treatments-xml/data/C7/40/D6/C740D6092A25BB0DFEC3E377E5BE298A.xml
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<document id="4B4C83F07D47DCDE735A4E32A8EFF8D8" ID-CLB-Dataset="24063" ID-DOI="10.1080/00222933.2019.1692088" ID-GBIF-Dataset="e6443eb6-bfcc-44a6-a547-7af9a94d4a7d" ID-ISSN="1464-5262" ID-Zenodo-Dep="3665519" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1581529324984" checkinUser="valdenar" docAuthor="Nunes, Rafael V. &amp; Vaz-de-Mello, Fernando Z." docDate="2019" docId="C740D6092A25BB0DFEC3E377E5BE298A" docLanguage="en" docName="J_NAT_HIST.53.3738.22312351.pdf.imf" docOrigin="Journal of Natural History 53 (37)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Dichotomius (Cephagonus) seag Nunes &amp; Vaz-de-Mello 2019, new species" docType="treatment" docVersion="18" lastPageNumber="2302" masterDocId="3B79AE712A62BB45FFA2E57EE46C2C23" masterDocTitle="Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)" masterLastPageNumber="2351" masterPageNumber="2231" pageNumber="2301" updateTime="1698782751744" updateUser="ExternalLinkService">
<mods:mods id="4ED590CFEC4E36746E823EC4E2382F79" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="7BAC00CA03CEA03F5D60423DF6365BED">Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)</mods:title>
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<mods:name id="323097B3AB1AA2ECD4D3F349A06D69AC" type="personal">
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<mods:namePart id="E204DC30476D4DE0A8678056F3DE28C5">Nunes, Rafael V.</mods:namePart>
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<mods:namePart id="1140CC817DE241309A432F704916C46C">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:title id="1FD987271029A9A2FB2D7AB82890029D">Journal of Natural History</mods:title>
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<mods:date id="2D5B60E29502C365E4B1F8D18F7D03FF">2019</mods:date>
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<mods:number id="2448E9E7D87466FB7C2E70030A27CB27">2020-01-13</mods:number>
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<mods:number id="206E3B3DBE04D2CDDEE0B16517F9775D">53</mods:number>
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<mods:number id="1BDC21B8558E296FF1A281C46AEADA31">37</mods:number>
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<treatment id="C740D6092A25BB0DFEC3E377E5BE298A" ID-DOI="http://doi.org/10.5281/zenodo.3671892" ID-GBIF-Taxon="162116289" ID-Zenodo-Dep="3671892" LSID="urn:lsid:plazi:treatment:C740D6092A25BB0DFEC3E377E5BE298A" httpUri="http://treatment.plazi.org/id/C740D6092A25BB0DFEC3E377E5BE298A" lastPageId="72" lastPageNumber="2302" pageId="71" pageNumber="2301">
<subSubSection id="07F334942A25BB02FEC3E377E6B82A65" pageId="71" pageNumber="2301" type="nomenclature">
<paragraph id="4F56671F2A25BB02FEC3E377E7AE2A00" blockId="71.[353,962,1545,1606]" box="[353,962,1545,1571]" pageId="71" pageNumber="2301">
<heading id="141ED0732A25BB02FEC3E377E7AE2A00" box="[353,962,1545,1571]" centered="true" fontSize="11" level="2" pageId="71" pageNumber="2301" reason="2">
2.6.:
<taxonomicName id="88E91C9C2A25BB02FE3BE377E7702A00" ID-CoL="6CRRD" authority="Nunes &amp; Vaz-de-Mello, 2019" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[409,796,1545,1571]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="2301" phylum="Arthropoda" rank="species" species="seag" status="sp. nov." subGenus="Cephagonus">
<emphasis id="7D9DBB0D2A25BB02FE3BE377E7702A00" box="[409,796,1545,1571]" italics="true" pageId="71" pageNumber="2301">Dichotomius (Cephagonus) seag</emphasis>
</taxonomicName>
<emphasis id="7D9DBB0D2A25BB02FC86E377E7AE2A00" bold="true" box="[804,962,1545,1571]" pageId="71" pageNumber="2301">
<taxonomicNameLabel id="66AE06762A25BB02FC86E377E7AE2A00" box="[804,962,1545,1571]" pageId="71" pageNumber="2301" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4F56671F2A25BB02FDEDE352E6B82A65" blockId="71.[353,962,1545,1606]" box="[591,724,1580,1606]" pageId="71" pageNumber="2301">
<heading id="141ED0732A25BB02FDEDE352E6B82A65" box="[591,724,1580,1606]" centered="true" fontSize="11" level="2" pageId="71" pageNumber="2301" reason="5">
(
<figureCitation id="D7D27B9A2A25BB02FDF5E352E6A02A65" box="[599,716,1580,1606]" captionStart="Figure 25" captionStartId="40.[160,225,1337,1360]" captionTargetBox="[163,1153,142,1305]" captionTargetId="figure@40.[163,1153,142,1305]" captionTargetPageId="40" captionText="Figure 25. D. seag: (a) male dorsal habitus (Holotype); (b) male ventral habitus; (c) paramera dorsal view; (d) paramera ventral view; (f) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3665565" httpUri="https://zenodo.org/record/3665565/files/figure.png" pageId="71" pageNumber="2301">Figure 25</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="07F334942A25BB0DFF02E3FAE7382D2D" lastPageId="72" lastPageNumber="2302" pageId="71" pageNumber="2301" type="diagnosis">
<paragraph id="4F56671F2A25BB0DFF02E3FAE7382D2D" blockId="71.[160,1155,1668,1727]" lastBlockId="72.[160,1156,142,270]" lastPageId="72" lastPageNumber="2302" pageId="71" pageNumber="2301">
<emphasis id="7D9DBB0D2A25BB02FF02E3FAE5762ABE" bold="true" box="[160,282,1668,1693]" italics="true" pageId="71" pageNumber="2301">Diagnosis.</emphasis>
distinguished from the other species in the group by the following combination of characteristics: males
<emphasis id="7D9DBB0D2A25BB02FE78E3D9E58C2A9C" box="[474,480,1703,1727]" italics="true" pageId="71" pageNumber="2301"></emphasis>
cephalic carina longer than wideı with a rounded apex (as in medium males of
<taxonomicName id="88E91C9C2A2ABB0DFECBE5F0E5B72C85" box="[361,475,142,166]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="2302" phylum="Arthropoda" rank="species" species="bicuspis">
<emphasis id="7D9DBB0D2A2ABB0DFECBE5F0E5B72C85" box="[361,475,142,166]" italics="true" pageId="72" pageNumber="2302">D. bicuspis</emphasis>
</taxonomicName>
of
<taxonomicName id="88E91C9C2A2ABB0DFE5DE5F0E6382C85" box="[511,596,142,166]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="2302" phylum="Arthropoda" rank="species" species="bicuspis">
<emphasis id="7D9DBB0D2A2ABB0DFE5DE5F0E6382C85" box="[511,596,142,166]" italics="true" pageId="72" pageNumber="2302">bicuspis</emphasis>
</taxonomicName>
group); anterior and posterior pronotum portions with very fine punctuation (viewed under 30x magnification); males
<emphasis id="7D9DBB0D2A2ABB0DFC04E5CFE7C02CEA" box="[934,940,177,201]" italics="true" pageId="72" pageNumber="2302"></emphasis>
pronotal disc with a single-rounded lobe at the anterior portion (as in
<taxonomicName id="88E91C9C2A2ABB0DFD40E5AAE7552CCF" authorityName="Haroldi" authorityYear="1867" box="[738,825,211,236]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="2302" phylum="Arthropoda" rank="species" species="fissus">
<emphasis id="7D9DBB0D2A2ABB0DFD40E5AAE7552CCF" box="[738,825,211,236]" italics="true" pageId="72" pageNumber="2302">D. fissus</emphasis>
</taxonomicName>
of the same group); pronotal margin smooth and; elytral striae with indistinct punctuation.
</paragraph>
</subSubSection>
<subSubSection id="07F334942A2ABB0DFF1CE442E7622D9F" pageId="72" pageNumber="2302" type="materials_examined">
<paragraph id="4F56671F2A2ABB0DFF1CE442E6C72D55" blockId="72.[190,1157,316,444]" pageId="72" pageNumber="2302">
<emphasis id="7D9DBB0D2A2ABB0DFF1CE442E4A02D73" bold="true" box="[190,204,316,336]" pageId="72" pageNumber="2302"></emphasis>
<materialsCitation id="FF816D422A2ABB0DFF7AE442E6CF2D55" ID-GBIF-Occurrence="2570409556" collectingDate="1997-03" collectionCode="CEMT" location="Nouragues Biological Reserve. Fezes Allouatta" municipality="Cayenne" pageId="72" pageNumber="2302" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="9052D9BD2A2ABB0DFF7AE442E53B2D77" box="[216,343,316,340]" pageId="72" pageNumber="2302" type="holotype">HOLOTYPEı</typeStatus>
<specimenCount id="59EFAC962A2ABB0DFEC0E442E5F72D77" box="[354,411,316,340]" pageId="72" pageNumber="2302" type="male">male</specimenCount>
: GUYANNE FRANÇAISE:
<emphasis id="7D9DBB0D2A2ABB0DFD11E442E7762D77" bold="true" box="[691,794,316,340]" pageId="72" pageNumber="2302">
<collectingMunicipality id="AF32FD652A2ABB0DFD11E442E7762D77" box="[691,794,316,340]" pageId="72" pageNumber="2302">Cayenne</collectingMunicipality>
</emphasis>
:
<location id="4A3631C42A2ABB0DFC89E442E5ED2D55" LSID="urn:lsid:plazi:treatment:C740D6092A25BB0DFEC3E377E5BE298A:4A3631C42A2ABB0DFC89E442E5ED2D55" municipality="Cayenne" name="Nouragues Biological Reserve. Fezes Allouatta" pageId="72" pageNumber="2302">
Nouragues Biological Reserve. Fezes
<emphasis id="7D9DBB0D2A2ABB0DFEBFE420E5ED2D55" box="[285,385,350,374]" italics="true" pageId="72" pageNumber="2302">Allouatta</emphasis>
</location>
.
<date id="3B5741DF2A2ABB0DFE2CE420E58E2D55" box="[398,482,350,374]" pageId="72" pageNumber="2302" value="1997-03">
<collectingDate id="2B13B8372A2ABB0DFE2CE420E58E2D55" box="[398,482,350,374]" pageId="72" pageNumber="2302" value="1997-03">iii-1997</collectingDate>
</date>
. F
<location id="4A3631C42A2ABB0DFDA3E420E65D2D55" LSID="urn:lsid:plazi:treatment:C740D6092A25BB0DFEC3E377E5BE298A:4A3631C42A2ABB0DFDA3E420E65D2D55" box="[513,561,350,374]" municipality="Cayenne" name="Feer" pageId="72" pageNumber="2302">Feer</location>
[at
<collectionCode id="29F8FFDA2A2ABB0DFDFCE420E6CF2D55" box="[606,675,350,374]" pageId="72" pageNumber="2302">CEMT</collectionCode>
</materialsCitation>
].
</paragraph>
<paragraph id="4F56671F2A2ABB0DFF1CE4FFE7622D9F" blockId="72.[190,1157,316,444]" pageId="72" pageNumber="2302">
<emphasis id="7D9DBB0D2A2ABB0DFF1CE4FFE4A02DB6" bold="true" box="[190,204,385,405]" pageId="72" pageNumber="2302"></emphasis>
<typeStatus id="9052D9BD2A2ABB0DFF7AE4FFE5302DBA" box="[216,348,385,409]" pageId="72" pageNumber="2302" type="paratype">PARATYPES</typeStatus>
: same data of
<typeStatus id="9052D9BD2A2ABB0DFDB0E4FFE6152DBA" box="[530,633,385,409]" pageId="72" pageNumber="2302" type="holotype">Holotype</typeStatus>
[1Ƌ];
<collectingCountry id="37FE278F2A2ABB0DFD6AE4FFE72F2DBA" box="[712,835,385,409]" name="Suriname" pageId="72" pageNumber="2302">SURINAME</collectingCountry>
:
<emphasis id="7D9DBB0D2A2ABB0DFCF3E4FFE7AA2DBA" bold="true" box="[849,966,385,409]" pageId="72" pageNumber="2302">Sipawilini</emphasis>
. Palumeu River. 2³28
<emphasis id="7D9DBB0D2A2ABB0DFEA8E4DDE5632D98" box="[266,271,419,443]" italics="true" pageId="72" pageNumber="2302"></emphasis>
Sı 55
<geoCoordinate id="2ADD01D82A2ABB0DFEE4E4DAE5E52D9F" box="[326,393,419,444]" degrees="3" direction="west" minutes="37" orientation="longitude" pageId="72" pageNumber="2302" precision="925" value="-3.6166668">
³37
<emphasis id="7D9DBB0D2A2ABB0DFEC8E4DDE51C2D98" bold="true" box="[362,368,419,443]" italics="true" pageId="72" pageNumber="2302">ʹ</emphasis>
W
</geoCoordinate>
.
<date id="3B5741DF2A2ABB0DFE36E4DAE5842D9F" box="[404,488,420,444]" pageId="72" pageNumber="2302" value="2012-03">iii-2012</date>
.
<collectionCode id="29F8FFDA2A2ABB0DFE50E4DAE5932D9F" box="[498,511,420,444]" name="Tavera, Department of Geology and Geophysics" pageId="72" pageNumber="2302">T</collectionCode>
Larsen [2 ƋƋ at
<collectionCode id="29F8FFDA2A2ABB0DFD63E4DAE76A2D9F" box="[705,774,420,444]" pageId="72" pageNumber="2302">CEMT</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="07F334942A2ABB0DFF02E496E62E28DF" pageId="72" pageNumber="2302" type="description">
<paragraph id="4F56671F2A2ABB0DFF02E496E013282A" blockId="72.[160,1156,488,1033]" pageId="72" pageNumber="2302">
<emphasis id="7D9DBB0D2A2ABB0DFF02E496E5412E22" bold="true" box="[160,301,488,513]" italics="true" pageId="72" pageNumber="2302">Description.</emphasis>
(maleı
<typeStatus id="9052D9BD2A2ABB0DFE2AE497E59C2E22" box="[392,496,489,513]" pageId="72" pageNumber="2302" type="holotype">holotype</typeStatus>
): BL: 23 mmı PW:
<quantity id="8811CAFA2A2ABB0DFD1BE497E7672E22" box="[697,779,489,513]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="72" pageNumber="2302" unit="mm" value="13.0">13 mm</quantity>
. Colour: blackı with brown elytra.
<emphasis id="7D9DBB0D2A2ABB0DFF02E772E4B22E07" bold="true" box="[160,222,524,548]" pageId="72" pageNumber="2302">Head</emphasis>
: fronto-clypealı genal and inter-ocular surfaces smooth. Gena borders slightly curved inwards. Cephalic carina longer than wideı rounded at the apex.
<emphasis id="7D9DBB0D2A2ABB0DFBA7E750E0112E65" bold="true" box="[1029,1149,558,582]" pageId="72" pageNumber="2302">Pronotum</emphasis>
: borders smoothı not serrated. Pronotal disc shinyı lacking punctuation and delimited anteriorly by a singleı rounded lobe. Anterior portionı including excavation and anterior angles with very fineı equally spaced punctures (viewed under 20x magnification). Posterior margin with two rows of small ocellate punctures.
<emphasis id="7D9DBB0D2A2ABB0DFCC7E7C7E0692EF2" bold="true" box="[869,1029,697,721]" pageId="72" pageNumber="2302">Metasternum</emphasis>
: sides and anterior portion of anterior lobe covered by dense red setae. Setigerous punctuation equally spacedı where present.
<emphasis id="7D9DBB0D2A2ABB0DFDA7E780E6262F35" bold="true" box="[517,586,766,790]" pageId="72" pageNumber="2302">Elytra</emphasis>
: striae weakly impressed lacking distinct punctuation. Striae expanding near the base forming foveae. Interstriae brown colourı lacking microsculpture.
<emphasis id="7D9DBB0D2A2ABB0DFEC7E63AE5B62F7F" bold="true" box="[357,474,836,860]" pageId="72" pageNumber="2302">Abdomen</emphasis>
: glabrousı punctuation of ventrites anterior margininterrupted medially.
<emphasis id="7D9DBB0D2A2ABB0DFE3CE618E6782F5D" bold="true" box="[414,532,870,894]" pageId="72" pageNumber="2302">Aedeagus</emphasis>
(
<figureCitation id="D7D27B9A2A2ABB0DFD8AE618E6F92F5D" box="[552,661,870,894]" captionStart="Figure 21" captionStartId="36.[160,225,1600,1623]" captionTargetBox="[163,1152,142,1569]" captionTargetId="figure@36.[163,1153,142,1569]" captionTargetPageId="36" captionText="Figure 21. D. fonsecae: (a) male dorsal habitus; (b) Holotype dorsal habitus; (c) holotype labels; (d) female dorsal view; (e) female ventral view; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3665557" httpUri="https://zenodo.org/record/3665557/files/figure.png" pageId="72" pageNumber="2302">Figure 21</figureCitation>
(c-e)): parameraı apexes rounded dorsallyı convergent; longitudinal excavation taking 3/4 of paramera length; basal excavations rounded. Laterallyı subtle declivity towards apex; posterior margin with an emargination near junction with phalobasis. Ventrallyı sclerotised portion excavatedı forming a groove constrained at mid-portion; subgenital plate rounded at basis and emarginated at apex.
</paragraph>
<paragraph id="4F56671F2A2ABB0DFF02E148E62E28DF" blockId="72.[160,1156,1078,1276]" pageId="72" pageNumber="2302">
<emphasis id="7D9DBB0D2A2ABB0DFF02E148E5AB286C" bold="true" box="[160,455,1078,1103]" italics="true" pageId="72" pageNumber="2302">Morphological variation.</emphasis>
male BL: 17
<emphasis id="7D9DBB0D2A2ABB0DFDF8E148E60B286D" box="[602,615,1078,1102]" italics="true" pageId="72" pageNumber="2302"></emphasis>
23 mmı PW:
<quantity id="8811CAFA2A2ABB0DFD5AE149E709286C" box="[760,869,1078,1103]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="72" pageNumber="2302" unit="mm" value="11.0" valueMax="13.0" valueMin="9.0">
9
<emphasis id="7D9DBB0D2A2ABB0DFCA7E148E77E286D" box="[773,786,1078,1102]" italics="true" pageId="72" pageNumber="2302"></emphasis>
13 mm
</quantity>
. Female BL: 17 mmı`PW:
<quantity id="8811CAFA2A2ABB0DFF02E127E48C2852" box="[160,224,1113,1137]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="72" pageNumber="2302" unit="mm" value="9.0">9 mm</quantity>
and is distinguished from males as follows: Head: fronto-clypeal surface with coarse puncture. Cephalic carina transversalı forming a weak pair of tubercles. Pronotum: simply convexı lacking lobes. Elytra: striae deeply impressedı punctures separated by twice their diameter. Abdomen: sixth ventrite bearing a pair of triangular projections reaching pygidium apical margin (
<figureCitation id="D7D27B9A2A2ABB0DFE18E19AE67428DF" box="[442,536,1252,1276]" captionStart="Figure 9" captionStartId="22.[160,225,1548,1571]" captionTargetBox="[163,1153,142,1516]" captionTargetId="figure@22.[163,1153,142,1516]" captionTargetPageId="22" captionText="Figure 9. (a) Abdominal ventrites of a male of D. spadiceus. (b) ventrites of a female of D. spadiceus. (c) abdominal ventrites of a female of D. itatiaiaensisı (d) detail of C; (e) abdominal ventrites of a female of D. fissus. (f) Adbominal ventrites of a female of D. reichei. (g) Abdominal ventrites of a female of D. lucianomourai." figureDoi="http://doi.org/10.5281/zenodo.3665537" httpUri="https://zenodo.org/record/3665537/files/figure.png" pageId="72" pageNumber="2302">Figure 9</figureCitation>
(b)).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A2ABB0DFF02E056E6852962" box="[160,745,1320,1345]" pageId="72" pageNumber="2302" type="distribution">
<paragraph id="4F56671F2A2ABB0DFF02E056E6852962" blockId="72.[160,745,1320,1345]" box="[160,745,1320,1345]" pageId="72" pageNumber="2302">
<emphasis id="7D9DBB0D2A2ABB0DFF02E056E55F2962" bold="true" box="[160,307,1320,1345]" italics="true" pageId="72" pageNumber="2302">Distribution.</emphasis>
French Guyanaı
<collectingCountry id="37FE278F2A2ABB0DFE5BE057E60F2962" box="[505,611,1321,1345]" name="Suriname" pageId="72" pageNumber="2302">Suriname</collectingCountry>
(
<figureCitation id="D7D27B9A2A2ABB0DFDD0E057E6B72962" box="[626,731,1321,1345]" captionStart="Figure 20" captionStartId="35.[160,225,1019,1042]" captionTargetBox="[163,1152,142,987]" captionTargetId="figure@35.[163,1153,142,988]" captionTargetPageId="35" captionText="Figure 20. Distribution map of the species in the D. fissus group." figureDoi="http://doi.org/10.5281/zenodo.3665555" httpUri="https://zenodo.org/record/3665555/files/figure.png" pageId="72" pageNumber="2302">Figure 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A2ABB0DFF02E010E5BE298A" pageId="72" pageNumber="2302" type="nomenclature">
<paragraph id="4F56671F2A2ABB0DFF02E010E5BE298A" blockId="72.[160,1155,1390,1449]" pageId="72" pageNumber="2302">
<emphasis id="7D9DBB0D2A2ABB0DFF02E010E54F29A4" bold="true" box="[160,291,1390,1415]" italics="true" pageId="72" pageNumber="2302">Etymology.</emphasis>
named after the Société Entomologique Antilles
<collectingCountry id="37FE278F2A2ABB0DFCF2E011E7C929A4" box="[848,933,1391,1415]" name="French Guiana" pageId="72" pageNumber="2302">Guyane</collectingCountry>
(SEAG)ı responsible for the
<typeStatus id="9052D9BD2A2ABB0DFF51E0EFE53B298A" box="[243,343,1425,1449]" pageId="72" pageNumber="2302" type="holotype">holotype</typeStatus>
collection.
</paragraph>
</subSubSection>
</treatment>
</document>