249 lines
31 KiB
XML
249 lines
31 KiB
XML
<document id="D44172E17095AC112A4C1EED5A9D2E13" ID-DOI="10.5852/cr-palevol2020v19a1" ID-ISSN="1777-571X" ID-Zenodo-Dep="14203901" ID-ZooBank="urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1732275478277" checkinUser="felipe" docAuthor="Van Den Hoek Ostende, Lars W., Casanovas-Vilar, Isaac & Furió, Marc" docDate="2020" docId="03F087FEFFD5FF80FEC1C2D32D46FEC5" docLanguage="en" docName="CRPalevol.19.1.1-25.pdf" docOrigin="Comptes Rendus Palevol 19 (1)" docSource="http://dx.doi.org/10.5852/cr-palevol2020v19a1" docStyle="DocumentStyle:DACF86F01658A8850E13A63D98C287FD.1:CRPalevol.2020-.journal_article" docStyleId="DACF86F01658A8850E13A63D98C287FD" docStyleName="CRPalevol.2020-.journal_article" docStyleVersion="1" docTitle="Soricinae Fischer 1814" docType="treatment" docVersion="3" lastPageNumber="20" masterDocId="FFC9FF86FFC1FF95FFF1C0262F68FFB4" masterDocTitle="Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="19" updateTime="1732302480368" updateUser="guilherme" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="AEA73E1883F8E4389A7FC12834D9020D">
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<mods:title id="6FFA5C5C9962B77382FB8835B6DF483D">Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain)</mods:title>
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<mods:name id="84C3936AB160F08456FF24864960AD97" type="personal">
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<mods:roleTerm id="744F7943558BFF104285A92038DCD008">Author</mods:roleTerm>
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<mods:namePart id="8C08293FCAD505131C9F7F3D1EB2125C">Van Den Hoek Ostende, Lars W.</mods:namePart>
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<mods:affiliation id="F2595ED4770CA6986DA453561BB96A95">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands)</mods:affiliation>
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<mods:nameIdentifier id="C1F653ADD0148FC9304831AB7DE5C43B" type="email">lars.vandenhoekostende@naturalis.nl</mods:nameIdentifier>
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<mods:name id="339BC19512A0E008012CE5277E35AE3B" type="personal">
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<mods:roleTerm id="9E5C77FD76B4CA69C6533BFCB9CC6547">Author</mods:roleTerm>
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<mods:namePart id="F639DC9F39801801E778008617EDFFA8">Casanovas-Vilar, Isaac</mods:namePart>
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<mods:affiliation id="AECD566CBF047616459DF7E8CD14056D">Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain)</mods:affiliation>
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<mods:nameIdentifier id="7D464015DE168B2C11D01070EB34FC53" type="email">isaac.casanovas@icp.cat</mods:nameIdentifier>
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<mods:namePart id="C6785C865811C11A7B8E1C695DA87EA8">Furió, Marc</mods:namePart>
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<mods:affiliation id="2B6B271F5AFE157EF5C6E7AF1E94009F">Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain)</mods:affiliation>
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<mods:nameIdentifier id="28E23EDA2DA63092B440F14BEDA53333" type="email">marc.furio@uab.cat</mods:nameIdentifier>
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<mods:title id="5E44391E9FCB5A964B7540034C18BCBC">Comptes Rendus Palevol</mods:title>
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</mods:titleInfo>
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<mods:part id="AFFC1B52581B72CD28A6E72149E403A5">
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<mods:date id="A94CFD262E3A2BE7A99D7E56B71858EC">2020</mods:date>
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<mods:number id="335953F046AC2545672BC8D1FC27F003">2020-07-22</mods:number>
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<mods:detail id="8ADDE6A4634229BE2E289660F9F4EE2F" type="volume">
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<mods:number id="8AC83F9B7C5AEABF0DE80AB65CADD19B">19</mods:number>
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<mods:url id="96A975A31F71E3CA858BD5A84CFCCAE7">http://dx.doi.org/10.5852/cr-palevol2020v19a1</mods:url>
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<mods:classification id="12330BF6370DCE32AE7DB81C8002E89C">journal article</mods:classification>
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<mods:identifier id="052C4944740E8B437E47AEE3C994EA12" type="DOI">10.5852/cr-palevol2020v19a1</mods:identifier>
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<mods:identifier id="812C7B1D984344ADF857771A804829DA" type="ISSN">1777-571X</mods:identifier>
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</mods:mods>
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<treatment id="03F087FEFFD5FF80FEC1C2D32D46FEC5" LSID="urn:lsid:plazi:treatment:03F087FEFFD5FF80FEC1C2D32D46FEC5" httpUri="http://treatment.plazi.org/id/03F087FEFFD5FF80FEC1C2D32D46FEC5" lastPageId="21" lastPageNumber="20" pageId="20" pageNumber="19">
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<subSubSection id="C3436563FFD5FF81FEC1C2D32D34FCBB" box="[304,604,757,784]" pageId="20" pageNumber="19" type="nomenclature">
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<paragraph id="8BE636E8FFD5FF81FEC1C2D32D34FCBB" blockId="20.[304,604,757,784]" box="[304,604,757,784]" pageId="20" pageNumber="19">
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<heading id="D0AE8184FFD5FF81FEC1C2D32D34FCBB" box="[304,604,757,784]" centered="true" fontSize="9" level="2" pageId="20" pageNumber="19" reason="7">
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<taxonomicName id="4C594D6BFFD5FF81FEC1C2D32ECAFCBA" ID-CoL="KCG" authorityName="Fischer" authorityYear="1814" box="[304,418,757,783]" class="Mammalia" family="Soricidae" kingdom="Animalia" order="Soricomorpha" pageId="20" pageNumber="19" phylum="Chordata" rank="subFamily" subFamily="Soricinae">
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</taxonomicName>
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gen. et sp. indet.
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3436563FFD5FF81FE79C3332D6AFC9B" box="[392,514,789,816]" pageId="20" pageNumber="19" type="description">
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<paragraph id="8BE636E8FFD5FF81FE79C3332D6AFC9B" blockId="20.[392,514,789,816]" box="[392,514,789,816]" pageId="20" pageNumber="19">
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(
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<figureCitation id="13622A6DFFD5FF81FE60C3332EA7FC84" box="[401,463,789,816]" captionStart="FIG" captionStartId="16.[132,157,1872,1889]" captionTargetBox="[165,1422,214,1829]" captionTargetId="figure-252@16.[703,1243,783,1355]" captionTargetPageId="16" captionText="FIG. 5. — Heterosoricid and soricids from the early Miocene of the Vallès-Penedès:Heterosorex neumayrianus Schlosser,1887 (A-G). A, IPS86252.Ax sin. CMV-I. B, IPS21023. P4 dext. (rev). CMV-II.C, IPS21025. M1+M2 dext.(rev). CMV-I. D, IPS86241. M2+M3 sin. CMV-I. E, IPS86251. ax sin. CMV-I. (ocl). F, IPS86254. i1 sin.CMV-I.(lab).G, IPS86244.m1-m3 sin. CMV-I.(G1-ocl; G2-lab).cf.Clapasorex alvarezae (H). H, IPS86819.m1 dext. (rev).TFR.(H1-ocl; H2-lab).cf.Oligosorex thauensis (I). I, IPS86866.m2 sin. VdP. (a-ocl; b-lab). Miosorex sp.(J-M). J, IPS86865. I1 sin. VdP. (lab). K, IPS86242.M1 sin.CMV-I.L, IPS86263.m1 sin. CMV-II. (L1-ocl; L2-lab).M, IPS90090.m3 sin.CMV-III.(M1-ocl; M2-lab).cf. Florinia (N-O). N, IPS86186.m1+m2 dext.(rev).SAB1.(N1-ocl; N2-lab).O, IPS85665.P4 dext. (rev) CB2. Paenelimnoecus (P-R). P, IPS86422. P4 sin. EPapiol. Q, IPS86425. m1 sin. EPapiol. (Q1-ocl; Q2-lab). R, IPS86242. m2 dext. (rev). EPapiol. (R1-ocl; R2-lab). Soricinae indet. (S-U). S, IPS85553. I1 sin. EC. (lab). T, IPS85554. i1 sin. EC. (lab). U, IPS85551. mand a1-m1 sin. EC. (U1-ocl; U2-lab). Scale bar: 1 mm." pageId="20" pageNumber="19">Fig. 5</figureCitation>
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S-U)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3436563FFD5FF81FF75C3692C6AFBB1" pageId="20" pageNumber="19" type="materials_examined">
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<paragraph id="8BE636E8FFD5FF81FF75C3692C6AFBB1" blockId="20.[131,777,846,1029]" pageId="20" pageNumber="19">
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<smallCapsWord id="8D00A034FFD5FF81FF75C3692F81FCD1" baselines="864,865" box="[132,233,847,870]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Material" pageId="20" pageNumber="19">MATERIAL</smallCapsWord>
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<smallCapsWord id="8D00A034FFD5FF81FF00C3722E73FCD1" baselines="865" box="[241,283,852,869]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="and" pageId="20" pageNumber="19">AND</smallCapsWord>
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<smallCapsWord id="8D00A034FFD5FF81FED2C3722ED5FCD1" baselines="865" box="[291,445,852,869]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="measurements" pageId="20" pageNumber="19">MEASUREMENTS</smallCapsWord>
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(in cm). —
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<materialsCitation id="3B313CB5FFD5FF81FDB1C3762D96FBB1" collectionCode="IPS" country="Spain" location="El Canyet" pageId="20" pageNumber="19" specimenCode="IPS85553, IPS85552, IPS85555, IPS85554, IPS85556, IPS85557, IPS20979, IPS85550, IPS85551" specimenCount="1">
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<emphasis id="B92DEAFAFFD5FF81FDB1C3762DE8FCDC" bold="true" box="[576,640,848,872]" pageId="20" pageNumber="19">
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||
<collectingCountry id="F34E7678FFD5FF81FDB1C3762D13FCDC" box="[576,635,848,872]" name="Spain" pageId="20" pageNumber="19">Spain</collectingCountry>
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||
.
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||
</emphasis>
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||
<location id="8E866033FFD5FF81FD79C3682D86FCD2" LSID="urn:lsid:plazi:treatment:03F087FEFFD5FF80FEC1C2D32D46FEC5:8E866033FFD5FF81FD79C3682D86FCD2" box="[648,750,846,870]" country="Spain" name="El Canyet" pageId="20" pageNumber="19">El Canyet</location>
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, 1 I1 sin.,
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<specimenCode id="DBFF9E93FFD5FF81FF27C34E2E54FC34" box="[214,316,872,896]" pageId="20" pageNumber="19">IPS85553</specimenCode>
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, 1.81 × 075 × 1.30; 2 i1 dext.,
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<specimenCode id="DBFF9E93FFD5FF81FD72C34E2D82FC34" box="[643,746,872,896]" pageId="20" pageNumber="19">IPS85552</specimenCode>
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, –,
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<specimenCode id="DBFF9E93FFD5FF81FF77C3A52F83FC2F" box="[134,235,899,923]" pageId="20" pageNumber="19">IPS85555</specimenCode>
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, 3.20; 2 i1 sin.,
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<specimenCode id="DBFF9E93FFD5FF81FE72C3A52E8FFC2F" box="[387,487,899,923]" pageId="20" pageNumber="19">IPS85554</specimenCode>
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, 3.60,
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<specimenCode id="DBFF9E93FFD5FF81FDD9C3A52DE4FC2F" box="[552,652,899,923]" pageId="20" pageNumber="19">IPS85556</specimenCode>
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, –,> 2.94; 1 mandible with i1 and p4 sin.,
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<specimenCode id="DBFF9E93FFD5FF81FE59C3BB2D64FC01" box="[424,524,925,949]" pageId="20" pageNumber="19">IPS85557</specimenCode>
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, i1 3.35, p4 1.27 × 0.90); 1 mandible with i1 and m1 dext.,
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<specimenCode id="DBFF9E93FFD5FF81FE2BC39E2D56FC64" box="[474,574,952,976]" pageId="20" pageNumber="19">IPS20979</specimenCode>
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, i1 3.18, m1 1.55 × 0.88 × 0.98; 1 p4 sin.,
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<specimenCode id="DBFF9E93FFD5FF81FEAFC3F42EAAFC5E" box="[350,450,978,1002]" pageId="20" pageNumber="19">IPS85550</specimenCode>
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, 1.20 × 0.82; 1 mandible with a1- m1 sin.,
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<specimenCode id="DBFF9E93FFD5FF81FF2BC3CB2E56FBB1" box="[218,318,1005,1029]" pageId="20" pageNumber="19">IPS85551</specimenCode>
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, 0.81 × 0.65, 1.10 × 0.79, 1.54 × 0.86 × 0.98
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3436563FFD5FF81FF75C4132BEEFEE5" pageId="20" pageNumber="19" type="description">
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<paragraph id="8BE636E8FFD5FF81FF75C4132E7DFBF9" blockId="20.[130,775,1077,1294]" box="[132,277,1077,1103]" pageId="20" pageNumber="19">
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<smallCapsWord id="8D00A034FFD5FF81FF75C4132E7DFBF9" baselines="1096,1096" box="[132,277,1077,1103]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="20" pageNumber="19">DESCRIPTION</smallCapsWord>
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</paragraph>
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<paragraph id="8BE636E8FFD5FF81FF75C4722FF4FBDA" blockId="20.[130,775,1077,1294]" box="[132,156,1108,1134]" pageId="20" pageNumber="19">
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<emphasis id="B92DEAFAFFD5FF81FF75C4722FF4FBDA" box="[132,156,1108,1134]" italics="true" pageId="20" pageNumber="19">I1</emphasis>
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</paragraph>
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<paragraph id="8BE636E8FFD5FF81FF75C4522EEBFABA" blockId="20.[130,775,1077,1294]" pageId="20" pageNumber="19">The upper incisor is not fissident, but there is a medial cuspule visible in dorsal view. The dorsal margin is regularly curved. The talon is not much developed. There is a narrow cingulum covering the base of the tooth. The root is only somewhat shorter than the crown.</paragraph>
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<paragraph id="8BE636E8FFD5FF81FF75C5152F81FAF9" blockId="20.[130,775,1331,1454]" box="[132,233,1331,1357]" pageId="20" pageNumber="19">
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<emphasis id="B92DEAFAFFD5FF81FF75C5152F81FAF9" box="[132,233,1331,1357]" italics="true" pageId="20" pageNumber="19">Mandible</emphasis>
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</paragraph>
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<paragraph id="8BE636E8FFD5FF81FF75C5752C6DFA19" blockId="20.[130,775,1331,1454]" pageId="20" pageNumber="19">The mandible shows that the mental foramen is placed below the reentrant valley of the m1. It is also noteworthy that the i1 extends back labially as much as below the paraconid of the m1.</paragraph>
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<paragraph id="8BE636E8FFD5FF81FF75C5F52FF2FA59" blockId="20.[131,775,1491,1613]" box="[132,154,1491,1517]" pageId="20" pageNumber="19">
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<emphasis id="B92DEAFAFFD5FF81FF75C5F52FF2FA59" box="[132,154,1491,1517]" italics="true" pageId="20" pageNumber="19">i1</emphasis>
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</paragraph>
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<paragraph id="8BE636E8FFD5FF81FF75C5D52E57F9F8" blockId="20.[131,775,1491,1613]" pageId="20" pageNumber="19">The lower incisor is bicuspulate. The anterior tip and the posterior cuspule have a similar height, whereas the intermediate one is a bit lower.</paragraph>
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<paragraph id="8BE636E8FFD5FF81FF75C6542FF7F938" blockId="20.[130,775,1650,1772]" box="[132,159,1650,1676]" pageId="20" pageNumber="19">
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||
<emphasis id="B92DEAFAFFD5FF81FF75C6542FF7F938" box="[132,159,1650,1676]" italics="true" pageId="20" pageNumber="19">a1</emphasis>
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||
</paragraph>
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<paragraph id="8BE636E8FFD5FF81FF75C6B42DAFF958" blockId="20.[130,775,1650,1772]" pageId="20" pageNumber="19">The antemolar has a heart-shaped occlusal outline. It is smaller in size than its posterior counterpart, the p4. There is no posterolingual basin, but it has a tetrahedral morphology.</paragraph>
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<paragraph id="8BE636E8FFD5FF81FF75C7342FF6F898" blockId="20.[132,774,1810,1900]" box="[132,158,1810,1836]" pageId="20" pageNumber="19">
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<emphasis id="B92DEAFAFFD5FF81FF75C7342FF6F898" box="[132,158,1810,1836]" italics="true" pageId="20" pageNumber="19">p4</emphasis>
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</paragraph>
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<paragraph id="8BE636E8FFD5FF81FF75C7172DA0F8DF" blockId="20.[132,774,1810,1900]" pageId="20" pageNumber="19">The last antemolar has a heart-shaped occlusal outline and a ‘paralophid-like’ cristid, which defines a lingual valley.</paragraph>
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<paragraph id="8BE636E8FFD5FF81FF75C7B72FCFF81F" blockId="20.[132,777,1937,2027]" box="[132,167,1937,1963]" pageId="20" pageNumber="19">
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||
<emphasis id="B92DEAFAFFD5FF81FF75C7B72FCFF81F" box="[132,167,1937,1963]" italics="true" pageId="20" pageNumber="19">m1</emphasis>
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</paragraph>
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<paragraph id="8BE636E8FFD5FF81FF75C7972BEEFEE5" blockId="20.[132,777,1937,2027]" lastBlockId="20.[812,1457,215,337]" pageId="20" pageNumber="19">The most significant characters are the presence of a high entoconid and its corresponding cristid, the oblique cristid ending much labially and a continuous cingulid covering all the base of the tooth, also by the lingual side. The rest of the characters (or their ‘absence’) are detailed in the discussion about the taxonomic ascription.</paragraph>
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||
</subSubSection>
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<subSubSection id="C3436563FFD5FF80FCDCC1512D46FEC5" lastPageId="21" lastPageNumber="20" pageId="20" pageNumber="19" type="discussion">
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<paragraph id="8BE636E8FFD5FF81FCDCC1512CF9FE3B" blockId="20.[810,1458,375,2027]" box="[813,913,375,401]" pageId="20" pageNumber="19">
|
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|
||
</paragraph>
|
||
<paragraph id="8BE636E8FFD5FF81FCDCC1B02BE6FD44" blockId="20.[810,1458,375,2027]" pageId="20" pageNumber="19">
|
||
Surprisingly, this species does not show the typical characters of the
|
||
<taxonomicName id="4C594D6BFFD5FF81FC9AC1902B48FE64" authorityName="Reumer" authorityYear="1987" box="[875,1056,438,464]" class="Mammalia" family="Soricidae" kingdom="Animalia" order="Soricomorpha" pageId="20" pageNumber="16" phylum="Chordata" rank="subFamily" subFamily="Crocidosoricinae">Crocidosoricinae</taxonomicName>
|
||
, which were the dominant shrews during the early Miocene. Instead, the elements recovered have typical derived
|
||
<taxonomicName id="4C594D6BFFD5FF81FC21C1D02B5BFDA4" authorityName="Fischer" authorityYear="1814" box="[976,1075,502,528]" class="Mammalia" family="Soricidae" kingdom="Animalia" order="Soricomorpha" pageId="20" pageNumber="19" phylum="Chordata" rank="subFamily" subFamily="Soricinae">Soricinae</taxonomicName>
|
||
traits. This is something unusual in localities of this age, as red-toothed shrews become frequent in Eurasia at the end of the Miocene. Some anterior teeth are isolated elements which have been tentatively ascribed to this form because they all come from the same locality, and their general aspect fits well with those of the limited dental series recovered. The
|
||
<specimenCount id="9D5FFD61FFD5FF81FBE1C2902BC5FD7B" box="[1040,1197,693,720]" count="2" pageId="20" pageNumber="19" type="generic">two specimens</specimenCount>
|
||
with m1 show that this element is larger than in the rest.
|
||
</paragraph>
|
||
<paragraph id="8BE636E8FFD5FF81FCB2C2D32A6EFA19" blockId="20.[810,1458,375,2027]" pageId="20" pageNumber="19">
|
||
The most significant derived features are the presence of only two lower antemolars (including the p4), a mental foramen placed under the reentrant valley of m1 and a p4 with a posterolingual basin (instead of a two-branched protoconid). If not a soricine shrew in itself, this form could be also ascribed to one of those species purportedly leading to the
|
||
<taxonomicName id="4C594D6BFFD5FF81FABDC3B32AC7FC1B" authorityName="Fischer" authorityYear="1814" box="[1356,1455,917,943]" class="Mammalia" family="Soricidae" kingdom="Animalia" order="Soricomorpha" pageId="20" pageNumber="19" phylum="Chordata" rank="subFamily" subFamily="Soricinae">Soricinae</taxonomicName>
|
||
clade (according to
|
||
<bibRefCitation id="EFC84B19FFD5FF81FC0DC3932BC5FC7B" author="FURIO M. & SANTOS-CUBEDO A. & MINWER-BARAKAT R. & AGUSTI J." box="[1020,1197,949,975]" pageId="20" pageNumber="19" pagination="1018 - 1032" refId="ref21431" refString="FURIO M., SANTOS-CUBEDO A., MINWER-BARAKAT R. & AGUSTI J. 2007. - Evolutionary history of the African soricid Myosorex (Insectivora, Mammalia) out of Africa. Journal of Vertebrate Paleontology 27 (4): 1018 - 1032. www. jstor. org / stable / 30117466" type="journal article" year="2007">
|
||
Furió
|
||
<emphasis id="B92DEAFAFFD5FF81FBCCC3932B18FC7B" box="[1085,1136,949,975]" italics="true" pageId="20" pageNumber="19">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
), like
|
||
<taxonomicName id="4C594D6BFFD5FF81FB1EC3932ACFFC7B" box="[1263,1447,949,975]" class="Mammalia" family="Soricidae" genus="Carposorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="20" pageNumber="19" phylum="Chordata" rank="species" species="sylviae">
|
||
<emphasis id="B92DEAFAFFD5FF81FB1EC3932ACFFC7B" box="[1263,1447,949,975]" italics="true" pageId="20" pageNumber="19">Carposorex sylviae</emphasis>
|
||
</taxonomicName>
|
||
. Actually, the dimensions of the m1 match perfectly with that species, according to the data provided by
|
||
<bibRefCitation id="EFC84B19FFD5FF81FAF0C3D22AC0FBBB" author="CROCHET" box="[1281,1448,1012,1039]" pageId="20" pageNumber="19" pagination="631 - 652" refId="ref20715" refString="CROCHET J-. Y. 1975. - Diversite des Insectivores Soricides du Miocene inferieur de France. Colloques Internationaux, CNRS, Paris, 218: 631 - 652." type="journal article" year="1975">Crochet (1975)</bibRefCitation>
|
||
. Moreover, the teeth have a massive aspect, the p4 has a labial branch more developed than the lingual one, and the m1 has a developed entoconid. However, the position of the mental foramen (under the reentrant valley of the m1; more advanced in crocidosoricines, also in
|
||
<taxonomicName id="4C594D6BFFD5FF81FB5DC4B22A74FB1A" authorityName="Crochet" authorityYear="1975" box="[1196,1308,1172,1198]" class="Mammalia" family="Soricidae" genus="Carposorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="20" pageNumber="19" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92DEAFAFFD5FF81FB5DC4B22A74FB1A" box="[1196,1308,1172,1198]" italics="true" pageId="20" pageNumber="19">Carposorex</emphasis>
|
||
</taxonomicName>
|
||
, in which it is placed under the p4), the absence of any connection between posterolabial crest of the protoconid and the labial cingulid, and the smooth enamel (instead of wrinkled as expected for the genus) discards such generic and specific allocation. In the other known species of the genus,
|
||
<taxonomicName id="4C594D6BFFD5FF81FB2CC5122AD9FAF9" box="[1245,1457,1331,1358]" class="Mammalia" family="Soricidae" genus="Carposorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="20" pageNumber="19" phylum="Chordata" rank="species" species="burkarti">
|
||
<emphasis id="B92DEAFAFFD5FF81FB2CC5122AD9FAF9" box="[1245,1457,1331,1358]" italics="true" pageId="20" pageNumber="19">Carposorex burkarti</emphasis>
|
||
</taxonomicName>
|
||
described by
|
||
<bibRefCitation id="EFC84B19FFD5FF81FC47C5722BC6FADA" author="HUGUENEY M. & MEIN P. & MARIDET O." box="[950,1198,1363,1390]" pageId="20" pageNumber="19" pagination="23 - 49" refId="ref22142" refString="HUGUENEY M., MEIN P. & MARIDET O. 2012. - Revision and new data on the Early and Middle Miocene soricids (Soricomorpha, Mammalia) from central and southeastern France. Swiss Journal of Palaeontology 131: 23 - 49 https: // doi. org / 10.1007 / s 13358 - 011 - 0036 - 1" type="journal article" year="2012">
|
||
Hugueney
|
||
<emphasis id="B92DEAFAFFD5FF81FBDCC5722B37FAD9" box="[1069,1119,1363,1389]" italics="true" pageId="20" pageNumber="19">et al.</emphasis>
|
||
(2012)
|
||
</bibRefCitation>
|
||
, the differences are even more, because of the higher number of lower antemolars (at least three in
|
||
<taxonomicName id="4C594D6BFFD5FF81FC4DC5B52B76FA19" box="[956,1054,1427,1453]" class="Mammalia" family="Soricidae" genus="Carposorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="20" pageNumber="19" phylum="Chordata" rank="species" species="sylviae">
|
||
<emphasis id="B92DEAFAFFD5FF81FC4DC5B52B76FA19" box="[956,1054,1427,1453]" italics="true" pageId="20" pageNumber="19">C. sylviae</emphasis>
|
||
</taxonomicName>
|
||
) and the smaller size.
|
||
</paragraph>
|
||
<paragraph id="8BE636E8FFD5FF81FCB2C5952AC7F8FF" blockId="20.[810,1458,375,2027]" pageId="20" pageNumber="19">
|
||
The oldest representative of the subfamily
|
||
<taxonomicName id="4C594D6BFFD5FF81FB19C5952A2EFA79" authorityName="Fischer" authorityYear="1814" box="[1256,1350,1459,1485]" class="Mammalia" family="Soricidae" kingdom="Animalia" order="Soricomorpha" pageId="20" pageNumber="19" phylum="Chordata" rank="subFamily" subFamily="Soricinae">Soricinae</taxonomicName>
|
||
in Europe is
|
||
<taxonomicName id="4C594D6BFFD5FF81FCB5C5F22B63FA59" box="[836,1035,1491,1517]" class="Mammalia" family="Soricidae" genus="Hemisorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="20" pageNumber="19" phylum="Chordata" rank="species" species="robustus">
|
||
<emphasis id="B92DEAFAFFD5FF81FCB5C5F22B63FA59" box="[836,1035,1491,1517]" italics="true" pageId="20" pageNumber="19">Hemisorex robustus</emphasis>
|
||
</taxonomicName>
|
||
. This species has been described in the French sites of Vieux-Collonges, Sansan and La Grive L3, L7 and M (
|
||
<bibRefCitation id="EFC84B19FFD5FF81FC56C6342B2EF998" author="BAUDELOT S." box="[935,1094,1554,1580]" pageId="20" pageNumber="19" pagination="290 - 291" refId="ref19878" refString="BAUDELOT S. 1967. - Sur quelques Soricides (Insectivores) miocenes de Sansan (Gers). Comptes Rendus Sommaire des Seances de la Societe Geologique de France, 7: 290 - 291." type="journal article" year="1967">Baudelot 1967</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC84B19FFD5FF81FBA2C6352B86F999" author="ENGESSER B." box="[1107,1262,1555,1581]" pageId="20" pageNumber="19" pagination="35 - 363" refId="ref21191" refString="ENGESSER B. 1972. - Die obermiozane Saugetierfauna von Anwil (Baselland). Tatigkeitsberichte der Naturforschenden Gesellschaft Baselland 28: 35 - 363." type="journal article" year="1972">Engesser 1972</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC84B19FFD5FF81FB0BC6352A5AF999" author="ENGESSER B." box="[1274,1330,1555,1581]" pageId="20" pageNumber="19" pagination="1 - 91" refId="ref21244" refString="ENGESSER B. 2009. - The Insectivores (Mammalia) from Sansan (Middle Miocene south-western France). Schweizerische Palaontologische Abhandlungen 128: 1 - 91." type="journal article" year="2009">2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC84B19FFD5FF81FACEC6352CF3F9F9" author="HUGUENEY M. & MEIN P. & MARIDET O." pageId="20" pageNumber="19" pagination="23 - 49" refId="ref22142" refString="HUGUENEY M., MEIN P. & MARIDET O. 2012. - Revision and new data on the Early and Middle Miocene soricids (Soricomorpha, Mammalia) from central and southeastern France. Swiss Journal of Palaeontology 131: 23 - 49 https: // doi. org / 10.1007 / s 13358 - 011 - 0036 - 1" type="journal article" year="2012">
|
||
Hugueney
|
||
<emphasis id="B92DEAFAFFD5FF81FCDCC6152C36F9F8" box="[813,862,1586,1612]" italics="true" pageId="20" pageNumber="19">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
). In fact, the measurements of the m
|
||
<quantity id="4CA19B0DFFD5FF81FAD2C6152A21F9F8" box="[1315,1353,1586,1613]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="19" unit="in" value="1.0">1 in</quantity>
|
||
this form also fit quite well within the ranges provided by
|
||
<bibRefCitation id="EFC84B19FFD5FF81FAA3C6752C1EF939" author="ENGESSER B." pageId="20" pageNumber="19" pagination="1 - 91" refId="ref21244" refString="ENGESSER B. 2009. - The Insectivores (Mammalia) from Sansan (Middle Miocene south-western France). Schweizerische Palaontologische Abhandlungen 128: 1 - 91." type="journal article" year="2009">Engesser (2009)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFC84B19FFD5FF81FC41C6552BC5F939" author="HUGUENEY M. & MEIN P. & MARIDET O." box="[944,1197,1650,1677]" pageId="20" pageNumber="19" pagination="23 - 49" refId="ref22142" refString="HUGUENEY M., MEIN P. & MARIDET O. 2012. - Revision and new data on the Early and Middle Miocene soricids (Soricomorpha, Mammalia) from central and southeastern France. Swiss Journal of Palaeontology 131: 23 - 49 https: // doi. org / 10.1007 / s 13358 - 011 - 0036 - 1" type="journal article" year="2012">
|
||
Hugueney
|
||
<emphasis id="B92DEAFAFFD5FF81FBD6C6552B33F938" box="[1063,1115,1650,1676]" italics="true" pageId="20" pageNumber="19">et al.</emphasis>
|
||
(2012)
|
||
</bibRefCitation>
|
||
, with a length of about
|
||
<quantity id="4CA19B0DFFD5FF81FCDFC6B42CFAF919" box="[814,914,1682,1709]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="19" unit="mm" value="1.5">1.50 mm</quantity>
|
||
and a width a bit less than 1.00 mm. The massive aspect, the bicuspulate i1, the p4 with a posterolingual basin and the position of the mental foramen would also support such ascription. However, the occlusal outline of the m1 (not as much rectangular as in
|
||
<taxonomicName id="4C594D6BFFD5FF81FBCFC7342BC1F89F" authorityName="Baudelot" authorityYear="1967" box="[1086,1193,1810,1835]" class="Mammalia" family="Soricidae" genus="Hemisorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="20" pageNumber="19" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92DEAFAFFD5FF81FBCFC7342BC1F89F" box="[1086,1193,1810,1835]" italics="true" pageId="20" pageNumber="19">Hemisorex</emphasis>
|
||
</taxonomicName>
|
||
) and the entoconid cristids (more developed than in
|
||
<taxonomicName id="4C594D6BFFD5FF81FB98C7142BBCF8FF" authorityName="Baudelot" authorityYear="1967" box="[1129,1236,1842,1867]" class="Mammalia" family="Soricidae" genus="Hemisorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="20" pageNumber="19" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92DEAFAFFD5FF81FB98C7142BBCF8FF" box="[1129,1236,1842,1867]" italics="true" pageId="20" pageNumber="19">Hemisorex</emphasis>
|
||
</taxonomicName>
|
||
), make a difference.
|
||
</paragraph>
|
||
<paragraph id="8BE636E8FFD5FF81FCB2C7742AC1F85F" blockId="20.[810,1458,375,2027]" pageId="20" pageNumber="19">
|
||
<taxonomicName id="4C594D6BFFD5FF81FCB2C7742B6AF8DF" authorityName="Ziegler" authorityYear="2003" box="[835,1026,1873,1899]" class="Mammalia" family="Soricidae" genus="Paenesorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="20" pageNumber="19" phylum="Chordata" rank="species" species="bicuspis">
|
||
<emphasis id="B92DEAFAFFD5FF81FCB2C7742B6AF8DF" box="[835,1026,1873,1899]" italics="true" pageId="20" pageNumber="19">Paenesorex bicuspis</emphasis>
|
||
</taxonomicName>
|
||
from Petersbuch 31 described byZiegler (2003) could be another possible candidate.However, it differs from the species found in El Canyet in the presence of a vestigial a2 between a1 and p4. Moreover, the overall size of teeth in
|
||
<taxonomicName id="4C594D6BFFD5FF81FCDCC7F42CF1F85F" authorityName="Ziegler" authorityYear="2003" box="[813,921,2002,2027]" class="Mammalia" family="Soricidae" genus="Paenesorex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="20" pageNumber="19" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92DEAFAFFD5FF81FCDCC7F42CF1F85F" box="[813,921,2002,2027]" italics="true" pageId="20" pageNumber="19">Paenesorex</emphasis>
|
||
</taxonomicName>
|
||
is smaller than that of this undetermined soricid.
|
||
</paragraph>
|
||
<paragraph id="8BE636E8FFD4FF80FF6AC0F12D46FEC5" blockId="21.[131,775,215,369]" pageId="21" pageNumber="20">
|
||
As there is hitherto no soricid species described in the early or middle Miocene matching all these characters, and in the absence of sufficient material we refrain from describing a new species, the most sensible option is leaving the identification of this form as
|
||
<taxonomicName id="4C594D6BFFD4FF80FE73C1712E8EFEC5" authorityName="Fischer" authorityYear="1814" box="[386,486,343,369]" class="Mammalia" family="Soricidae" kingdom="Animalia" order="Soricomorpha" pageId="21" pageNumber="19" phylum="Chordata" rank="subFamily" subFamily="Soricinae">Soricinae</taxonomicName>
|
||
indet.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |