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<document id="E1840B93DE63BD5CE0AE9213B8CB10AF" ID-DOI="10.5852/ejt.2024.956.2657" ID-ISSN="2118-9773" ID-Zenodo-Dep="13742850" ID-ZooBank="9DCDD521-909D-4436-8495-086A11C50DA0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1725988271530" checkinUser="plazi" docAuthor="Karimbumkara, Seena Narayanan &amp; Priyadarsanan, Dharma Rajan" docDate="2024" docId="557187D2FFE589129D69FE6CFCB4FB1F" docLanguage="en" docName="EJT.2024.956.1-61.pdf" docOrigin="European Journal of Taxonomy 956" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2657/12287" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Onitis excavatus Arrow 1931" docType="treatment" docVersion="4" lastPageNumber="21" masterDocId="A948FFAAFFF689069F01FFB1FFF9FF8A" masterDocTitle="Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India" masterLastPageNumber="61" masterPageNumber="1" pageNumber="20" updateTime="1726486743805" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="1BD680F7A9CF73329B0212A0CC55930C">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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<mods:namePart id="D70D5F58B01838DDB3D8F33BB646EFE3">Karimbumkara, Seena Narayanan</mods:namePart>
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<mods:namePart id="FE3467E9811FF5DA013E22F57FF830B1">Priyadarsanan, Dharma Rajan</mods:namePart>
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<mods:affiliation id="F09C6D464F63D66BD96E7097CEC5AA95">Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Jakkur P. O., Bangalore 560064, Karnataka, India.</mods:affiliation>
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<mods:date id="F7032FE661BDC791C840A1D7D18A4D33">2024</mods:date>
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<paragraph id="DD6736C4FFE589159D69FE6CFC35FE7D" blockId="19.[616,972,477,539]" box="[616,972,477,503]" pageId="19" pageNumber="20">
<heading id="862F81A8FFE589159D69FE6CFC35FE7D" box="[616,972,477,503]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="20" reason="2">
<taxonomicName id="1AD84D47FFE589159D69FE6CFC35FE7D" ID-CoL="49KG2" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[616,972,477,503]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="EFACEAD6FFE589159D69FE6CFCC9FE7D" bold="true" box="[616,816,477,503]" italics="true" pageId="19" pageNumber="20">Onitis excavatus</emphasis>
<bibRefCitation id="B9494B35FFE589159C36FE6CFC35FE7D" author="Arrow G. J." box="[823,972,477,503]" pageId="19" pageNumber="20" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE589159DAAFDB1FC71FD90" box="[683,904,512,539]" pageId="19" pageNumber="20" type="description">
<paragraph id="DD6736C4FFE589159DAAFDB1FC71FD90" blockId="19.[616,972,477,539]" box="[683,904,512,539]" pageId="19" pageNumber="20">
<figureCitation id="45E32A41FFE589159DAAFDB1FCF0FD91" box="[683,777,512,539]" captionStart="Fig" captionStartId="14.[189,233,1811,1837]" captionTargetBox="[327,1251,267,1773]" captionTargetId="figure-84@14.[189,1398,262,1777]" captionTargetPageId="14" captionText="Fig. 5. A. Onitis bordati Cambefort, 1988, holotype, ♂, MNHN EC1187, (image: Antoine Mantilleri, MNHN). B. O. brahma Lansberge, 1875, ♂, BMNH (E) 1237157, (image: BMNH). C. O.castaneus Redtenbacher, 1848, syntype, (image: Harald Schillhammer, NHMW). D. O. crassus Sharp, 1875, holotype, ♂, MNHN EC4813, (image: Antoine Mantilleri, MNHN). E. O. excavatus Arrow, 1931, lectotype, ♂, BMNH (E) 1237173, (image: BMNH). F. O. falcatus (Wulfen, 1786), ♂, AIMB/Co/ Sc1000070, (image: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742863" httpUri="https://zenodo.org/record/13742863/files/figure.png" pageId="19" pageNumber="20">Figs 5E</figureCitation>
,
<figureCitation id="45E32A41FFE589159C15FDB1FCA6FD90" box="[788,863,512,538]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. EH. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. IL. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. MP. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742867" httpUri="https://zenodo.org/record/13742867/files/figure.png" pageId="19" pageNumber="20">6AD</figureCitation>
,
<figureCitation id="45E32A41FFE589159C6AFDB1FC71FD90" box="[875,904,512,538]" captionStart="Fig" captionStartId="48.[190,233,1927,1953]" captionTargetBox="[224,1364,251,1878]" captionTargetId="figure-1@48.[209,1378,238,1890]" captionTargetPageId="48" captionText="Fig. 18. Map showing the distribution of Onitis crassus Sharp, 1875; O. excavatus Arrow, 1931; O. falcatus(Wulfen, 1789); O. feae Felsche, 1907; O. humerosus (Pallas, 1771) and O. kethai sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742903" httpUri="https://zenodo.org/record/13742903/files/figure.png" pageId="19" pageNumber="20">18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE589159FBCFDF3FC33FCA0" pageId="19" pageNumber="20" type="reference_group">
<paragraph id="DD6736C4FFE589159FBCFDF3FCA4FDD6" blockId="19.[189,861,578,604]" box="[189,861,578,604]" pageId="19" pageNumber="20">
<emphasis id="EFACEAD6FFE589159FBCFDF3FE79FDD6" box="[189,384,578,604]" italics="true" pageId="19" pageNumber="20">Onitisexcavatus</emphasis>
<bibRefCitation id="B9494B35FFE589159E84FDF3FDACFDD6" author="Arrow G. J." box="[389,597,578,604]" pageId="19" pageNumber="20" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931: 391</bibRefCitation>
(original description).
</paragraph>
<paragraph id="DD6736C4FFE589159FBCFD35FC33FCA0" blockId="19.[189,1399,643,811]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="FDC811EAFFE589159FBCFD35FC33FCA0" pageId="19" pageNumber="20">
<taxonomicName id="1AD84D47FFE589159FBCFD35FE7CFD17" ID-CoL="49KG2" authorityName="Arrow" authorityYear="1931" box="[189,389,644,670]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="excavatus">
<emphasis id="EFACEAD6FFE589159FBCFD35FE7CFD17" box="[189,389,644,670]" italics="true" pageId="19" pageNumber="20">Onitis excavatus</emphasis>
</taxonomicName>
<treatmentCitation id="5C7910D5FFE589159EAAFD32FD6BFD17" author="Balthasar V." box="[427,658,643,670]" page="93" pageId="19" pageNumber="20" year="1935">
<bibRefCitation id="B9494B35FFE589159EAAFD32FD6BFD17" author="Balthasar V." box="[427,658,643,670]" pageId="19" pageNumber="20" refId="ref27158" refString="Balthasar V. 1935. Scarabaeidae des palaarktischen Faunengebietes. Monographische Bestimmungstabelle. I. Coprinae I. Teil. Scarabaeini, Sisyphini, Panelini, Coprini, Onitini, Oniticellini. Edm. Reitter's Nachf. Emmerich Reitter, Troppau." type="book" year="1935">Balthasar 1935: 93</bibRefCitation>
</treatmentCitation>
(monograph); 1963: 37 (monograph). —
<treatmentCitation id="5C7910D5FFE589159B98FD35FA8FFD14" author="Janssens A." box="[1177,1398,643,670]" page="50" pageId="19" pageNumber="20" year="1937">
<bibRefCitation id="B9494B35FFE589159B98FD35FA8FFD14" author="Janssens A." box="[1177,1398,643,670]" pageId="19" pageNumber="20" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 50</bibRefCitation>
</treatmentCitation>
(revision); 1938: 2 (notes). —
<treatmentCitation id="5C7910D5FFE589159D61FD17FC25FD4B" author="Biswas S. &amp; Chatterjee S. K." box="[608,988,678,705]" page="158" pageId="19" pageNumber="20" year="1985">
<bibRefCitation id="B9494B35FFE589159D61FD17FC25FD4B" author="Biswas S. &amp; Chatterjee S. K." box="[608,988,678,705]" pageId="19" pageNumber="20" pagination="1 - 4" refId="ref27511" refString="Biswas S. &amp; Chatterjee S. K. 1985. Insecta: Coleoptera: Scarabaeidae: Coprinae. Records of the Zoological Survey of India, Fauna of Namdapha, Arunachal Pradesh (Special Issue), 82 (1 - 4): 147 - 177. https: // doi. org / 10.26515 / rzsi / v 82 / i 1 - 4 / 1984 / 161291" type="journal article" year="1985">Biswas &amp; Chatterjee 1985: 158</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFE589159BBBFD17FA8FFD4A" author="Mittal I. C." box="[1210,1398,678,705]" page="46" pageId="19" pageNumber="20" year="2005">
<bibRefCitation id="B9494B35FFE589159BBBFD17FA8FFD4A" author="Mittal I. C." box="[1210,1398,678,705]" pageId="19" pageNumber="20" pagination="43 - 51" refId="ref30210" refString="Mittal I. C. 2005. Diversity and conservation status of dung beetles (Laparosticti: Scarabaeidae: Coleoptera) in North India. Bulletin of National Institute of Ecology 15: 43 - 51." type="journal article" year="2005">Mittal 2005: 46</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="5C7910D5FFE589159E61FD7BFD55FD6E" author="Bezdek A. &amp; Krell F. - T." box="[352,684,713,740]" page="159" pageId="19" pageNumber="20" year="2006">
<bibRefCitation id="B9494B35FFE589159E61FD7BFD55FD6E" author="Bezdek A. &amp; Krell F. - T." box="[352,684,713,740]" pageId="19" pageNumber="20" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezdek &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(distribution). — Chandra
<emphasis id="EFACEAD6FFE589159B00FD7BFBB8FD6E" box="[1025,1089,714,740]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2012a: 52 (distribution). —
<treatmentCitation id="5C7910D5FFE589159E0EFD5CFDCBFC8D" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[271,562,749,775]" page="300" pageId="19" pageNumber="20" year="2014">
<bibRefCitation id="B9494B35FFE589159E0EFD5CFDCBFC8D" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[271,562,749,775]" pageId="19" pageNumber="20" pagination="295 - 307" refId="ref31267" refString="Siddiqui H., Ahmed Z. &amp; Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46 (2): 295 - 307." type="journal article" year="2014">
Siddiqui
<emphasis id="EFACEAD6FFE589159E7BFD5FFE4CFC8D" box="[378,437,749,775]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2014: 300
</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFE589159C09FD5CFBE7FC8D" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[776,1054,749,776]" page="1037" pageId="19" pageNumber="20" year="2015">
<bibRefCitation id="B9494B35FFE589159C09FD5CFBE7FC8D" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[776,1054,749,776]" pageId="19" pageNumber="20" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFE589159C56FD5FFC6BFC8D" box="[855,914,749,775]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
2015: 1037
</bibRefCitation>
</treatmentCitation>
(checklist). —
<treatmentCitation id="5C7910D5FFE589159BD5FD5CFEEFFCA0" author="Bezdek A." page="179" pageId="19" pageNumber="20" year="2016">
<bibRefCitation id="B9494B35FFE589159BD5FD5CFEEFFCA0" author="Bezdek A." pageId="19" pageNumber="20" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
</treatmentCitation>
(distribution). —
<bibRefCitation id="B9494B35FFE589159EEDFCA1FD1FFCA0" author="Schoolmeesters P." box="[492,742,784,810]" pageId="19" pageNumber="20" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE589159FBCFCE0FCA7FB8E" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="DD6736C4FFE589159FBCFCE0FE3EFCE1" blockId="19.[189,455,849,875]" box="[189,455,849,875]" pageId="19" pageNumber="20">
<heading id="862F81A8FFE589159FBCFCE0FE3EFCE1" bold="true" box="[189,455,849,875]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="EFACEAD6FFE589159FBCFCE0FE3EFCE1" bold="true" box="[189,455,849,875]" pageId="19" pageNumber="20">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE589159FBCFC31FCA7FB8E" blockId="19.[189,1399,896,1028]" pageId="19" pageNumber="20">
<emphasis id="EFACEAD6FFE589159FBCFC31FE86FC10" box="[189,383,896,922]" italics="true" pageId="19" pageNumber="20">Onitisexcavatus</emphasis>
closely resembles
<taxonomicName id="1AD84D47FFE589159D5DFC31FD24FC10" baseAuthorityName="Wulfen" baseAuthorityYear="1786" box="[604,733,896,922]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="falcatus">
<emphasis id="EFACEAD6FFE589159D5DFC31FD24FC10" box="[604,733,896,922]" italics="true" pageId="19" pageNumber="20">O. falcatus</emphasis>
</taxonomicName>
, both placed in Group I. These two species are similar as the pronotum is smooth or sparingly punctured, and the former differs from the latter in having larger and broader body, a deep transverse excavation behind the middle of the metasternum and presence of strong sharp teeth at the extremity of mesofemur in male.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE589159FBCFB9AFCD5F855" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="DD6736C4FFE589159FBCFB9AFE15FBCF" blockId="19.[189,492,1067,1093]" box="[189,492,1067,1093]" pageId="19" pageNumber="20">
<heading id="862F81A8FFE589159FBCFB9AFE15FBCF" bold="true" box="[189,492,1067,1093]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="EFACEAD6FFE589159FBCFB9AFE15FBCF" bold="true" box="[189,492,1067,1093]" pageId="19" pageNumber="20">Type material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="6DB03C99FFE589159FEBFBEBFB41FB30" collectionCode="BMNH" collectorName="Atkinson" country="Myanmar" determinerName="M. E. Bacchus" location="Tenasserim" municipality="Tenasserim" pageId="19" pageNumber="20" specimenCode="(E) 1237173." specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<paragraph id="DD6736C4FFE589159FEBFBEBFE9FFBFE" blockId="19.[189,1398,1114,1211]" box="[234,358,1114,1140]" pageId="19" pageNumber="20">
<emphasis id="EFACEAD6FFE589159FEBFBEBFE9FFBFE" bold="true" box="[234,358,1114,1140]" pageId="19" pageNumber="20">
<typeStatus id="02638866FFE589159FEBFBEBFE9FFBFE" box="[234,358,1114,1140]" pageId="19" pageNumber="20" type="lectotype">Lectotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE589159FBCFBCCFB41FB30" blockId="19.[189,1398,1114,1211]" pageId="19" pageNumber="20">
<collectingCountry id="A5CF7654FFE589159FBCFBCCFEAFFB1D" box="[189,342,1149,1175]" name="Myanmar" pageId="19" pageNumber="20">MYANMAR</collectingCountry>
<specimenCount id="CBDEFD4DFFE589159E7AFBCCFE6DFB1D" box="[379,404,1149,1175]" count="1" pageId="19" pageNumber="20" type="male"></specimenCount>
; “
<collectingMunicipality id="3D03ACBEFFE589159EB1FBCCFDC2FB1D" box="[432,571,1149,1175]" pageId="19" pageNumber="20">Tenasserim</collectingMunicipality>
,
<collectorName id="702D5312FFE589159D4AFBCCFD4EFB1D" box="[587,695,1149,1175]" pageId="19" pageNumber="20">Atkinson</collectorName>
Coll.923, near
<taxonomicName id="1AD84D47FFE589159C8DFBCCFB77FB1D" authority=", Lansb." authorityName="Lansb." box="[908,1166,1149,1175]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ludekingi">
<emphasis id="EFACEAD6FFE589159C8DFBCCFBC8FB1D" box="[908,1073,1149,1175]" italics="true" pageId="19" pageNumber="20">O. Ludekingi,</emphasis>
Lansb.
</taxonomicName>
, but quite diff. by sculptures,
<taxonomicName id="1AD84D47FFE589159E42FB11FDFDFB30" authorityName="Arrow" authorityYear="1931" box="[323,516,1184,1210]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="excavates">
<emphasis id="EFACEAD6FFE589159E42FB11FDFDFB30" box="[323,516,1184,1210]" italics="true" pageId="19" pageNumber="20">Onitis excavates</emphasis>
</taxonomicName>
male Arr.,
<determinerName id="28BBDA00FFE589159D86FB10FCD5FB30" box="[647,812,1184,1211]" pageId="19" pageNumber="20">M.E. Bacchus</determinerName>
det. 1975”;
<collectionCode id="BBC9AE01FFE589159CBCFB10FBECFB31" box="[957,1045,1185,1211]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="19" pageNumber="20">BMNH</collectionCode>
<specimenCode id="8D7E9EBFFFE589159B1CFB11FB41FB30" box="[1053,1208,1184,1210]" pageId="19" pageNumber="20">(E) 1237173.</specimenCode>
</paragraph>
</materialsCitation>
<paragraph id="DD6736C4FFE589159FEBFB50FCE2FB71" blockId="19.[189,1399,1249,2015]" box="[234,795,1249,1276]" pageId="19" pageNumber="20">
<emphasis id="EFACEAD6FFE589159FEBFB50FD9BFB71" bold="true" box="[234,610,1249,1275]" pageId="19" pageNumber="20">Additional material examined</emphasis>
(
<specimenCount id="CBDEFD4DFFE589159D70FB53FCEAFB71" box="[625,787,1249,1276]" count="29" pageId="19" pageNumber="20" type="generic">29 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFE589159FBCFAB4FCD5F855" blockId="19.[189,1399,1249,2015]" pageId="19" pageNumber="20">
<materialsCitation id="6DB03C99FFE589159FBCFAB4FE07FAEF" collectingDate="2018-09-24" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan" country="India" elevation="877" latitude="27.238258" location="Pakke" longLatPrecision="1" longitude="93.5373" pageId="19" pageNumber="20" specimenCode="Co/Sc1000038, Co/Sc1000039, Co/Sc1000040, Co/Sc1000041, Co/Sc1000042, Co/Sc1000043, Co/Sc1000044" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Arunachal Pradesh">
<collectingCountry id="A5CF7654FFE589159FBCFAB4FEF7FA95" box="[189,270,1285,1311]" name="India" pageId="19" pageNumber="20">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFE589159E2BFAB4FE96FA95" box="[298,367,1285,1311]" count="2" pageId="19" pageNumber="20" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE589159E7CFAB5FE38FA95" box="[381,449,1284,1311]" count="5" pageId="19" pageNumber="20" type="female">5 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE589159ECEFAB4FD4BFA95" box="[463,690,1285,1311]" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<location id="D807601FFFE589159DBFFAB4FCF3FA95" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159DBFFAB4FCF3FA95" box="[702,778,1285,1311]" country="India" latitude="27.238258" longLatPrecision="1" longitude="93.5373" name="Pakke" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Pakke</location>
;
<geoCoordinate id="B8EC5003FFE589159C16FAB2FC2BFA95" box="[791,978,1283,1311]" degrees="27" direction="north" minutes="14" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="17.73" value="27.238258">27°14ʹ17.73″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159CDEFAB2FB61FA95" box="[991,1176,1283,1311]" degrees="93" direction="east" minutes="32" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="14.28" value="93.5373">93°32ʹ14.28″ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159BE5FAB4FA89FA95" box="[1252,1392,1285,1311]" metricMagnitude="2" metricUnit="m" metricValue="8.77" pageId="19" pageNumber="20" unit="m" value="877.0">
<quantity id="1A209B21FFE589159BE5FAB4FAD6FA95" box="[1252,1327,1285,1311]" metricMagnitude="2" metricUnit="m" metricValue="8.77" pageId="19" pageNumber="20" unit="m" value="877.0">877 m</quantity>
a.s.l.
</elevation>
; hand-picked from elephant dung;
<date id="A9661004FFE589159D54FA99FD01FAC8" box="[597,760,1320,1346]" pageId="19" pageNumber="20" value="2018-09-24">
<collectingDate id="B922E9ECFFE589159D54FA99FD01FAC8" box="[597,760,1320,1346]" pageId="19" pageNumber="20" value="2018-09-24">24 Sep. 2018</collectingDate>
</date>
;
<collectorName id="702D5312FFE589159C06FA99FB9DFAC8" box="[775,1124,1320,1346]" pageId="19" pageNumber="20">Priyadarsanan Dharma Rajan</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159A23FA99FA97FAC8" box="[1314,1390,1320,1346]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFE589159FBCFAFAFE07FAEF" box="[189,510,1355,1382]" pageId="19" pageNumber="20">Co/Sc1000037 to 1000044</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159D1DFAFDFDF2FA21" collectingDate="2018-10-03" collectingMethod="hand-picked" collectionCode="AIMB" collectorName="Harsha Malhotra" country="India" elevation="956" latitude="28.362144" location="Sibuk" longLatPrecision="1" longitude="95.265884" pageId="19" pageNumber="20" specimenCode="Co/Sc1000045, Co/Sc1000046" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159D1DFAFDFDCDFAEC" box="[540,564,1356,1382]" count="1" pageId="19" pageNumber="20" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFE589159D42FAFDFDA3FAEC" box="[579,602,1356,1382]" count="1" pageId="19" pageNumber="20" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159D6BFAFAFCB7FAEF" box="[618,846,1355,1381]" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<location id="D807601FFFE589159C5CFAFAFC5CFAEF" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159C5CFAFAFC5CFAEF" box="[861,933,1355,1381]" country="India" latitude="28.362144" longLatPrecision="1" longitude="95.265884" name="Sibuk" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Sibuk</location>
;
<geoCoordinate id="B8EC5003FFE589159CB5FAF8FB74FAEC" box="[948,1165,1353,1382]" degrees="28" direction="north" minutes="21" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="43.7184" value="28.362144">28°21ʹ43.7184ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159B9AFAF8FA88FAEC" box="[1179,1393,1353,1382]" degrees="95" direction="east" minutes="15" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="57.1968" value="95.265884">95°15ʹ57.1968ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159FFCFADFFE72FA02" box="[253,395,1390,1417]" metricMagnitude="2" metricUnit="m" metricValue="9.559999999999999" pageId="19" pageNumber="20" unit="m" value="956.0">
<quantity id="1A209B21FFE589159FFCFADFFEB0FA03" box="[253,329,1390,1417]" metricMagnitude="2" metricUnit="m" metricValue="9.559999999999999" pageId="19" pageNumber="20" unit="m" value="956.0">956 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE589159E9AFADFFDD5FA02" box="[411,556,1390,1416]" pageId="19" pageNumber="20" value="2018-10-03">
<collectingDate id="B922E9ECFFE589159E9AFADFFDD5FA02" box="[411,556,1390,1416]" pageId="19" pageNumber="20" value="2018-10-03">3 Oct. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE589159D3AFADFFD35FA02" box="[571,716,1390,1416]" pageId="19" pageNumber="20">hand-picked</collectingMethod>
from cow dung;
<collectorName id="702D5312FFE589159C9FFADFFB9EFA02" box="[926,1127,1390,1416]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159A23FADFFA97FA02" box="[1314,1390,1390,1416]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFE589159FBCFA20FDF2FA21" box="[189,523,1425,1452]" pageId="19" pageNumber="20">Co/Sc1000045 to 1000046</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159D36FA23FCB5FA78" collectingDate="2018-10-09" collectingMethod="hand-picked" collectionCode="AIMB" collectorName="Harsha Malhotra" country="India" elevation="1557" latitude="27.524204" location="Rani Village" longLatPrecision="1" longitude="27.524204" municipality="Pasighat" pageId="19" pageNumber="20" specimenCode="Co/Sc1000047" specimenCount="1" specimenCount-female="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159D36FA23FDB7FA26" box="[567,590,1426,1452]" count="1" pageId="19" pageNumber="20" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159D65FA20FCB6FA21" box="[612,847,1425,1451]" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<locationDeviation id="CCD52FE6FFE589159C64FA20FBB8FA21" box="[869,1089,1425,1451]" pageId="19" pageNumber="20">community forest</locationDeviation>
,
<collectingMunicipality id="3D03ACBEFFE589159B57FA20FB45FA21" box="[1110,1212,1425,1451]" pageId="19" pageNumber="20">Pasighat</collectingMunicipality>
,
<location id="D807601FFFE589159BD3FA20FA8BFA21" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159BD3FA20FA8BFA21" box="[1234,1394,1425,1451]" country="India" latitude="27.524204" longLatPrecision="1" longitude="27.524204" municipality="Pasighat" name="Rani Village" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFE589159FBCFA02FE7EFA45" box="[189,391,1459,1487]" degrees="27" direction="north" minutes="31" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="27.138" value="27.524204">27°31ʹ27.138ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159E94FA02FDA2FA45" box="[405,603,1459,1487]" degrees="27" direction="east" minutes="31" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="27.138" value="27.524204">27°31ʹ27.138ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159DABFA05FCBEFA45" box="[682,839,1460,1487]" metricMagnitude="3" metricUnit="m" metricValue="1.557" pageId="19" pageNumber="20" unit="m" value="1557.0">
<quantity id="1A209B21FFE589159DABFA05FCFCFA45" box="[682,773,1460,1487]" metricMagnitude="3" metricUnit="m" metricValue="1.557" pageId="19" pageNumber="20" unit="m" value="1557.0">1557 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE589159C56FA04FC11FA45" box="[855,1000,1461,1487]" pageId="19" pageNumber="20" value="2018-10-09">
<collectingDate id="B922E9ECFFE589159C56FA04FC11FA45" box="[855,1000,1461,1487]" pageId="19" pageNumber="20" value="2018-10-09">9 Oct. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE589159CF6FA04FB71FA45" box="[1015,1160,1461,1487]" pageId="19" pageNumber="20">hand-picked</collectingMethod>
from donkey dung;
<collectorName id="702D5312FFE589159FBCFA69FE7EFA78" box="[189,391,1496,1522]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159D4BFA69FD6EFA78" box="[586,663,1496,1522]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFE589159D9EFA69FCB5FA78" box="[671,844,1496,1522]" pageId="19" pageNumber="20">Co/Sc1000047</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159C6FFA69FC6BF9B2" collectingDate="2018-10-10" collectingMethod="hand-picked" collectionCode="AIMB" collectorName="Harsha Malhotra" country="India" elevation="1532" latitude="27.523193" location="Talley Valley" longLatPrecision="1" longitude="93.839455" municipality="Ziro" pageId="19" pageNumber="20" specimenCode="Co/Sc1000048" specimenCount="1" specimenCount-female="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159C6FFA69FC7CFA78" box="[878,901,1496,1522]" count="1" pageId="19" pageNumber="20" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159C97FA69FB82FA78" box="[918,1147,1496,1522]" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE589159B8AFA69FB39FA78" box="[1163,1216,1496,1522]" pageId="19" pageNumber="20">Ziro</collectingMunicipality>
,
<location id="D807601FFFE589159BD1FA69FA8BFA78" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159BD1FA69FA8BFA78" box="[1232,1394,1496,1522]" country="India" latitude="27.523193" longLatPrecision="1" longitude="93.839455" municipality="Ziro" name="Talley Valley" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Talley Valley</location>
;
<geoCoordinate id="B8EC5003FFE589159FBCFA48FE62F99C" box="[189,411,1529,1558]" degrees="27" direction="north" minutes="31" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="23.4984" value="27.523193">27°31ʹ23.4984ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159EAFFA48FD70F99C" box="[430,649,1529,1558]" degrees="93" direction="east" minutes="50" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="22.0308" value="93.839455">93°50ʹ22.0308ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159DE3FA4AFC73F99F" box="[738,906,1531,1558]" metricMagnitude="3" metricUnit="m" metricValue="1.5319999999999998" pageId="19" pageNumber="20" unit="m" value="1532.0">
<quantity id="1A209B21FFE589159DE3FA4AFCBAF99C" box="[738,835,1531,1558]" metricMagnitude="3" metricUnit="m" metricValue="1.5319999999999998" pageId="19" pageNumber="20" unit="m" value="1532.0">1532 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE589159C9EFA4AFBB3F99F" box="[927,1098,1531,1557]" pageId="19" pageNumber="20" value="2018-10-10">
<collectingDate id="B922E9ECFFE589159C9EFA4AFBB3F99F" box="[927,1098,1531,1557]" pageId="19" pageNumber="20" value="2018-10-10">10 Oct. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE589159B5FFA4AFB16F99F" box="[1118,1263,1531,1557]" pageId="19" pageNumber="20">hand-picked</collectingMethod>
from cow dung;
<collectorName id="702D5312FFE589159E0BF9AFFE2AF9B2" box="[266,467,1566,1592]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159D91F9AFFD24F9B2" box="[656,733,1566,1592]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFE589159DE7F9AFFC6BF9B2" box="[742,914,1566,1592]" pageId="19" pageNumber="20">Co/Sc1000048</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159CB0F9AEFCB1F9F5" collectingDate="2018-10-18" collectingMethod="hand-picked" collectionCode="AIMB" collectorName="Harsha Malhotra" country="India" elevation="805" latitude="27.233759" location="Mobu" longLatPrecision="1" longitude="93.53302" municipality="Sangri" pageId="19" pageNumber="20" specimenCode="Co/Sc1000049" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159CB0F9AEFC33F9B3" box="[945,970,1567,1593]" count="1" pageId="19" pageNumber="20" type="male"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159CD9F9AFFB45F9B2" box="[984,1212,1566,1592]" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE589159BCBF9AFFAE0F9B2" box="[1226,1305,1566,1592]" pageId="19" pageNumber="20">Sangri</collectingMunicipality>
,
<location id="D807601FFFE589159A29F9AFFA8BF9B2" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159A29F9AFFA8BF9B2" box="[1320,1394,1566,1592]" country="India" latitude="27.233759" longLatPrecision="1" longitude="93.53302" municipality="Sangri" name="Mobu" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Mobu</location>
;
<geoCoordinate id="B8EC5003FFE589159FBCF9F1FE7FF9D6" box="[189,390,1600,1628]" degrees="27" direction="north" minutes="14" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="1.5324" value="27.233759">27°14ʹ1.5324ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159E92F9F1FD91F9D6" box="[403,616,1600,1628]" degrees="93" direction="east" minutes="31" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="58.8576" value="93.53302">93°31ʹ58.8576ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159DB5F9F0FCB8F9D1" box="[692,833,1601,1628]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="19" pageNumber="20" unit="m" value="805.0">
<quantity id="1A209B21FFE589159DB5F9F0FCF9F9D6" box="[692,768,1601,1628]" metricMagnitude="2" metricUnit="m" metricValue="8.05" pageId="19" pageNumber="20" unit="m" value="805.0">805 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE589159C51F9F3FC16F9D6" box="[848,1007,1602,1628]" pageId="19" pageNumber="20" value="2018-10-18">
<collectingDate id="B922E9ECFFE589159C51F9F3FC16F9D6" box="[848,1007,1602,1628]" pageId="19" pageNumber="20" value="2018-10-18">18 Oct. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE589159CFDF9F0FB74F9D1" box="[1020,1165,1601,1627]" pageId="19" pageNumber="20">hand-picked</collectingMethod>
from mithun dung;
<collectorName id="702D5312FFE589159FBCF9D4FE7FF9F5" box="[189,390,1637,1663]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159D45F9D4FD68F9F5" box="[580,657,1637,1663]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFE589159D9BF9D4FCB1F9F5" box="[666,840,1637,1663]" pageId="19" pageNumber="20">Co/Sc1000049</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159C67F9D4FC1BF94C" collectingDate="2018-09-23" collectingMethod="hand-picked" collectionCode="AIMB" collectorName="Harsha Malhotra" country="India" elevation="164" latitude="26.94896" location="Pakke Tiger Reserve" longLatPrecision="1" longitude="92.98454" municipality="Seijosa" pageId="19" pageNumber="20" specimenCode="Co/Sc1000050" specimenCount="1" specimenCount-female="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159C67F9D4FC84F9F5" box="[870,893,1637,1663]" count="1" pageId="19" pageNumber="20" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159C8CF9D4FB88F9F5" box="[909,1137,1637,1663]" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE589159B81F9D4FB21F9F5" box="[1152,1240,1637,1663]" pageId="19" pageNumber="20">Seijosa</collectingMunicipality>
,
<location id="D807601FFFE589159BE7F9D4FEE6F928" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159BE7F9D4FEE6F928" country="India" latitude="26.94896" longLatPrecision="1" longitude="92.98454" municipality="Seijosa" name="Pakke Tiger Reserve" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Pakke Tiger Reserve</location>
;
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,
<geoCoordinate id="B8EC5003FFE589159D1FF937FD11F928" box="[542,744,1670,1698]" degrees="92" direction="east" minutes="59" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="4.3764" value="92.98454">92°59ʹ4.3764ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159C3CF939FC2BF928" box="[829,978,1672,1698]" metricMagnitude="2" metricUnit="m" metricValue="1.64" pageId="19" pageNumber="20" unit="m" value="164.0">
<quantity id="1A209B21FFE589159C3CF939FC74F928" box="[829,909,1672,1698]" metricMagnitude="2" metricUnit="m" metricValue="1.64" pageId="19" pageNumber="20" unit="m" value="164.0">164 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE589159CE4F939FB77F928" box="[997,1166,1672,1698]" pageId="19" pageNumber="20" value="2018-09-23">
<collectingDate id="B922E9ECFFE589159CE4F939FB77F928" box="[997,1166,1672,1698]" pageId="19" pageNumber="20" value="2018-09-23">23 Sep. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE589159B9EF939FAC9F928" box="[1183,1328,1672,1698]" pageId="19" pageNumber="20">hand-picked</collectingMethod>
from elephant dung;
<collectorName id="702D5312FFE589159E6EF91AFDCAF94F" box="[367,563,1707,1733]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159DDEF91AFCD5F94C" box="[735,812,1707,1734]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFE589159C34F91AFC1BF94C" box="[821,994,1707,1734]" pageId="19" pageNumber="20">Co/Sc1000050</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159CF9F91AFB47F881" collectingDate="2018-09-24" collectingMethod="hand-picked" collectionCode="AIMB" collectorName="Harsha Malhotra" country="India" elevation="190" latitude="26.941978" location="Pakke Tiger Reserve" longLatPrecision="1" longitude="92.9716" municipality="Seijosa" pageId="19" pageNumber="20" specimenCode="Co/Sc1000051, Co/Sc1000052, Co/Sc1000053, Co/Sc1000054, Co/Sc1000055, Co/Sc1000056, Co/Sc1000057, Co/Sc1000058, Co/Sc1000059, Co/Sc1000060" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159CF9F91AFBC3F94C" box="[1016,1082,1707,1734]" count="6" pageId="19" pageNumber="20" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFE589159B44F91AFB7EF94C" box="[1093,1159,1707,1734]" count="4" pageId="19" pageNumber="20" type="female">4 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFE589159B93F91AFA8BF94F" box="[1170,1394,1707,1733]" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE589159FBCF97FFEECF962" box="[189,277,1742,1768]" pageId="19" pageNumber="20">Seijosa</collectingMunicipality>
,
<location id="D807601FFFE589159E1EF97FFDEBF963" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159E1EF97FFDEBF963" box="[287,530,1742,1769]" country="India" latitude="26.941978" longLatPrecision="1" longitude="92.9716" municipality="Seijosa" name="Pakke Tiger Reserve" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Pakke Tiger Reserve</location>
;
<geoCoordinate id="B8EC5003FFE589159D1CF97CFD1AF963" box="[541,739,1741,1769]" degrees="26" direction="north" minutes="56" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="31.128" value="26.941978">26°56ʹ31.128ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159DEFF97CFC5BF963" box="[750,930,1741,1769]" degrees="92" direction="east" minutes="58" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="17.76" value="92.9716">92°58ʹ17.76ʺ E</geoCoordinate>
; elev.
<quantity id="1A209B21FFE589159CE8F97FFBCCF963" box="[1001,1077,1742,1769]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="19" pageNumber="20" unit="m" value="190.0">
<elevation id="56F5D1F7FFE589159CE8F97FFBCCF963" box="[1001,1077,1742,1769]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="19" pageNumber="20" unit="m" value="190.0">190 m</elevation>
</quantity>
;
<date id="A9661004FFE589159B41F97FFB22F962" box="[1088,1243,1742,1768]" pageId="19" pageNumber="20" value="2018-09-24">
<collectingDate id="B922E9ECFFE589159B41F97FFB22F962" box="[1088,1243,1742,1768]" pageId="19" pageNumber="20" value="2018-09-24">24 Sep. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE589159BE4F97FFA8FF962" box="[1253,1398,1742,1768]" pageId="19" pageNumber="20">hand-picked</collectingMethod>
from elephant dung;
<collectorName id="702D5312FFE589159EB2F940FD83F881" box="[435,634,1777,1803]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159C2FF940FC82F886" box="[814,891,1777,1804]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFE589159C85F940FB47F881" box="[900,1214,1777,1804]" pageId="19" pageNumber="20">Co/Sc1000051 to 1000060</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159BD9F943FB61F8D8" collectingDate="2018-09-29" collectingMethod="hand-picked" collectionCode="AIMB" collectorName="Harsha Malhotra" country="India" elevation="135" latitude="27.968342" location="Rani village" longLatPrecision="1" longitude="95.320145" municipality="Pasighat" pageId="19" pageNumber="20" specimenCode="Co/Sc1000061" specimenCount="1" specimenCount-male="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159BD9F943FB08F886" box="[1240,1265,1778,1804]" count="1" pageId="19" pageNumber="20" type="male"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159BFDF940FEE4F8A5" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE589159E2BF8A4FE69F8A5" box="[298,400,1813,1839]" pageId="19" pageNumber="20">Pasighat</collectingMunicipality>
,
<location id="D807601FFFE589159E9CF8A4FDC9F8A5" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159E9CF8A4FDC9F8A5" box="[413,560,1813,1839]" country="India" latitude="27.968342" longLatPrecision="1" longitude="95.320145" municipality="Pasighat" name="Rani village" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Rani village</location>
;
<geoCoordinate id="B8EC5003FFE589159D3FF8A2FCFEF8A5" box="[574,775,1811,1839]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="6.0276" value="27.968342">27°58ʹ6.0276ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159C15F8A2FC10F8A5" box="[788,1001,1811,1839]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="12.5328" value="95.320145">95°19ʹ12.5328ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159B37F8A5FB3AF8A5" box="[1078,1219,1812,1839]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="19" pageNumber="20" unit="m" value="135.0">
<quantity id="1A209B21FFE589159B37F8A5FB78F8A5" box="[1078,1153,1812,1839]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="19" pageNumber="20" unit="m" value="135.0">135 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE589159BD0F8A4FA88F8A5" box="[1233,1393,1813,1839]" pageId="19" pageNumber="20" value="2018-09-29">
<collectingDate id="B922E9ECFFE589159BD0F8A4FA88F8A5" box="[1233,1393,1813,1839]" pageId="19" pageNumber="20" value="2018-09-29">29 Sep. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE589159FBCF889FEB7F8D8" box="[189,334,1848,1874]" pageId="19" pageNumber="20">hand-picked</collectingMethod>
from cow dung;
<collectorName id="702D5312FFE589159D1BF889FD18F8D8" box="[538,737,1848,1874]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159C94F889FC1BF8D8" box="[917,994,1848,1874]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFE589159CEDF889FB61F8D8" box="[1004,1176,1848,1874]" pageId="19" pageNumber="20">Co/Sc1000061</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159BB3F889FB89F812" collectingDate="2018-09-30" collectingMethod="cow dung" collectionCode="AIMB" collectorName="Harsha Malhotra" country="India" elevation="125" latitude="27.970259" location="Rani village" longLatPrecision="1" longitude="95.322365" municipality="Pasighat" pageId="19" pageNumber="20" specimenCode="Co/Sc1000061, Co/Sc1000062" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFE589159BB3F889FB33F8D8" box="[1202,1226,1848,1874]" count="1" pageId="19" pageNumber="20" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFE589159BD9F889FB16F8D8" box="[1240,1263,1848,1874]" count="1" pageId="19" pageNumber="20" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159BFDF889FEE4F8FF" country="India" name="Arunachal Pradesh" pageId="19" pageNumber="20">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE589159E28F8EAFE76F8FF" box="[297,399,1883,1909]" pageId="19" pageNumber="20">Pasighat</collectingMunicipality>
,
<location id="D807601FFFE589159E9AF8EAFDD4F8FF" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159E9AF8EAFDD4F8FF" box="[411,557,1883,1909]" country="India" latitude="27.970259" longLatPrecision="1" longitude="95.322365" municipality="Pasighat" name="Rani village" pageId="19" pageNumber="20" stateProvince="Arunachal Pradesh">Rani village</location>
;
<geoCoordinate id="B8EC5003FFE589159D3BF8E8FCE9F8FC" box="[570,784,1881,1910]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="19" pageNumber="20" precision="1" seconds="12.9324" value="27.970259">27°58ʹ12.9324ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFE589159C1CF8E8FC09F8FC" box="[797,1008,1881,1910]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="19" pageNumber="20" precision="1" seconds="20.5284" value="95.322365">95°19ʹ20.5284ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFE589159B3AF8EAFB3FF8FF" box="[1083,1222,1883,1910]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="19" pageNumber="20" unit="m" value="125.0">
<quantity id="1A209B21FFE589159B3AF8EAFB7FF8FC" box="[1083,1158,1883,1910]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="19" pageNumber="20" unit="m" value="125.0">125 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFE589159BD2F8EAFA8BF8FF" box="[1235,1394,1883,1909]" pageId="19" pageNumber="20" value="2018-09-30">
<collectingDate id="B922E9ECFFE589159BD2F8EAFA8BF8FF" box="[1235,1394,1883,1909]" pageId="19" pageNumber="20" value="2018-09-30">30 Sep. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE589159FBCF8CEFEC1F812" box="[189,312,1918,1945]" pageId="19" pageNumber="20">cow dung</collectingMethod>
;
<collectorName id="702D5312FFE589159E49F8CFFDEAF812" box="[328,531,1918,1944]" pageId="19" pageNumber="20">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159DD6F8CFFCDDF812" box="[727,804,1918,1944]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFE589159C2FF8CFFB89F812" box="[814,1136,1918,1945]" pageId="19" pageNumber="20">Co/Sc1000061 to 1000062</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFE589159B90F8CEFCD1F855" collectingDate="2021-04-25" collectingMethod="hand-picked" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara" country="India" location="Singphan Elephant Reserve" municipality="Mon District" pageId="19" pageNumber="20" specimenCode="Co/Sc1000063" specimenCount="1" specimenCount-female="1" stateProvince="Nagaland">
<specimenCount id="CBDEFD4DFFE589159B90F8CEFB51F813" box="[1169,1192,1919,1945]" count="1" pageId="19" pageNumber="20" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFE589159BBAF8CFFAD7F812" box="[1211,1326,1918,1944]" country="India" name="Nagaland" pageId="19" pageNumber="20">Nagaland</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFE589159A3EF8CEFEE1F831" pageId="19" pageNumber="20">Mon District</collectingMunicipality>
,
<location id="D807601FFFE589159E29F810FD8EF836" LSID="urn:lsid:plazi:treatment:557187D2FFE589129D69FE6CFCB4FB1F:D807601FFFE589159E29F810FD8EF836" box="[296,631,1953,1980]" country="India" municipality="Mon District" name="Singphan Elephant Reserve" pageId="19" pageNumber="20" stateProvince="Nagaland">Singphan Elephant Reserve</location>
;
<date id="A9661004FFE589159D86F810FCD2F836" box="[647,811,1953,1980]" pageId="19" pageNumber="20" value="2021-04-25">
<collectingDate id="B922E9ECFFE589159D86F810FCD2F836" box="[647,811,1953,1980]" pageId="19" pageNumber="20" value="2021-04-25">25 Apr. 2021</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFE589159C3AF810FC35F831" box="[827,972,1953,1979]" pageId="19" pageNumber="20">hand-picked</collectingMethod>
from cow dung;
<collectorName id="702D5312FFE589159BA7F813FE96F855" pageId="19" pageNumber="20">Seena Narayanan Karimbumkara</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFE589159D20F874FD96F855" box="[545,623,1989,2015]" pageId="19" pageNumber="20">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFE589159D79F874FCD1F855" box="[632,808,1989,2015]" pageId="19" pageNumber="20">Co/Sc1000063</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE289129FBCFEB9FC66FC36" pageId="20" pageNumber="21" type="description">
<paragraph id="DD6736C4FFE289129FBCFEB9FEB7FEA8" blockId="20.[189,334,264,290]" box="[189,334,264,290]" pageId="20" pageNumber="21">
<heading id="862F81A8FFE289129FBCFEB9FEB7FEA8" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="EFACEAD6FFE289129FBCFEB9FEB7FEA8" bold="true" box="[189,334,264,290]" pageId="20" pageNumber="21">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE289129FBCFE86FD11FEDB" blockId="20.[189,744,311,338]" box="[189,744,311,338]" pageId="20" pageNumber="21">MEASUREMENTS (in mm). TL=1925; BW= 13.</paragraph>
<paragraph id="DD6736C4FFE289129FEBFEC8FED3FE19" blockId="20.[189,1399,377,720]" box="[234,298,377,403]" pageId="20" pageNumber="21">
<emphasis id="EFACEAD6FFE289129FEBFEC8FED3FE19" bold="true" box="[234,298,377,403]" pageId="20" pageNumber="21">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE289129FBCFE2DFB4EFD45" blockId="20.[189,1399,377,720]" pageId="20" pageNumber="21">Black, shining, elytra and pygidium subopaque; clypeus short, elliptical, moderately closely and finely rugulose; frontal carina widely interrupted in the middle, a short transverse clypeal carina anteriorly, a small tubercle behind, vertex irregularly granular; pronotum very smooth with irregular minute punctures anteriorly, longitudinal groove absent; elytra finely striate with 1st, 3rd and 5th intervals elevated, narrower, rest of the intervals flat; pygidium smooth, unpunctured; metasternum with very deep transverse excavation in the middle, metasternal shield thickly hairy anteriorly, smooth, shining behind; protibia elongate with a serrate carina upon anterior half beneath, four short external teeth and a sharp incurved terminal process; mesofemur with a rounded lobe in the middle of hind edge, two sharp teeth near extremity above and one beneath; trochanters of hind legs acutely produced.</paragraph>
<paragraph id="DD6736C4FFE289129FBCFD46FAE3FCBE" blockId="20.[189,1399,759,820]" pageId="20" pageNumber="21">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFE289129EA8FD46FDD0FC9B" box="[425,553,759,785]" captionStart="Fig" captionStartId="18.[190,234,1893,1919]" captionTargetBox="[295,1246,340,1832]" captionTargetId="figure-1@18.[189,1398,311,1837]" captionTargetPageId="18" captionText="Fig. 6. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitisexcavatus Arrow, 1931, AIMB/Co/Sc1000037. EH. O. falcatus (Wulfen, 1786), AIMB/Co/Sc1000070. IL. O. bhomorensis sp. nov., NIM/NBAIR/COL/ONIT/H-271023A. MP. O. punctatostriatus Janssens, 1937, AIMB/ Co/Sc1000492. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742867" httpUri="https://zenodo.org/record/13742867/files/figure.png" pageId="20" pageNumber="21">Fig. 6AD</figureCitation>
). LP=3.03, Lp=2.93; BP=1.5, BpB=1.14, BpT=0.46. Parameres shorter than phallobase, straight above and slightly curved outwards below and hooked slightly at the tip.
</paragraph>
<paragraph id="DD6736C4FFE289129FEBFCEAFEBCFCFF" blockId="20.[189,1399,859,956]" box="[234,325,859,885]" pageId="20" pageNumber="21">
<emphasis id="EFACEAD6FFE289129FEBFCEAFEBCFCFF" bold="true" box="[234,325,859,885]" pageId="20" pageNumber="21">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFE289129FBCFCCEFC66FC36" blockId="20.[189,1399,859,956]" pageId="20" pageNumber="21">Legs simple, frontal carina slightly diagonal; punctation of the pronotum heavier and denser, elytral striae deeper than in male; metasternum without transverse pit.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFE289129FBCFC52FCB4FB1F" pageId="20" pageNumber="21" type="distribution">
<paragraph id="DD6736C4FFE289129FBCFC52FEAFFC77" blockId="20.[189,342,995,1021]" box="[189,342,995,1021]" pageId="20" pageNumber="21">
<heading id="862F81A8FFE289129FBCFC52FEAFFC77" bold="true" box="[189,342,995,1021]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="EFACEAD6FFE289129FBCFC52FEAFFC77" bold="true" box="[189,342,995,1021]" pageId="20" pageNumber="21">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFE289129FBCFBA3FCB4FB1F" blockId="20.[189,1399,1042,1174]" pageId="20" pageNumber="21">
<collectingCountry id="A5CF7654FFE289129FBCFBA3FF07FBA6" box="[189,254,1042,1068]" name="India" pageId="20" pageNumber="21">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFE289129E04FBA3FE1DFBA6" box="[261,484,1042,1068]" country="India" name="Arunachal Pradesh" pageId="20" pageNumber="21">Arunachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFE289129EEDFBA3FDC6FBA6" box="[492,575,1042,1068]" country="India" name="Assam" pageId="20" pageNumber="21">Assam</collectingRegion>
,
<collectingRegion id="1F1CF826FFE289129D49FBA3FCE6FBA6" box="[584,799,1042,1068]" country="India" name="Himachal Pradesh" pageId="20" pageNumber="21">Himachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFE289129C29FBA3FC54FBA6" box="[808,941,1042,1068]" country="India" name="Meghalaya" pageId="20" pageNumber="21">Meghalaya</collectingRegion>
,
<collectingRegion id="1F1CF826FFE289129CB6FBA3FBD3FBA6" box="[951,1066,1042,1068]" country="India" name="Nagaland" pageId="20" pageNumber="21">Nagaland</collectingRegion>
,
<collectingRegion id="1F1CF826FFE289129B35FBA3FB31FBA6" box="[1076,1224,1042,1068]" country="India" name="Uttarakhand" pageId="20" pageNumber="21">Uttarakhand</collectingRegion>
,
<collectingRegion id="1F1CF826FFE289129BD0FBA3FA8AFBA6" box="[1233,1395,1042,1068]" country="India" name="Uttar Pradesh" pageId="20" pageNumber="21">Uttar Pradesh</collectingRegion>
. Other countries:
<collectingCountry id="A5CF7654FFE289129E84FB84FE36FBC5" box="[389,463,1077,1103]" name="China" pageId="20" pageNumber="21">China</collectingCountry>
;
<collectingCountry id="A5CF7654FFE289129EDCFB84FDAAFBC5" box="[477,595,1077,1103]" name="Myanmar" pageId="20" pageNumber="21">Myanmar</collectingCountry>
;
<collectingCountry id="A5CF7654FFE289129D61FB84FD53FBC5" box="[608,682,1077,1103]" name="Nepal" pageId="20" pageNumber="21">Nepal</collectingCountry>
;
<collectingCountry id="A5CF7654FFE289129DB6FB84FCE7FBC5" box="[695,798,1077,1103]" name="Pakistan" pageId="20" pageNumber="21">Pakistan</collectingCountry>
;
<collectingCountry id="A5CF7654FFE289129C2AFB84FC7CFBC5" box="[811,901,1077,1103]" name="Taiwan" pageId="20" pageNumber="21">Taiwan</collectingCountry>
;
<collectingCountry id="A5CF7654FFE289129C93FB84FC07FBC5" box="[914,1022,1077,1103]" name="Thailand" pageId="20" pageNumber="21">Thailand</collectingCountry>
;
<collectingCountry id="A5CF7654FFE289129B0BFB84FB94FBC5" box="[1034,1133,1077,1103]" name="Vietnam" pageId="20" pageNumber="21">Vietnam</collectingCountry>
(
<bibRefCitation id="B9494B35FFE289129B7EFB84FF02FBF9" author="Biswas S. &amp; Chatterjee S. K." pageId="20" pageNumber="21" pagination="1 - 4" refId="ref27511" refString="Biswas S. &amp; Chatterjee S. K. 1985. Insecta: Coleoptera: Scarabaeidae: Coprinae. Records of the Zoological Survey of India, Fauna of Namdapha, Arunachal Pradesh (Special Issue), 82 (1 - 4): 147 - 177. https: // doi. org / 10.26515 / rzsi / v 82 / i 1 - 4 / 1984 / 161291" type="journal article" year="1985">Biswas &amp; Chatterjee 1985</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE289129E08FBE9FE61FBF9" author="Mittal I. C." box="[265,408,1112,1139]" pageId="20" pageNumber="21" pagination="43 - 51" refId="ref30210" refString="Mittal I. C. 2005. Diversity and conservation status of dung beetles (Laparosticti: Scarabaeidae: Coleoptera) in North India. Bulletin of National Institute of Ecology 15: 43 - 51." type="journal article" year="2005">Mittal 2005</bibRefCitation>
;
<bibRefCitation id="B9494B35FFE289129EA7FBE9FD53FBF8" author="Bezdek A. &amp; Krell F. - T." box="[422,682,1112,1138]" pageId="20" pageNumber="21" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006</bibRefCitation>
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;
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;
<bibRefCitation id="B9494B35FFE289129FBCFBCAFE75FB1C" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[189,396,1147,1174]" pageId="20" pageNumber="21" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
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;
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;
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).
</paragraph>
</subSubSection>
</treatment>
</document>