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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.199.2631" ID-GBIF-Dataset="a65a90f9-192a-41a7-9f33-10f5f2f2bab2" ID-PMC="PMC3368281" ID-Pensoft-Pub="1313-2970-199-23" ID-PubMed="22711995" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-199-23" ModsDocOrigin="ZooKeys 199" ModsDocTitle="Taxonomy and species-groups of the subgenus Crematogaster ( Orthocrema) in the Malagasy region (Hymenoptera, Formicidae)" checkinTime="1451248974654" checkinUser="pensoft" docAuthor="Blaimer, Bonnie B." docDate="2012" docId="3C901B7F5D87ED1CDC8DF6014FB14009" docLanguage="en" docName="ZooKeys 199: 23-70" docOrigin="ZooKeys 199" docSource="http://dx.doi.org/10.3897/zookeys.199.2631" docTitle="Crematogaster madecassa Emery" docType="treatment" docVersion="3" lastPageNumber="42" masterDocId="FFE8E629DF34847E021D117DFFA0FFA1" masterDocTitle="Taxonomy and species-groups of the subgenus Crematogaster (Orthocrema) in the Malagasy region (Hymenoptera, Formicidae)" masterLastPageNumber="70" masterPageNumber="23" pageNumber="39" updateTime="1668153978640" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Taxonomy and species-groups of the subgenus Crematogaster (Orthocrema) in the Malagasy region (Hymenoptera, Formicidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Blaimer, Bonnie B.</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>199</mods:number>
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<mods:start>23</mods:start>
<mods:end>70</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.199.2631</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.199.2631</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-199-23</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152035990" LSID="urn:lsid:plazi:treatment:3C901B7F5D87ED1CDC8DF6014FB14009" httpUri="http://treatment.plazi.org/id/3C901B7F5D87ED1CDC8DF6014FB14009" lastPageId="19" lastPageNumber="42" pageId="16" pageNumber="39">
<subSubSection pageId="16" pageNumber="39" type="nomenclature">
<paragraph pageId="16" pageNumber="39">
<taxonomicName LSID="http://species-id.net/wiki/Crematogaster_madecassa" authority="Emery" class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster madecassa" order="Hymenoptera" pageId="16" pageNumber="39" phylum="Arthropoda" rank="species" species="madecassa">Crematogaster madecassa Emery</taxonomicName>
Figures 2728
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="39" type="reference_group">
<paragraph pageId="16" pageNumber="39">
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster sordidula" order="Hymenoptera" pageId="16" pageNumber="39" phylum="Arthropoda" rank="species" species="sordidula">Crematogaster sordidula</taxonomicName>
var. madecassa Emery, 1895: 342. Worker and queen syntypes from MADAGASCAR: Diego-Suarez (Ch. Alluaud) [MSNG, examined]. Combination in
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster" order="Hymenoptera" pageId="16" pageNumber="39" phylum="Arthropoda" rank="genus">Crematogaster</taxonomicName>
(
<taxonomicName class="Insecta" family="Formicidae" genus="Orthocrema" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Orthocrema" order="Hymenoptera" pageId="16" pageNumber="39" phylum="Arthropoda" rank="genus">Orthocrema</taxonomicName>
): Wheeler, W.M. 1922:1024. Subspecies of sordidula: Wheeler, W.M. 1922:1024. Raised to species: Emery, 1912: 668; Emery, 1922:131.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="39" type="type material examined">
<paragraph pageId="16" pageNumber="39">Type material examined</paragraph>
<paragraph pageId="16" pageNumber="39">(MSNG). MADAGASCAR: Antsiranana: Diego-Suarez: [-12.26670, 49.28330] (Ch. Alluaud), CASENT0102053, CASENT0102054 and CASENT0101933. Lectotype worker by present designation: lower specimen of 2 workers on one pin, CASENT0102054 (image on AntWeb).</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="41" pageId="16" pageNumber="39" type="other material examined">
<paragraph pageId="16" pageNumber="39">Other material examined</paragraph>
<paragraph lastPageId="18" lastPageNumber="41" pageId="16" pageNumber="39">
(CASC, PSWC, MSNG, MCZC). MADAGASCAR: Antsiranana: Sakalava Beach: -12.26278, 49.39750, 10m (R.
<normalizedToken originalValue="HarinHala">Harin'Hala</normalizedToken>
); 7 km N Joffreville: -12.33333, 49.25000, 360m (R.
<normalizedToken originalValue="HarinHala">Harin'Hala</normalizedToken>
); R.S. Ambre: -12.46889, 49.24217, 325m (B.L.Fisher et al.); P.N. Montagne
<normalizedToken originalValue="dAmbre">d'Ambre</normalizedToken>
: -12.50035, 49.17500, 885m; -12.53444, 49.17950, 925m (B.L.Fisher et al.); R.S. Manongarivo: -13.96167, 48.43333, 400m; -13.97667, 48.42333, 780m; -13.99833, 48.42833, 1175m (B.L.Fisher et al.); Ampasindava, Ambilanivy: -13.79861, 48.16167, 600m (B.L.Fisher et al.); Nosy
<normalizedToken originalValue="Bé">Be</normalizedToken>
, R.N.I.
<normalizedToken originalValue="Lokobé">Lokobe</normalizedToken>
: -13.41944, 48.33117, 30m (B.L.Fisher et al.); F Andavakoera: -13.11833, 49.23000, 425m (B.L.Fisher et al.); F Antsahabe: -13.21167, 49.55667, 550m (B.L.Fisher et al.); F Binara: -13.25500, 49.61667, 375m; -13.26333, 49.60333, 650-800m (B.L.Fisher et al.); F Analabe: -13.08333, 49.90833, 30m (B.L.Fisher et al.); F Bekaraoka: -13.16667, 49.71000; 150m (B.L.Fisher et al.); F Ampondrabe: -12.97000, 49.70000, 175m (B.L.Fisher et al.); Montagne
<normalizedToken originalValue="dAkirindro">d'Akirindro</normalizedToken>
: -15.28833, 49.54833, 600m (B.L.Fisher et al.); 6.9 km NE Ambanizana: -15.56667, 50.00000, 825m (B.L.Fisher et al.); Montagne
<normalizedToken originalValue="dAnjanaharibe">d'Anjanaharibe</normalizedToken>
: -15.18833, 49.61500, 470-1100m (B.L.Fisher et al.); P.N. Marojejy: -14.43333, 49.78333, 450m; -14.43817, 49.77400, 488m; -14.43500, 49.76000, 775m (B.L.Fisher et al.); P.N. Marojejy [Manantenina]: -14.43667, 49.77500, 450m (B.L.Fisher et al.); R.N.I. Marojejy: -14.43583, 49.76056, 610m (G. Alpert); F Ambanitaza: -14.67933, 50.18367, 240m (B.L.Fisher et al.); F Betaolana: -14.52996, 49.44039, 880m (B.L.Fisher et al.); P.N. Ankarana: -12.86361, 49.22583, 210m (B.L.Fisher); F Ambato: -13.46450, 48.55167, 150m (B.L.Fisher); F Anabohazo: -14.30889, 47.91433, 120m (B.L.Fisher et al.); 30km N Antalaha, Amboangy, -14.66480, 50.19070, 130m (G.Alpert); Fianarantsoa: P.N. Andringitra: -22.23333, 47.0000, 825m (B.L.Fisher et al.); F Vevembe: -22.79100, 47.18183, 600m
<pageBreakToken pageId="17" pageNumber="40" start="start">(</pageBreakToken>
B.L.Fisher et al.);
<normalizedToken originalValue="Rés">Res</normalizedToken>
. Marotandrano: -16.28322, 48.81443, 865m (B.L.Fisher et al.); R.S. Manombo: -23.01580, 47.71900, 30m (B.L.Fisher et al.); Mahabo [
<normalizedToken originalValue="Rés">Res</normalizedToken>
.
<normalizedToken originalValue="Forestière">Forestiere</normalizedToken>
<normalizedToken originalValue="dAgnalazaha">d'Agnalazaha</normalizedToken>
]: -23.19383, 47.72300, 20m (B.L.Fisher et al.); Mahajanga: PN Ankarafantsika (Ampijora): -16.32083, 46.81067, 130m (B.L.Fisher et al.); Toamasina: P.N. Mananara-Nord: -16.45500, 49.78750, 225m (B.L.Fisher et al.); RS Ambatovaky: -16.81739, 49.29402, 360m; -16.77274, 49.26551, 450m; -16.81209, 49.29216, 460m; -16.77020, 49.26638, 470m; -16.76330, 49.26692, 520m (B.L.Fisher et al); F Ambatovy: -18.84950, 48.29470, 1010m; F.C. Sandranantitra: -18.04833, 49.09167, 450m (B.L.Fisher et al.);
<normalizedToken originalValue="Rés">Res</normalizedToken>
. Betampona: -17.92400, 49.19967, 390m; -17.88667, 49.20250, 520m (B.L.Fisher et al.); F Kalalao [Ile St.Marie]: -16.92250, 49.88733, 100m (B.L.Fisher et al.); F Sahafina: -18.81445, 48.96205, 100m;
<normalizedToken originalValue="Rés">Res</normalizedToken>
. Ambodiriana: -16.67233, 49.70117, 125m (B.L.Fisher et al.);
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="dAnalava">d'Analava</normalizedToken>
Mandrisy: -16.48567, 49.84700, 10m (B.L.Fisher et al.); S.F. Tampolo: -17.28250, 49.43000, 10m (B.L.Fisher et al.); Toliara: F Ivohibe: -24.56900, 47.20400, 200m (B.L.Fisher et al.); P.N. Andohahela: -24.75850, 46.85370, 275m (B.L.Fisher et al.); 10km NW Enakara,
<normalizedToken originalValue="Rés">Res</normalizedToken>
. Andohahela: -24.56667, 46.81667, 430m; 11km NW Enakara,
<normalizedToken originalValue="Rés">Res</normalizedToken>
. Andohahela: -24.56667, 46.83333, 800m (B.L.Fisher); 6km SSW Eminiminy,
<normalizedToken originalValue="Rés">Res</normalizedToken>
. Andohahela: -24.75000, 46.78330, 500m (P.S.Ward); 9km SSW Eminiminy,
<normalizedToken originalValue="Rés">Res</normalizedToken>
. Andohahela: -24.73330, 46.80000, 330m (P.S.Ward); 2.7km WNW 302
<normalizedToken originalValue="º">°</normalizedToken>
St.Luce: -24.77167, 47.17167, 20m (B.L.Fisher et al.); F Mandena: -24.95167, 47.00167, 20m (B.L.Fisher)
<pageBreakToken pageId="18" pageNumber="41" start="start">.</pageBreakToken>
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="41" type="worker measurements">
<paragraph pageId="18" pageNumber="41">Worker measurements</paragraph>
<paragraph pageId="18" pageNumber="41">(n=21). Lectotype worker: HW 0.54; HL 0.51; EL 0.13; SL 0.43; WL 0.54; SPL 0.13; PTH n.a.; PTL 0.17; PTW 0.16; PPL 0.10; PPW 0.17; LHT 0.40; CI 1.05; OI 0.26; SI 0.85; SPI 0.23; PTHI n.a.; PTWI 0.95; PPI 1.67; LBI 1.36.</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="41" type="other material">
<paragraph pageId="18" pageNumber="41">Other material.</paragraph>
<paragraph pageId="18" pageNumber="41">HW 0.48-0.60; HL 0.43-0.51; EL 0.10-0.14; SL 0.37-0.51; WL 0.44-0.63; SPL 0.08-0.14; PTH 0.09-0.14; PTL 0.14-0.22; PTW 0.12-0.22; PPL 0.09-0.14; PPW 0.13-0.21; LHT 0.33-0.52; CI 1.00-1.12; OI 0.22-0.27; SI 0.85-1.01; SPI 0.17-0.26; PTHI 0.55-0.72; PTWI 0.78-1.04; PPI 1.35-1.98; LBI 1.22-1.48.</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="41" type="diagnosis">
<paragraph pageId="18" pageNumber="41">Diagnosis.</paragraph>
<paragraph pageId="18" pageNumber="41">
Workers of
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster madecassa" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="madecassa">Crematogaster madecassa</taxonomicName>
can be distinguished from all other species treated here except
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster telolafy" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="telolafy">Crematogaster telolafy</taxonomicName>
by the presence of two distinct vertical carinae on the clypeus (Figure 16). From workers of
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster telolafy" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="telolafy">Crematogaster telolafy</taxonomicName>
,
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster madecassa" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="madecassa">Crematogaster madecassa</taxonomicName>
workers are distinguishable by their longer, more spiniform propodeal spines (Figure 18A) and longer antennal scapes. Queens of
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster madecassa" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="madecassa">Crematogaster madecassa</taxonomicName>
can be easily identified from queens of most species (
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster rasoherinae" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="rasoherinae">Crematogaster rasoherinae</taxonomicName>
,
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster volamena" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="volamena">Crematogaster volamena</taxonomicName>
,
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster mpanjono" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="mpanjono">Crematogaster mpanjono</taxonomicName>
) by the presence of propodeal spines.
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster madecassa" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="madecassa">Crematogaster madecassa</taxonomicName>
queens can be distinguished from
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster razana" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="razana">Crematogaster razana</taxonomicName>
queens by the absence of a median clypeal notch (present in
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster razana" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="razana">Crematogaster razana</taxonomicName>
queens). Note however that queens of
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster telolafy" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="telolafy">Crematogaster telolafy</taxonomicName>
are currently unknown and could be very similar morphologically to
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster madecassa" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="madecassa">Crematogaster madecassa</taxonomicName>
queens.
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="41" type="worker description">
<paragraph pageId="18" pageNumber="41">Worker description</paragraph>
<paragraph pageId="18" pageNumber="41">
(Figure 26
<normalizedToken originalValue="AC">A-C</normalizedToken>
). Very small species (HW 0.48-0.60, WL 0.44-0.63), with characters of the
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster madecassa" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="madecassa">Crematogaster madecassa</taxonomicName>
-group, in addition to the following. Clypeus with two distinct median vertical carinae; antennal scapes well surpassing posterior margin of head.
</paragraph>
<paragraph pageId="18" pageNumber="41">Metanotal groove constricted to less than half as wide as pronotal width; propodeal spines medium-sized (SPI 0.17-0.26) and straight, usually thin and acute, in lateral view directed upwards, in dorsal view moderately diverging; subpostpetiolar process usually present, often as acute minute tooth.</paragraph>
<paragraph pageId="18" pageNumber="41">Promesonotum usually with 6 erect, long flexuous setae: 2 humeral setae, and 2 setae each at anterior and posterior end of mesonotal carinae. Color pale to medium yellow.</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="41" type="intermediate worker measurements">
<paragraph pageId="18" pageNumber="41">Intermediate worker measurements</paragraph>
<paragraph pageId="18" pageNumber="41">(n=2). HW 0.82-0.85, HL 0.73-0.78, EL 0.18-0.19, SL 0.55-0.59, WL 0.89-1.01, SPL 0.20, PTH 0.18-0.19, PTL 0.31-0.33, PTW 0.27-0.28, PPL 0.19, PPW 0.28-0.31, LHT 0.60, CI 1.09-1.12, OI 0.24, SI 0.75, SPI 0.20-0.22, PTHI 0.59, PTWI 0.86, PPI 1.46-1.66, LBI 1.48-1.68.</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="41" type="intermediate worker description">
<paragraph pageId="18" pageNumber="41">Intermediate worker description</paragraph>
<paragraph pageId="18" pageNumber="41">
(Figures 27
<normalizedToken originalValue="EG">E-G</normalizedToken>
). Intermediate between workers and queens in size. Head, petiole and postpetiole characters similar to queens; ocelli present, but smaller than in queens; mesonotum is to various extent raised and fused dorsally over pronotum and has wing attachment sutures; otherwise mesosomal characters more similar to worker characters.
</paragraph>
<caption pageId="18" pageNumber="41">
<paragraph pageId="18" pageNumber="41">
Figure 27.
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster madecassa" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="madecassa">Crematogaster madecassa</taxonomicName>
, worker, intermorph and distribution.
<normalizedToken originalValue="AC">A-C</normalizedToken>
worker (CASENT0038498) A full face B dorsal C lateral D distribution
<normalizedToken originalValue="EG">E-G</normalizedToken>
intermorph (CASENT0436247) E full face F lateral G dorsal.
</paragraph>
</caption>
</subSubSection>
<subSubSection lastPageId="19" lastPageNumber="42" pageId="18" pageNumber="41" type="queen measurements">
<paragraph pageId="18" pageNumber="41">Queen measurements</paragraph>
<paragraph pageId="18" pageNumber="41">(n=11). HW 0.87-1.03, HL 0.72-0.86, EL 0.24-0.32, SL 0.54-0.62, MSNW 0.66-0.92, MSNL 0.72-0.98, WL 1.28-1.53, SPL 0.06-0.21, PTH 0.19-0.26, PTL 0.38-0.49, PTW 0.30-0.39, PPL 0.21-0.28, PPW 0.34-0.42, LHT 0.65-0.81, CI 1.11-1.21, OI 0.29-0.37, SI 0.68-0.76, MSNI 1.55-1.87, SPI 0.04-0.14, PTHI 0.45-0.61, PTWI 0.68-0.91, PPI 1.42-1.69, LBI 1.87-2.08.</paragraph>
<paragraph pageId="18" pageNumber="41">
Queen description (Figure 28
<normalizedToken originalValue="AC">A-C</normalizedToken>
). Very small (HW 0.87-1.03, WL 1.28-1.53), with characters of the
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster madecassa" order="Hymenoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="madecassa">Crematogaster madecassa</taxonomicName>
-group, in addition to the following.
</paragraph>
<paragraph pageId="19" pageNumber="42">
<pageBreakToken pageId="19" pageNumber="42" start="start">Mesosoma</pageBreakToken>
more compact (MSNI 1.55-1.87, WL 1.28-1.53), mesoscutum in dorsal view almost as wide as long; propodeal spines present, much shorter than in workers (SPI 0.04-0.14).
</paragraph>
<paragraph pageId="19" pageNumber="42">Male unknown.</paragraph>
<caption pageId="19" pageNumber="42">
<paragraph pageId="19" pageNumber="42">
Figure 28.
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster madecassa" order="Hymenoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="madecassa">Crematogaster madecassa</taxonomicName>
, queen. A full face, B lateral, C dorsal (CASENT0040391).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="19" pageNumber="42" type="distribution">
<paragraph pageId="19" pageNumber="42">Distribution and biology.</paragraph>
<paragraph pageId="19" pageNumber="42">
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster madecassa" order="Hymenoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="madecassa">Crematogaster madecassa</taxonomicName>
is fairly widespread throughout the rainforests and littoral forests of northern, eastern and southeastern Madagascar (Figure 27D) and is found up to medium elevations (highest record from 1175m). It occurs widely in sympatry with
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster rasoherinae" order="Hymenoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="rasoherinae">Crematogaster rasoherinae</taxonomicName>
, and at a few localities also with
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster volamena" order="Hymenoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="volamena">Crematogaster volamena</taxonomicName>
,
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster mpanjono" order="Hymenoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="mpanjono">Crematogaster mpanjono</taxonomicName>
and
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster telolafy" order="Hymenoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="telolafy">Crematogaster telolafy</taxonomicName>
. There is evidence that
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster madecassa" order="Hymenoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="madecassa">Crematogaster madecassa</taxonomicName>
nests both on the ground and arboreally, as collections have been made from rotten logs, as well as from dead twigs above the ground. As in
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster rasoherinae" order="Hymenoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="rasoherinae">Crematogaster rasoherinae</taxonomicName>
(see above), intermediate workers are found in
<taxonomicName class="Insecta" family="Formicidae" genus="Crematogaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crematogaster madecassa" order="Hymenoptera" pageId="19" pageNumber="42" phylum="Arthropoda" rank="species" species="madecassa">Crematogaster madecassa</taxonomicName>
(Figure 27
<normalizedToken originalValue="EF">E-F</normalizedToken>
).
</paragraph>
</subSubSection>
</treatment>
</document>