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<mods:title id="85B0F8B076F9B6F6FAF3F7820245A120">Redescription of the Holotype of Argulus floridensis Meehean, 1940</mods:title>
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<taxonomicName id="2CE981E59776FFBDFFDFFEB5D22BC078" authority="Meehean, 1940" authorityName="Meehean" authorityYear="1940" box="[123,512,351,375]" class="Maxillopoda" family="Argulidae" genus="Argulus" kingdom="Animalia" order="Arguloida" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="floridensis">
<emphasis id="D99D26749776FFBDFFDFFEB5D22BC078" bold="true" box="[123,512,351,375]" pageId="1" pageNumber="134">
<emphasis id="D99D26749776FFBDFFDFFEB5D164C078" bold="true" box="[123,335,351,375]" italics="true" pageId="1" pageNumber="134">Argulus floridensis</emphasis>
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</emphasis>
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<collectingRegion id="292D34849776FFBDFFDFFE95D0EBC09A" box="[123,192,383,405]" country="United States of America" name="Florida" pageId="1" pageNumber="134">Florida</collectingRegion>
fishlouse Figures 16.
</paragraph>
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.—
<typeStatus id="345244C49776FFBDFE1EFE42D236C0CF" box="[442,541,424,448]" pageId="1" pageNumber="134" type="holotype">Holotype</typeStatus>
(unique), adult male,
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total length [
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Meehean 1940:
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.85 mm],
<collectionCode id="8DF862A39776FFBDFF50FE22D16AC0EF" box="[244,321,456,480]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="134" type="Museum">USNM</collectionCode>
77810 [see note], Key West,
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, 1884, collector: A.E. Verrill [= Addison Emery Verrill]. Note:
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, p. 467) listed the catalog number as “77810 (old number 60589);” mentioned here in case of relevant information in historical records under the old number.
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<emphasis id="D99D26749776FFBDFF0EFDA2D10BC350" bold="true" box="[170,288,584,608]" pageId="1" pageNumber="134">DIAGNOSIS</emphasis>
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.— Two respiratory areas with smaller oblong “area” anterior to slightly larger oblong posterior “area;” single postantennal spines very broad; male with 77 support rods in suction cup; 1419 sclerites per rod in male; mouth tube with at least 19 scales on basal half; basal plate of second maxilla with 3 stout spines, scales covering nearly entire raised pad, one stout, naked seta on posterior margin of pad; accessory spines on thorax large, very broad base on raised pad with many scales, tapering to rounded tips; first 2 pairs of legs lacking flagella; posteroventral surface of coxae of second legs of male expanded into a broadly-rounded lobe that is covered with many small and closely-arranged scales; third legs of male with two pegs issuing from cleft on dorsal side of legs and two fingerlike papillae; small fleshy lobe at posterior of thorax; male abdomen bilobate with testes extending to the level of the anal sinus base; caudal rami long, slen- der, basal in anal sinus.
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.— Total length
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[
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Meehean 1940:
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.85 mm]. Carapace shape is “ovate, sinuses shallow, cephalic areas broad...; alae reaching abdomen [
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].” Carapace length [
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Meehean 1940:
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.88 mm]. Maximum carapace width [
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Meehean 1940:
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.28 mm]. Pair of compound eyes anteriorly with diameters 250 (left) and 220 (right) µm. Transverse distance between eyes
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(830 µm). Nauplius eye with one anterior and two posterior ocelli. Sclerotized dorsal ridges not forked anterior of eyes. Ventrally, carapace with small, posteriorly-projecting spines along outer margin [due to the deteriorated condition of the carapace alae, additional details cannot be included here]. Respiratory areas consist of two oblong-shaped areas, anterior one slightly smaller than posterior [from
<bibRefCitation id="8F7887979776FFBDFE8CFAC2D1E3C44F" author="MEEHEAN, O. L." box="[296,456,1320,1344]" pageId="1" pageNumber="134" pagination="459 - 522" refId="ref3178" refString="MEEHEAN, O. L. 1940. A review of the parasitic Crustacea of the genus Argulus in the collections of the United States National Museum. Proceedings of the United States National Museum 88 (3087): 459 - 522." type="journal article" year="1940">Meehean 1940</bibRefCitation>
] (Fig. 1). No pigmentation present (but probably lost in preservative and/or due to clearing). Thorax compressed dorsoventrally, 4-segmented, with 2 pairs of posteriorly-projecting spines ventrally. Spines digitate with broad base, anterior pair (accessory spines) larger than posterior pair (postmaxillary spines), both pairs angled mesially. Accessory spines between basal segments of second maxillae; very broad base on a raised pad covered with many scales; spines tapering to round- ed tips (
<figureCitation id="73D2E6E39776FFBDFF77F9C2D138C74F" box="[211,275,1576,1600]" captionStart="FIGURE" captionStartId="2.[155,167,605,624]" captionTargetBox="[123,1159,219,1131]" captionTargetId="figure-1945@2.[575,1176,216,1134]" captionTargetPageId="2" captionText="FIGURE 2. Argulus floridensis: Second maxil- la, raised pad with scales (adjacent on thorax), accessory spine, and postmaxillary spine. Scale bar = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13155433" httpUri="https://zenodo.org/record/13155433/files/figure.png" pageId="1" pageNumber="134">Fig. 2</figureCitation>
). Male with ovoid fleshy lobe at posterior of fourth thoracic segment between natatory lobes (Figs. 3A, 4). Thorax with
<superScript id="1C9C572E9776FFBDFC2CF9AAD4BEC75C" attach="right" box="[904,1173,1600,1619]" fontSize="8" pageId="1" pageNumber="134">
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1.
<taxonomicName id="2CE981E59776FFBDFC4CF9AAD4A4C75C" authorityName="Meehean" authorityYear="1940" box="[1000,1167,1600,1619]" class="Maxillopoda" family="Argulidae" genus="Argulus" kingdom="Animalia" order="Arguloida" pageId="1" pageNumber="134" phylum="Arthropoda" rank="species" species="floridensis">
<emphasis id="D99D26749776FFBDFC4CF9AAD4A4C75C" box="[1000,1167,1600,1619]" italics="true" pageId="1" pageNumber="134">Argulus floridensis</emphasis>
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:
</superScript>
</paragraph>
<paragraph id="EB56FA669776FFBEFCC3F9B3D4BDC5A5" blockId="1.[122,1174,424,1696]" lastBlockId="2.[122,1174,723,1708]" lastPageId="2" lastPageNumber="135" pageId="1" pageNumber="134">
shape of anterior and posterior scales scattered on ventral surface; thoracic segment at base of third respiratory areas (redrawn from leg produced laterally into expanded anterolateral lobe with scales
<bibRefCitation id="8F7887979776FFBDFCC3F967D3CDC7AF" author="MEEHEAN, O. L." box="[871,998,1677,1696]" pageId="1" pageNumber="134" pagination="459 - 522" refId="ref3178" refString="MEEHEAN, O. L. 1940. A review of the parasitic Crustacea of the genus Argulus in the collections of the United States National Museum. Proceedings of the United States National Museum 88 (3087): 459 - 522." type="journal article" year="1940">Meehean 1940</bibRefCitation>
). with secondary sexual structures (Figs.
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3.
<taxonomicName id="2CE981E59775FFBEFD18FB94D374C59E" authorityName="Meehean" authorityYear="1940" box="[700,863,1150,1169]" class="Maxillopoda" family="Argulidae" genus="Argulus" kingdom="Animalia" order="Arguloida" pageId="2" pageNumber="135" phylum="Arthropoda" rank="species" species="floridensis">
<emphasis id="D99D26749775FFBEFD18FB94D374C59E" box="[700,863,1150,1169]" italics="true" pageId="2" pageNumber="135">Argulus floridensis</emphasis>
</taxonomicName>
: A) ventral view of legs 2, 3, and 4 3A, B). (precoxa, coxa, and basis only); exopods, endopods, setae, and sensil-
</paragraph>
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2.
<taxonomicName id="2CE981E59775FFBEFF52FDB7D1B1C37F" authorityName="Meehean" authorityYear="1940" box="[246,410,605,624]" class="Maxillopoda" family="Argulidae" genus="Argulus" kingdom="Animalia" order="Arguloida" pageId="2" pageNumber="135" phylum="Arthropoda" rank="species" species="floridensis">
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: Second maxilla, raised pad with scales (adjacent on thorax),
</paragraph>
<paragraph id="EB56FA669775FFBEFFDFFD7BD0DEC3B2" blockId="2.[123,549,605,701]" pageId="2" pageNumber="135">accessory spine, and postmaxillary spine. Scale bar = 200 µm.</paragraph>
</caption>
<paragraph id="EB56FA669775FFBEFF0EFB59D2DBC5D1" blockId="2.[122,1174,723,1708]" pageId="2" pageNumber="135">First legs lack secondary modifica- lae omitted. B) dorsal view of leg 3 (precoxa, coxa, and basis only). tions. Second legs, the posterior ventral Scale bar = 500 µm.</paragraph>
<paragraph id="EB56FA669775FFBEFFDEFB19D0CEC484" blockId="2.[122,1174,723,1708]" pageId="2" pageNumber="135">
margin of coxae expanded into broadly-rounded lobe with many small, closely-arranged scales. Third legs, dorsal anterior coxae with broadly-rounded hump armed with many scales directed proximally; distal to this hump are two smaller, unarmed protuberances; at anterior distal margin of coxae an anteriorly-directed, fleshy projection (“upright fingerlike papilla” of
<bibRefCitation id="8F7887979775FFBEFC48FAB9D4A7C463" author="MEEHEAN, O. L." box="[1004,1164,1363,1388]" pageId="2" pageNumber="135" pagination="459 - 522" refId="ref3178" refString="MEEHEAN, O. L. 1940. A review of the parasitic Crustacea of the genus Argulus in the collections of the United States National Museum. Proceedings of the United States National Museum 88 (3087): 459 - 522." type="journal article" year="1940">Meehean 1940</bibRefCitation>
) (Fig. 3A).
</paragraph>
<paragraph id="EB56FA669775FFBEFF0EFA79D3B9C784" blockId="2.[122,1174,723,1708]" pageId="2" pageNumber="135">
Third legs with two sclerotized pegs in cleft on dorsal side at distal margins of coxae (or fused coxae+bases); pegs with a single, large horn and orifice distally; sclerotized ridge at anterior dorsal of coxa/basis junction; one fleshy papilla extends proximally (mesially) from sclerotized ridge anterior of two pegs and overhanging cleft (“saddle-shaped depression” of
<bibRefCitation id="8F7887979775FFBEFC42FA19D4AEC703" author="MEEHEAN, O. L." box="[998,1157,1523,1548]" pageId="2" pageNumber="135" pagination="459 - 522" refId="ref3178" refString="MEEHEAN, O. L. 1940. A review of the parasitic Crustacea of the genus Argulus in the collections of the United States National Museum. Proceedings of the United States National Museum 88 (3087): 459 - 522." type="journal article" year="1940">Meehean 1940</bibRefCitation>
). Posterior to pegs and cleft a triangular-shaped lobe covered with scales (Fig. 3B). Posterior margins of both precoxae+coxae and bases of fourth legs with natatory lobes bearing plumose setae. Anterior of bases a sclerotized ridge lies on basal half, opposed by recurved, finger-like appendage covered with fine scales distally; tiny spines or scales on its tip (Fig. 3A).
</paragraph>
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Abdomen bilobate. Abdomen length
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[
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Meehean 1940:
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.58 mm]; maximum width
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[
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Meehean 1940:
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.35 mm]. Anal sinus length
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(720 µm) [
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:
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]. Caudal rami paired, long, slender, near base of anal sinus (could not see the setae clearly, usually there are 5 simple setae at the tip of each). Tips of abdominal lobes narrowly rounded, not pointy. Testes extending to level of anal sinus base (
<figureCitation id="73D2E6E39774FFBFFD19FC4BD2D6C2B5" box="[701,765,929,954]" captionStart="FIGURE" captionStartId="3.[155,167,707,726]" captionTargetBox="[183,1171,224,678]" captionTargetId="figure-2254@3.[653,1176,245,651]" captionTargetPageId="3" captionText="FIGURE 4. Argulus floridensis: abdomen; fleshy lobe at FIGURE 5. Argulus floridensis: first and second antennae. posterior of thorax is shown as a dashed outline at anterior of The fleshy segments of the first antennae were obscured or abdomen. Setae not shown on caudal rami. Scale bar = 500 missing on this structure but are included as dashed outlines. µm. Scale bar = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.13155435" httpUri="https://zenodo.org/record/13155435/files/figure.png" pageId="3" pageNumber="136">Fig. 4</figureCitation>
). Scales absent on ventral surface of abdomen; spines or scales absent along margins of abdominal lobes.
</paragraph>
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4.
<taxonomicName id="2CE981E59774FFBFFF5DFD29D1B5C3D9" authorityName="Meehean" authorityYear="1940" box="[249,414,707,726]" class="Maxillopoda" family="Argulidae" genus="Argulus" kingdom="Animalia" order="Arguloida" pageId="3" pageNumber="136" phylum="Arthropoda" rank="species" species="floridensis">
<emphasis id="D99D26749774FFBFFF5DFD29D1B5C3D9" box="[249,414,707,726]" italics="true" pageId="3" pageNumber="136">Argulus floridensis</emphasis>
</taxonomicName>
: abdomen; fleshy lobe at
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5.
<taxonomicName id="2CE981E59774FFBFFCACFD29D381C3D9" authorityName="Meehean" authorityYear="1940" box="[776,938,707,726]" class="Maxillopoda" family="Argulidae" genus="Argulus" kingdom="Animalia" order="Arguloida" pageId="3" pageNumber="136" phylum="Arthropoda" rank="species" species="floridensis">
<emphasis id="D99D26749774FFBFFCACFD29D381C3D9" box="[776,938,707,726]" italics="true" pageId="3" pageNumber="136">Argulus floridensis</emphasis>
</taxonomicName>
: first and second antennae. posterior of thorax is shown as a dashed outline at anterior of The fleshy segments of the first antennae were obscured or abdomen. Setae not shown on caudal rami. Scale bar = 500 missing on this structure but are included as dashed outlines. µm. Scale bar = 200 µm.
</paragraph>
</caption>
<paragraph id="EB56FA669774FFBFFF0EFC0BD363C2F6" blockId="3.[121,1174,833,1722]" box="[170,840,993,1018]" pageId="3" pageNumber="136">Mouth tube long, slender, with at least 19 scales on basal half.</paragraph>
<paragraph id="EB56FA669774FFBFFF0EFBEBD196C436" blockId="3.[121,1174,833,1722]" pageId="3" pageNumber="136">
First antennae 4-segmented. First segment (basal segment) sclerotized, large with stout posteriorly-projecting posterior spine, narrowly rounded at tip; second segment sclerotized with small recurved spine anteriorly with narrow, blunt tip, posteriorly-projecting medial spine with rounded tip, and large recurved terminal spine; third and fourth fleshy segments missing or obscured on the
<typeStatus id="345244C49774FFBFFFDEFB6BD0FCC596" box="[122,215,1153,1177]" pageId="3" pageNumber="136" type="holotype">holotype</typeStatus>
(Fig. 5). Width of first antennae (mean of both antennae, µm) 530540 (535) in male. Second antennae 5-segmented, fleshy. First two segments larger; remaining three thin, cylindrical; basal segment bears posteriorly-projecting posterior spine with narrowly-rounded tip. Postantennal spines single (as opposed to double in some taxa), very wide at base, tapering to narrowly-rounded tip (Fig. 5).
</paragraph>
<paragraph id="EB56FA669774FFBFFF0EFAABD14BC716" blockId="3.[121,1174,833,1722]" pageId="3" pageNumber="136">First maxillae modified into suction cups in adult. In male, first maxillae inner diameter 680 µm and outer diameter 1,050 µm (n = 1). Number of sclerites per support rod in male 1419 (n = 25 support rods) [excludes abnormal rods with missing sclerites]. Basal (proximal) sclerite usually subquadrangular, shape variable; distal sclerites imbricate bowls, plates (Fig. 6). Suction cup with 29 sensilla on inside circumference.</paragraph>
<paragraph id="EB56FA669774FFBFFF0EF9CBD4BEC762" blockId="3.[121,1174,833,1722]" pageId="3" pageNumber="136">
Second maxillae 5-segmented with broad basal plate bearing three stout spines with the mesial spine broader than the other two (
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). Basal plate with elevated pad bearing 43 scales and one stout
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6.
<taxonomicName id="2CE981E59774FFBFFC4AF9B0D4A4C762" authorityName="Meehean" authorityYear="1940" box="[1006,1167,1626,1645]" class="Maxillopoda" family="Argulidae" genus="Argulus" kingdom="Animalia" order="Arguloida" pageId="3" pageNumber="136" phylum="Arthropoda" rank="species" species="floridensis">
<emphasis id="D99D26749774FFBFFC4AF9B0D4A4C762" box="[1006,1167,1626,1645]" italics="true" pageId="3" pageNumber="136">Argulus floridensis</emphasis>
</taxonomicName>
:
</superScript>
</paragraph>
<paragraph id="EB56FA669774FFBFFCD3F99ED3FBC7B5" blockId="3.[121,1174,833,1722]" pageId="3" pageNumber="136">Two support rods from suction seta posteriorly. Bi-dentate to multidentate scales on ventral surfaces cup (right first maxilla), Scale bar of segments 2 and 3; segment 4 lacking armature. Distal segment with = 100 µm,</paragraph>
<paragraph id="EB56FA669773FFB8FFDFFF31D1F9C01C" blockId="4.[123,1174,219,374]" pageId="4" pageNumber="137">2 sharp claws (1 broken/missing) and one blunt, elongate lobe positioned above claws (did not see a small sensillum at tip of lobe).</paragraph>
</subSubSection>
<subSubSection id="A3F3A9ED9773FFB8FF0FFEF6D1A8C03B" box="[171,387,284,308]" pageId="4" pageNumber="137" type="biology_ecology">
<paragraph id="EB56FA669773FFB8FF0FFEF6D1A8C03B" blockId="4.[123,1174,219,374]" box="[171,387,284,308]" pageId="4" pageNumber="137">
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<emphasis id="D99D26749773FFB8FF0FFEF6D0C2C03C" bold="true" box="[171,233,284,308]" pageId="4" pageNumber="137">HOST</emphasis>
</smallCapsWord>
.— Unknown.
</paragraph>
</subSubSection>
<subSubSection id="A3F3A9ED9773FFB8FF0FFED7D41AC079" pageId="4" pageNumber="137" type="etymology">
<paragraph id="EB56FA669773FFB8FF0FFED7D41AC079" blockId="4.[123,1174,219,374]" pageId="4" pageNumber="137">
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<emphasis id="D99D26749773FFB8FF0FFED7D11EC05C" bold="true" box="[171,309,317,341]" pageId="4" pageNumber="137">ETYMOLOGY</emphasis>
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.— The specific name,
<taxonomicName id="2CE981E59773FFB8FD89FED7D2B5C05A" authorityName="Meehean" authorityYear="1940" box="[557,670,317,341]" class="Maxillopoda" family="Argulidae" genus="Argulus" kingdom="Animalia" order="Arguloida" pageId="4" pageNumber="137" phylum="Arthropoda" rank="species" species="floridensis">
<emphasis id="D99D26749773FFB8FD89FED7D2B5C05A" box="[557,670,317,341]" italics="true" pageId="4" pageNumber="137">floridensis</emphasis>
</taxonomicName>
, is derived from the state,
<collectingRegion id="292D34849773FFB8FC1CFED7D42EC05A" box="[952,1029,317,341]" country="United States of America" name="Florida" pageId="4" pageNumber="137">Florida</collectingRegion>
,
<materialsCitation id="5B81F03B9773FFB8FBB6FED7D41AC079" collectionCode="Host" location="Florida" pageId="4" pageNumber="137" specimenCount="1" stateProvince="Florida">
in which the
<typeStatus id="345244C49773FFB8FFDFFEB4D082C079" box="[123,169,350,374]" pageId="4" pageNumber="137">type</typeStatus>
locality is located and has an adjectival suffix, ensis, meaning “of or from
<collectingRegion id="292D34849773FFB8FC75FEB7D435C07A" box="[977,1054,349,373]" country="United States of America" name="Florida" pageId="4" pageNumber="137">Florida</collectingRegion>
.”
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>