364 lines
44 KiB
XML
364 lines
44 KiB
XML
<document id="94F8545217AF8AB23F73140344B48521" ID-DOI="10.11646/phytotaxa.392.1.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="13717373" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725651628762" checkinUser="felipe" docAuthor="Ovrebo, Clark L., Hughes, Karen W. & Halling, Roy E." docDate="2019" docId="03EA87808430FF92FF47CC53FAC3FE09" docLanguage="en" docName="phytotaxa.392.1.3.pdf" docOrigin="Phytotaxa 392 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.392.1.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Tricholoma atratum Ovrebo, Hughes and Halling 2019, sp. nov." docType="treatment" docVersion="1" lastPageNumber="42" masterDocId="FFD3FFF88437FF9BFFCFCD27FF99FFBC" masterDocTitle="Three new species of Tricholoma from Costa Rica" masterLastPageNumber="44" masterPageNumber="33" pageNumber="40" updateTime="1725659538788" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="48CC2BC2209516FA1FD41CA21E8FCB32" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="33D7FA74A49DDAC057EB2E0FCD733CB2">
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<mods:title id="6CB96474F81EED21E09AB0AD54CA9EA2">Three new species of Tricholoma from Costa Rica</mods:title>
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</mods:titleInfo>
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<mods:name id="331AC7638B415FC833F5A2E764F7A964" type="personal">
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<mods:role id="E43CE663524DA53B8D686C0FB54C979E">
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<mods:roleTerm id="0B3E8E341BAC33944D8FC25E4ADB38A0">Author</mods:roleTerm>
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<mods:namePart id="8384F2C5031035911E45E12A3EA94CCD">Ovrebo, Clark L.</mods:namePart>
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<mods:affiliation id="320F98E50934AF26A5E23166B1DF79D1">Department of Biology, University of Central Oklahoma, Edmond, OK 73034, USA</mods:affiliation>
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</mods:name>
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<mods:name id="F0C397E9782235FDD2B7D40B02FA53B0" type="personal">
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<mods:roleTerm id="A00EFDFFC8952199D18FDACEDF3CEFAF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1A387D1ABCABDD98D7036381ABFFAF88">Hughes, Karen W.</mods:namePart>
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<mods:affiliation id="C761B335F31B99DA9F271E9409B3E5DE">Department of Ecology and Evolutionary Biology, University of Tennessee, Knoxville, TN 37996 USA</mods:affiliation>
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</mods:name>
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<mods:name id="CC8E2E20CB9FA3B3796492441F197242" type="personal">
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<mods:roleTerm id="0E1F12253415AC723110A852F260D38A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="68AB2D8F1C6A7AD97580F2A2A02A5D08">Halling, Roy E.</mods:namePart>
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<mods:affiliation id="97CEF10ABCD31943D2C9C76F4240ACBA">Institute of Systematic Botany, The New York Botanical Garden, Bronx, NY 10458 USA</mods:affiliation>
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<mods:typeOfResource id="523A348292117AC28FD709196ED1C84B">text</mods:typeOfResource>
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<mods:titleInfo id="D409F1B8A3789613F5431EEB5EE1DAC3">
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<mods:title id="2316DB198C2F8673EE38B7242C7149F5">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="4CC07D5776A02DC3C0760D1718640D02">
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<mods:date id="253638E52CDF02E0B23263E3DF15590D">2019</mods:date>
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<mods:detail id="27FCE80A5CFB12657D17C9980E5A6AA2" type="pubDate">
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<mods:number id="787708379803B66C101AF023FC320D60">2019-02-12</mods:number>
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<mods:detail id="774E2E6295165D7316B87863EA30B2FA" type="volume">
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<mods:number id="1E634203B4F7168391840AA54954C849">392</mods:number>
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</mods:detail>
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<mods:number id="5F0DE48DF8AA5A36AF1F5D53923175CA">1</mods:number>
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<mods:start id="0E4A57800F128083B58228146E90685A">33</mods:start>
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<mods:location id="D38D47DE01DBC49CB2DA7CB895B60DEE">
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<mods:url id="74D4D2448D7295C7C49A6097378A0A75">http://dx.doi.org/10.11646/phytotaxa.392.1.3</mods:url>
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<mods:classification id="656B98BDB565E5176E7F88F428573335">journal article</mods:classification>
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<mods:identifier id="3C12DD7B8F39CC8F386553AA8C7A45CC" type="DOI">10.11646/phytotaxa.392.1.3</mods:identifier>
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<mods:identifier id="E9BF45E0D4FCE31AAB679B729A0FCA01" type="ISSN">1179-3163</mods:identifier>
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<mods:identifier id="E57FA998AC24E33D647BAE33ECFFE35E" type="Zenodo-Dep">13717373</mods:identifier>
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</mods:mods>
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<treatment id="03EA87808430FF92FF47CC53FAC3FE09" LSID="urn:lsid:plazi:treatment:03EA87808430FF92FF47CC53FAC3FE09" httpUri="http://treatment.plazi.org/id/03EA87808430FF92FF47CC53FAC3FE09" lastPageId="9" lastPageNumber="42" pageId="7" pageNumber="40">
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<subSubSection id="C359651D8430FF9CFF47CC53FE3CFE0C" pageId="7" pageNumber="40" type="nomenclature">
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||
<paragraph id="8BFC36968430FF9CFF47CC53FC2DFE31" blockId="7.[136,948,372,397]" box="[136,948,372,397]" pageId="7" pageNumber="40">
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<heading id="D0B481FA8430FF9CFF47CC53FC2DFE31" box="[136,948,372,397]" fontSize="11" level="2" pageId="7" pageNumber="40" reason="3">
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<taxonomicName id="4C434D158430FF9CFF47CC53FD23FE31" ID-CoL="B32H2" authority="Ovrebo, Hughes and Halling" authorityName="Ovrebo, Hughes and Halling" authorityYear="2019" box="[136,698,372,397]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="40" phylum="Basidiomycota" rank="species" species="atratum" status="sp. nov.">
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<emphasis id="B937EA848430FF9CFF47CC53FEE8FE31" bold="true" box="[136,369,372,397]" italics="true" pageId="7" pageNumber="40">Tricholoma atratum</emphasis>
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Ovrebo, Hughes and Halling
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</taxonomicName>
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<emphasis id="B937EA848430FF9CFD0ECC53FC8FFE31" box="[705,790,372,397]" italics="true" pageId="7" pageNumber="40">
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<taxonomicNameLabel id="A20457FF8430FF9CFD0ECC53FC8FFE31" box="[705,790,372,397]" pageId="7" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
(
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<figureCitation id="13782A138430FF9CFCEACC53FCEDFE31" box="[805,884,372,397]" captionStart="FIGURE 2" captionStartId="4.[136,229,1504,1525]" captionTargetBox="[166,1420,269,1422]" captionTargetId="figure-16@4.[151,1437,247,1442]" captionTargetPageId="4" captionText="FIGURE 2. PhyML analysis of the nuclear ribosomal ITS region. Bootstrap values are given to the left of the supported node." figureDoi="http://doi.org/10.5281/zenodo.13717377" httpUri="https://zenodo.org/record/13717377/files/figure.png" pageId="7" pageNumber="40">Figs. 2</figureCitation>
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,
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<figureCitation id="13782A138430FF9CFC4FCC53FC09FE31" box="[896,912,372,397]" captionStart="FIGURES 5" captionStartId="6.[136,243,1207,1228]" captionTargetBox="[219,1368,775,1182]" captionTargetId="figure-41@6.[219,1368,775,1182]" captionTargetPageId="6" captionText="FIGURES 5. Basidiomes. Tricholoma atratum. A. Halling 8263, holotype. B. Halling 8158. Photos by R. E. Halling. Scale line = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.13717384" httpUri="https://zenodo.org/record/13717384/files/figure.png" pageId="7" pageNumber="40">5</figureCitation>
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,
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<figureCitation id="13782A138430FF9CFC53CC53FC32FE31" box="[924,939,372,397]" captionStart="FIGURE 7" captionStartId="9.[136,229,1279,1300]" captionTargetBox="[201,1389,481,1239]" captionTargetId="figure-161@9.[157,1430,461,1254]" captionTargetPageId="9" captionText="FIGURE 7. Microscopic features of Tricholoma atratum. holotype. A. basidiospores, B. pileocystidia. Scale line=10 μm. Drawings by C. L. Ovrebo." figureDoi="http://doi.org/10.5281/zenodo.13717388" httpUri="https://zenodo.org/record/13717388/files/figure.png" pageId="7" pageNumber="40">7</figureCitation>
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)
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</heading>
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</paragraph>
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<paragraph id="8BFC36968430FF9CFF47CCBCFE3CFE0C" blockId="7.[136,421,411,432]" box="[136,421,411,432]" pageId="7" pageNumber="40">Mycobank no.:—MB 827633</paragraph>
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</subSubSection>
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<subSubSection id="C359651D8430FF92FF47CCC7FAC3FE09" lastPageId="9" lastPageNumber="42" pageId="7" pageNumber="40" type="diagnosis">
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<paragraph id="8BFC36968430FF9CFF47CCC7FDF5FDA1" blockId="7.[136,1452,480,2089]" pageId="7" pageNumber="40">
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Diagnosis:—Characterized by the dark gray dry pileus, lack of cheilocystidia, presence of pileocystidia, relatively large spores, and occurrence in
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<collectingCountry id="F35476068430FF9CFE25CF23FDF1FDA1" box="[490,616,516,541]" name="Costa Rica" pageId="7" pageNumber="40">Costa Rica</collectingCountry>
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.
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</paragraph>
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<paragraph id="8BFC36968430FF9CFF72CF0FFBFCFDD9" blockId="7.[136,1452,480,2089]" pageId="7" pageNumber="40">
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<materialsCitation id="3B2B3CCB8430FF9CFF72CF0FFBFCFDD9" collectingDate="2001-07-06" collectionCode="N, W, NY" collectorName="El Guarco & Macizo de la Muerte" country="Costa Rica" elevation="2600" location="Cartago Prov." pageId="7" pageNumber="40" specimenCode="USJ 109186" specimenCount="2" stateProvince="Cartago" typeStatus="holotype">
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<typeStatus id="54F888348430FF9CFF72CF0FFEBCFDFD" box="[189,293,552,577]" pageId="7" pageNumber="40" type="holotype">Holotype</typeStatus>
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:—
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<collectingCountry id="F35476068430FF9CFE8ACF0FFE68FDFD" box="[325,497,552,577]" name="Costa Rica" pageId="7" pageNumber="40">COSTA RICA</collectingCountry>
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.
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<collectingRegion id="4987F8748430FF9CFE32CF0FFD04FDFD" box="[509,669,552,577]" country="Costa Rica" name="Cartago" pageId="7" pageNumber="40">Cartago Prov.</collectingRegion>
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:
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<collectorName id="26B653408430FF9CFD66CF0FFCB9FDFD" box="[681,800,552,577]" pageId="7" pageNumber="40">El Guarco</collectorName>
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, Tapanti, Parque Nacional Tapanti,
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<collectorName id="26B653408430FF9CFB78CF0FFA31FDFD" box="[1207,1448,552,577]" pageId="7" pageNumber="40">Macizo de la Muerte</collectorName>
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, 9˚41’6”
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<collectionCode id="ED52AE538430FF9CFF29CF6BFF64FDD9" box="[230,253,588,613]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="7" pageNumber="40" type="Herbarium">N</collectionCode>
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, 83˚52’30”
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<collectionCode id="ED52AE538430FF9CFE4DCF6BFE04FDD9" box="[386,413,588,613]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="7" pageNumber="40" type="Herbarium">W</collectionCode>
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,
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<quantity id="4CBB9B738430FF9CFE67CF6BFD98FDD9" box="[424,513,588,613]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="40" unit="m" value="2600.0">
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||
<elevation id="006ED1A58430FF9CFE67CF6BFD98FDD9" box="[424,513,588,613]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="40" unit="m" value="2600.0">2600 m</elevation>
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</quantity>
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:
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<date id="FFFD10568430FF9CFDC4CF6BFD17FDD9" box="[523,654,588,613]" pageId="7" pageNumber="40" value="2001-07-06">
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<collectingDate id="EFB9E9BE8430FF9CFDC4CF6BFD17FDD9" box="[523,654,588,613]" pageId="7" pageNumber="40" value="2001-07-06">6 July 2001</collectingDate>
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</date>
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<emphasis id="B937EA848430FF9CFD5ACF6BFCB3FDD9" box="[661,810,588,613]" italics="true" pageId="7" pageNumber="40">Halling 8263</emphasis>
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(
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<specimenCode id="DBE59EED8430FF9CFCF7CF6BFC51FDD9" box="[824,968,588,613]" collectionCode="USJ" country="Costa Rica" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15819" lsid="urn:lsid:biocol.org:col:15819" name="Universidad de Costa Rica" pageId="7" pageNumber="40" type="University or college">USJ 109186</specimenCode>
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;
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<typeStatus id="54F888348430FF9CFC1DCF6BFBBDFDD9" box="[978,1060,588,613]" pageId="7" pageNumber="40" type="isotype">Isotype</typeStatus>
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<collectionCode id="ED52AE538430FF9CFBE3CF6BFBC0FDD9" box="[1068,1113,588,613]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="7" pageNumber="40" type="Herbarium">NY</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BFC36968430FF9CFF72CF57FE0AFD11" blockId="7.[136,1452,480,2089]" pageId="7" pageNumber="40">
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Etymology:—From Latin
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<emphasis id="B937EA848430FF9CFE23CF57FDA2FD35" box="[492,571,624,649]" italics="true" pageId="7" pageNumber="40">atratus</emphasis>
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meaning dark, black, clothed in black; referring to the dark gray to black coloration of the pileus.
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</paragraph>
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<paragraph id="8BFC36968430FF9CFF72CF9FFE8AFC71" blockId="7.[136,1452,480,2089]" pageId="7" pageNumber="40">
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Description:—
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<emphasis id="B937EA848430FF9CFEABCF9FFE33FD6D" box="[356,426,696,721]" italics="true" pageId="7" pageNumber="40">Pileus</emphasis>
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<quantity id="4CBB9B738430FF9CFE7BCF9FFDA9FD6D" box="[436,560,696,721]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="7" pageNumber="40" unit="mm" value="60.0" valueMax="80.0" valueMin="40.0">40–80 mm</quantity>
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wide, convex to plano-convex, sometimes with a low umbo, dry, appressedfibrillose, becoming radially rugulose often with some moisture loss, black on disc and dark gray toward margin or dark gray overall, with undertones of pale greenish yellow; context
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<quantity id="4CBB9B738430FF9CFCBDCE27FC2BFCA5" box="[882,946,768,793]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="40" unit="mm" value="7.0">7 mm</quantity>
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thick (on one collection) white to pale watery gray, unchanging, odor and taste farinaceous.
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<emphasis id="B937EA848430FF9CFD48CE03FD76FC81" box="[647,751,804,829]" italics="true" pageId="7" pageNumber="40">Lamellae</emphasis>
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adnexed-emarginate, broad, grayish yellow (4A3, 4B4), close, edges even to uneven.
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<emphasis id="B937EA848430FF9CFE49CE6FFE27FCDD" box="[390,446,840,865]" italics="true" pageId="7" pageNumber="40">Stipe</emphasis>
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70–110 (150) mm long,
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<quantity id="4CBB9B738430FF9CFD1ACE6FFCD9FCDD" box="[725,832,840,865]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="13.0" metricValueMin="6.0" pageId="7" pageNumber="40" unit="mm" value="9.5" valueMax="13.0" valueMin="6.0">6–13 mm</quantity>
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wide, equal but sometimes tapered or enlarged at base, fibrillose, whitish with pale gray tints, some staining slightly orangish brown from handling with age, white at base and often matted-fibrillose, context becoming hollow, white, unchanging.
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<emphasis id="B937EA848430FF9CFC69CEB7FAB5FC15" box="[934,1324,912,937]" italics="true" pageId="7" pageNumber="40">Macrochemical color reactions:—</emphasis>
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no reaction with PDAB.
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</paragraph>
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<paragraph id="8BFC36968430FF9CFF72CEFFFF4EFA1D" blockId="7.[136,1452,480,2089]" pageId="7" pageNumber="40">
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<emphasis id="B937EA848430FF9CFF72CEFFFEC2FC4D" box="[189,347,984,1009]" italics="true" pageId="7" pageNumber="40">Basidiospores</emphasis>
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8–9(–9.5) × 6–7.5 μm (mean=8.53 × 6.54 μm, Q=1.21–1.42, mean Q=1.3), elliptic, broadly elliptic to subovate in profile and face views, smooth, thin-walled, colourless, inamyloid.
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<emphasis id="B937EA848430FF9CFB83CEDBFB3BFBA9" box="[1100,1186,1020,1045]" italics="true" pageId="7" pageNumber="40">Basidia</emphasis>
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39–50 × 6–10 μm, 4- sterigmate, clavate, colourless, many with granular-guttulate contents.
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<emphasis id="B937EA848430FF9CFC0EC907FB09FB85" box="[961,1168,1056,1081]" italics="true" pageId="7" pageNumber="40">Hymenial cystidia</emphasis>
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absent.
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<emphasis id="B937EA848430FF9CFB39C907FA32FB85" box="[1270,1451,1056,1081]" italics="true" pageId="7" pageNumber="40">Lamellar trama</emphasis>
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hyphae 4–10 μm diam, parallel, pale yellow.
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<emphasis id="B937EA848430FF9CFDB6C963FC8FFBE1" box="[633,790,1092,1117]" italics="true" pageId="7" pageNumber="40">Subhymenium</emphasis>
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composed of hyphae 2.5–3 μm diam, colourless.
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<emphasis id="B937EA848430FF9CFAF5C963FA35FBE1" box="[1338,1452,1092,1117]" italics="true" pageId="7" pageNumber="40">Pileipellis</emphasis>
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as a cutis, hyphae radially arranged to slightly interwoven, not embedded in a gelatinous matrix, 3–8 μm diam, cylindric to slightly inflated, smooth or roughened and thickened with brownish incrustations, the outer wall surface sometimes fracturing to
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<taxonomicName id="4C434D158430FF9CFED3C997FE02FB75" box="[284,411,1200,1225]" form="bands" pageId="7" pageNumber="40" rank="form">form bands</taxonomicName>
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of varying pigment intensity, colourless to brown, brown as a layer.
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<emphasis id="B937EA848430FF9CFB68C997FAA0FB75" box="[1191,1337,1200,1225]" italics="true" pageId="7" pageNumber="40">Pileocystidia</emphasis>
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scattered, formed by recurved end-cells, 22–37 × 5–13 μm, cylindric to clavate, occasionally two-celled, brown.
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<emphasis id="B937EA848430FF9CFA83C9F3FA32FB51" box="[1356,1451,1236,1261]" italics="true" pageId="7" pageNumber="40">Subcutis</emphasis>
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composed of pseudoparenchymatous hyphae, cells 12–23 μm diam, broadly inflated, cells of the layer often collapsed and the layer is then difficult to discern, colourless to brown as a layer.
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<emphasis id="B937EA848430FF9CFC69C83BFB87FA89" box="[934,1054,1308,1333]" italics="true" pageId="7" pageNumber="40">Pileitrama</emphasis>
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made up of hyphae 4–12 μm diam, pale yellow.
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<emphasis id="B937EA848430FF9CFED6C867FE00FAE5" box="[281,409,1344,1369]" italics="true" pageId="7" pageNumber="40">Stipitipellis</emphasis>
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made up of hyphae 3–5 μm wide, appressed or sometimes clumped or entangled in places, smooth, thin-walled, very pale yellow.
|
||
<emphasis id="B937EA848430FF9CFD8CC843FD50FAC1" box="[579,713,1380,1405]" italics="true" pageId="7" pageNumber="40">Stipititrama</emphasis>
|
||
hyphae 4–14 μm diam, parallel, pale yellow. Clamp connections absent.
|
||
</paragraph>
|
||
<paragraph id="8BFC36968430FF9CFF72C88BFB97FA79" blockId="7.[136,1452,480,2089]" box="[189,1038,1452,1477]" pageId="7" pageNumber="40">
|
||
Habitat:—Gregarious, on soil, under
|
||
<taxonomicName id="4C434D158430FF9CFD91C88BFCA5FA79" box="[606,828,1452,1477]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="40" phylum="Tracheophyta" rank="species" species="copeyensis">
|
||
<emphasis id="B937EA848430FF9CFD91C88BFCA5FA79" box="[606,828,1452,1477]" italics="true" pageId="7" pageNumber="40">Quercus copeyensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B937EA848430FF9CFCBCC88BFB97FA79" box="[883,1038,1452,1477]" italics="true" pageId="7" pageNumber="40">
|
||
<taxonomicName id="4C434D158430FF9CFCBCC88BFB93FA79" box="[883,1034,1452,1477]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="40" phylum="Tracheophyta" rank="species" species="seemannii">Q. seemannii</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BFC36968430FF9CFF72C8F7FD5FF98D" blockId="7.[136,1452,480,2089]" pageId="7" pageNumber="40">
|
||
<materialsCitation id="3B2B3CCB8430FF9CFF72C8F7FD58F98D" collectingDate="2000-07-10" collectingDateMax="2001-06-09" collectingDateMin="2000-07-10" collectionCode="N, W, NY" country="Costa Rica" county="San Gerardo de Dota" elevation="2200" location="Savegre" municipality="Albergue de la Montana" pageId="7" pageNumber="40" specimenCode="USJ 72083, USJ 74107" specimenCount="1" stateProvince="San Jose">
|
||
Additional collections studied:—
|
||
<collectingCountry id="F35476068430FF9CFDE2C8F7FD4EFA55" box="[557,727,1488,1513]" name="Costa Rica" pageId="7" pageNumber="40">COSTA RICA</collectingCountry>
|
||
.
|
||
<collectingRegion id="4987F8748430FF9CFD2FC8F7FC1EFA55" box="[736,903,1488,1513]" country="Costa Rica" name="San Jose" pageId="7" pageNumber="40">San Jose Prov.</collectingRegion>
|
||
:
|
||
<collectingCounty id="629D4E1A8430FF9CFC5EC8F7FBE6FA55" box="[913,1151,1488,1513]" pageId="7" pageNumber="40">San Gerardo de Dota</collectingCounty>
|
||
, ±
|
||
<locationDeviation id="9A4E2FB48430FF9CFB6FC8F7FF45F9B1" location="Cerro de la Muerte" pageId="7" pageNumber="40">
|
||
<quantity id="4CBB9B738430FF9CFB6FC8F7FB40FA55" box="[1184,1241,1488,1513]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="40" unit="km" value="5.0">5 km</quantity>
|
||
SW of Cerro de la Muerte
|
||
</locationDeviation>
|
||
,
|
||
<collectingMunicipality id="6B98ACEC8430FF9CFF2BC8D3FE6BF9B1" box="[228,498,1524,1549]" pageId="7" pageNumber="40">Albergue de la Montaña</collectingMunicipality>
|
||
,
|
||
<location id="8E9C604D8430FF9CFE33C8D3FDC0F9B1" LSID="urn:lsid:plazi:treatment:03EA87808430FF92FF47CC53FAC3FE09:8E9C604D8430FF9CFE33C8D3FDC0F9B1" box="[508,601,1524,1549]" country="Costa Rica" county="San Gerardo de Dota" municipality="Albergue de la Montana" name="Savegre" pageId="7" pageNumber="40" stateProvince="San Jose">Savegre</location>
|
||
, 9˚32’02”
|
||
<collectionCode id="ED52AE538430FF9CFD01C8D3FD7CF9B1" box="[718,741,1524,1549]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="7" pageNumber="40" type="Herbarium">N</collectionCode>
|
||
, 83˚48’27”
|
||
<collectionCode id="ED52AE538430FF9CFCA8C8D3FC1BF9B1" box="[871,898,1524,1549]" country="Austria" lsid="urn:lsid:biocol.org:col:15588" name="Naturhistorisches Museum Wien" pageId="7" pageNumber="40" type="Herbarium">W</collectionCode>
|
||
,
|
||
<quantity id="4CBB9B738430FF9CFC43C8D3FC78F9B1" box="[908,993,1524,1549]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="40" unit="m" value="2200.0">
|
||
<elevation id="006ED1A58430FF9CFC43C8D3FC78F9B1" box="[908,993,1524,1549]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="40" unit="m" value="2200.0">2200 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFFD10568430FF9CFC23C8D3FBE4F9B1" box="[1004,1149,1524,1549]" pageId="7" pageNumber="40" value="2000-07-10">
|
||
<collectingDate id="EFB9E9BE8430FF9CFC23C8D3FBE4F9B1" box="[1004,1149,1524,1549]" pageId="7" pageNumber="40" value="2000-07-10">10 July 2000</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="B937EA848430FF9CFB48C8D3FA83F9B1" box="[1159,1306,1524,1549]" italics="true" pageId="7" pageNumber="40">Halling 8008</emphasis>
|
||
(
|
||
<specimenCode id="DBE59EED8430FF9CFAE7C8D3FA31F9B1" box="[1320,1448,1524,1549]" collectionCode="USJ" country="Costa Rica" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15819" lsid="urn:lsid:biocol.org:col:15819" name="Universidad de Costa Rica" pageId="7" pageNumber="40" type="University or college">USJ 72083</specimenCode>
|
||
;
|
||
<collectionCode id="ED52AE538430FF9CFF47CB3FFF2DF98D" box="[136,180,1560,1585]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="7" pageNumber="40" type="Herbarium">NY</collectionCode>
|
||
),
|
||
<date id="FFFD10568430FF9CFF07CB3FFECBF98D" box="[200,338,1560,1585]" pageId="7" pageNumber="40" value="2001-06-09">
|
||
<collectingDate id="EFB9E9BE8430FF9CFF07CB3FFECBF98D" box="[200,338,1560,1585]" pageId="7" pageNumber="40" value="2001-06-09">9 June 2001</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="B937EA848430FF9CFE92CB3FFE6BF98D" box="[349,498,1560,1585]" italics="true" pageId="7" pageNumber="40">Halling 8158</emphasis>
|
||
(
|
||
<specimenCode id="DBE59EED8430FF9CFDCFCB3FFD1BF98D" box="[512,642,1560,1585]" collectionCode="USJ" country="Costa Rica" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15819" lsid="urn:lsid:biocol.org:col:15819" name="Universidad de Costa Rica" pageId="7" pageNumber="40" type="University or college">USJ 74107</specimenCode>
|
||
;
|
||
<collectionCode id="ED52AE538430FF9CFD43CB3FFD20F98D" box="[652,697,1560,1585]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="7" pageNumber="40" type="Herbarium">NY</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFC36968430FF9CFF72CB1BFD14F8B5" blockId="7.[136,1452,480,2089]" pageId="7" pageNumber="40">
|
||
Discussion:—
|
||
<taxonomicName id="4C434D158430FF9CFE93CB1BFDDAF9E9" authorityName="Ovrebo, Hughes and Halling" authorityYear="2019" box="[348,579,1596,1621]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="40" phylum="Basidiomycota" rank="species" species="atratum">
|
||
<emphasis id="B937EA848430FF9CFE93CB1BFDDAF9E9" box="[348,579,1596,1621]" italics="true" pageId="7" pageNumber="40">Tricholoma atratum</emphasis>
|
||
</taxonomicName>
|
||
is characterized by the dark gray, dry pileus, farinaceous odor and taste, pseudoparenchymatous pileal subcutis, lack of cheilocystidia, presence of scattered pileocystidia, and relatively large spores. The yellowish undertones on the pileus surface and grayish yellow pigmentation of the lamellae may also serve as distinguishing features of this species. The recurved end cells of the pileus surface we interpret as pileocystidia (
|
||
<figureCitation id="13782A138430FF9CFF5FCBEBFF6AF959" box="[144,243,1740,1765]" captionStart="FIGURE 7" captionStartId="9.[136,229,1279,1300]" captionTargetBox="[201,1389,481,1239]" captionTargetId="figure-161@9.[157,1430,461,1254]" captionTargetPageId="9" captionText="FIGURE 7. Microscopic features of Tricholoma atratum. holotype. A. basidiospores, B. pileocystidia. Scale line=10 μm. Drawings by C. L. Ovrebo." figureDoi="http://doi.org/10.5281/zenodo.13717388" httpUri="https://zenodo.org/record/13717388/files/figure.png" pageId="7" pageNumber="40">Figure 7</figureCitation>
|
||
). Pileal surface hyphae can be recurved in
|
||
<taxonomicName id="4C434D158430FF9CFD10CBEBFC23F959" box="[735,954,1740,1765]" pageId="7" pageNumber="40">
|
||
<emphasis id="B937EA848430FF9CFD10CBEBFCF9F959" box="[735,864,1740,1765]" italics="true" pageId="7" pageNumber="40">Tricholoma</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
but they are usually clumped and represent macromorphologically the surface squamules.
|
||
</paragraph>
|
||
<paragraph id="8BFC36968430FF93FF72CA33FCAAFEBD" blockId="7.[136,1452,480,2089]" lastBlockId="8.[136,1452,160,257]" lastPageId="8" lastPageNumber="41" pageId="7" pageNumber="40">
|
||
The rugulose pileus, farinaceous odor and taste and pseudoparenchymatous subcutis are features that
|
||
<taxonomicName id="4C434D158430FF9CFAF8CA33FA35F891" authorityName="Ovrebo, Hughes and Halling" authorityYear="2019" box="[1335,1452,1812,1837]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="40" phylum="Basidiomycota" rank="species" species="atratum">
|
||
<emphasis id="B937EA848430FF9CFAF8CA33FA35F891" box="[1335,1452,1812,1837]" italics="true" pageId="7" pageNumber="40">T. atratum</emphasis>
|
||
</taxonomicName>
|
||
shares with the other taxa described here as well as taxa found in
|
||
<bibRefCitation id="EFD24B678430FF9CFC5FCA1FFBA0F8ED" author="Ovrebo, C. L." box="[912,1081,1848,1873]" pageId="7" pageNumber="40" pagination="418 - 425" refId="ref6106" refString="Ovrebo, C. L. (1986) Three new species of Tricholoma with a description of Tricholoma luteomaculosum. Mycologia 78 (3): 418 - 425. https: // doi. org / 10.2307 / 3793045" type="journal article" year="1986">Ovrebo (1986)</bibRefCitation>
|
||
and several species included in
|
||
<bibRefCitation id="EFD24B678430FF9CFF47CA7BFEB5F8C9" author="Shanks, K. M." box="[136,300,1884,1909]" pageId="7" pageNumber="40" refId="ref6448" refString="Shanks, K. M. (1997) The Agaricales (gilled fungi) of California. 11. Tricholomataceae II. Tricholoma. Mad River Press, Eureka, California. 54 pp. + 9 pls." type="book" year="1997">Shanks (1997)</bibRefCitation>
|
||
, so the initial hypothesis was that species sharing these characteristics would all be related. For many of the putatively related taxa found in these publications, sequence data are not yet available so molecular comparisons can only be made among a few species. It now appears unlikely, however, that
|
||
<taxonomicName id="4C434D158430FF9CFBD0CA83FB0FF801" authorityName="Ovrebo, Hughes and Halling" authorityYear="2019" box="[1055,1174,1956,1981]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="40" phylum="Basidiomycota" rank="species" species="atratum">
|
||
<emphasis id="B937EA848430FF9CFBD0CA83FB0FF801" box="[1055,1174,1956,1981]" italics="true" pageId="7" pageNumber="40">T. atratum</emphasis>
|
||
</taxonomicName>
|
||
is closely related to the phenotypically similar taxa as shown in
|
||
<figureCitation id="13782A138430FF9CFD85CAEFFD2AF85D" box="[586,691,1992,2017]" captionStart="FIGURE 1" captionStartId="3.[136,229,1609,1630]" captionTargetBox="[324,1260,260,1475]" captionTargetId="figure-14@3.[259,1329,187,1587]" captionTargetPageId="3" captionText="FIGURE 1. PhyML analysis of the nuclear ribosomal ITS region. Collection locations where known are given by the US Postal code for states. GSM=Great Smoky Mountains NationalPark,Tennessee side; BC=Canada British Columbia; CR=Costa Rica; ENV=environmental sequence. Ecto=ectomycorrhizal root tip. Bootstrap values are given to the left of the supported node. Percent divergence calculations were calculated using all base pair differences (i.e., a 13bp insertion in CLO3549 and AGF21 was counted as 13 bp differences, not a single difference)." figureDoi="http://doi.org/10.5281/zenodo.13717375" httpUri="https://zenodo.org/record/13717375/files/figure.png" pageId="7" pageNumber="40">Figures 1</figureCitation>
|
||
&
|
||
<figureCitation id="13782A138430FF9CFD18CAEFFD71F85D" box="[727,744,1992,2017]" captionStart="FIGURE 2" captionStartId="4.[136,229,1504,1525]" captionTargetBox="[166,1420,269,1422]" captionTargetId="figure-16@4.[151,1437,247,1442]" captionTargetPageId="4" captionText="FIGURE 2. PhyML analysis of the nuclear ribosomal ITS region. Bootstrap values are given to the left of the supported node." figureDoi="http://doi.org/10.5281/zenodo.13717377" httpUri="https://zenodo.org/record/13717377/files/figure.png" pageId="7" pageNumber="40">2</figureCitation>
|
||
. The phylogeny in
|
||
<figureCitation id="13782A138430FF9CFC71CAEFFB85F85D" box="[958,1052,1992,2017]" captionStart="FIGURE 1" captionStartId="3.[136,229,1609,1630]" captionTargetBox="[324,1260,260,1475]" captionTargetId="figure-14@3.[259,1329,187,1587]" captionTargetPageId="3" captionText="FIGURE 1. PhyML analysis of the nuclear ribosomal ITS region. Collection locations where known are given by the US Postal code for states. GSM=Great Smoky Mountains NationalPark,Tennessee side; BC=Canada British Columbia; CR=Costa Rica; ENV=environmental sequence. Ecto=ectomycorrhizal root tip. Bootstrap values are given to the left of the supported node. Percent divergence calculations were calculated using all base pair differences (i.e., a 13bp insertion in CLO3549 and AGF21 was counted as 13 bp differences, not a single difference)." figureDoi="http://doi.org/10.5281/zenodo.13717375" httpUri="https://zenodo.org/record/13717375/files/figure.png" pageId="7" pageNumber="40">Figure 1</figureCitation>
|
||
shows a close relationship between
|
||
<taxonomicName id="4C434D158430FF9CFF47CACBFE28F7B9" authority="(Shanks 1997)" baseAuthorityName="Shanks" baseAuthorityYear="1997" box="[136,433,2028,2053]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="40" phylum="Basidiomycota" rank="species" species="mutabile">
|
||
<emphasis id="B937EA848430FF9CFF47CACBFE9CF7B9" box="[136,261,2028,2053]" italics="true" pageId="7" pageNumber="40">T. mutabile</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD24B678430FF9CFEDBCACBFE30F7B9" author="Shanks, K. M." box="[276,425,2028,2053]" pageId="7" pageNumber="40" refId="ref6448" refString="Shanks, K. M. (1997) The Agaricales (gilled fungi) of California. 11. Tricholomataceae II. Tricholoma. Mad River Press, Eureka, California. 54 pp. + 9 pls." type="book" year="1997">Shanks 1997</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C434D158430FF9CFE72CACBFC82F7B9" authority="(Ovrebo 1986)" baseAuthorityName="Ovrebo" baseAuthorityYear="1986" box="[445,795,2028,2053]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="40" phylum="Basidiomycota" rank="species" species="marquettense">
|
||
<emphasis id="B937EA848430FF9CFE72CACBFDF7F7B9" box="[445,622,2028,2053]" italics="true" pageId="7" pageNumber="40">T. marquettense</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD24B678430FF9CFDB2CACBFC8BF7B9" author="Ovrebo, C. L." box="[637,786,2028,2053]" pageId="7" pageNumber="40" pagination="418 - 425" refId="ref6106" refString="Ovrebo, C. L. (1986) Three new species of Tricholoma with a description of Tricholoma luteomaculosum. Mycologia 78 (3): 418 - 425. https: // doi. org / 10.2307 / 3793045" type="journal article" year="1986">Ovrebo 1986</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C434D158430FF9CFCE6CACBFC0CF7B9" box="[809,917,2028,2053]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="40" phylum="Basidiomycota" rank="species" species="felschii">
|
||
<emphasis id="B937EA848430FF9CFCE6CACBFC0CF7B9" box="[809,917,2028,2053]" italics="true" pageId="7" pageNumber="40">T. felschii</emphasis>
|
||
</taxonomicName>
|
||
and the European
|
||
<emphasis id="B937EA848430FF9CFBA8CACBFA9CF7B9" box="[1127,1285,2028,2053]" italics="true" pageId="7" pageNumber="40">
|
||
<taxonomicName id="4C434D158430FF9CFBA8CACBFA98F7B9" authorityName="Bon" authorityYear="1975" box="[1127,1281,2028,2053]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="40" phylum="Basidiomycota" rank="species" species="josserandii">T. josserandii</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but
|
||
<taxonomicName id="4C434D158430FF9CFAF8CACBFA35F7B9" authorityName="Ovrebo, Hughes and Halling" authorityYear="2019" box="[1335,1452,2028,2053]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="40" phylum="Basidiomycota" rank="species" species="atratum">
|
||
<emphasis id="B937EA848430FF9CFAF8CACBFA35F7B9" box="[1335,1452,2028,2053]" italics="true" pageId="7" pageNumber="40">T. atratum</emphasis>
|
||
</taxonomicName>
|
||
does not cluster with these taxa. Rather,
|
||
<figureCitation id="13782A138430FF9CFD85C537FD31F795" box="[586,680,2064,2089]" captionStart="FIGURE 2" captionStartId="4.[136,229,1504,1525]" captionTargetBox="[166,1420,269,1422]" captionTargetId="figure-16@4.[151,1437,247,1442]" captionTargetPageId="4" captionText="FIGURE 2. PhyML analysis of the nuclear ribosomal ITS region. Bootstrap values are given to the left of the supported node." figureDoi="http://doi.org/10.5281/zenodo.13717377" httpUri="https://zenodo.org/record/13717377/files/figure.png" pageId="7" pageNumber="40">Figure 2</figureCitation>
|
||
indicates a relationship of
|
||
<taxonomicName id="4C434D158430FF9CFC1AC537FBD0F795" authorityName="Ovrebo, Hughes and Halling" authorityYear="2019" box="[981,1097,2064,2089]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="7" pageNumber="40" phylum="Basidiomycota" rank="species" species="atratum">
|
||
<emphasis id="B937EA848430FF9CFC1AC537FBD0F795" box="[981,1097,2064,2089]" italics="true" pageId="7" pageNumber="40">T. atratum</emphasis>
|
||
</taxonomicName>
|
||
with the European
|
||
<taxonomicName id="4C434D158430FF93FAECC537FEC3FF05" authority="Quel. (1873: 327)" authorityName="Quel." authorityPageNumber="327" authorityYear="1873" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" lastPageId="8" lastPageNumber="41" order="Agaricales" pageId="7" pageNumber="40" phylum="Basidiomycota" rank="species" species="orirubens">
|
||
<emphasis id="B937EA848430FF9CFAECC537FA32F795" box="[1315,1451,2064,2089]" italics="true" pageId="7" pageNumber="40">T. orirubens</emphasis>
|
||
Quél. (1873: 327)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C434D15843FFF93FEA5CD87FD77FF05" authority="Bres. (1898: 47)" authorityName="Bres." authorityPageNumber="47" authorityYear="1898" box="[362,750,160,185]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="8" pageNumber="41" phylum="Basidiomycota" rank="species" species="squarrulosum">
|
||
<emphasis id="B937EA84843FFF93FEA5CD87FDBCFF05" box="[362,549,160,185]" italics="true" pageId="8" pageNumber="41">T. squarrulosum</emphasis>
|
||
Bres. (1898: 47)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C434D15843FFF93FCE2CD87FB42FF05" authority="Sacc. (1887: 104)" authorityName="Sacc." authorityPageNumber="104" authorityYear="1887" box="[813,1243,160,185]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="8" pageNumber="41" phylum="Basidiomycota" rank="species" species="atrosquamosum">
|
||
<emphasis id="B937EA84843FFF93FCE2CD87FC67FF05" box="[813,1022,160,185]" italics="true" pageId="8" pageNumber="41">T. atrosquamosum</emphasis>
|
||
Sacc. (1887: 104)
|
||
</taxonomicName>
|
||
. According to the descriptions provided by
|
||
<bibRefCitation id="EFD24B67843FFF93FE6CCDE3FCE8FF61" author="Christensen, M. & Heilmann-Clausen, J." box="[419,881,196,221]" pageId="8" pageNumber="41" refId="ref5762" refString="Christensen, M. & Heilmann-Clausen, J. (2013) The genus Tricholoma. Fungi of Northern Europe, vol. 4. Svampetryk, Denmark, 228 pp." type="book" year="2013">Christensen & Heilmann-Clausen (2013)</bibRefCitation>
|
||
, the latter species have smaller spores, the subcutis is poorly differentiated and their pilei tend to be squamulose.
|
||
</paragraph>
|
||
<caption id="DF3C661E843FFF93FF47CAECFB18F7B8" ID-DOI="http://doi.org/10.5281/zenodo.13717386" ID-Zenodo-Dep="13717386" httpUri="https://zenodo.org/record/13717386/files/figure.png" pageId="8" pageNumber="41" startId="8.[136,229,1995,2016]" targetBox="[190,1406,295,1925]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8BFC3696843FFF93FF47CAECFB18F7B8" blockId="8.[136,1452,1995,2052]" pageId="8" pageNumber="41">
|
||
<emphasis id="B937EA84843FFF93FF47CAECFF67F85C" bold="true" box="[136,254,1995,2016]" pageId="8" pageNumber="41">FIGURE 6.</emphasis>
|
||
Microscopic features of
|
||
<taxonomicName id="4C434D15843FFF93FE3ECAECFD2AF85C" box="[497,691,1995,2016]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="8" pageNumber="41" phylum="Basidiomycota" rank="species" species="undetermined">
|
||
<emphasis id="B937EA84843FFF93FE3ECAECFDF9F85C" box="[497,608,1995,2016]" italics="true" pageId="8" pageNumber="41">Tricholoma</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
<taxonomicName id="4C434D15843FFF93FD76CAECFCEDF85C" box="[697,884,1995,2016]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="8" pageNumber="41" phylum="Basidiomycota" rank="species" species="felschii">
|
||
<emphasis id="B937EA84843FFF93FD76CAECFCEDF85C" box="[697,884,1995,2016]" italics="true" pageId="8" pageNumber="41">Tricholoma felschii</emphasis>
|
||
</taxonomicName>
|
||
, holotype, A. basidiospores, B. cheilocystidia.
|
||
<taxonomicName id="4C434D15843FFF93FAF2CAECFE99F7B8" authorityName="Ovrebo, Hughes and Halling" authorityYear="2019" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="8" pageNumber="41" phylum="Basidiomycota" rank="species" species="costaricense">
|
||
<emphasis id="B937EA84843FFF93FAF2CAECFE99F7B8" italics="true" pageId="8" pageNumber="41">Tricholoma costaricense</emphasis>
|
||
</taxonomicName>
|
||
, holotype, C. basidiospores, D. cheilocystidia. Scale line=10 μm. Drawings by C. L. Ovrebo.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFC3696843EFF92FF72CD87FAC3FE09" blockId="9.[136,1452,160,437]" pageId="9" pageNumber="42">
|
||
Based on morphological features,
|
||
<taxonomicName id="4C434D15843EFF92FD8ECD87FD2EFF05" authorityName="Ovrebo, Hughes and Halling" authorityYear="2019" box="[577,695,160,185]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="9" pageNumber="42" phylum="Basidiomycota" rank="species" species="atratum">
|
||
<emphasis id="B937EA84843EFF92FD8ECD87FD2EFF05" box="[577,695,160,185]" italics="true" pageId="9" pageNumber="42">T. atratum</emphasis>
|
||
</taxonomicName>
|
||
differs from species treated by
|
||
<bibRefCitation id="EFD24B67843EFF92FBEFCD87FB5EFF05" author="Ovrebo, C. L." box="[1056,1223,160,185]" pageId="9" pageNumber="42" pagination="418 - 425" refId="ref6106" refString="Ovrebo, C. L. (1986) Three new species of Tricholoma with a description of Tricholoma luteomaculosum. Mycologia 78 (3): 418 - 425. https: // doi. org / 10.2307 / 3793045" type="journal article" year="1986">Ovrebo (1986)</bibRefCitation>
|
||
by the darker pileal coloration, the lack of cheilocystidia and the larger spores. Of the species treated by
|
||
<bibRefCitation id="EFD24B67843EFF92FBF8CDE3FB45FF61" author="Shanks, K. M." box="[1079,1244,196,221]" pageId="9" pageNumber="42" refId="ref6448" refString="Shanks, K. M. (1997) The Agaricales (gilled fungi) of California. 11. Tricholomataceae II. Tricholoma. Mad River Press, Eureka, California. 54 pp. + 9 pls." type="book" year="1997">Shanks (1997)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C434D15843EFF92FB28CDE3FE1EFEBD" authority="Shanks & Ovrebo (1996: 500)" authorityName="Shanks & Ovrebo" authorityPageNumber="500" authorityYear="1996" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="9" pageNumber="42" phylum="Basidiomycota" rank="species" species="nigrum">
|
||
<emphasis id="B937EA84843EFF92FB28CDE3FACDFF61" box="[1255,1364,196,221]" italics="true" pageId="9" pageNumber="42">T. nigrum</emphasis>
|
||
Shanks & Ovrebo (1996: 500)
|
||
</taxonomicName>
|
||
also has a dark gray pileus, but it possesses cheilocystidia, a pileipellis as an ixocutis and the spores are smaller.
|
||
<taxonomicName id="4C434D15843EFF92FE91CC2BFDAEFE99" box="[350,567,268,293]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="9" pageNumber="42" phylum="Basidiomycota" rank="species" species="felschii">
|
||
<emphasis id="B937EA84843EFF92FE91CC2BFDAEFE99" box="[350,567,268,293]" italics="true" pageId="9" pageNumber="42">Tricholoma felschii</emphasis>
|
||
</taxonomicName>
|
||
has a dark gray pileal coloration, but has cheilocystidia and the pileipellis is an ixocutis.
|
||
<taxonomicName id="4C434D15843EFF92FF3FCC17FD59FEF5" authority="Ovrebo (1989: 3139)" authorityName="Ovrebo" authorityPageNumber="3139" authorityYear="1989" box="[240,704,304,329]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="9" pageNumber="42" phylum="Basidiomycota" rank="species" species="pullum">
|
||
<emphasis id="B937EA84843EFF92FF3FCC17FE51FEF5" box="[240,456,304,329]" italics="true" pageId="9" pageNumber="42">Tricholoma pullum</emphasis>
|
||
<bibRefCitation id="EFD24B67843EFF92FE1FCC17FD59FEF5" author="Ovrebo, C. L." box="[464,704,304,329]" pageId="9" pageNumber="42" pagination="3134 - 3152" refId="ref6147" refString="Ovrebo, C. L. (1989) Tricholoma, subgenus Tricholoma, section Albidogrisea: North American species found principally in the Great Lakes Region. Canadian Journal of Botany 67: 3134 - 3152. https: // doi. org / 10.1139 / b 89 - 393" type="journal article" year="1989">Ovrebo (1989: 3139)</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C434D15843EFF92FD36CC17FB15FEF5" authority="Ovrebo (1989: 3140)" authorityName="Ovrebo" authorityPageNumber="3140" authorityYear="1989" box="[761,1164,304,329]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="9" pageNumber="42" phylum="Basidiomycota" rank="species" species="atrodiscum">
|
||
<emphasis id="B937EA84843EFF92FD36CC17FC0DFEF5" box="[761,916,304,329]" italics="true" pageId="9" pageNumber="42">T. atrodiscum</emphasis>
|
||
<bibRefCitation id="EFD24B67843EFF92FC53CC17FB15FEF5" author="Ovrebo, C. L." box="[924,1164,304,329]" pageId="9" pageNumber="42" pagination="3134 - 3152" refId="ref6147" refString="Ovrebo, C. L. (1989) Tricholoma, subgenus Tricholoma, section Albidogrisea: North American species found principally in the Great Lakes Region. Canadian Journal of Botany 67: 3134 - 3152. https: // doi. org / 10.1139 / b 89 - 393" type="journal article" year="1989">Ovrebo (1989: 3140)</bibRefCitation>
|
||
</taxonomicName>
|
||
have dark gray pilei, but possess cheilocystidia, have a bitter to acrid taste and lack the pseudoparenchymatous subcutis. The dark pileus and yellowish tones of the lamellae are found in
|
||
<taxonomicName id="4C434D15843EFF92FDBCCC5FFB86FE2D" authority="(Fr.) Quel. (1873: 338)" authorityName="Quel." authorityPageNumber="338" authorityYear="1873" baseAuthorityName="Fr." box="[627,1055,376,401]" class="Agaricomycetes" family="Tricholomataceae" genus="Tricholoma" kingdom="Fungi" order="Agaricales" pageId="9" pageNumber="42" phylum="Basidiomycota" rank="species" species="portentosum">
|
||
<emphasis id="B937EA84843EFF92FDBCCC5FFC80FE2D" box="[627,793,376,401]" italics="true" pageId="9" pageNumber="42">T. portentosum</emphasis>
|
||
(Fr.) Quél. (1873: 338)
|
||
</taxonomicName>
|
||
, but the latter is viscid, has stronger yellow coloration to the pileus and stipe, lacks the pseudoparenchymatous subcutis and the spores are smaller.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |