FADE81CFFA2FB61EBDEFADF8C0A" box="[1187,1310,1770,1796]" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="19" pageNumber="320">Fig. 9BE</figureCitation>
); paramere long, slender, clavate, somewhat incurved and larger in apical part, with round apex (
<figureCitation id="A8E97FADE81CFFA2FBBFEA3AFB388C26" box="[1149,1273,1806,1832]" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="19" pageNumber="320">Fig. 9FG</figureCitation>
). Aedeagus in dorsal view ovoid, dorsally sclerotized and in lateral view long and narrow (
<figureCitation id="A8E97FADE81CFFA2FC2AEA06FBA08C42" box="[1000,1121,1842,1868]" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="19" pageNumber="320">Figs. 9H, I</figureCitation>
); articulatory apparatus (aa) in ventral view with basal plate arms relatively slender and jointly forming a V-shape, and in lateral view arched strongly (
<figureCitation id="A8E97FADE81CFFA2FEC6EA4EFEB68C9A" box="[260,375,1914,1940]" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="19" pageNumber="320">Fig. 9H, I</figureCitation>
); dorsal phallothecal sclerite (dps) in lateral view with posteromedian weakly produced (
<figureCitation id="A8E97FADE81CFFA2FAADEA4EFF738CB6" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="19" pageNumber="320">Fig. 9I</figureCitation>
); spoon-like sclerites (sps) weakly anteriorly produced, hyaline and glabrous (
<figureCitation id="A8E97FADE81CFFA2FB81EAAAFB7B8CB6" box="[1091,1210,1950,1976]" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="19" pageNumber="320">Fig. 9H, I</figureCitation>
); membranous sac-like lobes almost disappeared (
<figureCitation id="A8E97FADE81CFFA2FE34EAF6FD938CD2" box="[502,594,1986,2012]" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="19" pageNumber="320">Fig. 9H</figureCitation>
); distal dorsal lobe of endosoma (ddl) round and weakly bulged, centrally covered longitudinally with tiny prickles, surrounded with four large and long horn-shaped prickles centrally, and a few smaller prickles (
<figureCitation id="A8E97FADE81BFFA5FE48ED8FFDC68BDB" box="[394,519,187,213]" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="20" pageNumber="321">Fig. 9HK</figureCitation>
).
</paragraph>
<caption id="64AD33A0E81BFFA5FF55EA5CFB878CE2" ID-DOI="http://doi.org/10.5281/zenodo.12758831" ID-Zenodo-Dep="12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="20" pageNumber="321" startId="20.[151,250,1896,1920]" targetBox="[328,1261,237,1872]" targetPageId="20" targetType="figure">
<paragraph id="306D6328E81BFFA5FF55EA5CFB878CE2" blockId="20.[151,1436,1896,2028]" pageId="20" pageNumber="321">
<emphasis id="02A6BF3AE81BFFA5FF55EA5CFED48C8E" bold="true" box="[151,277,1896,1920]" pageId="20" pageNumber="321">FIGURE 9.</emphasis>
<emphasis id="02A6BF3AE81BFFA5FEDFEA5CFD6C8C8E" box="[285,685,1896,1920]" italics="true" pageId="20" pageNumber="321">
<taxonomicName id="F7D218ABE81BFFA5FEDFEA5CFD688C8E" authority="Truong &amp; Ha" authorityName="Truong &amp; Giang &amp; Nguyen &amp; Chau &amp; Ha" authorityYear="2024" box="[285,681,1896,1920]" class="Insecta" family="Reduviidae" genus="Sycanus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="taynguyenensis" status="sp. nov.">Sycanus taynguyenensis Truong &amp; Ha</taxonomicName>
,
</emphasis>
<emphasis id="02A6BF3AE81BFFA5FD77EA5CFCC68C8E" bold="true" box="[693,775,1896,1920]" pageId="20" pageNumber="321">
<taxonomicNameLabel id="19950241E81BFFA5FD77EA5CFCC68C8E" box="[693,775,1896,1920]" pageId="20" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia,
<emphasis id="02A6BF3AE81BFFA5FEB3EAB8FD528CAA" box="[369,659,1932,1958]" italics="true" pageId="20" pageNumber="321">holotype, ♂, NDD2019-285.</emphasis>
BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl).
</paragraph>
</caption>
<paragraph id="306D6328E81AFFA4FF05EDA2FF7D8AFB" blockId="21.[151,1437,150,501]" pageId="21" pageNumber="322">
<emphasis id="02A6BF3AE81AFFA4FF05EDA2FEFB8BBE" bold="true" box="[199,314,150,176]" pageId="21" pageNumber="322">Vestiture.</emphasis>
Body clothed with yellow, shining, griseous setae interleaved with black setae. Head with moderately densely bent setae (
<figureCitation id="A8E97FADE81AFFA4FEB9ED8FFE3B8BDB" box="[379,506,187,213]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="21" pageNumber="322">Fig. 8D, E</figureCitation>
); first visible labial segment somewhat with a few bent setae; remaining labium without setae (
<figureCitation id="A8E97FADE81AFFA4FEF9EDEBFE548BF7" box="[315,405,223,249]" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="21" pageNumber="322">Fig. 8D</figureCitation>
). Scape with some short, slender, oblique setae, denser apically; pedicel with short, oblique, setae, denser than first joint, and denser apically; first and second flagellomeres with abundantly short, slender, more oblique than second joint setae. Collar, anterolateral angles, posterior pronotal lobe, scutellum, pleuron, sternum, coxae, coria with dense of bent, yellowish griseous setae, interleaved with long, erect, vertical, black setae (
<figureCitation id="A8E97FADE81AFFA4FAADEC7FFF7C8A87" captionStart="FIGURE 8" captionStartId="18.[151,250,1942,1966]" captionTargetBox="[183,1403,355,1919]" captionTargetId="figure-101@18.[183,1403,345,1919]" captionTargetPageId="18" captionText="FIGURE 8. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., holotype, ♂, NDD2019-285. A, body in dorsal view; B, body in lateral view; C, body in ventral view; D, head in lateral view; E, head in dorsal view; F, pronotum and scutellum in dorsal view; G, right hemelytra." figureDoi="http://doi.org/10.5281/zenodo.12758829" httpUri="https://zenodo.org/record/12758829/files/figure.png" pageId="21" pageNumber="322">Fig. 8H</figureCitation>
); abdomen beneath with straight, slender, oblique griseous setae; trochanter, femora, tibia with long, slender, erect, vertical, black setae; Male pygophore ventrally moderately densely with bent, slender, yellowish griseous setae, somewhat with black setae (
<figureCitation id="A8E97FADE81AFFA4FDD5EC83FD4F8ADF" box="[535,654,439,465]" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="21" pageNumber="322">Fig. 9BE</figureCitation>
); paramere apically with some long, yellowish griseous setae (
<figureCitation id="A8E97FADE81AFFA4FA84EC83FF6E8AFB" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="21" pageNumber="322">Fig. 9F, G</figureCitation>
).
</paragraph>
<paragraph id="306D6328E81AFFA4FF55EF17FE438932" blockId="21.[151,784,546,609]" box="[151,386,546,573]" pageId="21" pageNumber="322">
<emphasis id="02A6BF3AE81AFFA4FF55EF17FE438932" bold="true" box="[151,386,546,573]" pageId="21" pageNumber="322">Female description.</emphasis>
</paragraph>
<paragraph id="306D6328E81AFFA4FF55EF73FCD1896F" blockId="21.[151,784,546,609]" box="[151,784,583,609]" pageId="21" pageNumber="322">General external morphology similar to that of the male.</paragraph>
<paragraph id="306D6328E81AFFA4FF55EFBAFDBC89FF" blockId="21.[151,1437,654,1725]" pageId="21" pageNumber="322">
<emphasis id="02A6BF3AE81AFFA4FF55EFBAFEDE89A6" bold="true" box="[151,287,654,680]" pageId="21" pageNumber="322">Coloration.</emphasis>
Almost similar to male but differ in the following characters. Abdominal sternites wholly black; Gonocoxa VIII black with a reddish-brown spot near posterior apex; abdominal laterotergite VIII, abdominal tergite IX, and gonapophysis VIII black (
<figureCitation id="A8E97FADE81AFFA4FDD4EFE3FDB189FF" box="[534,624,727,753]" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="21" pageNumber="322">Fig. 9A</figureCitation>
).
</paragraph>
<paragraph id="306D6328E81AFFA4FF05EFCEFE7C8F73" blockId="21.[151,1437,654,1725]" pageId="21" pageNumber="322">
<emphasis id="02A6BF3AE81AFFA4FF05EFCEFE80881A" bold="true" box="[199,321,762,788]" pageId="21" pageNumber="322">Structure.</emphasis>
Almost the same as male but larger than male and differ in the following characters. Body large-sized (
<quantity id="F72ACECDE81AFFA4FF21EE2BFE9F8837" box="[227,350,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="2.134" pageId="21" pageNumber="322" unit="mm" value="21.34">21.34 mm</quantity>
), elongated, and somewhat robust. Head together with neck shorter than or almost as long as pronotum. Proportional length of first to third visible labial segments 2.2:4.1:0.9. Scape about 1.8 times as long as head, about 2.5 times as long as pedicel, longer than second flagellomere; first flagellomere about half as long as pedicel and about 1/5 times as long as second flagellomere; proportional average length of scape, pedicel, first and second flagellomeres 6.4:2.5:1.3:5.9. Abdominal laterotergite VIII with thin posterior margin, not exceed the posterior margin of abdominal tergite IX; abdominal sternite VII forming a wide sub-pentagonal median concavity, with posteromedian margin gently U-shaped, with inner posterolateral margin slightly concave but weakly convex medially; gonocoxa VIII subtriangular, produced mesad and forming a pointed apex at apical inner corner, and with inner margin weakly incurved in posterior 3/4; gonapophysis VIII subtriangular, posteriorly produced; abdominal tergite IX small (
<figureCitation id="A8E97FADE81AFFA4FE94E957FE718F73" box="[342,432,1123,1149]" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="21" pageNumber="322">Fig. 9A</figureCitation>
).
</paragraph>
<paragraph id="306D6328E81AFFA4FF05E9B2FD2C8E03" blockId="21.[151,1437,654,1725]" pageId="21" pageNumber="322">
<emphasis id="02A6BF3AE81AFFA4FF05E9B2FEFB8FAE" bold="true" box="[199,314,1158,1184]" pageId="21" pageNumber="322">Vestiture.</emphasis>
Almost the same as male except for the following characters. Anterior 1/3 of gonocoxa VIII almost no setae; posterior 2/3 of gonocoxa VIII covered with short and long slender black setae; posterior margin of gonapophysis VIII densely covered with long erect black setae; abdominal laterotergite VIII and abdominal tergite IX covered with short and long black setae (
<figureCitation id="A8E97FADE81AFFA4FD44E9C7FD218E03" box="[646,736,1267,1293]" captionStart="FIGURE 9" captionStartId="20.[151,250,1896,1920]" captionTargetBox="[328,1261,237,1872]" captionTargetId="figure-45@20.[327,1261,237,1872]" captionTargetPageId="20" captionText="FIGURE 9. Sycanus taynguyenensis Truong &amp; Ha, sp. nov., A, female genitalia in ventral view, paratype, ♀, EG2018-395; BK, male genitalia, holotype, ♂, NDD2019-285. BE, pygophore with parameres; B, dorsal view; C, lateral view; D, ventral view; E, caudal view; FG, left paramere; F, dorsal view; G, lateral view; HJ, aedeagus with endosoma semi-everted dorsal view; H, dorsal view; I, lateral view; J, ventral view; K, distal dorsal lobe of endosoma (ddl)." figureDoi="http://doi.org/10.5281/zenodo.12758831" httpUri="https://zenodo.org/record/12758831/files/figure.png" pageId="21" pageNumber="322">Fig. 9A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="78C830A3E81AFFA4FF05E823FE7C8D23" pageId="21" pageNumber="322" type="description">
<paragraph id="306D6328E81AFFA4FF05E823FE7C8D23" blockId="21.[151,1437,654,1725]" pageId="21" pageNumber="322">
<emphasis id="02A6BF3AE81AFFA4FF05E823FEBF8E3F" bold="true" box="[199,382,1303,1329]" pageId="21" pageNumber="322">Measurements.</emphasis>
All dimensions are given in mm.
<emphasis id="02A6BF3AE81AFFA4FD31E823FCAF8E3E" bold="true" box="[755,878,1302,1329]" pageId="21" pageNumber="322">
<typeStatus id="EF69DD8AE81AFFA4FD31E823FCA18E3F" box="[755,864,1303,1329]" pageId="21" pageNumber="322" type="holotype">Holotype</typeStatus>
(
</emphasis>
<specimenCount id="26D4A8A1E81AFFA4FCADE823FC458E3F" box="[879,900,1303,1329]" count="1" pageId="21" pageNumber="322" type="male"></specimenCount>
<emphasis id="02A6BF3AE81AFFA4FC46E822FC578E3E" bold="true" box="[900,918,1302,1328]" pageId="21" pageNumber="322">):</emphasis>
BL 15.50; HL 3.29; AoL 1.47; AoW 0.77; PoL 1.02; PoW 1.08; NL 1.16; OE 1.42; IE 0.63; ED 0.89; OD 0.16; OCD 0.43; COD 0.17; R1L 2.22; R2L 3.68; R3L 0.71; A1L 5.47; A2L 2.25; A3L 1.33; A4L 4.80; PnL 2.82; PnW 3.36; APL 0.87; PPL 1.96; SPL 1.45; HeL 10.39; HeW 3.52; Sc 7.66; R+M 5.32; AFL 5.60; ATL 6.31; MFL 5.37; MTL 6.21; PFL 6.79; PTL 8.30.
<emphasis id="02A6BF3AE81AFFA4FB0DE8B7FA8A8E92" bold="true" box="[1231,1355,1410,1437]" pageId="21" pageNumber="322">
<typeStatus id="EF69DD8AE81AFFA4FB0DE8B7FAFD8E93" box="[1231,1340,1411,1437]" pageId="21" pageNumber="322" type="paratype">Paratype</typeStatus>
(
</emphasis>
<specimenCount id="26D4A8A1E81AFFA4FA89E8B7FAA18E93" box="[1355,1376,1411,1437]" count="1" pageId="21" pageNumber="322" type="female"></specimenCount>
<emphasis id="02A6BF3AE81AFFA4FAA2E8B6FAB28E92" bold="true" box="[1376,1395,1410,1436]" pageId="21" pageNumber="322">):</emphasis>
BL 21.34; HL 3.50; AoL 1.74; AoW 0.89; PoL 0.95; PoW 1.16; NL 1.37; OE 1.56; IE 0.70; ED 0.91; OD 0.15; OCD 0.42; COD 0.15; R1L 2.21; R2L 4.07; R3L 0.90; A1L 6.39; A2L 2.53; A3L 1.32; A4L 5.86; PnL 3.89; PnW 4.54; APL 1.17; PPL 2.72; SPL 2.31; HeL 12.24; HeW 4.58; Sc 9.43; R+M 6.53; AFL 6.20; ATL 7.24; MFL 5.63; MTL 6.87; PFL 7.36; PTL 9.67.
</paragraph>
</subSubSection>
<subSubSection id="78C830A3E81AFFA4FF05EB02FBE58D5F" box="[199,1060,1590,1617]" pageId="21" pageNumber="322" type="distribution">
<paragraph id="306D6328E81AFFA4FF05EB02FBE58D5F" blockId="21.[151,1437,654,1725]" box="[199,1060,1590,1617]" pageId="21" pageNumber="322">
<emphasis id="02A6BF3AE81AFFA4FF05EB02FEA08D5E" bold="true" box="[199,353,1590,1616]" pageId="21" pageNumber="322">Distribution.</emphasis>
<collectingCountry id="48C523B8E81AFFA4FEA5EB03FE088D5F" box="[359,457,1591,1617]" name="Vietnam" pageId="21" pageNumber="322">Vietnam</collectingCountry>
, Central Highlands (
<collectingRegion id="F216ADCAE81AFFA4FD6DEB03FCD28D5F" box="[687,787,1591,1617]" country="Vietnam" name="Dak Lak" pageId="21" pageNumber="322">Dak Lak</collectingRegion>
and
<collectingRegion id="F216ADCAE81AFFA4FC8BEB03FC5C8D5F" box="[841,925,1591,1617]" country="Vietnam" name="Gia Lai" pageId="21" pageNumber="322">Gia Lai</collectingRegion>
Provinces).
</paragraph>
</subSubSection>
<subSubSection id="78C830A3E81AFFA4FF05EB6FFBE88D7B" box="[199,1065,1626,1653]" pageId="21" pageNumber="322" type="materials_examined">
<paragraph id="306D6328E81AFFA4FF05EB6FFBE88D7B" blockId="21.[151,1437,654,1725]" box="[199,1065,1626,1653]" pageId="21" pageNumber="322">
<materialsCitation id="80BA6975E81AFFA4FF05EB6FFBE48D7B" box="[199,1061,1626,1653]" country="Vietnam" location="Provinces" municipality="Central Highlands" pageId="21" pageNumber="322" specimenCount="1" stateProvince="Dak Lak" typeStatus="holotype">
<emphasis id="02A6BF3AE81AFFA4FF05EB6FFEA78D7A" bold="true" box="[199,358,1626,1653]" pageId="21" pageNumber="322">
<typeStatus id="EF69DD8AE81AFFA4FF05EB6FFEC38D7B" box="[199,258,1627,1653]" pageId="21" pageNumber="322">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="48C523B8E81AFFA4FEAEEB6FFE0F8D7B" box="[364,462,1627,1653]" name="Vietnam" pageId="21" pageNumber="322">Vietnam</collectingCountry>
,
<collectingMunicipality id="D009F952E81AFFA4FE1BEB6FFD678D7B" box="[473,678,1627,1653]" pageId="21" pageNumber="322">Central Highlands</collectingMunicipality>
(
<collectingRegion id="F216ADCAE81AFFA4FD76EB6FFCD98D7B" box="[692,792,1627,1653]" country="Vietnam" name="Dak Lak" pageId="21" pageNumber="322">Dak Lak</collectingRegion>
and
<collectingRegion id="F216ADCAE81AFFA4FC8CEB6FFC638D7B" box="[846,930,1627,1653]" country="Vietnam" name="Gia Lai" pageId="21" pageNumber="322">Gia Lai</collectingRegion>
<location id="350D35F3E81AFFA4FC6BEB6FFBDC8D7B" LSID="urn:lsid:plazi:treatment:B87BD23EE81EFFA4FF55EE6BFE528DB3:350D35F3E81AFFA4FC6BEB6FFBDC8D7B" box="[937,1053,1627,1653]" country="Vietnam" municipality="Central Highlands" name="Provinces" pageId="21" pageNumber="322" stateProvince="Dak Lak">Provinces</location>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="78C830A3E81AFFA4FF05EB4BFE528DB3" pageId="21" pageNumber="322" type="etymology">
<paragraph id="306D6328E81AFFA4FF05EB4BFE528DB3" blockId="21.[151,1437,654,1725]" pageId="21" pageNumber="322">
<emphasis id="02A6BF3AE81AFFA4FF05EB4BFE8F8D97" bold="true" box="[199,334,1663,1689]" pageId="21" pageNumber="322">Etymology.</emphasis>
This new species is named after the Tay Nguyen region, the local name of Central Highlands,
<collectingCountry id="48C523B8E81AFFA4FF55EB97FF388DB3" box="[151,249,1699,1725]" name="Vietnam" pageId="21" pageNumber="322">Vietnam</collectingCountry>
; an adjective.
</paragraph>
</subSubSection>
</treatment>
</document>

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