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<document id="A05D52987D30D260DC241565F126DCA8" ID-DOI="10.5852/cr-palevol2020v19a1" ID-ISSN="1777-571X" ID-Zenodo-Dep="14203901" ID-ZooBank="urn:lsid:zoobank.org:pub:9B7BA215-F7E3-4AF0-B764-43F0DD3EADB3" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1732275478277" checkinUser="felipe" docAuthor="Van Den Hoek Ostende, Lars W., Casanovas-Vilar, Isaac &amp; Furió, Marc" docDate="2020" docId="03F087FEFFCAFF98FF5EC6D72C6CF9F8" docLanguage="en" docName="CRPalevol.19.1.1-25.pdf" docOrigin="Comptes Rendus Palevol 19 (1)" docSource="http://dx.doi.org/10.5852/cr-palevol2020v19a1" docStyle="DocumentStyle:DACF86F01658A8850E13A63D98C287FD.1:CRPalevol.2020-.journal_article" docStyleId="DACF86F01658A8850E13A63D98C287FD" docStyleName="CRPalevol.2020-.journal_article" docStyleVersion="1" docTitle="Desmanodon daamsi van den Hoek Ostende 1997" docType="treatment" docVersion="3" lastPageNumber="12" masterDocId="FFC9FF86FFC1FF95FFF1C0262F68FFB4" masterDocTitle="Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="10" updateTime="1732302480368" updateUser="guilherme" zenodo-license-document="CC-BY-4.0">
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<mods:title id="BDFF60C62D5463773AD56959A3BD62E6">Stuck in the middle. A geographical appraisal of the oldest insectivores - and a marsupial - from the Vallès-Penedès Basin (early Miocene, Catalonia, Spain)</mods:title>
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<mods:namePart id="641886C16FE19B77CADA0DF518AB85AB">Van Den Hoek Ostende, Lars W.</mods:namePart>
<mods:affiliation id="4CB48897344C6B808B64AD75936899A5">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden (The Netherlands)</mods:affiliation>
<mods:nameIdentifier id="DDEF4DB957730986FBD4436431A1AF4C" type="email">lars.vandenhoekostende@naturalis.nl</mods:nameIdentifier>
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<mods:namePart id="00AB529AC1F73DBBD6B454DFC5CA649D">Casanovas-Vilar, Isaac</mods:namePart>
<mods:affiliation id="8743FBB2E23E0C1229E9AB340603F31A">Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain)</mods:affiliation>
<mods:nameIdentifier id="B1CB46104A4375574343B39DB0B41EFB" type="email">isaac.casanovas@icp.cat</mods:nameIdentifier>
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<mods:namePart id="735897B8BB5A68D787F3188FB6C9EDC1">Furió, Marc</mods:namePart>
<mods:affiliation id="904C65814E921F03B5215046007C17FA">Institut Català de Paleontologia Miquel Crusafont. Edifici Z, c / de les Columnes, Campus de la UAB, 08193 Cerdanyola del Vallès, Barcelona (Spain) and Departament de Geologia, Universitat Autònoma de Barcelona (08193), Cerdanyola del Vallès, Barcelona (Spain)</mods:affiliation>
<mods:nameIdentifier id="512981A3A361780194463888BA5239AD" type="email">marc.furio@uab.cat</mods:nameIdentifier>
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<mods:title id="BAAB4ACF55B7DC095119618676EC032E">Comptes Rendus Palevol</mods:title>
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<mods:part id="13B68E6BE834BB388D92B8FAC6EBAD77">
<mods:date id="22E7AAC4747210961D35F56BC0007A7E">2020</mods:date>
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<mods:number id="DDD0C50A1820DC15781A1A274F8DD7C2">2020-07-22</mods:number>
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<mods:number id="8A77C44BE077BFD700CF234F7C9AFAC3">19</mods:number>
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<subSubSection id="C3436563FFCAFF9EFF5EC6D72DB4F8B8" box="[175,732,1777,1804]" pageId="11" pageNumber="10" type="nomenclature">
<paragraph id="8BE636E8FFCAFF9EFF5EC6D72DB4F8B8" blockId="11.[175,732,1777,1804]" box="[175,732,1777,1804]" pageId="11" pageNumber="10">
<heading id="D0AE8184FFCAFF9EFF5EC6D72DB4F8B8" box="[175,732,1777,1804]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="10" reason="2">
<taxonomicName id="4C594D6BFFCAFF9EFF5EC6D72DB4F8B8" authority="Van den Hoek Ostende, 1997" authorityName="van den Hoek Ostende" authorityYear="1997" box="[175,732,1777,1804]" class="Mammalia" family="Talpidae" genus="Desmanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="10" phylum="Chordata" rank="species" species="daamsi">
<emphasis id="B92DEAFAFFCAFF9EFF5EC6D72EF9F8B8" bold="true" box="[175,401,1777,1804]" italics="true" pageId="11" pageNumber="10">Desmanodon daamsi</emphasis>
<bibRefCitation id="EFC84B19FFCAFF9EFE69C6D42DB4F8B8" author="VAN DEN HOEK OSTENDE L. W." box="[408,732,1778,1804]" pageId="11" pageNumber="10" pagination="1 - 2" refId="ref23776" refString="VAN DEN HOEK OSTENDE L. W. 1997. - Insectivore faunas from the Lower Miocene of Anatolia. Part 4: The genus Desmanodon (Talpidae) with the description of a new species from the Lower Miocene of Spain. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen, vol. 100, 1 - 2: 27 - 65." type="journal article" year="1997">Van den Hoek Ostende, 1997</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3436563FFCAFF9EFE78C7342D6AF898" box="[393,514,1810,1836]" pageId="11" pageNumber="10" type="description">
<paragraph id="8BE636E8FFCAFF9EFE78C7342D6AF898" blockId="11.[393,514,1810,1836]" box="[393,514,1810,1836]" pageId="11" pageNumber="10">
(
<figureCitation id="13622A6DFFCAFF9EFE63C7342EB8F898" box="[402,464,1810,1836]" captionStart="FIG" captionStartId="12.[132,157,1938,1955]" captionTargetBox="[140,1447,215,1895]" captionTargetId="figure-156@12.[699,1261,773,1336]" captionTargetPageId="12" captionText="FIG. 4. — Talpid and dimylids from the early Miocene of the Vallès-Penedès. Desmanodon daamsi Van den Hoek Ostende, 1997 (A-F). A, IPS106596. P4 dext. (rev). TFR. B, IPS86816. M1 sin. TFR. C, IPS86818. M2 dext. (rev). TFR. D, IPS86203. M3 sin. SAB-3. E, IPS86232. m2 dext. (rev). Palau 3B. (E1-ocl; E2-lab). F, IPS86278.m3 sin.CMV-III.(F1-ocl;F2-lab).Chainodus sp.(G). G, IPS116324.m1+m2 dext.(rev).CB2.(G1-ocl; G2-lab).Plesiodimylus sp.(H-K). H, IPS86256.M2 dext. (rev).CMV-II.I, IPS85481. M2 dext. (rev). TFR. J, IPS85668.m2 sin. CB2. (ocl). K, IPS86185. mand. with c-m2 sin.SAB1.(K1-ocl; K2-lab). Scale bar:1 mm." pageId="11" pageNumber="10">Fig. 4</figureCitation>
A-F)
</paragraph>
</subSubSection>
<subSubSection id="C3436563FFCAFF9EFF75C76D2A6DFE8A" pageId="11" pageNumber="10" type="materials_examined">
<paragraph id="8BE636E8FFCAFF9EFF75C76D2A6DFE8A" blockId="11.[131,776,1867,2023]" lastBlockId="11.[812,1456,215,318]" pageId="11" pageNumber="10">
<smallCapsWord id="8D00A034FFCAFF9EFF75C76D2F8FF8D5" baselines="1884,1885" box="[132,231,1867,1890]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Material" pageId="11" pageNumber="10">MATERIAL</smallCapsWord>
<smallCapsWord id="8D00A034FFCAFF9EFF1EC7762E71F8D5" baselines="1885" box="[239,281,1872,1889]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="and" pageId="11" pageNumber="10">AND</smallCapsWord>
<smallCapsWord id="8D00A034FFCAFF9EFEEEC7762EDFF8D5" baselines="1885" box="[287,439,1872,1889]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="measurements" pageId="11" pageNumber="10">MEASUREMENTS</smallCapsWord>
(in cm). —
<materialsCitation id="3B313CB5FFCAFF9EFDC4C76A2EECF8C9" collectionCode="IPS" country="Spain" location="El Palau" pageId="11" pageNumber="10" specimenCode="IPS86232" specimenCount="1">
<emphasis id="B92DEAFAFFCAFF9EFDC4C76A2D1BF8D0" bold="true" box="[565,627,1868,1892]" pageId="11" pageNumber="10">
<collectingCountry id="F34E7678FFCAFF9EFDC4C76A2D18F8D0" box="[565,624,1868,1892]" name="Spain" pageId="11" pageNumber="10">Spain</collectingCountry>
.
</emphasis>
<location id="8E866033FFCAFF9EFD8BC76D2DA3F8D7" LSID="urn:lsid:plazi:treatment:03F087FEFFCAFF98FF5EC6D72C6CF9F8:8E866033FFCAFF9EFD8BC76D2DA3F8D7" box="[634,715,1867,1891]" country="Spain" name="El Palau" pageId="11" pageNumber="10">El Palau</location>
3B,
<quantity id="4CA19B0DFFCAFF9EFD0BC76D2FF1F8C9" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="10" unit="m" value="1.0">1 m</quantity>
2 dext.,
<specimenCode id="DBFF9E93FFCAFF9EFF12C7432E2EF8C9" box="[227,326,1893,1917]" pageId="11" pageNumber="10">IPS86232</specimenCode>
, × ;
</materialsCitation>
<materialsCitation id="3B313CB5FFCAFF9EFE79C7432F9AF82C" collectionCode="IPS" country="Spain" elevation="1" location="Can Marti Vell" pageId="11" pageNumber="10" specimenCode="IPS86278" specimenCount="1">
<location id="8E866033FFCAFF9EFE79C7432D7EF8C9" LSID="urn:lsid:plazi:treatment:03F087FEFFCAFF98FF5EC6D72C6CF9F8:8E866033FFCAFF9EFE79C7432D7EF8C9" box="[392,534,1893,1917]" country="Spain" name="Can Marti Vell" pageId="11" pageNumber="10">Can Martí Vell</location>
III,
<quantity id="4CA19B0DFFCAFF9EFDCFC7432D0CF8C9" box="[574,612,1893,1917]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="10" unit="m" value="1.0">1 m</quantity>
3 sin.,
<specimenCode id="DBFF9E93FFCAFF9EFD51C7432C6BF8C9" box="[672,771,1893,1917]" pageId="11" pageNumber="10">IPS86278</specimenCode>
, 1.24 × 0.78
</materialsCitation>
;
<materialsCitation id="3B313CB5FFCAFF9EFF0BC7A62DA6F853" collectionCode="IPS" location="Sant Andreu de la Barca" municipality="Sant Andreu de la Barca" pageId="11" pageNumber="10" specimenCode="IPS86203, IPS86195, IPS86192-93, IPS86183, IPS86181" specimenCount="1">
<collectingMunicipality id="6B82AC92FFCAFF9EFF0BC7A62EB3F823" box="[250,475,1920,1944]" pageId="11" pageNumber="10">Sant Andreu de la Barca</collectingMunicipality>
3, 1 M3 sin.,
<specimenCode id="DBFF9E93FFCAFF9EFDADC7592DA8F823" box="[604,704,1919,1943]" pageId="11" pageNumber="10">IPS86203</specimenCode>
, 1.12 × 1.67; Sant Andreu de la Barca 1, 1 P? dext.,
<specimenCode id="DBFF9E93FFCAFF9EFDD2C7BC2DEFF806" box="[547,647,1946,1970]" pageId="11" pageNumber="10">IPS86195</specimenCode>
, 1.45 × 0.69; 2 P4 dext.,
<specimenCode id="DBFF9E93FFCAFF9EFF1EC7922E1EF878" box="[239,374,1972,1996]" pageId="11" pageNumber="10">IPS86192-93</specimenCode>
, 1.43 × 1.17; × ; 2 M1 sin.,
<specimenCode id="DBFF9E93FFCAFF9EFD6AC7922D96F878" box="[667,766,1972,1996]" pageId="11" pageNumber="10">IPS86183</specimenCode>
- 84, × ; 1.62 × 1.50; 1 M2 dext.,
<specimenCode id="DBFF9E93FFCAFF9EFE18C7E92D26F853" box="[489,590,1999,2023]" pageId="11" pageNumber="10">IPS86181</specimenCode>
, 1.77 × 2.25
</materialsCitation>
;
<materialsCitation id="3B313CB5FFCAFF9EFD29C7E92AC4FE90" collectionCode="IPS" location="Turo de les Forques" pageId="11" pageNumber="10" specimenCode="IPS85483, IPS85488, IPS86816, IPS86817, IPS86818" specimenCount="1">
<location id="8E866033FFCAFF9EFD29C7E92CD0FF5A" LSID="urn:lsid:plazi:treatment:03F087FEFFCAFF98FF5EC6D72C6CF9F8:8E866033FFCAFF9EFD29C7E92CD0FF5A" box="[728,952,215,2023]" name="Turo de les Forques" pageId="11" pageNumber="10">Turó de les Forques</location>
, 1 P4 dext.,
<specimenCode id="DBFF9E93FFCAFF9EFBC0C0F12BFDFF5B" box="[1073,1173,215,239]" pageId="11" pageNumber="10">IPS85483</specimenCode>
, 1.36 × 1.24; 1 damaged M1 dext.,
<specimenCode id="DBFF9E93FFCAFF9EFC98C0D72CA6FEBD" box="[873,974,241,265]" pageId="11" pageNumber="10">IPS85488</specimenCode>
, × ; 1 M1 sin.,
<specimenCode id="DBFF9E93FFCAFF9EFB76C0D72B84FEBD" box="[1159,1260,241,265]" pageId="11" pageNumber="10">IPS86816</specimenCode>
, 1.73 × 2.32; 1 M2 sin.,
<specimenCode id="DBFF9E93FFCAFF9EFCACC12A2CA9FE90" box="[861,961,268,292]" pageId="11" pageNumber="10">IPS86817</specimenCode>
, 1.74 × 1.97; 1 M2 dext.,
<specimenCode id="DBFF9E93FFCAFF9EFB36C12A2A44FE90" box="[1223,1324,268,292]" pageId="11" pageNumber="10">IPS86818</specimenCode>
, 1.76 × 2.13
</materialsCitation>
;
<materialsCitation id="3B313CB5FFCAFF9EFCDCC1002A68FE8A" box="[813,1280,294,318]" collectionCode="IPS" location="Costablanca" pageId="11" pageNumber="10" specimenCode="IPS19372" specimenCount="1">
<location id="8E866033FFCAFF9EFCDCC1002CCFFE8A" LSID="urn:lsid:plazi:treatment:03F087FEFFCAFF98FF5EC6D72C6CF9F8:8E866033FFCAFF9EFCDCC1002CCFFE8A" box="[813,935,294,318]" name="Costablanca" pageId="11" pageNumber="10">Costablanca</location>
, 1 M2 sin.,
<specimenCode id="DBFF9E93FFCAFF9EFBECC1002BEAFE8A" box="[1053,1154,294,318]" pageId="11" pageNumber="10">IPS19372</specimenCode>
, 1.91 × 1.84
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3436563FFCAFF98FCDCC1512C6FFC5A" lastPageId="13" lastPageNumber="12" pageId="11" pageNumber="10" type="description">
<paragraph id="8BE636E8FFCAFF9EFCDCC1512CD5FE3B" blockId="11.[811,1458,375,752]" box="[813,957,375,401]" pageId="11" pageNumber="10">
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</paragraph>
<paragraph id="8BE636E8FFCAFF9EFCDCC1B12C2DFE05" blockId="11.[811,1458,375,752]" box="[813,837,407,433]" pageId="11" pageNumber="10">
<emphasis id="B92DEAFAFFCAFF9EFCDCC1B12C2DFE05" box="[813,837,407,433]" italics="true" pageId="11" pageNumber="10">P?</emphasis>
</paragraph>
<paragraph id="8BE636E8FFCAFF9EFCDCC1912A48FD5B" blockId="11.[811,1458,375,752]" pageId="11" pageNumber="10">
One premolar from Sant Andreu de la Barca 1 is tentatively assigned to
<taxonomicName id="4C594D6BFFCAFF9EFC5FC1F02B5CFE44" authorityName="Engesser" authorityYear="1980" box="[942,1076,470,496]" class="Mammalia" family="Talpidae" genus="Desmanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="11" pageNumber="10" phylum="Chordata" rank="genus">
<emphasis id="B92DEAFAFFCAFF9EFC5FC1F02B5CFE44" box="[942,1076,470,496]" italics="true" pageId="11" pageNumber="10">Desmanodon</emphasis>
</taxonomicName>
based on its high tip. The occlusal outline is teardrop-shaped, with the point directed backwards. The tip of this unicuspid lies in the front part of the premolar at about two thirds its length. It bears a sharp posterocrista and has a round anterior face. A narrow cingulum surrounds the tooth, being interrupted at the anterolabial flank of the paracone only. The cingulum is a bit thicker at the posterior end of the premolar.There are two plank-shaped roots of about equal size, which stand parallel to one another.
</paragraph>
<paragraph id="8BE636E8FFCAFF9EFCDCC3332C22FC9B" blockId="11.[811,1455,789,1039]" box="[813,842,789,815]" pageId="11" pageNumber="10">
<emphasis id="B92DEAFAFFCAFF9EFCDCC3332C22FC9B" box="[813,842,789,815]" italics="true" pageId="11" pageNumber="10">P4</emphasis>
</paragraph>
<paragraph id="8BE636E8FFCAFF9EFCDCC3132BFDFBBA" blockId="11.[811,1455,789,1039]" pageId="11" pageNumber="10">
The outline of the occlusal surface is triangular; the premolar is somewhat longer than wide. Both parastyle and protocone are small, the posterocrista is S-curved (Morphotype C sensu
<bibRefCitation id="EFC84B19FFCAFF9EFCDAC3B32B02FC1B" author="VAN DEN HOEK OSTENDE L. W." box="[811,1130,917,943]" pageId="11" pageNumber="10" pagination="1 - 29" refId="ref23680" refString="VAN DEN HOEK OSTENDE L. W. 1989. - The Talpidae (Insectivora, Mammalia) of Eggingen-Mittelhart Baden-Wurttemberg, (F. R. G.) with special reference to the Paratalpa - Desmanodon lineage. Stuttgarter Beitrage zur Naturkunde, B 152: 1 - 29." type="journal article" year="1989">
<collectingRegion id="499DF80AFFCAFF9EFCDAC3B32C3DFC1B" box="[811,853,917,943]" country="Turkey" name="Van" pageId="11" pageNumber="10">Van</collectingRegion>
den Hoek Ostende 1989
</bibRefCitation>
). The paracone is high and its posterocrista is curved just behind its tip. The narrow cingulum is continuous on the lingual side, interrupted at the base of the paracone on the labial side.
</paragraph>
<paragraph id="8BE636E8FFCAFF9EFCDCC4122C3BFBFA" blockId="11.[811,1457,1076,1581]" box="[813,851,1076,1102]" pageId="11" pageNumber="10">
<emphasis id="B92DEAFAFFCAFF9EFCDCC4122C3BFBFA" box="[813,851,1076,1102]" italics="true" pageId="11" pageNumber="10">M1</emphasis>
</paragraph>
<paragraph id="8BE636E8FFCAFF9EFCDCC4722AD8F999" blockId="11.[811,1457,1076,1581]" pageId="11" pageNumber="10">
The labial cusps, and particularly the metacone, are very high. The occlusal surface is irregularly quadrangular, with a long lingual side. The posterior arm of the paracone is relatively straight. The posterior arm of the metacone is just somewhat longer than the anterior arm. The mesostyle is incompletely divided. There is a small, slightly protruding parastyle in the anterolabial corner of the molar. The low protocone lies in the anterolingual corner of the molar. Its short anterior arm ends against the base of the paracone. The posterior arm follows the contours of the outline lingually, and continues as the well-developed posterior cingulum. In one of the
<specimenCount id="9D5FFD61FFCAFF9EFA76C5B22CF2FA79" count="2" pageId="11" pageNumber="10" type="generic">two specimens</specimenCount>
from Turó de les Forques, a slight thickening in the posterior arm reveals the position of the hypocone. There is no protoconule. In one of the
<specimenCount id="9D5FFD61FFCAFF9EFB82C5D52A7AF9B9" box="[1139,1298,1523,1549]" count="2" pageId="11" pageNumber="10" type="generic">two specimens</specimenCount>
, there is a very short portion of lingual cingulum just behind the protocone.
</paragraph>
<paragraph id="8BE636E8FFCAFF9EFCDCC6742C3BF9D8" blockId="11.[811,1457,1618,1900]" box="[813,851,1618,1644]" pageId="11" pageNumber="10">
<emphasis id="B92DEAFAFFCAFF9EFCDCC6742C3BF9D8" box="[813,851,1618,1644]" italics="true" pageId="11" pageNumber="10">M2</emphasis>
</paragraph>
<paragraph id="8BE636E8FFCAFF9EFCDCC6552A09F8DF" blockId="11.[811,1457,1618,1900]" pageId="11" pageNumber="10">It is slightly asymmetrical due to the anterior position of the protocone and the small hypocone. There is no protoconule. The high labial cusps occupy about two thirds of the occlusal surface. The mesostyle is poorly divided. The small parastyle is slightly protruding; the metastyle is more developed as a labial thickening of the posterior cingulum. There are short stretches of cingulum along the anterior flank of the paracone and the posterior flank of the metacone, respectively.</paragraph>
<paragraph id="8BE636E8FFCAFF9EFCDCC7B72C3BF81F" blockId="11.[811,1456,1937,2028]" box="[813,851,1937,1963]" pageId="11" pageNumber="10">
<emphasis id="B92DEAFAFFCAFF9EFCDCC7B72C3BF81F" box="[813,851,1937,1963]" italics="true" pageId="11" pageNumber="10">M3</emphasis>
</paragraph>
<paragraph id="8BE636E8FFCAFF98FCDCC7942EC7FE25" blockId="11.[811,1456,1937,2028]" lastBlockId="13.[131,777,215,401]" lastPageId="13" lastPageNumber="12" pageId="11" pageNumber="10">It has an incompletely divided mesostyle, with the two cusplets clearly separated in the unworn specimen from Sant Andreu de la Barca 3. The anterior arm of the protocone is only slightly longer than the posterior arm, and bends sharply at its end to form a protruding parastyle. There is a short stretch of cingulum between the parastyle and the anterior flank of the paracone. The hypocone is a small but distinct cusp, well isolated from the metacone.</paragraph>
<caption id="DF266660FFCDFF99FF75C7B42AD9F853" pageId="12" pageNumber="11" startId="12.[132,157,1938,1955]" targetBox="[140,1447,215,1895]" targetPageId="12" targetType="figure">
<paragraph id="8BE636E8FFCDFF99FF75C7B42AD9F853" blockId="12.[132,1457,1938,2023]" pageId="12" pageNumber="11">
<smallCapsWord id="8D00A034FFCDFF99FF75C7B42FF5F817" baselines="1951,1952" box="[132,157,1938,1955]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="12" pageNumber="11">FIG</smallCapsWord>
. 4. — Talpid and dimylids from the early Miocene of the Vallès-Penedès.
<taxonomicName id="4C594D6BFFCDFF99FCF7C7B42BDBF817" authority="Van den Hoek Ostende, 1997" authorityName="van den Hoek Ostende" authorityYear="1997" box="[774,1203,1938,1955]" class="Mammalia" family="Talpidae" genus="Desmanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="12" pageNumber="11" phylum="Chordata" rank="species" species="daamsi">
<emphasis id="B92DEAFAFFCDFF99FCF7C7B42CDDF817" box="[774,949,1938,1955]" italics="true" pageId="12" pageNumber="11">Desmanodon daamsi</emphasis>
<bibRefCitation id="EFC84B19FFCDFF99FC4AC7B42BDBF817" author="VAN DEN HOEK OSTENDE L. W." box="[955,1203,1938,1955]" pageId="12" pageNumber="11" pagination="1 - 2" refId="ref23776" refString="VAN DEN HOEK OSTENDE L. W. 1997. - Insectivore faunas from the Lower Miocene of Anatolia. Part 4: The genus Desmanodon (Talpidae) with the description of a new species from the Lower Miocene of Spain. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen, vol. 100, 1 - 2: 27 - 65." type="journal article" year="1997">Van den Hoek Ostende, 1997</bibRefCitation>
</taxonomicName>
(
<emphasis id="B92DEAFAFFCDFF99FB4CC7B42BB4F817" bold="true" box="[1213,1244,1938,1955]" pageId="12" pageNumber="11">A -F</emphasis>
).
<emphasis id="B92DEAFAFFCDFF99FB1DC7B42B91F817" bold="true" box="[1260,1273,1938,1955]" pageId="12" pageNumber="11">A</emphasis>
, IPS106596. P4 dext. (rev). TFR.
<emphasis id="B92DEAFAFFCDFF99FF2FC78F2F83F80E" bold="true" box="[222,235,1961,1978]" pageId="12" pageNumber="11">B</emphasis>
, IPS86816. M1 sin. TFR.
<emphasis id="B92DEAFAFFCDFF99FE4EC78F2EA5F80E" bold="true" box="[447,461,1961,1978]" pageId="12" pageNumber="11">C</emphasis>
, IPS86818. M2 dext. (rev). TFR.
<emphasis id="B92DEAFAFFCDFF99FD2DC78F2D82F80E" bold="true" box="[732,746,1961,1978]" pageId="12" pageNumber="11">D</emphasis>
, IPS86203. M3 sin. SAB-3.
<emphasis id="B92DEAFAFFCDFF99FC23C78F2CB6F80E" bold="true" box="[978,990,1961,1978]" pageId="12" pageNumber="11">E</emphasis>
, IPS86232. m2 dext. (rev). Palau 3B. (E1-ocl; E2-lab).
<emphasis id="B92DEAFAFFCDFF99FA6EC78F2AC2F80E" bold="true" box="[1439,1450,1961,1978]" pageId="12" pageNumber="11">F</emphasis>
, IPS86278.m3 sin.CMV-III.(F1-ocl;F2-lab).
<taxonomicName id="4C594D6BFFCDFF99FE1DC7E62D0CF865" box="[492,612,1984,2001]" class="Mammalia" family="Dimylidae" genus="Chainodus" kingdom="Animalia" order="Erinaceomorpha" pageId="12" pageNumber="11" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B92DEAFAFFCDFF99FE1DC7E62D2DF865" box="[492,581,1984,2001]" italics="true" pageId="12" pageNumber="11">Chainodus</emphasis>
sp.
</taxonomicName>
(
<emphasis id="B92DEAFAFFCDFF99FD9AC7E62D11F865" bold="true" box="[619,633,1984,2001]" pageId="12" pageNumber="11">G</emphasis>
).
<emphasis id="B92DEAFAFFCDFF99FD76C7E62DFDF865" bold="true" box="[647,661,1984,2001]" pageId="12" pageNumber="11">G</emphasis>
, IPS116324.m1+m2 dext.(rev).CB2.(G1-ocl; G2-lab).
<taxonomicName id="4C594D6BFFCDFF99FBAAC7E62B84F865" box="[1115,1260,1984,2001]" class="Mammalia" family="Dimylidae" genus="Plesiodimylus" kingdom="Animalia" order="Erinaceomorpha" pageId="12" pageNumber="11" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B92DEAFAFFCDFF99FBAAC7E62BA5F865" box="[1115,1229,1984,2001]" italics="true" pageId="12" pageNumber="11">Plesiodimylus</emphasis>
sp.
</taxonomicName>
(
<emphasis id="B92DEAFAFFCDFF99FB02C7E62A7DF865" bold="true" box="[1267,1301,1984,2001]" pageId="12" pageNumber="11">H -K</emphasis>
).
<emphasis id="B92DEAFAFFCDFF99FAD5C7E62A5AF865" bold="true" box="[1316,1330,1984,2001]" pageId="12" pageNumber="11">H</emphasis>
, IPS86256.M2 dext. (rev).CMV-II.
<emphasis id="B92DEAFAFFCDFF99FED2C7F02E41F853" bold="true" box="[291,297,2006,2023]" pageId="12" pageNumber="11">I</emphasis>
, IPS85481. M2 dext. (rev). TFR.
<emphasis id="B92DEAFAFFCDFF99FDC2C7F02D55F853" bold="true" box="[563,573,2006,2023]" pageId="12" pageNumber="11">J</emphasis>
, IPS85668.m2 sin. CB2. (ocl).
<emphasis id="B92DEAFAFFCDFF99FCCDC7F02C21F853" bold="true" box="[828,841,2006,2023]" pageId="12" pageNumber="11">K</emphasis>
, IPS86185. mand. with c-m2 sin.SAB1.(K1-ocl; K2-lab). Scale bar:1 mm.
</paragraph>
</caption>
<paragraph id="8BE636E8FFCCFF98FF75C1912FCEFE65" blockId="13.[130,776,439,751]" box="[132,166,439,465]" pageId="13" pageNumber="12">
<emphasis id="B92DEAFAFFCCFF98FF75C1912FCEFE65" box="[132,166,439,465]" italics="true" pageId="13" pageNumber="12">m2</emphasis>
</paragraph>
<paragraph id="8BE636E8FFCCFF98FF75C1F02DD1FD5B" blockId="13.[130,776,439,751]" pageId="13" pageNumber="12">
The specimen from El
<collectingCountry id="F34E7678FFCCFF98FE97C1F02EF5FE44" box="[358,413,470,496]" name="Palau" pageId="13" pageNumber="12">Palau</collectingCountry>
3B has been restored after breakage. The trigonid is somewhat narrower and clearly shorter than the talonid. The cuspids are very high. The oblique cristid ends against the posterior wall of the trigonid, just labially to the metaconid. The entocristid is high, and quickly slopes down. The cingulid is well developed on the anterior and posterior sides. The lingual cingulid cannot be assessed properly because of the damage, but is certainly narrower than the anterior and posterior cingulids. The entostylid protrudes sharply.
</paragraph>
<paragraph id="8BE636E8FFCCFF98FF75C3332FCFFC9B" blockId="13.[132,775,789,1006]" box="[132,167,789,815]" pageId="13" pageNumber="12">
<emphasis id="B92DEAFAFFCCFF98FF75C3332FCFFC9B" box="[132,167,789,815]" italics="true" pageId="13" pageNumber="12">m3</emphasis>
</paragraph>
<paragraph id="8BE636E8FFCCFF98FF75C3102C6FFC5A" blockId="13.[132,775,789,1006]" pageId="13" pageNumber="12">It has a talonid which is only slightly reduced and somewhat shorter and narrower than the trigonid. The two arms of the protoconid form a narrow V-shape. The oblique cristid ends halfway the posterior wall of the trigonid. The entocristid is well developed. The anterior cingulid is narrow but distinct and rounds the paraconid, ending below the trigonid valley.</paragraph>
</subSubSection>
<subSubSection id="C3436563FFCCFF98FF75C4132C6CF9F8" pageId="13" pageNumber="12" type="discussion">
<paragraph id="8BE636E8FFCCFF98FF75C4132F80FBF9" blockId="13.[130,778,1077,1613]" box="[132,232,1077,1103]" pageId="13" pageNumber="12">
<smallCapsWord id="8D00A034FFCCFF98FF75C4132F80FBF9" baselines="1096,1096" box="[132,232,1077,1103]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="13" pageNumber="12">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="8BE636E8FFCCFF98FF75C4722C6CF9F8" blockId="13.[130,778,1077,1613]" pageId="13" pageNumber="12">
The early history of the genus
<taxonomicName id="4C594D6BFFCCFF98FE26C4722D36FBDA" authorityName="Engesser" authorityYear="1980" box="[471,606,1108,1134]" class="Mammalia" family="Talpidae" genus="Desmanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="13" pageNumber="12" phylum="Chordata" rank="genus">
<emphasis id="B92DEAFAFFCCFF98FE26C4722D36FBDA" box="[471,606,1108,1134]" italics="true" pageId="13" pageNumber="12">Desmanodon</emphasis>
</taxonomicName>
mirrors that of
<emphasis id="B92DEAFAFFCCFF98FF77C4522FBFFB3A" box="[134,215,1140,1166]" italics="true" pageId="13" pageNumber="12">
<taxonomicName id="4C594D6BFFCCFF98FF77C4522FBBFB3A" authorityName="Pomel" authorityYear="1848" box="[134,211,1140,1166]" class="Mammalia" family="Erinaceidae" genus="Galerix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Erinaceomorpha" pageId="13" pageNumber="12" phylum="Chordata" rank="genus">Galerix</taxonomicName>
.
</emphasis>
This talpid also had its oldest known records in
<collectingCountry id="F34E7678FFCCFF98FD41C4522C6FFB3A" box="[688,775,1140,1166]" name="Turkey" pageId="13" pageNumber="12">Anatolia</collectingCountry>
and it purportedly dispersed into Europe around the MN2/ MN3 transition (
<bibRefCitation id="EFC84B19FFCCFF98FEC9C4922D0EFB7A" author="VAN DEN HOEK OSTENDE L. W." box="[312,614,1204,1230]" pageId="13" pageNumber="12" pagination="1 - 2" refId="ref23776" refString="VAN DEN HOEK OSTENDE L. W. 1997. - Insectivore faunas from the Lower Miocene of Anatolia. Part 4: The genus Desmanodon (Talpidae) with the description of a new species from the Lower Miocene of Spain. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen, vol. 100, 1 - 2: 27 - 65." type="journal article" year="1997">
<collectingRegion id="499DF80AFFCCFF98FEC9C4922E0AFB7A" box="[312,354,1204,1230]" country="Turkey" name="Van" pageId="13" pageNumber="12">Van</collectingRegion>
den Hoek Ostende 1997
</bibRefCitation>
;
<bibRefCitation id="EFC84B19FFCCFF98FD80C4922E3CFB5A" author="VAN DEN HOEK OSTENDE L. W. &amp; FURIO M. &amp; MADERN A. &amp; PRIETO J." pageId="13" pageNumber="12" pagination="813 - 823" refId="ref24280" refString="VAN DEN HOEK OSTENDE L. W., FURIO M., MADERN A. &amp; PRIETO J. 2016. - Enters the shrew, some considerations on the Miocene palaeobiogeography of Iberian insectivores. Comptes Rendus Palevol 15: 813 - 823. https: // doi. org / 10.1016 / j. crpv. 2016.03.006" type="journal article" year="2016">
<collectingRegion id="499DF80AFFCCFF98FD80C4922DF3FB7A" box="[625,667,1204,1230]" country="Turkey" name="Van" pageId="13" pageNumber="12">Van</collectingRegion>
den Hoek Ostende
<emphasis id="B92DEAFAFFCCFF98FF14C4F22E7FFB5A" box="[229,279,1236,1262]" italics="true" pageId="13" pageNumber="12">et al.</emphasis>
2016
</bibRefCitation>
). The only species known so far from the Iberian Peninsula is
<taxonomicName id="4C594D6BFFCCFF98FEAFC4D22EA2FAB9" authorityName="van den Hoek Ostende" authorityYear="1997" box="[350,458,1267,1293]" class="Mammalia" family="Talpidae" genus="Desmanodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Soricomorpha" pageId="13" pageNumber="12" phylum="Chordata" rank="species" species="daamsi">
<emphasis id="B92DEAFAFFCCFF98FEAFC4D22EA2FAB9" box="[350,458,1267,1293]" italics="true" pageId="13" pageNumber="12">D. daamsi</emphasis>
</taxonomicName>
, which is characterized by an incompletely divided mesostyle in the upper molars and poor hypocone development on the M1 and M2. In these characters, as well as in size, the material from the Vallès-Penedès fits well with the earlier described assemblages from the basins of Calatayud-Teruel, Rubielos de Mora, and Ribesalbes-Alcora (
<bibRefCitation id="EFC84B19FFCCFF98FF7CC5952ED6FA79" author="VAN DEN HOEK OSTENDE L. W." box="[141,446,1459,1485]" pageId="13" pageNumber="12" pagination="1 - 2" refId="ref23776" refString="VAN DEN HOEK OSTENDE L. W. 1997. - Insectivore faunas from the Lower Miocene of Anatolia. Part 4: The genus Desmanodon (Talpidae) with the description of a new species from the Lower Miocene of Spain. Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen, vol. 100, 1 - 2: 27 - 65." type="journal article" year="1997">
<collectingRegion id="499DF80AFFCCFF98FF7CC5952FDFFA79" box="[141,183,1459,1485]" country="Turkey" name="Van" pageId="13" pageNumber="12">Van</collectingRegion>
den Hoek Ostende 1997
</bibRefCitation>
, 2003;
<bibRefCitation id="EFC84B19FFCCFF98FDFDC5952DBAFA7A" author="HORDIJK K. &amp; BOSMA A. A. &amp; DE BRUIJN H. &amp; VAN DAM J. &amp; GERAEDTS C. &amp; VAN DEN HOEK OSTENDE L. W. &amp; REUMER J. W. W. &amp; WESSELS W." box="[524,722,1459,1486]" pageId="13" pageNumber="12" pagination="321 - 346" refId="ref22043" refString="HORDIJK K., BOSMA A. A., DE BRUIJN H., VAN DAM J., GERAEDTS C., VAN DEN HOEK OSTENDE L. W., REUMER J. W. W. &amp; WESSELS W. 2015. - Biostratigraphic and paleoecologic implications of the small mammal assemblage from the late early Miocene of Montalvos 2, Teruel Basin, Spain. Palaeobiodiversity and Palaeoenvironment 95 (3): 321 - 346. https: // doi. org / 10.1007 / s 12549 - 015 - 0203 - 2" type="journal article" year="2015">
Hordijk
<emphasis id="B92DEAFAFFCCFF98FD97C5922DFEFA79" box="[614,662,1459,1485]" italics="true" pageId="13" pageNumber="12">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="EFC84B19FFCCFF98FD2CC5952EDEFA59" author="VAN DEN HOEK OSTENDE L. W. &amp; ALVAREZ-SIERRA M. A. &amp; GARCIA-PAREDES I. &amp; MONTOYA P. &amp; RUIZ-SANCHEZ F. J. &amp; PELAEZ-CAMPOMANES P." pageId="13" pageNumber="12" pagination="127 - 175" refId="ref24340" refString="VAN DEN HOEK OSTENDE L. W., ALVAREZ-SIERRA M. A., GARCIA-PAREDES I., MONTOYA P., RUIZ-SANCHEZ F. J. &amp; PELAEZ-CAMPOMANES P. 2017. - Alto de Ballester, biogeographical consequences of atypical MN 3 micromammal assemblages from eastern Spain. Palaeontographica Abteilung, A 308: 127 - 175. https: // doi. org / 10.1127 / pala / 308 / 2017 / 127" type="journal article" year="2017">
<collectingRegion id="499DF80AFFCCFF98FD2CC5952C62FA79" box="[733,778,1459,1485]" country="Turkey" name="Van" pageId="13" pageNumber="12">Van</collectingRegion>
den Hoek Ostende
<emphasis id="B92DEAFAFFCCFF98FEBDC5F22E14FA59" box="[332,380,1491,1517]" italics="true" pageId="13" pageNumber="12">et al.</emphasis>
2017
</bibRefCitation>
;
<bibRefCitation id="EFC84B19FFCCFF98FE30C5F52DA2FA59" author="CRESPO-ROURES V. D. &amp; MARQUINA-BLASCO R. &amp; MONTOYA P." box="[449,714,1491,1517]" pageId="13" pageNumber="12" pagination="407 - 416" refId="ref20645" refString="CRESPO-ROURES V. D., MARQUINA-BLASCO R., RUIZ-SANCHEZ, F. J. &amp; MONTOYA P. 2019. - An unusual insectivore assemblage from the early Miocene of southwestern Europe: The talpids and dimylids from the Ribesalbes-Alcora Basin (Spain). Comptes Rendus Palevol 18: 407 - 416. https: // doi. org / 10.1016 / j. crpv. 2019.03.003" type="journal article" year="2019">
Crespo-Roures
<emphasis id="B92DEAFAFFCCFF98FD90C5F22DF9FA59" box="[609,657,1491,1517]" italics="true" pageId="13" pageNumber="12">et al.</emphasis>
2019
</bibRefCitation>
). The most characteristic feature of the Iberian talpid, its relatively slender humerus, could not be ascertained for in the Vallès-Penedès assemblages as to date no humeri have been found.
</paragraph>
</subSubSection>
</treatment>
</document>