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<document id="C28E9D8EA444E4BB3E4619498F7A4DDF" ID-CLB-Dataset="304139" ID-DOI="10.1590/1982-0224-2022-0002" ID-GBIF-Dataset="1f6c7a0d-e5f8-479d-be08-03fd894b7cfe" ID-ISSN="1982-0224" ID-Zenodo-Dep="13350428" ID-ZooBank="0A755C8D-B807-41CF-825B-BD3209119D54" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1720669642593" checkinUser="felipe" docAuthor="Reis, Roberto E. &amp; Lehmann A., Pablo" docDate="2022" docId="7F0187BBE848427CFDAFFB695F44FB9E" docLanguage="en" docName="NeotropIchthyol.20.2.e220002.pdf" docOrigin="Neotropical Ichthyology (e 220002) 20 (2)" docSource="http://dx.doi.org/10.1590/1982-0224-2022-0002" docStyle="DocumentStyle:FC0C3FD10650AF21F8DA93D98D275823.3:NeotropIchthyol.2020-.journal_article" docStyleId="FC0C3FD10650AF21F8DA93D98D275823" docStyleName="NeotropIchthyol.2020-.journal_article" docStyleVersion="3" docTitle="Rhinotocinclus hardmani Reis &amp; Lehmann A. 2022, new combination" docType="treatment" docVersion="4" lastPageNumber="63" masterDocId="8338FFC3E8744242FFBCFFBA5B18FFAE" masterDocTitle="A new genus of armored catfish (Siluriformes: Loricariidae) from the Greater Amazon, with a review of the species and description of five new species" masterLastPageNumber="100" masterPageNumber="1" pageNumber="61" updateTime="1728051297399" updateUser="julia" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8C4B44747278347E502CBD5C27464CAD">A new genus of armored catfish (Siluriformes: Loricariidae) from the Greater Amazon, with a review of the species and description of five new species</mods:title>
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<mods:name id="78FAE0462C7B4AEAF05E0D6577E2E3D8" type="personal">
<mods:role id="0C24281A2513284E9EA5C3A5D8D31F39">
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<mods:namePart id="3727BA60241350C9F2E66D559FC0FED4">Reis, Roberto E.</mods:namePart>
<mods:affiliation id="71BDDFFBC70D58035BA876288143A3CA">PontifÍcia Universidade Católica do Rio Grande do Sul. Av. Ipiranga, 6681, P. O. Box 1429, 90619 - 900 Porto Alegre, RS, Brazil.</mods:affiliation>
<mods:nameIdentifier id="ECC563FAB44D8B5373C396FB4CF35501" type="email">reis@pucrs.br</mods:nameIdentifier>
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<mods:namePart id="D3D6AA6F3092E00A275C0C884E7D8689">Lehmann A., Pablo</mods:namePart>
<mods:affiliation id="F1CC104FC7D191D266F2E0F25A7FDD96">Laboratório de Ictiologia, Universidade do Vale do Rio dos Sinos. Av. Unisinos, 950, 93022 - 000 São Leopoldo, RS, Brazil.</mods:affiliation>
<mods:nameIdentifier id="D4663E91E14FDD7104BDBD96FF1C823E" type="email">pablole@unisinos.br.</mods:nameIdentifier>
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<mods:title id="5905F497C5CBB381EAD5E04774E4DD32">Neotropical Ichthyology</mods:title>
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<mods:date id="E3D61FF81805E27CCBC02922F4AB89F4">2022</mods:date>
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<mods:number id="7C3D8F57F0DA79F9B11686DFBA709F18">2022-07-08</mods:number>
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<mods:classification id="9103D2479AE23A0B3D7281EBC3806845">journal article</mods:classification>
<mods:identifier id="0E18FB1E7BF8478DF0232AB5611F1298" type="CLB-Dataset">304139</mods:identifier>
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<treatment id="7F0187BBE848427CFDAFFB695F44FB9E" LSID="urn:lsid:plazi:treatment:7F0187BBE848427CFDAFFB695F44FB9E" httpUri="http://treatment.plazi.org/id/7F0187BBE848427CFDAFFB695F44FB9E" lastPageId="62" lastPageNumber="63" pageId="60" pageNumber="61">
<subSubSection id="BFB26526E848427EFDAFFB695E90FB45" box="[531,1416,1235,1259]" pageId="60" pageNumber="61" type="nomenclature">
<paragraph id="F71736ADE848427EFDAFFB695E90FB45" blockId="60.[531,1416,1235,1259]" box="[531,1416,1235,1259]" pageId="60" pageNumber="61">
<taxonomicName id="30A84D2EE848427EFDAFFB695FAEFB45" authority="(Lehmann, Lujan &amp; Reis, 2022)" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Lehmann, Lujan &amp; Reis" baseAuthorityYear="2022" box="[531,1206,1235,1259]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="hardmani" status="comb. nov.">
<emphasis id="C5DCEABFE848427EFDAFFB695853FB45" bold="true" box="[531,843,1235,1259]" pageId="60" pageNumber="61">Rhinotocinclus hardmani</emphasis>
(Lehmann, Lujan &amp; Reis, 2022)
</taxonomicName>
,
<taxonomicNameLabel id="DEEF57C4E848427EFB7DFB6E5E90FB45" box="[1217,1416,1235,1259]" pageId="60" pageNumber="61" rank="species">new combination</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="BFB26526E848427EFCCEFAA45F30FA98" box="[882,1064,1310,1334]" pageId="60" pageNumber="61" type="description">
<paragraph id="F71736ADE848427EFCCEFAA45F30FA98" blockId="60.[882,1064,1310,1334]" box="[882,1064,1310,1334]" pageId="60" pageNumber="61">
(
<figureCitation id="6F932A28E848427EFCC6FAA458D4FA98" box="[890,972,1310,1334]" captionStart="FIGURE 34" captionStartId="61.[136,209,193,213]" captionTargetBox="[488,1459,171,1194]" captionTargetId="figure-205@61.[488,1459,171,1194]" captionTargetPageId="61" captionText="FIGURE 34 | Rhinotocinclus hardmani, holotype, CSBD F3618, 22.7 mm SL, female, Kuribrong River at Grass Shoals Rapids, Potaro-Siparuni, Guyana." figureDoi="http://doi.org/10.5281/zenodo.13887672" httpUri="https://zenodo.org/record/13887672/files/figure.png" pageId="60" pageNumber="61">Fig. 34</figureCitation>
;
<tableCitation id="BA2A0316E848427EFC6BFAA45F39FA98" box="[983,1057,1310,1334]" captionStart="TABLE 6" captionStartId="55.[136,198,193,213]" captionTargetBox="[150,1444,299,1541]" captionTargetId="graphics-51@55.[136,1459,283,1553]" captionTargetPageId="55" captionText="TABLE 6 | Descriptive morphometrics of Rhinotocinclus species. Values given as percent of standard length or head length. Range includes the holotype (Hol), SD = standard deviation." httpUri="http://table.plazi.org/id/A3D76625E8434275FF34FF7B59E6FF55" pageId="60" pageNumber="61" tableUuid="A3D76625E8434275FF34FF7B59E6FF55">Tab. 6</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BFB26526E848427EFE54FAD65E36FA0A" pageId="60" pageNumber="61" type="reference_group">
<paragraph id="F71736ADE848427EFE54FAD65E36FA0A" blockId="60.[488,1459,1388,1445]" pageId="60" pageNumber="61">
<taxonomicName id="30A84D2EE848427EFE54FAD658FEFAD1" authority="Lehmann, Lujan &amp; Reis, 2022: 2" authorityName="Lehmann, Lujan &amp; Reis" authorityPageNumber="2" authorityYear="2022" box="[488,998,1388,1407]" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="hardmani">
<emphasis id="C5DCEABFE848427EFE54FAD659B5FAD1" box="[488,685,1388,1407]" italics="true" pageId="60" pageNumber="61">Parotocinclus hardmani</emphasis>
Lehmann, Lujan &amp; Reis, 2022:2
</taxonomicName>
(Type-locality: Kuribrong River at Grass Shoals Rapids,
<collectingRegion id="356CF84FE848427EFDF1FA2B59C7FA0A" box="[589,735,1425,1444]" country="Guyana" name="Potaro-Siparuni" pageId="60" pageNumber="61">Potaro-Siparuni</collectingRegion>
,
<collectingCountry id="8FBF763DE848427EFD54FA285828FA0B" box="[744,816,1426,1445]" name="Guyana" pageId="60" pageNumber="61">Guyana</collectingCountry>
,
<geoCoordinate id="929C506AE848427EFC85FA2B58ADFA0A" box="[825,949,1425,1444]" degrees="05" direction="north" minutes="28" orientation="latitude" pageId="60" pageNumber="61" precision="1" seconds="44.7" value="5.4790835">05°2844.7”N</geoCoordinate>
<geoCoordinate id="929C506AE848427EFC07FA2B5F27FA0A" box="[955,1087,1425,1444]" degrees="59" direction="west" minutes="31" orientation="longitude" pageId="60" pageNumber="61" precision="1" seconds="54.4" value="-59.531776">59°3154.4”W</geoCoordinate>
.
<typeStatus id="2813880FE848427EFBF4FA2B5FB8FA0A" box="[1096,1184,1425,1444]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
: CSBD F3618).
</paragraph>
</subSubSection>
<subSubSection id="BFB26526E848427FFDBAFA625984F928" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="diagnosis">
<paragraph id="F71736ADE848427FFDBAFA625984F928" blockId="60.[486,1460,1496,1856]" lastBlockId="61.[487,1461,1310,1670]" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
<emphasis id="C5DCEABFE848427EFDBAFA62599EFA5E" bold="true" box="[518,646,1496,1520]" pageId="60" pageNumber="61">Diagnosis.</emphasis>
<taxonomicName id="30A84D2EE848427EFD33FA6258BAFA5E" box="[655,930,1496,1520]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="hardmanni">
<emphasis id="C5DCEABFE848427EFD33FA6258BAFA5E" box="[655,930,1496,1520]" italics="true" pageId="60" pageNumber="61">Rhinotocinclus hardmanni</emphasis>
</taxonomicName>
is distinguished from all congeners, except
<taxonomicName id="30A84D2EE848427EFA27FA635927F9B8" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Schmidt &amp; Ferraris" baseAuthorityYear="1985" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="collinsae">
<emphasis id="C5DCEABFE848427EFA27FA635927F9B8" italics="true" pageId="60" pageNumber="61">R. collinsae</emphasis>
</taxonomicName>
and
<taxonomicName id="30A84D2EE848427EFDC2FA4559E4F9B8" box="[638,764,1534,1558]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="halbolthi">
<emphasis id="C5DCEABFE848427EFDC2FA4559E4F9B8" box="[638,764,1534,1558]" italics="true" pageId="60" pageNumber="61">R. halbolthi</emphasis>
</taxonomicName>
, by having accessory teeth on both premaxilla and dentary (Fig. 5A; accessory teeth absent, Figs. 5B,C); the odontodes on the ventral surface of first pelvic-fin ray aligned with main ray axis (Fig. 9B;
<emphasis id="C5DCEABFE848427EFBF1F9F35F70F9CE" box="[1101,1128,1609,1632]" italics="true" pageId="60" pageNumber="61">vs.</emphasis>
odontodes bent and pointing mesially, Fig. 9A); and lacking a light mark from the snout tip to nostrils (
<emphasis id="C5DCEABFE848427EFAE6F9D55E6CF928" box="[1370,1396,1647,1670]" italics="true" pageId="60" pageNumber="61">vs.</emphasis>
light mark present and Y-, V-shaped or as two separate lines).
<taxonomicName id="30A84D2EE848427EFB39F9295E80F905" box="[1157,1432,1683,1707]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="hardmanni">
<emphasis id="C5DCEABFE848427EFB39F9295E80F905" box="[1157,1432,1683,1707]" italics="true" pageId="60" pageNumber="61">Rhinotocinclus hardmanni</emphasis>
</taxonomicName>
is further distinguished from
<taxonomicName id="30A84D2EE848427EFCA2F9035890F97E" authorityName="Boeseman" authorityYear="1974" box="[798,904,1720,1744]" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="britskii">
<emphasis id="C5DCEABFE848427EFCA2F9035890F97E" box="[798,904,1720,1744]" italics="true" pageId="60" pageNumber="61">R. britskii</emphasis>
</taxonomicName>
,
<taxonomicName id="30A84D2EE848427EFC25F9035F11F97E" box="[921,1033,1720,1744]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="discolor">
<emphasis id="C5DCEABFE848427EFC25F9035F11F97E" box="[921,1033,1720,1744]" italics="true" pageId="60" pageNumber="61">R. discolor</emphasis>
</taxonomicName>
,
<taxonomicName id="30A84D2EE848427EFBA6F9035F9DF97E" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Schaefer &amp; Provenzano" baseAuthorityYear="1993" box="[1050,1157,1720,1744]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="eppleyi">
<emphasis id="C5DCEABFE848427EFBA6F9035F9DF97E" box="[1050,1157,1720,1744]" italics="true" pageId="60" pageNumber="61">R. eppleyi</emphasis>
</taxonomicName>
,
<taxonomicName id="30A84D2EE848427EFB2AF9035E10F97E" box="[1174,1288,1720,1744]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="isabelae">
<emphasis id="C5DCEABFE848427EFB2AF9035E10F97E" box="[1174,1288,1720,1744]" italics="true" pageId="60" pageNumber="61">R. isabelae</emphasis>
</taxonomicName>
,
<taxonomicName id="30A84D2EE848427EFAA5F9035E92F97E" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Lehmann &amp; Reis" baseAuthorityYear="2021" box="[1305,1418,1720,1744]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="kwarup">
<emphasis id="C5DCEABFE848427EFAA5F9035E92F97E" box="[1305,1418,1720,1744]" italics="true" pageId="60" pageNumber="61">R. kwarup</emphasis>
</taxonomicName>
,
<taxonomicName id="30A84D2EE848427EFA27F9035941F958" authorityName="Garavello" authorityYear="1988" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="longirostris">
<emphasis id="C5DCEABFE848427EFA27F9035941F958" italics="true" pageId="60" pageNumber="61">R. longirostris</emphasis>
</taxonomicName>
,
<taxonomicName id="30A84D2EE848427EFDDBF96559D6F958" box="[615,718,1758,1782]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="pilosus">
<emphasis id="C5DCEABFE848427EFDDBF96559D6F958" box="[615,718,1758,1782]" italics="true" pageId="60" pageNumber="61">R. pilosus</emphasis>
</taxonomicName>
,
<taxonomicName id="30A84D2EE848427EFD61F9655874F958" authorityName="Schaefer" authorityYear="1988" box="[733,876,1758,1782]" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="polyochrus">
<emphasis id="C5DCEABFE848427EFD61F9655874F958" box="[733,876,1758,1782]" italics="true" pageId="60" pageNumber="61">R. polyochrus</emphasis>
</taxonomicName>
,
<taxonomicName id="30A84D2EE848427EFCC6F96558F9F958" box="[890,993,1758,1782]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="variola">
<emphasis id="C5DCEABFE848427EFCC6F96558F9F958" box="[890,993,1758,1782]" italics="true" pageId="60" pageNumber="61">R. variola</emphasis>
</taxonomicName>
, and
<taxonomicName id="30A84D2EE848427EFB9CF9655F6AF958" box="[1056,1138,1758,1782]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="yaka">
<emphasis id="C5DCEABFE848427EFB9CF9655F6AF958" box="[1056,1138,1758,1782]" italics="true" pageId="60" pageNumber="61">R. yaka</emphasis>
</taxonomicName>
by lacking a triangular dark spot on the anterior portion of the dorsal-fin membrane (
<emphasis id="C5DCEABFE848427EFBE3F8BE5F61F8B5" box="[1119,1145,1796,1819]" italics="true" pageId="60" pageNumber="61">vs.</emphasis>
dorsal-fin spot present); and from
<taxonomicName id="30A84D2EE848427EFD9AF8935999F8EE" box="[550,641,1833,1856]" family="Loricariidae" genus="Hisonotus" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="acuen">
<emphasis id="C5DCEABFE848427EFD9AF8935999F8EE" box="[550,641,1833,1856]" italics="true" pageId="60" pageNumber="61">R. acuen</emphasis>
</taxonomicName>
,
<taxonomicName id="30A84D2EE848427EFD33F8935807F8EE" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Carvalho &amp; Datovo" baseAuthorityYear="2012" box="[655,799,1832,1856]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="bockmanni">
<emphasis id="C5DCEABFE848427EFD33F8935807F8EE" box="[655,799,1832,1856]" italics="true" pageId="60" pageNumber="61">R. bockmanni</emphasis>
</taxonomicName>
,
<taxonomicName id="30A84D2EE848427EFC92F89358C6F8EE" box="[814,990,1832,1856]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="chromodontus">
<emphasis id="C5DCEABFE848427EFC92F89358C6F8EE" box="[814,990,1832,1856]" italics="true" pageId="60" pageNumber="61">R. chromodontus</emphasis>
</taxonomicName>
,
<taxonomicName id="30A84D2EE848427EFC50F8935F22F8EE" box="[1004,1082,1832,1856]" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="dani">
<emphasis id="C5DCEABFE848427EFC50F8935F22F8EE" box="[1004,1082,1832,1856]" italics="true" pageId="60" pageNumber="61">R. dani</emphasis>
</taxonomicName>
,
<taxonomicName id="30A84D2EE848427EFBF5F8935FAEF8EE" authorityName="Ribeiro-Silva, Silva, Venere, Silva &amp; Roxo" authorityYear="2020" box="[1097,1206,1832,1856]" family="Loricariidae" genus="Hisonotus" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="dinizae">
<emphasis id="C5DCEABFE848427EFBF5F8935FAEF8EE" box="[1097,1206,1832,1856]" italics="true" pageId="60" pageNumber="61">R. dinizae</emphasis>
</taxonomicName>
,
<taxonomicName id="30A84D2EE848427EFB79F8935E09F8EE" authorityName="Gamarra, Calegari &amp; Reis" authorityYear="2019" box="[1221,1297,1832,1856]" family="Loricariidae" genus="Curculionichthys" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="hera">
<emphasis id="C5DCEABFE848427EFB79F8935E09F8EE" box="[1221,1297,1832,1856]" italics="true" pageId="60" pageNumber="61">R. hera</emphasis>
</taxonomicName>
,
<taxonomicName id="30A84D2EE848427EFAA3F8935EB4F8EE" box="[1311,1452,1833,1856]" family="Loricariidae" genus="Hisonotus" kingdom="Animalia" order="Siluriformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="jumaorum">
<emphasis id="C5DCEABFE848427EFAA3F8935EB4F8EE" box="[1311,1452,1833,1856]" italics="true" pageId="60" pageNumber="61">R. jumaorum</emphasis>
</taxonomicName>
,
<taxonomicName id="30A84D2EE849427FFE54FAA55968FA98" authorityName="Reis &amp; Lehmann A." authorityYear="2022" box="[488,624,1310,1334]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="loxochelis" status="n. sp.">
<emphasis id="C5DCEABFE849427FFE54FAA55968FA98" box="[488,624,1310,1334]" italics="true" pageId="61" pageNumber="62">R. loxochelis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEEF57C4E849427FFDC5FAA459A1FA9B" box="[633,697,1310,1333]" pageId="61" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
,
<taxonomicName id="30A84D2EE849427FFD7BFAA5584DFA98" authorityName="Reis &amp; Lehmann A." authorityYear="2022" box="[711,853,1310,1334]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="marginalis" status="n. sp.">
<emphasis id="C5DCEABFE849427FFD7BFAA5584DFA98" box="[711,853,1310,1334]" italics="true" pageId="61" pageNumber="62">R. marginalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEEF57C4E849427FFCE3FAA45886FA9B" box="[863,926,1310,1333]" pageId="61" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
, and
<taxonomicName id="30A84D2EE849427FFC5CFAA55F70FA98" authorityName="Roxo, Messias &amp; Silva" authorityYear="2019" box="[992,1128,1310,1334]" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="pentakelis">
<emphasis id="C5DCEABFE849427FFC5CFAA55F70FA98" box="[992,1128,1310,1334]" italics="true" pageId="61" pageNumber="62">R. pentakelis</emphasis>
</taxonomicName>
by possessing an adipose fin (
<emphasis id="C5DCEABFE849427FFE4CFAFE5912FAF5" box="[496,522,1348,1371]" italics="true" pageId="61" pageNumber="62">vs.</emphasis>
adipose fin absent).
<taxonomicName id="30A84D2EE849427FFD44FAF95F12FAF5" box="[760,1034,1347,1371]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="hardmanni">
<emphasis id="C5DCEABFE849427FFD44FAF95F12FAF5" box="[760,1034,1347,1371]" italics="true" pageId="61" pageNumber="62">Rhinotocinclus hardmanni</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="30A84D2EE849427FFAB6FAFE5E9BFAF5" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Schmidt &amp; Ferraris" baseAuthorityYear="1985" box="[1290,1411,1347,1371]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="collinsae">
<emphasis id="C5DCEABFE849427FFAB6FAFE5E9BFAF5" box="[1290,1411,1347,1371]" italics="true" pageId="61" pageNumber="62">R. collinsae</emphasis>
</taxonomicName>
and
<taxonomicName id="30A84D2EE849427FFE56FAD3597AFA2E" authorityName="Lehmann, Lazzarotto &amp; Reis" authorityYear="2014" box="[490,610,1384,1408]" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="halbothi">
<emphasis id="C5DCEABFE849427FFE56FAD3597AFA2E" box="[490,610,1384,1408]" italics="true" pageId="61" pageNumber="62">R. halbothi</emphasis>
</taxonomicName>
by having few and minute teeth accessory teeth (
<emphasis id="C5DCEABFE849427FFB0EFAD35FD4FA2E" box="[1202,1228,1385,1408]" italics="true" pageId="61" pageNumber="62">vs.</emphasis>
many and large), a large urogenital papilla in males, 34 times bigger than normal (Fig. 2C;
<emphasis id="C5DCEABFE849427FFA91FA355E5FFA08" box="[1325,1351,1423,1446]" italics="true" pageId="61" pageNumber="62">vs.</emphasis>
normally developed urogenital papilla, Fig. 2B), by lacking a skin flap on first pelvic-fin ray of males (
<emphasis id="C5DCEABFE849427FFD88FA635957FA5E" box="[564,591,1497,1520]" italics="true" pageId="61" pageNumber="62">vs.</emphasis>
skin flap present), and having the belly naked or almost naked between lateral abdominal plates (
<emphasis id="C5DCEABFE849427FFD0AFA4559C8F9B8" box="[694,720,1535,1558]" italics="true" pageId="61" pageNumber="62">vs.</emphasis>
fully plated belly). It is further distinguished from
<taxonomicName id="30A84D2EE849427FFAABFA455E88F9B8" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Schmidt &amp; Ferraris" baseAuthorityYear="1985" box="[1303,1424,1534,1558]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="collinsae">
<emphasis id="C5DCEABFE849427FFAABFA455E88F9B8" box="[1303,1424,1534,1558]" italics="true" pageId="61" pageNumber="62">R. collinsae</emphasis>
</taxonomicName>
by having the adipose-fin spine coalesced to the dorsum (Fig. 4C;
<emphasis id="C5DCEABFE849427FFB11F99E5FDFF995" box="[1197,1223,1572,1595]" italics="true" pageId="61" pageNumber="62">vs.</emphasis>
adipose fin normally developed, Figs. 4A,B), and from
<taxonomicName id="30A84D2EE849427FFCE2F9F358C9F9CE" authorityName="Lehmann, Lazzarotto &amp; Reis" authorityYear="2014" box="[862,977,1608,1632]" family="Loricariidae" genus="Parotocinclus" kingdom="Animalia" order="Siluriformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="halbothi">
<emphasis id="C5DCEABFE849427FFCE2F9F358C9F9CE" box="[862,977,1608,1632]" italics="true" pageId="61" pageNumber="62">R. halbothi</emphasis>
</taxonomicName>
by having fewer premaxillary teeth (1822
<emphasis id="C5DCEABFE849427FFE5BF9D55919F928" box="[487,513,1647,1670]" italics="true" pageId="61" pageNumber="62">vs.</emphasis>
2832 teeth).
</paragraph>
</subSubSection>
<caption id="A3D76625E849427FFF34FF7B5A6FFEC5" ID-DOI="http://doi.org/10.5281/zenodo.13887672" ID-Zenodo-Dep="13887672" httpUri="https://zenodo.org/record/13887672/files/figure.png" pageId="61" pageNumber="62" startId="61.[136,209,193,213]" targetBox="[488,1459,171,1194]" targetPageId="61">
<paragraph id="F71736ADE849427FFF34FF7B5A6FFEC5" blockId="61.[136,441,193,363]" pageId="61" pageNumber="62">
<emphasis id="C5DCEABFE849427FFF34FF7B5BC7FF55" bold="true" pageId="61" pageNumber="62">
<collectionCode id="91B9AE68E849427FFF34FF7B5BC9FF7B" box="[136,209,193,213]" pageId="61" pageNumber="62">FIGURE</collectionCode>
34 |
<taxonomicName id="30A84D2EE849427FFEB4FF7B5BC7FF55" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Lehmann, Lujan &amp; Reis" baseAuthorityYear="2022" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="hardmani">
<emphasis id="C5DCEABFE849427FFEB4FF7B5BC7FF55" bold="true" italics="true" pageId="61" pageNumber="62">Rhinotocinclus hardmani</emphasis>
</taxonomicName>
</emphasis>
,
<typeStatus id="2813880FE849427FFF55FF5D5A25FF55" box="[233,317,231,251]" pageId="61" pageNumber="62" type="holotype">holotype</typeStatus>
,
<collectionCode id="91B9AE68E849427FFEF9FF5D5A62FF55" box="[325,378,231,251]" pageId="61" pageNumber="62">CSBD</collectionCode>
F3618,
<quantity id="30509B48E849427FFF34FEB65BCDFE8E" box="[136,213,268,288]" metricMagnitude="-2" metricUnit="m" metricValue="2.27" pageId="61" pageNumber="62" unit="mm" value="22.7">22.7 mm</quantity>
SL, female, Kuribrong River at Grass Shoals Rapids,
<collectingRegion id="356CF84FE849427FFF34FEED5A07FEC5" box="[136,287,343,363]" country="Guyana" name="Potaro-Siparuni" pageId="61" pageNumber="62">Potaro-Siparuni</collectingRegion>
,
<collectingCountry id="8FBF763DE849427FFE94FEED5A6BFEC5" box="[296,371,343,363]" name="Guyana" pageId="61" pageNumber="62">Guyana</collectingCountry>
.
</paragraph>
</caption>
<subSubSection id="BFB26526E849427FFDBAF9025E33F958" pageId="61" pageNumber="62" type="distribution">
<paragraph id="F71736ADE849427FFDBAF9025E33F958" blockId="61.[488,1459,1720,1782]" pageId="61" pageNumber="62">
<emphasis id="C5DCEABFE849427FFDBAF9025854F97E" bold="true" box="[518,844,1720,1744]" pageId="61" pageNumber="62">Geographical distribution.</emphasis>
<taxonomicName id="30A84D2EE849427FFCE5F9025F7BF97E" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Lehmann, Lujan &amp; Reis" baseAuthorityYear="2022" box="[857,1123,1720,1744]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="hardmani">
<emphasis id="C5DCEABFE849427FFCE5F9025F7BF97E" box="[857,1123,1720,1744]" italics="true" pageId="61" pageNumber="62">Rhinotocinclus hardmani</emphasis>
</taxonomicName>
occurs in the Potaro River,
<collectingCountry id="8FBF763DE849427FFE54F9645940F958" box="[488,600,1758,1782]" name="Guyana" pageId="61" pageNumber="62">Essequibo</collectingCountry>
River basin in the Region of
<collectingRegion id="356CF84FE849427FFC1FF9645F41F958" box="[931,1113,1758,1782]" country="Guyana" name="Potaro-Siparuni" pageId="61" pageNumber="62">Potaro-Siparuni</collectingRegion>
,
<collectingCountry id="8FBF763DE849427FFBD9F9645FA5F958" box="[1125,1213,1758,1782]" name="Guyana" pageId="61" pageNumber="62">Guyana</collectingCountry>
(
<figureCitation id="6F932A28E849427FFB70F9645E07F958" box="[1228,1311,1758,1782]" captionStart="FIGURE 32" captionStartId="58.[136,209,1276,1296]" captionTargetBox="[138,1450,171,1231]" captionTargetId="figure-156@58.[136,1459,171,1244]" captionTargetPageId="58" captionText="FIGURE 32 | Geographical distribution of Rhinotocinclus species of the R. collinsae group in southeastern Guiana Shield. Rhinotocinclus collinsae (yellow dots); R. halbothi (red dots); R. hardmani (blue dots). T = Type-locality" figureDoi="http://doi.org/10.5281/zenodo.13887666" httpUri="https://zenodo.org/record/13887666/files/figure.png" pageId="61" pageNumber="62">Fig. 32</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BFB26526E849427CFDBAF8935F74FDAE" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62" type="discussion">
<paragraph id="F71736ADE849427CFDBAF8935F74FDAE" blockId="61.[489,1460,1832,1894]" lastBlockId="62.[487,1460,190,513]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
<emphasis id="C5DCEABFE849427FFDBAF8935961F8EF" bold="true" box="[518,633,1833,1857]" pageId="61" pageNumber="62">Remarks.</emphasis>
<taxonomicName id="30A84D2EE849427FFD3DF892589EF8EE" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Lehmann, Lujan &amp; Reis" baseAuthorityYear="2022" box="[641,902,1832,1856]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="hardmani">
<emphasis id="C5DCEABFE849427FFD3DF892589EF8EE" box="[641,902,1832,1856]" italics="true" pageId="61" pageNumber="62">Rhinotocinclus hardmani</emphasis>
</taxonomicName>
is sympatric and syntopic with
<taxonomicName id="30A84D2EE849427FFB4EF8935E72F8EE" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Schmidt &amp; Ferraris" baseAuthorityYear="1985" box="[1266,1386,1832,1856]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="collinsae">
<emphasis id="C5DCEABFE849427FFB4EF8935E72F8EE" box="[1266,1386,1832,1856]" italics="true" pageId="61" pageNumber="62">R. collinsae</emphasis>
</taxonomicName>
in the Potaro River, having been collected together at the same spot. Despite both species are superficially similar, they have many distinguishing characters.
<taxonomicName id="30A84D2EE84A427CFB0CFF045EACFF78" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Lehmann, Lujan &amp; Reis" baseAuthorityYear="2022" box="[1200,1460,190,214]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="hardmani">
<emphasis id="C5DCEABFE84A427CFB0CFF045EACFF78" box="[1200,1460,190,214]" italics="true" pageId="62" pageNumber="63">Rhinotocinclus hardmani</emphasis>
</taxonomicName>
is easily diagnosed from
<taxonomicName id="30A84D2EE84A427CFCBFFF5E5864FF55" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Schmidt &amp; Ferraris" baseAuthorityYear="1985" box="[771,892,227,251]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="collinsae">
<emphasis id="C5DCEABFE84A427CFCBFFF5E5864FF55" box="[771,892,227,251]" italics="true" pageId="62" pageNumber="63">R. collinsae</emphasis>
</taxonomicName>
by having few and minute teeth accessory teeth, while they are numerous and large in
<taxonomicName id="30A84D2EE84A427CFC16FEB35F3CFE8E" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Schmidt &amp; Ferraris" baseAuthorityYear="1985" box="[938,1060,264,288]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="collinsae">
<emphasis id="C5DCEABFE84A427CFC16FEB35F3CFE8E" box="[938,1060,264,288]" italics="true" pageId="62" pageNumber="63">R. collinsae</emphasis>
</taxonomicName>
, by the large urogenital papilla of males, which is 34 times bigger than that of
<taxonomicName id="30A84D2EE84A427CFC61FE955F4BFEE8" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Schmidt &amp; Ferraris" baseAuthorityYear="1985" box="[989,1107,302,326]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="collinsae">
<emphasis id="C5DCEABFE84A427CFC61FE955F4BFEE8" box="[989,1107,302,326]" italics="true" pageId="62" pageNumber="63">R. collinsae</emphasis>
</taxonomicName>
, by males lacking a skin flap on first pelvic-fin ray, which is present in
<taxonomicName id="30A84D2EE84A427CFC27FEEE5F0AFEC5" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Schmidt &amp; Ferraris" baseAuthorityYear="1985" box="[923,1042,339,363]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="collinsae">
<emphasis id="C5DCEABFE84A427CFC27FEEE5F0AFEC5" box="[923,1042,339,363]" italics="true" pageId="62" pageNumber="63">R. collinsae</emphasis>
</taxonomicName>
, by having the belly naked or almost naked between lateral abdominal plates, while it is fully plated in
<taxonomicName id="30A84D2EE84A427CFB67FEC35E4CFE3E" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Schmidt &amp; Ferraris" baseAuthorityYear="1985" box="[1243,1364,376,400]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="collinsae">
<emphasis id="C5DCEABFE84A427CFB67FEC35E4CFE3E" box="[1243,1364,376,400]" italics="true" pageId="62" pageNumber="63">R. collinsae</emphasis>
</taxonomicName>
, and by having the adipose-fin spine coalesced to the dorsum, while it is free from the dorsum on
<taxonomicName id="30A84D2EE84A427CFDACFE7E5992FE75" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Schmidt &amp; Ferraris" baseAuthorityYear="1985" box="[528,650,451,475]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="collinsae">
<emphasis id="C5DCEABFE84A427CFDACFE7E5992FE75" box="[528,650,451,475]" italics="true" pageId="62" pageNumber="63">R. collinsae</emphasis>
</taxonomicName>
.
<taxonomicName id="30A84D2EE84A427CFD27FE7958BAFE75" authorityName="Reis &amp; Lehmann A." authorityYear="2022" baseAuthorityName="Lehmann, Lujan &amp; Reis" baseAuthorityYear="2022" box="[667,930,451,475]" family="Loricariidae" genus="Rhinotocinclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="hardmani">
<emphasis id="C5DCEABFE84A427CFD27FE7958BAFE75" box="[667,930,451,475]" italics="true" pageId="62" pageNumber="63">Rhinotocinclus hardmani</emphasis>
</taxonomicName>
was tentatively categorized as Least Concern (LC) by Lehmann
<emphasis id="C5DCEABFE84A427CFD06FE5359F4FDAE" box="[698,748,488,512]" italics="true" pageId="62" pageNumber="63">et al.</emphasis>
(2022) in the original description.
</paragraph>
</subSubSection>
<subSubSection id="BFB26526E84A427CFDBAFD8F5F44FB9E" pageId="62" pageNumber="63" type="materials_examined">
<paragraph id="F71736ADE84A427CFDBAFD8F5F44FB9E" blockId="62.[486,1461,565,1072]" pageId="62" pageNumber="63">
<materialsCitation id="47C03CF0E84A427CFDBAFD8F5EADFDE5" ID-GBIF-Occurrence="4949514304" box="[518,1461,565,587]" collectionCode="CSBD" country="Guyana" location="Essequibo River basin" pageId="62" pageNumber="63" specimenCode="CSBD F3618" specimenCount="1" stateProvince="Essequibo River basin" typeStatus="holotype">
<emphasis id="C5DCEABFE84A427CFDBAFD8F5F26FDE5" bold="true" box="[518,1086,565,587]" pageId="62" pageNumber="63">
Material examined.
<collectingRegion id="356CF84FE84A427CFD5EFD8F58CCFDE5" box="[738,980,565,587]" pageId="62" pageNumber="63">Essequibo River basin</collectingRegion>
,
<collectingCountry id="8FBF763DE84A427CFC5DFD8F5F21FDE5" box="[993,1081,565,587]" name="Guyana" pageId="62" pageNumber="63">Guyana</collectingCountry>
:
</emphasis>
<specimenCode id="A70E9ED6E84A427CFBFBFD8F5FCBFDE5" box="[1095,1235,565,587]" pageId="62" pageNumber="63">CSBD F3618</specimenCode>
,
<typeStatus id="2813880FE84A427CFB63FD8F5E20FDE5" box="[1247,1336,565,587]" pageId="62" pageNumber="63" type="holotype">holotype</typeStatus>
(measured),
</materialsCitation>
<materialsCitation id="47C03CF0E84A427CFE55FDE15860FDDE" ID-GBIF-Occurrence="4949514320" box="[489,888,602,624]" collectionCode="AUM" pageId="62" pageNumber="63" specimenCode="AUM 62850" specimenCount="78" typeStatus="paratype">
<specimenCode id="A70E9ED6E84A427CFE55FDE15971FDDE" box="[489,617,602,624]" pageId="62" pageNumber="63">AUM 62850</specimenCode>
,
<specimenCount id="E1AEFD24E84A427CFDCEFDE05993FDC1" box="[626,651,602,623]" pageId="62" pageNumber="63" type="generic">78</specimenCount>
<typeStatus id="2813880FE84A427CFD33FDE159F5FDDE" box="[655,749,603,624]" pageId="62" pageNumber="63">paratypes</typeStatus>
(9 measured),
</materialsCitation>
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<specimenCode id="A70E9ED6E84A427CFCC0FDE05F1CFDDE" box="[892,1028,602,624]" pageId="62" pageNumber="63">CSBD F3619</specimenCode>
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,
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<specimenCode id="A70E9ED6E84A427CFB2FFDE05E17FDDE" box="[1171,1295,602,624]" pageId="62" pageNumber="63">MCP 54588</specimenCode>
, 17 (9 measured) + 3 cs
<typeStatus id="2813880FE84A427CFD97FD3A5994FD3B" box="[555,652,640,661]" pageId="62" pageNumber="63" type="paratype">paratypes</typeStatus>
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,
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<specimenCode id="A70E9ED6E84A427CFD24FD3A5831FD3B" box="[664,809,640,661]" pageId="62" pageNumber="63">ROM 110801</specimenCode>
,
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<typeStatus id="2813880FE84A427CFCEBFD3A58A0FD3B" box="[855,952,640,661]" pageId="62" pageNumber="63">paratypes</typeStatus>
,
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<materialsCitation id="47C03CF0E84A427CFC78FD3A5FA7FD15" ID-GBIF-Occurrence="4949514328" collectionCode="UMMZ" latitude="5.4790835" location="Kuribrong River at Grass Shoals Rapids" longLatPrecision="1" longitude="-59.531776" pageId="62" pageNumber="63" specimenCode="UMMZ 252792" specimenCount="10" stateProvince="Potaro-Siparuni" typeStatus="paratype">
<specimenCode id="A70E9ED6E84A427CFC78FD3A5F72FD38" box="[964,1130,640,662]" pageId="62" pageNumber="63">UMMZ 252792</specimenCode>
,
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<typeStatus id="2813880FE84A427CFB24FD3A5FE1FD3B" box="[1176,1273,640,661]" pageId="62" pageNumber="63">paratypes</typeStatus>
,
<location id="F2776076E84A427CFAB9FD3A59CAFD15" LSID="urn:lsid:plazi:treatment:7F0187BBE848427CFDAFFB695F44FB9E:F2776076E84A427CFAB9FD3A59CAFD15" latitude="5.4790835" longLatPrecision="1" longitude="-59.531776" name="Kuribrong River at Grass Shoals Rapids" pageId="62" pageNumber="63" stateProvince="Potaro-Siparuni">Kuribrong River at Grass Shoals Rapids</location>
,
<collectingRegion id="356CF84FE84A427CFD63FD1F589BFD15" box="[735,899,677,699]" country="Guyana" name="Potaro-Siparuni" pageId="62" pageNumber="63">Potaro-Siparuni</collectingRegion>
,
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<geoCoordinate id="929C506AE84A427CFB9AFD1F5FA3FD15" box="[1062,1211,677,699]" degrees="59" direction="west" minutes="31" orientation="longitude" pageId="62" pageNumber="63" precision="1" seconds="54.4" value="-59.531776">59°3154.4”W</geoCoordinate>
.
</materialsCitation>
<materialsCitation id="47C03CF0E84A427CFB7BFD1F5EACFD4E" ID-GBIF-Occurrence="4949514305" collectionCode="AUM" latitude="-5.4423614" location="Kuribrong River at Ram Sheep Rapids" longLatPrecision="1" longitude="-59.502" pageId="62" pageNumber="63" specimenCode="AUM 62879" specimenCount="33" stateProvince="Potaro-Siparuni" typeStatus="paratype">
<specimenCode id="A70E9ED6E84A427CFB7BFD1F5E55FD15" box="[1223,1357,677,699]" pageId="62" pageNumber="63">AUM 62879</specimenCode>
, 17 of
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<typeStatus id="2813880FE84A427CFE54FD715951FD4E" box="[488,585,715,736]" pageId="62" pageNumber="63">paratypes</typeStatus>
,
<location id="F2776076E84A427CFDEEFD7058C1FD4E" LSID="urn:lsid:plazi:treatment:7F0187BBE848427CFDAFFB695F44FB9E:F2776076E84A427CFDEEFD7058C1FD4E" box="[594,985,714,736]" latitude="-5.4423614" longLatPrecision="1" longitude="-59.502" name="Kuribrong River at Ram Sheep Rapids" pageId="62" pageNumber="63" stateProvince="Potaro-Siparuni">Kuribrong River at Ram Sheep Rapids</location>
,
<collectingRegion id="356CF84FE84A427CFC5FFD705F9FFD4E" box="[995,1159,714,736]" country="Guyana" name="Potaro-Siparuni" pageId="62" pageNumber="63">Potaro-Siparuni</collectingRegion>
,
<geoCoordinate id="929C506AE84A427CFB2DFD705E0DFD4E" box="[1169,1301,714,736]" degrees="05" direction="south" minutes="26" orientation="latitude" pageId="62" pageNumber="63" precision="1" seconds="32.5" value="-5.4423614">05°2632.5”S</geoCoordinate>
<geoCoordinate id="929C506AE84A427CFAA7FD705EB7FD4E" box="[1307,1455,714,736]" degrees="59" direction="west" minutes="30" orientation="longitude" pageId="62" pageNumber="63" precision="1" seconds="07.2" value="-59.502">59°3007.2”W</geoCoordinate>
.
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<materialsCitation id="47C03CF0E84A427CFE54FD4A5FC1FC85" ID-GBIF-Occurrence="4949514325" collectionCode="AUM" latitude="-5.4065" location="Falls Creek (Kiwikparu Creek), just upstream from mouth of Kuribrong River" longLatPrecision="15" longitude="-59.53361" pageId="62" pageNumber="63" specimenCode="AUM 62895" specimenCount="26" stateProvince="Potaro-Siparuni" typeStatus="paratype">
<specimenCode id="A70E9ED6E84A427CFE54FD4A5976FCA8" box="[488,622,752,774]" pageId="62" pageNumber="63">AUM 62895</specimenCode>
, 14 of
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<typeStatus id="2813880FE84A427CFD58FD4A585DFCAB" box="[740,837,752,773]" pageId="62" pageNumber="63">paratypes</typeStatus>
, Grass
<location id="F2776076E84A427CFC29FD4A581DFC85" LSID="urn:lsid:plazi:treatment:7F0187BBE848427CFDAFFB695F44FB9E:F2776076E84A427CFC29FD4A581DFC85" latitude="-5.4065" longLatPrecision="15" longitude="-59.53361" name="Falls Creek (Kiwikparu Creek), just upstream from mouth of Kuribrong River" pageId="62" pageNumber="63" stateProvince="Potaro-Siparuni">Falls Creek (Kiwikparu Creek), just upstream from mouth of Kuribrong River</location>
,
<collectingRegion id="356CF84FE84A427CFCAFFCAF58AFFC85" box="[787,951,789,811]" country="Guyana" name="Potaro-Siparuni" pageId="62" pageNumber="63">Potaro-Siparuni</collectingRegion>
,
<geoCoordinate id="929C506AE84A427CFC79FCAF5F50FC85" box="[965,1096,789,811]" degrees="05" direction="south" minutes="24" orientation="latitude" pageId="62" pageNumber="63" precision="1" seconds="23.4" value="-5.4065">05°2423.4”S</geoCoordinate>
<geoCoordinate id="929C506AE84A427CFBEFFCAF5FCDFC85" box="[1107,1237,789,811]" degrees="59" direction="west" minutes="32" orientation="longitude" pageId="62" pageNumber="63" precision="15" seconds="01" value="-59.53361">59°3201”W</geoCoordinate>
.
</materialsCitation>
<materialsCitation id="47C03CF0E84A427CFB5FFCAF5EACFCFE" ID-GBIF-Occurrence="4949514343" collectionCode="INHS" latitude="5.30375" location="Potaro River at Amatuk Cataract" longLatPrecision="1" longitude="-59.311165" pageId="62" pageNumber="63" specimenCode="INHS 49522" specimenCount="1" stateProvince="Potaro-Siparuni" typeStatus="paratype">
<specimenCode id="A70E9ED6E84A427CFB5FFCAF5E73FC85" box="[1251,1387,789,811]" pageId="62" pageNumber="63">INHS 49522</specimenCode>
, 5 + 1 cs
<typeStatus id="2813880FE84A427CFDB8FC81597DFCFE" box="[516,613,827,848]" pageId="62" pageNumber="63" type="paratype">paratypes</typeStatus>
,
<location id="F2776076E84A427CFDCDFC8158D1FCE1" LSID="urn:lsid:plazi:treatment:7F0187BBE848427CFDAFFB695F44FB9E:F2776076E84A427CFDCDFC8158D1FCE1" box="[625,969,826,848]" latitude="5.30375" longLatPrecision="1" longitude="-59.311165" name="Potaro River at Amatuk Cataract" pageId="62" pageNumber="63" stateProvince="Potaro-Siparuni">Potaro River at Amatuk Cataract</location>
,
<collectingRegion id="356CF84FE84A427CFC6AFC805F62FCFE" box="[982,1146,826,848]" country="Guyana" name="Potaro-Siparuni" pageId="62" pageNumber="63">Potaro-Siparuni</collectingRegion>
,
<geoCoordinate id="929C506AE84A427CFB3BFC805E0AFCFE" box="[1159,1298,826,848]" degrees="05" direction="north" minutes="18" orientation="latitude" pageId="62" pageNumber="63" precision="1" seconds="13.5" value="5.30375">05°1813.5”N</geoCoordinate>
<geoCoordinate id="929C506AE84A427CFAA7FC805EA8FCFE" box="[1307,1456,826,848]" degrees="59" direction="west" minutes="18" orientation="longitude" pageId="62" pageNumber="63" precision="1" seconds="40.2" value="-59.311165">59°1840.2”W</geoCoordinate>
.
</materialsCitation>
<materialsCitation id="47C03CF0E84A427CFE54FCDA58EDFCDB" ID-GBIF-Occurrence="4949514330" box="[488,1013,864,886]" collectionCode="INHS" pageId="62" pageNumber="63" specimenCode="INHS 49555" specimenCount="2" typeStatus="paratype">
<specimenCode id="A70E9ED6E84A427CFE54FCDA5974FCD8" box="[488,620,864,886]" pageId="62" pageNumber="63">INHS 49555</specimenCode>
,
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<typeStatus id="2813880FE84A427CFD37FCDA59F4FCDB" box="[651,748,864,885]" pageId="62" pageNumber="63">paratypes</typeStatus>
,
<quantity id="30509B48E84A427CFD4BFCDA5893FCDB" box="[759,907,864,885]" metricMagnitude="-2" metricUnit="m" metricValue="2.095" metricValueMax="2.28" metricValueMin="1.91" pageId="62" pageNumber="63" unit="mm" value="20.95" valueMax="22.8" valueMin="19.1">19.122.8 mm</quantity>
SL + 1 cs,
</materialsCitation>
<materialsCitation id="47C03CF0E84A427CFC41FCDA5E1EFC34" ID-GBIF-Occurrence="4949514306" collectionCode="INHS" country="Guyana" latitude="5.258861" location="Potaro River at Waratuk Cataract" longLatPrecision="1" longitude="-59.40028" pageId="62" pageNumber="63" specimenCode="INHS 49556" specimenCount="1" stateProvince="Potaro-Siparuni">
<specimenCode id="A70E9ED6E84A427CFC41FCDA5F99FCD8" box="[1021,1153,864,886]" pageId="62" pageNumber="63">INHS 49556</specimenCode>
,
<specimenCount id="E1AEFD24E84A427CFB30FCDA5F80FCDB" box="[1164,1176,864,885]" pageId="62" pageNumber="63">1</specimenCount>
,
<quantity id="30509B48E84A427CFB1AFCDA5E1AFCDB" box="[1190,1282,864,886]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="62" pageNumber="63" unit="mm" value="23.5">23.5 mm</quantity>
SL,
<location id="F2776076E84A427CFA8DFCDA59A3FC34" LSID="urn:lsid:plazi:treatment:7F0187BBE848427CFDAFFB695F44FB9E:F2776076E84A427CFA8DFCDA59A3FC34" country="Guyana" latitude="5.258861" longLatPrecision="1" longitude="-59.40028" name="Potaro River at Waratuk Cataract" pageId="62" pageNumber="63" stateProvince="Potaro-Siparuni">Potaro River at Waratuk Cataract</location>
,
<collectingRegion id="356CF84FE84A427CFD7AFC3F5872FC35" box="[710,874,901,923]" country="Guyana" name="Potaro-Siparuni" pageId="62" pageNumber="63">Potaro-Siparuni</collectingRegion>
,
<collectingCountry id="8FBF763DE84A427CFCC9FC3F58DFFC34" box="[885,967,901,922]" name="Guyana" pageId="62" pageNumber="63">Guyana</collectingCountry>
,
<geoCoordinate id="929C506AE84A427CFC6EFC3F5F46FC35" box="[978,1118,901,923]" degrees="05" direction="north" minutes="15" orientation="latitude" pageId="62" pageNumber="63" precision="1" seconds="31.9" value="5.258861">05°1531.9”N</geoCoordinate>
<geoCoordinate id="929C506AE84A427CFBDAFC3F5E1AFC34" box="[1126,1282,901,923]" degrees="59" direction="west" minutes="24" orientation="longitude" pageId="62" pageNumber="63" precision="1" seconds="01.0" value="-59.40028">59°24 01.0”W</geoCoordinate>
.
</materialsCitation>
<materialsCitation id="47C03CF0E84A427CFAB1FC3F5E0CFC48" ID-GBIF-Occurrence="4949514311" collectionCode="ROM" latitude="5.4139585" location="Mikobe Creek" longLatPrecision="1" longitude="-59.470253" pageId="62" pageNumber="63" specimenCode="ROM 91423" specimenCount="17" stateProvince="Potaro-Siparuni" typeStatus="paratype">
<specimenCode id="A70E9ED6E84A427CFAB1FC3F5E88FC34" box="[1293,1424,901,922]" pageId="62" pageNumber="63">ROM 91423</specimenCode>
,
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<typeStatus id="2813880FE84A427CFE54FC115951FC6E" box="[488,585,939,960]" pageId="62" pageNumber="63">paratypes</typeStatus>
,
<location id="F2776076E84A427CFDEAFC1059F6FC6E" LSID="urn:lsid:plazi:treatment:7F0187BBE848427CFDAFFB695F44FB9E:F2776076E84A427CFDEAFC1059F6FC6E" box="[598,750,938,960]" latitude="5.4139585" longLatPrecision="1" longitude="-59.470253" name="Mikobe Creek" pageId="62" pageNumber="63" stateProvince="Potaro-Siparuni">Mikobe Creek</location>
,
<locationDeviation id="E6A52F8FE84A427CFD40FC115800FC4B" pageId="62" pageNumber="63">
approx.
<quantity id="30509B48E84A427CFCEDFC105883FC6E" box="[849,923,938,960]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="62" pageNumber="63" unit="km" value="0.5">0.5 km</quantity>
upstream from mouth, at rapids beyond first rapid blocking upstream boat entry
</locationDeviation>
,
<collectingRegion id="356CF84FE84A427CFC98FC6A58D0FC48" box="[804,968,976,998]" country="Guyana" name="Potaro-Siparuni" pageId="62" pageNumber="63">Potaro-Siparuni</collectingRegion>
,
<geoCoordinate id="929C506AE84A427CFC68FC6A5F73FC48" box="[980,1131,976,998]" degrees="05" direction="north" minutes="24" orientation="latitude" pageId="62" pageNumber="63" precision="1" seconds="50.25" value="5.4139585">05°2450.25”N</geoCoordinate>
<geoCoordinate id="929C506AE84A427CFBCFFC6A5E0CFC48" box="[1139,1300,976,998]" degrees="59" direction="west" minutes="28" orientation="longitude" pageId="62" pageNumber="63" precision="1" seconds="12.91" value="-59.470253">59°2812.91”W</geoCoordinate>
</materialsCitation>
.
<materialsCitation id="47C03CF0E84A427CFAA3FC6A5F44FB9E" ID-GBIF-Occurrence="4949514318" collectionCode="ROM" latitude="5.407375" location="creek entering Kuribrong River at upper Grass Falls, at lowermost series of rapids near mouth" longLatPrecision="1" longitude="-59.532513" pageId="62" pageNumber="63" specimenCode="ROM 111040" specimenCount="49" stateProvince="Potaro-Siparuni" typeStatus="paratype">
<specimenCode id="A70E9ED6E84A427CFAA3FC6A5EB7FC4B" box="[1311,1455,976,997]" pageId="62" pageNumber="63">ROM 111040</specimenCode>
,
<specimenCount id="E1AEFD24E84A427CFE54FC4F5919FBA4" box="[488,513,1013,1034]" pageId="62" pageNumber="63" type="generic">49</specimenCount>
<typeStatus id="2813880FE84A427CFDB4FC4F5971FBA4" box="[520,617,1013,1034]" pageId="62" pageNumber="63">paratypes</typeStatus>
,
<location id="F2776076E84A427CFDCFFC4F5978FB9E" LSID="urn:lsid:plazi:treatment:7F0187BBE848427CFDAFFB695F44FB9E:F2776076E84A427CFDCFFC4F5978FB9E" latitude="5.407375" longLatPrecision="1" longitude="-59.532513" name="creek entering Kuribrong River at upper Grass Falls, at lowermost series of rapids near mouth" pageId="62" pageNumber="63" stateProvince="Potaro-Siparuni">creek entering Kuribrong River at upper Grass Falls, at lowermost series of rapids near mouth</location>
,
<collectingRegion id="356CF84FE84A427CFDD6FBA05816FB9E" box="[618,782,1050,1072]" country="Guyana" name="Potaro-Siparuni" pageId="62" pageNumber="63">Potaro-Siparuni</collectingRegion>
,
<geoCoordinate id="929C506AE84A427CFCA5FBA058A8FB9E" box="[793,944,1050,1072]" degrees="05" direction="north" minutes="24" orientation="latitude" pageId="62" pageNumber="63" precision="1" seconds="26.55" value="5.407375">05°2426.55”N</geoCoordinate>
<geoCoordinate id="929C506AE84A427CFC0BFBA05F40FB9E" box="[951,1112,1050,1072]" degrees="59" direction="west" minutes="31" orientation="longitude" pageId="62" pageNumber="63" precision="1" seconds="57.05" value="-59.532513">59°3157.05”W</geoCoordinate>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>