285 lines
43 KiB
XML
285 lines
43 KiB
XML
<document id="B47B735A7EDE5C299D881E40B6767C9F" ID-DOI="10.11646/zootaxa.3847.1.1" ID-GBIF-Dataset="774be4c8-f95f-4d06-a555-dbe3f1aa2f21" ID-ISSN="1175-5326" ID-Zenodo-Dep="286815" ID-ZooBank="C7BDD136-2686-4049-B395-684797B26406" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1460469621688" checkinUser="plazi" docAuthor="Van Der Geer, Alexandra A. E." docDate="2014" docId="03D287A2FD11371DFEF7FD91FF66FDA5" docLanguage="en" docName="zt03847p032.pdf" docOrigin="Zootaxa 3847 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hoplitomeryx macpheei Van Der Geer 2014, sp. nov." docType="treatment" docVersion="10" lastPageNumber="26" masterDocId="FFEBFFDAFD093704FE60FFB5FF9CFFC5" masterDocTitle="Systematic revision of the family Hoplitomerycidae Leinders, 1984 (Artiodactyla: Cervoidea), with the description of a new genus and four new species" masterLastPageNumber="32" masterPageNumber="1" pageNumber="25" updateTime="1730803629954" updateUser="ExternalLinkService">
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<mods:title id="E042B8CF9DE8C3CA741767DC179BEA96">Systematic revision of the family Hoplitomerycidae Leinders, 1984 (Artiodactyla: Cervoidea), with the description of a new genus and four new species</mods:title>
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<mods:roleTerm id="A5FF92A19116D911596CD9D0A82AD9E9">Author</mods:roleTerm>
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<mods:namePart id="008F7889BD2193C6960ADEB69B98368A">Van Der Geer, Alexandra A. E.</mods:namePart>
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<mods:title id="2C983C91D5A823158E5F9067F510576D">Zootaxa</mods:title>
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<mods:part id="ECF81EB27BE9775A4450A302753B14D0">
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<mods:date id="E0A7C17ED61D1CD49DAE1943FC1377F5">2014</mods:date>
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<mods:number id="D52FD91D2B1ADCBB89949CDB183C2E8F">2014-08-06</mods:number>
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<mods:detail id="716FA7DA8E0A289B9C306FE1BB239D0F" type="volume">
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<mods:number id="4008786EF3C7E9E998D321D5D76FD5AF">3847</mods:number>
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<treatment id="03D287A2FD11371DFEF7FD91FF66FDA5" ID-DOI="http://doi.org/10.5281/zenodo.5630052" ID-GBIF-Taxon="127668901" ID-Zenodo-Dep="5630052" LSID="urn:lsid:plazi:treatment:03D287A2FD11371DFEF7FD91FF66FDA5" httpUri="http://treatment.plazi.org/id/03D287A2FD11371DFEF7FD91FF66FDA5" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
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<subSubSection id="C361653FFD11371CFEF7FD91FF63FD9B" pageId="24" pageNumber="25" type="nomenclature">
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<paragraph id="8BC436B4FD11371CFEF7FD91FD81FDFB" blockId="24.[151,541,548,607]" box="[151,541,548,574]" pageId="24" pageNumber="25">
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<heading id="D08C81D8FD11371CFEF7FD91FD81FDFB" bold="true" box="[151,541,548,574]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
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<emphasis id="B90FEAA6FD11371CFEF7FD91FD81FDFB" bold="true" box="[151,541,548,574]" pageId="24" pageNumber="25">
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<taxonomicName id="4C7B4D37FD11371CFEF7FD91FE24FDFB" authority="Van Der Geer, 2014" authorityName="Van Der Geer" authorityYear="2014" box="[151,440,548,574]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="macpheei" status="sp. nov.">
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<emphasis id="B90FEAA6FD11371CFEF7FD91FE24FDFB" bold="true" box="[151,440,548,574]" italics="true" pageId="24" pageNumber="25">Hoplitomeryx macpheei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23C57DDFD11371CFFDFFD91FD81FDFB" box="[447,541,548,574]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC436B4FD11371CFEF7FDF3FF63FD9B" blockId="24.[151,541,548,607]" box="[151,255,582,607]" pageId="24" pageNumber="25">
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<figureCitation id="13402A31FD11371CFEF7FDF3FF4BFD9B" box="[151,215,582,607]" captionStart="FIGURE 7" captionStartId="23.[151,250,1887,1909]" captionTargetBox="[239,1344,201,1864]" captionTargetId="figure@23.[220,1375,193,1881]" captionTargetPageId="23" captionText="FIGURE 7. Holotypes of three new Hoplitomeryx species from Gargano, South Italy (Late Miocene; Middle or Late Turolian, MN 12 - 13). A – E Hoplitomeryx devosi sp. n., left metacarpal RGM 178.517; F – I Hoplitomeryx macpheei sp. n., left metacarpal RGM 260.918; J – M Hoplitomeryx kriegsmani sp. n., left metacarpal RGM 178.516. A, F, J proximal view. B, G, K dorsal view. C, H, L palmar view. D, I, M medial view. E distal view. Note that the sideward and backward bending of RGM 178.516 is a post-mortem defect." figureDoi="http://doi.org/10.5281/zenodo.286822" httpUri="https://zenodo.org/record/286822/files/figure.png" pageId="24" pageNumber="25">Fig. 7</figureCitation>
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F–I
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</paragraph>
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</subSubSection>
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<subSubSection id="C361653FFD11371CFEF7FD3BFEB5FCDA" pageId="24" pageNumber="25" type="reference_group">
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||
<paragraph id="8BC436B4FD11371CFEF7FD3BFB1EFD61" blockId="24.[151,1436,654,799]" box="[151,1154,654,676]" pageId="24" pageNumber="25">
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<taxonomicName id="4C7B4D37FD11371CFEF7FD3BFE5EFD61" authority="Leinders, 1984" authorityName="Leinders" authorityYear="1984" box="[151,450,654,676]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
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<emphasis id="B90FEAA6FD11371CFEF7FD3BFEBEFD61" box="[151,290,654,676]" italics="true" pageId="24" pageNumber="25">Hoplitomeryx</emphasis>
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||
<bibRefCitation id="EFEA4B45FD11371CFF48FD3BFE5EFD61" author="Leinders, J." box="[296,450,654,676]" firstAuthor="Leinders" journalOrPublisher="Scripta Geologica" pageId="24" pageNumber="25" pagination="1 - 68" part="70" refId="ref24476" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." title="Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy)" type="journal article" year="1984">Leinders, 1984</bibRefCitation>
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</taxonomicName>
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, horn core
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<typeStatus id="54C08816FD11371CFC53FD3AFDC3FD61" box="[563,607,655,676]" pageId="24" pageNumber="25">type</typeStatus>
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II—Leinders 1984: p. 21, 22, pl. 5B [RGM 260.902].
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</paragraph>
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<paragraph id="8BC436B4FD11371CFEF7FD18FAF8FD06" blockId="24.[151,1436,654,799]" box="[151,1380,685,707]" pageId="24" pageNumber="25">
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<taxonomicName id="4C7B4D37FD11371CFEF7FD18FE5EFD06" authority="Leinders, 1984" authorityName="Leinders" authorityYear="1984" box="[151,450,685,707]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
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<emphasis id="B90FEAA6FD11371CFEF7FD18FEBEFD06" box="[151,290,685,707]" italics="true" pageId="24" pageNumber="25">Hoplitomeryx</emphasis>
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<bibRefCitation id="EFEA4B45FD11371CFF48FD18FE5EFD06" author="Leinders, J." box="[296,450,685,707]" firstAuthor="Leinders" journalOrPublisher="Scripta Geologica" pageId="24" pageNumber="25" pagination="1 - 68" part="70" refId="ref24476" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." title="Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy)" type="journal article" year="1984">Leinders, 1984</bibRefCitation>
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||
</taxonomicName>
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||
, size 3—
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||
<bibRefCitation id="EFEA4B45FD11371CFC7EFD18FD43FD06" author="Van der Geer, A. A. E." box="[542,735,685,707]" firstAuthor="Van der Geer" journalOrPublisher="Monografies de la Societat d'Historia Natural de les Balears" pageId="24" pageNumber="25" pagination="325 - 336" part="12" refId="ref26530" refString="Van der Geer, A. A. E. (2005) The postcranial of the deer Hoplitomeryx (Mio-Pliocene; Italy): another example of adaptive radiation on Eastern Mediterranean Islands. Monografies de la Societat d'Historia Natural de les Balears, 12, 325 - 336." title="The postcranial of the deer Hoplitomeryx (Mio-Pliocene; Italy): another example of adaptive radiation on Eastern Mediterranean Islands" type="journal article" year="2005">Van der Geer 2005</bibRefCitation>
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(partim): p. 331, 332;
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<bibRefCitation id="EFEA4B45FD11371CFDA5FD1BFB1AFD06" author="Van der Geer, A. A. E." box="[965,1158,685,707]" firstAuthor="Van der Geer" journalOrPublisher="Quaternary International" pageId="24" pageNumber="25" pagination="145 - 159" part="182" publicationUrl="http://dx.doi.org/10.1016/j.quatint.2007.09.021" refId="ref26581" refString="Van der Geer, A. A. E. (2008) The effect of insularity on the Eastern Mediterranean early cervoid Hoplitomeryx: the study of the forelimb. Quaternary International, 182, 145 - 159. http: // dx. doi. org / 10.1016 / j. quatint. 2007.09.021" title="The effect of insularity on the Eastern Mediterranean early cervoid Hoplitomeryx: the study of the forelimb" type="journal article" year="2008">Van der Geer 2008</bibRefCitation>
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: p. 153, 154, fig. 4, 6.
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</paragraph>
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<paragraph id="8BC436B4FD11371CFEF7FD79FB4AFD27" blockId="24.[151,1436,654,799]" box="[151,1238,716,738]" pageId="24" pageNumber="25">
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||
<taxonomicName id="4C7B4D37FD11371CFEF7FD79FD8DFD27" authority="Leinders, 1984" authorityName="Leinders" authorityYear="1984" box="[151,529,716,738]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="matthei">
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<emphasis id="B90FEAA6FD11371CFEF7FD79FEE8FD27" box="[151,372,716,738]" italics="true" pageId="24" pageNumber="25">Hoplitomeryx matthei</emphasis>
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<bibRefCitation id="EFEA4B45FD11371CFF1BFD79FD8DFD27" author="Leinders, J." box="[379,529,716,738]" firstAuthor="Leinders" journalOrPublisher="Scripta Geologica" pageId="24" pageNumber="25" pagination="1 - 68" part="70" refId="ref24476" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." title="Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy)" type="journal article" year="1984">Leinders, 1984</bibRefCitation>
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||
</taxonomicName>
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||
—
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||
<bibRefCitation id="EFEA4B45FD11371CFC4BFD79FC82FD27" author="Mazza, P. P. A. & Rustioni, M." box="[555,798,716,738]" firstAuthor="Mazza" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="24" pageNumber="25" pagination="1304 - 1333" part="163" refId="ref25321" refString="Mazza, P. P. A. & Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333." title="Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983" type="journal article" year="2011">Mazza & Rustioni 2011</bibRefCitation>
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(partim): p. 1320, fig. 3, 6 [RGM 261.141].
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</paragraph>
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<paragraph id="8BC436B4FD11371CFEF7FD5FFEB5FCDA" blockId="24.[151,1436,654,799]" pageId="24" pageNumber="25">
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<taxonomicName id="4C7B4D37FD11371CFEF7FD5FFDE8FCC5" authority="Mazza et Rustioni, 2011" authorityName="Mazza et Rustioni" authorityYear="2011" box="[151,628,746,768]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="magnus">
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<emphasis id="B90FEAA6FD11371CFEF7FD5FFEEBFCC5" box="[151,375,746,768]" italics="true" pageId="24" pageNumber="25">Hoplitomeryx magnus</emphasis>
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<bibRefCitation id="EFEA4B45FD11371CFF1DFD5EFDE8FCC5" author="Mazza, P. P. A. & Rustioni, M." box="[381,628,746,768]" firstAuthor="Mazza" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="24" pageNumber="25" pagination="1304 - 1333" part="163" refId="ref25321" refString="Mazza, P. P. A. & Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333." title="Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983" type="journal article" year="2011">Mazza et Rustioni, 2011</bibRefCitation>
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</taxonomicName>
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—
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<bibRefCitation id="EFEA4B45FD11371CFCEEFD5FFC1CFCC5" author="Mazza, P. P. A. & Rustioni, M." box="[654,896,746,768]" firstAuthor="Mazza" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="24" pageNumber="25" pagination="1304 - 1333" part="163" refId="ref25321" refString="Mazza, P. P. A. & Rustioni, M. (2011) Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983. Zoological Journal of the Linnean Society, 163, 1304 - 1333." title="Five new species of Hoplitomeryx from the Neogene of Abruzzo and Apulia (central and southern Italy) with revision of the genus and of Hoplitomeryx matthei Leinders, 1983" type="journal article" year="2011">Mazza & Rustioni 2011</bibRefCitation>
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(partim): p. 1324, 1325, fig. 4 [RGM 260.951, RGM 261.135].
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</paragraph>
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</subSubSection>
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<subSubSection id="C361653FFD11371CFEF7FCF9FC87FBA5" pageId="24" pageNumber="25" type="materials_examined">
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<paragraph id="8BC436B4FD11371CFEF7FCF9FDEFFCA0" blockId="24.[151,1437,844,2021]" box="[151,627,844,869]" pageId="24" pageNumber="25">
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<materialsCitation id="3B133CE9FD11371CFEF7FCF9FDF3FCA0" box="[151,623,844,869]" pageId="24" pageNumber="25" typeStatus="holotype">
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<emphasis id="B90FEAA6FD11371CFEF7FCF9FE97FCA0" bold="true" box="[151,267,844,869]" pageId="24" pageNumber="25">
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<typeStatus id="54C08816FD11371CFEF7FCF9FE9AFCA0" box="[151,262,844,869]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Left metacarpal
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<collectionCode id="ED6AAE71FD11371CFFA9FCF9FD95FCA1" box="[457,521,844,868]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
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260.918
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BC436B4FD11371CFEA7FCDAFBEAFBF8" blockId="24.[151,1437,844,2021]" pageId="24" pageNumber="25">
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||
<emphasis id="B90FEAA6FD11371CFEA7FCDAFEDAFC4D" bold="true" box="[199,326,879,904]" pageId="24" pageNumber="25">
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||
<typeStatus id="54C08816FD11371CFEA7FCDAFEDDFC4D" box="[199,321,879,904]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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Proximal humerus
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<collectionCode id="ED6AAE71FD11371CFC45FCC5FDF9FC4D" box="[549,613,880,904]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
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260.950, distal humerus
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<collectionCode id="ED6AAE71FD11371CFDE4FCC5FC58FC4D" box="[900,964,880,904]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
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||
425.278, radius-ulna
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<collectionCode id="ED6AAE71FD11371CFADCFCC5FB60FC4D" box="[1212,1276,880,904]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
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||
425.282 (
|
||
<figureCitation id="13402A31FD11371CFB0FFCDAFF3AFC69" captionStart="FIGURE 4" captionStartId="10.[151,250,1633,1655]" captionTargetBox="[304,1286,188,1606]" captionTargetId="figure@10.[297,1295,177,1620]" captionTargetPageId="10" captionText="FIGURE 4. Postcranial elements of Hoplitomeryx. A, right distal humerus (RGM 178.487; Chiro 20 E) showing conical aspect of the trochlea: anterior view; B, distal left humerus (RGM 425.409; Fina N) without a supracondylar foramen: posterior view; C, distal right humerus (RGM 178.483; Chiro 20 A) with a small foramen: posterior view; D, right radius-ulna (RGM 260.916; San Giovannino) with high degree of synostotic fusion: lateral view; E, right radius-ulna (RGM 425.282; San Giovannino) with no fusion: medial view. Variations observed in the distal (left) radius: F, typical cervid pattern with the radial facet extending further than the intermedial facet; both facets are elongated (after Heintz 1970); G, both facets are tear-shaped and equally developed (after Zeder & Lapham 2010); H, the intermedial facet is much smaller than the radial facet and is almost round (after Heintz 1970). I, left metacarpal (RGM 425.350; Gervasio): anterior (dorsal) view; J, right metacarpal (RGM 425.351; Gervasio): posterior (palmar) view; K, right metacarpal (RGM 425.300; San Giovannino): anterior (dorsal) view; L, right metacarpal (RGM 425.300; San Giovannino): posterior (palmar) view. Schematic drawings of metacarpals showing the variation in morphology of the proximal articulation in proximal view: M, pattern seen in RGM 260.919 (San Giovannino), the typically cervid pattern; N, pattern seen in RGM 425.322 (San Giovannino); O, pattern seen in RGM 262.000 (Gargano, unknown fissure); P, pattern seen in RGM 425.350 (Gervasio)." figureDoi="http://doi.org/10.5281/zenodo.286819" httpUri="https://zenodo.org/record/286819/files/figure.png" pageId="24" pageNumber="25">Fig. 4</figureCitation>
|
||
E), radius
|
||
<collectionCode id="ED6AAE71FD11371CFF49FC21FEF5FC69" box="[297,361,916,940]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
260.866, unciform
|
||
<collectionCode id="ED6AAE71FD11371CFC3EFC21FD02FC69" box="[606,670,916,940]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
425.264, anterior first phalanx
|
||
<collectionCode id="ED6AAE71FD11371CFA48FC21FBF4FC69" box="[1064,1128,916,940]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
260.861, anterior second phalanges
|
||
<collectionCode id="ED6AAE71FD11371CFF6FFC0DFED3FC15" box="[271,335,952,976]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
260.886 and
|
||
<collectionCode id="ED6AAE71FD11371CFF8BFC0DFDB7FC15" box="[491,555,952,976]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
261.143, femur
|
||
<collectionCode id="ED6AAE71FD11371CFC88FC0DFCB4FC15" box="[744,808,952,976]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
425.245 (
|
||
<figureCitation id="13402A31FD11371CFDFBFC02FC43FC15" box="[923,991,951,976]" captionStart="FIGURE 5" captionStartId="11.[151,250,1725,1747]" captionTargetBox="[290,1308,202,1695]" captionTargetId="figure@11.[276,1315,193,1705]" captionTargetPageId="11" captionText="FIGURE 5. Postcranial elements of Hoplitomeryx. A, first posterior phalanx (RGM 425.269; San Giovannino) showing a deerlike morphology: abaxial view; B, same specimen, showing triangular sesamoid facets but no ligament attachment areas: volar view; C, first anterior phalanx (RGM 178.503; San Giovannino): abaxial view; D, same specimen, showing pronounced ligament attachments in the form of ridges but no sesamoid facet: volar view; E, posterior second phalanx (RGM 261.251; San Giovannino): abaxial view; F, anterior second phalanx (RGM 260.937; San Giovannino): axial view; G, third phalanx (RGM 178.683; Trefossi 2 A): abaxial view; H, third phalanx (RGM 260.877; San Giovannino): axial view; I, third phalanx (RGM 261.531; Nazario 4): abaxial view; J, third phalanx (RGM 425.386; Falcone 2 B); K, right femur (RGM 425.314; San Giovannino): anterior view; L, left femur (RGM 425.207; Nazario 4): anterior view; M, right femur (RGM 425.245; San Giovannino): anterior view; N, left patella (RGM 425.246; San Giovannino): anterior (dorsal) view; O, same specimen: lateral view; P, same specimen, showing the hook-like extension at the medial side: inner view." figureDoi="http://doi.org/10.5281/zenodo.286820" httpUri="https://zenodo.org/record/286820/files/figure.png" pageId="24" pageNumber="25">Fig. 5</figureCitation>
|
||
M), patella
|
||
<collectionCode id="ED6AAE71FD11371CFA03FC0DFB3FFC15" box="[1123,1187,952,976]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
425.246 (
|
||
<figureCitation id="13402A31FD11371CFB77FC02FAC5FC15" box="[1303,1369,951,976]" captionStart="FIGURE 5" captionStartId="11.[151,250,1725,1747]" captionTargetBox="[290,1308,202,1695]" captionTargetId="figure@11.[276,1315,193,1705]" captionTargetPageId="11" captionText="FIGURE 5. Postcranial elements of Hoplitomeryx. A, first posterior phalanx (RGM 425.269; San Giovannino) showing a deerlike morphology: abaxial view; B, same specimen, showing triangular sesamoid facets but no ligament attachment areas: volar view; C, first anterior phalanx (RGM 178.503; San Giovannino): abaxial view; D, same specimen, showing pronounced ligament attachments in the form of ridges but no sesamoid facet: volar view; E, posterior second phalanx (RGM 261.251; San Giovannino): abaxial view; F, anterior second phalanx (RGM 260.937; San Giovannino): axial view; G, third phalanx (RGM 178.683; Trefossi 2 A): abaxial view; H, third phalanx (RGM 260.877; San Giovannino): axial view; I, third phalanx (RGM 261.531; Nazario 4): abaxial view; J, third phalanx (RGM 425.386; Falcone 2 B); K, right femur (RGM 425.314; San Giovannino): anterior view; L, left femur (RGM 425.207; Nazario 4): anterior view; M, right femur (RGM 425.245; San Giovannino): anterior view; N, left patella (RGM 425.246; San Giovannino): anterior (dorsal) view; O, same specimen: lateral view; P, same specimen, showing the hook-like extension at the medial side: inner view." figureDoi="http://doi.org/10.5281/zenodo.286820" httpUri="https://zenodo.org/record/286820/files/figure.png" pageId="24" pageNumber="25">Fig. 5</figureCitation>
|
||
N–P), tibia
|
||
<collectionCode id="ED6AAE71FD11371CFEB4FC69FE88FC31" box="[212,276,988,1012]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
260.854, astragals
|
||
<collectionCode id="ED6AAE71FD11371CFF9AFC69FDA6FC31" box="[506,570,988,1012]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
260.863 (
|
||
<figureCitation id="13402A31FD11371CFCD4FC69FD67FC30" box="[692,763,988,1013]" captionStart="FIGURE 6" captionStartId="12.[151,250,1824,1846]" captionTargetBox="[205,1378,205,1796]" captionTargetId="figure@12.[196,1392,193,1803]" captionTargetPageId="12" captionText="FIGURE 6. Postcranial elements of Hoplitomeryx. A, left tibia (RGM 335.882 + 425.328; San Giovannino): medial view; B, right tibia (RGM 425.144; Chiro D 1): anterior view. Astragals of different size in anterior (dorsal) view: C, left astragal (RGM 425.385; Falcone 2 B); D, left astragal (RGM 260.863; San Giovannino); E, right astragal (RGM 425.421; Fina N). F, left astragal (RGM 263.951; Gargano, unknown fissure): lateral view; G, left calcaneum (RGM 260.901; San Giovannino): anterior view; H, same specimen: posterior view; I, same specimen: medial view; J,; K, detail of distal right metatarsal (RGM 178.518; San Giovannino) showing the proximal extension of the dorsal gully into the (fused) cubonavicular." figureDoi="http://doi.org/10.5281/zenodo.286821" httpUri="https://zenodo.org/record/286821/files/figure.png" pageId="24" pageNumber="25">Fig. 6</figureCitation>
|
||
D) and
|
||
<collectionCode id="ED6AAE71FD11371CFD37FC69FC0BFC31" box="[855,919,988,1012]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
260.890, calcanei
|
||
<collectionCode id="ED6AAE71FD11371CFA17FC69FB2BFC31" box="[1143,1207,988,1012]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
260.883 and
|
||
<collectionCode id="ED6AAE71FD11371CFB3DFC69FA01FC31" box="[1373,1437,988,1012]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
425.313, first phalanges
|
||
<collectionCode id="ED6AAE71FD11371CFFD3FBB5FE6FFBDD" box="[435,499,1024,1048]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
260.913 and
|
||
<collectionCode id="ED6AAE71FD11371CFCF4FBB5FD48FBDD" box="[660,724,1024,1048]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
260.909, seventh cervical vertebra
|
||
<collectionCode id="ED6AAE71FD11371CFA10FBB5FB2CFBDD" box="[1136,1200,1024,1048]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
425.307, nasal horn core
|
||
<collectionCode id="ED6AAE71FD11371CFEAEFB91FE92FBF9" box="[206,270,1060,1084]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
260.902, associated left and right hemimandibles and maxilla
|
||
<collectionCode id="ED6AAE71FD11371CFDACFB91FB90FBF9" box="[972,1036,1060,1084]" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="24" pageNumber="25">RGM</collectionCode>
|
||
260.951.
|
||
</paragraph>
|
||
<paragraph id="8BC436B4FD11371CFEA7FBF2FC87FBA5" blockId="24.[151,1437,844,2021]" box="[199,795,1095,1120]" pageId="24" pageNumber="25">
|
||
<emphasis id="B90FEAA6FD11371CFEA7FBF2FE25FBA5" bold="true" box="[199,441,1095,1120]" pageId="24" pageNumber="25">Referred specimens.</emphasis>
|
||
See Appendix III, and
|
||
<figureCitation id="13402A31FD11371CFCDEFBF2FC9CFBA5" box="[702,768,1095,1120]" captionStart="FIGURE 4" captionStartId="10.[151,250,1633,1655]" captionTargetBox="[304,1286,188,1606]" captionTargetId="figure@10.[297,1295,177,1620]" captionTargetPageId="10" captionText="FIGURE 4. Postcranial elements of Hoplitomeryx. A, right distal humerus (RGM 178.487; Chiro 20 E) showing conical aspect of the trochlea: anterior view; B, distal left humerus (RGM 425.409; Fina N) without a supracondylar foramen: posterior view; C, distal right humerus (RGM 178.483; Chiro 20 A) with a small foramen: posterior view; D, right radius-ulna (RGM 260.916; San Giovannino) with high degree of synostotic fusion: lateral view; E, right radius-ulna (RGM 425.282; San Giovannino) with no fusion: medial view. Variations observed in the distal (left) radius: F, typical cervid pattern with the radial facet extending further than the intermedial facet; both facets are elongated (after Heintz 1970); G, both facets are tear-shaped and equally developed (after Zeder & Lapham 2010); H, the intermedial facet is much smaller than the radial facet and is almost round (after Heintz 1970). I, left metacarpal (RGM 425.350; Gervasio): anterior (dorsal) view; J, right metacarpal (RGM 425.351; Gervasio): posterior (palmar) view; K, right metacarpal (RGM 425.300; San Giovannino): anterior (dorsal) view; L, right metacarpal (RGM 425.300; San Giovannino): posterior (palmar) view. Schematic drawings of metacarpals showing the variation in morphology of the proximal articulation in proximal view: M, pattern seen in RGM 260.919 (San Giovannino), the typically cervid pattern; N, pattern seen in RGM 425.322 (San Giovannino); O, pattern seen in RGM 262.000 (Gargano, unknown fissure); P, pattern seen in RGM 425.350 (Gervasio)." figureDoi="http://doi.org/10.5281/zenodo.286819" httpUri="https://zenodo.org/record/286819/files/figure.png" pageId="24" pageNumber="25">Fig. 4</figureCitation>
|
||
A.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361653FFD11371CFEA7FBD9FDD5FA0D" pageId="24" pageNumber="25" type="diagnosis">
|
||
<paragraph id="8BC436B4FD11371CFEA7FBD9FF46FB08" blockId="24.[151,1437,844,2021]" pageId="24" pageNumber="25">
|
||
<emphasis id="B90FEAA6FD11371CFEA7FBD9FEDDFB40" bold="true" box="[199,321,1132,1157]" pageId="24" pageNumber="25">Diagnosis.</emphasis>
|
||
A large-sized hoplitomerycid. Estimated body mass 78.0 kg. The fusion between radius and ulna is weak and the ulna leaves no trace on the radius. The shaft of the tibia is almost straight. The astragal is non-parallel sided.
|
||
</paragraph>
|
||
<paragraph id="8BC436B4FD11371CFEA7FB62FB82FAD1" blockId="24.[151,1437,844,2021]" pageId="24" pageNumber="25">
|
||
<emphasis id="B90FEAA6FD11371CFEA7FB62FE53FB35" bold="true" box="[199,463,1239,1264]" pageId="24" pageNumber="25">Differential diagnosis.</emphasis>
|
||
About twice the body size of
|
||
<taxonomicName id="4C7B4D37FD11371CFD48FB6CFC3CFB35" box="[808,928,1240,1264]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="matthei">
|
||
<emphasis id="B90FEAA6FD11371CFD48FB6CFC3CFB35" box="[808,928,1240,1264]" italics="true" pageId="24" pageNumber="25">H. matthei</emphasis>
|
||
</taxonomicName>
|
||
. The lateral horn core is compressed anteroposteriorly unlike the circular cross section seen in
|
||
<taxonomicName id="4C7B4D37FD11371CFCB6FB48FCD0FAD1" box="[726,844,1276,1300]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="matthei">
|
||
<emphasis id="B90FEAA6FD11371CFCB6FB48FCD0FAD1" box="[726,844,1276,1300]" italics="true" pageId="24" pageNumber="25">H. matthei</emphasis>
|
||
</taxonomicName>
|
||
(horn core
|
||
<typeStatus id="54C08816FD11371CFDAEFB48FC62FAD0" box="[974,1022,1277,1301]" pageId="24" pageNumber="25">type</typeStatus>
|
||
I).
|
||
</paragraph>
|
||
<paragraph id="8BC436B4FD11371CFEA7FAAAFEC3FA45" blockId="24.[151,1437,844,2021]" pageId="24" pageNumber="25">
|
||
<emphasis id="B90FEAA6FD11371CFEA7FAAAFE20FAF2" bold="true" box="[199,444,1311,1336]" pageId="24" pageNumber="25">Derivation of name.</emphasis>
|
||
Named after Ross D.E. MacPhee, curator of mammals at the American Museum of Natural History in New York, in honour of his contributions to the knowledge of the evolution and extinction of insular mammals.
|
||
</paragraph>
|
||
<paragraph id="8BC436B4FD11371CFEA7FA39FDD5FA0D" blockId="24.[151,1437,844,2021]" pageId="24" pageNumber="25">
|
||
<emphasis id="B90FEAA6FD11371CFEA7FA39FD81FA60" bold="true" box="[199,541,1420,1445]" pageId="24" pageNumber="25">Preservation and deposition.</emphasis>
|
||
Naturalis Biodiversity Center, Leiden,
|
||
<collectingCountry id="F36C7624FD11371CFDB7FA39FB14FA60" box="[983,1160,1420,1445]" name="Netherlands" pageId="24" pageNumber="25">the Netherlands</collectingCountry>
|
||
(formerly Rijksmuseum van Geologie en Mineralogie (RGM)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361653FFD11371CFEA7FA61FD4DF965" pageId="24" pageNumber="25" type="materials_examined">
|
||
<paragraph id="8BC436B4FD11371CFEA7FA61FC71F9F0" blockId="24.[151,1437,844,2021]" pageId="24" pageNumber="25">
|
||
<emphasis id="B90FEAA6FD11371CFEA7FA61FD9EFA28" bold="true" box="[199,514,1492,1517]" pageId="24" pageNumber="25">
|
||
<typeStatus id="54C08816FD11371CFEA7FA61FE9EFA28" box="[199,258,1492,1517]" pageId="24" pageNumber="25">Type</typeStatus>
|
||
locality and horizon.
|
||
</emphasis>
|
||
Late Miocene (Middle or Late Turolian, MN12-13) fissure filling with code San Giovannino in an abandoned limestone quarry near the farm of San Giovannino south of the provincial road between Poggio Imperiale and Apricena (Province of Foggia, Apulia,
|
||
<collectingCountry id="F36C7624FD11371CFDC9F9A9FC7DF9F0" box="[937,993,1564,1589]" name="Italy" pageId="24" pageNumber="25">Italy</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC436B4FD11371CFEA7F98AFD4DF965" blockId="24.[151,1437,844,2021]" pageId="24" pageNumber="25">
|
||
<emphasis id="B90FEAA6FD11371CFEA7F98AFE05F99D" bold="true" box="[199,409,1599,1624]" pageId="24" pageNumber="25">Studied localities.</emphasis>
|
||
Fissure fillings with codes Biancone 2, Chirò 2, 3, 5A, 13, 20A and E, 28 and D3, Fina H and N, Gervasio, Pizzicoli 1, 4 and 12, San Giovannino. All localities are located in the north-western portion of the Gargano Peninsula, Apulia, south-eastern
|
||
<collectingCountry id="F36C7624FD11371CFCF8F932FD51F965" box="[664,717,1671,1696]" name="Italy" pageId="24" pageNumber="25">Italy</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361653FFD11371DFEA7F919FCC2FF3D" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
|
||
<paragraph id="8BC436B4FD11371CFEA7F919FC7AF858" blockId="24.[151,1437,844,2021]" pageId="24" pageNumber="25">
|
||
<emphasis id="B90FEAA6FD11371CFEA7F919FE7AF900" bold="true" box="[199,486,1708,1733]" pageId="24" pageNumber="25">
|
||
Description of
|
||
<typeStatus id="54C08816FD11371CFF18F919FE7DF900" box="[376,481,1708,1733]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
RGM 260.918 is an almost complete metacarpal of which only the distal trochlea is missing distalward of the (fused) epiphyseal line. Its general morphology corresponds to that of
|
||
<taxonomicName id="4C7B4D37FD11371CFABEF965FAB1F922" box="[1246,1325,1744,1767]" class="Mammalia" family="Cervidae" genus="Cervus" kingdom="Animalia" order="Artiodactyla" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90FEAA6FD11371CFABEF965FAB1F922" box="[1246,1325,1744,1767]" italics="true" pageId="24" pageNumber="25">Cervus</emphasis>
|
||
</taxonomicName>
|
||
but more slender. The crest that separates the medial and lateral articulation is broken but seems to have ended in the central fossa, making an angle of about twenty degrees with the parasagittal plane. The central fossa is large, situated more or less centrally and makes only minimal contact with the palmar surface. The attachment area for the carpal ligaments on the dorsal surface is weakly developed. The palmar groove for the M. interosseus palmaris is only proximally expressed. The scars for the lateral metacarpals are pronounced.
|
||
</paragraph>
|
||
<paragraph id="8BC436B4FD11371DFEA7F81DFCC2FF3D" blockId="24.[151,1437,844,2021]" lastBlockId="25.[151,1436,151,608]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
|
||
<emphasis id="B90FEAA6FD11371CFEA7F81DFEE1F804" bold="true" box="[199,381,1960,1985]" pageId="24" pageNumber="25">Measurements.</emphasis>
|
||
<typeStatus id="54C08816FD11371CFFE6F81DFE6FF804" box="[390,499,1960,1985]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
|
||
: maximal length =
|
||
<quantity id="4C839B51FD11371CFCB4F81DFCAAF805" box="[724,822,1960,1984]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" pageId="24" pageNumber="25" unit="mm" value="190.0">190 mm</quantity>
|
||
, proximal depth (DAPP) =
|
||
<quantity id="4C839B51FD11371CFA17F81DFB7CF805" box="[1143,1248,1960,1985]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="24" pageNumber="25" unit="mm" value="17.5">17.5 mm</quantity>
|
||
, proximal width (DTP) =
|
||
<quantity id="4C839B51FD11371CFE9EF879FEF9F821" box="[254,357,1996,2020]" metricMagnitude="-2" metricUnit="m" metricValue="3.01" pageId="24" pageNumber="25" unit="mm" value="30.1">30.1 mm</quantity>
|
||
, distal depth (DAPD; estimated at epiphyseal line) =
|
||
<quantity id="4C839B51FD11371CFDA3F879FBB6F821" box="[963,1066,1996,2020]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="24" pageNumber="25" unit="mm" value="13.1">13.1 mm</quantity>
|
||
, distal width (DTD; estimated at epiphyseal line) =
|
||
<quantity id="4C839B51FD10371DFF08FF2DFE26FF6A" box="[360,442,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="25" pageNumber="26" unit="mm" value="27.0">27 mm</quantity>
|
||
. The distal diameters are confirmed by referred distal metacarpal RGM 261.530: DAPD =
|
||
<quantity id="4C839B51FD10371DFED7FF09FEBAFF11" box="[183,294,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="25" pageNumber="26" unit="mm" value="14.7">14.7 mm</quantity>
|
||
, DTD =
|
||
<quantity id="4C839B51FD10371DFFC4FF09FD88FF11" box="[420,532,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="2.82" pageId="25" pageNumber="26" unit="mm" value="28.2">28.2 mm</quantity>
|
||
. For other measurements of referred specimens, see Appendix III (linear measurements) and Appendix I and II (body mass estimations).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C361653FFD10371DFEA7FEB1FF66FDA5" pageId="25" pageNumber="26" type="discussion">
|
||
<paragraph id="8BC436B4FD10371DFEA7FEB1FF66FDA5" blockId="25.[151,1436,151,608]" pageId="25" pageNumber="26">
|
||
<emphasis id="B90FEAA6FD10371DFEA7FEB1FEA7FED8" bold="true" box="[199,315,260,285]" pageId="25" pageNumber="26">Remarks.</emphasis>
|
||
Skull fragment RGM 261.099 (horn core
|
||
<typeStatus id="54C08816FD10371DFD54FEB0FCF8FED8" box="[820,868,261,285]" pageId="25" pageNumber="26">type</typeStatus>
|
||
IV in
|
||
<bibRefCitation id="EFEA4B45FD10371DFDDBFEB1FBFFFED9" author="Leinders, J." box="[955,1123,260,285]" firstAuthor="Leinders" journalOrPublisher="Scripta Geologica" pageId="25" pageNumber="26" pagination="1 - 68" part="70" refId="ref24476" refString="Leinders, J. (1984) Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy). Scripta Geologica, 70, 1 - 68." title="Hoplitomerycidae fam. nov. (Ruminantia, Mammalia) from Neogene fissure fillings in Gargano (Italy)" type="journal article" year="1984">Leinders 1984</bibRefCitation>
|
||
; Pizzicoli 5) represents an animal larger than the
|
||
<typeStatus id="54C08816FD10371DFFF8FE9DFE54FE85" box="[408,456,296,320]" pageId="25" pageNumber="26">type</typeStatus>
|
||
species (
|
||
<taxonomicName id="4C7B4D37FD10371DFC53FE9CFD36FE85" box="[563,682,296,320]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="matthei">
|
||
<emphasis id="B90FEAA6FD10371DFC53FE9CFD36FE85" box="[563,682,296,320]" italics="true" pageId="25" pageNumber="26">H. matthei</emphasis>
|
||
</taxonomicName>
|
||
) but seems not large enough for the largest size group and is here attributed to
|
||
<taxonomicName id="4C7B4D37FD10371DFF54FEF8FE56FEA1" authority="Van Der Geer, 2014" authorityName="Van Der Geer" authorityYear="2014" box="[308,458,332,356]" class="Mammalia" family="Hoplitomerycidae" genus="Hoplitomeryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="macpheei" status="sp. nov.">
|
||
<emphasis id="B90FEAA6FD10371DFF54FEF8FE56FEA1" box="[308,458,332,356]" italics="true" pageId="25" pageNumber="26">H. macpheei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90FEAA6FD10371DFFB8FEF9FD82FEA0" bold="true" box="[472,542,332,357]" pageId="25" pageNumber="26">
|
||
<taxonomicNameLabel id="A23C57DDFD10371DFFB8FEF9FD82FEA0" box="[472,542,332,357]" pageId="25" pageNumber="26" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
. RGM 261.099 might further be an indication that the configuration and morphology of the horn cores may differ between the species in addition to between the sexes. The lack of associated posterior skull parts and / or maxillae with horn cores hampers a more precise diagnosis based on horn core
|
||
<typeStatus id="54C08816FD10371DFEB2FE0DFE8DFE15" box="[210,273,440,464]" pageId="25" pageNumber="26">types</typeStatus>
|
||
. The
|
||
<typeStatus id="54C08816FD10371DFF36FE02FE24FE15" box="[342,440,439,464]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
is more slender than the referred proximal metacarpal RGM 425.322 from the same fissure filling. They likely represent a female and a male individual respectively. The morphology of the proximal articulation of both specimens was earlier described as morphotype 1 (
|
||
<bibRefCitation id="EFEA4B45FD10371DFDC7FDB5FBE3FDDD" author="Van der Geer, A. A. E." box="[935,1151,511,536]" firstAuthor="Van der Geer" journalOrPublisher="Monografies de la Societat d'Historia Natural de les Balears" pageId="25" pageNumber="26" pagination="325 - 336" part="12" refId="ref26530" refString="Van der Geer, A. A. E. (2005) The postcranial of the deer Hoplitomeryx (Mio-Pliocene; Italy): another example of adaptive radiation on Eastern Mediterranean Islands. Monografies de la Societat d'Historia Natural de les Balears, 12, 325 - 336." title="The postcranial of the deer Hoplitomeryx (Mio-Pliocene; Italy): another example of adaptive radiation on Eastern Mediterranean Islands" type="journal article" year="2005">Van der Geer 2005</bibRefCitation>
|
||
), but RGM 425.322 has a more pronounced crest than the
|
||
<typeStatus id="54C08816FD10371DFC76FD91FDE0FDF8" box="[534,636,548,573]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
. Patella RGM 425.246 fits perfectly well on the distal femur RGM 425.245.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |