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<document id="BBE99DF63B979DDA2979FAC2156CB724" ID-DOI="10.5852/ejt.2024.956.2657" ID-ISSN="2118-9773" ID-Zenodo-Dep="13742850" ID-ZooBank="9DCDD521-909D-4436-8495-086A11C50DA0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1725988271530" checkinUser="plazi" docAuthor="Karimbumkara, Seena Narayanan &amp; Priyadarsanan, Dharma Rajan" docDate="2024" docId="557187D2FFF389019D5AFB71FABDF814" docLanguage="en" docName="EJT.2024.956.1-61.pdf" docOrigin="European Journal of Taxonomy 956" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onitis assamensis Biswas 1980" docType="treatment" docVersion="4" lastPageNumber="8" masterDocId="A948FFAAFFF689069F01FFB1FFF9FF8A" masterDocTitle="Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India" masterLastPageNumber="61" masterPageNumber="1" pageNumber="6" updateTime="1726486743805" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7DD3ACC707977762F428119793D243B5">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
</mods:titleInfo>
<mods:name id="0148D68FB0BEBF5D68B1E98BBF080AD6" type="personal">
<mods:role id="23B65764AC28538FBB9C09A67A26D378">
<mods:roleTerm id="86282C8313DA5FEB9ECE08116DA336A6">Author</mods:roleTerm>
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<mods:namePart id="F0F16037D6021ACC0A994D0F8232340F">Karimbumkara, Seena Narayanan</mods:namePart>
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<mods:affiliation id="69B0B32114ED876A123B896CC221D9A5">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
<mods:nameIdentifier id="C8B02BE6403FD3E6A1035C8DD03CA5B9" type="email">seena.narayanan@atree.org</mods:nameIdentifier>
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<mods:namePart id="E9DFEDE594AE65877CB136086E13A445">Priyadarsanan, Dharma Rajan</mods:namePart>
<mods:nameIdentifier id="9B75F24CC31D7C4712C301DFCDCD371D" type="LSID">03907E93-F783-4720-9F7B-CB662E7FC745</mods:nameIdentifier>
<mods:affiliation id="3136DDFEAD34BC779EAB6ACAFA1C4450">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
<mods:nameIdentifier id="886C02EA38D538DCA2985EB80301C1C8" type="email">priyan@atree.org</mods:nameIdentifier>
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<mods:title id="37AF929970819C6401767421E16142C4">European Journal of Taxonomy</mods:title>
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<mods:date id="3EC9690E03B6D7F20D32849D2A8CEED0">2024</mods:date>
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<mods:number id="6BA4301EA8AB4C006887DD919083E847">2024-09-10</mods:number>
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<mods:number id="48BA8E2EFFC56B37630B0F268927BE27">956</mods:number>
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<treatment id="557187D2FFF389019D5AFB71FABDF814" ID-DOI="http://doi.org/10.5281/zenodo.13742857" ID-Zenodo-Dep="13742857" LSID="urn:lsid:plazi:treatment:557187D2FFF389019D5AFB71FABDF814" httpUri="http://treatment.plazi.org/id/557187D2FFF389019D5AFB71FABDF814" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
<subSubSection id="95C2654FFFF389039D5AFB71FC20FB50" box="[603,985,1216,1242]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="DD6736C4FFF389039D5AFB71FC20FB50" blockId="5.[603,985,1216,1277]" box="[603,985,1216,1242]" pageId="5" pageNumber="6">
<heading id="862F81A8FFF389039D5AFB71FC20FB50" box="[603,985,1216,1242]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
<taxonomicName id="1AD84D47FFF389039D5AFB71FC20FB50" ID-CoL="6SMW9" authority="Biswas, 1980" authorityName="Biswas" authorityYear="1980" box="[603,985,1216,1242]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF389039D5AFB71FCCAFB50" bold="true" box="[603,819,1216,1242]" italics="true" pageId="5" pageNumber="6">Onitis assamensis</emphasis>
<bibRefCitation id="B9494B35FFF389039C3BFB71FC20FB50" author="Biswas S." box="[826,985,1216,1242]" pageId="5" pageNumber="6" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas, 1980</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF389039DB5FB52FC86FB77" box="[692,895,1251,1277]" pageId="5" pageNumber="6" type="description">
<paragraph id="DD6736C4FFF389039DB5FB52FC86FB77" blockId="5.[603,985,1216,1277]" box="[692,895,1251,1277]" pageId="5" pageNumber="6">
<figureCitation id="45E32A41FFF389039DB5FB52FD04FB77" box="[692,765,1251,1277]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[262,1340,366,1854]" captionTargetId="figure-1@6.[189,1398,315,1882]" captionTargetPageId="6" captionText="Fig. 1. Onitis assamensis Biswas, 1980, ♂ (A–D), AIMB/Co/Sc1000001; ♀ (E–F), AIMB/Co/ Sc1000009. A. Habitus, dorsal view. B. Habitus, ventral view. C. Protibia. D. Pygidium. E. Habitus, dorsal view. F. Habitus, ventral view. (Images: AIMB)" figureDoi="http://doi.org/10.5281/zenodo.13742852" httpUri="https://zenodo.org/record/13742852/files/figure.png" pageId="5" pageNumber="6">Figs 1</figureCitation>
,
<figureCitation id="45E32A41FFF389039C0AFB52FCAFFB77" box="[779,854,1251,1277]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. A–D. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). E–F. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). G–J. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). K–N. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742861" httpUri="https://zenodo.org/record/13742861/files/figure.png" pageId="5" pageNumber="6">4A–D</figureCitation>
,
<figureCitation id="45E32A41FFF389039C63FB52FC86FB77" box="[866,895,1251,1277]" captionStart="Fig" captionStartId="47.[189,232,1930,1956]" captionTargetBox="[223,1365,249,1879]" captionTargetId="figure-1@47.[208,1379,236,1891]" captionTargetPageId="47" captionText="Fig. 17. Map showing the distribution of OnitisassamensisBiswas, 1980; O. bhomorensis sp. nov.; O. bordatiCambefort, 1988; O. brahma Lansberge, 1875 and O. castaneus Redtenbacher, 1848 in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742901" httpUri="https://zenodo.org/record/13742901/files/figure.png" pageId="5" pageNumber="6">17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF389039FBCFA94FAAEFA0A" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="DD6736C4FFF389039FBCFA94FC81FAB4" blockId="5.[189,888,1316,1343]" box="[189,888,1316,1343]" pageId="5" pageNumber="6">
<emphasis id="EFACEAD6FFF389039FBCFA94FE69FAB5" box="[189,400,1317,1343]" italics="true" pageId="5" pageNumber="6">Onitisassamensis</emphasis>
<bibRefCitation id="B9494B35FFF389039E96FA95FD88FAB5" author="Biswas S." box="[407,625,1316,1343]" pageId="5" pageNumber="6" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas, 1980: 339</bibRefCitation>
(original description).
</paragraph>
<paragraph id="DD6736C4FFF389039FBCFAD7FAAEFA0A" blockId="5.[189,1367,1382,1409]" box="[189,1367,1382,1409]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="FDC811EAFFF389039FBCFAD7FAAEFA0A" box="[189,1367,1382,1409]" pageId="5" pageNumber="6">
<taxonomicName id="1AD84D47FFF389039FBCFAD7FE69FA0A" ID-CoL="6SMW9" authorityName="Biswas" authorityYear="1980" box="[189,400,1382,1408]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF389039FBCFAD7FE69FA0A" box="[189,400,1382,1408]" italics="true" pageId="5" pageNumber="6">Onitis assamensis</emphasis>
</taxonomicName>
–
<treatmentCitation id="5C7910D5FFF389039EACFAD7FD3AFA0A" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[429,707,1382,1409]" page="1037" pageId="5" pageNumber="6" year="2015">
<bibRefCitation id="B9494B35FFF389039EACFAD7FD3AFA0A" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[429,707,1382,1409]" pageId="5" pageNumber="6" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFF389039EFDFAD6FDCEFA0A" box="[508,567,1382,1408]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
2015: 1037
</bibRefCitation>
</treatmentCitation>
(checklist). —
<bibRefCitation id="B9494B35FFF389039C7BFAD7FB8DFA0A" author="Schoolmeesters P." box="[890,1140,1382,1408]" pageId="5" pageNumber="6" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF389039FBCFA16FDECF883" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="DD6736C4FFF389039FBCFA16FE3FFA4B" blockId="5.[189,454,1447,1473]" box="[189,454,1447,1473]" pageId="5" pageNumber="6">
<heading id="862F81A8FFF389039FBCFA16FE3FFA4B" bold="true" box="[189,454,1447,1473]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="EFACEAD6FFF389039FBCFA16FE3FFA4B" bold="true" box="[189,454,1447,1473]" pageId="5" pageNumber="6">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFF389039FBCFA67FDECF883" blockId="5.[189,1399,1494,1802]" pageId="5" pageNumber="6">
<taxonomicName id="1AD84D47FFF389039FBCFA67FE6CFA7A" authorityName="Biswas" authorityYear="1980" box="[189,405,1494,1520]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF389039FBCFA67FE6CFA7A" box="[189,405,1494,1520]" italics="true" pageId="5" pageNumber="6">Onitis assamensis</emphasis>
</taxonomicName>
and
<taxonomicName id="1AD84D47FFF389039ECFFA67FDA3FA7A" authorityName="Cambefort" authorityYear="1988" box="[462,602,1494,1520]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="naviauxi">
<emphasis id="EFACEAD6FFF389039ECFFA67FDA3FA7A" box="[462,602,1494,1520]" italics="true" pageId="5" pageNumber="6">O. naviauxi</emphasis>
</taxonomicName>
differ from each other in the number of teeth on the anteriomedian ridge of the protibia in the male. Clypeal carina short in male of
<taxonomicName id="1AD84D47FFF389039CF5FA4BFB5CF999" authorityName="Biswas" authorityYear="1980" box="[1012,1189,1530,1556]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF389039CF5FA4BFB5CF999" box="[1012,1189,1530,1556]" italics="true" pageId="5" pageNumber="6">O. assamensis</emphasis>
</taxonomicName>
and
<taxonomicName id="1AD84D47FFF389039BE7FA4BFA8FF999" authorityName="Cambefort" authorityYear="1988" box="[1254,1398,1530,1556]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="naviauxi">
<emphasis id="EFACEAD6FFF389039BE7FA4BFA8FF999" box="[1254,1398,1530,1556]" italics="true" pageId="5" pageNumber="6">O. naviauxi</emphasis>
</taxonomicName>
while it is longer in female; clypeus in female pointed in both species; metasternal shield hairy in males and smooth in females. In the original description of
<taxonomicName id="1AD84D47FFF389039CCBF9F1FB80F9D3" authorityName="Biswas" authorityYear="1980" box="[970,1145,1600,1626]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF389039CCBF9F1FB80F9D3" box="[970,1145,1600,1626]" italics="true" pageId="5" pageNumber="6">O. assamensis</emphasis>
</taxonomicName>
the frontal carina is considered entire, but on observing the specimen in detail, it is confirmed that it is interrupted due to the placement of tubercle on it. The teeth on the infero-internal ridge in the profemur is a very prominent character of the species and the arrangement of teeth, their number and size may vary with individuals. In a specimen we collected, an extra small tooth was present near the femoro-tibial joint on one of the fore legs.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF389019FBCF880FEB3FE07" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="DD6736C4FFF389039FBCF880FE53F8C1" blockId="5.[189,426,1841,1867]" box="[189,426,1841,1867]" pageId="5" pageNumber="6">
<heading id="862F81A8FFF389039FBCF880FE53F8C1" bold="true" box="[189,426,1841,1867]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="EFACEAD6FFF389039FBCF880FE53F8C1" bold="true" box="[189,426,1841,1867]" pageId="5" pageNumber="6">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="6DB03C99FFF389039FEBF8D1FE0FF84A" collectingDate="1972-12-21" collectionCode="ZSIM" collectorName="A. K. Ghosh" country="India" location="Kaziranga National Park" pageId="5" pageNumber="6" specimenCode="A1/4397" specimenCount="1" specimenCount-female="1" stateProvince="Assam" typeStatus="holotype">
<paragraph id="DD6736C4FFF389039FEBF8D1FEA5F8F0" blockId="5.[189,1399,1888,1984]" box="[234,348,1888,1914]" pageId="5" pageNumber="6">
<emphasis id="EFACEAD6FFF389039FEBF8D1FEA5F8F0" bold="true" box="[234,348,1888,1914]" pageId="5" pageNumber="6">
<typeStatus id="02638866FFF389039FEBF8D1FEA5F8F0" box="[234,348,1888,1914]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFF389039FBCF832FE0FF84A" blockId="5.[189,1399,1888,1984]" pageId="5" pageNumber="6">
<collectingCountry id="A5CF7654FFF389039FBCF832FEF7F817" box="[189,270,1923,1949]" name="India" pageId="5" pageNumber="6">INDIA</collectingCountry>
•
<specimenCount id="CBDEFD4DFFF389039E2AF832FEBBF817" box="[299,322,1923,1949]" count="1" pageId="5" pageNumber="6" type="female">♀</specimenCount>
;
<collectingRegion id="1F1CF826FFF389039E51F832FE5AF817" box="[336,419,1923,1949]" country="India" name="Assam" pageId="5" pageNumber="6">Assam</collectingRegion>
,
<location id="D807601FFFF389039EB0F832FD22F817" LSID="urn:lsid:plazi:treatment:557187D2FFF389019D5AFB71FABDF814:D807601FFFF389039EB0F832FD22F817" box="[433,731,1923,1949]" country="India" name="Kaziranga National Park" pageId="5" pageNumber="6" stateProvince="Assam">Kaziranga National Park</location>
;
<date id="A9661004FFF389039DE9F832FC75F817" box="[744,908,1923,1949]" pageId="5" pageNumber="6" value="1972-12-21">
<collectingDate id="B922E9ECFFF389039DE9F832FC75F817" box="[744,908,1923,1949]" pageId="5" pageNumber="6" value="1972-12-21">21 Dec. 1972</collectingDate>
</date>
;
<collectorName id="702D5312FFF389039C99F832FBDEF817" box="[920,1063,1923,1949]" pageId="5" pageNumber="6">A.K. Ghosh</collectorName>
leg.; from rhinoceros dung;
<collectionCode id="BBC9AE01FFF389039FBCF817FEFAF84A" box="[189,259,1958,1984]" pageId="5" pageNumber="6">ZSIM</collectionCode>
Regd. No.
<specimenCode id="8D7E9EBFFFF389039E89F817FE0BF84A" box="[392,498,1958,1984]" pageId="5" pageNumber="6">A1/4397</specimenCode>
.
</paragraph>
</materialsCitation>
<caption id="89A7664CFFF089009FBCF839FCC7F862" ID-DOI="http://doi.org/10.5281/zenodo.13742852" ID-Zenodo-Dep="13742852" httpUri="https://zenodo.org/record/13742852/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1928,1954]" targetBox="[262,1340,366,1854]" targetPageId="6">
<paragraph id="DD6736C4FFF089009FBCF839FCC7F862" blockId="6.[189,1398,1928,2025]" pageId="6" pageNumber="7">
<emphasis id="EFACEAD6FFF089009FBCF839FEECF828" bold="true" box="[189,277,1928,1954]" pageId="6" pageNumber="7">Fig. 1.</emphasis>
<taxonomicName id="1AD84D47FFF089009E24F839FD40F828" authority="Biswas, 1980" authorityName="Biswas" authorityYear="1980" box="[293,697,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF089009E24F839FDF8F828" box="[293,513,1928,1954]" italics="true" pageId="6" pageNumber="7">Onitis assamensis</emphasis>
<bibRefCitation id="B9494B35FFF089009D13F839FD40F828" author="Biswas S." box="[530,697,1928,1954]" pageId="6" pageNumber="7" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas, 1980</bibRefCitation>
</taxonomicName>
, ♂ (A–D), AIMB/Co/Sc1000001; ♀ (E–F), AIMB/Co/ Sc1000009.
<emphasis id="EFACEAD6FFF089009E53F81AFE9EF84F" bold="true" box="[338,359,1963,1989]" pageId="6" pageNumber="7">A</emphasis>
. Habitus, dorsal view.
<emphasis id="EFACEAD6FFF089009D7FF81AFD6BF84F" bold="true" box="[638,658,1963,1989]" pageId="6" pageNumber="7">B</emphasis>
. Habitus, ventral view.
<emphasis id="EFACEAD6FFF089009CB3F81AFC3EF84F" bold="true" box="[946,967,1963,1989]" pageId="6" pageNumber="7">C</emphasis>
. Protibia.
<emphasis id="EFACEAD6FFF089009B47F81AFBA2F84F" bold="true" box="[1094,1115,1963,1989]" pageId="6" pageNumber="7">D</emphasis>
. Pygidium.
<emphasis id="EFACEAD6FFF089009BEEF81AFAFAF84F" bold="true" box="[1263,1283,1963,1989]" pageId="6" pageNumber="7">E</emphasis>
. Habitus, dorsal view.
<emphasis id="EFACEAD6FFF089009E53F87FFE9DF862" bold="true" box="[338,356,1998,2024]" pageId="6" pageNumber="7">F</emphasis>
. Habitus, ventral view. (Images: AIMB)
</paragraph>
</caption>
<paragraph id="DD6736C4FFF189019FEBFEB8FD62FEA9" blockId="7.[189,1398,265,397]" box="[234,667,265,292]" pageId="7" pageNumber="8">
<emphasis id="EFACEAD6FFF189019FEBFEB8FE18FEA9" bold="true" box="[234,481,265,291]" pageId="7" pageNumber="8">Additional material</emphasis>
(
<specimenCount id="CBDEFD4DFFF189019EF3FEB8FD6AFEA9" box="[498,659,265,292]" count="15" pageId="7" pageNumber="8" type="generic">15 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFF189019FBCFE9DFEB3FE07" blockId="7.[189,1398,265,397]" pageId="7" pageNumber="8">
<materialsCitation id="6DB03C99FFF189019FBCFE9DFEBCFE07" collectingDate="2019-05-21" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan &amp; Karimbumkara Seena Narayanan" country="India" location="Kaziranga National Park" pageId="7" pageNumber="8" specimenCode="AIMB/Co/Sc1000001, AIMB/Co/Sc1000002, AIMB/Co/Sc1000003, AIMB/Co/Sc1000004, AIMB/Co/Sc1000005, AIMB/Co/Sc1000006, AIMB/Co/Sc1000007, AIMB/Co/Sc1000008, AIMB/Co/Sc1000009, AIMB/Co/Sc1000010, AIMB/Co/Sc1000011, AIMB/Co/Sc1000012, AIMB/Co/Sc1000013, AIMB/Co/Sc1000014, AIMB/Co/Sc1000015" specimenCount="15" specimenCount-female="7" specimenCount-male="8" stateProvince="Assam">
<collectingCountry id="A5CF7654FFF189019FBCFE9DFEF7FECC" box="[189,270,300,326]" name="India" pageId="7" pageNumber="8">INDIA</collectingCountry>
•
<specimenCount id="CBDEFD4DFFF189019E2AFE9DFE88FECD" box="[299,369,300,327]" count="8" pageId="7" pageNumber="8" type="male">8 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFF189019E7EFE9CFE3DFECD" box="[383,452,301,327]" count="7" pageId="7" pageNumber="8" type="female">7 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFF189019ED2FE9DFDDFFECC" box="[467,550,300,326]" country="India" name="Assam" pageId="7" pageNumber="8">Assam</collectingRegion>
,
<location id="D807601FFFF189019D32FE9DFCA4FECC" LSID="urn:lsid:plazi:treatment:557187D2FFF389019D5AFB71FABDF814:D807601FFFF189019D32FE9DFCA4FECC" box="[563,861,300,326]" country="India" name="Kaziranga National Park" pageId="7" pageNumber="8" stateProvince="Assam">Kaziranga National Park</location>
;
<date id="A9661004FFF189019C6BFE9DFBF4FECC" box="[874,1037,300,327]" pageId="7" pageNumber="8" value="2019-05-21">
<collectingDate id="B922E9ECFFF189019C6BFE9DFBF4FECC" box="[874,1037,300,327]" pageId="7" pageNumber="8" value="2019-05-21">21 May 2019</collectingDate>
</date>
;
<collectorName id="702D5312FFF189019B1AFE9DFA8FFECC" box="[1051,1398,300,326]" pageId="7" pageNumber="8">Priyadarsanan Dharma Rajan</collectorName>
&amp;
<collectorName id="702D5312FFF189019FDCFEFEFD9CFEE0" box="[221,613,335,362]" pageId="7" pageNumber="8">Karimbumkara Seena Narayanan</collectorName>
leg.; hand–picked, rhinoceros dung;
<collectionCode id="BBC9AE01FFF189019B1EFEE1FB95FEE0" box="[1055,1132,336,362]" pageId="7" pageNumber="8">AIMB</collectionCode>
<specimenCode id="8D7E9EBFFFF189019B72FEFEFEBCFE07" pageId="7" pageNumber="8">AIMB/Co/Sc1000001 to 1000015</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF189019FBCFE05FED5F984" pageId="7" pageNumber="8" type="description">
<paragraph id="DD6736C4FFF189019FBCFE05FEB7FE44" blockId="7.[189,334,436,462]" box="[189,334,436,462]" pageId="7" pageNumber="8">
<heading id="862F81A8FFF189019FBCFE05FEB7FE44" bold="true" box="[189,334,436,462]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="EFACEAD6FFF189019FBCFE05FEB7FE44" bold="true" box="[189,334,436,462]" pageId="7" pageNumber="8">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFF189019FBCFE52FDD8FDAB" blockId="7.[189,1399,482,545]" pageId="7" pageNumber="8">MEASUREMENTS (in mm). TL=21.5–24.8; BW =10.7–12.5; PL=8.5–11; PW= 10.5–13.5; EL=10.5– 12.5; HL= 5–6; HW=5.5–6.8.</paragraph>
<paragraph id="DD6736C4FFF189019FEBFDF6FED3FDEB" blockId="7.[189,1399,583,1103]" box="[234,298,583,609]" pageId="7" pageNumber="8">
<emphasis id="EFACEAD6FFF189019FEBFDF6FED3FDEB" bold="true" box="[234,298,583,609]" pageId="7" pageNumber="8">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFF189019FBCFDDBFAAFFBC5" blockId="7.[189,1399,583,1103]" pageId="7" pageNumber="8">Body shiny green or bronzy, broadly oval. Head rounded, clypeus straight in the middle anteriorly, finely rugose with scattered punctures; clypeal carina short, rugose behind; frontal carina almost straight, narrowly interrupted in the middle by the placement of a strong conical blunt tubercle; vertex sparsely punctured, elevated towards the eyes with a depression in the middle devoid of any punctures. Pronotum with small, close dense punctures between the basal foveae, punctures larger and less close anterior to the foveae, finer and sparse anteriorly and at the sides; basal foveae oval, deep, connected by a bordered base; scutellum triangular visibly large. Elytra broadly curved at the sides; striae deep, broader at the base, narrow and smooth towards the tip; elytral intervals smooth without punctures, sutural interval slightly convex. Pygidium margined, lesser sheen, imperceptibly punctured. Metasternal shield with minute punctures, front angles with minute granules and fine yellow hairs; sides of metasternum with reddish hairs, densely granular. Fore legs with rows of reddish long hairs or setae on the ridges; a strong curved tooth on the anteromedian ridge of profemur with a minute oblique denticle near the base; protibia with four distantly placed external teeth, the infero-medial ridge with 6–7 teeth, small proximally and stouter distally; mid femur unarmed; hind femur with a strong curved lateral tooth towards the apex.</paragraph>
<paragraph id="DD6736C4FFF189019FBCFBC7FD7EFB5C" blockId="7.[189,1399,1141,1238]" pageId="7" pageNumber="8">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFF189019EBFFBC4FDBFFB1A" box="[446,582,1141,1168]" captionStart="Fig" captionStartId="13.[189,232,1856,1882]" captionTargetBox="[300,1241,290,1779]" captionTargetId="figure-75@13.[189,1398,262,1786]" captionTargetPageId="13" captionText="Fig. 4. A–D. Onitis assamensis Biswas, 1980, Aedeagus, lateral, dorsal, ventral and apical views, AIMB/ Co/Sc1000001 (images: AIMB). E–F. O. naviauxi Cambefort, 1988, lateral and dorsal views, MNHN EC4807, (images: Antoine Mantilleri, MNHN). G–J. O. bordati Cambefort, 1988, aedeagus; lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000016, (images AIMB). K–N. O. siva Gillet, 1911, aedagus, lateral, dorsal, ventral and apical views, AIMB/Co/Sc1000704, (images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742861" httpUri="https://zenodo.org/record/13742861/files/figure.png" pageId="7" pageNumber="8">Fig. 4A–D</figureCitation>
). LP=3.6, Lp =2.86; BP=1.86, BpB=1.43, BpT= 0.71. Phallobase longer than parameres, broader at base and further tapers, becomes blunt and slightly curved at the tip. Phallobase slightly broader at the base.
</paragraph>
<paragraph id="DD6736C4FFF189019FEBFB4CFEBCFA9D" blockId="7.[189,1399,1277,1550]" box="[234,325,1277,1303]" pageId="7" pageNumber="8">
<emphasis id="EFACEAD6FFF189019FEBFB4CFEBCFA9D" bold="true" box="[234,325,1277,1303]" pageId="7" pageNumber="8">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFF189019FBCFA91FED5F984" blockId="7.[189,1399,1277,1550]" pageId="7" pageNumber="8">Body greenish, broadly oval, not very convex; head sub triangular, frontal carina entire with a tubercle. Clypeal carina long and curved placed almost midway between the clypeal margin and the frontal carina; clypeus parabolic, rugulose. Pronotum strongly and closely punctured; front angles sharp; basal foveae rounded. Elytra: dull, striae shallow and broad, intervals flat, not perceptibly punctate. Ventral surface: metasternum flat, smooth with two deep pits behind and a shallow longitudinal groove between; sides of metasternum closely granular and hairy. Pygidium without hair, slightly shining and coarse; legs unarmed.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF189019FBCF984FC0CF9F4" box="[189,1013,1589,1662]" pageId="7" pageNumber="8" type="distribution">
<paragraph id="DD6736C4FFF189019FBCF984FEAFF9C5" blockId="7.[189,342,1589,1615]" box="[189,342,1589,1615]" pageId="7" pageNumber="8">
<heading id="862F81A8FFF189019FBCF984FEAFF9C5" bold="true" box="[189,342,1589,1615]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="EFACEAD6FFF189019FBCF984FEAFF9C5" bold="true" box="[189,342,1589,1615]" pageId="7" pageNumber="8">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFF189019FBCF9D5FC0CF9F4" blockId="7.[189,1013,1635,1662]" box="[189,1013,1635,1662]" pageId="7" pageNumber="8">
<collectingCountry id="A5CF7654FFF189019FBCF9D5FF07F9F4" box="[189,254,1636,1662]" name="India" pageId="7" pageNumber="8">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFF189019E06F9D5FEAEF9F4" box="[263,343,1636,1662]" country="India" name="Assam" pageId="7" pageNumber="8">Assam</collectingRegion>
(
<bibRefCitation id="B9494B35FFF189019E67F9D5FDFBF9F4" author="Biswas S." box="[358,514,1636,1662]" pageId="7" pageNumber="8" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas 1980</bibRefCitation>
;
<bibRefCitation id="B9494B35FFF189019D0FF9D5FD24F9F4" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[526,733,1635,1662]" pageId="7" pageNumber="8" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFF189019D5CF9D5FD61F9F4" box="[605,664,1636,1662]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
2015
</bibRefCitation>
;
<bibRefCitation id="B9494B35FFF189019DE8F9D5FC10F9F4" author="Schoolmeesters P." box="[745,1001,1636,1662]" pageId="7" pageNumber="8" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFF189019FBCF914FABDF814" pageId="7" pageNumber="8" type="discussion">
<paragraph id="DD6736C4FFF189019FBCF914FED6F935" blockId="7.[189,303,1701,1727]" box="[189,303,1701,1727]" pageId="7" pageNumber="8">
<heading id="862F81A8FFF189019FBCF914FED6F935" bold="true" box="[189,303,1701,1727]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="EFACEAD6FFF189019FBCF914FED6F935" bold="true" box="[189,303,1701,1727]" pageId="7" pageNumber="8">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFF189019FBCF965FABDF814" blockId="7.[189,1399,1748,1950]" pageId="7" pageNumber="8">
A single female specimen of
<taxonomicName id="1AD84D47FFF189019D14F965FD47F967" authorityName="Biswas" authorityYear="1980" box="[533,702,1748,1774]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="EFACEAD6FFF189019D14F965FD47F967" box="[533,702,1748,1774]" italics="true" pageId="7" pageNumber="8">O. assamensis</emphasis>
</taxonomicName>
was collected by
<bibRefCitation id="B9494B35FFF189019C93F965FBC4F964" author="Biswas S." box="[914,1085,1748,1774]" pageId="7" pageNumber="8" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas (1980)</bibRefCitation>
from rhinoceros dung and described as the
<typeStatus id="02638866FFF189019E7FF946FE10F89B" box="[382,489,1783,1809]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
. During our visit to Kaziranga National Park, we have collected
<specimenCount id="CBDEFD4DFFF189019BD9F946FA8FF89B" box="[1240,1398,1783,1810]" count="15" pageId="7" pageNumber="8" type="generic">15 specimens</specimenCount>
of the species of which eight were males and
<specimenCount id="CBDEFD4DFFF189019DEBF8AAFC6FF8BE" box="[746,918,1818,1845]" count="7" pageId="7" pageNumber="8" type="female">seven females</specimenCount>
. The rarity of the species mentioned by Biswas can be attributed to the winter in December when he had made the collection. The species was common and active in May (pre-monsoon), when we collected them at the same location. We have described the male of
<emphasis id="EFACEAD6FFF189019EC1F835FD6AF814" box="[448,659,1924,1950]" italics="true" pageId="7" pageNumber="8">Onitisassamensis</emphasis>
which had never previously been discovered or described.
</paragraph>
</subSubSection>
</treatment>
</document>
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