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<document id="044C26E2348E8F26BEB09B8DA1B2BAF0" ID-DOI="10.5281/zenodo.206618" ID-GBIF-Dataset="f5b1d7ac-ca42-4e4c-b2d0-b96771bc085c" ID-ISSN="1175-5326" ID-Zenodo-Dep="206618" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460257628293" checkinUser="plazi" docAuthor="Li, Chun-Lin, Wang, Chuan-Chan &amp; Chen, Hou-Jay" docDate="2011" docId="039387C6803DFFF861D1DDE159E4FA44" docLanguage="en" docName="zt02790p034.pdf" docOrigin="Zootaxa 2790" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Amphicoma Latreille 1807" docType="treatment" docVersion="9" lastPageNumber="25" masterDocId="FFAAFFBE803FFFFA6146DF04580DFFBE" masterDocTitle="Synopsis of the genus Amphicoma Latreille (Coleoptera: Glaphyridae) of Taiwan with special reference to the male genitalia" masterLastPageNumber="34" masterPageNumber="23" pageNumber="25" updateTime="1698243705731" updateUser="plazi">
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<mods:title id="894B9ED58FF9E327CB63FCE9255DB97C">Synopsis of the genus Amphicoma Latreille (Coleoptera: Glaphyridae) of Taiwan with special reference to the male genitalia</mods:title>
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<mods:namePart id="40A36AB55980FEC95AD09A3D40794FD7">Li, Chun-Lin</mods:namePart>
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<mods:namePart id="67194B4662EC7A30F67C38640B898871">Chen, Hou-Jay</mods:namePart>
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<mods:date id="A8CA651EB712AD497ADE2080B726EC10">2011</mods:date>
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<treatment id="039387C6803DFFF861D1DDE159E4FA44" ID-DOI="http://doi.org/10.5281/zenodo.6193206" ID-GBIF-Taxon="119392669" ID-Zenodo-Dep="6193206" LSID="urn:lsid:plazi:treatment:039387C6803DFFF861D1DDE159E4FA44" httpUri="http://treatment.plazi.org/id/039387C6803DFFF861D1DDE159E4FA44" lastPageNumber="25" pageId="2" pageNumber="25">
<subSubSection id="C320655B803DFFF861D1DDE15A36FD41" box="[151,571,741,767]" pageId="2" pageNumber="25" type="nomenclature">
<paragraph id="8B8536D0803DFFF861D1DDE15A36FD41" blockId="2.[151,571,741,767]" box="[151,571,741,767]" pageId="2" pageNumber="25">
<heading id="D0CD81BC803DFFF861D1DDE15A36FD41" bold="true" box="[151,571,741,767]" fontSize="11" level="1" pageId="2" pageNumber="25" reason="1">
<emphasis id="B94EEAC2803DFFF861D1DDE15A36FD41" bold="true" box="[151,571,741,767]" pageId="2" pageNumber="25">
Genus
<taxonomicName id="4C3A4D53803DFFF861A8DDE15A36FD41" ID-CoL="629ZQ" authority="Latreille, 1807" authorityName="Latreille" authorityYear="1807" box="[238,571,741,767]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC2803DFFF861A8DDE1598CFD41" bold="true" box="[238,385,741,767]" italics="true" pageId="2" pageNumber="25">Amphicoma</emphasis>
Latreille, 1807
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320655B803DFFF861D1DC285AB4FB67" pageId="2" pageNumber="25" type="reference_group">
<paragraph id="8B8536D0803DFFF861D1DC285A8DFCDE" blockId="2.[151,1292,811,957]" pageId="2" pageNumber="25">
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<taxonomicName id="4C3A4D53803DFFF861D1DC2859ECFCFF" ID-CoL="629ZQ" authority="Latreille, 1807: 118" authorityName="Latreille" authorityPageNumber="118" authorityYear="1807" box="[151,481,811,834]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC2803DFFF861D1DC28591DFCFC" box="[151,272,812,834]" italics="true" pageId="2" pageNumber="25">Amphicoma</emphasis>
Latreille, 1807: 118
</taxonomicName>
.
<typeStatus id="54818872803DFFF860ACDC295A11FCFC" box="[490,540,813,834]" pageId="2" pageNumber="25">Type</typeStatus>
species:
<taxonomicName id="4C3A4D53803DFFF8633FDC285C1BFCFF" authority="Fabricius, 1781" authorityName="Fabricius" authorityYear="1781" box="[633,1046,811,834]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="B94EEAC2803DFFF8633FDC285B7CFCFC" box="[633,881,812,834]" italics="true" pageId="2" pageNumber="25">Melolontha abdominalis</emphasis>
Fabricius, 1781
</taxonomicName>
by original designation.
<taxonomicName id="4C3A4D53803DFFF861D1DC4E59C4FCDE" authority="Gistel, 1848: 162" authorityName="Gistel" authorityPageNumber="162" authorityYear="1848" box="[151,457,842,864]" class="Insecta" family="Glaphyridae" genus="Amphitriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC2803DFFF861D1DC4E5919FCDE" box="[151,276,842,864]" italics="true" pageId="2" pageNumber="25">Amphitriche</emphasis>
<treatmentCitation id="0A9B10C1803DFFF8605CDC4E59C4FCDE" author="Gistel" box="[282,457,842,864]" page="162" pageId="2" pageNumber="25" year="1848">
<bibRefCitation id="EFAB4B21803DFFF8605CDC4E5997FCDE" author="Gistel" box="[282,410,842,864]" pageId="2" pageNumber="25" refString="Gistel, J. N. F. X. (1848) Naturgeschichte des Thierreichs. Fur hohere Schulen. Hohhmann'sche Verlags, Stuttgart, xvi + 216 + 4 pp." type="book" year="1848">Gistel, 1848</bibRefCitation>
: 162
</treatmentCitation>
</taxonomicName>
(junior synonym)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8536D0803DFFF861D1DC6D5A82FCC1" blockId="2.[151,1292,811,957]" box="[151,655,873,895]" pageId="2" pageNumber="25">
<treatmentCitationGroup id="AB2A11FE803DFFF861D1DC6D5A82FCC1" box="[151,655,873,895]" pageId="2" pageNumber="25">
<taxonomicName id="4C3A4D53803DFFF861D1DC6D59DAFCC1" ID-CoL="XJC" authority="Yawata, 1942: 34" authorityName="Yawata" authorityPageNumber="34" authorityYear="1942" box="[151,471,873,895]" class="Insecta" family="Glaphyridae" genus="Anthypnoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC2803DFFF861D1DC6D592CFCC1" box="[151,289,873,895]" italics="true" pageId="2" pageNumber="25">Anthypnoides</emphasis>
<treatmentCitation id="0A9B10C1803DFFF86061DC6E59DAFCC1" author="Yawata" box="[295,471,874,895]" page="34" pageId="2" pageNumber="25" year="1942">
<bibRefCitation id="EFAB4B21803DFFF86061DC6E59B9FCC1" author="Yawata" box="[295,436,874,895]" pageId="2" pageNumber="25" refString="Yawata, H. (1942) Notes on the Glaphyrinae of Japan with descriptions of a new genus and two new species. Transactions of the Kansai Entomological Society, 12, 33 - 37." type="journal article" year="1942">Yawata, 1942</bibRefCitation>
: 34
</treatmentCitation>
</taxonomicName>
(junior synonym)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8536D0803DFFF861D1DC8C5A8AFC20" blockId="2.[151,1292,811,957]" box="[151,647,904,926]" pageId="2" pageNumber="25">
<treatmentCitationGroup id="AB2A11FE803DFFF861D1DC8C5A8AFC20" box="[151,647,904,926]" pageId="2" pageNumber="25">
<taxonomicName id="4C3A4D53803DFFF861D1DC8C59DDFC23" ID-CoL="3383" authority="Fairmaire, 1891: 8" authorityName="Fairmaire" authorityPageNumber="8" authorityYear="1891" box="[151,464,904,926]" class="Insecta" family="Glaphyridae" genus="Arrhephora" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC2803DFFF861D1DC8C5903FC20" box="[151,270,904,926]" italics="true" pageId="2" pageNumber="25">Arrhephora</emphasis>
<treatmentCitation id="0A9B10C1803DFFF86052DC8C59DDFC23" author="Fairmaire" box="[276,464,904,926]" page="8" pageId="2" pageNumber="25" year="1891">
<bibRefCitation id="EFAB4B21803DFFF86052DC8C59B4FC23" author="Fairmaire" box="[276,441,904,926]" pageId="2" pageNumber="25" refString="Fairmaire, L. (1891) Descriptions de Coleopteres de l'interieur de la Chine. Annales de la Societe Entomologique de Belgique, 35, 6 - 24." type="journal article" year="1891">Fairmaire, 1891</bibRefCitation>
: 8
</treatmentCitation>
</taxonomicName>
(junior synonym)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8536D0803DFFF861D1DCAC5A7AFC02" blockId="2.[151,1292,811,957]" box="[151,631,934,957]" pageId="2" pageNumber="25">
<treatmentCitationGroup id="AB2A11FE803DFFF861D1DCAC5A7AFC02" box="[151,631,934,957]" pageId="2" pageNumber="25">
<taxonomicName id="4C3A4D53803DFFF861D1DCAC59CDFC02" ID-CoL="7Y4H" authority="Fairmaire, 1891: 7" authorityName="Fairmaire" authorityPageNumber="7" authorityYear="1891" box="[151,448,934,957]" class="Insecta" family="Glaphyridae" genus="Toxocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC2803DFFF861D1DCAC58F3FC03" box="[151,254,936,957]" italics="true" pageId="2" pageNumber="25">Toxocerus</emphasis>
<treatmentCitation id="0A9B10C1803DFFF86042DCA259CDFC02" author="Fairmaire" box="[260,448,934,956]" page="7" pageId="2" pageNumber="25" year="1891">
<bibRefCitation id="EFAB4B21803DFFF86042DCA259A4FC02" author="Fairmaire" box="[260,425,934,956]" pageId="2" pageNumber="25" refString="Fairmaire, L. (1891) Descriptions de Coleopteres de l'interieur de la Chine. Annales de la Societe Entomologique de Belgique, 35, 6 - 24." type="journal article" year="1891">Fairmaire, 1891</bibRefCitation>
: 7
</treatmentCitation>
</taxonomicName>
(junior synonym)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8536D0803DFFF861D1DCED5AB4FB67" blockId="2.[151,1437,1001,1530]" pageId="2" pageNumber="25">
<taxonomicName id="4C3A4D53803DFFF861D1DCED5910FBBF" ID-CoL="629ZQ" box="[151,285,1001,1025]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC2803DFFF861D1DCED5910FBBF" box="[151,285,1001,1025]" italics="true" pageId="2" pageNumber="25">Amphicoma</emphasis>
</taxonomicName>
can be separated from other glaphyrid genera by the following combination of characters: body surface pilose, setae moderate in length; male antennal club elongated, not cupuliform, antennal club outwardly/ inwardly curved; clypeus widened at base; mesoepimeron not visible dorsally; scutellum elongate-triangular; abdomen not elongate, prepygidium/tergite rarely exposed when viewed dorsally; protibia bidentate or tridentate, basal teeth weakly developed/vestigial; apex of male mesotibia with incision; metafemora not broadly enlarged; metatibia straight, not curved inwardly; protarsomeres 14 without comb-like spines and apical protarsomere longer than the basal three protarsomeres measured together.
</paragraph>
</subSubSection>
<subSubSection id="C320655B803DFFF86183DBE15BAEFA9F" pageId="2" pageNumber="25" type="distribution">
<paragraph id="8B8536D0803DFFF86183DBE15BAEFA9F" blockId="2.[151,1437,1001,1530]" pageId="2" pageNumber="25">
<emphasis id="B94EEAC2803DFFF86183DBE15953FB40" bold="true" box="[197,350,1253,1278]" pageId="2" pageNumber="25">Distribution.</emphasis>
Eastern to southwestern
<collectingCountry id="F32D7640803DFFF86333DBE15AB6FB43" box="[629,699,1253,1277]" name="China" pageId="2" pageNumber="25">China</collectingCountry>
, Indochina,
<collectingCountry id="F32D7640803DFFF86205DBE25B8BFB43" box="[835,902,1254,1277]" name="Japan" pageId="2" pageNumber="25">Japan</collectingCountry>
, Ryukyu Archipelago (both islands of Iriomotejima and Ishigakijima),
<collectingCountry id="F32D7640803DFFF860E4DA0D59FAFA9F" box="[418,503,1289,1313]" name="Taiwan" pageId="2" pageNumber="25">Taiwan</collectingCountry>
,
<collectingCountry id="F32D7640803DFFF86344DA0D5A86FA9F" box="[514,651,1289,1313]" name="Switzerland" pageId="2" pageNumber="25">Switzerland</collectingCountry>
,
<collectingCountry id="F32D7640803DFFF863D0DA0D5AE5FA9F" box="[662,744,1289,1313]" name="Greece" pageId="2" pageNumber="25">Greece</collectingCountry>
,
<collectingCountry id="F32D7640803DFFF863B5DA0D5B3EFA9F" box="[755,819,1289,1313]" name="Spain" pageId="2" pageNumber="25">Spain</collectingCountry>
and
<collectingCountry id="F32D7640803DFFF8622FDA0D5B93FA9F" box="[873,926,1289,1313]" name="Italy" pageId="2" pageNumber="25">Italy</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C320655B803DFFF86183DA2959E4FA44" pageId="2" pageNumber="25" type="discussion">
<paragraph id="8B8536D0803DFFF86183DA2959E4FA44" blockId="2.[151,1437,1001,1530]" pageId="2" pageNumber="25">
<emphasis id="B94EEAC2803DFFF86183DA295934FAFB" bold="true" box="[197,313,1325,1349]" pageId="2" pageNumber="25">Remarks.</emphasis>
Nikodým (2005) divided the genus
<taxonomicName id="4C3A4D53803DFFF8638ADA295B5FFAFB" box="[716,850,1325,1349]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC2803DFFF8638ADA295B5FFAFB" box="[716,850,1325,1349]" italics="true" pageId="2" pageNumber="25">Amphicoma</emphasis>
</taxonomicName>
into four species groups:
<taxonomicName id="4C3A4D53803DFFF86537DA295CFAFAFB" box="[1137,1271,1325,1349]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="corinthia">
<emphasis id="B94EEAC2803DFFF86537DA295CFAFAFB" box="[1137,1271,1325,1349]" italics="true" pageId="2" pageNumber="25">A. corinthia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D53803DFFF86442DA2958BFFAD7" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="B94EEAC2803DFFF86442DA295D18FAFB" box="[1284,1301,1325,1349]" italics="true" pageId="2" pageNumber="25">A</emphasis>
.
<emphasis id="B94EEAC2803DFFF86465DA2958BFFAD7" italics="true" pageId="2" pageNumber="25">abdominalis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3A4D53803DFFF86187DA555945FAD7" box="[193,328,1361,1385]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="florentini">
<emphasis id="B94EEAC2803DFFF86187DA555945FAD7" box="[193,328,1361,1385]" italics="true" pageId="2" pageNumber="25">A. florentini</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D53803DFFF860C4DA5559EEFAD7" box="[386,483,1361,1385]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="dubia">
<emphasis id="B94EEAC2803DFFF860C4DA5559EEFAD7" box="[386,483,1361,1385]" italics="true" pageId="2" pageNumber="25">A. dubia</emphasis>
</taxonomicName>
, based on three male character states of the mesotibial spur, the curviform antennal club and the length of antennal club. This delimitation of the species group concepts is roughly applicable to a local fauna like in
<collectingCountry id="F32D7640803DFFF8606BDA9D598FFA0F" box="[301,386,1433,1457]" name="Taiwan" pageId="2" pageNumber="25">Taiwan</collectingCountry>
, but it may not be appropriate when considering phylogenetic relationships within the entire genus. We therefore will refrain from using
<taxonomicName id="4C3A4D53803DFFF863C3DAB95B06FA6B" box="[645,779,1469,1493]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="B94EEAC2803DFFF863C3DAB95B06FA6B" box="[645,779,1469,1493]" italics="true" pageId="2" pageNumber="25">Amphicoma</emphasis>
</taxonomicName>
species groups until a comprehensive phylogenetic analysis is conducted for the genus.
</paragraph>
</subSubSection>
</treatment>
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