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<mods:title id="53CC2F8694A853CD1627B0D0037D8BB1">The Phalangopsidae crickets (Orthoptera, Grylloidea) of the Seychelles Archipelago: Taxonomy of an ecological radiation</mods:title>
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<taxonomicName id="A5E4A9E1B945FF8A12D2591F3789FC82" ID-CoL="8NWTD" authority="Karny, 1909" authorityName="Karny" authorityYear="1909" box="[151,480,824,851]" class="Insecta" family="Phalangopsidae" genus="Paragryllodes" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="genus">
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<emphasis id="50900E70B945FF8A12D2591F3756FC83" bold="true" box="[151,319,824,851]" italics="true" pageId="5" pageNumber="206">Paragryllodes</emphasis>
<bibRefCitation id="0675AF93B945FF8A1303591E3789FC82" author="Karny, H. H." box="[326,480,824,851]" pageId="5" pageNumber="206" pagination="477 - 480" refId="ref26601" refString="Karny, H. H. (1909) Ostafrikanische Orthopteren. Sammelausbeute von A. Borgert, 1904 - 1905. 9. Mitteilung. Zoologische Jahrbucher, Abteilung fur Systematik, Oekologie und Geographie der Tiere, 27, 477 - 480." type="journal article" year="1909">Karny, 1909</bibRefCitation>
</emphasis>
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<paragraph id="625BD262B945FF8A12D2597B3749FCA7" blockId="5.[151,480,824,887]" box="[151,288,860,887]" pageId="5" pageNumber="206">
(
<figureCitation id="FADFCEE7B945FF8A12DA597B3695FCA7" box="[159,252,860,887]" captionStart="FIGURE 1" captionStartId="6.[151,250,1896,1920]" captionTargetBox="[151,1436,179,1870]" captionTargetId="figure-21@6.[151,1436,179,1870]" captionTargetPageId="6" captionText="FIGURE 1. Phalangopsidae crickets from the Seychelles in the wild. A, Paragryllodes nitidula (Bolivar, 1912) n. comb., female. B, Phaloria (Papuloria) bolivari Hugel &amp; Desutter-Grandcolas, n. sp., male. C, same, female. D, Phaeogryllus fuscus Bolivar, 1912, male. E, same, female. F, Phalangacris ferlegro Hugel &amp; Desutter-Grandcolas, n. sp. male, G, same, female." figureDoi="http://doi.org/10.5281/zenodo.5540769" httpUri="https://zenodo.org/record/5540769/files/figure.png" pageId="5" pageNumber="206">Figs 1A</figureCitation>
,
<figureCitation id="FADFCEE7B945FF8A134D597A377EFCA7" box="[264,279,861,887]" captionStart="FIGURE 2" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[169,1430,200,1820]" captionTargetId="figure-21@8.[157,1430,181,1838]" captionTargetPageId="8" captionText="FIGURE 2. Paragryllodes Karny, 1909: AD, Paragryllodes nitidula (Bolivar, 1912) n. comb.A, head and pronotum in dorsal view; B, head in face view; some of the light patterns on the black shiny cuticula are reflections and not real patterns; C, hindfemur outer side; D, copulatory papilla in ventral view. E, Paragryllodes milloti (Chopard, 1958), copulatory papilla in ventral view. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5540773" httpUri="https://zenodo.org/record/5540773/files/figure.png" pageId="5" pageNumber="206">2</figureCitation>
)
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<paragraph id="625BD262B945FF8A12D25986378FFC69" blockId="5.[151,555,929,984]" box="[151,486,929,953]" pageId="5" pageNumber="206">
<treatmentCitationGroup id="42F4F54CB945FF8A12D25986378FFC69" box="[151,486,929,953]" pageId="5" pageNumber="206">
<treatmentCitation id="E345F473B945FF8A12D25986378BFC69" author="Karny, H. H." box="[151,482,929,953]" page="478" pageId="5" pageNumber="206" year="1909">
<taxonomicName id="A5E4A9E1B945FF8A12D25986378BFC69" ID-CoL="8NWTD" authority="Karny, 1909: 478" authorityName="Karny" authorityPageNumber="478" authorityYear="1909" box="[151,482,929,953]" class="Insecta" family="Phalangopsidae" genus="Paragryllodes" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="genus">
<emphasis id="50900E70B945FF8A12D259863741FC69" box="[151,296,929,953]" italics="true" pageId="5" pageNumber="206">Paragryllodes</emphasis>
<bibRefCitation id="0675AF93B945FF8A136B5986378BFC69" author="Karny, H. H." box="[302,482,929,953]" pageId="5" pageNumber="206" pagination="477 - 480" refId="ref26601" refString="Karny, H. H. (1909) Ostafrikanische Orthopteren. Sammelausbeute von A. Borgert, 1904 - 1905. 9. Mitteilung. Zoologische Jahrbucher, Abteilung fur Systematik, Oekologie und Geographie der Tiere, 27, 477 - 480." type="journal article" year="1909">Karny, 1909: 478</bibRefCitation>
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.
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</paragraph>
<paragraph id="625BD262B945FF8A12D259E73442FC08" blockId="5.[151,555,929,984]" box="[151,555,959,984]" pageId="5" pageNumber="206">
<treatmentCitationGroup id="42F4F54CB945FF8A12D259E73442FC08" box="[151,555,959,984]" pageId="5" pageNumber="206">
<taxonomicName id="A5E4A9E1B945FF8A12D259E737B3FC07" ID-CoL="8NZ4Z" authority="Bolivar, 1912: 286" authorityName="Bolivar" authorityPageNumber="286" authorityYear="1912" box="[151,474,959,984]" class="Insecta" family="Phalangopsidae" genus="Seychellesia" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="genus">
<emphasis id="50900E70B945FF8A12D259E7377DFC08" box="[151,276,960,984]" italics="true" pageId="5" pageNumber="206">Seychellesia</emphasis>
<treatmentCitation id="E345F473B945FF8A135E59E737B3FC07" author="Bolivar, I." box="[283,474,959,984]" page="286" pageId="5" pageNumber="206" year="1912">
<bibRefCitation id="0675AF93B945FF8A135E59E737B3FC07" author="Bolivar, I." box="[283,474,959,984]" pageId="5" pageNumber="206" pagination="263 - 292" refId="ref24320" refString="Bolivar, I. (1912) Orthoptera Acrydiidae, Phasgonuridae, Gryllidae of the Seychelles. Transactions of the Linnean Society of London, Series 2, Zoologie, 15, 263 - 292. https: // doi. org / 10.1111 / j. 1096 - 3642.1912. tb 00101. x" type="journal article" year="1912">
<collectingRegion id="A0201C80B945FF8A135E59E73700FC08" box="[283,361,960,984]" country="Colombia" name="Bolivar" pageId="5" pageNumber="206">Bolivar</collectingRegion>
, 1912: 286
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,
<emphasis id="50900E70B945FF8A13A059E73442FC08" bold="true" box="[485,555,960,984]" pageId="5" pageNumber="206">n. syn.</emphasis>
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<emphasis id="50900E70B945FF8A12D259DA374AFBC5" bold="true" box="[151,291,1021,1045]" pageId="5" pageNumber="206">
<typeStatus id="BD5F6CC0B945FF8A12D259DA36A5FBC5" box="[151,204,1021,1045]" pageId="5" pageNumber="206">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="A5E4A9E1B945FF8A136F59DA34FEFBC5" ID-CoL="4CWDK" authority="Karny, 1909" authorityName="Karny" authorityYear="1909" box="[298,663,1021,1045]" class="Insecta" family="Phalangopsidae" genus="Paragryllodes" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="borgerti">
<emphasis id="50900E70B945FF8A136F59DA347AFBC5" box="[298,531,1021,1045]" italics="true" pageId="5" pageNumber="206">Paragryllodes borgerti</emphasis>
<bibRefCitation id="0675AF93B945FF8A105C59DA34FEFBC5" author="Karny, H. H." box="[537,663,1021,1045]" pageId="5" pageNumber="206" pagination="477 - 480" refId="ref26601" refString="Karny, H. H. (1909) Ostafrikanische Orthopteren. Sammelausbeute von A. Borgert, 1904 - 1905. 9. Mitteilung. Zoologische Jahrbucher, Abteilung fur Systematik, Oekologie und Geographie der Tiere, 27, 477 - 480." type="journal article" year="1909">Karny, 1909</bibRefCitation>
</taxonomicName>
by original monotypy.
</paragraph>
<paragraph id="625BD262B945FF8A12D25E6435A8F849" blockId="5.[151,1437,1091,1946]" pageId="5" pageNumber="206">
<emphasis id="50900E70B945FF8A12D25E6437EEFB8D" bold="true" box="[151,391,1091,1117]" pageId="5" pageNumber="206">Emended diagnosis.</emphasis>
Size variable. Cuticula shiny. Coloration black brown mottled with yellow (
<figureCitation id="FADFCEE7B945FF8A16B75E643324FB8D" box="[1266,1357,1091,1117]" captionStart="FIGURE 1" captionStartId="6.[151,250,1896,1920]" captionTargetBox="[151,1436,179,1870]" captionTargetId="figure-21@6.[151,1436,179,1870]" captionTargetPageId="6" captionText="FIGURE 1. Phalangopsidae crickets from the Seychelles in the wild. A, Paragryllodes nitidula (Bolivar, 1912) n. comb., female. B, Phaloria (Papuloria) bolivari Hugel &amp; Desutter-Grandcolas, n. sp., male. C, same, female. D, Phaeogryllus fuscus Bolivar, 1912, male. E, same, female. F, Phalangacris ferlegro Hugel &amp; Desutter-Grandcolas, n. sp. male, G, same, female." figureDoi="http://doi.org/10.5281/zenodo.5540769" httpUri="https://zenodo.org/record/5540769/files/figure.png" pageId="5" pageNumber="206">Fig. 1A</figureCitation>
); main yellow line on the face transverse or vertical. Fastigium (
<figureCitation id="FADFCEE7B945FF8A116D5E4035EBFB51" box="[808,898,1127,1153]" captionStart="FIGURE 2" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[169,1430,200,1820]" captionTargetId="figure-21@8.[157,1430,181,1838]" captionTargetPageId="8" captionText="FIGURE 2. Paragryllodes Karny, 1909: AD, Paragryllodes nitidula (Bolivar, 1912) n. comb.A, head and pronotum in dorsal view; B, head in face view; some of the light patterns on the black shiny cuticula are reflections and not real patterns; C, hindfemur outer side; D, copulatory papilla in ventral view. E, Paragryllodes milloti (Chopard, 1958), copulatory papilla in ventral view. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5540773" httpUri="https://zenodo.org/record/5540773/files/figure.png" pageId="5" pageNumber="206">Fig. 2A</figureCitation>
) very narrow, grooved, sharply separated from the vertex, lower than head dorsum and separated from it by a transverse furrow; lateral ocelli close to each other, compared to their distance to the median ocellus; median ocellus in frontal position, apical on fastigium. Eyes strongly protruding (
<figureCitation id="FADFCEE7B945FF8A13C75EF43797FB3E" box="[386,510,1235,1262]" captionStart="FIGURE 2" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[169,1430,200,1820]" captionTargetId="figure-21@8.[157,1430,181,1838]" captionTargetPageId="8" captionText="FIGURE 2. Paragryllodes Karny, 1909: AD, Paragryllodes nitidula (Bolivar, 1912) n. comb.A, head and pronotum in dorsal view; B, head in face view; some of the light patterns on the black shiny cuticula are reflections and not real patterns; C, hindfemur outer side; D, copulatory papilla in ventral view. E, Paragryllodes milloti (Chopard, 1958), copulatory papilla in ventral view. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5540773" httpUri="https://zenodo.org/record/5540773/files/figure.png" pageId="5" pageNumber="206">Fig. 2A, B</figureCitation>
). Scapes very large, longer than wide. Pedicels with a tooth of variable size. Last article of maxillary palpi moderately elongate, slightly widened toward the apex and with apical truncation (
<figureCitation id="FADFCEE7B945FF8A172A5ED036D2FAE5" captionStart="FIGURE 2" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[169,1430,200,1820]" captionTargetId="figure-21@8.[157,1430,181,1838]" captionTargetPageId="8" captionText="FIGURE 2. Paragryllodes Karny, 1909: AD, Paragryllodes nitidula (Bolivar, 1912) n. comb.A, head and pronotum in dorsal view; B, head in face view; some of the light patterns on the black shiny cuticula are reflections and not real patterns; C, hindfemur outer side; D, copulatory papilla in ventral view. E, Paragryllodes milloti (Chopard, 1958), copulatory papilla in ventral view. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5540773" httpUri="https://zenodo.org/record/5540773/files/figure.png" pageId="5" pageNumber="206">Fig. 2B</figureCitation>
). Legs usually patterned with yellow and dark blown. Tibiae I with two apical, ventral spurs; inner tympanum present, outer tympanum absent. Tibiae II with three apical spurs. Tibiae III strongly serrulated on their basal two thirds and without spines between the first subapical spurs; with 3-4/4 subapical spurs, and three outer (the median the longest) and three inner (the median and dorsal subequal in length) apical spurs. Basitarsomeres III serrulated. FWs always present in both males and females, not covering the whole abdomen. Wings lacking. Cerci very long.
<emphasis id="50900E70B945FF8A12D25FE836B2FA39" bold="true" box="[151,219,1487,1513]" pageId="5" pageNumber="206">Male.</emphasis>
Metanotum and tergites without clear glandular structure (in particular, no metanotal area covered with dense setae and no eversible tergal region). Apical spurs of tibiae III not modified. Posterior angles of supra-anal plate not elongate. Length of subgenital plate variable. FWs usually completely overlapping, sometimes scaliform; not corneous. Stridulum variably developed, absent in micropterous species; when present, harp with many oblique, parallel veins; mirror usually short, often not clearly separated from distal field venation. Male genitalia characterized by the development of ectophallic dorsal valves and shape of pseudepiphallic sclerite: pseudepiphallic sclerite transverse or quadrate, with raised basal part; rami variable in length but not separate from the main sclerite. Pseudepiphallic parameres well developed, often partly covered with long setae, massive, not hook-shaped. Epi-ectophallic apodemes long and thick. Ectophallic dorsal valves elongate and acute, most often extending beyond pseudepiphallic distal margin, their shape differing in each species group. Endophallic apodeme comprising a small, median and vertical crest and a variable apodeme on each side of the endophallic sclerite; without a large dorsal cavity.
<emphasis id="50900E70B945FF8A177B5D1033F5F881" bold="true" box="[1342,1436,1847,1873]" pageId="5" pageNumber="206">Female.</emphasis>
FWs always present, overlapping or not. Length of ovipositor variable. Female genitalia: copulatory papilla forming a sclerotized chamber, long and tubular, or short and triangular in shape.
</paragraph>
</subSubSection>
<caption id="369B82EAB946FF8912D25D4F3317F818" ID-DOI="http://doi.org/10.5281/zenodo.5540769" ID-Zenodo-Dep="5540769" httpUri="https://zenodo.org/record/5540769/files/figure.png" pageId="6" pageNumber="207" startId="6.[151,250,1896,1920]" targetBox="[151,1436,179,1870]" targetPageId="6">
<paragraph id="625BD262B946FF8912D25D4F3317F818" blockId="6.[151,1436,1896,1992]" pageId="6" pageNumber="207">
<emphasis id="50900E70B946FF8912D25D4F377FF850" bold="true" box="[151,278,1896,1920]" pageId="6" pageNumber="207">FIGURE 1.</emphasis>
<taxonomicName id="A5E4A9E1B946FF8913655D4F37D6F850" authorityName="Blanchard" authorityYear="1845" box="[288,447,1896,1920]" class="Insecta" family="Phalangopsidae" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="family">Phalangopsidae</taxonomicName>
crickets from the Seychelles in the wild. A,
<taxonomicName id="A5E4A9E1B946FF8911D95D4F3344F850" authority="(Bolivar, 1912)" authorityName="Hugel &amp; Warren &amp; Desutter-Grandcolas" authorityYear="2021" baseAuthorityName="Bolivar" baseAuthorityYear="1912" box="[924,1325,1896,1920]" class="Insecta" family="Phalangopsidae" genus="Paragryllodes" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="nitidula">
<emphasis id="50900E70B946FF8911D95D4F32EFF850" box="[924,1158,1896,1920]" italics="true" pageId="6" pageNumber="207">Paragryllodes nitidula</emphasis>
(
<bibRefCitation id="0675AF93B946FF8916D35D4F334FF850" author="Bolivar, I." box="[1174,1318,1896,1920]" pageId="6" pageNumber="207" pagination="263 - 292" refId="ref24320" refString="Bolivar, I. (1912) Orthoptera Acrydiidae, Phasgonuridae, Gryllidae of the Seychelles. Transactions of the Linnean Society of London, Series 2, Zoologie, 15, 263 - 292. https: // doi. org / 10.1111 / j. 1096 - 3642.1912. tb 00101. x" type="journal article" year="1912">Bolivar, 1912</bibRefCitation>
)
</taxonomicName>
<emphasis id="50900E70B946FF8917725D4F33FFF850" bold="true" box="[1335,1430,1896,1920]" pageId="6" pageNumber="207">n. comb.</emphasis>
, female. B,
<taxonomicName id="A5E4A9E1B946FF8913425DAB3519F874" authority="Hugel &amp; Desutter-Grandcolas" authorityName="Hugel &amp; Desutter-Grandcolas" authorityYear="2021" box="[263,880,1932,1956]" class="Insecta" family="Phalangopsidae" genus="Phaloria" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="bolivari" status="sp. nov." subGenus="Papuloria">
<emphasis id="50900E70B946FF8913425DAB345FF874" box="[263,566,1932,1956]" italics="true" pageId="6" pageNumber="207">Phaloria (Papuloria) bolivari</emphasis>
Hugel &amp; Desutter-Grandcolas
</taxonomicName>
,
<emphasis id="50900E70B946FF89113F5DAB35DDF874" bold="true" box="[890,948,1932,1956]" pageId="6" pageNumber="207">
<taxonomicNameLabel id="4BA3B30BB946FF89113F5DAB35DDF874" box="[890,948,1932,1956]" pageId="6" pageNumber="207" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male. C, same, female. D,
<taxonomicName id="A5E4A9E1B946FF89168A5DAB374DF818" authority="Bolivar, 1912" authorityName="Bolivar" authorityYear="1912" class="Insecta" family="Phalangopsidae" genus="Phaeogryllus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="50900E70B946FF89168A5DAB33F5F874" box="[1231,1436,1932,1956]" italics="true" pageId="6" pageNumber="207">Phaeogryllus fuscus</emphasis>
<bibRefCitation id="0675AF93B946FF8912D25D97374DF818" author="Bolivar, I." box="[151,292,1968,1992]" pageId="6" pageNumber="207" pagination="263 - 292" refId="ref24320" refString="Bolivar, I. (1912) Orthoptera Acrydiidae, Phasgonuridae, Gryllidae of the Seychelles. Transactions of the Linnean Society of London, Series 2, Zoologie, 15, 263 - 292. https: // doi. org / 10.1111 / j. 1096 - 3642.1912. tb 00101. x" type="journal article" year="1912">Bolivar, 1912</bibRefCitation>
</taxonomicName>
, male. E, same, female. F,
<taxonomicName id="A5E4A9E1B946FF8910765D973222F818" authority="Hugel &amp; Desutter-Grandcolas" authorityName="Hugel &amp; Warren &amp; Desutter-Grandcolas" authorityYear="2021" box="[563,1099,1968,1992]" class="Insecta" family="Phalangopsidae" genus="Phalangacris" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="ferlegro" status="sp. nov.">
<emphasis id="50900E70B946FF8910765D97357AF818" box="[563,787,1968,1992]" italics="true" pageId="6" pageNumber="207">Phalangacris ferlegro</emphasis>
Hugel &amp; Desutter-Grandcolas
</taxonomicName>
,
<emphasis id="50900E70B946FF8916105D9732E7F818" bold="true" box="[1109,1166,1968,1992]" pageId="6" pageNumber="207">
<taxonomicNameLabel id="4BA3B30BB946FF8916105D9732E7F818" box="[1109,1166,1968,1992]" pageId="6" pageNumber="207" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
male, G, same, female.
</paragraph>
</caption>
<subSubSection id="2AFE81E9B947FF8812825AB0326FFE8F" pageId="7" pageNumber="208" type="discussion">
<paragraph id="625BD262B947FF8812825AB0326FFE8F" blockId="7.[151,1437,151,351]" pageId="7" pageNumber="208">
<emphasis id="50900E70B947FF8812825AB03759FF61" bold="true" box="[199,304,151,177]" pageId="7" pageNumber="208">Remark.</emphasis>
In
<taxonomicName id="A5E4A9E1B947FF8813165AB0379AFF61" authorityName="Karny" authorityYear="1909" box="[339,499,151,177]" class="Insecta" family="Phalangopsidae" genus="Paragryllodes" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="50900E70B947FF8813165AB0379AFF61" box="[339,499,151,177]" italics="true" pageId="7" pageNumber="208">Paragryllodes</emphasis>
</taxonomicName>
, female copulatory papilla forms an elongated sclerotized chamber of various width;
<taxonomicName id="A5E4A9E1B947FF8812D25A9E37F0FF03" authorityName="Hugel &amp; Warren &amp; Desutter-Grandcolas" authorityYear="2021" baseAuthorityName="Hugel &amp; Warren &amp; Desutter-Grandcolas" baseAuthorityYear="2021" box="[151,409,185,211]" class="Insecta" family="Phalangopsidae" genus="Paragryllodes" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="nitidula" status="comb. nov.">
<emphasis id="50900E70B947FF8812D25A9E37F0FF03" box="[151,409,185,211]" italics="true" pageId="7" pageNumber="208">Paragryllodes nitidula</emphasis>
</taxonomicName>
<emphasis id="50900E70B947FF8813E65A9E3465FF03" bold="true" box="[419,524,185,211]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="4BA3B30BB947FF8813E65A9E3465FF03" box="[419,524,185,211]" pageId="7" pageNumber="208" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
papilla (
<figureCitation id="FADFCEE7B947FF8810305A9E34B9FF03" box="[629,720,185,211]" captionStart="FIGURE 2" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[169,1430,200,1820]" captionTargetId="figure-21@8.[157,1430,181,1838]" captionTargetPageId="8" captionText="FIGURE 2. Paragryllodes Karny, 1909: AD, Paragryllodes nitidula (Bolivar, 1912) n. comb.A, head and pronotum in dorsal view; B, head in face view; some of the light patterns on the black shiny cuticula are reflections and not real patterns; C, hindfemur outer side; D, copulatory papilla in ventral view. E, Paragryllodes milloti (Chopard, 1958), copulatory papilla in ventral view. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5540773" httpUri="https://zenodo.org/record/5540773/files/figure.png" pageId="7" pageNumber="208">Fig. 2D</figureCitation>
) is similar to that of African species (see for example
<bibRefCitation id="0675AF93B947FF8817175A9D37EAFF26" author="Desutter-Grandcolas, L." pageId="7" pageNumber="208" pagination="495 - 524" refId="ref25086" refString="Desutter-Grandcolas, L. (1999) The genus Paragryllodes Karny, 1909 in tropical Africa (Orthoptera, Grylloidea, Phalangopsidae): new taxa and field observations on the habitat of the species. Zoosystema, 21, 495 - 524." type="journal article" year="1999">Desutter-Grandcolas 1999</bibRefCitation>
). We illustrate here the variation of
<taxonomicName id="A5E4A9E1B947FF8811605AFB35ACFF26" authorityName="Karny" authorityYear="1909" box="[805,965,220,246]" class="Insecta" family="Phalangopsidae" genus="Paragryllodes" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="50900E70B947FF8811605AFB35ACFF26" box="[805,965,220,246]" italics="true" pageId="7" pageNumber="208">Paragryllodes</emphasis>
</taxonomicName>
copulatory papilla with that of
<taxonomicName id="A5E4A9E1B947FF8817705AFA3756FEC9" authority="Chopard, 1958" authorityName="Chopard" authorityYear="1958" class="Insecta" family="Phalangopsidae" genus="Paragryllodes" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="milloti">
<emphasis id="50900E70B947FF8817705AFA33F5FF26" box="[1333,1436,220,246]" italics="true" pageId="7" pageNumber="208">P. milloti</emphasis>
Chopard, 1958
</taxonomicName>
since it is particularly wide in this species (
<figureCitation id="FADFCEE7B947FF8811775AD835E2FEC9" box="[818,907,255,281]" captionStart="FIGURE 2" captionStartId="8.[151,250,1862,1886]" captionTargetBox="[169,1430,200,1820]" captionTargetId="figure-21@8.[157,1430,181,1838]" captionTargetPageId="8" captionText="FIGURE 2. Paragryllodes Karny, 1909: AD, Paragryllodes nitidula (Bolivar, 1912) n. comb.A, head and pronotum in dorsal view; B, head in face view; some of the light patterns on the black shiny cuticula are reflections and not real patterns; C, hindfemur outer side; D, copulatory papilla in ventral view. E, Paragryllodes milloti (Chopard, 1958), copulatory papilla in ventral view. Scales 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5540773" httpUri="https://zenodo.org/record/5540773/files/figure.png" pageId="7" pageNumber="208">Fig. 2E</figureCitation>
). This species is not related to other species of the subgenus
<taxonomicName id="A5E4A9E1B947FF8813755B0637D0FEEB" authorityName="Gorochov" authorityYear="2015" box="[304,441,289,315]" class="Insecta" family="Phalangopsidae" genus="Cortigryllus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="50900E70B947FF8813755B0637D0FEEB" box="[304,441,289,315]" italics="true" pageId="7" pageNumber="208">Cortigryllus</emphasis>
</taxonomicName>
where it has recently been tentatively placed (
<bibRefCitation id="0675AF93B947FF8811855B06321FFEEC" author="Gorochov, A. V." box="[960,1142,289,316]" pageId="7" pageNumber="208" pagination="600 - 611" refId="ref25951" refString="Gorochov, A. V. (2015) New and little-known crickets of the subfamily Phalangopsinae (Orthoptera, Gryllidae). 8. The Genus Paragryllodes and notes on the subfamily classification. Zoologicheskii Zhurnal, 94, 905 - 916. [in Russian, Entomological Review, 95, 600 - 611 (2015)] https: // doi. org / 10.1134 / S 001387381505005 X" type="journal article" year="2015">Gorochov 2015</bibRefCitation>
); a review of species from
<collectingCountry id="1AF392F2B947FF8812D25B633697FE8E" box="[151,254,324,350]" name="Comoros" pageId="7" pageNumber="208">Comoros</collectingCountry>
will be addressed separately (Desutter-Grandcolas &amp; Hugel in prep).
</paragraph>
</subSubSection>
</treatment>
</document>