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<mods:title id="5F439C784C0EC08E185D7F669F795FCC">Study on the genus Onitis (Coleoptera: Scarabaeidae: Scarabaeinae) of the Indian Subcontinent, with three new species from India</mods:title>
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<mods:namePart id="6F4134C747732F954B916EF6CA2C12ED">Karimbumkara, Seena Narayanan</mods:namePart>
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<mods:namePart id="50E164C3E59914E511DE889D25A9E9D3">Priyadarsanan, Dharma Rajan</mods:namePart>
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<taxonomicName id="1AD84D47FFDA892A9D65FDD4FC29FDF5" ID-CoL="49KLJ" authority="Arrow, 1931" authorityName="Arrow" authorityYear="1931" box="[612,976,612,639]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="EFACEAD6FFDA892A9D65FDD4FCCDFDF5" bold="true" box="[612,820,612,639]" italics="true" pageId="44" pageNumber="45">Onitis subopacus</emphasis>
<bibRefCitation id="B9494B35FFDA892A9C3AFDD4FC29FDF5" author="Arrow G. J." box="[827,976,613,639]" pageId="44" pageNumber="45" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931</bibRefCitation>
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<figureCitation id="45E32A41FFDA892A9D9DFD39FCF0FD28" box="[668,777,648,674]" captionStart="Fig" captionStartId="34.[189,231,1828,1854]" captionTargetBox="[309,1270,282,1772]" captionTargetId="figure-77@34.[189,1398,262,1780]" captionTargetPageId="34" captionText="Fig. 12.A. Onitissinghalensis Lansberge, 1875, holotype, ♂, MNHN EC4812, (image:Antoine Mantilleri, MNHN.) B. Onitissiva Gillet, 1911, holotype, ♀, (image: Jonathan Brecko, IRSNB). C. O. subopacus Arrow, 1931, holotype, ♂, BMNH(E)1237174, (image: BMNH). D. O.virens Lansberge, 1875, paralectotype, ♂, (image: Jonathan Brecko, IRSNB). E–F Onitisvisthara sp. nov., holotype, ♂, NIM/ NBAIR/COL/ONIT/H-271023C (images: AIMB). E. Habitus, dorsal view. F. Habitus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13742883" httpUri="https://zenodo.org/record/13742883/files/figure.png" pageId="44" pageNumber="45">Figs 12C</figureCitation>
, 
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, 
<figureCitation id="45E32A41FFDA892A9C7BFD39FC6EFD28" box="[890,919,648,674]" captionStart="Fig" captionStartId="50.[189,232,1930,1956]" captionTargetBox="[230,1361,265,1880]" captionTargetId="figure-1@50.[214,1374,251,1892]" captionTargetPageId="50" captionText="Fig. 20. Map showing the distribution of Onitissiva Gillet, 1911; O. subopacus Arrow, 1931; O. virens Lansberge, 1875 and O. visthara sp. nov. in the Indian Subcontinent." figureDoi="http://doi.org/10.5281/zenodo.13742911" httpUri="https://zenodo.org/record/13742911/files/figure.png" pageId="44" pageNumber="45">20</figureCitation>
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<emphasis id="EFACEAD6FFDA892A9FBCFD7BFE71FD6E" box="[189,392,714,740]" italics="true" pageId="44" pageNumber="45">Onitissubopacus</emphasis>
<bibRefCitation id="B9494B35FFDA892A9E8FFD7BFDA4FD6E" author="Arrow G. J." box="[398,605,713,740]" pageId="44" pageNumber="45" refId="ref26985" refString="Arrow G. J. 1931. The fauna of British India including Ceylon and Burma, Coleoptera Lamellicornia Part III, (Coprinae). Taylor &amp; Francis, London." type="book" year="1931">Arrow, 1931: 395</bibRefCitation>
(original description).
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<taxonomicName id="1AD84D47FFDA892A9FBCFCBAFE75FCAF" ID-CoL="49KLJ" authorityName="Arrow" authorityYear="1931" box="[189,396,779,805]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="EFACEAD6FFDA892A9FBCFCBAFE75FCAF" box="[189,396,779,805]" italics="true" pageId="44" pageNumber="45">Onitis subopacus</emphasis>
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– 
<treatmentCitation id="5C7910D5FFDA892A9EB0FCBAFD6EFCAF" author="Balthasar V." box="[433,663,779,806]" page="94" pageId="44" pageNumber="45" year="1935">
<bibRefCitation id="B9494B35FFDA892A9EB0FCBAFD6EFCAF" author="Balthasar V." box="[433,663,779,806]" pageId="44" pageNumber="45" refId="ref27158" refString="Balthasar V. 1935. Scarabaeidae des palaarktischen Faunengebietes. Monographische Bestimmungstabelle. I. Coprinae I. Teil. Scarabaeini, Sisyphini, Panelini, Coprini, Onitini, Oniticellini. Edm. Reitter's Nachf. Emmerich Reitter, Troppau." type="book" year="1935">Balthasar 1935: 94</bibRefCitation>
</treatmentCitation>
(monograph); 1963: 53 (monograph). — 
<treatmentCitation id="5C7910D5FFDA892A9B9AFCBDFA8FFCAC" author="Janssens A." box="[1179,1398,779,806]" page="51" pageId="44" pageNumber="45" year="1937">
<bibRefCitation id="B9494B35FFDA892A9B9AFCBDFA8FFCAC" author="Janssens A." box="[1179,1398,779,806]" pageId="44" pageNumber="45" refId="ref29633" refString="Janssens A. 1937. Revision des Onitides. Deuxieme serie, fascicule 11. Memoires du Musee royal d'Histoire naturelle de Belgique, Brussels 11 (2)." type="journal volume" year="1937">Janssens 1937: 51</bibRefCitation>
</treatmentCitation>
(revision). — 
<treatmentCitation id="5C7910D5FFDA892A9E95FC9FFCF3FCC2" author="Biswas S. &amp; Chatterjee S. K." box="[404,778,814,840]" page="67" pageId="44" pageNumber="45" year="1986">
<bibRefCitation id="B9494B35FFDA892A9E95FC9FFCF3FCC2" author="Biswas S. &amp; Chatterjee S. K." box="[404,778,814,840]" pageId="44" pageNumber="45" pagination="57 - 67" refId="ref27582" refString="Biswas S. &amp; Chatterjee S. K. 1986 a. Dung beetle fauna (Coleoptera: Scarabaeidae: Scarabaeinae) of Palamou Tiger Reserve, Bihar, with description of a new species. Records of the Zoological Survey of India 83 (3 - 4): 57 - 67. https: // doi. org / 10.26515 / rzsi / v 83 / i 3 - 4 / 1986 / 161311" type="journal article" year="1986">Biswas &amp; Chatterjee 1986a: 67</bibRefCitation>
</treatmentCitation>
(distribution). — 
<treatmentCitation id="5C7910D5FFDA892A9CE4FC9FFB33FCC2" author="Chandra K." box="[997,1226,814,840]" page="360" pageId="44" pageNumber="45" year="2000">
<bibRefCitation id="B9494B35FFDA892A9CE4FC9FFB33FCC2" author="Chandra K." box="[997,1226,814,840]" pageId="44" pageNumber="45" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000: 360</bibRefCitation>
</treatmentCitation>
(distribution); 2005: 150 (distribution). — 
<treatmentCitation id="5C7910D5FFDA892A9D39FCE0FD09FCE1" author="Mittal I. C." box="[568,752,849,876]" page="46" pageId="44" pageNumber="45" year="2005">
<bibRefCitation id="B9494B35FFDA892A9D39FCE0FD09FCE1" author="Mittal I. C." box="[568,752,849,876]" pageId="44" pageNumber="45" pagination="43 - 51" refId="ref30210" refString="Mittal I. C. 2005. Diversity and conservation status of dung beetles (Laparosticti: Scarabaeidae: Coleoptera) in North India. Bulletin of National Institute of Ecology 15: 43 - 51." type="journal article" year="2005">Mittal 2005: 46</bibRefCitation>
</treatmentCitation>
(list). — 
<treatmentCitation id="5C7910D5FFDA892A9C65FCE0FB35FCE6" author="Chandra K. &amp; Ahirwar S. C." box="[868,1228,849,876]" page="149" pageId="44" pageNumber="45" year="2005">
<bibRefCitation id="B9494B35FFDA892A9C65FCE0FB35FCE6" author="Chandra K. &amp; Ahirwar S. C." box="[868,1228,849,876]" pageId="44" pageNumber="45" pagination="147 - 155" refId="ref28418" refString="Chandra K. &amp; Ahirwar S. C. 2005. Scarabaeid Beetles (Coleoptera) of Kanha Tiger Reserve, Madhya Pradesh. Records Zoological Survey of India 105 (1 - 2): 147 - 155." type="journal article" year="2005">Chandra &amp; Ahirwar 2005: 149</bibRefCitation>
</treatmentCitation>
(distribution). — 
<treatmentCitation id="5C7910D5FFDA892A9E0FFCC4FDBAFC05" author="Bezdek A. &amp; Krell F. - T." box="[270,579,884,911]" page="159" pageId="44" pageNumber="45" year="2006">
<bibRefCitation id="B9494B35FFDA892A9E0FFCC4FDBAFC05" author="Bezdek A. &amp; Krell F. - T." box="[270,579,884,911]" pageId="44" pageNumber="45" pagination="158 - 159" refId="ref27342" refString="Bezdek A. &amp; Krell F. - T. 2006. Tribe Onitini. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. 158 - 159. Apollo Books, Stenstrup." type="book chapter" year="2006">Bezděk &amp; Krell 2006: 159</bibRefCitation>
</treatmentCitation>
(distribution). — 
<treatmentCitation id="5C7910D5FFDA892A9C17FCC4FC08FC05" author="Sabu T. K. &amp; Vinod K. V. &amp; Vineesh P. J." box="[790,1009,884,911]" page="5" pageId="44" pageNumber="45" year="2006">
<bibRefCitation id="B9494B35FFDA892A9C17FCC4FC08FC05" author="Sabu T. K. &amp; Vinod K. V. &amp; Vineesh P. J." box="[790,1009,884,911]" pageId="44" pageNumber="45" pagination="1 - 17" refId="ref30840" refString="Sabu T. K., Vinod K. V. &amp; Vineesh P. J. 2006. Guild structure, diversity and succession of dung beetles associated with Indian elephant dung in South Western Ghats forests. Journal of Insect Science 6 (1): 1 - 17. https: // doi. org / 10.1673 / 2006 _ 06 _ 17.1" type="journal article" year="2006">
Sabu 
<emphasis id="EFACEAD6FFDA892A9C59FCC4FC6BFC05" box="[856,914,885,911]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2006: 5
</bibRefCitation>
</treatmentCitation>
(list); 2011: 34 (list). — 
<treatmentCitation id="5C7910D5FFDA892A9A12FCC4FE1EFC38" author="Chandra K. &amp; Ahirwar S. C." page="276" pageId="44" pageNumber="45" year="2007">
<bibRefCitation id="B9494B35FFDA892A9A12FCC4FE1EFC38" author="Chandra K. &amp; Ahirwar S. C." pageId="44" pageNumber="45" pagination="273 - 300" refId="ref28458" refString="Chandra K. &amp; Ahirwar S. C. 2007. Insecta: Coleoptera: Scarabaeidae. Zoological Survey of India, Fauna of Madhya Pradesh (Including Chhattisgarh), State Fauna Series 15 (Part 1): 273 - 300." type="book chapter" year="2007">Chandra &amp; Ahirwar 2007: 276</bibRefCitation>
</treatmentCitation>
(distribution). — 
<treatmentCitation id="5C7910D5FFDA892A9DBFFC29FC73FC38" author="Sewak R." box="[702,906,920,946]" page="60" pageId="44" pageNumber="45" year="2009">
<bibRefCitation id="B9494B35FFDA892A9DBFFC29FC73FC38" author="Sewak R." box="[702,906,920,946]" pageId="44" pageNumber="45" pagination="1 - 106" refId="ref31232" refString="Sewak R. 2009 b. Dung beetles (Coleoptera: Scarabaeidae: Coprinae) of Rajasthan. Records of the Zoological Survey of India, Occassional Paper No 296: 1 - 106." type="journal article" year="2009">Sewak 2009b: 60</bibRefCitation>
</treatmentCitation>
(key, diagnosis, distribution). — 
<treatmentCitation id="5C7910D5FFDA892A9A17FC29FE5DFC5C" author="Thakare V. &amp; Zade V. &amp; Chandra K. &amp; Gupta D." page="75" pageId="44" pageNumber="45" year="2012">
<bibRefCitation id="B9494B35FFDA892A9A17FC29FE5DFC5C" author="Thakare V. &amp; Zade V. &amp; Chandra K. &amp; Gupta D." pageId="44" pageNumber="45" pagination="73 - 80" refId="ref31359" refString="Thakare V., Zade V., Chandra K. &amp; Gupta D. 2012. Scarab beetles (Coleoptera: Scarabaeoidea) of Melghat Tiger Reserve, Central India. Academic Journal of Entomology 5 (2): 73 - 80." type="journal article" year="2012">
Thakare 
<emphasis id="EFACEAD6FFDA892A9FEBFC0DFED2FC5F" box="[234,299,955,981]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2012: 75
</bibRefCitation>
</treatmentCitation>
(distribution). — 
<treatmentCitation id="5C7910D5FFDA892A9D8BFC0AFC14FC5F" author="Chandra K. &amp; Gupta D." box="[650,1005,955,981]" page="274" pageId="44" pageNumber="45" year="2011">
<bibRefCitation id="B9494B35FFDA892A9D8BFC0AFC14FC5F" author="Chandra K. &amp; Gupta D." box="[650,1005,955,981]" pageId="44" pageNumber="45" pagination="272 - 278" refId="ref28502" refString="Chandra K. &amp; Gupta D. 2011. Study of scarabaeid beetles (Coleoptera) of Veerangana Durgavati Wildlife Sanctuary, Damoh, Madhya Pradesh, India. Deccan Current Science 5: 272 - 278." type="journal article" year="2011">Chandra &amp; Gupta 2011: 274</bibRefCitation>
</treatmentCitation>
(distribution); 2012a: 889 (list) Chandra 
<emphasis id="EFACEAD6FFDA892A9E55FC6EFE77FC72" box="[340,398,990,1016]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2012a: 52 (list); 2012b: 33 (distribution). — 
<treatmentCitation id="5C7910D5FFDA892A9C9BFC6FFB09FC72" author="Chandra K. &amp; Gupta D." box="[922,1264,990,1016]" page="349" pageId="44" pageNumber="45" year="2013">
<bibRefCitation id="B9494B35FFDA892A9C9BFC6FFB09FC72" author="Chandra K. &amp; Gupta D." box="[922,1264,990,1016]" pageId="44" pageNumber="45" pagination="331 - 360" refId="ref28634" refString="Chandra K. &amp; Gupta D. 2013 a. Taxonomic studies on dung beetles (Coleoptera: Scarabaeidae, Geotrupidae, Hybosoridae) of Chhattisgarh, India. Munis Entomology and Zoology 8 (1): 331 - 360." type="journal article" year="2013">Chandra &amp; Gupta 2013a: 349</bibRefCitation>
</treatmentCitation>
(diagnosis, distribution); 2013b: 4667 (distribution). — Chandra 
<emphasis id="EFACEAD6FFDA892A9C67FBB3FC5BFB91" box="[870,930,1025,1051]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2011: 67 (distribution). — 
<treatmentCitation id="5C7910D5FFDA892A9BEAFBB3FE98FBB5" author="Gupta D. &amp; Chandra K. &amp; Khan S." page="230" pageId="44" pageNumber="45" year="2014">
<bibRefCitation id="B9494B35FFDA892A9BEAFBB3FE98FBB5" author="Gupta D. &amp; Chandra K. &amp; Khan S." pageId="44" pageNumber="45" pagination="225 - 240" refId="ref29198" refString="Gupta D., Chandra K. &amp; Khan S. 2014. An updated checklist of Scarabaeoid beetles (Coleoptera: Scarabaeoidea) of Pench Tiger Reserve, Madhya Pradesh, India. Journal of Entomology and Zoology Studies 2 (5): 225 - 240. Available from https: // www. entomoljournal. com / vol 2 Issue 5 / 57.1. html [accessed 12 Jun. 2024]" type="journal article" year="2014">
Gupta 
<emphasis id="EFACEAD6FFDA892A9A3AFBB3FA8EFB91" box="[1339,1399,1025,1051]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2014: 230
</bibRefCitation>
</treatmentCitation>
(distribution); 2015: 1037 (list, distribution). — 
<treatmentCitation id="5C7910D5FFDA892A9CA7FB94FB35FBB5" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[934,1228,1061,1087]" page="300" pageId="44" pageNumber="45" year="2014">
<bibRefCitation id="B9494B35FFDA892A9CA7FB94FB35FBB5" author="Siddiqui H. &amp; Ahmed Z. &amp; Khatri I." box="[934,1228,1061,1087]" pageId="44" pageNumber="45" pagination="295 - 307" refId="ref31267" refString="Siddiqui H., Ahmed Z. &amp; Khatri I. 2014. Distributional notes and new records for the dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Pakistan. Pakistan Journal of Zoology 46 (2): 295 - 307." type="journal article" year="2014">
Siddiqui 
<emphasis id="EFACEAD6FFDA892A9B10FB94FBB4FBB5" box="[1041,1101,1061,1087]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2014: 300
</bibRefCitation>
</treatmentCitation>
(distribution). — 
<treatmentCitation id="5C7910D5FFDA892A9E13FBF9FE14FBE8" author="Bezdek A." box="[274,493,1096,1122]" page="179" pageId="44" pageNumber="45" year="2016">
<bibRefCitation id="B9494B35FFDA892A9E13FBF9FE14FBE8" author="Bezdek A." box="[274,493,1096,1122]" pageId="44" pageNumber="45" pagination="177 - 180" refId="ref27284" refString="Bezdek A. 2016. Scarabaeidae, subfamily Scarabaeinae, tribe Onitini. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 177 - 180. Brill, Leiden, Boston." type="book chapter" year="2016">Bezděk 2016: 179</bibRefCitation>
</treatmentCitation>
(distribution). — 
<treatmentCitation id="5C7910D5FFDA892A9DCBFBF9FC18FBE8" author="Ghosh J. &amp; Das, P. &amp; Ghosh S. K. &amp; Bhunia D. &amp; Kushwaha R. K. &amp; Gupta D. &amp; Chandra K." box="[714,993,1096,1122]" page="239" pageId="44" pageNumber="45" year="2020">
<bibRefCitation id="B9494B35FFDA892A9DCBFBF9FC18FBE8" author="Ghosh J. &amp; Das, P. &amp; Ghosh S. K. &amp; Bhunia D. &amp; Kushwaha R. K. &amp; Gupta D. &amp; Chandra K." box="[714,993,1096,1122]" pageId="44" pageNumber="45" pagination="221 - 275" refId="ref29141" refString="Ghosh J., Das, P., Ghosh S. K., Bhunia D., Kushwaha R. K., Gupta D., &amp; Chandra K. 2020. Insecta: Coleoptera. Zoological Survey of India, Fauna of Haryana, State Fauna Series 24: 221 - 275." type="book chapter" year="2020">
Ghosh 
<emphasis id="EFACEAD6FFDA892A9C20FBF8FCA7FBE8" box="[801,862,1096,1122]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2020: 239
</bibRefCitation>
</treatmentCitation>
(distribution). — 
<bibRefCitation id="B9494B35FFDA892A9BBFFBF9FEDCFB0F" author="Schoolmeesters P." pageId="44" pageNumber="45" refId="ref31037" refString="Schoolmeesters P. 2022. Scarabs: World Scarabaeidae Database. In: Banki O., Roskov Y., Doring M., Ower G., Hernandez Robles D. R., Plata Corredor C. A., Stjernegaard Jeppesen T., Orn A., Vandepitte L., Hobern D., Schalk P., DeWalt R. E., Ma K., Miller J., Orrell T., Aalbu R., Abbott J., Adlard R., C. Aedo C. et al. (Version 2024 - 03 - 05). https: // doi. org / 10.48580 / dfz 6 w- 38 g" type="book" year="2022">Schoolmeesters 2022</bibRefCitation>
(online catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="DD6736C4FFDA892A9FBCFB3FFD0DFB41" blockId="44.[189,1406,779,1228]" pageId="44" pageNumber="45">
<emphasis id="EFACEAD6FFDA892A9FBCFB3FFE85FB22" box="[189,380,1166,1192]" italics="true" pageId="44" pageNumber="45">Onitisphilemon</emphasis>
– Lansberge (nec Fabricius) 1875: 133. — 
<bibRefCitation id="B9494B35FFDA892A9C94FB3EFB5BFB22" author="Boucomont A." box="[917,1186,1166,1193]" pageId="44" pageNumber="45" pagination="238 - 350" refId="ref27893" refString="Boucomont A. 1914. Les Coprophages de I'Archipel Malais. Annales de la Societe entomologique de France. LXXXIII: 238 - 350." type="book chapter" year="1914">Boucomont 1914: 336</bibRefCitation>
(distribution). — 
<bibRefCitation id="B9494B35FFDA892A9FEBFB03FDB3FB46" author="Boucomont A. &amp; Gillet J." box="[234,586,1201,1228]" pageId="44" pageNumber="45" pagination="1 - 76" refId="ref27918" refString="Boucomont A. &amp; Gillet J. 1921. Faune entomologique de l'Indochine francaise. Famille Scarabaeidae Laparosticti (Coleopteres). Portail. Saigon 4: 1 - 76." type="journal article" year="1921">Boucomont &amp; Gillet 1921: 19</bibRefCitation>
(distribution).
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDA892A9FBCFB42FAFFFAD5" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="DD6736C4FFDA892A9FBCFB42FE3FFA87" blockId="44.[189,454,1267,1293]" box="[189,454,1267,1293]" pageId="44" pageNumber="45">
<heading id="862F81A8FFDA892A9FBCFB42FE3FFA87" bold="true" box="[189,454,1267,1293]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="EFACEAD6FFDA892A9FBCFB42FE3FFA87" bold="true" box="[189,454,1267,1293]" pageId="44" pageNumber="45">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDA892A9FBCFA90FAFFFAD5" blockId="44.[189,1399,1313,1375]" pageId="44" pageNumber="45">
<emphasis id="EFACEAD6FFDA892A9FBCFA90FE71FAB1" box="[189,392,1313,1339]" italics="true" pageId="44" pageNumber="45">Onitissubopacus</emphasis>
belongs to Group III and closely resembles 
<taxonomicName id="1AD84D47FFDA892A9C93FA93FC04FAB1" authorityName="Lansberge" authorityYear="1875" box="[914,1021,1314,1340]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="virens">
<emphasis id="EFACEAD6FFDA892A9C93FA93FC04FAB1" box="[914,1021,1314,1340]" italics="true" pageId="44" pageNumber="45">O. virens</emphasis>
</taxonomicName>
of the same group but differs in that the former has a less strongly punctured pronotum and broadly interrupted frontal carina.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDA892B9FBCFA37FA8EFD90" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45" type="materials_examined">
<paragraph id="DD6736C4FFDA892A9FBCFA37FE15FA2A" blockId="44.[189,492,1414,1440]" box="[189,492,1414,1440]" pageId="44" pageNumber="45">
<heading id="862F81A8FFDA892A9FBCFA37FE15FA2A" bold="true" box="[189,492,1414,1440]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="EFACEAD6FFDA892A9FBCFA37FE15FA2A" bold="true" box="[189,492,1414,1440]" pageId="44" pageNumber="45">Type material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="6DB03C99FFDA892A9FEBFA04FC2EF99F" collectionCode="BMNH" country="Sri Lanka" location="SRI LANKA" pageId="44" pageNumber="45" specimenCode="(E) 1237174" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="DD6736C4FFDA892A9FEBFA04FEA5FA45" blockId="44.[189,1398,1461,1558]" box="[234,348,1461,1487]" pageId="44" pageNumber="45">
<emphasis id="EFACEAD6FFDA892A9FEBFA04FEA5FA45" bold="true" box="[234,348,1461,1487]" pageId="44" pageNumber="45">
<typeStatus id="02638866FFDA892A9FEBFA04FEA5FA45" box="[234,348,1461,1487]" pageId="44" pageNumber="45" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDA892A9FBCFA69FC2EF99F" blockId="44.[189,1398,1461,1558]" pageId="44" pageNumber="45">
<collectingCountry id="A5CF7654FFDA892A9FBCFA69FEACFA78" box="[189,341,1496,1522]" name="Sri Lanka" pageId="44" pageNumber="45">SRI LANKA</collectingCountry>
• 
<specimenCount id="CBDEFD4DFFDA892A9E6BFA69FE7AFA78" box="[362,387,1496,1522]" count="1" pageId="44" pageNumber="45" type="male">♂</specimenCount>
; “Kalupahani, Haldumulle Ceylon, 1904–171, 
<taxonomicName id="1AD84D47FFDA892A9CA2FA69FB95FA78" authorityName="Arrow" authorityYear="1931" box="[931,1132,1496,1522]" class="Insecta" family="Scarabaeidae" genus="Onitis" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="subopacus">
<emphasis id="EFACEAD6FFDA892A9CA2FA69FB95FA78" box="[931,1132,1496,1522]" italics="true" pageId="44" pageNumber="45">Onitis subopacus</emphasis>
</taxonomicName>
type, Arrow (aedeagus point-mounted and attached with specimen)”; 
<collectionCode id="BBC9AE01FFDA892A9DDDFA4DFCCDF99C" box="[732,820,1532,1558]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="44" pageNumber="45">BMNH</collectionCode>
<specimenCode id="8D7E9EBFFFDA892A9C3DFA4AFC2AF99F" box="[828,979,1531,1557]" pageId="44" pageNumber="45">(E) 1237174</specimenCode>
.
</paragraph>
</materialsCitation>
<paragraph id="DD6736C4FFDA892A9FEBF98DFCE2F9DC" blockId="44.[189,1400,1596,2010]" box="[234,795,1596,1623]" pageId="44" pageNumber="45">
<emphasis id="EFACEAD6FFDA892A9FEBF98DFD9BF9DC" bold="true" box="[234,610,1596,1622]" pageId="44" pageNumber="45">Additional material examined</emphasis>
(
<specimenCount id="CBDEFD4DFFDA892A9D70F98CFCEAF9DC" box="[625,787,1596,1623]" count="39" pageId="44" pageNumber="45" type="generic">39 specimens</specimenCount>
)
</paragraph>
<paragraph id="DD6736C4FFDA892B9FBCF9D1FA8EFD90" blockId="44.[189,1400,1596,2010]" lastBlockId="45.[189,1400,263,538]" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">
<materialsCitation id="6DB03C99FFDA892A9FBCF9D1FC4AF914" collectingDate="1928-05-18" collectionCode="BMNH" collectorName="C. F. C. Beeson" country="India" county="Kashmir" location="Haran Pltn" municipality="Haran Pltn." pageId="44" pageNumber="45" specimenCode="(E) 1237175" specimenCount="1" specimenCount-female="1">
<collectingCountry id="A5CF7654FFDA892A9FBCF9D1FEF2F9F0" box="[189,267,1632,1658]" name="India" pageId="44" pageNumber="45">INDIA</collectingCountry>
• 
<specimenCount id="CBDEFD4DFFDA892A9E25F9D1FEC3F9F0" box="[292,314,1632,1658]" count="1" pageId="44" pageNumber="45" type="female">♀</specimenCount>
; “ 
<collectingCounty id="34064E48FFDA892A9E52F9EEFE40F9F3" box="[339,441,1631,1657]" pageId="44" pageNumber="45">Kashmir</collectingCounty>
; 
<location id="D807601FFFDA892A9EC5F9D1FDB9F9F3" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDA892A9EC5F9D1FDB9F9F3" box="[452,576,1631,1658]" country="India" county="Kashmir" municipality="Haran Pltn." name="Haran Pltn" pageId="44" pageNumber="45">Haran Pltn</location>
. 5500, 
<location id="D807601FFFDA892A9D93F9EEFCDBF9F3" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDA892A9D93F9EEFCDBF9F3" box="[658,802,1631,1657]" country="India" county="Kashmir" municipality="Haran Pltn." name="Sindh Valley" pageId="44" pageNumber="45">Sindh Valley</location>
, 
<date id="A9661004FFDA892A9C2CF9D1FC64F9F0" box="[813,925,1632,1658]" pageId="44" pageNumber="45" value="1928-05-18">
<collectingDate id="B922E9ECFFDA892A9C2CF9D1FC64F9F0" box="[813,925,1632,1658]" pageId="44" pageNumber="45" value="1928-05-18">18.v.1928</collectingDate>
</date>
, 
<collectorName id="702D5312FFDA892A9CA8F9D1FBB4F9F0" box="[937,1101,1632,1658]" pageId="44" pageNumber="45">C.F.C. Beeson</collectorName>
; ex. coll., Dehradun, B.M. 1928-472; 
<emphasis id="EFACEAD6FFDA892A9E38F932FE05F917" box="[313,508,1667,1693]" italics="true" pageId="44" pageNumber="45">Onitissubopacus</emphasis>
co-type, Arrow”; 
<collectionCode id="BBC9AE01FFDA892A9DCEF932FCDEF917" box="[719,807,1667,1693]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="44" pageNumber="45">BMNH</collectionCode>
<specimenCode id="8D7E9EBFFFDA892A9C27F932FC4AF914" box="[806,947,1667,1694]" pageId="44" pageNumber="45">(E) 1237175</specimenCode>
</materialsCitation>
• 
<materialsCitation id="6DB03C99FFDA892A9CD1F932FE65F94A" collectionCode="ZSIM" country="India" determinerName="G. J. Arrow" location="Arrow Collection" municipality="Roorkee" pageId="44" pageNumber="45" specimenCount="1" specimenCount-male="1">
<specimenCount id="CBDEFD4DFFDA892A9CD1F932FC11F917" box="[976,1000,1667,1693]" count="1" pageId="44" pageNumber="45" type="male">♂</specimenCount>
; “ 
<collectingMunicipality id="3D03ACBEFFDA892A9B01F932FB9DF917" box="[1024,1124,1667,1693]" pageId="44" pageNumber="45">Roorkee</collectingMunicipality>
, 
<location id="D807601FFFDA892A9B71F932FAC0F917" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDA892A9B71F932FAC0F917" box="[1136,1337,1667,1693]" country="India" municipality="Roorkee" name="Arrow Collection" pageId="44" pageNumber="45">Arrow Collection</location>
, Det. 
<determinerName id="28BBDA00FFDA892A9FBCF917FEC3F94A" box="[189,314,1702,1728]" pageId="44" pageNumber="45">G.J. Arrow</determinerName>
”; 
<collectionCode id="BBC9AE01FFDA892A9E59F917FE65F94A" box="[344,412,1702,1728]" pageId="44" pageNumber="45">ZSIM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="6DB03C99FFDA892A9EB3F916FDE0F94A" box="[434,537,1702,1729]" collectionCode="ZSIM" country="India" pageId="44" pageNumber="45" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBDEFD4DFFDA892A9EB3F916FE31F94B" box="[434,456,1703,1729]" count="1" pageId="44" pageNumber="45" type="female">♀</specimenCount>
; 
<collectionCode id="BBC9AE01FFDA892A9ED4F917FDE0F94A" box="[469,537,1702,1728]" pageId="44" pageNumber="45">ZSIM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="6DB03C99FFDA892A9D2EF917FCBFF88C" collectingDate="2009-05-08" collectingMethod="open cow dung bait traps" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara" country="India" elevation="839" latitude="8.861611" location="Rockwood" longLatPrecision="1" longitude="77.13347" municipality="Shendurney Wildlife Sanctuary" pageId="44" pageNumber="45" specimenCode="Co/Sc1000711 to 1000716" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFDA892A9D2EF917FD88F94B" box="[559,625,1702,1729]" count="4" pageId="44" pageNumber="45" type="male">4 ♂♂</specimenCount>
, 
<specimenCount id="CBDEFD4DFFDA892A9D7AF917FD45F94B" box="[635,700,1702,1729]" count="2" pageId="44" pageNumber="45" type="female">2 ♀♀</specimenCount>
; 
<collectingRegion id="1F1CF826FFDA892A9DC9F917FCECF94A" box="[712,789,1702,1728]" country="India" name="Kerala" pageId="44" pageNumber="45">Kerala</collectingRegion>
, 
<collectingMunicipality id="3D03ACBEFFDA892A9C1EF917FB85F94A" box="[799,1148,1702,1728]" pageId="44" pageNumber="45">Shendurney Wildlife Sanctuary</collectingMunicipality>
, 
<location id="D807601FFFDA892A9B87F917FAF9F94A" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDA892A9B87F917FAF9F94A" box="[1158,1280,1702,1728]" country="India" latitude="8.861611" longLatPrecision="1" longitude="77.13347" municipality="Shendurney Wildlife Sanctuary" name="Rockwood" pageId="44" pageNumber="45" stateProvince="Kerala">Rockwood</location>
grassland; 
<geoCoordinate id="B8EC5003FFDA892A9FBCF979FE99F96E" box="[189,352,1736,1764]" degrees="08" direction="north" minutes="51" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="41.8" value="8.861611">08°51ʹ41.8ʺ N</geoCoordinate>
, 
<geoCoordinate id="B8EC5003FFDA892A9E6DF979FDF5F96E" box="[364,524,1736,1764]" degrees="77" direction="east" minutes="08" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="00.5" value="77.13347">77°08ʹ00.5ʺ E</geoCoordinate>
; elev. 
<elevation id="56F5D1F7FFDA892A9D54F978FD25F969" box="[597,732,1737,1764]" metricMagnitude="2" metricUnit="m" metricValue="8.39" pageId="44" pageNumber="45" unit="m" value="839.0">
<quantity id="1A209B21FFDA892A9D54F978FD66F96E" box="[597,671,1737,1764]" metricMagnitude="2" metricUnit="m" metricValue="8.39" pageId="44" pageNumber="45" unit="m" value="839.0">839 m</quantity>
a.s.l.
</elevation>
; 
<date id="A9661004FFDA892A9DE8F978FC81F969" box="[745,888,1737,1764]" pageId="44" pageNumber="45" value="2009-05-08">
<collectingDate id="B922E9ECFFDA892A9DE8F978FC81F969" box="[745,888,1737,1764]" pageId="44" pageNumber="45" value="2009-05-08">8 May 2009</collectingDate>
</date>
; 
<collectingMethod id="04994ED3FFDA892A9C85F97BFB5CF96E" box="[900,1189,1737,1764]" pageId="44" pageNumber="45">open cow dung bait traps</collectingMethod>
; 
<collectorName id="702D5312FFDA892A9BB3F978FE9FF88C" pageId="44" pageNumber="45">Seena Narayanan Karimbumkara</collectorName>
leg.; 
<collectionCode id="BBC9AE01FFDA892A9EB7F95CFE06F88D" box="[438,511,1773,1799]" pageId="44" pageNumber="45">AIMB</collectionCode>
/ 
<specimenCode id="8D7E9EBFFFDA892A9D09F95CFCBFF88C" box="[520,838,1772,1799]" pageId="44" pageNumber="45">Co/Sc1000711 to 1000716</specimenCode>
</materialsCitation>
• 
<materialsCitation id="6DB03C99FFDA892A9C70F95CFA8EF8A0" collectingDate="2009-12-04" collectionCode="AIMB" collectorName="Co &amp; Sc" country="India" elevation="173" latitude="8.912666" location="Kattilappara" longLatPrecision="1" longitude="77.09592" municipality="Shendurney Wildlife Sanctuary" pageId="44" pageNumber="45" specimenCode="Co/Sc1000717" specimenCount="1" specimenCount-female="1" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFDA892A9C70F95CFC7EF88D" box="[881,903,1773,1799]" count="1" pageId="44" pageNumber="45" type="female">♀</specimenCount>
; 
<collectingRegion id="1F1CF826FFDA892A9C9EF95DFC15F88C" box="[927,1004,1772,1798]" country="India" name="Kerala" pageId="44" pageNumber="45">Kerala</collectingRegion>
, 
<collectingMunicipality id="3D03ACBEFFDA892A9B01F95DFA8AF88D" box="[1024,1395,1772,1799]" pageId="44" pageNumber="45">Shendurney Wildlife Sanctuary</collectingMunicipality>
, 
<location id="D807601FFFDA892A9FBCF8BEFEBFF8A3" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDA892A9FBCF8BEFEBFF8A3" box="[189,326,1807,1833]" country="India" latitude="8.912666" longLatPrecision="1" longitude="77.09592" municipality="Shendurney Wildlife Sanctuary" name="Kattilappara" pageId="44" pageNumber="45" stateProvince="Kerala">Kattilappara</location>
evergreen forest; 
<geoCoordinate id="B8EC5003FFDA892A9D0DF8BFFD55F8A0" box="[524,684,1806,1834]" degrees="08" direction="north" minutes="54" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="45.6" value="8.912666">08°54ʹ45.6ʺ N</geoCoordinate>
, 
<geoCoordinate id="B8EC5003FFDA892A9DB7F8BFFCADF8A0" box="[694,852,1806,1834]" degrees="77" direction="east" minutes="05" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="45.3" value="77.09592">77°05ʹ45.3ʺ E</geoCoordinate>
; elev. 
<quantity id="1A209B21FFDA892A9C99F8BEFC24F8A3" box="[920,989,1807,1834]" metricMagnitude="2" metricUnit="m" metricValue="1.725168" pageId="44" pageNumber="45" unit="ft" value="566.0">
<elevation id="56F5D1F7FFDA892A9C99F8BEFC24F8A3" box="[920,989,1807,1834]" metricMagnitude="2" metricUnit="m" metricValue="1.725168" pageId="44" pageNumber="45" unit="ft" value="566.0">566 ft</elevation>
</quantity>
, 
<date id="A9661004FFDA892A9CECF8A1FB8EF8A0" box="[1005,1143,1808,1834]" pageId="44" pageNumber="45" value="2009-12-04">
<collectingDate id="B922E9ECFFDA892A9CECF8A1FB8EF8A0" box="[1005,1143,1808,1834]" pageId="44" pageNumber="45" value="2009-12-04">4 Dec. 2009</collectingDate>
</date>
; 
<collectionCode id="BBC9AE01FFDA892A9B7EF8A1FB31F8A0" box="[1151,1224,1808,1834]" pageId="44" pageNumber="45">AIMB</collectionCode>
/ 
<specimenCode id="8D7E9EBFFFDA892A9BD0F8A1FA8EF8A0" box="[1233,1399,1808,1834]" pageId="44" pageNumber="45">Co/Sc1000717</specimenCode>
</materialsCitation>
• 
<materialsCitation id="6DB03C99FFDA892A9FCFF882FD8CF819" collectingDate="2009-12-03" collectingMethod="open cow dung bait traps" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara" country="India" elevation="185" latitude="8.9151945" location="Kattilappara" longLatPrecision="1" longitude="77.10797" municipality="Shendurney Wildlife Sanctuary" pageId="44" pageNumber="45" specimenCode="Co/Sc1000718 to 1000724" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFDA892A9FCFF882FEE8F8C7" box="[206,273,1843,1869]" count="2" pageId="44" pageNumber="45" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="CBDEFD4DFFDA892A9E1CF882FEA6F8C7" box="[285,351,1843,1870]" count="5" pageId="44" pageNumber="45" type="female">5 ♀♀</specimenCount>
; 
<collectingRegion id="1F1CF826FFDA892A9E6CF882FE43F8C7" box="[365,442,1843,1869]" country="India" name="Kerala" pageId="44" pageNumber="45">Kerala</collectingRegion>
, 
<collectingMunicipality id="3D03ACBEFFDA892A9EC7F882FCD1F8C7" box="[454,808,1843,1869]" pageId="44" pageNumber="45">Shendurney Wildlife Sanctuary</collectingMunicipality>
; 
<location id="D807601FFFDA892A9C32F882FC45F8C7" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDA892A9C32F882FC45F8C7" box="[819,956,1843,1869]" country="India" latitude="8.9151945" longLatPrecision="1" longitude="77.10797" municipality="Shendurney Wildlife Sanctuary" name="Kattilappara" pageId="44" pageNumber="45" stateProvince="Kerala">Kattilappara</location>
moist deciduous forest; 
<geoCoordinate id="B8EC5003FFDA892A9BD1F880FA8AF8C7" box="[1232,1395,1841,1869]" degrees="08" direction="north" minutes="54" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="54.7" value="8.9151945">08°54ʹ54.7ʺ N</geoCoordinate>
, 
<geoCoordinate id="B8EC5003FFDA892A9FBCF8E5FEA4F8FB" box="[189,349,1876,1905]" degrees="77" direction="east" minutes="06" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="28.7" value="77.10797">77°06ʹ28.7ʺ E</geoCoordinate>
; elev. 
<elevation id="56F5D1F7FFDA892A9EA4F8E7FDD2F8FA" box="[421,555,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="1.85" pageId="44" pageNumber="45" unit="m" value="185.0">
<quantity id="1A209B21FFDA892A9EA4F8E7FE16F8FB" box="[421,495,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="1.85" pageId="44" pageNumber="45" unit="m" value="185.0">185 m</quantity>
a.s.l.
</elevation>
; 
<date id="A9661004FFDA892A9D38F8E7FD31F8FA" box="[569,712,1878,1905]" pageId="44" pageNumber="45" value="2009-12-03">
<collectingDate id="B922E9ECFFDA892A9D38F8E7FD31F8FA" box="[569,712,1878,1905]" pageId="44" pageNumber="45" value="2009-12-03">3 Dec. 2009</collectingDate>
</date>
; 
<collectingMethod id="04994ED3FFDA892A9DD2F8E6FC0DF8FB" box="[723,1012,1878,1905]" pageId="44" pageNumber="45">open cow dung bait traps</collectingMethod>
; 
<collectorName id="702D5312FFDA892A9B00F8E7FA8EF8FA" box="[1025,1399,1878,1905]" pageId="44" pageNumber="45">Seena Narayanan Karimbumkara</collectorName>
leg.; 
<collectionCode id="BBC9AE01FFDA892A9FF5F8C8FEC4F819" box="[244,317,1913,1939]" pageId="44" pageNumber="45">AIMB</collectionCode>
/ 
<specimenCode id="8D7E9EBFFFDA892A9E47F8C8FD8CF819" box="[326,629,1913,1940]" pageId="44" pageNumber="45">Co/Sc1000718 to 1000724</specimenCode>
</materialsCitation>
• 
<materialsCitation id="6DB03C99FFDA892A9D91F8C8FC0AF850" collectingDate="2009-12-03" collectingMethod="open cow dung bait traps" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara" country="India" elevation="207" latitude="8.918222" location="Kattilapara" longLatPrecision="1" longitude="77.09575" municipality="Shendurney Wildlife Sanctuary" pageId="44" pageNumber="45" specimenCode="Co/Sc1000725 to 1000738" specimenCount="15" specimenCount-female="6" specimenCount-male="9" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFDA892A9D91F8C8FD2DF81E" box="[656,724,1913,1940]" count="9" pageId="44" pageNumber="45" type="male">9 ♂♂</specimenCount>
, 
<specimenCount id="CBDEFD4DFFDA892A9DE0F8C8FCDDF81E" box="[737,804,1913,1940]" count="6" pageId="44" pageNumber="45" type="female">6 ♀♀</specimenCount>
; 
<collectingRegion id="1F1CF826FFDA892A9C32F8C8FC79F819" box="[819,896,1913,1939]" country="India" name="Kerala" pageId="44" pageNumber="45">Kerala</collectingRegion>
, 
<collectingMunicipality id="3D03ACBEFFDA892A9C8CF8C8FB17F819" box="[909,1262,1913,1939]" pageId="44" pageNumber="45">Shendurney Wildlife Sanctuary</collectingMunicipality>
, 
<location id="D807601FFFDA892A9BFAF8C8FA8EF819" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDA892A9BFAF8C8FA8EF819" box="[1275,1399,1913,1939]" country="India" latitude="8.918222" longLatPrecision="1" longitude="77.09575" municipality="Shendurney Wildlife Sanctuary" name="Kattilapara" pageId="44" pageNumber="45" stateProvince="Kerala">Kattilapara</location>
degraded forest; 
<geoCoordinate id="B8EC5003FFDA892A9E7DF82AFDD9F83D" box="[380,544,1947,1975]" degrees="08" direction="north" minutes="55" orientation="latitude" pageId="44" pageNumber="45" precision="1" seconds="05.6" value="8.918222">08°55ʹ05.6ʺ N</geoCoordinate>
, 
<geoCoordinate id="B8EC5003FFDA892A9D2CF82AFD37F83D" box="[557,718,1947,1975]" degrees="77" direction="east" minutes="05" orientation="longitude" pageId="44" pageNumber="45" precision="1" seconds="44.7" value="77.09575">77°05ʹ44.7ʺ E</geoCoordinate>
; elev. 
<elevation id="56F5D1F7FFDA892A9C18F82CFC58F83C" box="[793,929,1948,1975]" metricMagnitude="2" metricUnit="m" metricValue="2.07" pageId="44" pageNumber="45" unit="m" value="207.0">
<quantity id="1A209B21FFDA892A9C18F82CFC9AF83D" box="[793,867,1949,1975]" metricMagnitude="2" metricUnit="m" metricValue="2.07" pageId="44" pageNumber="45" unit="m" value="207.0">207 m</quantity>
a.s.l.
</elevation>
; 
<date id="A9661004FFDA892A9CAEF82CFBB9F83D" box="[943,1088,1949,1975]" pageId="44" pageNumber="45" value="2009-12-03">
<collectingDate id="B922E9ECFFDA892A9CAEF82CFBB9F83D" box="[943,1088,1949,1975]" pageId="44" pageNumber="45" value="2009-12-03">3 Dec. 2009</collectingDate>
</date>
; 
<collectingMethod id="04994ED3FFDA892A9B4CF82CFA8AF83D" box="[1101,1395,1948,1975]" pageId="44" pageNumber="45">open cow dung bait traps</collectingMethod>
; 
<collectorName id="702D5312FFDA892A9FBCF871FDCAF853" box="[189,563,1983,2010]" pageId="44" pageNumber="45">Seena Narayanan Karimbumkara</collectorName>
leg.; 
<collectionCode id="BBC9AE01FFDA892A9D72F80EFD45F850" box="[627,700,1983,2010]" pageId="44" pageNumber="45">AIMB</collectionCode>
/ 
<specimenCode id="8D7E9EBFFFDA892A9DC4F80EFC0AF850" box="[709,1011,1983,2010]" pageId="44" pageNumber="45">Co/Sc1000725 to 1000738</specimenCode>
</materialsCitation>
• 
<materialsCitation id="6DB03C99FFDA892B9B0FF871FB4DFECC" collectingDate="2009-11-29" collectingMethod="open cow dung bait traps" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara" country="India" elevation="1234" lastPageId="45" lastPageNumber="46" latitude="8.827528" location="Pandimotta" longLatPrecision="1" longitude="77.21719" municipality="Shendurney Wildlife Sanctuary" pageId="44" pageNumber="45" specimenCode="Co/Sc1000739" specimenCount="1" specimenCount-male="1" stateProvince="Kerala">
<specimenCount id="CBDEFD4DFFDA892A9B0FF871FBDFF850" box="[1038,1062,1984,2010]" count="1" pageId="44" pageNumber="45" type="male">♂</specimenCount>
; 
<collectingRegion id="1F1CF826FFDA892A9B35F80EFB78F853" box="[1076,1153,1983,2009]" country="India" name="Kerala" pageId="44" pageNumber="45">Kerala</collectingRegion>
, 
<collectingMunicipality id="3D03ACBEFFDA892B9B8CF80EFED4FEA9" lastPageId="45" lastPageNumber="46" pageId="44" pageNumber="45">Shendurney Wildlife Sanctuary</collectingMunicipality>
, 
<location id="D807601FFFDB892B9E38FEB8FE41FEA9" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDB892B9E38FEB8FE41FEA9" box="[313,440,265,291]" country="India" latitude="8.827528" longLatPrecision="1" longitude="77.21719" municipality="Shendurney Wildlife Sanctuary" name="Pandimotta" pageId="45" pageNumber="46" stateProvince="Kerala">Pandimotta</location>
montane evergreen; 
<geoCoordinate id="B8EC5003FFDB892B9DA2FEB6FCBCFEA9" box="[675,837,263,291]" degrees="08" direction="north" minutes="49" orientation="latitude" pageId="45" pageNumber="46" precision="1" seconds="39.1" value="8.827528">08°49ʹ39.1ʺ N</geoCoordinate>
, 
<geoCoordinate id="B8EC5003FFDB892B9C51FEB6FC16FEA9" box="[848,1007,263,291]" degrees="77" direction="east" minutes="13" orientation="longitude" pageId="45" pageNumber="46" precision="1" seconds="01.9" value="77.21719">77°13ʹ01.9ʺ E</geoCoordinate>
; elev. 
<elevation id="56F5D1F7FFDB892B9B36FEB8FB30FEA9" box="[1079,1225,265,291]" metricMagnitude="3" metricUnit="m" metricValue="1.234" pageId="45" pageNumber="46" unit="m" value="1234.0">
<quantity id="1A209B21FFDB892B9B36FEB8FB74FEA9" box="[1079,1165,265,291]" metricMagnitude="3" metricUnit="m" metricValue="1.234" pageId="45" pageNumber="46" unit="m" value="1234.0">1234 m</quantity>
a.s.l.
</elevation>
; 
<date id="A9661004FFDB892B9BD7FEB8FA8AFEA9" box="[1238,1395,265,291]" pageId="45" pageNumber="46" value="2009-11-29">
<collectingDate id="B922E9ECFFDB892B9BD7FEB8FA8AFEA9" box="[1238,1395,265,291]" pageId="45" pageNumber="46" value="2009-11-29">29 Nov. 2009</collectingDate>
</date>
; 
<collectingMethod id="04994ED3FFDB892B9FBCFE9CFE12FECD" box="[189,491,300,327]" pageId="45" pageNumber="46">open cow dung bait traps</collectingMethod>
; 
<collectorName id="702D5312FFDB892B9EFAFE9DFC8EFECC" box="[507,887,300,327]" pageId="45" pageNumber="46">Seena Narayanan Karimbumkara</collectorName>
leg.; 
<collectionCode id="BBC9AE01FFDB892B9CBCFE9DFBFFFECC" box="[957,1030,300,326]" pageId="45" pageNumber="46">AIMB</collectionCode>
/ 
<specimenCode id="8D7E9EBFFFDB892B9B0EFE9DFB4DFECC" box="[1039,1204,300,326]" pageId="45" pageNumber="46">Co/Sc1000739</specimenCode>
</materialsCitation>
• 
<materialsCitation id="6DB03C99FFDB892B9BD4FE9CFC48FE07" collectingMethod="open cow dung bait traps" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan" country="India" latitude="11.974199" location="Sungathakatte" longLatPrecision="1" longitude="76.1856" municipality="Nagarhole National Park" pageId="45" pageNumber="46" specimenCode="/Co/Sc1000740" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka">
<specimenCount id="CBDEFD4DFFDB892B9BD4FE9CFB14FECD" box="[1237,1261,301,327]" count="1" pageId="45" pageNumber="46" type="male">♂</specimenCount>
; 
<collectingRegion id="1F1CF826FFDB892B9BFCFE9DFA8AFECC" box="[1277,1395,300,326]" country="India" name="Karnataka" pageId="45" pageNumber="46">Karnataka</collectingRegion>
, 
<collectingMunicipality id="3D03ACBEFFDB892B9FBCFEFEFE2BFEE3" box="[189,466,335,361]" pageId="45" pageNumber="46">Nagarhole National Park</collectingMunicipality>
, 
<location id="D807601FFFDB892B9EDDFEFEFD83FEE3" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDB892B9EDDFEFEFD83FEE3" box="[476,634,335,361]" country="India" latitude="11.974199" longLatPrecision="1" longitude="76.1856" municipality="Nagarhole National Park" name="Sungathakatte" pageId="45" pageNumber="46" stateProvince="Karnataka">Sungathakatte</location>
dry deciduous forest; 
<geoCoordinate id="B8EC5003FFDB892B9C6CFEFFFBE2FEE0" box="[877,1051,334,362]" degrees="11" direction="north" minutes="58" orientation="latitude" pageId="45" pageNumber="46" precision="1" seconds="27.12" value="11.974199">11°58ʹ27.12″ N</geoCoordinate>
, 
<geoCoordinate id="B8EC5003FFDB892B9B24FEFFFB3AFEE0" box="[1061,1219,334,362]" degrees="76" direction="east" minutes="11" orientation="longitude" pageId="45" pageNumber="46" precision="1" seconds="8.16" value="76.1856">76°11ʹ8.16″ E</geoCoordinate>
; 
<collectingMethod id="04994ED3FFDB892B9BCCFEE1FEDEFE07" pageId="45" pageNumber="46">open cow dung bait traps</collectingMethod>
; 
<collectorName id="702D5312FFDB892B9E32FEC3FD82FE06" box="[307,635,370,396]" pageId="45" pageNumber="46">Priyadarsanan Dharma Rajan</collectorName>
leg.; 
<collectionCode id="BBC9AE01FFDB892B9DBBFEC2FCFAFE07" box="[698,771,371,397]" pageId="45" pageNumber="46">AIMB</collectionCode>
<specimenCode id="8D7E9EBFFFDB892B9C02FEC2FC48FE07" box="[771,945,371,397]" pageId="45" pageNumber="46">/Co/Sc1000740</specimenCode>
</materialsCitation>
• 
<materialsCitation id="6DB03C99FFDB892B9CCBFEC2FEC2FE59" collectingMethod="open cattle dung baited trap" collectionCode="AIMB" collectorName="Priyadarsanan Dharma Rajan" country="India" latitude="12.0634" location="Sarathi Nulla" longLatPrecision="1" longitude="76.17799" pageId="45" pageNumber="46" specimenCode="Co/ Sc1000741" specimenCount="1" specimenCount-female="1">
<specimenCount id="CBDEFD4DFFDB892B9CCBFEC2FC19FE07" box="[970,992,371,397]" count="1" pageId="45" pageNumber="46" type="female">♀</specimenCount>
; 
<location id="D807601FFFDB892B9CEFFEC3FB7AFE06" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDB892B9CEFFEC3FB7AFE06" box="[1006,1155,370,396]" country="India" latitude="12.0634" longLatPrecision="1" longitude="76.17799" name="Sarathi Nulla" pageId="45" pageNumber="46">Sarathi Nulla</location>
dry deciduous forest; 
<geoCoordinate id="B8EC5003FFDB892B9FBCFE25FE9BFE3A" box="[189,354,404,432]" degrees="12" direction="north" minutes="3" orientation="latitude" pageId="45" pageNumber="46" precision="1" seconds="48.24" value="12.0634">12°3ʹ48.24″ N</geoCoordinate>
, 
<geoCoordinate id="B8EC5003FFDB892B9E71FE25FDEBFE3A" box="[368,530,404,432]" degrees="76" direction="east" minutes="10" orientation="longitude" pageId="45" pageNumber="46" precision="1" seconds="40.8" value="76.17799">76°10ʹ40.8″ E</geoCoordinate>
; 
<collectingMethod id="04994ED3FFDB892B9D20FE27FC9AFE3A" box="[545,867,406,432]" pageId="45" pageNumber="46">open cattle dung baited trap</collectingMethod>
; 
<collectorName id="702D5312FFDB892B9C71FE27FB42FE3A" box="[880,1211,406,432]" pageId="45" pageNumber="46">Priyadarsanan Dharma Rajan</collectorName>
leg.; 
<collectionCode id="BBC9AE01FFDB892B9BFCFE27FABFFE3A" box="[1277,1350,406,432]" pageId="45" pageNumber="46">AIMB</collectionCode>
/ 
<specimenCode id="8D7E9EBFFFDB892B9A4FFE27FEC2FE59" pageId="45" pageNumber="46">Co/ Sc1000741</specimenCode>
</materialsCitation>
• 
<materialsCitation id="6DB03C99FFDB892B9E57FE08FD5FFE7C" collectingDate="2019-02-18" collectionCode="AIMB" collectorName="Harsha Malhotra" country="India" location="South Tripura" municipality="Eden of Bison" pageId="45" pageNumber="46" specimenCode="Co/Sc1000742" specimenCount="1" specimenCount-female="1" stateProvince="Tripura">
<specimenCount id="CBDEFD4DFFDB892B9E57FE08FE95FE59" box="[342,364,441,467]" count="1" pageId="45" pageNumber="46" type="female">♀</specimenCount>
; 
<collectingRegion id="1F1CF826FFDB892B9E7AFE08FE29FE59" box="[379,464,441,467]" country="India" name="Tripura" pageId="45" pageNumber="46">Tripura</collectingRegion>
, 
<collectingMunicipality id="3D03ACBEFFDB892B9EDDFE08FD7BFE59" box="[476,642,441,467]" pageId="45" pageNumber="46">Eden of Bison</collectingMunicipality>
, 
<location id="D807601FFFDB892B9D8EFE08FCD4FE59" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDB892B9D8EFE08FCD4FE59" box="[655,813,441,467]" country="India" municipality="Eden of Bison" name="South Tripura" pageId="45" pageNumber="46" stateProvince="Tripura">South Tripura</location>
, forest; 
<date id="A9661004FFDB892B9C89FE08FBDBFE59" box="[904,1058,441,467]" pageId="45" pageNumber="46" value="2019-02-18">
<collectingDate id="B922E9ECFFDB892B9C89FE08FBDBFE59" box="[904,1058,441,467]" pageId="45" pageNumber="46" value="2019-02-18">18 Feb. 2019</collectingDate>
</date>
; hand-picked from cow dung; 
<collectorName id="702D5312FFDB892B9FBCFE6DFE8EFE7C" box="[189,375,476,502]" pageId="45" pageNumber="46">Harsha Malhotra</collectorName>
leg.; 
<collectionCode id="BBC9AE01FFDB892B9EAEFE6DFE01FE7C" box="[431,504,476,502]" pageId="45" pageNumber="46">AIMB</collectionCode>
/ 
<specimenCode id="8D7E9EBFFFDB892B9D00FE6DFD5FFE7C" box="[513,678,476,502]" pageId="45" pageNumber="46">Co/Sc1000742</specimenCode>
</materialsCitation>
• 
<materialsCitation id="6DB03C99FFDB892B9DB8FE6DFA8AFD90" collectingDate="2021-04-24" collectionCode="AIMB" collectorName="Seena Narayanan Karimbumkara" country="India" location="Forest" municipality="Singphan Elephant Reserve" pageId="45" pageNumber="46" specimenCode="Co/Sc1000743 to 1000745" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Nagaland">
<specimenCount id="CBDEFD4DFFDB892B9DB8FE6DFD00FE7D" box="[697,761,476,503]" count="2" pageId="45" pageNumber="46" type="male">2 ♂♂</specimenCount>
, 
<specimenCount id="CBDEFD4DFFDB892B9C03FE6CFCE1FE7D" box="[770,792,477,503]" count="1" pageId="45" pageNumber="46" type="female">♀</specimenCount>
; 
<collectingRegion id="1F1CF826FFDB892B9C25FE6DFC6BFE7C" box="[804,914,476,502]" country="India" name="Nagaland" pageId="45" pageNumber="46">Nagaland</collectingRegion>
, 
<collectingMunicipality id="3D03ACBEFFDB892B9C9BFE6DFB35FE7D" box="[922,1228,476,503]" pageId="45" pageNumber="46">Singphan Elephant Reserve</collectingMunicipality>
, 
<location id="D807601FFFDB892B9BD4FE6CFAE7FE7D" LSID="urn:lsid:plazi:treatment:557187D2FFDA892B9D65FDD4FC1FFA7B:D807601FFFDB892B9BD4FE6CFAE7FE7D" box="[1237,1310,477,503]" country="India" municipality="Singphan Elephant Reserve" name="Forest" pageId="45" pageNumber="46" stateProvince="Nagaland">Forest</location>
; 
<date id="A9661004FFDB892B9A27FE6DFF01FD93" pageId="45" pageNumber="46" value="2021-04-24">
<collectingDate id="B922E9ECFFDB892B9A27FE6DFF01FD93" pageId="45" pageNumber="46" value="2021-04-24">24Apr. 2021</collectingDate>
</date>
; hand-picked from cow dung; 
<collectorName id="702D5312FFDB892B9D48FE4EFC43FD93" box="[585,954,511,538]" pageId="45" pageNumber="46">Seena Narayanan Karimbumkara</collectorName>
leg.; 
<collectionCode id="BBC9AE01FFDB892B9CF4FE4EFBC7FD93" box="[1013,1086,511,537]" pageId="45" pageNumber="46">AIMB</collectionCode>
/ 
<specimenCode id="8D7E9EBFFFDB892B9B46FE4EFA8AFD90" box="[1095,1395,511,538]" pageId="45" pageNumber="46">Co/Sc1000743 to 1000745</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDB892B9FBCFDF1FCE8FB00" pageId="45" pageNumber="46" type="description">
<paragraph id="DD6736C4FFDB892B9FBCFDF1FEB7FDD0" blockId="45.[189,334,576,602]" box="[189,334,576,602]" pageId="45" pageNumber="46">
<heading id="862F81A8FFDB892B9FBCFDF1FEB7FDD0" bold="true" box="[189,334,576,602]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="EFACEAD6FFDB892B9FBCFDF1FEB7FDD0" bold="true" box="[189,334,576,602]" pageId="45" pageNumber="46">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDB892B9FBCFDDEFCFCFD03" blockId="45.[189,773,623,650]" box="[189,773,623,650]" pageId="45" pageNumber="46">MEASUREMENTS (in mm). TL=16–23, BW= 8–12.</paragraph>
<paragraph id="DD6736C4FFDB892B9FEBFD00FED3FD41" blockId="45.[189,1399,689,962]" box="[234,298,689,715]" pageId="45" pageNumber="46">
<emphasis id="EFACEAD6FFDB892B9FEBFD00FED3FD41" bold="true" box="[234,298,689,715]" pageId="45" pageNumber="46">Male</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDB892B9FBCFD65FBFDFC4B" blockId="45.[189,1399,689,962]" pageId="45" pageNumber="46">
Black with a slight metallic lustre; clypeus elliptical, rugulose, vertex granular, more shining; frontal carina widely interrupted in the middle with a conical tubercle behind and a curved clypeal carina in front; pronotum not strongly but rather closely punctured, with an incomplete smooth median longitudinal line, basal foveae deep; pygidium opaque scarcely punctured; elytra finely striate with sutural, 3 
<superScript id="2AAD9B8CFFDB892B9E2BFCD1FEC1FCE4" attach="left" box="[298,312,864,878]" fontSize="6" pageId="45" pageNumber="46">rd</superScript>
and 5 
<superScript id="2AAD9B8CFFDB892B9E7EFCD1FE75FCE4" attach="left" box="[383,396,864,878]" fontSize="6" pageId="45" pageNumber="46">th</superScript>
intervals slightly convex; metasternal shield smooth and shining, the front angles and sides of metasternum closely clothed with reddish hairs. Protibia toothed, mid femur lobed near middle with tooth on posterior edge, trochanter of the hind leg sharply toothed.
</paragraph>
<paragraph id="DD6736C4FFDB892B9FBCFC58FA98FBAC" blockId="45.[189,1399,1000,1062]" pageId="45" pageNumber="46">
AEDEAGUS (in mm) (
<figureCitation id="45E32A41FFDB892B9EB5FC58FDB1FB89" box="[436,584,1001,1027]" captionStart="Fig" captionStartId="36.[189,232,1964,1990]" captionTargetBox="[275,1278,1055,1924]" captionTargetId="figure-228@36.[189,1398,1033,1927]" captionTargetPageId="36" captionText="Fig. 13. Aedeagus, lateral, dorsal, ventral and apical views. A–D. Onitissubopacus Arrow, 1931, AIMB/ Co/Sc1000711. E–H. O. virens Lansberge, 1875, AIMB/Co/Sc1000765. (Images: AIMB)." figureDoi="http://doi.org/10.5281/zenodo.13742887" httpUri="https://zenodo.org/record/13742887/files/figure.png" pageId="45" pageNumber="46">Fig. 13A–D</figureCitation>
). LP=3, Lp=2.5; BP=1.25, BpB= 0.5, BpT =0.4. Phallobase longer than parameres. Parameres curved outwards at the base and then curved inwards with the tip pointed.
</paragraph>
<paragraph id="DD6736C4FFDB892B9FEBFBFCFEBCFBED" blockId="45.[189,785,1101,1163]" box="[234,325,1101,1127]" pageId="45" pageNumber="46">
<emphasis id="EFACEAD6FFDB892B9FEBFBFCFEBCFBED" bold="true" box="[234,325,1101,1127]" pageId="45" pageNumber="46">Female</emphasis>
</paragraph>
<paragraph id="DD6736C4FFDB892B9FBCFBC1FCE8FB00" blockId="45.[189,785,1101,1163]" box="[189,785,1136,1163]" pageId="45" pageNumber="46">Protibia toothed, middle and hind legs not toothed.</paragraph>
</subSubSection>
<subSubSection id="95C2654FFFDB892B9FBCFB03FC1FFA7B" pageId="45" pageNumber="46" type="distribution">
<paragraph id="DD6736C4FFDB892B9FBCFB03FEAFFB46" blockId="45.[189,342,1202,1228]" box="[189,342,1202,1228]" pageId="45" pageNumber="46">
<heading id="862F81A8FFDB892B9FBCFB03FEAFFB46" bold="true" box="[189,342,1202,1228]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="EFACEAD6FFDB892B9FBCFB03FEAFFB46" bold="true" box="[189,342,1202,1228]" pageId="45" pageNumber="46">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DD6736C4FFDB892B9FBCFB51FC1FFA7B" blockId="45.[189,1399,1248,1522]" pageId="45" pageNumber="46">
<collectingCountry id="A5CF7654FFDB892B9FBCFB51FF07FB70" box="[189,254,1248,1274]" name="India" pageId="45" pageNumber="46">India</collectingCountry>
: 
<collectingRegion id="1F1CF826FFDB892B9E0BFB51FE16FB70" box="[266,495,1248,1274]" country="India" name="Arunachal Pradesh" pageId="45" pageNumber="46">Arunachal Pradesh</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9EFDFB50FDB6FB71" box="[508,591,1249,1275]" country="India" name="Assam" pageId="45" pageNumber="46">Assam</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9D5FFB51FD58FB70" box="[606,673,1248,1274]" country="India" name="Bihar" pageId="45" pageNumber="46">Bihar</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9DB1FB51FCB0FB70" box="[688,841,1248,1274]" country="India" name="Chhattisgarh" pageId="45" pageNumber="46">Chhattisgarh</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9C56FB51FC4BFB70" box="[855,946,1248,1274]" country="India" name="Gujarat" pageId="45" pageNumber="46">Gujarat</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9CC0FB50FBDEFB71" box="[961,1063,1249,1275]" country="India" name="Haryana" pageId="45" pageNumber="46">Haryana</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9B37FB51FAEAFB70" box="[1078,1299,1248,1274]" country="India" name="Himachal Pradesh" pageId="45" pageNumber="46">Himachal Pradesh</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9A23FB50FEA2FA94" country="India" name="Jammu and Kashmir" pageId="45" pageNumber="46">Jammu and Kashmir</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9E6AFAB5FE1EFA94" box="[363,487,1284,1310]" country="India" name="Jharkhand" pageId="45" pageNumber="46">Jharkhand</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9EF6FAB5FD8AFA94" box="[503,627,1284,1310]" country="India" name="Karnataka" pageId="45" pageNumber="46">Karnataka</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9D85FAB5FD2CFA94" box="[644,725,1284,1310]" country="India" name="Kerala" pageId="45" pageNumber="46">Kerala</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9DE7FAB5FC4BFA94" box="[742,946,1284,1310]" country="India" name="Madhya Pradesh" pageId="45" pageNumber="46">Madhya Pradesh</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9CC2FAB5FBA0FA94" box="[963,1113,1284,1310]" country="India" name="Maharashtra" pageId="45" pageNumber="46">Maharashtra</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9B68FAB5FB17FA94" box="[1129,1262,1284,1310]" country="India" name="Meghalaya" pageId="45" pageNumber="46">Meghalaya</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9BFEFAB5FA8BFA94" box="[1279,1394,1284,1310]" country="India" name="Nagaland" pageId="45" pageNumber="46">Nagaland</collectingRegion>
, New 
<collectingRegion id="1F1CF826FFDB892B9FFFFA96FEBBFACB" box="[254,322,1319,1345]" country="India" name="Delhi" pageId="45" pageNumber="46">Delhi</collectingRegion>
, Punjab, 
<collectingRegion id="1F1CF826FFDB892B9EB0FA96FDD1FACB" box="[433,552,1319,1345]" country="India" name="Rajasthan" pageId="45" pageNumber="46">Rajasthan</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9D34FA96FD3CFACB" box="[565,709,1319,1345]" country="India" name="Tamil Nadu" pageId="45" pageNumber="46">Tamil Nadu</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9DD3FA96FCD5FACB" box="[722,812,1319,1345]" country="India" name="Tripura" pageId="45" pageNumber="46">Tripura</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9C38FA96FC34FACB" box="[825,973,1319,1345]" country="India" name="Uttarakhand" pageId="45" pageNumber="46">Uttarakhand</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9CDAFA96FB79FACB" box="[987,1152,1319,1345]" country="India" name="Uttar Pradesh" pageId="45" pageNumber="46">Uttar Pradesh</collectingRegion>
, 
<collectingRegion id="1F1CF826FFDB892B9B8CFA96FADEFACB" box="[1165,1319,1319,1345]" country="India" name="West Bengal" pageId="45" pageNumber="46">West Bengal</collectingRegion>
. Other countries: 
<collectingCountry id="A5CF7654FFDB892B9E39FAFBFE33FAEE" box="[312,458,1354,1380]" name="Afghanistan" pageId="45" pageNumber="46">Afghanistan</collectingCountry>
, 
<collectingCountry id="A5CF7654FFDB892B9ED9FAFBFD9DFAEE" box="[472,612,1354,1380]" name="Bangladesh" pageId="45" pageNumber="46">Bangladesh</collectingCountry>
, 
<collectingCountry id="A5CF7654FFDB892B9D70FAFBFD14FAEE" box="[625,749,1354,1380]" name="Cambodia" pageId="45" pageNumber="46">Cambodia</collectingCountry>
, 
<collectingCountry id="A5CF7654FFDB892B9DF8FAFBFCBBFAEE" box="[761,834,1354,1380]" name="China" pageId="45" pageNumber="46">China</collectingCountry>
, 
<collectingCountry id="A5CF7654FFDB892B9C51FAFBFC3CFAEE" box="[848,965,1354,1380]" name="Indonesia" pageId="45" pageNumber="46">Indonesia</collectingCountry>
, Malay Peninsula, 
<collectingCountry id="A5CF7654FFDB892B9BA9FAFAFAE5FAEF" box="[1192,1308,1355,1381]" name="Myanmar" pageId="45" pageNumber="46">Myanmar</collectingCountry>
, 
<collectingCountry id="A5CF7654FFDB892B9A28FAFBFA8BFAEE" box="[1321,1394,1354,1380]" name="Nepal" pageId="45" pageNumber="46">Nepal</collectingCountry>
, 
<collectingCountry id="A5CF7654FFDB892B9FBCFADCFEDAFA0D" box="[189,291,1389,1415]" name="Pakistan" pageId="45" pageNumber="46">Pakistan</collectingCountry>
, 
<collectingCountry id="A5CF7654FFDB892B9E2EFADCFE5CFA0D" box="[303,421,1389,1415]" name="Sri Lanka" pageId="45" pageNumber="46">Sri Lanka</collectingCountry>
, 
<collectingCountry id="A5CF7654FFDB892B9EB0FADCFDE5FA0D" box="[433,540,1389,1415]" name="Thailand" pageId="45" pageNumber="46">Thailand</collectingCountry>
, 
<collectingCountry id="A5CF7654FFDB892B9D29FADCFD76FA0D" box="[552,655,1389,1415]" name="Vietnam" pageId="45" pageNumber="46">Vietnam</collectingCountry>
. (
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; 
<bibRefCitation id="B9494B35FFDB892B9CF7FADCFB66FA0D" author="Chandra K." box="[1014,1183,1389,1415]" pageId="45" pageNumber="46" pagination="359 - 362" refId="ref28226" refString="Chandra K. 2000. Inventory of scarabaeid beetles (Coleoptera) from Madhya Pradesh India. Zoos' Print Journal 15 (11): 359 - 362. https: // doi. org / 10.11609 / JoTT. ZPJ. 15.11.359 - 62" type="journal article" year="2000">Chandra 2000</bibRefCitation>
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).
</paragraph>
</subSubSection>
</treatment>
</document>