A revision of the Thyropygus allevatus group. Part 1: the T. opinatus subgroup (Diplopoda: Spirostreptida: Harpagophoridae) Author Pimvichai, Piyatida Author Enghoff, Henrik Author Panha, Somsak text Zootaxa 2009 2016 17 50 journal article 10.5281/zenodo.185971 1db39382-b4f0-46fa-ae08-dd5e57e0e26b 1175-5326 185971 Thyropygus bispinispatula n. sp. ( Figs. 11 A–D, 18C) Material: HOLOTYPE male THAILAND , Chumphon Province, Lang Suan district, Khao Krieab Temple, 9° 49ˏ 4˝ N, 99° 2ˏ 17˝ E. 11 October 2008 . P. Prasankok and members of Animal Systematics Research Unit leg., ( CUMZ ). – Paratypes : 2 females and 3 juveniles , same data as holotype ( CUMZ ). Etymology: The species epithet is a Latin noun in apposition and refers to the spinose spatulate lobe of the gonopod telopodite. FIGURE 11. Thyropygus bispinispatula , holotype, gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C: left telopodite, anterior view. D: left telopodite, posterior view. Diagnosis: A species of the opinatus subgroup. Differing from all other species in the subgroup by having the spatulate lobe ( sl ) at the apical part of telopodite terminating in two sharp brown spines. Description: Adult males with 67 podous rings, no apodous rings. Length ca. 12 cm , width ca. 7.4 mm . Adult females with 64 podous rings, no apodous rings. Length ca. 13–16 cm , width ca. 7.6–9.3 mm . Overall color of living animal ( Fig. 18 C) yellowish brown. Legs and antennae brownish orange. Paraprocts dark brown. Gonopods ( Figs. 11 A–D): Anterior coxal fold ( ac ) ( Fig. 11 A): lateral process ( alp ) slender, slightly curved mesad; mesal process ( amp ) almost as long as alp , straight, flattened, parallel-sided, directed straight distad. Posterior coxal fold ( pc ) ( Fig. 11 B) basally with moderately high lateral paracoxites ( px ), distally with two processes: mesal process ( pmp ) slightly shorter than plp , and curving behind plp ; lateral process ( plp ) a rounded lobe, curving caudad. Telopodite ( Figs. 11 C–D) leaving coxite between pmp and plp ; Femoral spine ( fe ) very long, directed obliquely distad, crenulated along inner curvature, in situ resting against posterior surface of amp ; telopodite distally to fe with a vertical, smooth lobe ( lo ) projecting distolaterad; tibial spine ( ti ) very long, curving in horizontal plane close to the middle part of fe ; spatulate lobe ( sl ) terminating in two sharp brown spines, the outer spine slightly smaller and shorter than the inner one; palette ( pa ) simple, gutterlike, distally with about seven brownish blepharochaetae ( bp ). Distribution ( Fig. 20 ): Known only from the type locality.