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<document id="0C0558A6AD522753178B38E0C1B60169" ID-DOI="10.5281/zenodo.185971" ID-GBIF-Dataset="1db39382-b4f0-46fa-ae08-dd5e57e0e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="185971" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460034098030" checkinUser="plazi" docAuthor="Pimvichai, Piyatida, Enghoff, Henrik &amp; Panha, Somsak" docDate="2009" docId="1644D538F450FFD6FF49F9519C11FE0D" docLanguage="en" docName="zt02016p050.pdf" docOrigin="Zootaxa 2016" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Thyropygus bispinispatula Pimvichai, Enghoff &amp; Panha, 2009, n. sp." docType="treatment" docVersion="11" lastPageNumber="37" masterDocId="EA7DAD40F442FFC2FFDEFFED9F66FFCD" masterDocTitle="A revision of the Thyropygus allevatus group. Part 1: the T. opinatus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)" masterLastPageNumber="50" masterPageNumber="17" pageNumber="35" updateTime="1698234296060" updateUser="plazi">
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<mods:title id="0803F5533E9FABFEC3EFF4759DB7312C">A revision of the Thyropygus allevatus group. Part 1: the T. opinatus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)</mods:title>
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<mods:name id="2C248621A209283C39BC24BA4B109F5D" type="personal">
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<mods:namePart id="BF0BC7FB8BDADFD9DB0ED353B60CD1CB">Pimvichai, Piyatida</mods:namePart>
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<mods:namePart id="EA6BDAB6ED757EEBB45A293BA5ADC5C9">Enghoff, Henrik</mods:namePart>
</mods:name>
<mods:name id="906A0FEA7C63926FB6C4DA304008F8EC" type="personal">
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<mods:namePart id="2597525DD68184E4A1CC00546648C265">Panha, Somsak</mods:namePart>
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<mods:typeOfResource id="AE27569E163730922C93C23EEEAC23C4">text</mods:typeOfResource>
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<mods:title id="3231C501BF56226A96711F19535E9AEA">Zootaxa</mods:title>
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<mods:part id="E008BD5906692E7ACB54469CD4762C9D">
<mods:date id="9E98EDC06A122CB6013C74C4AC9559F4">2009</mods:date>
<mods:detail id="DF391113523D0C1A72A397CB10FC4C6A" type="volume">
<mods:number id="CB651342598613B69565E4CC72CD1F6E">2016</mods:number>
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<mods:identifier id="44625F91E90B82C2BA718FDBF0D52C90" type="DOI">10.5281/zenodo.185971</mods:identifier>
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<treatment id="1644D538F450FFD6FF49F9519C11FE0D" ID-DOI="http://doi.org/10.5281/zenodo.6218628" ID-GBIF-Taxon="119382577" ID-Zenodo-Dep="6218628" LSID="urn:lsid:plazi:treatment:1644D538F450FFD6FF49F9519C11FE0D" httpUri="http://treatment.plazi.org/id/1644D538F450FFD6FF49F9519C11FE0D" lastPageId="20" lastPageNumber="37" pageId="18" pageNumber="35">
<subSubSection id="D6F737A5F450FFD0FF49F9519EE6F934" pageId="18" pageNumber="35" type="nomenclature">
<paragraph id="9E52642EF450FFD0FF49F9519D46F91B" blockId="18.[151,544,1724,1785]" box="[151,544,1724,1750]" pageId="18" pageNumber="35">
<heading id="C51AD342F450FFD0FF49F9519D46F91B" bold="true" box="[151,544,1724,1750]" fontSize="11" level="1" pageId="18" pageNumber="35" reason="1">
<emphasis id="AC99B83CF450FFD0FF49F9519D46F91B" bold="true" box="[151,544,1724,1750]" pageId="18" pageNumber="35">
<taxonomicName id="59ED1FADF450FFD0FF49F9519EB0F91B" ID-CoL="56SYV" box="[151,470,1724,1750]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="18" pageNumber="35" phylum="Arthropoda" rank="species" species="bispinispatula" status="sp. nov.">
<emphasis id="AC99B83CF450FFD0FF49F9519EB0F91B" bold="true" box="[151,470,1724,1750]" italics="true" pageId="18" pageNumber="35">Thyropygus bispinispatula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7AA0547F450FFD0FE00F9519D46F91B" box="[478,544,1724,1750]" pageId="18" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9E52642EF450FFD0FF49F9329EE6F934" blockId="18.[151,544,1724,1785]" box="[151,384,1759,1785]" pageId="18" pageNumber="35">
(
<figureCitation id="06D678ABF450FFD0FF41F9329F9DF934" box="[159,251,1759,1785]" captionStart="FIGURE 11" captionStartId="19.[151,257,1579,1603]" captionTargetBox="[278,1326,209,1549]" captionTargetId="figure@19.[226,1360,194,1555]" captionTargetPageId="19" captionText="FIGURE 11. Thyropygus bispinispatula, holotype, gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C: left telopodite, anterior view. D: left telopodite, posterior view." httpUri="https://zenodo.org/record/185982/files/figure.png" pageId="18" pageNumber="35">Figs. 11</figureCitation>
AD, 18C)
</paragraph>
</subSubSection>
<subSubSection id="D6F737A5F450FFD0FF49F8C19BF6F859" pageId="18" pageNumber="35" type="materials_examined">
<paragraph id="9E52642EF450FFD0FF49F8C19BF6F859" blockId="18.[151,1436,1836,2017]" pageId="18" pageNumber="35">
<emphasis id="AC99B83CF450FFD0FF49F8C19E68F88B" bold="true" box="[151,270,1836,1862]" pageId="18" pageNumber="35">Material:</emphasis>
<typeStatus id="4156DA8CF450FFD0FECAF8C19EC9F88B" box="[276,431,1836,1862]" pageId="18" pageNumber="35" type="holotype">HOLOTYPE</typeStatus>
male
<collectingCountry id="E6FA24BEF450FFD0FE28F8C19DF7F88B" box="[502,657,1836,1862]" name="Thailand" pageId="18" pageNumber="35">THAILAND</collectingCountry>
, Chumphon Province, Lang Suan district, Khao Krieab Temple, 9° 49ˏ 4˝ N, 99° 2ˏ 17˝ E.
<date id="EA5342EEF450FFD0FE6DF8BE9DE7F8A0" box="[435,641,1875,1901]" pageId="18" pageNumber="35" value="2008-10-11">11 October 2008</date>
. P. Prasankok and members of Animal Systematics Research Unit leg., (
<collectionCode id="F8FCFCEBF450FFD0FF07F8979E52F859" box="[217,308,1914,1940]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="18" pageNumber="35">CUMZ</collectionCode>
).
<typeStatus id="4156DA8CF450FFD0FE80F8979EB2F859" box="[350,468,1914,1940]" pageId="18" pageNumber="35" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="88EBAFA7F450FFD0FE01F8979D36F859" box="[479,592,1914,1940]" pageId="18" pageNumber="35" type="female">2 females</specimenCount>
and
<specimenCount id="88EBAFA7F450FFD0FD57F8979C6AF859" box="[649,780,1914,1940]" pageId="18" pageNumber="35" type="juvenile">3 juveniles</specimenCount>
, same data as
<typeStatus id="4156DA8CF450FFD0FC6CF8979B7FF859" box="[946,1049,1914,1940]" pageId="18" pageNumber="35" type="holotype">holotype</typeStatus>
(
<collectionCode id="F8FCFCEBF450FFD0FBF6F8979BE2F859" box="[1064,1156,1914,1940]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="18" pageNumber="35">CUMZ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D6F737A5F450FFD0FF1BF84D9ECBF82C" pageId="18" pageNumber="35" type="etymology">
<paragraph id="9E52642EF450FFD0FF1BF84D9ECBF82C" blockId="18.[151,1436,1836,2017]" pageId="18" pageNumber="35">
<emphasis id="AC99B83CF450FFD0FF1BF84D9E31F877" bold="true" box="[197,343,1952,1978]" pageId="18" pageNumber="35">Etymology:</emphasis>
The species epithet is a Latin noun in apposition and refers to the spinose spatulate lobe of the gonopod telopodite.
</paragraph>
</subSubSection>
<caption id="CA9234A6F451FFD1FF49F9C69BFCF9A9" httpUri="https://zenodo.org/record/185982/files/figure.png" pageId="19" pageNumber="36" targetBox="[278,1326,209,1549]" targetPageId="19">
<paragraph id="9E52642EF451FFD1FF49F9C69BFCF9A9" blockId="19.[151,1436,1579,1636]" pageId="19" pageNumber="36">
<emphasis id="AC99B83CF451FFD1FF49F9C69E4EF98E" bold="true" box="[151,296,1579,1603]" pageId="19" pageNumber="36">FIGURE 11.</emphasis>
<taxonomicName id="59ED1FADF451FFD1FEEEF9C69D34F98F" box="[304,594,1579,1602]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="36" phylum="Arthropoda" rank="species" species="bispinispatula">
<emphasis id="AC99B83CF451FFD1FEEEF9C69D34F98F" box="[304,594,1579,1602]" italics="true" pageId="19" pageNumber="36">Thyropygus bispinispatula</emphasis>
</taxonomicName>
, holotype, gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C: left telopodite, anterior view. D: left telopodite, posterior view.
</paragraph>
</caption>
<subSubSection id="D6F737A5F451FFD1FF1BF97A9BAFF915" pageId="19" pageNumber="36" type="diagnosis">
<paragraph id="9E52642EF451FFD1FF1BF97A9BAFF915" blockId="19.[151,1436,1687,2022]" pageId="19" pageNumber="36">
<emphasis id="AC99B83CF451FFD1FF1BF97A9E21F97C" bold="true" box="[197,327,1687,1713]" pageId="19" pageNumber="36">Diagnosis:</emphasis>
A species of the
<taxonomicName id="59ED1FADF451FFD1FDD1F97A9D12F97D" box="[527,628,1687,1712]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="36" phylum="Arthropoda" rank="species" species="opinatus">
<emphasis id="AC99B83CF451FFD1FDD1F97A9D12F97D" box="[527,628,1687,1712]" italics="true" pageId="19" pageNumber="36">opinatus</emphasis>
</taxonomicName>
subgroup. Differing from all other species in the subgroup by having the spatulate lobe (
<emphasis id="AC99B83CF451FFD1FEABF9539EEFF91A" box="[373,393,1726,1751]" italics="true" pageId="19" pageNumber="36">sl</emphasis>
) at the apical part of telopodite terminating in two sharp brown spines.
</paragraph>
</subSubSection>
<subSubSection id="D6F737A5F451FFD6FF1BF9099C08FE57" lastPageId="20" lastPageNumber="37" pageId="19" pageNumber="36" type="description">
<paragraph id="9E52642EF451FFD1FF1BF9099F8CF8BF" blockId="19.[151,1436,1687,2022]" pageId="19" pageNumber="36">
<emphasis id="AC99B83CF451FFD1FF1BF9099E06F933" bold="true" box="[197,352,1764,1790]" pageId="19" pageNumber="36">Description:</emphasis>
Adult males with 67 podous rings, no apodous rings. Length ca.
<quantity id="5915C9CBF451FFD1FBB5F9099BD1F933" box="[1131,1207,1764,1790]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="19" pageNumber="36" unit="cm" value="12.0">12 cm</quantity>
, width ca.
<quantity id="5915C9CBF451FFD1FAE7F9099AFEF933" box="[1337,1432,1764,1790]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="19" pageNumber="36" unit="mm" value="7.4">7.4 mm</quantity>
. Adult females with 64 podous rings, no apodous rings. Length ca.
<quantity id="5915C9CBF451FFD1FC72F8E69B43F8E8" box="[940,1061,1803,1829]" metricMagnitude="-1" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="19" pageNumber="36" unit="cm" value="14.5" valueMax="16.0" valueMin="13.0">1316 cm</quantity>
, width ca.
<quantity id="5915C9CBF451FFD1FB78F8E69A5EF8E8" box="[1190,1336,1803,1829]" metricMagnitude="-3" metricUnit="m" metricValue="8.45" metricValueMax="9.3" metricValueMin="7.6" pageId="19" pageNumber="36" unit="mm" value="8.45" valueMax="9.3" valueMin="7.6">7.69.3 mm</quantity>
. Overall color of living animal (
<figureCitation id="06D678ABF451FFD1FE6BF8DF9D6BF881" box="[437,525,1842,1868]" captionStart="FIGURE 18. A" captionStartId="29.[151,256,1564,1588]" captionTargetBox="[163,1421,241,1493]" captionTargetId="figure@29.[151,1436,229,1506]" captionTargetPageId="29" captionText="FIGURE 18. A: Thyropygus opinatus, living male. B: Thyropygus bearti, living male. C: Thyropygus bispinispatula, living male. D: Thyropygus brachyacanthus, living male." httpUri="https://zenodo.org/record/185989/files/figure.png" pageId="19" pageNumber="36">Fig. 18</figureCitation>
C) yellowish brown. Legs and antennae brownish orange. Paraprocts dark brown.
</paragraph>
<paragraph id="9E52642EF451FFD6FF1BF8929C08FE57" blockId="19.[151,1436,1687,2022]" lastBlockId="20.[151,1436,152,448]" lastPageId="20" lastPageNumber="37" pageId="19" pageNumber="36">
Gonopods (
<figureCitation id="06D678ABF451FFD1FE8BF8929EDEF854" box="[341,440,1919,1945]" captionStart="FIGURE 11" captionStartId="19.[151,257,1579,1603]" captionTargetBox="[278,1326,209,1549]" captionTargetId="figure@19.[226,1360,194,1555]" captionTargetPageId="19" captionText="FIGURE 11. Thyropygus bispinispatula, holotype, gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C: left telopodite, anterior view. D: left telopodite, posterior view." httpUri="https://zenodo.org/record/185982/files/figure.png" pageId="19" pageNumber="36">Figs. 11</figureCitation>
AD): Anterior coxal fold (
<emphasis id="AC99B83CF451FFD1FCCCF8929C49F855" box="[786,815,1919,1944]" italics="true" pageId="19" pageNumber="36">ac</emphasis>
) (
<figureCitation id="06D678ABF451FFD1FC95F8929CC2F854" box="[843,932,1919,1945]" captionStart="FIGURE 11" captionStartId="19.[151,257,1579,1603]" captionTargetBox="[278,1326,209,1549]" captionTargetId="figure@19.[226,1360,194,1555]" captionTargetPageId="19" captionText="FIGURE 11. Thyropygus bispinispatula, holotype, gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C: left telopodite, anterior view. D: left telopodite, posterior view." httpUri="https://zenodo.org/record/185982/files/figure.png" pageId="19" pageNumber="36">Fig. 11</figureCitation>
A): lateral process (
<emphasis id="AC99B83CF451FFD1FB42F8929BA5F855" box="[1180,1219,1919,1944]" italics="true" pageId="19" pageNumber="36">alp</emphasis>
) slender, slightly curved mesad; mesal process (
<emphasis id="AC99B83CF451FFD1FDDFF84B9D52F872" box="[513,564,1958,1983]" italics="true" pageId="19" pageNumber="36">amp</emphasis>
) almost as long as
<emphasis id="AC99B83CF451FFD1FCC4F84B9C59F872" box="[794,831,1958,1983]" italics="true" pageId="19" pageNumber="36">alp</emphasis>
, straight, flattened, parallel-sided, directed straight distad. Posterior coxal fold (
<emphasis id="AC99B83CF451FFD1FE3AF8209D66F82B" box="[484,512,1997,2022]" italics="true" pageId="19" pageNumber="36">pc</emphasis>
) (
<figureCitation id="06D678ABF451FFD1FDC4F8219D0BF82B" box="[538,621,1996,2022]" captionStart="FIGURE 11" captionStartId="19.[151,257,1579,1603]" captionTargetBox="[278,1326,209,1549]" captionTargetId="figure@19.[226,1360,194,1555]" captionTargetPageId="19" captionText="FIGURE 11. Thyropygus bispinispatula, holotype, gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C: left telopodite, anterior view. D: left telopodite, posterior view." httpUri="https://zenodo.org/record/185982/files/figure.png" pageId="19" pageNumber="36">Fig. 11</figureCitation>
B) basally with moderately high lateral paracoxites (
<emphasis id="AC99B83CF451FFD1FB08F8209B94F82B" box="[1238,1266,1997,2022]" italics="true" pageId="19" pageNumber="36">px</emphasis>
), distally with two processes: mesal process (
<emphasis id="AC99B83CF456FFD6FDDBFF759D5EFF7C" box="[517,568,152,177]" italics="true" pageId="20" pageNumber="37">pmp</emphasis>
) slightly shorter than
<emphasis id="AC99B83CF456FFD6FC9EFF759C00FF7C" box="[832,870,152,177]" italics="true" pageId="20" pageNumber="37">plp</emphasis>
, and curving behind
<emphasis id="AC99B83CF456FFD6FBBAFF759BECFF7C" box="[1124,1162,152,177]" italics="true" pageId="20" pageNumber="37">plp</emphasis>
; lateral process (
<emphasis id="AC99B83CF456FFD6FA89FF759A1BFF7C" box="[1367,1405,152,177]" italics="true" pageId="20" pageNumber="37">plp</emphasis>
) a rounded lobe, curving caudad. Telopodite (
<figureCitation id="06D678ABF456FFD6FD4CFF539D89FF15" box="[658,751,190,216]" captionStart="FIGURE 11" captionStartId="19.[151,257,1579,1603]" captionTargetBox="[278,1326,209,1549]" captionTargetId="figure@19.[226,1360,194,1555]" captionTargetPageId="19" captionText="FIGURE 11. Thyropygus bispinispatula, holotype, gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C: left telopodite, anterior view. D: left telopodite, posterior view." httpUri="https://zenodo.org/record/185982/files/figure.png" pageId="20" pageNumber="37">Figs. 11</figureCitation>
CD) leaving coxite between
<emphasis id="AC99B83CF456FFD6FB8CFF529BE3FF15" box="[1106,1157,191,216]" italics="true" pageId="20" pageNumber="37">pmp</emphasis>
and
<emphasis id="AC99B83CF456FFD6FB61FF529B82FF15" box="[1215,1252,191,216]" italics="true" pageId="20" pageNumber="37">plp</emphasis>
; Femoral spine (
<emphasis id="AC99B83CF456FFD6FF7FFF089FD0FF33" box="[161,182,229,254]" italics="true" pageId="20" pageNumber="37">fe</emphasis>
) very long, directed obliquely distad, crenulated along inner curvature,
<emphasis id="AC99B83CF456FFD6FBFAFF089B16FF33" box="[1060,1136,229,254]" italics="true" pageId="20" pageNumber="37">in situ</emphasis>
resting against posterior surface of
<emphasis id="AC99B83CF456FFD6FECBFEE19E2EFEE8" box="[277,328,268,293]" italics="true" pageId="20" pageNumber="37">amp</emphasis>
; telopodite distally to
<emphasis id="AC99B83CF456FFD6FD88FEE19D0DFEE8" box="[598,619,268,293]" italics="true" pageId="20" pageNumber="37">fe</emphasis>
with a vertical, smooth lobe (
<emphasis id="AC99B83CF456FFD6FC0AFEE19C8DFEE8" box="[980,1003,268,293]" italics="true" pageId="20" pageNumber="37">lo</emphasis>
) projecting distolaterad; tibial spine (
<emphasis id="AC99B83CF456FFD6FF7FFEDF9FD7FE86" box="[161,177,306,331]" italics="true" pageId="20" pageNumber="37">ti</emphasis>
) very long, curving in horizontal plane close to the middle part of
<emphasis id="AC99B83CF456FFD6FC1AFEDF9CBFFE86" box="[964,985,306,331]" italics="true" pageId="20" pageNumber="37">fe</emphasis>
; spatulate lobe (
<emphasis id="AC99B83CF456FFD6FB43FEDF9BD6FE86" box="[1181,1200,306,331]" italics="true" pageId="20" pageNumber="37">sl</emphasis>
) terminating in two sharp brown spines, the outer spine slightly smaller and shorter than the inner one; palette (
<emphasis id="AC99B83CF456FFD6FB1DFEB49B86FEBF" box="[1219,1248,345,370]" italics="true" pageId="20" pageNumber="37">pa</emphasis>
) simple, gutterlike, distally with about seven brownish blepharochaetae (
<emphasis id="AC99B83CF456FFD6FC9FFE6D9C38FE54" box="[833,862,384,409]" italics="true" pageId="20" pageNumber="37">bp</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="D6F737A5F456FFD6FF1BFE4B9C11FE0D" box="[197,887,422,448]" pageId="20" pageNumber="37" type="distribution">
<paragraph id="9E52642EF456FFD6FF1BFE4B9C11FE0D" blockId="20.[151,1436,152,448]" box="[197,887,422,448]" pageId="20" pageNumber="37">
<emphasis id="AC99B83CF456FFD6FF1BFE4B9E38FE0D" bold="true" box="[197,350,422,448]" pageId="20" pageNumber="37">Distribution</emphasis>
(
<figureCitation id="06D678ABF456FFD6FEB0FE4B9EA0FE0D" box="[366,454,422,448]" captionStart="FIGURE 20" captionStartId="32.[151,255,1208,1232]" captionText="FIGURE 20. Known distribution of the species of the T. opinatus subgroup." pageId="20" pageNumber="37">Fig. 20</figureCitation>
): Known only from the
<typeStatus id="4156DA8CF456FFD6FD3EFE4B9C74FE0D" box="[736,786,422,448]" pageId="20" pageNumber="37">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>