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<mods:title id="F419A06FF7684C74B4B89361FC55FC7F">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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<mods:namePart id="D928E5FC96A14724AE9656503A84C2C8">Karimbumkara, Seena Narayanan</mods:namePart>
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<mods:namePart id="CF49833A0C2799F4DC28295105E27065">Priyadarsanan, Dharma Rajan</mods:namePart>
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<heading id="862F81A8FFDB892B9D69F987FC35F9DB" box="[616,972,1590,1617]" centered="true" fontSize="11" level="2" pageId="45" pageNumber="46" reason="2">
<taxonomicName id="1AD84D47FFDB892B9D69F987FC35F9DB" ID-CoL="49KMB" authority="Lansberge, 1875" authorityName="Lansberge" authorityYear="1875" box="[616,972,1590,1617]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFDB892B9D69F987FCF8F9DA" bold="true" box="[616,769,1590,1616]" italics="true" pageId="45" pageNumber="46">Onitis virens</emphasis>
<bibRefCitation id="B9494B35FFDB892B9C08F987FC35F9DB" author="Lansberge G." box="[777,972,1590,1617]" pageId="45" pageNumber="46" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
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<subSubSection id="95C2654FFFDB892B9D9CF9EBFC6EF9FE" box="[669,919,1626,1652]" pageId="45" pageNumber="46" type="description">
<paragraph id="DD6736C4FFDB892B9D9CF9EBFC6EF9FE" blockId="45.[616,972,1590,1652]" box="[669,919,1626,1652]" pageId="45" pageNumber="46">
<figureCitation id="45E32A41FFDB892B9D9CF9EBFCF2F9FE" box="[669,779,1626,1652]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). EF Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13742883" httpUri="https://zenodo.org/record/13742883/files/figure.png" pageId="45" pageNumber="46">Figs 12D</figureCitation>
,
<figureCitation id="45E32A41FFDB892B9C19F9EBFC97F9FE" box="[792,878,1626,1652]" captionStart="Fig" captionStartId="36.[189,232,1964,1990]" captionTargetBox="[275,1278,1055,1924]" captionTargetId="figure-228@36.[189,1398,1033,1927]" captionTargetPageId="36" captionText="Fig. 13. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitissubopacus Arrow, 1931, AIMB/ Co/Sc1000711. EH. O. virens Lansberge, 1875, AIMB/Co/Sc1000765. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742887" httpUri="https://zenodo.org/record/13742887/files/figure.png" pageId="45" pageNumber="46">13EH</figureCitation>
,
<figureCitation id="45E32A41FFDB892B9C7BF9EBFC6EF9FE" box="[890,919,1626,1652]" captionStart="Fig" captionStartId="50.[189,232,1930,1956]" captionTargetBox="[230,1361,265,1880]" captionTargetId="figure-1@50.[214,1374,251,1892]" captionTargetPageId="50" captionText="Fig. 20. Map showing the distribution of Onitissiva Gillet, 1911; O. subopacus Arrow, 1931; O. virens Lansberge, 1875 and O. visthara sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742911" httpUri="https://zenodo.org/record/13742911/files/figure.png" pageId="45" pageNumber="46">20</figureCitation>
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<emphasis id="EFACEAD6FFDB892B9FBCF92DFEAAF93C" box="[189,339,1692,1718]" italics="true" pageId="45" pageNumber="46">Onitisvirens</emphasis>
<bibRefCitation id="B9494B35FFDB892B9E5BF92AFDA0F93C" author="Lansberge G." box="[346,601,1691,1718]" pageId="45" pageNumber="46" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875: 135</bibRefCitation>
(original description).
</paragraph>
<paragraph id="DD6736C4FFDB892B9FBCF90FFD64F952" blockId="45.[189,864,1691,1753]" box="[189,669,1726,1753]" pageId="45" pageNumber="46">
<emphasis id="EFACEAD6FFDB892B9FBCF90FFE69F952" box="[189,400,1726,1752]" italics="true" pageId="45" pageNumber="46">Onitisamplectens</emphasis>
<bibRefCitation id="B9494B35FFDB892B9E96F90FFD60F952" author="Lansberge G." box="[407,665,1726,1753]" pageId="45" pageNumber="46" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875: 136</bibRefCitation>
.
</paragraph>
<paragraph id="DD6736C4FFDB89289FBCF8B1FC33FEE3" blockId="45.[189,1399,1792,1995]" lastBlockId="46.[234,1399,265,362]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<treatmentCitationGroup id="FDC811EAFFDB89289FBCF8B1FC33FEE3" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<taxonomicName id="1AD84D47FFDB892B9FBCF8B1FEAFF890" ID-CoL="49KMB" authorityName="Lansberge" authorityYear="1875" box="[189,342,1792,1818]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFDB892B9FBCF8B1FEAFF890" box="[189,342,1792,1818]" italics="true" pageId="45" pageNumber="46">Onitis virens</emphasis>
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<treatmentCitation id="5C7910D5FFDB892B9E7BF8B1FD1FF890" author="Boucomont A. &amp; Gillet J." box="[378,742,1792,1818]" page="19" pageId="45" pageNumber="46" year="1921">
<bibRefCitation id="B9494B35FFDB892B9E7BF8B1FD1FF890" author="Boucomont A. &amp; Gillet J." box="[378,742,1792,1818]" pageId="45" pageNumber="46" pagination="1 - 76" refId="ref27918" refString="Boucomont A. &amp; Gillet J. 1921. Faune entomologique de l'Indochine francaise. Famille Scarabaeidae Laparosticti (Coleopteres). Portail. Saigon 4: 1 - 76." type="journal article" year="1921">Boucomont &amp; Gillet 1921: 19</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDB892B9CC5F8B1FB6DF890" author="Arrow G. J." box="[964,1172,1792,1818]" page="396" pageId="45" pageNumber="46" year="1931">
<bibRefCitation id="B9494B35FFDB892B9CC5F8B1FB6DF890" author="Arrow G. J." box="[964,1172,1792,1818]" pageId="45" pageNumber="46" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow 1931: 396</bibRefCitation>
</treatmentCitation>
(key, description). —
<treatmentCitation id="5C7910D5FFDB892B9E14F892FE06F8B4" author="Balthasar V." box="[277,511,1827,1854]" page="52" pageId="45" pageNumber="46" year="1935">
<bibRefCitation id="B9494B35FFDB892B9E14F892FE06F8B4" author="Balthasar V." box="[277,511,1827,1854]" pageId="45" pageNumber="46" refId="ref27158" refString="Balthasar V. 1935. Scarabaeidae des palaarktischen Faunengebietes. Monographische Bestimmungstabelle. I. Coprinae I. Teil. Scarabaeini, Sisyphini, Panelini, Coprini, Onitini, Oniticellini. Edm. Reitter's Nachf. Emmerich Reitter, Troppau." type="book" year="1935">Balthasar 1935: 52</bibRefCitation>
</treatmentCitation>
(monograph); 1963: 40 (monograph). —
<treatmentCitation id="5C7910D5FFDB892B9B0FF892FB09F8B7" author="Paulian R." box="[1038,1264,1827,1854]" page="144" pageId="45" pageNumber="46" year="1945">
<bibRefCitation id="B9494B35FFDB892B9B0FF892FB09F8B7" author="Paulian R." box="[1038,1264,1827,1854]" pageId="45" pageNumber="46" refId="ref30535" refString="Paulian R. 1945. Faune de l'empire francais: Coleopteres Scarabeides de l'Indochine, Vol. III: 1 - 227. Librairie Larose, Paris." type="book" year="1945">Paulian 1945: 144</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="5C7910D5FFDB892B9E13F8F7FD74F8EA" author="Biswas S. &amp; Chatterjee S. K." box="[274,653,1862,1888]" page="260" pageId="45" pageNumber="46" year="1991">
<bibRefCitation id="B9494B35FFDB892B9E13F8F7FD74F8EA" author="Biswas S. &amp; Chatterjee S. K." box="[274,653,1862,1888]" pageId="45" pageNumber="46" pagination="243 - 262" refId="ref27741" refString="Biswas S. &amp; Chatterjee S. K. 1991. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, State Fauna Series 1: Fauna of Orissa (Part 3): 243 - 262. Available from: https: // shorturl. at / rZQQv [accessed 15 Oct. 2018]" type="book chapter" year="1991">Biswas &amp; Chatterjee 1991: 260</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDB892B9C6AF8F7FBA8F8EA" author="Chandra K." box="[875,1105,1862,1888]" page="360" pageId="45" pageNumber="46" year="2000">
<bibRefCitation id="B9494B35FFDB892B9C6AF8F7FBA8F8EA" author="Chandra K." box="[875,1105,1862,1888]" pageId="45" pageNumber="46" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000: 360</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDB892B9A2EF8F7FEA8F809" author="Mittal I. C." page="46" pageId="45" pageNumber="46" year="2005">
<bibRefCitation id="B9494B35FFDB892B9A2EF8F7FEA8F809" author="Mittal I. C." pageId="45" pageNumber="46" pagination="43 - 51" refId="ref30210" refString="Mittal I. C. 2005. Diversity and conservation status of dung beetles (Laparosticti: Scarabaeidae: Coleoptera) in North India. Bulletin of National Institute of Ecology 15: 43 - 51." type="journal article" year="2005">Mittal 2005: 46</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="5C7910D5FFDB892B9EC0F8D8FCDCF80E" author="Chandra K. &amp; Ahirwar S. C." box="[449,805,1897,1924]" page="149" pageId="45" pageNumber="46" year="2005">
<bibRefCitation id="B9494B35FFDB892B9EC0F8D8FCDCF80E" author="Chandra K. &amp; Ahirwar S. C." box="[449,805,1897,1924]" pageId="45" pageNumber="46" pagination="147 - 155" refId="ref28418" refString="Chandra K. &amp; Ahirwar S. C. 2005. Scarabaeid Beetles (Coleoptera) of Kanha Tiger Reserve, Madhya Pradesh. Records Zoological Survey of India 105 (1 - 2): 147 - 155." type="journal article" year="2005">Chandra &amp; Ahirwar 2005: 149</bibRefCitation>
</treatmentCitation>
(distribution); 2007: 276 (distribution). —
<treatmentCitation id="5C7910D5FFDB892B9A1EF8D8FE38F82D" author="Bezdek A. &amp; Krell F. - T." page="159" pageId="45" pageNumber="46" year="2006">
<bibRefCitation id="B9494B35FFDB892B9A1EF8D8FE38F82D" author="Bezdek A. &amp; Krell F. - T." pageId="45" pageNumber="46" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFDB892B9D95F83CFCA4F82C" author="Sewak R." box="[660,861,1932,1959]" page="60" pageId="45" pageNumber="46" year="2009">
<bibRefCitation id="B9494B35FFDB892B9D95F83CFCA4F82C" author="Sewak R." box="[660,861,1932,1959]" pageId="45" pageNumber="46" pagination="1 - 48" refId="ref31194" refString="Sewak R. 2009 a. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Thar desert of Gujarat. Records of the Zoological Survey of India, Occassional Paper No 295: 1 - 48." type="journal article" year="2009">Sewak 2009a: 60</bibRefCitation>
</treatmentCitation>
(key, diagnosis, distribution), 2009b: 60 (key, diagnosis, distribution). —
<treatmentCitation id="5C7910D5FFDB892B9D32F801FCDEF840" author="Sabu T. K. &amp; Nithya S. &amp; Vinod K. V." box="[563,807,1968,1994]" page="34" pageId="45" pageNumber="46" year="2011">
<bibRefCitation id="B9494B35FFDB892B9D32F801FCDEF840" author="Sabu T. K. &amp; Nithya S. &amp; Vinod K. V." box="[563,807,1968,1994]" pageId="45" pageNumber="46" pagination="29 - 38" refId="ref30906" refString="Sabu T. K., Nithya S. &amp; Vinod K. V. 2011. Faunal survey, endemism and possible species loss of Scarabaeinae (Coleoptera: Scarabaeidae) in the western slopes of the moist South Western Ghats, South India. Zootaxa 2830: 29 - 38. https: // doi. org / 10.11646 / ZOOTAXA. 2830.1.3" type="book chapter" year="2011">
Sabu
<emphasis id="EFACEAD6FFDB892B9D79F800FD4CF840" box="[632,693,1968,1994]" italics="true" pageId="45" pageNumber="46">et al.</emphasis>
2011: 34
</bibRefCitation>
</treatmentCitation>
(list). — Chandra
<emphasis id="EFACEAD6FFDB892B9B0CF800FBB3F840" box="[1037,1098,1968,1994]" italics="true" pageId="45" pageNumber="46">et al.</emphasis>
2012a: 52 (distribution). —
<treatmentCitation id="5C7910D5FFD889289E16FEB8FD37FEAE" author="Karimbumkara S. N. &amp; Rajan P. D." box="[279,718,265,292]" page="175" pageId="46" pageNumber="47" year="2013">
<bibRefCitation id="B9494B35FFD889289E16FEB8FD37FEAE" author="Karimbumkara S. N. &amp; Rajan P. D." box="[279,718,265,292]" pageId="46" pageNumber="47" pagination="173 - 178" refId="ref29842" refString="Karimbumkara S. N. &amp; Rajan P. D. 2013. Insecta: Coleoptera: Scarabaeidae: Scarabaeinae, Dung Beetles. Zoological Survey of India, Fauna of Karnataka, State Fauna Series 21: 173 - 178." type="book chapter" year="2013">Karimbumkara &amp; Rajan 2013: 175</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="5C7910D5FFD889289C56FEB8FB75FEA9" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[855,1164,265,292]" page="1037" pageId="46" pageNumber="47" year="2015">
<bibRefCitation id="B9494B35FFD889289C56FEB8FB75FEA9" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[855,1164,265,292]" pageId="46" pageNumber="47" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
Gupta
<emphasis id="EFACEAD6FFD889289CAFFEBBFC09FEA9" box="[942,1008,265,291]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
2015: 1037
</bibRefCitation>
</treatmentCitation>
(list, distribution). —
<treatmentCitation id="5C7910D5FFD889289E0FFE9DFDD2FECC" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[270,555,300,326]" page="301" pageId="46" pageNumber="47" year="2014">
<bibRefCitation id="B9494B35FFD889289E0FFE9DFDD2FECC" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[270,555,300,326]" pageId="46" pageNumber="47" pagination="295 - 307" refId="ref31267" refString="Siddiqui H., Ahmed Z. &amp; Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46 (2): 295 - 307." type="journal article" year="2014">
Siddiqui
<emphasis id="EFACEAD6FFD889289E76FE9CFE48FECC" box="[375,433,300,326]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
2014: 301
</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFD889289DFCFE9DFC29FECC" author="Bezdek A." box="[765,976,300,326]" page="180" pageId="46" pageNumber="47" year="2016">
<bibRefCitation id="B9494B35FFD889289DFCFE9DFC29FECC" author="Bezdek A." box="[765,976,300,326]" pageId="46" pageNumber="47" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 180</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="5C7910D5FFD889289BA3FE9DFEEFFEE0" author="Ghosh J. &amp; Das, P. &amp; Ghosh S. K. &amp; Bhunia D. &amp; Kushwaha R. K. &amp; Gupta D. &amp; Chandra K." page="239" pageId="46" pageNumber="47" year="2020">
<bibRefCitation id="B9494B35FFD889289BA3FE9DFEEFFEE0" author="Ghosh J. &amp; Das, P. &amp; Ghosh S. K. &amp; Bhunia D. &amp; Kushwaha R. K. &amp; Gupta D. &amp; Chandra K." pageId="46" pageNumber="47" pagination="221 - 275" refId="ref29141" refString="Ghosh J., Das, P., Ghosh S. K., Bhunia D., Kushwaha R. K., Gupta D., &amp; Chandra K. 2020. Insecta: Coleoptera. Zoological Survey of India, Fauna of Haryana, State Fauna Series 24: 221 - 275." type="book chapter" year="2020">
Ghosh
<emphasis id="EFACEAD6FFD889289BF5FE9CFAD7FECC" box="[1268,1326,300,326]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
2020: 239
</bibRefCitation>
</treatmentCitation>
(distribution). —
<bibRefCitation id="B9494B35FFD889289EEDFEFEFD1FFEE0" author="Schoolmeesters P." box="[492,742,335,362]" pageId="46" pageNumber="47" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFD889289FBCFE20FD53FDCA" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="DD6736C4FFD889289FBCFE20FE3FFE21" blockId="46.[189,454,401,427]" box="[189,454,401,427]" pageId="46" pageNumber="47">
<heading id="862F81A8FFD889289FBCFE20FE3FFE21" bold="true" box="[189,454,401,427]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="EFACEAD6FFD889289FBCFE20FE3FFE21" bold="true" box="[189,454,401,427]" pageId="46" pageNumber="47">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFD889289FBCFE71FA8EFE77" blockId="46.[189,1399,448,510]" pageId="46" pageNumber="47">
<emphasis id="EFACEAD6FFD889289FBCFE71FEB6FE50" box="[189,335,448,474]" italics="true" pageId="46" pageNumber="47">Onitisvirens</emphasis>
belongs to Group III and closely resembles
<taxonomicName id="1AD84D47FFD889289C4BFE71FC1FFE50" authorityName="Arrow" authorityYear="1931" box="[842,998,448,474]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="EFACEAD6FFD889289C4BFE71FC1FFE50" box="[842,998,448,474]" italics="true" pageId="46" pageNumber="47">O. subopacus</emphasis>
</taxonomicName>
of the same group but differs from the latter in having a strongly punctured pronotum and clypeo-frontal carina which is narrowly interrupted.
</paragraph>
<paragraph id="DD6736C4FFD889289FBCFD94FD53FDCA" blockId="46.[189,682,549,576]" box="[189,682,549,576]" pageId="46" pageNumber="47">
Also see remarks under
<emphasis id="EFACEAD6FFD889289ED9FD94FD5AFDB5" box="[472,675,549,575]" italics="true" pageId="46" pageNumber="47">Onitissubopacus</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFD889289FBCFDD6FB88F9B5" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="DD6736C4FFD889289FBCFDD6FE15FD0B" blockId="46.[189,492,615,641]" box="[189,492,615,641]" pageId="46" pageNumber="47">
<heading id="862F81A8FFD889289FBCFDD6FE15FD0B" bold="true" box="[189,492,615,641]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="EFACEAD6FFD889289FBCFDD6FE15FD0B" bold="true" box="[189,492,615,641]" pageId="46" pageNumber="47">Type material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="6DB03C99FFD889289FEBFD24FEA2FCB7" collectionCode="IRSNB" collectorName="A. Janssens &amp; Collection E. Candeze" determinerName="G. J. Arrow &amp; Van Lansberge" location="Van" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" stateProvince="Van" typeStatus="syntype">
<paragraph id="DD6736C4FFD889289FEBFD24FEA3FD25" blockId="46.[189,1399,661,900]" box="[234,346,661,687]" pageId="46" pageNumber="47">
<emphasis id="EFACEAD6FFD889289FEBFD24FEA3FD25" bold="true" box="[234,346,661,687]" pageId="46" pageNumber="47">
<typeStatus id="02638866FFD889289FEBFD24FEA3FD25" box="[234,346,661,687]" pageId="46" pageNumber="47" type="syntype">Syntypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFD889289FBCFD08FCD0FC09" blockId="46.[189,1399,661,900]" pageId="46" pageNumber="47">
UNKNOWN TYPE LOCALTY •
<specimenCount id="CBDEFD4DFFD889289D57FD0BFD94FD5E" box="[598,621,698,724]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
;
<emphasis id="EFACEAD6FFD889289D7FFD0BFCDFFD5E" box="[638,806,698,724]" italics="true" pageId="46" pageNumber="47">“Onitisvirens</emphasis>
Lansb.,
<determinerName id="28BBDA00FFD889289C8EFD08FBE4FD5E" box="[911,1053,697,724]" pageId="46" pageNumber="47">G. J. Arrow</determinerName>
det.;
<collectorName id="702D5312FFD889289B60FD0BFB15FD5E" box="[1121,1260,698,724]" pageId="46" pageNumber="47">A. Janssens</collectorName>
vid., 1936:
<taxonomicName id="1AD84D47FFD889289FBCFD6CFE38FD7C" authority="Lansb." authorityName="Lansberge" authorityYear="1875" box="[189,449,732,759]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFD889289FBCFD6CFE90FD7C" box="[189,361,733,759]" italics="true" pageId="46" pageNumber="47">ONITIS virens</emphasis>
Lansb.
</taxonomicName>
; Syntype; det
<determinerName id="28BBDA00FFD889289D71FD6CFCDCFD7C" box="[624,805,732,759]" pageId="46" pageNumber="47">Van Lansberge</determinerName>
.
<emphasis id="EFACEAD6FFD889289C30FD6DFBFFFD7C" box="[817,1030,732,758]" italics="true" pageId="46" pageNumber="47">Onitisamplectens</emphasis>
Lansb.;
<taxonomicName id="1AD84D47FFD889289B71FD6DFB0EFD7C" box="[1136,1271,732,758]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="amplectens">
<emphasis id="EFACEAD6FFD889289B71FD6DFB0EFD7C" box="[1136,1271,732,758]" italics="true" pageId="46" pageNumber="47">Amplectens</emphasis>
</taxonomicName>
Type Lsb. Mercara;
<emphasis id="EFACEAD6FFD889289E2AFCB1FE47FC90" box="[299,446,768,794]" italics="true" pageId="46" pageNumber="47">Onitisvirens</emphasis>
Lansb.Révis.Arrow 1927; Ex-Typis de
<taxonomicName id="1AD84D47FFD889289C8BFCB1FB98FC90" authority="Lansb." authorityName="Lansb." box="[906,1121,768,794]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="amplectens">
<emphasis id="EFACEAD6FFD889289C8BFCB1FBF7FC90" box="[906,1038,768,794]" italics="true" pageId="46" pageNumber="47">amplectens</emphasis>
Lansb.
</taxonomicName>
;
<collectorName id="702D5312FFD889289B6CFCB1FA8AFC90" box="[1133,1395,768,794]" pageId="46" pageNumber="47">Collection E. Candeze</collectorName>
: 109”;
<collectionCode id="BBC9AE01FFD889289E04FC92FEA2FCB7" box="[261,347,803,829]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="46" pageNumber="47">IRSNB</collectionCode>
<materialsCitation id="6DB03C99FFD889289E74FC92FCDDFC09" collectionCode="IRSNB" collectorName="Collection E. Candeze" determinerName="Van Lansberge" location="Van" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" stateProvince="Van" typeStatus="syntype">
<specimenCount id="CBDEFD4DFFD889289E74FC92FE75FCB7" box="[373,396,803,829]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
; “A. Janssens vid., 1936:
<taxonomicName id="1AD84D47FFD889289DBCFC92FC44FCB7" authority="Lansb." authorityName="Lansberge" authorityYear="1875" box="[701,957,803,829]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFD889289DBCFC92FC9EFCB7" box="[701,871,803,829]" italics="true" pageId="46" pageNumber="47">ONITIS virens</emphasis>
Lansb.
</taxonomicName>
; cf. Ann. Soc. Ent. Belg. Xviii. 1875, p.135; Syntype; Ex-Typis de
<taxonomicName id="1AD84D47FFD889289D14FCF7FD09FCEA" authority="Lansb." authorityName="Lansb." box="[533,752,838,864]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="amplectens">amplectens Lansb.</taxonomicName>
; AmplectensInd br Lansb.; Det.
<determinerName id="28BBDA00FFD889289B79FCF6FADEFCEA" box="[1144,1319,838,865]" pageId="46" pageNumber="47">Van Lansberge</determinerName>
<taxonomicName id="1AD84D47FFD889289A2EFCF6FE61FC09" authority="Lansb." authorityName="Lansb." class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="amplectens">
<emphasis id="EFACEAD6FFD889289A2EFCF6FEB8FC09" italics="true" pageId="46" pageNumber="47">Onitis amplectens</emphasis>
Lansb.
</taxonomicName>
;
<collectorName id="702D5312FFD889289EA6FCD8FD4DFC09" box="[423,692,873,900]" pageId="46" pageNumber="47">Collection E. Candeze</collectorName>
”;
<collectionCode id="BBC9AE01FFD889289DCBFCD8FCDDFC09" box="[714,804,873,899]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="46" pageNumber="47">IRSNB</collectionCode>
</materialsCitation>
.
</paragraph>
</materialsCitation>
<paragraph id="DD6736C4FFD889289FEBFC1AFCE2FC4F" blockId="46.[189,1400,939,1600]" box="[234,795,939,965]" pageId="46" pageNumber="47">
<emphasis id="EFACEAD6FFD889289FEBFC1AFD9BFC4F" bold="true" box="[234,610,939,965]" pageId="46" pageNumber="47">Additional material examined</emphasis>
(
<specimenCount id="CBDEFD4DFFD889289D70FC1AFCEAFC4F" box="[625,787,939,965]" count="47" pageId="46" pageNumber="47" type="generic">47 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFD889289FBCFC7EFB88F9B5" blockId="46.[189,1400,939,1600]" pageId="46" pageNumber="47">
<materialsCitation id="6DB03C99FFD889289FBCFC7EFB62FC63" box="[189,1179,974,1001]" collectionCode="BMNH" country="India" location="INDIA" pageId="46" pageNumber="47" specimenCode="(E) 1237153" specimenCount="1" specimenCount-male="1">
<collectingCountry id="A5CF7654FFD889289FBCFC7EFEF2FC63" box="[189,267,975,1001]" name="India" pageId="46" pageNumber="47">INDIA</collectingCountry>
<specimenCount id="CBDEFD4DFFD889289E2BFC7EFEBBFC63" box="[298,322,975,1001]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
; “
<taxonomicName id="1AD84D47FFD889289E5CFC7EFD33FC63" authority="Lansb., Compared" authorityName="Lansb., Compared" box="[349,714,975,1001]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFD889289E5CFC7EFE17FC62" box="[349,494,975,1001]" italics="true" pageId="46" pageNumber="47">Onitis virens</emphasis>
Lansb., Compared
</taxonomicName>
with type, G.J.A.”;
<collectionCode id="BBC9AE01FFD889289CB6FC7EFBF6FC63" box="[951,1039,975,1001]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="46" pageNumber="47">BMNH</collectionCode>
<specimenCode id="8D7E9EBFFFD889289B0FFC7EFB62FC63" box="[1038,1179,974,1001]" pageId="46" pageNumber="47">(E) 1237153</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289BBBFC7EFC4DFB87" collectionCode="BMNH" collectorName="H. Stevens." country="India" location="Rungbong Vall." municipality="Gopaldhara" pageId="46" pageNumber="47" specimenCode="(E)1237154" specimenCount="1" specimenCount-female="1" stateProvince="Sikkim">
<specimenCount id="CBDEFD4DFFD889289BBBFC7EFB29FC63" box="[1210,1232,975,1001]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
; “
<collectingMunicipality id="3D03ACBEFFD889289BEAFC7EFA8AFC63" box="[1259,1395,975,1001]" pageId="46" pageNumber="47">Gopaldhara</collectingMunicipality>
,
<location id="D807601FFFD889289FBCFC43FE92FB86" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289FBCFC43FE92FB86" box="[189,363,1010,1036]" country="India" municipality="Gopaldhara" name="Rungbong Vall." pageId="46" pageNumber="47" stateProvince="Sikkim">Rungbong Vall.</location>
,
<collectingRegion id="1F1CF826FFD889289E73FC43FE3FFB86" box="[370,454,1010,1036]" country="India" name="Sikkim" pageId="46" pageNumber="47">Sikkim</collectingRegion>
,
<collectorName id="702D5312FFD889289ECCFC43FDB5FB86" box="[461,588,1010,1036]" pageId="46" pageNumber="47">H.Stevens.</collectorName>
, 1916-218;
<collectionCode id="BBC9AE01FFD889289DC8FC43FCD8FB86" box="[713,801,1010,1036]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="46" pageNumber="47">BMNH</collectionCode>
<specimenCode id="8D7E9EBFFFD889289C20FC43FC50FB87" box="[801,937,1010,1037]" pageId="46" pageNumber="47">(E)1237154</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289CC5FC43FE42FBA5" collectingDate="1927-02-23" country="India" county="Nerbuda Survey" elevation="823" location="Bewa State" municipality="Harai" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1">
<specimenCount id="CBDEFD4DFFD889289CC5FC43FC25FB86" box="[964,988,1010,1036]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
; “
<collectingCounty id="34064E48FFD889289CEEFC43FB5CFB86" box="[1007,1189,1010,1036]" pageId="46" pageNumber="47">Nerbuda Survey</collectingCounty>
,
<collectingMunicipality id="3D03ACBEFFD889289BADFC43FB14FB86" box="[1196,1261,1010,1036]" pageId="46" pageNumber="47">Harai</collectingMunicipality>
,
<location id="D807601FFFD889289BF5FC43FA8AFB86" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289BF5FC43FA8AFB86" box="[1268,1395,1010,1036]" country="India" county="Nerbuda Survey" municipality="Harai" name="Bewa State" pageId="46" pageNumber="47">Bewa State</location>
,
<quantity id="1A209B21FFD889289FBCFBA4FEECFBA5" box="[189,277,1045,1071]" metricMagnitude="2" metricUnit="m" metricValue="8.2296" pageId="46" pageNumber="47" unit="ft" value="2700.0">
<elevation id="56F5D1F7FFD889289FBCFBA4FEECFBA5" box="[189,277,1045,1071]" metricMagnitude="2" metricUnit="m" metricValue="8.2296" pageId="46" pageNumber="47" unit="ft" value="2700.0">2700 ft.</elevation>
</quantity>
;
<date id="A9661004FFD889289E1FFBA4FE49FBA5" box="[286,432,1045,1071]" pageId="46" pageNumber="47" value="1927-02-23">
<collectingDate id="B922E9ECFFD889289E1FFBA4FE49FBA5" box="[286,432,1045,1071]" pageId="46" pageNumber="47" value="1927-02-23">23 Feb. 1927</collectingDate>
</date>
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289ECFFBA7FDCDFBA5" box="[462,564,1045,1072]" collectionCode="ZSIM" country="India" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBDEFD4DFFD889289ECFFBA7FE1DFBBA" box="[462,484,1046,1072]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
;
<collectionCode id="BBC9AE01FFD889289EF1FBA4FDCDFBA5" box="[496,564,1045,1071]" pageId="46" pageNumber="47">ZSIM</collectionCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289D49FBA4FD83FBFC" collectingDate="1999-05-14" collectingMethod="open cow dung bait traps" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan" country="India" latitude="11.837999" location="Budipadaga" longLatPrecision="1" longitude="77.07755" municipality="Biligiri Rangaswamy Tiger Reserve" pageId="46" pageNumber="47" specimenCode="Co/Sc1000746 to 1000749" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Karnataka">
<specimenCount id="CBDEFD4DFFD889289D49FBA4FD71FBBA" box="[584,648,1045,1072]" count="2" pageId="46" pageNumber="47" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFD889289D93FBA4FD28FBBA" box="[658,721,1045,1072]" count="2" pageId="46" pageNumber="47" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFD889289DDCFBA4FCABFBA5" box="[733,850,1045,1071]" country="India" name="Karnataka" pageId="46" pageNumber="47">Karnataka</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFD889289C5AFBA4FB11FBBA" box="[859,1256,1045,1072]" pageId="46" pageNumber="47">Biligiri Rangaswamy Tiger Reserve</collectingMunicipality>
,
<location id="D807601FFFD889289BF0FBA4FA8EFBA5" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289BF0FBA4FA8EFBA5" box="[1265,1399,1045,1071]" country="India" latitude="11.837999" longLatPrecision="1" longitude="77.07755" municipality="Biligiri Rangaswamy Tiger Reserve" name="Budipadaga" pageId="46" pageNumber="47" stateProvince="Karnataka">Budipadaga</location>
scrub;
<geoCoordinate id="B8EC5003FFD889289E09FB86FE53FBD9" box="[264,426,1079,1107]" degrees="11" direction="north" minutes="50" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="16.8" value="11.837999">11°50ʹ16.8″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD889289EB7FB86FDAFFBD9" box="[438,598,1079,1107]" degrees="77" direction="east" minutes="4" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="39.18" value="77.07755">77°4ʹ39.18″ E</geoCoordinate>
;
<date id="A9661004FFD889289D63FB89FD02FBD8" box="[610,763,1080,1107]" pageId="46" pageNumber="47" value="1999-05-14">
<collectingDate id="B922E9ECFFD889289D63FB89FD02FBD8" box="[610,763,1080,1107]" pageId="46" pageNumber="47" value="1999-05-14">14 May 1999</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFD889289C06FB88FBDDFBD9" box="[775,1060,1080,1107]" pageId="46" pageNumber="47">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFD889289B2EFB89FA8EFBD8" box="[1071,1399,1080,1106]" pageId="46" pageNumber="47">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFD889289FF7FBEAFEC6FBFF" box="[246,319,1115,1141]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFD889289E49FBEAFD83FBFC" box="[328,634,1115,1142]" pageId="46" pageNumber="47">Co/Sc1000746 to 1000749</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289D9BFBEDFE3CFB36" collectingDate="1999-12-22" collectingMethod="open cow dung bait traps" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan" country="India" latitude="11.89545" location="Kanneri" longLatPrecision="1" longitude="77.13596" pageId="46" pageNumber="47" specimenCode="Co/ Sc1000750 to 1000762" specimenCount="13" specimenCount-female="5" specimenCount-male="8">
<specimenCount id="CBDEFD4DFFD889289D9BFBEDFD19FBFC" box="[666,736,1116,1142]" count="8" pageId="46" pageNumber="47" type="male">8 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFD889289DEEFBEAFCCDFBFC" box="[751,820,1115,1142]" count="5" pageId="46" pageNumber="47" type="female">5 ♀♀</specimenCount>
;
<location id="D807601FFFD889289C44FBEAFC66FBFF" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289C44FBEAFC66FBFF" box="[837,927,1115,1141]" country="India" latitude="11.89545" longLatPrecision="1" longitude="77.13596" name="Kanneri" pageId="46" pageNumber="47">Kanneri</location>
moist deciduous forest;
<geoCoordinate id="B8EC5003FFD889289BBFFBEBFA8AFBFC" box="[1214,1395,1114,1142]" degrees="11" direction="north" minutes="53" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="43.62" value="11.89545">11°53ʹ43.62″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD889289FBCFBCCFEACFB13" box="[189,341,1149,1177]" degrees="77" direction="east" minutes="8" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="9.48" value="77.13596">77°8ʹ9.48″ E</geoCoordinate>
;
<date id="A9661004FFD889289E66FBCEFDF5FB13" box="[359,524,1151,1177]" pageId="46" pageNumber="47" value="1999-12-22">
<collectingDate id="B922E9ECFFD889289E66FBCEFDF5FB13" box="[359,524,1151,1177]" pageId="46" pageNumber="47" value="1999-12-22">22 Dec. 1999</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFD889289D1CFBCEFCA8FB13" box="[541,849,1151,1177]" pageId="46" pageNumber="47">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFD889289C63FBCEFB4DFB13" box="[866,1204,1151,1177]" pageId="46" pageNumber="47">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFD889289BFCFBCEFABFFB13" box="[1277,1350,1151,1177]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFD889289A4FFBCEFE3CFB36" pageId="46" pageNumber="47">Co/ Sc1000750 to 1000762</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289EE3FB13FB11FB55" collectingDate="1999-05-11" collectingMethod="open cow dung bait traps" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan" country="India" latitude="12.032" location="Purani" longLatPrecision="1" longitude="77.1313" pageId="46" pageNumber="47" specimenCode="Co/Sc1000763 to 1000764" specimenCount="2" specimenCount-male="2">
<specimenCount id="CBDEFD4DFFD889289EE3FB13FDDEFB36" box="[482,551,1186,1212]" count="2" pageId="46" pageNumber="47" type="male">2 ♂♂</specimenCount>
,
<location id="D807601FFFD889289D35FB13FD84FB36" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289D35FB13FD84FB36" box="[564,637,1186,1212]" country="India" latitude="12.032" longLatPrecision="1" longitude="77.1313" name="Purani" pageId="46" pageNumber="47">Purani</location>
dry deciduous forest;
<geoCoordinate id="B8EC5003FFD889289C7EFB11FBEFFB36" box="[895,1046,1184,1212]" degrees="12" direction="north" minutes="1" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="55.2" value="12.032">12°1ʹ55.2″ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD889289B25FB11FB3EFB36" box="[1060,1223,1184,1212]" degrees="77" direction="east" minutes="7" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="52.68" value="77.1313">77°7ʹ52.68″ E</geoCoordinate>
;
<date id="A9661004FFD889289BD4FB13FA8AFB36" box="[1237,1395,1186,1212]" pageId="46" pageNumber="47" value="1999-05-11">
<collectingDate id="B922E9ECFFD889289BD4FB13FA8AFB36" box="[1237,1395,1186,1212]" pageId="46" pageNumber="47" value="1999-05-11">11 May 1999</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFD889289FBCFB77FE23FB6A" box="[189,474,1221,1248]" pageId="46" pageNumber="47">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFD889289EE7FB74FCD7FB55" box="[486,814,1221,1247]" pageId="46" pageNumber="47">Priyadarsanan Dharma Rajan</collectorName>
leg.;
<collectionCode id="BBC9AE01FFD889289C6AFB74FC4DFB55" box="[875,948,1221,1247]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFD889289CBCFB74FB11FB55" box="[957,1256,1221,1248]" pageId="46" pageNumber="47">Co/Sc1000763 to 1000764</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289A00FB77FAE3FAAC" collectingDate="2009-12-03" collectingMethod="open cow dung bait traps" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara" country="India" elevation="207" latitude="8.918222" location="Kattilapara" longLatPrecision="1" longitude="77.09575" municipality="Shendurney Wildlife Sanctuary" pageId="46" pageNumber="47" specimenCode="Co/Sc1000765" specimenCount="1" specimenCount-male="1" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFD889289A00FB77FAE0FB6A" box="[1281,1305,1222,1248]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
;
<collectingRegion id="1F1CF826FFD889289A24FB74FA8AFB55" box="[1317,1395,1221,1247]" country="India" name="Kerala" pageId="46" pageNumber="47">Kerala</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFD889289FBCFB59FDE3FA88" box="[189,538,1256,1282]" pageId="46" pageNumber="47">Shendurney Wildlife Sanctuary</collectingMunicipality>
,
<location id="D807601FFFD889289D24FB59FD59FA88" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289D24FB59FD59FA88" box="[549,672,1256,1282]" country="India" latitude="8.918222" longLatPrecision="1" longitude="77.09575" municipality="Shendurney Wildlife Sanctuary" name="Kattilapara" pageId="46" pageNumber="47" stateProvince="Kerala">Kattilapara</location>
degraded forest;
<geoCoordinate id="B8EC5003FFD889289C61FB56FBF9FA89" box="[864,1024,1255,1283]" degrees="08" direction="north" minutes="55" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="05.6" value="8.918222">08°55ʹ05.6ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD889289B0AFB56FB50FA89" box="[1035,1193,1255,1283]" degrees="77" direction="east" minutes="05" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="44.7" value="77.09575">77°05ʹ44.7ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFD889289BEFFB59FA8BFA88" box="[1262,1394,1256,1283]" metricMagnitude="2" metricUnit="m" metricValue="2.07" pageId="46" pageNumber="47" unit="m" value="207.0">
<quantity id="1A209B21FFD889289BEFFB59FACFFA89" box="[1262,1334,1256,1283]" metricMagnitude="2" metricUnit="m" metricValue="2.07" pageId="46" pageNumber="47" unit="m" value="207.0">207 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFD889289FBCFABDFEB1FAAC" box="[189,328,1292,1318]" pageId="46" pageNumber="47" value="2009-12-03">
<collectingDate id="B922E9ECFFD889289FBCFABDFEB1FAAC" box="[189,328,1292,1318]" pageId="46" pageNumber="47" value="2009-12-03">3 Dec. 2009</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFD889289E53FABDFD92FAAC" box="[338,619,1291,1318]" pageId="46" pageNumber="47">open cow dung bait traps</collectingMethod>
;
<collectorName id="702D5312FFD889289D77FABDFC1EFAAF" box="[630,999,1291,1318]" pageId="46" pageNumber="47">Seena Narayanan Karimbumkara</collectorName>
leg.;
<collectionCode id="BBC9AE01FFD889289B22FABAFB95FAAC" box="[1059,1132,1291,1318]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFD889289B74FABAFAE3FAAC" box="[1141,1306,1291,1318]" pageId="46" pageNumber="47">Co/Sc1000765</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289A31FABDFE38FA05" collectingDate="2018-09-30" collectingMethod="cow dung baited pitfall trap" collectionCode="AIMB" collectorName="Harsha Malhotra" country="India" elevation="127" latitude="27.96913" location="Rani Village" longLatPrecision="1" longitude="95.32144" municipality="Pasighat" pageId="46" pageNumber="47" specimenCode="Co/ Sc1000766 to 1000769" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Arunachal Pradesh">
<specimenCount id="CBDEFD4DFFD889289A31FABDFA8BFAAC" box="[1328,1394,1292,1318]" count="2" pageId="46" pageNumber="47" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFD889289FBCFA9EFEF8FAC3" box="[189,257,1327,1353]" count="2" pageId="46" pageNumber="47" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="1F1CF826FFD889289E0EFA9EFE1EFAC3" box="[271,487,1327,1353]" country="India" name="Arunachal Pradesh" pageId="46" pageNumber="47">Arunachal Pradesh</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFD889289EF5FA9EFDACFAC3" box="[500,597,1327,1353]" pageId="46" pageNumber="47">Pasighat</collectingMunicipality>
, community forest,
<location id="D807601FFFD889289C3BFA9EFC35FAC3" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289C3BFA9EFC35FAC3" box="[826,972,1327,1353]" country="India" latitude="27.96913" longLatPrecision="1" longitude="95.32144" municipality="Pasighat" name="Rani Village" pageId="46" pageNumber="47" stateProvince="Arunachal Pradesh">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFD889289CD8FA9CFB60FAC3" box="[985,1177,1325,1353]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="8.8644" value="27.96913">27°58ʹ8.8644ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD889289BA6FA9CFA8BFAC3" box="[1191,1394,1325,1353]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="17.1804" value="95.32144">95°19ʹ17.1804ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFD889289FFBFAE3FE7AFAE6" box="[250,387,1362,1388]" metricMagnitude="2" metricUnit="m" metricValue="1.27" pageId="46" pageNumber="47" unit="m" value="127.0">
<quantity id="1A209B21FFD889289FFBFAE3FEBDFAE6" box="[250,324,1362,1388]" metricMagnitude="2" metricUnit="m" metricValue="1.27" pageId="46" pageNumber="47" unit="m" value="127.0">127 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFD889289E90FAE3FDD7FAE6" box="[401,558,1362,1388]" pageId="46" pageNumber="47" value="2018-09-30">
<collectingDate id="B922E9ECFFD889289E90FAE3FDD7FAE6" box="[401,558,1362,1388]" pageId="46" pageNumber="47" value="2018-09-30">30 Sep. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFD889289D3AFAE3FC84FAE6" box="[571,893,1362,1388]" pageId="46" pageNumber="47">cow dung baited pitfall trap</collectingMethod>
;
<collectorName id="702D5312FFD889289C8BFAE3FBB0FAE6" box="[906,1097,1362,1388]" pageId="46" pageNumber="47">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFD889289BFCFAE3FABFFAE6" box="[1277,1350,1362,1388]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFD889289A4FFAE3FE38FA05" pageId="46" pageNumber="47">Co/ Sc1000766 to 1000769</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289ED9FAC4FE47FA5C" collectingDate="2018-09-30" collectionCode="AIMB" collectorName="Harsha Malhotra" country="India" elevation="131" latitude="27.969614" location="Rani Village" longLatPrecision="1" longitude="95.32392" pageId="46" pageNumber="47" specimenCode="Co/ Sc1000770 to 1000784" specimenCount="15" specimenCount-female="9" specimenCount-male="6">
<specimenCount id="CBDEFD4DFFD889289ED9FAC4FDE2FA1A" box="[472,539,1397,1424]" count="6" pageId="46" pageNumber="47" type="male">6 ♂♂</specimenCount>
,
<specimenCount id="CBDEFD4DFFD889289D27FAC4FD9EFA1A" box="[550,615,1397,1424]" count="9" pageId="46" pageNumber="47" type="female">9 ♀♀</specimenCount>
; agricultural field,
<location id="D807601FFFD889289C3BFAC4FC33FA05" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289C3BFAC4FC33FA05" box="[826,970,1397,1423]" country="India" latitude="27.969614" longLatPrecision="1" longitude="95.32392" name="Rani Village" pageId="46" pageNumber="47">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFD889289CD5FAC2FB66FA1A" box="[980,1183,1395,1424]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="10.6068" value="27.969614">27°58ʹ10.6068ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD889289BAAFAC2FA8BFA1A" box="[1195,1394,1395,1424]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="26.1156" value="95.32392">95°19ʹ26.1156ʺ E</geoCoordinate>
, elev.
<elevation id="56F5D1F7FFD889289FF8FA29FE7BFA38" box="[249,386,1432,1459]" metricMagnitude="2" metricUnit="m" metricValue="1.31" pageId="46" pageNumber="47" unit="m" value="131.0">
<quantity id="1A209B21FFD889289FF8FA29FEBDFA39" box="[249,324,1432,1459]" metricMagnitude="2" metricUnit="m" metricValue="1.31" pageId="46" pageNumber="47" unit="m" value="131.0">131 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFD889289E91FA29FDD2FA38" box="[400,555,1432,1458]" pageId="46" pageNumber="47" value="2018-09-30">
<collectingDate id="B922E9ECFFD889289E91FA29FDD2FA38" box="[400,555,1432,1458]" pageId="46" pageNumber="47" value="2018-09-30">30 Sep. 2018</collectingDate>
</date>
; hand-picked from cow dung;
<collectorName id="702D5312FFD889289C8DFA29FBB3FA38" box="[908,1098,1432,1458]" pageId="46" pageNumber="47">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFD889289BFCFA29FABFFA38" box="[1277,1350,1432,1458]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFD889289A4FFA29FE47FA5C" pageId="46" pageNumber="47">Co/ Sc1000770 to 1000784</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289ED5FA0DFEC2F996" collectingDate="2018-09-30" collectingMethod="cow dung baited pitfall trap" collectionCode="AIMB" collectorName="Harsha Malhotra" country="India" elevation="133" latitude="27.969172" location="Rani Village" longLatPrecision="1" longitude="95.32194" pageId="46" pageNumber="47" specimenCode="Co/Sc1000785 to 1000787" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount id="CBDEFD4DFFD889289ED5FA0DFE12FA5C" box="[468,491,1468,1494]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
,
<specimenCount id="CBDEFD4DFFD889289EF7FA0DFDCFFA5C" box="[502,566,1468,1494]" count="2" pageId="46" pageNumber="47" type="female">2 ♀♀</specimenCount>
; community forest,
<location id="D807601FFFD889289C12FA0AFC58FA5F" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289C12FA0AFC58FA5F" box="[787,929,1467,1493]" country="India" latitude="27.969172" longLatPrecision="1" longitude="95.32194" name="Rani Village" pageId="46" pageNumber="47">Rani Village</location>
;
<geoCoordinate id="B8EC5003FFD889289CAAFA0BFB9EFA5C" box="[939,1127,1466,1494]" degrees="27" direction="north" minutes="58" orientation="latitude" pageId="46" pageNumber="47" precision="1" seconds="9.0156" value="27.969172">27°58ʹ9.0156ʺ N</geoCoordinate>
,
<geoCoordinate id="B8EC5003FFD889289B70FA0BFAC1FA5C" box="[1137,1336,1466,1494]" degrees="95" direction="east" minutes="19" orientation="longitude" pageId="46" pageNumber="47" precision="1" seconds="18.9804" value="95.32194">95°19ʹ18.9804ʺ E</geoCoordinate>
; elev.
<elevation id="56F5D1F7FFD889289FBCFA6EFEC7FA73" box="[189,318,1503,1529]" metricMagnitude="2" metricUnit="m" metricValue="1.33" pageId="46" pageNumber="47" unit="m" value="133.0">
<quantity id="1A209B21FFD889289FBCFA6EFEFDFA73" box="[189,260,1503,1529]" metricMagnitude="2" metricUnit="m" metricValue="1.33" pageId="46" pageNumber="47" unit="m" value="133.0">133 m</quantity>
a.s.l.
</elevation>
;
<date id="A9661004FFD889289E49FA6EFE24FA73" box="[328,477,1503,1529]" pageId="46" pageNumber="47" value="2018-09-30">
<collectingDate id="B922E9ECFFD889289E49FA6EFE24FA73" box="[328,477,1503,1529]" pageId="46" pageNumber="47" value="2018-09-30">30 Sep. 2018</collectingDate>
</date>
;
<collectingMethod id="04994ED3FFD889289EE7FA6EFCEEFA73" box="[486,791,1503,1529]" pageId="46" pageNumber="47">cow dung baited pitfall trap</collectingMethod>
;
<collectorName id="702D5312FFD889289C20FA6EFC22FA73" box="[801,987,1503,1529]" pageId="46" pageNumber="47">Harsha Malhotra</collectorName>
and team leg.;
<collectionCode id="BBC9AE01FFD889289B7EFA6FFB30FA73" box="[1151,1225,1502,1529]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFD889289BD3FA6FFEC2F996" pageId="46" pageNumber="47">Co/Sc1000785 to 1000787</specimenCode>
</materialsCitation>
<materialsCitation id="6DB03C99FFD889289E51F9B3FB94F9B5" collectingDate="2021-04-15" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara" country="India" location="Intanki National Park" municipality="Bongkolong village" pageId="46" pageNumber="47" specimenCode="Co/Sc1000788" specimenCount="1" specimenCount-female="1" stateProvince="Nagaland">
<specimenCount id="CBDEFD4DFFD889289E51F9B3FE9FF996" box="[336,358,1538,1564]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
;
<collectingRegion id="1F1CF826FFD889289E72F9B3FE18F996" box="[371,481,1538,1564]" country="India" name="Nagaland" pageId="46" pageNumber="47">Nagaland</collectingRegion>
,
<collectingMunicipality id="3D03ACBEFFD889289EEBF9B3FD31F996" box="[490,712,1538,1564]" pageId="46" pageNumber="47">Bongkolong village</collectingMunicipality>
, near
<location id="D807601FFFD889289C09F9B3FC01F996" LSID="urn:lsid:plazi:treatment:557187D2FFDB89359D69F987FE3FFD64:D807601FFFD889289C09F9B3FC01F996" box="[776,1016,1538,1564]" country="India" municipality="Bongkolong village" name="Intanki National Park" pageId="46" pageNumber="47" stateProvince="Nagaland">Intanki National Park</location>
, open ground;
<date id="A9661004FFD889289B9BF9B3FAD5F996" box="[1178,1324,1538,1565]" pageId="46" pageNumber="47" value="2021-04-15">
<collectingDate id="B922E9ECFFD889289B9BF9B3FAD5F996" box="[1178,1324,1538,1565]" pageId="46" pageNumber="47" value="2021-04-15">15 Apr. 2021</collectingDate>
</date>
; hand-picked from cow dung;
<collectorName id="702D5312FFD889289EC7F994FCCEF9B5" box="[454,823,1573,1600]" pageId="46" pageNumber="47">Seena Narayanan Karimbumkara</collectorName>
leg.;
<collectionCode id="BBC9AE01FFD889289C73F994FC45F9B5" box="[882,956,1573,1599]" pageId="46" pageNumber="47">AIMB</collectionCode>
/
<specimenCode id="8D7E9EBFFFD889289CC7F994FB94F9B5" box="[966,1133,1573,1599]" pageId="46" pageNumber="47">Co/Sc1000788</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFD889359FBCF9D6FB94FE21" lastPageId="51" lastPageNumber="52" pageId="46" pageNumber="47" type="description">
<paragraph id="DD6736C4FFD889289FBCF9D6FEB7F90B" blockId="46.[189,334,1639,1665]" box="[189,334,1639,1665]" pageId="46" pageNumber="47">
<heading id="862F81A8FFD889289FBCF9D6FEB7F90B" bold="true" box="[189,334,1639,1665]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="EFACEAD6FFD889289FBCF9D6FEB7F90B" bold="true" box="[189,334,1639,1665]" pageId="46" pageNumber="47">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFD889289FBCF927FCE2F93B" blockId="46.[189,795,1686,1713]" box="[189,795,1686,1713]" pageId="46" pageNumber="47">MEASUREMENTS (in mm). TL=1723, BW= 9.513.</paragraph>
<paragraph id="DD6736C4FFD889289FEBF969FED3F978" blockId="46.[189,1399,1752,2025]" box="[234,298,1752,1778]" pageId="46" pageNumber="47">
<emphasis id="EFACEAD6FFD889289FEBF969FED3F978" bold="true" box="[234,298,1752,1778]" pageId="46" pageNumber="47">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFD889289FBCF94AFAA7F862" blockId="46.[189,1399,1752,2025]" pageId="46" pageNumber="47">Black with a very feeble metallic lustre; clypeus elliptical, margin entire, closely and finely rugulose, vertex rugulose and more shining, frontal carina interrupted by a very short medial gap, with a conical tubercle behind and an anterior short transverse clypeal carina; pronotum fairly closely and strongly punctured, with an anterior narrow smooth median line and slight narrow groove behind; elytra finely striate with intervals flat and imperceptibly punctured, the sutural interval slightly elevated and shining; pygidium flat, smooth and feebly punctured; metasternal shield smooth and shining behind, front angles and sides of metasternum clothed with red hairs. Protibia slender, curved, mid femur and hind trochanter toothed.</paragraph>
<caption id="89A7664CFFD989299FBCF83BFE4BF860" ID-DOI="http://doi.org/10.5281/zenodo.13742901" ID-Zenodo-Dep="13742901" httpUri="https://zenodo.org/record/13742901/files/figure.png" pageId="47" pageNumber="48" startId="47.[189,232,1930,1956]" targetBox="[223,1365,249,1879]" targetPageId="47">
<paragraph id="DD6736C4FFD989299FBCF83BFE4BF860" blockId="47.[189,1399,1930,2026]" pageId="47" pageNumber="48">
<emphasis id="EFACEAD6FFD989299FBCF83BFEE5F82E" bold="true" box="[189,284,1930,1956]" pageId="47" pageNumber="48">Fig. 17.</emphasis>
Map showing the distribution of
<emphasis id="EFACEAD6FFD989299DB6F83BFC6DF82E" box="[695,916,1930,1956]" italics="true" pageId="47" pageNumber="48">Onitisassamensis</emphasis>
<bibRefCitation id="B9494B35FFD989299C96F83BFBC0F82E" author="Biswas S." box="[919,1081,1930,1956]" pageId="47" pageNumber="48" pagination="338 - 342" refId="ref27446" refString="Biswas S. 1980. Studies on the scarab beetles (Coleoptera: Scarabaeidae) of North India Part II. Three new species and two new records from India. Journal of Bombay Natural History Society 76: 338 - 342. Available from: https: // www. biodiversitylibrary. org / part / 151549 [accessed 15 Oct. 2018]" type="journal article" year="1980">Biswas, 1980</bibRefCitation>
;
<taxonomicName id="1AD84D47FFD989299B4AF83BFAFEF82E" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[1099,1287,1930,1956]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bhomorensis" status="sp. nov.">
<emphasis id="EFACEAD6FFD989299B4AF83BFAFEF82E" box="[1099,1287,1930,1956]" italics="true" pageId="47" pageNumber="48">O. bhomorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFD989299A10F83BFA8BF82E" box="[1297,1394,1930,1956]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName id="1AD84D47FFD989299FBCF81CFDF1F84D" authority="Cambefort, 1988" authorityName="Cambefort" authorityYear="1988" box="[189,520,1965,1991]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bordati">
<emphasis id="EFACEAD6FFD989299FBCF81CFEC6F84D" box="[189,319,1965,1991]" italics="true" pageId="47" pageNumber="48">O. bordati</emphasis>
<bibRefCitation id="B9494B35FFD989299E41F81CFDF1F84D" author="Cambefort Y." box="[320,520,1965,1991]" pageId="47" pageNumber="48" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort, 1988</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="1AD84D47FFD989299D19F81CFC9CF842" authority="Lansberge, 1875" authorityName="Lansberge" authorityYear="1875" box="[536,869,1965,1992]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="brahma">
<emphasis id="EFACEAD6FFD989299D19F81CFD61F84D" box="[536,664,1965,1991]" italics="true" pageId="47" pageNumber="48">O. brahma</emphasis>
<bibRefCitation id="B9494B35FFD989299DA1F81CFC9CF842" author="Lansberge G." box="[672,869,1965,1992]" pageId="47" pageNumber="48" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="1AD84D47FFD989299CA1F81CFAD2F84D" authority="Redtenbacher, 1848" authorityName="Redtenbacher" authorityYear="1848" box="[928,1323,1965,1991]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="EFACEAD6FFD989299CA1F81CFBC0F84D" box="[928,1081,1965,1991]" italics="true" pageId="47" pageNumber="48">O. castaneus</emphasis>
Redtenbacher, 1848
</taxonomicName>
in the Indian Subcontinent.
</paragraph>
</caption>
<caption id="89A7664CFFC689369FBFF836FE04F862" ID-DOI="http://doi.org/10.5281/zenodo.13742903" ID-Zenodo-Dep="13742903" httpUri="https://zenodo.org/record/13742903/files/figure.png" pageId="48" pageNumber="49" startId="48.[190,233,1927,1953]" targetBox="[224,1364,251,1878]" targetPageId="48">
<paragraph id="DD6736C4FFC689369FBFF836FE04F862" blockId="48.[190,1400,1927,2024]" pageId="48" pageNumber="49">
<emphasis id="EFACEAD6FFC689369FBFF836FED8F82B" bold="true" box="[190,289,1927,1953]" pageId="48" pageNumber="49">Fig. 18.</emphasis>
Map showing the distribution of
<taxonomicName id="1AD84D47FFC689369DCAF839FBE2F828" authority="Sharp, 1875" authorityName="Sharp" authorityYear="1875" box="[715,1051,1927,1954]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="EFACEAD6FFC689369DCAF839FC8EF82B" box="[715,887,1928,1954]" italics="true" pageId="48" pageNumber="49">Onitis crassus</emphasis>
Sharp, 1875
</taxonomicName>
;
<taxonomicName id="1AD84D47FFC689369B2DF839FA8AF828" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[1068,1395,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="15" phylum="Arthropoda" rank="species" species="excavatus">
O.
<emphasis id="EFACEAD6FFC689369B57F839FB33F82B" box="[1110,1226,1928,1953]" italics="true" pageId="48" pageNumber="49">excavatus</emphasis>
<bibRefCitation id="B9494B35FFC689369BD7F839FA8AF828" author="Arrow G. J." box="[1238,1395,1928,1954]" pageId="48" pageNumber="49" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="1AD84D47FFC689369FBFF81AFE06F84F" authority="(Wulfen, 1789)" baseAuthorityName="Wulfen" baseAuthorityYear="1789" box="[190,511,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="falcatus">
<emphasis id="EFACEAD6FFC689369FBFF81AFEBFF84F" box="[190,326,1963,1989]" italics="true" pageId="48" pageNumber="49">O. falcatus</emphasis>
(Wulfen, 1789)
</taxonomicName>
;
<taxonomicName id="1AD84D47FFC689369D0AF81AFCEFF84F" authority="Felsche, 1907" authorityName="Felsche" authorityYear="1907" box="[523,790,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="feae">
<emphasis id="EFACEAD6FFC689369D0AF81AFD99F84F" box="[523,608,1963,1989]" italics="true" pageId="48" pageNumber="49">O. feae</emphasis>
<bibRefCitation id="B9494B35FFC689369D6EF81AFCEFF84F" author="Felsche C." box="[623,790,1963,1989]" pageId="48" pageNumber="49" pagination="273 - 296" refId="ref29092" refString="Felsche C. 1907. Coprophage Scarabaeiden. Deutsche Entomologische Zeitschrift: 273 - 296." type="book chapter" year="1907">Felsche, 1907</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="1AD84D47FFC689369C22F81AFB89F84F" authority="(Pallas, 1771)" baseAuthorityName="Pallas" baseAuthorityYear="1771" box="[803,1136,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="humerosus">
<emphasis id="EFACEAD6FFC689369C22F81AFC3CF84F" box="[803,965,1963,1989]" italics="true" pageId="48" pageNumber="49">O. humerosus</emphasis>
(
<bibRefCitation id="B9494B35FFC689369CD4F81AFB9EF84F" author="Pallas P. S." box="[981,1127,1963,1989]" pageId="48" pageNumber="49" refId="ref30468" refString="Pallas P. S. 1771. Reise durch verschiedene Provinzen des Russischen Reiches in den Jahren 1768 - 1774 I: 1 - 504 (453 - 504). Petersburg Akademische Buchhandlung, St Petersburg." type="book" year="1771">Pallas, 1771</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="1AD84D47FFC689369BABF81AFAEFF84F" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[1194,1302,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="kethai" status="sp. nov.">
<emphasis id="EFACEAD6FFC689369BABF81AFAEFF84F" box="[1194,1302,1963,1989]" italics="true" pageId="48" pageNumber="49">O. kethai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFC689369A1CF81AFA81F84F" box="[1309,1400,1963,1989]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
in the Indian Subcontinent.
</paragraph>
</caption>
<caption id="89A7664CFFC789379FBCF836FE02F862" ID-DOI="http://doi.org/10.5281/zenodo.13742907" ID-Zenodo-Dep="13742907" httpUri="https://zenodo.org/record/13742907/files/figure.png" pageId="49" pageNumber="50" startId="49.[189,232,1927,1953]" targetBox="[227,1362,257,1876]" targetPageId="49">
<paragraph id="DD6736C4FFC789379FBCF836FE02F862" blockId="49.[189,1399,1927,2024]" pageId="49" pageNumber="50">
<emphasis id="EFACEAD6FFC789379FBCF836FEE2F82B" bold="true" box="[189,283,1927,1953]" pageId="49" pageNumber="50">Fig. 19.</emphasis>
Map showing the distribution of
<emphasis id="EFACEAD6FFC789379DAAF839FCCFF828" box="[683,822,1928,1954]" italics="true" pageId="49" pageNumber="50">Onitislama</emphasis>
<bibRefCitation id="B9494B35FFC789379C3EF839FBFFF828" author="Lansberge G." box="[831,1030,1927,1954]" pageId="49" pageNumber="50" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
;
<taxonomicName id="1AD84D47FFC789379B15F839FA8BF828" authority="Cambefort, 1988" authorityName="Cambefort" authorityYear="1988" box="[1044,1394,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="naviauxi">
<emphasis id="EFACEAD6FFC789379B15F839FADAF828" box="[1044,1315,1928,1954]" italics="true" pageId="49" pageNumber="50">
O. naviauxi
<bibRefCitation id="B9494B35FFC789379BA7F839FADAF828" author="Cambefort Y." box="[1190,1315,1928,1954]" pageId="49" pageNumber="50" pagination="189 - 192" refId="ref28159" refString="Cambefort Y. 1988. Deux nouveaux Onitis d'Asie tropicale. Bulletin de la Societe entomologique de France, Paris 92 (5 - 6): 189 - 192. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1987 _ num _ 92 _ 5 _ 17495 [accessed 10 May 2017]" type="journal article" year="1988">Cambefort</bibRefCitation>
</emphasis>
, 1988
</taxonomicName>
;
<taxonomicName id="1AD84D47FFC789379FBCF81AFDEBF84F" authority="Fabricius, 1801" authorityName="Fabricius" authorityYear="1801" box="[189,530,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="philemon">
<emphasis id="EFACEAD6FFC789379FBCF81AFEB6F84F" box="[189,335,1963,1989]" italics="true" pageId="49" pageNumber="50">O. philemon</emphasis>
<bibRefCitation id="B9494B35FFC789379E56F81AFDEBF84F" author="Fabricius J. C." box="[343,530,1963,1989]" pageId="49" pageNumber="50" refId="ref29058" refString="Fabricius J. C. 1801. Systema Eleutheratorum secundum ordines, genera, species adjectis synonymis, locis, observationibus, descriptionibus. Volume 2. Impensis Bibliopolii novi academici, Kiel." type="book" year="1801">Fabricius, 1801</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="1AD84D47FFC789379D1EF81AFC43F84F" authority="Janssens, 1937" authorityName="Janssens" authorityYear="1937" box="[543,954,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="punctatostriatus">
<emphasis id="EFACEAD6FFC789379D1EF81AFCFAF84F" box="[543,771,1963,1989]" italics="true" pageId="49" pageNumber="50">O. punctatostriatus</emphasis>
<bibRefCitation id="B9494B35FFC789379C0BF81AFC43F84F" author="Janssens A." box="[778,954,1963,1989]" pageId="49" pageNumber="50" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens, 1937</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="1AD84D47FFC789379CF4F81AFA8EF84C" authority="Lansberge, 1875" authorityName="Lansberge, AIMB" authorityYear="1875" box="[1013,1399,1963,1990]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="singhalensis">
<emphasis id="EFACEAD6FFC789379CF4F81AFB53F84F" box="[1013,1194,1963,1989]" italics="true" pageId="49" pageNumber="50">O. singhalensis</emphasis>
<bibRefCitation id="B9494B35FFC789379BB2F81AFA8EF84C" author="Lansberge G." box="[1203,1399,1963,1990]" pageId="49" pageNumber="50" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
</taxonomicName>
in the Indian Subcontinent.
</paragraph>
</caption>
<caption id="89A7664CFFC489349FBCF83BFC1DF84D" ID-DOI="http://doi.org/10.5281/zenodo.13742911" ID-Zenodo-Dep="13742911" httpUri="https://zenodo.org/record/13742911/files/figure.png" pageId="50" pageNumber="51" startId="50.[189,232,1930,1956]" targetBox="[230,1361,265,1880]" targetPageId="50">
<paragraph id="DD6736C4FFC489349FBCF83BFC1DF84D" blockId="50.[189,1399,1930,1992]" pageId="50" pageNumber="51">
<emphasis id="EFACEAD6FFC489349FBCF83BFEE2F82E" bold="true" box="[189,283,1930,1956]" pageId="50" pageNumber="51">Fig. 20.</emphasis>
Map showing the distribution of
<emphasis id="EFACEAD6FFC489349DA2F83BFCD8F82E" box="[675,801,1930,1956]" italics="true" pageId="50" pageNumber="51">Onitissiva</emphasis>
<bibRefCitation id="B9494B35FFC489349C28F83BFC4EF82E" author="Gillet J. J. E." box="[809,951,1930,1956]" pageId="50" pageNumber="51" pagination="313 - 314" refId="ref29108" refString="Gillet J. J. E. 1911. Coprides nouveaux de la region orientale (etc.). Annales de la Societe entomologique de Belgique LV: 313 - 314." type="book chapter" year="1911">Gillet, 1911</bibRefCitation>
;
<taxonomicName id="1AD84D47FFC489349CC2F83BFAF9F82E" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[963,1280,1930,1956]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="EFACEAD6FFC489349CC2F83BFB9AF82E" box="[963,1123,1930,1956]" italics="true" pageId="50" pageNumber="51">O. subopacus</emphasis>
<bibRefCitation id="B9494B35FFC489349B68F83BFAF9F82E" author="Arrow G. J." box="[1129,1280,1930,1956]" pageId="50" pageNumber="51" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="1AD84D47FFC489349A0DF83BFE79F842" authority="Lansberge, 1875" authorityName="Lansberge" authorityYear="1875" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFC489349A0DF83BFA8EF82E" box="[1292,1399,1930,1956]" italics="true" pageId="50" pageNumber="51">O. virens</emphasis>
<bibRefCitation id="B9494B35FFC489349FBCF81CFE79F842" author="Lansberge G." box="[189,384,1965,1992]" pageId="50" pageNumber="51" pagination="1 - 200" refId="ref29884" refString="Lansberge G. 1875. Monographie des Onitides. Annales de la Societe entomologique de Belgique 11 (2): 1 - 200." type="journal article" year="1875">Lansberge, 1875</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="1AD84D47FFC489349EB8F81CFDC4F84D" authorityName="Karimbumkara &amp; Priyadarsanan" authorityYear="2024" box="[441,573,1965,1991]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="visthara" status="sp. nov.">
<emphasis id="EFACEAD6FFC489349EB8F81CFDC4F84D" box="[441,573,1965,1991]" italics="true" pageId="50" pageNumber="51">O. visthara</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F49F57ADFFC489349D45F81FFD66F842" box="[580,671,1966,1992]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
in the Indian Subcontinent.
</paragraph>
</caption>
<paragraph id="DD6736C4FFC589359FBCFEB8FC38FECC" blockId="51.[189,1398,265,327]" pageId="51" pageNumber="52">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFC589359EB2FEB8FDB2FEA9" box="[435,587,265,291]" captionStart="Fig" captionStartId="36.[189,232,1964,1990]" captionTargetBox="[275,1278,1055,1924]" captionTargetId="figure-228@36.[189,1398,1033,1927]" captionTargetPageId="36" captionText="Fig. 13. Aedeagus, lateral, dorsal, ventral and apical views. AD. Onitissubopacus Arrow, 1931, AIMB/ Co/Sc1000711. EH. O. virens Lansberge, 1875, AIMB/Co/Sc1000765. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742887" httpUri="https://zenodo.org/record/13742887/files/figure.png" pageId="51" pageNumber="52">Fig. 13E H</figureCitation>
). LP=3.5, Lp=3; BP=1.6, BpB=0.7, BpT=0.5. Phallobase slightly longer than the paramere, both straight below and sharp at the tip.
</paragraph>
<paragraph id="DD6736C4FFC589359FEBFEDFFEBCFE02" blockId="51.[189,1133,366,427]" box="[234,325,366,392]" pageId="51" pageNumber="52">
<emphasis id="EFACEAD6FFC589359FEBFEDFFEBCFE02" bold="true" box="[234,325,366,392]" pageId="51" pageNumber="52">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFC589359FBCFE20FB94FE21" blockId="51.[189,1133,366,427]" box="[189,1133,401,427]" pageId="51" pageNumber="52">Protibia short, broad with four stout teeth, the mid and hind femur without teeth.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFC589359FBCFE63FE3FFD64" pageId="51" pageNumber="52" type="distribution">
<paragraph id="DD6736C4FFC589359FBCFE63FEAFFE66" blockId="51.[189,342,466,492]" box="[189,342,466,492]" pageId="51" pageNumber="52">
<heading id="862F81A8FFC589359FBCFE63FEAFFE66" bold="true" box="[189,342,466,492]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="EFACEAD6FFC589359FBCFE63FEAFFE66" bold="true" box="[189,342,466,492]" pageId="51" pageNumber="52">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFC589359FBCFDB0FE3FFD64" blockId="51.[189,1399,513,750]" pageId="51" pageNumber="52">
<collectingCountry id="A5CF7654FFC589359FBCFDB0FF04FD91" box="[189,253,513,539]" name="India" pageId="51" pageNumber="52">India</collectingCountry>
:
<collectingRegion id="1F1CF826FFC589359E09FDB0FE10FD91" box="[264,489,513,539]" country="India" name="Arunachal Pradesh" pageId="51" pageNumber="52">Arunachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359EF5FDB0FDBFFD91" box="[500,582,513,539]" country="India" name="Assam" pageId="51" pageNumber="52">Assam</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359D52FDB0FD6CFD91" box="[595,661,513,539]" country="India" name="Bihar" pageId="51" pageNumber="52">Bihar</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359DA2FDB0FCD5FD91" box="[675,812,513,539]" country="India" name="Chhattisgarh" pageId="51" pageNumber="52">Chattisgarh</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359C38FDB0FC6AFD91" box="[825,915,513,539]" country="India" name="Gujarat" pageId="51" pageNumber="52">Gujarat</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359CA1FDB0FBFCFD91" box="[928,1029,513,539]" country="India" name="Haryana" pageId="51" pageNumber="52">Haryana</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359B13FDB0FB12FD91" box="[1042,1259,513,539]" country="India" name="Himachal Pradesh" pageId="51" pageNumber="52">Himachal Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359BF9FDB0FA8BFD91" box="[1272,1394,513,539]" country="India" name="Karnataka" pageId="51" pageNumber="52">Karnataka</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359FBCFD95FEF4FDB4" box="[189,269,548,574]" country="India" name="Kerala" pageId="51" pageNumber="52">Kerala</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359E1AFD95FE1AFDB4" box="[283,483,548,574]" country="India" name="Madhya Pradesh" pageId="51" pageNumber="52">Madhya Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359EF0FD95FD7CFDB4" box="[497,645,548,574]" country="India" name="Maharashtra" pageId="51" pageNumber="52">Maharashtra</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359D92FD95FCEEFDB4" box="[659,791,548,574]" country="India" name="Meghalaya" pageId="51" pageNumber="52">Meghalaya</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359C24FD95FC6EFDB4" box="[805,919,548,574]" country="India" name="Nagaland" pageId="51" pageNumber="52">Nagaland</collectingRegion>
, New
<collectingRegion id="1F1CF826FFC589359CE7FD95FBD0FDB4" box="[998,1065,548,574]" country="India" name="Delhi" pageId="51" pageNumber="52">Delhi</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359B39FD95FB74FDB4" box="[1080,1165,548,574]" country="India" name="Odisha" pageId="51" pageNumber="52">Odisha</collectingRegion>
, Punjab,
<collectingRegion id="1F1CF826FFC589359BFCFD95FA8AFDB4" box="[1277,1395,548,574]" country="India" name="Rajasthan" pageId="51" pageNumber="52">Rajasthan</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359FBCFDF6FEEDFDEB" box="[189,276,583,609]" country="India" name="Sikkim" pageId="51" pageNumber="52">Sikkim</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359E29FDF6FE44FDEB" box="[296,445,583,609]" country="India" name="Tamil Nadu" pageId="51" pageNumber="52">Tamil Nadu</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359ED1FDF6FDD0FDEB" box="[464,553,583,609]" country="India" name="Tripura" pageId="51" pageNumber="52">Tripura</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359D3FFDF6FD28FDEB" box="[574,721,583,609]" country="India" name="Uttarakhand" pageId="51" pageNumber="52">Uttarakhand</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359DE5FDF9FC76FDEB" box="[740,911,583,610]" country="India" name="Uttar Pradesh" pageId="51" pageNumber="52">Uttar Pradesh</collectingRegion>
,
<collectingRegion id="1F1CF826FFC589359CA2FDF9FBBBFDEB" box="[931,1090,583,610]" country="India" name="West Bengal" pageId="51" pageNumber="52">West Bengal</collectingRegion>
. Other countries:
<collectingCountry id="A5CF7654FFC589359A2BFDF6FA8BFDEB" box="[1322,1394,583,609]" name="China" pageId="51" pageNumber="52">China</collectingCountry>
,
<collectingCountry id="A5CF7654FFC589359FBCFDDBFEBEFD0E" box="[189,327,618,644]" name="Bangladesh" pageId="51" pageNumber="52">Bangladesh</collectingCountry>
,
<collectingCountry id="A5CF7654FFC589359E54FDDAFE69FD0F" box="[341,400,619,645]" name="Laos" pageId="51" pageNumber="52">Laos</collectingCountry>
,
<collectingCountry id="A5CF7654FFC589359E9FFDDAFDE8FD0F" box="[414,529,619,645]" name="Myanmar" pageId="51" pageNumber="52">Myanmar</collectingCountry>
,
<collectingCountry id="A5CF7654FFC589359D1EFDDBFD9EFD0E" box="[543,615,618,644]" name="Nepal" pageId="51" pageNumber="52">Nepal</collectingCountry>
,
<collectingCountry id="A5CF7654FFC589359D74FDDBFD23FD0E" box="[629,730,618,644]" name="Pakistan" pageId="51" pageNumber="52">Pakistan</collectingCountry>
,
<collectingCountry id="A5CF7654FFC589359DE6FDDBFCA8FD0E" box="[743,849,618,644]" name="Thailand" pageId="51" pageNumber="52">Thailand</collectingCountry>
,
<collectingCountry id="A5CF7654FFC589359C5EFDDBFC38FD0E" box="[863,961,618,644]" name="Vietnam" pageId="51" pageNumber="52">Vietnam</collectingCountry>
(
<bibRefCitation id="B9494B35FFC589359CD3FDDBFB87FD0E" author="Chandra K." box="[978,1150,618,644]" pageId="51" pageNumber="52" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000</bibRefCitation>
;
<bibRefCitation id="B9494B35FFC589359B8DFDDBFF03FD22" author="Chandra K. &amp; Ahirwar S. C." pageId="51" pageNumber="52" pagination="147 - 155" refId="ref28418" refString="Chandra K. &amp; Ahirwar S. C. 2005. Scarabaeid Beetles (Coleoptera) of Kanha Tiger Reserve, Madhya Pradesh. Records Zoological Survey of India 105 (1 - 2): 147 - 155." type="journal article" year="2005">Chandra &amp; Ahirwar 2005</bibRefCitation>
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<bibRefCitation id="B9494B35FFC589359E0AFD3FFEB0FD22" author="Chandra K. &amp; Ahirwar S. C." box="[267,329,654,680]" pageId="51" pageNumber="52" pagination="273 - 300" refId="ref28458" refString="Chandra K. &amp; Ahirwar S. C. 2007. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Madhya Pradesh (Including Chhattisgarh), State Fauna Series 15 (Part 1): 273 - 300." type="book chapter" year="2007">2007</bibRefCitation>
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<bibRefCitation id="B9494B35FFC589359E5BFD3CFD91FD2D" author="Bezdek A. &amp; Krell F. - T." box="[346,616,653,680]" pageId="51" pageNumber="52" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006</bibRefCitation>
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<bibRefCitation id="B9494B35FFC589359D79FD3CFCE4FD22" author="Sewak R." box="[632,797,653,680]" pageId="51" pageNumber="52" pagination="1 - 48" refId="ref31194" refString="Sewak R. 2009 a. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Thar desert of Gujarat. Records of the Zoological Survey of India, Occassional Paper No 295: 1 - 48." type="journal article" year="2009">Sewak 2009a</bibRefCitation>
,
<bibRefCitation id="B9494B35FFC589359C2FFD3CFC83FD2D" author="Sewak R." box="[814,890,653,679]" pageId="51" pageNumber="52" pagination="1 - 106" refId="ref31232" refString="Sewak R. 2009 b. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Rajasthan. Records of the Zoological Survey of India, Occassional Paper No 296: 1 - 106." type="journal article" year="2009">2009b</bibRefCitation>
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Sabu
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2012a;
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;
<bibRefCitation id="B9494B35FFC589359D26FD00FD0FFD46" author="Gupta D. &amp; Chandra K. &amp; Khan S." box="[551,758,689,716]" pageId="51" pageNumber="52" pagination="1025 - 1039" refId="ref29273" refString="Gupta D., Chandra K. &amp; Khan S. 2015. Studies on Tribe Onitini (Scarabaeinae: Scarabaeidae: Coleoptera) from Madhya Pradesh with a Checklist from India. Proceedings of the National Academy of Sciences, India Section B: Biological Sciences volume 87: 1025 - 1039. https: // doi. org / 10.1007 / s 40011 - 015 - 0676 - 9" type="journal article" year="2015">
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).
</paragraph>
</subSubSection>
</treatment>
</document>