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<mods:title id="877ABB570829B5CDEA81068C3B72D8AE">Antaganides n. gen., a new genus of Middle Jurassic nautiloid, ancestor of Pseudaganides Spath, 1927, with discussion on the phyletic origin of the family Pseudonautilidae</mods:title>
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<mods:namePart id="AC62C52D9A2B738FDB2F6D696DC80BB7">Branger, Patrick</mods:namePart>
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<heading id="D0D4592DFF99FFF90539A5F7FD88FB76" box="[269,638,1173,1200]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="480" reason="2">
<taxonomicName id="4C2395C2FF99FFF90539A5F7FE12FB69" authorityName="Branger" authorityYear="2024" box="[269,484,1174,1200]" class="Cephalopoda" family="Pseudonautilidae" genus="Antaganides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="11" pageNumber="480" phylum="Mollusca" rank="species" species="grulkei" status="n. gen., n. sp.">
<emphasis id="B9573253FF99FFF90539A5F7FE12FB69" bold="true" box="[269,484,1174,1200]" italics="true" pageId="11" pageNumber="480">Antaganides grulkei</emphasis>
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<taxonomicNameLabel id="A2648F28FF99FFF905DFA5F4FD88FB76" box="[491,638,1173,1199]" pageId="11" pageNumber="480" rank="species">n. gen., n. sp.</taxonomicNameLabel>
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<paragraph id="8B9CEE41FF99FFF9054AA5D5FDFBFB17" blockId="11.[269,638,1173,1230]" box="[382,525,1204,1230]" pageId="11" pageNumber="480">(Fig. 11A, B)</paragraph>
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urn:lsid:zoobank.org:act:
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<taxonomicName id="4C2395C2FF99FFF904B0A459FEC6FA89" baseAuthorityName="Crick" baseAuthorityYear="1898" box="[132,304,1336,1360]" class="Cephalopoda" family="Nautilidae" genus="Nautilus" isUncertain="true" kingdom="Animalia" order="Nautilida" pageId="11" pageNumber="480" phylum="Mollusca" rank="species" species="fuscus">
<emphasis id="B9573253FF99FFF904B0A459FF23FA89" box="[132,213,1336,1360]" italics="true" pageId="11" pageNumber="480">Nautilus</emphasis>
cf.
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<treatmentCitation id="0A82C850FF99FFF9057DA459FDF7FA89" author="LISSAJOUS M." box="[329,513,1336,1360]" page="48" pageId="11" pageNumber="480" year="1923">
<bibRefCitation id="EFB293B0FF99FFF9057DA459FDF7FA89" author="LISSAJOUS M." box="[329,513,1336,1360]" pageId="11" pageNumber="480" pagination="1 - 286" refId="ref15475" refString="LISSAJOUS M. 1923. - Etude sur la faune du Bathonien des environs de Macon. Travaux du Laboratoire de Geologie de la Faculte des Sciences de Lyon 3: 1 - 286, 33 pls." type="journal article" year="1923">Lissajous 1923: 48</bibRefCitation>
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, pl. II, fig. 2-2a.
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</paragraph>
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. — Small sized nautilus, maximum diameter
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, shell rather compressed on the last whorl with an oval section except on the body-chamber where the venter flattens. Umbilicus closed. Suture lines with a wide and shallow lateral lobe.
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ETYMOLOGY.
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<taxonomicName id="4C2395C2FF99FFF9051FA486FE12FA26" authorityName="Branger" authorityYear="2024" box="[299,484,1511,1535]" class="Cephalopoda" family="Pseudonautilidae" genus="Antaganides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="11" pageNumber="480" phylum="Mollusca" rank="species" species="grulkei" status="n. gen., n. sp.">Antaganides grulkei</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A2648F28FF99FFF905D8A486FD85FA27" box="[492,627,1511,1535]" pageId="11" pageNumber="480" rank="species">n. gen., n. sp.</taxonomicNameLabel>
is dedicated to Wolfgang Grulke, Sherborne,
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, who made a worldwide promotion for nautiloids studies through his magnificent book “
<emphasis id="B9573253FF99FFF904BBA757FE4CF997" box="[143,442,1590,1614]" italics="true" pageId="11" pageNumber="480">
<taxonomicName id="4C2395C2FF99FFF904BBA757FF15F997" authorityName="Linnaeus" authorityYear="1758" box="[143,227,1590,1614]" class="Cephalopoda" family="Nautilidae" genus="Nautilus" kingdom="Animalia" order="Nautilida" pageId="11" pageNumber="480" phylum="Mollusca" rank="genus">Nautilus</taxonomicName>
, the Beautiful Survivor
</emphasis>
” (
<bibRefCitation id="EFB293B0FF99FFF905E6A757FDB9F997" author="GRULKE W." box="[466,591,1590,1614]" pageId="11" pageNumber="480" refId="ref15094" refString="GRULKE W. 2016. - Nautilus, Beautiful Survivor. One Edition, Sherborne, 224 p." type="book" year="2016">Grulke 2016</bibRefCitation>
). Doing this work, he gave us the opportunity to meet or to discuss with most of the specialists of the world.
</paragraph>
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. —
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.
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<collectingCountry id="F334AED1FF99FFF90589A7FAFDF7F96A" box="[445,513,1691,1715]" name="France" pageId="11" pageNumber="480">France</collectingCountry>
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(complete);
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,
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,
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(
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); [
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,
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]; Base of the
<emphasis id="B9573253FF99FFF905A5A78BFE0EF8DB" box="[401,504,1770,1794]" italics="true" pageId="11" pageNumber="480">Progracilis</emphasis>
Zone,
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Subzone; Middle to Upper Bathonian;
<collectorName id="26D68B97FF99FFF90592A664FDB7F8C4" box="[422,577,1797,1821]" pageId="11" pageNumber="480">Patrick Branger</collectorName>
leg.;
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;
<emphasis id="B9573253FF99FFF906FFA664FF23F8EE" italics="true" pageId="11" pageNumber="480">2023. PB.N97</emphasis>
(Fig. 11A, B).
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</paragraph>
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. —
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(complete); same data as holotype;
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(complete, body-chambers poorly preserved); Nouvelle-Aquitaine, La Crèche, Champs-Albert; [
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,
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]; base of the
<emphasis id="B9573253FF99FFF906F9A6FEFF2FF808" italics="true" pageId="11" pageNumber="480">Retrocostatum</emphasis>
Zone;
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; Patrick Branger leg.;
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;
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.
</materialsCitation>
</paragraph>
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. —
<taxonomicName id="4C2395C2FF99FFF9008EA320FA91FD80" authorityName="Branger" authorityYear="2024" box="[1210,1383,577,601]" class="Cephalopoda" family="Pseudonautilidae" genus="Antaganides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="11" pageNumber="480" phylum="Mollusca" rank="species" species="grulkei" status="n. gen., n. sp.">
<emphasis id="B9573253FF99FFF9008EA320FA91FD80" box="[1210,1383,577,601]" italics="true" pageId="11" pageNumber="480">Antaganides grulkei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2648F28FF99FFF9015DA320FC95FDAA" pageId="11" pageNumber="480" rank="species">n. gen., n. sp.</taxonomicNameLabel>
has been mostly encountered within the Middle Bathonian of the Submediterranean Realm where it appears at the base of the
<emphasis id="B9573253FF99FFF90718A3F1FC7AFD71" box="[812,908,656,680]" italics="true" pageId="11" pageNumber="480">Progracilis</emphasis>
Zone,
<emphasis id="B9573253FF99FFF907E2A3F1FBDDFD71" box="[982,1067,656,680]" italics="true" pageId="11" pageNumber="480">Orbignyi</emphasis>
Subzone. The
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has been found there, in les Hauts-de-Rochefort quarry. It persists until the Upper Bathonian, at the base of the
<emphasis id="B9573253FF99FFF9000FA3A7FB4AFD04" box="[1083,1212,710,733]" italics="true" pageId="11" pageNumber="480">Retrocostatum</emphasis>
Zone where the youngest specimens have been found. In
<collectingRegion id="49E720A3FF99FFF90051A381FB3EFD21" box="[1125,1224,736,760]" country="France" name="Bourgogne" pageId="11" pageNumber="480">Burgundy</collectingRegion>
, the nautilus figured as
<taxonomicName id="4C2395C2FF99FFF9071AA39BFBA1FCCB" authority="(Crick, 1898)" baseAuthorityName="Crick" baseAuthorityYear="1898" box="[814,1111,762,786]" class="Cephalopoda" family="Nautilidae" genus="Nautilus" isUncertain="true" kingdom="Animalia" order="Nautilida" pageId="11" pageNumber="480" phylum="Mollusca" rank="species" species="fuscus">
<emphasis id="B9573253FF99FFF9071AA39BFC88FCCB" box="[814,894,762,786]" italics="true" pageId="11" pageNumber="480">Nautilus</emphasis>
cf.
<emphasis id="B9573253FF99FFF907A8A39BFC27FCCB" box="[924,977,762,786]" italics="true" pageId="11" pageNumber="480">fuscus</emphasis>
(
<bibRefCitation id="EFB293B0FF99FFF907E8A39BFBA6FCCB" author="CRICK G. C." box="[988,1104,762,786]" pageId="11" pageNumber="480" pagination="117 - 139" refId="ref14766" refString="CRICK G. C. 1898. - Description of new or imperfectly known species of Nautilus from the Inferior Oolite, preserved in the British Museum (Natural History). Proceedings of the Malacological Society of London 3: 117 - 139. https: // archive. org / details / biostor- 193383" type="journal article" year="1898">Crick, 1898</bibRefCitation>
)
</taxonomicName>
byLissajous (1923) also comes from the upper part of the Middle Bathonian, “
<emphasis id="B9573253FF99FFF90088A275FACDFCF5" box="[1212,1339,788,812]" italics="true" pageId="11" pageNumber="480">Arbustigerum</emphasis>
Zone” (that corresponds to
<emphasis id="B9573253FF99FFF907F5A251FBF1FC9E" box="[961,1031,816,839]" italics="true" pageId="11" pageNumber="480">Morrisi</emphasis>
and
<emphasis id="B9573253FF99FFF9000CA251FB74FC9E" box="[1080,1154,816,839]" italics="true" pageId="11" pageNumber="480">Bremeri</emphasis>
Zones [
<bibRefCitation id="EFB293B0FF99FFF900E7A24EFA6FFC9E" author="MANGOLD C. &amp; MARTIN A. &amp; PRIEUR A." box="[1235,1433,815,839]" pageId="11" pageNumber="480" pagination="1 - 155" refId="ref15590" refString="MANGOLD C., MARTIN A. &amp; PRIEUR A. 2012. - Les Perisphinctidae du Bathonien moyen et superieur du Maconnais (Saone-et- Loire, France). Travaux et Documents des Laboratoires de Geologie de Lyon 169: 1 - 155." type="journal article" year="2012">
Mangold
<emphasis id="B9573253FF99FFF90106A251FA97FC9E" box="[1330,1377,815,839]" italics="true" pageId="11" pageNumber="480">et al.</emphasis>
2012
</bibRefCitation>
]).
</paragraph>
</subSubSection>
<subSubSection id="C339BDCAFF99FFF90719A201FBF4FAF6" pageId="11" pageNumber="480" type="description">
<paragraph id="8B9CEE41FF99FFF90719A201FB9CFCAE" blockId="11.[813,1130,863,887]" box="[813,1130,863,887]" pageId="11" pageNumber="480">
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.
<emphasis id="B9573253FF99FFF907E3A201FC06FCAE" box="[983,1008,864,887]" italics="true" pageId="11" pageNumber="480"></emphasis>
See
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</paragraph>
<paragraph id="8B9CEE41FF99FFF90719A2F7FC4BFC77" blockId="11.[811,1456,918,1328]" box="[813,957,918,944]" pageId="11" pageNumber="480">
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</paragraph>
<paragraph id="8B9CEE41FF99FFF90719A2D7FBF4FAF6" blockId="11.[811,1456,918,1328]" pageId="11" pageNumber="480">
The
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(
<emphasis id="B9573253FF99FFF907FDA2D7FBA9FC09" box="[969,1119,950,976]" italics="true" pageId="11" pageNumber="480">2023.PB.N97</emphasis>
; Fig. 11A, B) and the specimen
<emphasis id="B9573253FF99FFF9071FA2B7FC39FC29" box="[811,975,982,1008]" italics="true" pageId="11" pageNumber="480">2023.PB.N133</emphasis>
are rather well preserved samples. They only reach a small size, a little more than
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, at the adult stage.
<emphasis id="B9573253FF99FFF90718A577FC38FBE9" box="[812,974,1046,1072]" italics="true" pageId="11" pageNumber="480">2023.PB.N134</emphasis>
is a young one with a diameter of
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, The shell is always compressed with dorsal sides slightly rounded showing a maximum width at about two third height. Then, the lateral-ventral sides flatten and converge. The ventral part, rounded on the inner whorls, flattens on the body-chamber, boarded by angular but not sharp shoulders. The umbilicus is very small, funnel-shaped. The lateral lobe of the suture line (
<figureCitation id="1318F2C4FF99FFF90702A594FC7BFAC9" box="[822,909,1269,1296]" captionStart="FIG" captionStartId="7.[132,143,1019,1036]" captionTargetBox="[226,1361,215,976]" captionTargetId="figure-370@7.[227,1361,215,975]" captionTargetPageId="7" captionText="FIG. 4. — Suture lines of Antaganides n. gen. and Pseudaganides Spath, 1927: A, Antaganides cf. erycinus (Tagliarini,1901), n. comb., 2023.PB.N117; B, Antaganides sauvageti n. gen., n. sp. 2023.PB.N64; C, Antaganides fournieri n. gen., n. sp., 2023.PB.N105; D, Pseudaganides aquitanense n. sp., 2023.PB.N96; E, Pseudaganides buffeventin. sp.,2023.PB.N76; F, Pseudaganides crechensisn. sp., 2023.PB.N107; G, Pseudaganides subbiangulatus (dOrbigny,1850),2023. PB.N73, Patina Zone; H, Pseudaganides subbiangulatus, 2023.PB.N112, Anceps Zone." figureDoi="http://doi.org/10.5281/zenodo.13884439" httpUri="https://zenodo.org/record/13884439/files/figure.png" pageId="11" pageNumber="480">Figs 4D</figureCitation>
;
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) is well marked (P/L &lt;30%) whereas the ventral lobe is very shallow.
</paragraph>
</subSubSection>
<subSubSection id="C339BDCAFF99FFF90719A437FBA6F9A9" pageId="11" pageNumber="480" type="discussion">
<paragraph id="8B9CEE41FF99FFF90719A437FC67FAB7" blockId="11.[811,1456,1366,1648]" box="[813,913,1366,1392]" pageId="11" pageNumber="480">
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</paragraph>
<paragraph id="8B9CEE41FF99FFF90719A417FBA6F9A9" blockId="11.[811,1456,1366,1648]" pageId="11" pageNumber="480">
<taxonomicName id="4C2395C2FF99FFF90719A417FC01FA49" authorityName="Branger" authorityYear="2024" box="[813,1015,1398,1424]" class="Cephalopoda" family="Pseudonautilidae" genus="Antaganides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="11" pageNumber="480" phylum="Mollusca" rank="species" species="grulkei" status="n. gen., n. sp.">
<emphasis id="B9573253FF99FFF90719A417FC01FA49" box="[813,1015,1398,1424]" italics="true" pageId="11" pageNumber="480">Antaganides grulkei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2648F28FF99FFF907CFA417FB7CFA49" box="[1019,1162,1398,1424]" pageId="11" pageNumber="480" rank="species">n. gen., n. sp.</taxonomicNameLabel>
is a very close species to the contemporary
<taxonomicName id="4C2395C2FF99FFF907F3A4F7FB54FA69" authorityName="Branger" authorityYear="2024" box="[967,1186,1430,1456]" class="Cephalopoda" family="Pseudonautilidae" genus="Antaganides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="11" pageNumber="480" phylum="Mollusca" rank="species" species="fournieri" status="n. gen., n. sp.">
<emphasis id="B9573253FF99FFF907F3A4F7FB54FA69" box="[967,1186,1430,1456]" italics="true" pageId="11" pageNumber="480">Antaganides fournieri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2648F28FF99FFF90090A4F7FAC7FA69" box="[1188,1329,1430,1456]" pageId="11" pageNumber="480" rank="species">n. gen., n. sp.</taxonomicNameLabel>
, comparing size and shape of the shell. In comparison, the umbilicus is funnel-like in one case and deep and boarded by subvertical sides for the other. The lateral lobe of the suture line also shows differences, especially close to the umbilicus where the suture line is more retrosiphonate in
<taxonomicName id="4C2395C2FF99FFF9005EA757FB14F996" authorityName="Branger" authorityYear="2024" box="[1130,1250,1589,1616]" class="Cephalopoda" family="Pseudonautilidae" genus="Antaganides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="11" pageNumber="480" phylum="Mollusca" rank="species" species="fournieri" status="n. gen., n. sp.">
<emphasis id="B9573253FF99FFF9005EA757FB14F996" box="[1130,1250,1589,1616]" italics="true" pageId="11" pageNumber="480">A. fournieri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2648F28FF99FFF900DDA757FA8FF989" box="[1257,1401,1590,1616]" pageId="11" pageNumber="480" rank="species">n. gen., n. sp.</taxonomicNameLabel>
than on
<taxonomicName id="4C2395C2FF99FFF90764A734FC40F9B6" authorityName="Branger" authorityYear="2024" box="[848,950,1621,1647]" class="Cephalopoda" family="Pseudonautilidae" genus="Antaganides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="11" pageNumber="480" phylum="Mollusca" rank="species" species="grulkei" status="n. gen., n. sp.">
<emphasis id="B9573253FF99FFF90764A734FC40F9B6" box="[848,950,1621,1647]" italics="true" pageId="11" pageNumber="480">A. grulkei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2648F28FF99FFF90789A737FBA6F9A9" box="[957,1104,1622,1648]" pageId="11" pageNumber="480" rank="species">n. gen., n. sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>