treatments-xml/data/BE/7C/63/BE7C6334FF8DFF8E2AE445068B37FB1B.xml

<document id="529279D437CC1B643D3CA298A0344B07" ID-DOI="10.5281/zenodo.4509187" ID-ISSN="2345-7600" ID-Zenodo-Dep="4509187" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612548490932" checkinUser="carolina" docAuthor="Husana, Daniel Edison M., Naruse, Tohru &amp; Kase, Tomoki" docDate="2010" docId="BE7C6334FF8DFF8E2AE445068B37FB1B" docLanguage="en" docName="RafflesBZool.58.1.51-55.pdf" docOrigin="Raffles Bulletin of Zoology 58 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Karstarma philippinarum Husana &amp; Naruse &amp; Kase 2010, new species" docType="treatment" docVersion="4" lastPageNumber="55" masterDocId="42451B4CFF8CFF8A2A3244788A61FFAE" masterDocTitle="A New Species Of The Genus Karstarma (Crustacea: Decapoda: Brachyura: Sesarmidae) From Anchialine Caves In The Philippines" masterLastPageNumber="55" masterPageNumber="51" pageNumber="52" updateTime="1698867934949" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="BD04DB036A776D71D8A8C2C942AF1FB8">A New Species Of The Genus Karstarma (Crustacea: Decapoda: Brachyura: Sesarmidae) From Anchialine Caves In The Philippines</mods:title>
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<mods:name id="3A4E486E2E4FF027FEFD92B31760C33E" type="personal">
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<mods:namePart id="57B05BA277717A2D316733A0E36D673C">Husana, Daniel Edison M.</mods:namePart>
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<mods:namePart id="8DA239651DADCADC8306CF2A33188531">Naruse, Tohru</mods:namePart>
</mods:name>
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<mods:namePart id="70EFB1DA5CA11BFFCE7B3E9DF50EEE71">Kase, Tomoki</mods:namePart>
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<mods:title id="A067CAE9AABA71C473C5AC465E8B8F93">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="5DAAE5E10FF15FE3C4A2A7D099348F10">
<mods:date id="46F9166F733DBF3133F44A8193512311">2010</mods:date>
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<mods:number id="98B21DBBD5F1DEAAEBE7DF01B712E50E">2010-02-28</mods:number>
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<mods:number id="9FFA530E804C9C5AF458E1F8E7B43EC7">58</mods:number>
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<mods:identifier id="B0D149129D88F81FC57F9FEFFAED5756" type="DOI">10.5281/zenodo.4509187</mods:identifier>
<mods:identifier id="CD530AA6B1692BD204AE609A2B6C7F7F" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="03473BD2E8E3C63AAE0DA6467B113D63" type="Zenodo-Dep">4509187</mods:identifier>
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<treatment id="BE7C6334FF8DFF8E2AE445068B37FB1B" LSID="urn:lsid:plazi:treatment:BE7C6334FF8DFF8E2AE445068B37FB1B" httpUri="http://treatment.plazi.org/id/BE7C6334FF8DFF8E2AE445068B37FB1B" lastPageId="4" lastPageNumber="55" pageId="1" pageNumber="52">
<subSubSection id="7ECF81A9FF8DFF8B2AE4450688E6FE38" box="[214,647,382,406]" pageId="1" pageNumber="52" type="nomenclature">
<paragraph id="366AD222FF8DFF8B2AE4450688E6FE38" blockId="1.[214,647,382,437]" box="[214,647,382,406]" pageId="1" pageNumber="52">
<heading id="6D22654EFF8DFF8B2AE4450688E6FE38" bold="true" box="[214,647,382,406]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="52" reason="2">
<emphasis id="04A10E30FF8DFF8B2AE4450688E6FE38" bold="true" box="[214,647,382,406]" pageId="1" pageNumber="52">
<taxonomicName id="F1D5A9A1FF8DFF8B2AE445068B93FE38" ID-CoL="3R3BY" authority="Husana &amp; Naruse &amp; Kase, 2010" authorityName="Husana &amp; Naruse &amp; Kase" authorityYear="2010" box="[214,498,382,406]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="philippinarum" status="sp. nov.">
<emphasis id="04A10E30FF8DFF8B2AE445068B93FE38" bold="true" box="[214,498,382,406]" italics="true" pageId="1" pageNumber="52">Karstarma philippinarum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="1F92B34BFF8DFF8B2833450688E6FE38" box="[513,647,382,406]" pageId="1" pageNumber="52" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7ECF81A9FF8DFF8B2B4045E68B8BFE1B" box="[370,490,414,437]" pageId="1" pageNumber="52" type="description">
<paragraph id="366AD222FF8DFF8B2B4045E68B8BFE1B" blockId="1.[214,647,382,437]" box="[370,490,414,437]" pageId="1" pageNumber="52">
(
<figureCitation id="AEEECEA7FF8DFF8B2B4845E68B81FE1B" box="[378,480,414,437]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[112,147,2060,2081]" captionStartId-1="1.[810,845,2060,2081]" captionStartId-2="2.[134,169,2003,2024]" captionTargetBox-0="[111,750,1526,2043]" captionTargetBox-1="[809,1448,1181,2042]" captionTargetBox-2="[142,1460,252,1964]" captionTargetId-0="figure-543@1.[111,750,1526,2043]" captionTargetId-1="figure-566@1.[809,1448,1181,2043]" captionTargetId-2="figure-89@2.[118,1481,230,1979]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionTargetPageId-2="2" captionText-0="Fig. 1. Live coloration of Karstarma philippinarum, new species, in its natural habitat, Bat Cave, Boracay Island, Philippines." captionText-1="Fig. 2. Karstarma philippinarum, new species. Holotype male (23.8 × 30.2 mm, NMCR 27063). a, habitus; b, carapace, dorsal view." captionText-2="Fig. 3. Karstarma philippinarum, new species. Paratype male NSMT-Cr2007.0105. a, carapace; b, abdomen and telson; c, cephalothorax, frontal view; d, left cheliped, dorsal view; e, left fourth ambulatory leg, dorsal view; f, left cheliped, carpus; g, left chela, outer view; h, left antenna, ventral view; i, G1, ventral and dorsal views; j, left third maxilliped, ventral view. Scale bars: a–e, g = 5.0 mm; f = 2.0 mm; h–j = 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4509189" figureDoi-1="http://doi.org/10.5281/zenodo.4509191" figureDoi-2="http://doi.org/10.5281/zenodo.4509193" httpUri-0="https://zenodo.org/record/4509189/files/figure.png" httpUri-1="https://zenodo.org/record/4509191/files/figure.png" httpUri-2="https://zenodo.org/record/4509193/files/figure.png" pageId="1" pageNumber="52">Figs. 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7ECF81A9FF8DFF892A5D45A5884CFB3B" lastPageId="3" lastPageNumber="54" pageId="1" pageNumber="52" type="materials_examined">
<paragraph id="366AD222FF8DFF8B2A5D45A58B39FD85" blockId="1.[111,750,477,555]" pageId="1" pageNumber="52">
<emphasis id="04A10E30FF8DFF8B2A5D45A58B26FE5C" bold="true" box="[111,327,477,498]" pageId="1" pageNumber="52">
<emphasis id="04A10E30FF8DFF8B2A5D45A58B4CFE5C" bold="true" box="[111,301,477,498]" italics="true" pageId="1" pageNumber="52">Material examined</emphasis>
. –
</emphasis>
<materialsCitation id="86BDD87FFF8DFF8B2B6245A58B39FD85" collectingDate="2004-12-05" collectionCode="NMCR" collectorName="E. Husana" latitude="11.999233" location="Bat Cave" longLatPrecision="1" longitude="121.924034" municipality="Boracay Island" pageId="1" pageNumber="52" specimenCode="NMCR 27063" specimenCount="1" typeStatus="holotype">
<typeStatus id="E96E6C80FF8DFF8B2B6245A58BCCFE5C" box="[336,429,477,498]" pageId="1" pageNumber="52" type="holotype">Holotype</typeStatus>
: male (23.8 × 
<quantity id="F12D7FC7FF8DFF8B287245A588FAFE5C" box="[576,667,477,498]" metricMagnitude="-2" metricUnit="m" metricValue="3.02" pageId="1" pageNumber="52" unit="mm" value="30.2">30.2 mm</quantity>
, 
<specimenCode id="66737A59FF8DFF8B289545A58ACFFDA1" collectionCode="NMCR" country="USA" lsid="urn:lsid:biocol.org:col:14936" name="New Mexico State University" pageId="1" pageNumber="52" type="Herbarium">NMCR 27063</specimenCode>
), 
<location id="330A84F9FF8DFF8B2A8F45828B76FDA1" LSID="urn:lsid:plazi:treatment:BE7C6334FF8DFF8E2AE445068B37FB1B:330A84F9FF8DFF8B2A8F45828B76FDA1" box="[189,279,506,527]" latitude="11.999233" longLatPrecision="1" longitude="121.924034" municipality="Boracay Island" name="Bat Cave" pageId="1" pageNumber="52">Bat Cave</location>
, 
<collectingMunicipality id="D60E4858FF8DFF8B2B2D45828BD0FDA1" box="[287,433,506,527]" pageId="1" pageNumber="52">Boracay Island</collectingMunicipality>
, 
<geoCoordinate id="53E1B4E5FF8DFF8B2B8B45828853FDA1" box="[441,562,506,527]" degrees="11" direction="north" minutes="59.954" orientation="latitude" pageId="1" pageNumber="52" precision="1" value="11.999233">11°59.954'N</geoCoordinate>
<geoCoordinate id="53E1B4E5FF8DFF8B2805458288DAFDA1" box="[567,699,506,527]" degrees="121" direction="east" minutes="55.442" orientation="longitude" pageId="1" pageNumber="52" precision="1" value="121.924034">121°55.442'E</geoCoordinate>
, coll. 
<collectorName id="9B20B7F4FF8DFF8B2A5D466E8AB4FD85" box="[111,213,534,555]" pageId="1" pageNumber="52">E. Husana</collectorName>
, 
<date id="426BF4E2FF8DFF8B2AD3466E8B35FD85" box="[225,340,534,555]" pageId="1" pageNumber="52" value="2004-12-05">
<collectingDate id="522F0D0AFF8DFF8B2AD3466E8B35FD85" box="[225,340,534,555]" pageId="1" pageNumber="52" value="2004-12-05">5 Dec.2004</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="366AD222FF8DFF8B2A5D46288B5DFB9C" blockId="1.[111,750,592,1074]" pageId="1" pageNumber="52">
<typeStatus id="E96E6C80FF8DFF8B2A5D46288AB4FDCB" box="[111,213,592,613]" pageId="1" pageNumber="52" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="20D319ABFF8DFF8B2AD046288B49FDCB" box="[226,296,592,613]" pageId="1" pageNumber="52" type="male">1 male</specimenCount>
(22.1 × 
<quantity id="F12D7FC7FF8DFF8B2BB846288B88FDCB" box="[394,489,592,613]" metricMagnitude="-2" metricUnit="m" metricValue="2.77" pageId="1" pageNumber="52" unit="mm" value="27.7">27.7 mm</quantity>
, NSMT-Cr2007.0105), 
<materialsCitation id="86BDD87FFF8DFF8B28D346288866FD2C" collectionCode="NMCR" pageId="1" pageNumber="52" specimenCode="NMCR 27064" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="20D319ABFF8DFF8B28D346288AD1FD2C" pageId="1" pageNumber="52" type="female">1 female</specimenCount>
(25.2 × 
<quantity id="F12D7FC7FF8DFF8B2B3846158B04FD2C" box="[266,357,621,642]" metricMagnitude="-2" metricUnit="m" metricValue="3.11" pageId="1" pageNumber="52" unit="mm" value="31.1">31.1 mm</quantity>
, 
<specimenCode id="66737A59FF8DFF8B2B4346158861FD2C" box="[369,512,621,642]" collectionCode="NMCR" country="USA" lsid="urn:lsid:biocol.org:col:14936" name="New Mexico State University" pageId="1" pageNumber="52" type="Herbarium">NMCR 27064</specimenCode>
)
</materialsCitation>
, 
<specimenCount id="20D319ABFF8DFF8B282146158808FD2C" box="[531,617,621,642]" pageId="1" pageNumber="52" type="female">1 female</specimenCount>
(18.9 × 24.0 mm, NSMT-Cr2007.0107), 
<specimenCount id="20D319ABFF8DFF8B2BBD46F28BB4FD31" box="[399,469,650,671]" pageId="1" pageNumber="52" type="male">1 male</specimenCount>
(19.8 × 
<quantity id="F12D7FC7FF8DFF8B280746F288F2FD31" box="[565,659,650,671]" metricMagnitude="-2" metricUnit="m" metricValue="2.46" pageId="1" pageNumber="52" unit="mm" value="24.6">24.6 mm</quantity>
, NSMT- Cr2007.0108), 
<specimenCount id="20D319ABFF8DFF8B2B3146DE8B27FD15" box="[259,326,678,699]" pageId="1" pageNumber="52" type="male">1 male</specimenCount>
(20.6 × 26.0 mm, NSMT-Cr2007.0109), 
<specimenCount id="20D319ABFF8DFF8B28D346DE8ADDFD76" pageId="1" pageNumber="52" type="juvenile">1 juvenile</specimenCount>
male (13.0 × 17.0 mm, NSMT-Cr2007.0110), 
<specimenCount id="20D319ABFF8DFF8B28BE46BB888CFD76" box="[652,749,707,728]" pageId="1" pageNumber="52" type="juvenile">1 juvenile</specimenCount>
male (11.4 × 
<quantity id="F12D7FC7FF8DFF8B2AC046988B2DFD5B" box="[242,332,736,757]" metricMagnitude="-2" metricUnit="m" metricValue="1.44" pageId="1" pageNumber="52" unit="mm" value="14.4">14.4 mm</quantity>
), 
<specimenCount id="20D319ABFF8DFF8B2B6C46988BC1FD5B" box="[350,416,736,757]" pageId="1" pageNumber="52" type="male">1 male</specimenCount>
(19.2 × 24.0 mm), 
<materialsCitation id="86BDD87FFF8DFF8B285246988A9EFCE5" collectingDate="2004-01-29" collectionCode="T" collectorName="E. Husana &amp; Kase &amp; S. Kinjo" latitude="7.0121665" location="Tagbaobo Cave" longLatPrecision="1" longitude="125.78017" municipality="Samal Island" pageId="1" pageNumber="52" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="20D319ABFF8DFF8B2852469888D5FD5B" box="[608,692,736,757]" pageId="1" pageNumber="52" type="female">1 female</specimenCount>
(20.7 × 
<quantity id="F12D7FC7FF8DFF8B2AB046858AB9FCBC" box="[130,216,765,786]" metricMagnitude="-2" metricUnit="m" metricValue="2.57" pageId="1" pageNumber="52" unit="mm" value="25.7">25.7 mm</quantity>
) (NSMT-Cr2007.0111), 
<location id="330A84F9FF8DFF8B2BFB46858803FCBC" LSID="urn:lsid:plazi:treatment:BE7C6334FF8DFF8E2AE445068B37FB1B:330A84F9FF8DFF8B2BFB46858803FCBC" box="[457,610,765,786]" latitude="7.0121665" longLatPrecision="1" longitude="125.78017" municipality="Samal Island" name="Tagbaobo Cave" pageId="1" pageNumber="52">Tagbaobo Cave</location>
, 
<collectingMunicipality id="D60E4858FF8DFF8B28594685888BFCBC" box="[619,746,765,786]" pageId="1" pageNumber="52">Samal Island</collectingMunicipality>
, 
<geoCoordinate id="53E1B4E5FF8DFF8B2A5D47628ABDFC81" box="[111,220,794,815]" degrees="7" direction="north" minutes="00.730" orientation="latitude" pageId="1" pageNumber="52" precision="1" value="7.0121665">7°00.730'N</geoCoordinate>
<geoCoordinate id="53E1B4E5FF8DFF8B2AD047628B06FC81" box="[226,359,794,815]" degrees="125" direction="east" minutes="46.810" orientation="longitude" pageId="1" pageNumber="52" precision="1" value="125.78017">125°46.810'E</geoCoordinate>
, coll. 
<collectorName id="9B20B7F4FF8DFF8B2B9347628867FC81" box="[417,518,794,815]" pageId="1" pageNumber="52">E. Husana</collectorName>
, 
<collectionCode id="50C44AE7FF8DFF8B282247628840FC81" box="[528,545,794,815]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="52">T</collectionCode>
. 
<collectorName id="9B20B7F4FF8DFF8B28184762883BFC81" box="[554,602,794,815]" pageId="1" pageNumber="52">Kase</collectorName>
&amp; 
<collectorName id="9B20B7F4FF8DFF8B284B476288ADFC81" box="[633,716,794,815]" pageId="1" pageNumber="52">S. Kinjo</collectorName>
, 28 &amp; 
<date id="426BF4E2FF8DFF8B2ABA474E8A9EFCE5" box="[136,255,822,843]" pageId="1" pageNumber="52" value="2004-01-29">
<collectingDate id="522F0D0AFF8DFF8B2ABA474E8A9EFCE5" box="[136,255,822,843]" pageId="1" pageNumber="52" value="2004-01-29">29 Jan.2004</collectingDate>
</date>
</materialsCitation>
; 
<specimenCount id="20D319ABFF8DFF8B2B3B474E8B3DFCE5" box="[265,348,822,843]" pageId="1" pageNumber="52" type="female">1 female</specimenCount>
(22.6 × 
<quantity id="F12D7FC7FF8DFF8B2B9D474E886CFCE5" box="[431,525,822,843]" metricMagnitude="-2" metricUnit="m" metricValue="2.92" pageId="1" pageNumber="52" unit="mm" value="29.2">29. 2 mm</quantity>
, NSMT-Cr2007.0096), 
<specimenCount id="20D319ABFF8DFF8B2A5D472B8AD0FCC6" box="[111,177,851,872]" pageId="1" pageNumber="52" type="male">1 male</specimenCount>
(28.3 × 
<quantity id="F12D7FC7FF8DFF8B2B35472B8B00FCC6" box="[263,353,851,872]" metricMagnitude="-2" metricUnit="m" metricValue="3.61" pageId="1" pageNumber="52" unit="mm" value="36.1">36.1 mm</quantity>
, NSMT-Cr2007.0097), 
<specimenCount id="20D319ABFF8DFF8B2878472B88FFFCC6" box="[586,670,851,872]" pageId="1" pageNumber="52" type="female">1 female</specimenCount>
(29.3 × 
<quantity id="F12D7FC7FF8DFF8B2A5D47088AA4FC2B" box="[111,197,880,901]" metricMagnitude="-2" metricUnit="m" metricValue="3.6799999999999997" pageId="1" pageNumber="52" unit="mm" value="36.8">36.8 mm</quantity>
, NSMT-Cr2007.0098), 
<specimenCount id="20D319ABFF8DFF8B2B9447088B99FC2B" box="[422,504,880,901]" pageId="1" pageNumber="52" type="female">1 female</specimenCount>
(24.1 × 
<quantity id="F12D7FC7FF8DFF8B2877470888FAFC2B" box="[581,667,880,901]" metricMagnitude="-2" metricUnit="m" metricValue="3.06" pageId="1" pageNumber="52" unit="mm" value="30.6">30.6 mm</quantity>
, NSMT- Cr2007.0099), 
<specimenCount id="20D319ABFF8DFF8B2B3147F58B27FC0C" box="[259,326,909,930]" pageId="1" pageNumber="52" type="male">1 male</specimenCount>
(25.7 × 
<quantity id="F12D7FC7FF8DFF8B2BAF47F58B96FC0C" box="[413,503,909,930]" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="1" pageNumber="52" unit="mm" value="31.8">31.8 mm</quantity>
, NSMT-Cr2007.0100), 
<specimenCount id="20D319ABFF8DFF8B28D347F58AFFFC11" pageId="1" pageNumber="52" type="male">1 male</specimenCount>
(25.0 × 
<quantity id="F12D7FC7FF8DFF8B2ACB47D28B34FC11" box="[249,341,938,959]" metricMagnitude="-2" metricUnit="m" metricValue="3.12" pageId="1" pageNumber="52" unit="mm" value="31.2">31.2 mm</quantity>
, NSMT-Cr2007.0101), 
<specimenCount id="20D319ABFF8DFF8B287647D288FAFC11" box="[580,667,938,959]" pageId="1" pageNumber="52" type="female">1 female</specimenCount>
(30.8 × 
<quantity id="F12D7FC7FF8DFF8B2A5D47BE8AA4FC75" box="[111,197,966,987]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="1" pageNumber="52" unit="mm" value="37.5">37.5 mm</quantity>
, NSMT-Cr2007.0102), 
<specimenCount id="20D319ABFF8DFF8B2B9447BE8B99FC75" box="[422,504,966,987]" pageId="1" pageNumber="52" type="female">1 female</specimenCount>
(28.2 × 
<quantity id="F12D7FC7FF8DFF8B287747BE88FAFC75" box="[581,667,966,987]" metricMagnitude="-2" metricUnit="m" metricValue="3.47" pageId="1" pageNumber="52" unit="mm" value="34.7">34.7 mm</quantity>
, NSMT- Cr2007.0103), 
<materialsCitation id="86BDD87FFF8DFF8B2B31479B8B56FB9C" collectingDate="2004-12-05" collectorName="E. Husana" latitude="11.999233" location="Bat Cave" longLatPrecision="1" longitude="121.924034" municipality="Boracay Island" pageId="1" pageNumber="52" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="20D319ABFF8DFF8B2B31479B8B39FC56" box="[259,344,995,1016]" pageId="1" pageNumber="52" type="female">1 female</specimenCount>
(29.4 × 
<quantity id="F12D7FC7FF8DFF8B2B83479B886AFC56" box="[433,523,995,1016]" metricMagnitude="-2" metricUnit="m" metricValue="3.69" pageId="1" pageNumber="52" unit="mm" value="36.9">36.9 mm</quantity>
, NSMT-Cr2007.0104), 
<location id="330A84F9FF8DFF8B2A5D40788AAEFBBB" LSID="urn:lsid:plazi:treatment:BE7C6334FF8DFF8E2AE445068B37FB1B:330A84F9FF8DFF8B2A5D40788AAEFBBB" box="[111,207,1024,1045]" latitude="11.999233" longLatPrecision="1" longitude="121.924034" municipality="Boracay Island" name="Bat Cave" pageId="1" pageNumber="52">Bat Cave</location>
, 
<collectingMunicipality id="D60E4858FF8DFF8B2AEF40788B19FBBB" box="[221,376,1024,1045]" pageId="1" pageNumber="52">Boracay Island</collectingMunicipality>
, 
<geoCoordinate id="53E1B4E5FF8DFF8B2BB440788863FBBB" box="[390,514,1024,1045]" degrees="11" direction="north" minutes="59.954" orientation="latitude" pageId="1" pageNumber="52" precision="1" value="11.999233">11°59.954'N</geoCoordinate>
<geoCoordinate id="53E1B4E5FF8DFF8B283E407888F4FBBB" box="[524,661,1024,1045]" degrees="121" direction="east" minutes="55.442" orientation="longitude" pageId="1" pageNumber="52" precision="1" value="121.924034">121°55.442'E</geoCoordinate>
, coll. 
<collectorName id="9B20B7F4FF8DFF8B28EA40788AD8FB9C" pageId="1" pageNumber="52">E. Husana</collectorName>
, 
<date id="426BF4E2FF8DFF8B2AF640658B56FB9C" box="[196,311,1053,1074]" pageId="1" pageNumber="52" value="2004-12-05">
<collectingDate id="522F0D0AFF8DFF8B2AF640658B56FB9C" box="[196,311,1053,1074]" pageId="1" pageNumber="52" value="2004-12-05">5 Dec.2004</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="366AD222FF8DFF8B2A5D402E8A87FB55" blockId="1.[111,750,1110,1275]" pageId="1" pageNumber="52">
<emphasis id="04A10E30FF8DFF8B2A5D402E8B12FBC5" bold="true" box="[111,371,1110,1131]" pageId="1" pageNumber="52">
<emphasis id="04A10E30FF8DFF8B2A5D402E8B34FBC5" bold="true" box="[111,341,1110,1131]" italics="true" pageId="1" pageNumber="52">Comparative material</emphasis>
. –
</emphasis>
<taxonomicName id="F1D5A9A1FF8DFF8B2B4F402F8A86FB26" authority="(Ng, Guinot &amp; Iliffe, 1994)" baseAuthorityName="Ng, Guinot &amp; Iliffe" baseAuthorityYear="1994" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="ultrapes">
<emphasis id="04A10E30FF8DFF8B2B4F402F8829FBC2" box="[381,584,1111,1132]" italics="true" pageId="1" pageNumber="52">Karstarma ultrapes</emphasis>
(Ng, Guinot &amp; Iliffe, 1994)
</taxonomicName>
. 
<materialsCitation id="86BDD87FFF8DFF8B2AC0400B888BFB6C" collectingDate="1988-08-15" collectionCode="T" collectorName="M. Iliffe &amp; S. Sarbu" country="Solomon Islands" county="Nggela Pile Island" location="Mbetibula Cave" municipality="Florida Islands" pageId="1" pageNumber="52" specimenCode="MNHN-B24796" specimenCount="1" typeStatus="holotype">
<typeStatus id="E96E6C80FF8DFF8B2AC0400B8B2EFB26" box="[242,335,1139,1160]" pageId="1" pageNumber="52" type="holotype">Holotype</typeStatus>
: male (22.9 × 
<quantity id="F12D7FC7FF8DFF8B2BEC400B8856FB26" box="[478,567,1139,1160]" metricMagnitude="-2" metricUnit="m" metricValue="2.8899999999999997" pageId="1" pageNumber="52" unit="mm" value="28.9">28.9 mm</quantity>
, 
<specimenCode id="66737A59FF8DFF8B2870400B8885FB26" box="[578,740,1139,1160]" collectionCode="MNHN-B" pageId="1" pageNumber="52">MNHN-B24796</specimenCode>
), 
<location id="330A84F9FF8DFF8B2A5D40E88B70FB0B" LSID="urn:lsid:plazi:treatment:BE7C6334FF8DFF8E2AE445068B37FB1B:330A84F9FF8DFF8B2A5D40E88B70FB0B" box="[111,273,1168,1189]" country="Solomon Islands" county="Nggela Pile Island" municipality="Florida Islands" name="Mbetibula Cave" pageId="1" pageNumber="52">Mbetibula Cave</location>
(station 88-083), 
<collectingMunicipality id="D60E4858FF8DFF8B2BFC40E88809FB0B" box="[462,616,1168,1189]" pageId="1" pageNumber="52">Florida Islands</collectingMunicipality>
, 
<collectingCounty id="DF0BAAAEFF8DFF8B284440E88ACDFB6C" pageId="1" pageNumber="52">Nggela Pile Island</collectingCounty>
, 
<collectingCountry id="4EC292B2FF8DFF8B2A8640D58B34FB6C" box="[180,341,1197,1218]" name="Solomon Islands" pageId="1" pageNumber="52">Solomon Islands</collectingCountry>
, coll. 
<collectionCode id="50C44AE7FF8DFF8B2BBE40D58BFCFB6C" box="[396,413,1197,1218]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="52">T</collectionCode>
. 
<collectorName id="9B20B7F4FF8DFF8B2B9740D58B94FB6C" box="[421,501,1197,1218]" pageId="1" pageNumber="52">M. Iliffe</collectorName>
&amp; 
<collectorName id="9B20B7F4FF8DFF8B282340D58802FB6C" box="[529,611,1197,1218]" pageId="1" pageNumber="52">S. Sarbu</collectorName>
, 
<date id="426BF4E2FF8DFF8B285940D5888BFB6C" box="[619,746,1197,1218]" pageId="1" pageNumber="52" value="1988-08-15">
<collectingDate id="522F0D0AFF8DFF8B285940D5888BFB6C" box="[619,746,1197,1218]" pageId="1" pageNumber="52" value="1988-08-15">15 Aug.1988</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="86BDD87FFF8DFF8B2A5D40B28A80FB55" accessionNumber="B24797" collectingDate="1988-08-15" collectionCode="T" collectorName="M. Iliffe &amp; S. Sarbu" country="Solomon Islands" county="Nggela Pile Island" location="Mbetibula Cave" municipality="Florida Islands" pageId="1" pageNumber="52" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="E96E6C80FF8DFF8B2A5D40B28AA6FB71" box="[111,199,1226,1247]" pageId="1" pageNumber="52" type="paratype">Paratype</typeStatus>
: 
<specimenCount id="20D319ABFF8DFF8B2AFD40B28B42FB71" box="[207,291,1226,1247]" pageId="1" pageNumber="52" type="female">1 female</specimenCount>
(32.5 × 
<quantity id="F12D7FC7FF8DFF8B2B4740B28BACFB71" box="[373,461,1226,1247]" metricMagnitude="-2" metricUnit="m" metricValue="3.94" pageId="1" pageNumber="52" unit="mm" value="39.4">39.4 mm</quantity>
, MNHN- B24797), data same as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="366AD222FF8DFF8B291844C6888CFA12" blockId="1.[810,1450,190,1109]" lastBlockId="1.[111,750,1316,1468]" pageId="1" pageNumber="52">
furrow present along median epi- and protogastric regions, diverging on meso- and metagastric regions and surrounding gastro-cardiac groove, with inward-curving setae along epi- and protogastric furrow, tips of setae partly interlocked especially in dorsal part; cervical furrow distinct but shallow on anterior part whereas indistinct on posterior part; gastrocardiac and cardio-intestinal furrows shallow; postorbital and epigastric protuberances slightly rugose. Frontal region gently granulose. Frontal margin deflexed, bilobed, medially separated by U-shaped concavity in dorsal view; downwardly inclined towards antennular septum in anterior view (
<figureCitation id="AEEECEA7FF8DFF8B2F4C45868927FD9B" captionStart="Fig" captionStartId="2.[134,169,2003,2024]" captionTargetBox="[142,1460,252,1964]" captionTargetId="figure-89@2.[118,1481,230,1979]" captionTargetPageId="2" captionText="Fig. 3. Karstarma philippinarum, new species. Paratype male NSMT-Cr2007.0105. a, carapace; b, abdomen and telson; c, cephalothorax, frontal view; d, left cheliped, dorsal view; e, left fourth ambulatory leg, dorsal view; f, left cheliped, carpus; g, left chela, outer view; h, left antenna, ventral view; i, G1, ventral and dorsal views; j, left third maxilliped, ventral view. Scale bars: a–e, g = 5.0 mm; f = 2.0 mm; h–j = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4509193" httpUri="https://zenodo.org/record/4509193/files/figure.png" pageId="1" pageNumber="52">Fig. 3c</figureCitation>
), ventral extensions of front embracing median tip of proepistome; supraorbital margin cristate, compressed Wshaped, submedian part gently produced; external orbital angle relatively produced, inner margin straight, base of inner margin concave; infraorbital margin medially broader, laterally reaching as far as below external orbital angle; suborbital crista present, lateral three-fifths thinly raised, forming keel, edge of keel lined with microscopical granules. Anterolateral and anterior half of posterolateral margin continuous, no clear demarcation, strongly diverging posteriorly, cristate, with 3 distinct teeth including external orbital angle, first and second teeth separated by deep, narrow fissure, third tooth separated from second tooth by shallow V-shaped notch, tip of second tooth slightly closer to that of first tooth than to third tooth, third tooth followed posteriorly by few small notches. Posterior half of posterolateral margin subcristate, converging abruptly towards gently sinuous posterior carapace margin. Epistome with posterior margin 
<emphasis id="04A10E30FF8DFF8B2A5D415D8B70FA92" bold="true" box="[111,273,1316,1341]" pageId="1" pageNumber="52">
<emphasis id="04A10E30FF8DFF8B2A5D415D8A93FA93" bold="true" box="[111,242,1317,1341]" italics="true" pageId="1" pageNumber="52">Description</emphasis>
. –
</emphasis>
Carapace (
<figureCitation id="AEEECEA7FF8DFF8B2BA7415D8873FA92" box="[405,530,1317,1340]" captionStart="Fig" captionStartId="1.[810,845,2060,2081]" captionTargetBox="[809,1448,1181,2042]" captionTargetId="figure-566@1.[809,1448,1181,2043]" captionTargetPageId="1" captionText="Fig. 2. Karstarma philippinarum, new species. Holotype male (23.8 × 30.2 mm, NMCR 27063). a, habitus; b, carapace, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4509191" httpUri="https://zenodo.org/record/4509191/files/figure.png" pageId="1" pageNumber="52">Figs. 2a, b</figureCitation>
, 
<figureCitation id="AEEECEA7FF8DFF8B2813415D885AFA92" box="[545,571,1317,1340]" captionStart="Fig" captionStartId="2.[134,169,2003,2024]" captionTargetBox="[142,1460,252,1964]" captionTargetId="figure-89@2.[118,1481,230,1979]" captionTargetPageId="2" captionText="Fig. 3. Karstarma philippinarum, new species. Paratype male NSMT-Cr2007.0105. a, carapace; b, abdomen and telson; c, cephalothorax, frontal view; d, left cheliped, dorsal view; e, left fourth ambulatory leg, dorsal view; f, left cheliped, carpus; g, left chela, outer view; h, left antenna, ventral view; i, G1, ventral and dorsal views; j, left third maxilliped, ventral view. Scale bars: a–e, g = 5.0 mm; f = 2.0 mm; h–j = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4509193" httpUri="https://zenodo.org/record/4509193/files/figure.png" pageId="1" pageNumber="52">3a</figureCitation>
) approximately trapezoidal, widest between bases of second and third ambulatory legs; dorsal surface distinctly rugose, especially on branchial regions, with scattered minute setae and short stiff black setae on entire surface. Deep longitudinal
</paragraph>
<caption id="62AA82AAFF8DFF8B2A424C7488DCF790" ID-DOI="http://doi.org/10.5281/zenodo.4509189" ID-Zenodo-Dep="4509189" httpUri="https://zenodo.org/record/4509189/files/figure.png" pageId="1" pageNumber="52" startId="1.[112,147,2060,2081]" targetBox="[111,750,1526,2043]" targetPageId="1">
<paragraph id="366AD222FF8DFF8B2A424C7488DCF790" blockId="1.[111,751,2060,2110]" pageId="1" pageNumber="52">
Fig. 1. Live coloration of 
<taxonomicName id="F1D5A9A1FF8DFF8B2B5C4C748806F78F" authority="Husana &amp; Naruse &amp; Kase, 2010" authorityName="Husana &amp; Naruse &amp; Kase" authorityYear="2010" box="[366,615,2060,2081]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="philippinarum" status="sp. nov.">
<emphasis id="04A10E30FF8DFF8B2B5C4C748806F78F" box="[366,615,2060,2081]" italics="true" pageId="1" pageNumber="52">Karstarma philippinarum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="1F92B34BFF8DFF8B28464C74888DF78F" box="[628,748,2060,2081]" pageId="1" pageNumber="52" rank="species">new species</taxonomicNameLabel>
, in its natural habitat, Bat Cave, Boracay Island, Philippines.
</paragraph>
</caption>
<caption id="62AA82AAFF8DFF8B29184C748FC4F790" ID-DOI="http://doi.org/10.5281/zenodo.4509191" ID-Zenodo-Dep="4509191" httpUri="https://zenodo.org/record/4509191/files/figure.png" pageId="1" pageNumber="52" startId="1.[810,845,2060,2081]" targetBox="[809,1448,1181,2042]" targetPageId="1">
<paragraph id="366AD222FF8DFF8B29184C748FC4F790" blockId="1.[810,1450,2060,2110]" pageId="1" pageNumber="52">
Fig. 2. 
<taxonomicName id="F1D5A9A1FF8DFF8B295E4C748E00F78F" authority="Husana &amp; Naruse &amp; Kase, 2010" authorityName="Husana &amp; Naruse &amp; Kase" authorityYear="2010" box="[876,1121,2060,2081]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="philippinarum" status="sp. nov.">
<emphasis id="04A10E30FF8DFF8B295E4C748E00F78F" box="[876,1121,2060,2081]" italics="true" pageId="1" pageNumber="52">Karstarma philippinarum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="1F92B34BFF8DFF8B2E594C748E81F78F" box="[1131,1248,2060,2081]" pageId="1" pageNumber="52" rank="species">new species</taxonomicNameLabel>
. Holotype male (23.8 × 30.2 mm, NMCR 27063). a, habitus; b, carapace, dorsal view.
</paragraph>
</caption>
<caption id="62AA82AAFF8EFF882AB443AB8B29F790" ID-DOI="http://doi.org/10.5281/zenodo.4509193" ID-Zenodo-Dep="4509193" httpUri="https://zenodo.org/record/4509193/files/figure.png" pageId="2" pageNumber="53" startId="2.[134,169,2003,2024]" targetBox="[142,1460,252,1964]" targetPageId="2">
<paragraph id="366AD222FF8EFF882AB443AB8B29F790" blockId="2.[134,1472,2003,2110]" pageId="2" pageNumber="53">
Fig. 3. 
<taxonomicName id="F1D5A9A1FF8EFF882AFE43AB8BA5F846" authority="Husana &amp; Naruse &amp; Kase, 2010" authorityName="Husana &amp; Naruse &amp; Kase" authorityYear="2010" box="[204,452,2003,2024]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="philippinarum" status="sp. nov.">
<emphasis id="04A10E30FF8EFF882AFE43AB8BA5F846" box="[204,452,2003,2024]" italics="true" pageId="2" pageNumber="53">Karstarma philippinarum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="1F92B34BFF8EFF882BE343AB8829F846" box="[465,584,2003,2024]" pageId="2" pageNumber="53" rank="species">new species</taxonomicNameLabel>
. Paratype male NSMT-Cr2007.0105. a, carapace; b, abdomen and telson; c, cephalothorax, frontal view; d, left cheliped, dorsal view; e, left fourth ambulatory leg, dorsal view; f, left cheliped, carpus; g, left chela, outer view; h, left antenna, ventral view; i, G1, ventral and dorsal views; j, left third maxilliped, ventral view. Scale bars: a–e, g = 5.0 mm; f = 2.0 mm; h–j = 1.0 mm.
</paragraph>
</caption>
<paragraph id="366AD222FF8FFF892A5D44C68ABFFF5B" blockId="3.[111,750,190,245]" pageId="3" pageNumber="54">cristate, bent anteriorly, trilobed, median projection longest, triangular.</paragraph>
<paragraph id="366AD222FF8FFF892A5D45668824FE1B" blockId="3.[111,750,286,437]" pageId="3" pageNumber="54">
Eyes (
<figureCitation id="AEEECEA7FF8FFF892A8745668B43FE9B" box="[181,290,286,309]" captionStart="Fig" captionStartId="2.[134,169,2003,2024]" captionTargetBox="[142,1460,252,1964]" captionTargetId="figure-89@2.[118,1481,230,1979]" captionTargetPageId="2" captionText="Fig. 3. Karstarma philippinarum, new species. Paratype male NSMT-Cr2007.0105. a, carapace; b, abdomen and telson; c, cephalothorax, frontal view; d, left cheliped, dorsal view; e, left fourth ambulatory leg, dorsal view; f, left cheliped, carpus; g, left chela, outer view; h, left antenna, ventral view; i, G1, ventral and dorsal views; j, left third maxilliped, ventral view. Scale bars: a–e, g = 5.0 mm; f = 2.0 mm; h–j = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4509193" httpUri="https://zenodo.org/record/4509193/files/figure.png" pageId="3" pageNumber="54">Fig. 3a, c</figureCitation>
) well developed, well pigmented, cornea wider than peduncle in dorsal view. Third maxilliped (
<figureCitation id="AEEECEA7FF8FFF8928F145468AE4FEDB" captionStart="Fig" captionStartId="2.[134,169,2003,2024]" captionTargetBox="[142,1460,252,1964]" captionTargetId="figure-89@2.[118,1481,230,1979]" captionTargetPageId="2" captionText="Fig. 3. Karstarma philippinarum, new species. Paratype male NSMT-Cr2007.0105. a, carapace; b, abdomen and telson; c, cephalothorax, frontal view; d, left cheliped, dorsal view; e, left fourth ambulatory leg, dorsal view; f, left cheliped, carpus; g, left chela, outer view; h, left antenna, ventral view; i, G1, ventral and dorsal views; j, left third maxilliped, ventral view. Scale bars: a–e, g = 5.0 mm; f = 2.0 mm; h–j = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4509193" httpUri="https://zenodo.org/record/4509193/files/figure.png" pageId="3" pageNumber="54">Fig. 3j</figureCitation>
) elongated; ischium as long as merus, with oblique median groove; merus with crest adjacent to inner margin, lined with setae; exopod slender, with long flagellum.
</paragraph>
<paragraph id="366AD222FF8FFF892A5D45A68859FD9B" blockId="3.[111,750,478,565]" pageId="3" pageNumber="54">Thoracic sternites 2 and 3 fused, lateral margin strongly convex outwards, suture indistinct; suture between 3 and 4 visible, concave towards the anterior part.</paragraph>
<paragraph id="366AD222FF8FFF892A5D4626884CFB3B" blockId="3.[111,750,606,1173]" pageId="3" pageNumber="54">
Chelipeds (
<figureCitation id="AEEECEA7FF8FFF892AD446268B51FDDB" box="[230,304,606,629]" captionStart="Fig" captionStartId="1.[810,845,2060,2081]" captionTargetBox="[809,1448,1181,2042]" captionTargetId="figure-566@1.[809,1448,1181,2043]" captionTargetPageId="1" captionText="Fig. 2. Karstarma philippinarum, new species. Holotype male (23.8 × 30.2 mm, NMCR 27063). a, habitus; b, carapace, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4509191" httpUri="https://zenodo.org/record/4509191/files/figure.png" pageId="3" pageNumber="54">Fig. 2a</figureCitation>
) equal, more robust in males than females, merus (
<figureCitation id="AEEECEA7FF8FFF892AF446068B7AFD3B" box="[198,283,638,661]" captionStart="Fig" captionStartId="2.[134,169,2003,2024]" captionTargetBox="[142,1460,252,1964]" captionTargetId="figure-89@2.[118,1481,230,1979]" captionTargetPageId="2" captionText="Fig. 3. Karstarma philippinarum, new species. Paratype male NSMT-Cr2007.0105. a, carapace; b, abdomen and telson; c, cephalothorax, frontal view; d, left cheliped, dorsal view; e, left fourth ambulatory leg, dorsal view; f, left cheliped, carpus; g, left chela, outer view; h, left antenna, ventral view; i, G1, ventral and dorsal views; j, left third maxilliped, ventral view. Scale bars: a–e, g = 5.0 mm; f = 2.0 mm; h–j = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4509193" httpUri="https://zenodo.org/record/4509193/files/figure.png" pageId="3" pageNumber="54">Fig. 3d</figureCitation>
) triangular in cross-section, all margins granulated, inner surface with two longitudinal rows of black fine setae, subdistal part of ventral inner margin with short, longitudinal non-corneous raise; carpus (
<figureCitation id="AEEECEA7FF8FFF89285546A688D1FD5B" box="[615,688,734,757]" captionStart="Fig" captionStartId="2.[134,169,2003,2024]" captionTargetBox="[142,1460,252,1964]" captionTargetId="figure-89@2.[118,1481,230,1979]" captionTargetPageId="2" captionText="Fig. 3. Karstarma philippinarum, new species. Paratype male NSMT-Cr2007.0105. a, carapace; b, abdomen and telson; c, cephalothorax, frontal view; d, left cheliped, dorsal view; e, left fourth ambulatory leg, dorsal view; f, left cheliped, carpus; g, left chela, outer view; h, left antenna, ventral view; i, G1, ventral and dorsal views; j, left third maxilliped, ventral view. Scale bars: a–e, g = 5.0 mm; f = 2.0 mm; h–j = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4509193" httpUri="https://zenodo.org/record/4509193/files/figure.png" pageId="3" pageNumber="54">Fig. 3f</figureCitation>
) with inner angle slightly foliaceous, fringed with small teeth, outer margin with several low teeth; chela (
<figureCitation id="AEEECEA7FF8FFF892877476688CEFC9B" box="[581,687,798,821]" captionStart="Fig" captionStartId="2.[134,169,2003,2024]" captionTargetBox="[142,1460,252,1964]" captionTargetId="figure-89@2.[118,1481,230,1979]" captionTargetPageId="2" captionText="Fig. 3. Karstarma philippinarum, new species. Paratype male NSMT-Cr2007.0105. a, carapace; b, abdomen and telson; c, cephalothorax, frontal view; d, left cheliped, dorsal view; e, left fourth ambulatory leg, dorsal view; f, left cheliped, carpus; g, left chela, outer view; h, left antenna, ventral view; i, G1, ventral and dorsal views; j, left third maxilliped, ventral view. Scale bars: a–e, g = 5.0 mm; f = 2.0 mm; h–j = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4509193" httpUri="https://zenodo.org/record/4509193/files/figure.png" pageId="3" pageNumber="54">Fig. 3d, g</figureCitation>
) with slightly inflated palm, palm sparsely granulated on lower half; movable finger slightly shorter than palm, immovable finger with 2 median longitudinal ridges along outer edge, cutting edges lined with numerous triangular teeth. Ambulatory legs (
<figureCitation id="AEEECEA7FF8FFF892A9F47C68A9BFC7B" box="[173,250,958,981]" captionStart="Fig" captionStartId="1.[112,147,2060,2081]" captionTargetBox="[111,750,1526,2043]" captionTargetId="figure-543@1.[111,750,1526,2043]" captionTargetPageId="1" captionText="Fig. 1. Live coloration of Karstarma philippinarum, new species, in its natural habitat, Bat Cave, Boracay Island, Philippines." figureDoi="http://doi.org/10.5281/zenodo.4509189" httpUri="https://zenodo.org/record/4509189/files/figure.png" pageId="3" pageNumber="54">Figs. 1</figureCitation>
, 
<figureCitation id="AEEECEA7FF8FFF892B3947C68B46FC7B" box="[267,295,958,981]" captionStart="Fig" captionStartId="1.[810,845,2060,2081]" captionTargetBox="[809,1448,1181,2042]" captionTargetId="figure-566@1.[809,1448,1181,2043]" captionTargetPageId="1" captionText="Fig. 2. Karstarma philippinarum, new species. Holotype male (23.8 × 30.2 mm, NMCR 27063). a, habitus; b, carapace, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4509191" httpUri="https://zenodo.org/record/4509191/files/figure.png" pageId="3" pageNumber="54">2a</figureCitation>
, 
<figureCitation id="AEEECEA7FF8FFF892B0747C68B2EFC7B" box="[309,335,958,981]" captionStart="Fig" captionStartId="2.[134,169,2003,2024]" captionTargetBox="[142,1460,252,1964]" captionTargetId="figure-89@2.[118,1481,230,1979]" captionTargetPageId="2" captionText="Fig. 3. Karstarma philippinarum, new species. Paratype male NSMT-Cr2007.0105. a, carapace; b, abdomen and telson; c, cephalothorax, frontal view; d, left cheliped, dorsal view; e, left fourth ambulatory leg, dorsal view; f, left cheliped, carpus; g, left chela, outer view; h, left antenna, ventral view; i, G1, ventral and dorsal views; j, left third maxilliped, ventral view. Scale bars: a–e, g = 5.0 mm; f = 2.0 mm; h–j = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4509193" httpUri="https://zenodo.org/record/4509193/files/figure.png" pageId="3" pageNumber="54">3e</figureCitation>
) very long, third pair longest, fourth pair shortest. Meri rugose on dorsal surfaces, merus of third ambulatory leg 3.83–5.12 times width (mean = 4.39, n = 14), 1.08–1.60 times CL (mean = 1.37, n = 14). Margins of propodi and dactyli lined with stiff and long black setae, inner and outer margins of dactyli of first and second ambulatory legs with mat of dense short black setae.
</paragraph>
</subSubSection>
<subSubSection id="7ECF81A9FF8FFF89291B44C68BC4F81B" pageId="3" pageNumber="54" type="discussion">
<paragraph id="366AD222FF8FFF89291B44C68986FC3B" blockId="3.[809,1450,190,917]" pageId="3" pageNumber="54">
<emphasis id="04A10E30FF8FFF89291B44C689C7FF78" bold="true" box="[809,934,190,214]" pageId="3" pageNumber="54">
<emphasis id="04A10E30FF8FFF89291B44C689EAFF78" bold="true" box="[809,907,190,214]" italics="true" pageId="3" pageNumber="54">Remarks</emphasis>
. –
</emphasis>
<taxonomicName id="F1D5A9A1FF8FFF89299F44C78EA0FF78" authority="Husana &amp; Naruse &amp; Kase, 2010" authorityName="Husana &amp; Naruse &amp; Kase" authorityYear="2010" box="[941,1217,191,214]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="philippinarum" status="sp. nov.">
<emphasis id="04A10E30FF8FFF89299F44C78EA0FF78" box="[941,1217,191,214]" italics="true" pageId="3" pageNumber="54">Karstarma philippinarum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="1F92B34BFF8FFF892EFD44C68F32FF7B" box="[1231,1363,190,213]" pageId="3" pageNumber="54" rank="species">new species</taxonomicNameLabel>
, closely resembles 
<taxonomicName id="F1D5A9A1FF8FFF8929A844A78E71FF58" baseAuthorityName="Ng, Guinot &amp; Iliffe" baseAuthorityYear="1994" box="[922,1040,223,246]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="ultrapes">
<emphasis id="04A10E30FF8FFF8929A844A78E71FF58" box="[922,1040,223,246]" italics="true" pageId="3" pageNumber="54">K. ultrapes</emphasis>
</taxonomicName>
. They could easily be misidentified as the same species because of their similar body pigmentations and appearances. But obvious distinction between the two species could easily be noticed if they are compared side by side. The clear difference between them is the proportions of the ambulatory legs; 
<taxonomicName id="F1D5A9A1FF8FFF892E3745078ED9FE38" authorityName="Husana &amp; Naruse &amp; Kase" authorityYear="2010" box="[1029,1208,383,406]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="philippinarum">
<emphasis id="04A10E30FF8FFF892E3745078ED9FE38" box="[1029,1208,383,406]" italics="true" pageId="3" pageNumber="54">K. philippinarum</emphasis>
</taxonomicName>
has shorter ambulatory legs than those of 
<taxonomicName id="F1D5A9A1FF8FFF8929CA45E78E13FE18" baseAuthorityName="Ng, Guinot &amp; Iliffe" baseAuthorityYear="1994" box="[1016,1138,415,438]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="ultrapes">
<emphasis id="04A10E30FF8FFF8929CA45E78E13FE18" box="[1016,1138,415,438]" italics="true" pageId="3" pageNumber="54">K. ultrapes</emphasis>
</taxonomicName>
. For example, the merus of the third ambulatory leg of 
<taxonomicName id="F1D5A9A1FF8FFF892E6C45C78F78FE78" authorityName="Husana &amp; Naruse &amp; Kase" authorityYear="2010" box="[1118,1305,447,470]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="philippinarum">
<emphasis id="04A10E30FF8FFF892E6C45C78F78FE78" box="[1118,1305,447,470]" italics="true" pageId="3" pageNumber="54">K. philippinarum</emphasis>
</taxonomicName>
is 3.83–5.12 times width (mean = 4.39, n = 14) and 1.08–1.60 times CL (mean = 1.37, n = 14), while that of 
<taxonomicName id="F1D5A9A1FF8FFF892EE745878F32FDB8" baseAuthorityName="Ng, Guinot &amp; Iliffe" baseAuthorityYear="1994" box="[1237,1363,511,534]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="ultrapes">
<emphasis id="04A10E30FF8FFF892EE745878F32FDB8" box="[1237,1363,511,534]" italics="true" pageId="3" pageNumber="54">K. ultrapes</emphasis>
</taxonomicName>
is 5.45 and 5.47 times width (mean = 5.46, n = 2) and 1.40 and 1.66 times CL (mean = 1.53, n = 2) (also see 
<figureCitation id="AEEECEA7FF8FFF892F1146468F1AFDFB" box="[1315,1403,574,597]" captionStart="Fig" captionStartId="1.[810,845,2060,2081]" captionTargetBox="[809,1448,1181,2042]" captionTargetId="figure-566@1.[809,1448,1181,2043]" captionTargetPageId="1" captionText="Fig. 2. Karstarma philippinarum, new species. Holotype male (23.8 × 30.2 mm, NMCR 27063). a, habitus; b, carapace, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4509191" httpUri="https://zenodo.org/record/4509191/files/figure.png" pageId="3" pageNumber="54">Figs. 2a</figureCitation>
, 
<figureCitation id="AEEECEA7FF8FFF892FBA46468FC5FDFB" box="[1416,1444,574,597]" captionStart="Fig" captionStartId="3.[809,844,2032,2053]" captionTargetBox="[809,1448,1124,2013]" captionTargetId="figure-761@3.[809,1448,1124,2013]" captionTargetPageId="3" captionText="Fig. 4. Karstarma ultrapes (Ng, Guinot &amp; Iliffe, 1994). Holotype male (22.9 × 28.9 mm, MNHN-B24796). a, habitus; b, carapace, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4509195" httpUri="https://zenodo.org/record/4509195/files/figure.png" pageId="3" pageNumber="54">4a</figureCitation>
; 
<bibRefCitation id="5244AFD3FF8FFF89291B462689B4FDDB" author="Ng, P" box="[809,981,606,629]" pageId="3" pageNumber="54" pagination="12 - 22" refId="ref3434" refString="Ng, P. K. L., D. Guinot &amp; T. M. Iliffe, 1994. Sesarmoides ultrapes new species, a remarkable sesarmine crab from caves in the Solomon Islands (Decapoda: Brachyura: Grapsidae). Crustacean Research, 23: 12 - 22." type="journal article" year="1994">Ng et al., 1994</bibRefCitation>
: fig. 5). In addition, 
<taxonomicName id="F1D5A9A1FF8FFF892EF246278F1AFDD8" authorityName="Husana &amp; Naruse &amp; Kase" authorityYear="2010" box="[1216,1403,607,630]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="philippinarum">
<emphasis id="04A10E30FF8FFF892EF246278F1AFDD8" box="[1216,1403,607,630]" italics="true" pageId="3" pageNumber="54">K. philippinarum</emphasis>
</taxonomicName>
has blunter external orbital angle and larger projection on the supraorbital margin. In 
<taxonomicName id="F1D5A9A1FF8FFF892E1946E78E84FD18" authorityName="Husana &amp; Naruse &amp; Kase" authorityYear="2010" box="[1067,1253,671,694]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="philippinarum">
<emphasis id="04A10E30FF8FFF892E1946E78E84FD18" box="[1067,1253,671,694]" italics="true" pageId="3" pageNumber="54">K. philippinarum</emphasis>
</taxonomicName>
, the inner margin of the external orbital angle is convex and the submedian convexity of the supraorbital margin is more produced, forming a rounded concavity between them. While in 
<taxonomicName id="F1D5A9A1FF8FFF892FA2468789E1FC98" baseAuthorityName="Ng, Guinot &amp; Iliffe" baseAuthorityYear="1994" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="ultrapes">
<emphasis id="04A10E30FF8FFF892FA2468789E1FC98" italics="true" pageId="3" pageNumber="54">K. ultrapes</emphasis>
</taxonomicName>
, the inner margin of the angle is gradually confluent with the supraorbital margin, the submedian convexity of the supraorbital margin is lower (
<figureCitation id="AEEECEA7FF8FFF892EA247268E8BFCDB" box="[1168,1258,862,885]" captionStart="Fig" captionStartId="1.[810,845,2060,2081]" captionTargetBox="[809,1448,1181,2042]" captionTargetId="figure-566@1.[809,1448,1181,2043]" captionTargetPageId="1" captionText="Fig. 2. Karstarma philippinarum, new species. Holotype male (23.8 × 30.2 mm, NMCR 27063). a, habitus; b, carapace, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4509191" httpUri="https://zenodo.org/record/4509191/files/figure.png" pageId="3" pageNumber="54">Figs. 2b</figureCitation>
, 
<figureCitation id="AEEECEA7FF8FFF892EC747268F70FCDB" box="[1269,1297,862,885]" captionStart="Fig" captionStartId="2.[134,169,2003,2024]" captionTargetBox="[142,1460,252,1964]" captionTargetId="figure-89@2.[118,1481,230,1979]" captionTargetPageId="2" captionText="Fig. 3. Karstarma philippinarum, new species. Paratype male NSMT-Cr2007.0105. a, carapace; b, abdomen and telson; c, cephalothorax, frontal view; d, left cheliped, dorsal view; e, left fourth ambulatory leg, dorsal view; f, left cheliped, carpus; g, left chela, outer view; h, left antenna, ventral view; i, G1, ventral and dorsal views; j, left third maxilliped, ventral view. Scale bars: a–e, g = 5.0 mm; f = 2.0 mm; h–j = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4509193" httpUri="https://zenodo.org/record/4509193/files/figure.png" pageId="3" pageNumber="54">3a</figureCitation>
, 
<figureCitation id="AEEECEA7FF8FFF892F2E47268F5BFCDB" box="[1308,1338,862,885]" captionStart="Fig" captionStartId="3.[809,844,2032,2053]" captionTargetBox="[809,1448,1124,2013]" captionTargetId="figure-761@3.[809,1448,1124,2013]" captionTargetPageId="3" captionText="Fig. 4. Karstarma ultrapes (Ng, Guinot &amp; Iliffe, 1994). Holotype male (22.9 × 28.9 mm, MNHN-B24796). a, habitus; b, carapace, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4509195" httpUri="https://zenodo.org/record/4509195/files/figure.png" pageId="3" pageNumber="54">4b</figureCitation>
; 
<bibRefCitation id="5244AFD3FF8FFF892F7447268900FC3B" author="Ng, P" pageId="3" pageNumber="54" pagination="12 - 22" refId="ref3434" refString="Ng, P. K. L., D. Guinot &amp; T. M. Iliffe, 1994. Sesarmoides ultrapes new species, a remarkable sesarmine crab from caves in the Solomon Islands (Decapoda: Brachyura: Grapsidae). Crustacean Research, 23: 12 - 22." type="journal article" year="1994">Ng et al., 1994</bibRefCitation>
, figs. 5–7a).
</paragraph>
<paragraph id="366AD222FF8FFF89291B47C68B77F9FB" blockId="3.[809,1449,958,1077]" lastBlockId="3.[111,750,1214,1621]" pageId="3" pageNumber="54">
When Davie &amp; Ng (2007) established 
<taxonomicName id="F1D5A9A1FF8FFF892EF147C78F56FC78" authorityName="Davie &amp; Ng" authorityYear="2007" box="[1219,1335,959,982]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="04A10E30FF8FFF892EF147C78F56FC78" box="[1219,1335,959,982]" italics="true" pageId="3" pageNumber="54">Karstarma</emphasis>
</taxonomicName>
, they used two spellings, “ 
<taxonomicName id="F1D5A9A1FF8FFF8929E147A78E26FC58" authorityName="Davie &amp; Ng" authorityYear="2007" box="[979,1095,991,1014]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="04A10E30FF8FFF8929E147A78E26FC58" box="[979,1095,991,1014]" italics="true" pageId="3" pageNumber="54">Karstarma</emphasis>
</taxonomicName>
” and “ 
<taxonomicName id="F1D5A9A1FF8FFF892EA447A78E9EFC58" authorityName="N.g.Davie" authorityYear="2007" box="[1174,1279,991,1014]" class="Malacostraca" family="Sesarmidae" genus="Karstama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="04A10E30FF8FFF892EA447A78E9EFC58" box="[1174,1279,991,1014]" italics="true" pageId="3" pageNumber="54">Karstama</emphasis>
</taxonomicName>
”, for the genus name. Although Davie &amp; Ng (2007: 229) clearly stated that the genus name is derived from an arbitrary combination Abdominal somites segments, all freely segmented. Male abdomen (
<figureCitation id="AEEECEA7FF8FFF892AD340A68B50FB5B" box="[225,305,1246,1269]" captionStart="Fig" captionStartId="2.[134,169,2003,2024]" captionTargetBox="[142,1460,252,1964]" captionTargetId="figure-89@2.[118,1481,230,1979]" captionTargetPageId="2" captionText="Fig. 3. Karstarma philippinarum, new species. Paratype male NSMT-Cr2007.0105. a, carapace; b, abdomen and telson; c, cephalothorax, frontal view; d, left cheliped, dorsal view; e, left fourth ambulatory leg, dorsal view; f, left cheliped, carpus; g, left chela, outer view; h, left antenna, ventral view; i, G1, ventral and dorsal views; j, left third maxilliped, ventral view. Scale bars: a–e, g = 5.0 mm; f = 2.0 mm; h–j = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4509193" httpUri="https://zenodo.org/record/4509193/files/figure.png" pageId="3" pageNumber="54">Fig. 3b</figureCitation>
) triangular, wide; somites 1 and 2 short; lateral margins of somites 3 strongly convex; somites 4 and 5 trapezoidal in shape, lateral margins straight or gently sinuous; lateral margins of somites 6 convex; telson rounded on distal margin. Female abdomen rounded, very wide, somites 3 trapezoidal in shape; somites 4 to 5 with convex lateral margins, almost reaching proximal half of coxa; lateral margin of somites 6 convex; telson distinctly sunken into distal margin of somites 6. G1 (
<figureCitation id="AEEECEA7FF8FFF89285341A688CAFA5B" box="[609,683,1502,1525]" captionStart="Fig" captionStartId="2.[134,169,2003,2024]" captionTargetBox="[142,1460,252,1964]" captionTargetId="figure-89@2.[118,1481,230,1979]" captionTargetPageId="2" captionText="Fig. 3. Karstarma philippinarum, new species. Paratype male NSMT-Cr2007.0105. a, carapace; b, abdomen and telson; c, cephalothorax, frontal view; d, left cheliped, dorsal view; e, left fourth ambulatory leg, dorsal view; f, left cheliped, carpus; g, left chela, outer view; h, left antenna, ventral view; i, G1, ventral and dorsal views; j, left third maxilliped, ventral view. Scale bars: a–e, g = 5.0 mm; f = 2.0 mm; h–j = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4509193" httpUri="https://zenodo.org/record/4509193/files/figure.png" pageId="3" pageNumber="54">Fig. 3i</figureCitation>
) very stout, almost straight, outer margin and subdistal area covered with dense long setae, covering pectinated distal part; distal part spade-like.
</paragraph>
<paragraph id="366AD222FF8FFF892A5D42068BC4F81B" blockId="3.[111,750,1662,1973]" pageId="3" pageNumber="54">
<emphasis id="04A10E30FF8FFF892A5D42068842F938" box="[111,547,1662,1686]" italics="true" pageId="3" pageNumber="54">
<emphasis id="04A10E30FF8FFF892A5D42068B6CF938" bold="true" box="[111,269,1662,1686]" italics="true" pageId="3" pageNumber="54">Colouration. –</emphasis>
<taxonomicName id="F1D5A9A1FF8FFF892B2142078842F938" authority="Husana &amp; Naruse &amp; Kase, 2010" authorityName="Husana &amp; Naruse &amp; Kase" authorityYear="2010" box="[275,547,1663,1686]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="philippinarum" status="sp. nov.">Karstarma philippinarum</taxonomicName>
</emphasis>
, 
<taxonomicNameLabel id="1F92B34BFF8FFF89281D420688D1F93B" box="[559,688,1662,1685]" pageId="3" pageNumber="54" rank="species">new species</taxonomicNameLabel>
, has a semi-circular bright orange pattern on the anterior half of the dorsal surface of the carapace (
<figureCitation id="AEEECEA7FF8FFF892B8E42C68B9AF97B" box="[444,507,1726,1749]" captionStart="Fig" captionStartId="1.[112,147,2060,2081]" captionTargetBox="[111,750,1526,2043]" captionTargetId="figure-543@1.[111,750,1526,2043]" captionTargetPageId="1" captionText="Fig. 1. Live coloration of Karstarma philippinarum, new species, in its natural habitat, Bat Cave, Boracay Island, Philippines." figureDoi="http://doi.org/10.5281/zenodo.4509189" httpUri="https://zenodo.org/record/4509189/files/figure.png" pageId="3" pageNumber="54">Fig. 1</figureCitation>
). The posterior part of the dorsal surfaces of the carapace, including the intestinal region, is brownish in male live specimen. Females, on the other hand, have whitish intestinal region. The meri and carpi of the ambulatory legs are orange, and the propodi and dactyli are white. The chelipeds are orangish, except for whitish distal ends of the fingers in males and whitish palm and fingers in females.
</paragraph>
</subSubSection>
<subSubSection id="7ECF81A9FF8FFF8E2A5D43A688EEFB9B" lastPageId="4" lastPageNumber="55" pageId="3" pageNumber="54" type="etymology">
<paragraph id="366AD222FF8FFF892A5D43A68AB6F79B" blockId="3.[111,750,2014,2101]" pageId="3" pageNumber="54">
<emphasis id="04A10E30FF8FFF892A5D43A68A9CF858" bold="true" box="[111,253,2014,2038]" pageId="3" pageNumber="54">
<emphasis id="04A10E30FF8FFF892A5D43A68A85F858" bold="true" box="[111,228,2014,2038]" italics="true" pageId="3" pageNumber="54">Etymology</emphasis>
. –
</emphasis>
The species name, 
<taxonomicName id="F1D5A9A1FF8FFF892BF843A78802F858" authorityName="Husana &amp; Naruse &amp; Kase" authorityYear="2010" box="[458,611,2015,2038]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="philippinarum">
<emphasis id="04A10E30FF8FFF892BF843A78802F858" box="[458,611,2015,2038]" italics="true" pageId="3" pageNumber="54">philippinarum</emphasis>
</taxonomicName>
, was derived from the 
<collectingCountry id="4EC292B2FF8FFF892AEB43868B37F7BB" box="[217,342,2046,2069]" name="Philippines" pageId="3" pageNumber="54">Philippines</collectingCountry>
, where the present new species was collected.
</paragraph>
<caption id="62AA82AAFF8FFF89291B438889C3F791" ID-DOI="http://doi.org/10.5281/zenodo.4509195" ID-Zenodo-Dep="4509195" httpUri="https://zenodo.org/record/4509195/files/figure.png" pageId="3" pageNumber="54" startId="3.[809,844,2032,2053]" targetBox="[809,1448,1124,2013]" targetPageId="3">
<paragraph id="366AD222FF8FFF89291B438889C3F791" blockId="3.[809,1449,2032,2111]" pageId="3" pageNumber="54">
Fig. 4. 
<taxonomicName id="F1D5A9A1FF8FFF89294243888F25F7AB" authority="(Ng, Guinot &amp; Iliffe, 1994)" baseAuthorityName="Ng, Guinot &amp; Iliffe" baseAuthorityYear="1994" box="[880,1348,2032,2053]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="ultrapes">
<emphasis id="04A10E30FF8FFF89294243888E4EF7AB" box="[880,1071,2032,2053]" italics="true" pageId="3" pageNumber="54">Karstarma ultrapes</emphasis>
(Ng, Guinot &amp; Iliffe, 1994)
</taxonomicName>
. Holotype male (22.9 × 28.9 mm, MNHN-B24796). a, habitus; b, carapace, dorsal view.
</paragraph>
</caption>
<paragraph id="366AD222FF88FF8E2AB444C68857FE3B" blockId="4.[134,773,190,405]" pageId="4" pageNumber="55">
of “karst” and “ 
<taxonomicName id="F1D5A9A1FF88FF8E2B7044C78BC3FF78" authority="Ng, Guinot &amp; Davie (2008)" authorityName="Say" authorityYear="1817" box="[322,418,191,214]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="04A10E30FF88FF8E2B7044C78BC3FF78" box="[322,418,191,214]" italics="true" pageId="4" pageNumber="55">Sesarma</emphasis>
</taxonomicName>
”, Ng, Guinot &amp; Davie (2008) consistently spelled the genus as “ 
<taxonomicName id="F1D5A9A1FF88FF8E283644A7880CFF58" authorityName="N.g.Davie" authorityYear="2007" box="[516,621,223,246]" class="Malacostraca" family="Sesarmidae" genus="Karstama" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="04A10E30FF88FF8E283644A7880CFF58" box="[516,621,223,246]" italics="true" pageId="4" pageNumber="55">Karstama</emphasis>
</taxonomicName>
” without any comment. Indeed, the authors have not been aware of the presence of the second spelling (P. J. F. Davie &amp; P. K. L. Ng, pers. comm.). According to the ICZN Article 24.2.3, we select the spelling “ 
<taxonomicName id="F1D5A9A1FF88FF8E2BA145278868FED8" authorityName="Davie &amp; Ng" authorityYear="2007" box="[403,521,351,374]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="04A10E30FF88FF8E2BA145278868FED8" box="[403,521,351,374]" italics="true" pageId="4" pageNumber="55">Karstarma</emphasis>
</taxonomicName>
” as a correct original spelling (see also Wowor &amp; Ng, 2009).
</paragraph>
<paragraph id="366AD222FF88FF8E2AB445C688EEFB9B" blockId="4.[134,773,446,1077]" pageId="4" pageNumber="55">
<emphasis id="04A10E30FF88FF8E2AB445C68B37FE78" bold="true" box="[134,342,446,470]" pageId="4" pageNumber="55">
<emphasis id="04A10E30FF88FF8E2AB445C68B5AFE78" bold="true" box="[134,315,446,470]" italics="true" pageId="4" pageNumber="55">Notes on habitat</emphasis>
. –
</emphasis>
<taxonomicName id="F1D5A9A1FF88FF8E2B6F45C78811FE78" authority="Husana &amp; Naruse &amp; Kase, 2010" authorityName="Husana &amp; Naruse &amp; Kase" authorityYear="2010" box="[349,624,447,470]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="philippinarum" status="sp. nov.">
<emphasis id="04A10E30FF88FF8E2B6F45C78811FE78" box="[349,624,447,470]" italics="true" pageId="4" pageNumber="55">Karstarma philippinarum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="1F92B34BFF88FF8E284C45C68960FE7B" box="[638,769,446,469]" pageId="4" pageNumber="55" rank="species">new species</taxonomicNameLabel>
, was found in two anchialine caves that are filled with normal high saline seawater. Tagbaobo Cave is located in a remote area on the eastern side of Samal Island just south of 
<collectingRegion id="F4111CC0FF88FF8E288D46668964FD9B" box="[703,773,542,565]" country="Philippines" name="Davao" pageId="4" pageNumber="55">Davao</collectingRegion>
City in Mindanao. The entrance of this cave is located on a limestone sea cliff about four meters above the high tide line and inside is gloomy, with a direct underground and below sea-level connection to the sea. The animals co-occurred with 
<taxonomicName id="F1D5A9A1FF88FF8E2A8E46C78B12FD78" authorityName="Husana &amp; Naruse &amp; Kase" authorityYear="2010" box="[188,371,702,726]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="philippinarum">
<emphasis id="04A10E30FF88FF8E2A8E46C78AAFFD78" box="[188,206,703,726]" italics="true" pageId="4" pageNumber="55">K</emphasis>
. 
<emphasis id="04A10E30FF88FF8E2AE846C78B12FD78" box="[218,371,703,726]" italics="true" pageId="4" pageNumber="55">philippinarum</emphasis>
</taxonomicName>
in this cave are the hyppolitid shrimp 
<taxonomicName id="F1D5A9A1FF88FF8E2AB446A78B9EFD5B" authority="(Clark, 1989)" baseAuthorityName="Clark" baseAuthorityYear="1989" box="[134,511,734,758]" class="Malacostraca" family="Hippolytidae" genus="Parhippolyte" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="misticia">
<emphasis id="04A10E30FF88FF8E2AB446A78B08FD58" box="[134,361,735,758]" italics="true" pageId="4" pageNumber="55">Parhippolyte misticia</emphasis>
(
<bibRefCitation id="5244AFD3FF88FF8E2B4446A68B99FD5B" author="Clark, J" box="[374,504,734,757]" pageId="4" pageNumber="55" pagination="445 - 452" refId="ref3194" refString="Clark, J., 1989. Koror mysticius, new genus, new species (Decapoda: Hippolytidae), a cave shrimp from Palau. Journal of Crustacean Biology, 9: 445 - 452." type="journal article" year="1989">Clark, 1989</bibRefCitation>
)
</taxonomicName>
, undetermined ostracods and mysids. Bat Cave is in a limestone body and its entrance is about 
<quantity id="F12D7FC7FF88FF8E2AEC47668B41FC9B" box="[222,288,798,821]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="55" unit="m" value="100.0">100 m</quantity>
away from the sea. The cave extends seaward and appears to connect to the sea because dim light is visible from the water at the innermost part of the cave. Other crustaceans collected inside this cave are the gecarcinid crabs 
<taxonomicName id="F1D5A9A1FF88FF8E2AB447E78BBFFC1B" authority="Ng, 2002" authorityName="Ng" authorityYear="2002" box="[134,478,926,950]" class="Malacostraca" family="Gecarcinidae" genus="Discoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilipes">
<emphasis id="04A10E30FF88FF8E2AB447E78B0AFC18" box="[134,363,927,950]" italics="true" pageId="4" pageNumber="55">Discoplax gracilipes</emphasis>
<bibRefCitation id="5244AFD3FF88FF8E2B4447E68BBFFC1B" author="Ng, P" box="[374,478,926,949]" pageId="4" pageNumber="55" pagination="419 - 435" refId="ref3379" refString="Ng, P. K. L., 2002. New species of cavernicolous crabs of the genus Sesarmoides from the western Pacific, with a key to the genus (Crustacea: Decapoda: Brchyura: Sesarmidae). Raffles Bulltein of Zoology, 50 (2): 419 - 435." type="journal article" year="2002">Ng, 2002</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="F1D5A9A1FF88FF8E282847E78BC0FC7B" authority="H. Milne Edwards, 1837" authorityName="H. Milne Edwards" authorityYear="1837" class="Malacostraca" family="Gecarcinidae" genus="Gecarcoidea" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="lalandii">
<emphasis id="04A10E30FF88FF8E282847E78965FC18" box="[538,772,927,950]" italics="true" pageId="4" pageNumber="55">Gecarcoidea lalandii</emphasis>
H. 
<bibRefCitation id="5244AFD3FF88FF8E2A9947C68BC0FC7B" author="Milne Edwards, H" box="[171,417,958,981]" pageId="4" pageNumber="55" pagination="1 - 531" refId="ref3338" refString="Milne Edwards, H. 1837. Histoire naturelle des Crustaces, comprenant l'anatpmie, la physiologie et la classification de ces animaux, 2: 1 - 531, atlas 1 - 32, pls. 1 - 42." type="journal article" year="1837">Milne Edwards, 1837</bibRefCitation>
</taxonomicName>
. Many bats inhabit both caves. 
<taxonomicName id="F1D5A9A1FF88FF8E2AB447A78BC1FC58" authorityName="Husana &amp; Naruse &amp; Kase" authorityYear="2010" box="[134,416,991,1014]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="philippinarum">
<emphasis id="04A10E30FF88FF8E2AB447A78BC1FC58" box="[134,416,991,1014]" italics="true" pageId="4" pageNumber="55">Karstarma philippinarum</emphasis>
</taxonomicName>
was observed in water and dry parts of the cave floors and walls, seemingly feeding on bat feces and other organic materials in both caves.
</paragraph>
</subSubSection>
<subSubSection id="7ECF81A9FF88FF8E2AB440268B37FB1B" pageId="4" pageNumber="55" type="distribution">
<paragraph id="366AD222FF88FF8E2AB440268B37FB1B" blockId="4.[134,772,1118,1205]" pageId="4" pageNumber="55">
<emphasis id="04A10E30FF88FF8E2AB440268800FBD8" box="[134,609,1118,1142]" italics="true" pageId="4" pageNumber="55">
<emphasis id="04A10E30FF88FF8E2AB440268B53FBD8" bold="true" box="[134,306,1118,1142]" italics="true" pageId="4" pageNumber="55">Distribution. –</emphasis>
<taxonomicName id="F1D5A9A1FF88FF8E2B0F40278800FBD8" authority="Husana &amp; Naruse &amp; Kase, 2010" authorityName="Husana &amp; Naruse &amp; Kase" authorityYear="2010" box="[317,609,1119,1142]" class="Malacostraca" family="Sesarmidae" genus="Karstarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="philippinarum" status="sp. nov.">Karstarma philippinarum</taxonomicName>
</emphasis>
, 
<taxonomicNameLabel id="1F92B34BFF88FF8E284640268961FBDB" box="[628,768,1118,1141]" pageId="4" pageNumber="55" rank="species">new species</taxonomicNameLabel>
, is currently known only from Boracay Island and Samal Island, 
<collectingCountry id="4EC292B2FF88FF8E2AE440E68B33FB1B" box="[214,338,1182,1205]" name="Philippines" pageId="4" pageNumber="55">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>