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<document id="DA8231907467CD94DBBE8503A3EB2C8E" ID-CLB-Dataset="301510" ID-DOI="10.1371/journal.pone.0209792" ID-GBIF-Dataset="447a44cd-2c7e-4499-b31b-6e529c586ed5" ID-ISSN="1932-6203" ID-PMC="PMC6334993" ID-PubMed="30650102" ID-Zenodo-Dep="12630921" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1719975332143" checkinUser="felipe" docAuthor="Barboza, Gloria E., García, Carolina Carrizo, González, Segundo Leiva, Scaldaferro, Marisel &amp; Reyes, Ximena" docDate="2019" docId="03F887D6FFBBFFFA307EF9486265F950" docLanguage="en" docName="PLoSONE.14.1.e0209792.pdf" docOrigin="PLoS ONE (e 0209792) 14 (1)" docSource="http://dx.doi.org/10.1371/journal.pone.0209792" docStyle="DocumentStyle:55318D21378FF0528C342C9740A1E1BE.2:PLoSONE.2014-.journal_article" docStyleId="55318D21378FF0528C342C9740A1E1BE" docStyleName="PLoSONE.2014-.journal_article" docStyleVersion="2" docTitle="Capsicum benoistii Barboza 2019, sp. nov." docType="treatment" docVersion="2" lastPageNumber="7" masterDocId="FFC1FFAEFFB8FFFC326BFF8E615BFFE8" masterDocTitle="Four new species of Capsicum (Solanaceae) from the tropical Andes and an update on the phylogeny of the genus" masterLastPageNumber="26" masterPageNumber="1" pageNumber="4" updateTime="1723726385848" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2EE81266BB6374A2E1C0607834282924">Four new species of Capsicum (Solanaceae) from the tropical Andes and an update on the phylogeny of the genus</mods:title>
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<mods:namePart id="2359D2E55D0CD1B71ABA6722B88DFAEB">Barboza, Gloria E.</mods:namePart>
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<mods:namePart id="5FE1126A21ACAEBA6E8EF1027FA57910">García, Carolina Carrizo</mods:namePart>
<mods:affiliation id="BA016D92679956D8421B4A6A1CD7E85E">Instituto Multidisciplinario de Biología Vegetal (IMBIV), CONICET, Córdoba, Argentina, &amp; Department of Botany and Biodiversity Research, University of Vienna, Vienna, Austria,</mods:affiliation>
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<mods:namePart id="04CC90539741D3AA11A1BF18056C282C">González, Segundo Leiva</mods:namePart>
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<mods:namePart id="B1B87E33C9926127A51A069C2E7A6565">Scaldaferro, Marisel</mods:namePart>
<mods:affiliation id="19A9049100EE14F3E0189255FBB5B363">Instituto Multidisciplinario de Biología Vegetal (IMBIV), CONICET, Córdoba, Argentina, &amp; Facultad de Ciencias Exactas, Físicas y Naturales, Universidad Nacional de Córdoba, Córdoba, Argentina,</mods:affiliation>
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<mods:namePart id="7EEC8D4DC06BF586D09EC99CAB8FB385">Reyes, Ximena</mods:namePart>
<mods:affiliation id="1993417FE609F5461FA54EA8360917A0">Centro de Investigaciones Fitoecogenéticas de Pairumani, Cochabamba, Bolivia</mods:affiliation>
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<mods:title id="0F88B21D61C7BA131F859A7786931928">PLoS ONE</mods:title>
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<mods:date id="61CCD12129FD7CB0BBC85FBE6FDB1E51">2019</mods:date>
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<mods:number id="22E4BFB7EFC9016EE0660C36C2192F84">2019-01-16</mods:number>
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<mods:identifier id="E4175ADD524BA8CBF5009A6D9F26AB8F" type="DOI">10.1371/journal.pone.0209792</mods:identifier>
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<treatment id="03F887D6FFBBFFFA307EF9486265F950" ID-DOI="http://doi.org/10.5281/zenodo.13326488" ID-Zenodo-Dep="13326488" LSID="urn:lsid:plazi:treatment:03F887D6FFBBFFFA307EF9486265F950" httpUri="http://treatment.plazi.org/id/03F887D6FFBBFFFA307EF9486265F950" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4">
<heading id="D0A681ACFFBBFFFF307EF9486402F8E9" fontSize="10" level="2" pageId="3" pageNumber="4" reason="2">
<subSubSection id="C34B654BFFBBFFFF307EF948654BF936" box="[533,1040,1733,1758]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BEE36C0FFBBFFFF307EF948654BF936" blockId="3.[533,1524,1685,1897]" box="[533,1040,1733,1758]" pageId="3" pageNumber="4">
<emphasis id="B925EAD2FFBBFFFF307EF948654BF936" bold="true" box="[533,1040,1733,1758]" pageId="3" pageNumber="4">
<taxonomicName id="4C514D43FFBBFFFF307EF94862EAF936" ID-CoL="7TBQJ" authority="Hunz. ex Barboza" authorityName="Barboza" authorityYear="2019" box="[533,945,1733,1758]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="benoistii" status="sp. nov.">
<emphasis id="B925EAD2FFBBFFFF307EF94863BFF935" bold="true" box="[533,740,1733,1758]" italics="true" pageId="3" pageNumber="4">Capsicum benoistii</emphasis>
Hunz. ex Barboza
</taxonomicName>
,
<taxonomicNameLabel id="A21657A9FFBBFFFF31D0F948654BF936" box="[955,1040,1734,1758]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="C34B654BFFBBFFFF367DF9486331F8CB" pageId="3" pageNumber="4" type="description">
<paragraph id="8BEE36C0FFBBFFFF367DF9486402F8E9" blockId="3.[533,1524,1685,1897]" pageId="3" pageNumber="4">
[
<materialsCitation id="3B393C9DFFBBFFFF3677F948640EF8E9" collectingDate="1931-04-03" collectorName="M. R. Benoist" country="Ecuador" location="Banos" municipality="Type" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Tungurahua" typeStatus="holotype">
urn:lsid:ipni.org:names: 77192556–1].
<collectingMunicipality id="6B8AACBAFFBBFFFF37DCF94864B4F936" box="[1463,1519,1734,1758]" pageId="3" pageNumber="4">Type</collectingMunicipality>
:
<collectingCountry id="F3467650FFBBFFFF307EF967632AF8E9" box="[533,625,1769,1793]" name="Ecuador" pageId="3" pageNumber="4">Ecuador</collectingCountry>
.
<collectingRegion id="4995F822FFBBFFFF3010F9676259F8E9" box="[635,770,1769,1793]" country="Ecuador" name="Tungurahua" pageId="3" pageNumber="4">Tungurahua</collectingRegion>
:
<location id="8E8E601BFFBBFFFF3160F9676215F8E9" LSID="urn:lsid:plazi:treatment:03F887D6FFBBFFFA307EF9486265F950:8E8E601BFFBBFFFF3160F9676215F8E9" box="[779,846,1769,1793]" country="Ecuador" municipality="Type" name="Banos" pageId="3" pageNumber="4" stateProvince="Tungurahua">Baños</location>
,
<date id="FFEF1000FFBBFFFF3133F9676296F8E9" box="[856,973,1769,1793]" pageId="3" pageNumber="4" value="1931-04-03">
<collectingDate id="EFABE9E8FFBBFFFF3133F9676296F8E9" box="[856,973,1769,1793]" pageId="3" pageNumber="4" value="1931-04-03">3 Apr 1931</collectingDate>
</date>
(fl),
<collectorName id="26A45316FFBBFFFF3194F96765D0F8E8" box="[1023,1163,1769,1793]" pageId="3" pageNumber="4">
<emphasis id="B925EAD2FFBBFFFF3194F967654DF8E8" box="[1023,1046,1769,1792]" italics="true" pageId="3" pageNumber="4">M</emphasis>
.
<emphasis id="B925EAD2FFBBFFFF3649F9676569F8E8" box="[1058,1074,1769,1792]" italics="true" pageId="3" pageNumber="4">R</emphasis>
.
<emphasis id="B925EAD2FFBBFFFF3655F9676598F8E9" box="[1086,1219,1769,1793]" italics="true" pageId="3" pageNumber="4">Benoist 4204</emphasis>
</collectorName>
(
<typeStatus id="54EA8862FFBBFFFF36BBF967646AF8E9" box="[1232,1329,1769,1793]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
, P)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36C0FFBBFFFF305EF8856331F8CB" blockId="3.[533,1524,1685,1897]" box="[565,618,1803,1827]" pageId="3" pageNumber="4">
<figureCitation id="136A2A45FFBBFFFF305EF8856331F8CB" box="[565,618,1803,1827]" captionStart="Fig 1" captionStartId="5.[454,482,203,222]" captionTargetId="figure-1@4.[454,1536,208,1872]" captionTargetPageId="4" captionText="Fig 1. Capsicum benoistii Hunz. ex Barboza. (A) Flowering branch. (B) Flower. (C) Calyx. (D) Section of the calyx showing venation in tube and appendages. (E) Trichome of the calyx. (F) Opened corolla. (G, H) Gynoecium, with long and short style, respectively. Drawn by N. de Flury." figureDoi="http://doi.org/10.5281/zenodo.13326484" httpUri="https://zenodo.org/record/13326484/files/figure.png" pageId="3" pageNumber="4">Fig 1</figureCitation>
</paragraph>
</subSubSection>
</heading>
<subSubSection id="C34B654BFFBBFFF9305EF8A06205FED2" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BEE36C0FFBBFFFF305EF8A06484F881" blockId="3.[533,1524,1685,1897]" pageId="3" pageNumber="4">
<emphasis id="B925EAD2FFBBFFFF305EF8A063F8F8AE" bold="true" box="[565,675,1838,1862]" pageId="3" pageNumber="4">Diagnosis</emphasis>
. Similar to
<taxonomicName id="4C514D43FFBBFFFF3176F8A0658CF8AE" authority="(Dammer) Hunz." authorityName="Hunz." baseAuthorityName="Dammer" box="[797,1239,1838,1862]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="geminifolium">
<emphasis id="B925EAD2FFBBFFFF3176F8A0654DF8AE" box="[797,1046,1838,1862]" italics="true" pageId="3" pageNumber="4">Capsicum geminifolium</emphasis>
(Dammer) Hunz.
</taxonomicName>
but differing in the length of the flowering pedicels, the shape of the corolla, and the presence of heterostylous flowers.
</paragraph>
<caption id="DF2E6648FFBDFFF933ADFF4563AAFEF9" ID-DOI="http://doi.org/10.5281/zenodo.13326484" ID-Zenodo-Dep="13326484" httpUri="https://zenodo.org/record/13326484/files/figure.png" pageId="5" pageNumber="6" startId="5.[454,482,203,222]" targetBox="[454,1532,208,1872]" targetPageId="4" targetType="figure">
<paragraph id="8BEE36C0FFBDFFF933ADFF4563AAFEF9" blockId="5.[454,1518,202,273]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF933ADFF456212FF36" bold="true" box="[454,841,202,222]" pageId="5" pageNumber="6">
Fig 1.
<taxonomicName id="4C514D43FFBDFFF93395FF456212FF36" authority="Hunz. ex Barboza." authorityName="Barboza." box="[510,841,202,222]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="benoistii">
<emphasis id="B925EAD2FFBDFFF93395FF4563F8FF35" bold="true" box="[510,675,202,222]" italics="true" pageId="5" pageNumber="6">Capsicum benoistii</emphasis>
Hunz. ex Barboza.
</taxonomicName>
</emphasis>
(A) Flowering branch. (B) Flower. (C) Calyx. (D) Section of the calyx showing venation in tube and appendages. (E) Trichome of the calyx. (F) Opened corolla. (G, H) Gynoecium, with long and short style, respectively. Drawn by N. de Flury.
</paragraph>
</caption>
<paragraph id="8BEE36C0FFBDFFF933ADFEA96205FED2" blockId="5.[454,862,295,314]" box="[454,862,295,314]" pageId="5" pageNumber="6">https://doi.org/10.1371/journal.pone.0209792.g001</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFBDFFF9305EFEED64ACFB70" pageId="5" pageNumber="6" type="description">
<paragraph id="8BEE36C0FFBDFFF9305EFEED64ACFB70" blockId="5.[533,1536,355,1869]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF9305EFEED63E3FE93" bold="true" box="[565,696,355,379]" pageId="5" pageNumber="6">Description</emphasis>
. Shrubs few branched. Young stems light brown, glabrous or pubescent, striate; bark of older stems dark brown, glabrous, striate; lenticels absent. Sympodial units difoliate, geminate, leaf pair markedly anisophyllous in size and shape. Leaves simple, membranaceous, discolorous, adaxial surface dark green, abaxial surface light green, glabrous or with simple antrorse trichomes
<quantity id="4CA99B25FFBDFFF9310EFE6362BFFDED" box="[869,996,493,517]" metricMagnitude="-4" metricUnit="m" metricValue="7" metricValueMax="11.0" metricValueMin="3.0" pageId="5" pageNumber="6" unit="mm" value="0.7" valueMax="1.1" valueMin="0.3">0.3–1.1 mm</quantity>
long adaxially and abaxially, trichomes more abundant on main veins; the larger leaves with blades
<quantity id="4CA99B25FFBDFFF9363BFD9E65E6FDC0" box="[1104,1213,528,552]" metricMagnitude="-1" metricUnit="m" metricValue="1.025" metricValueMax="1.2" metricValueMin="0.85" pageId="5" pageNumber="6" unit="cm" value="10.25" valueMax="12.0" valueMin="8.5">8.5–12 cm</quantity>
long,
<quantity id="4CA99B25FFBDFFF93695FD9E6404FDC0" box="[1278,1375,528,552]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="6.0" metricValueMin="2.8" pageId="5" pageNumber="6" unit="cm" value="4.4" valueMax="6.0" valueMin="2.8">2.8–6 cm</quantity>
wide, ovate or elliptic, major veins 4–5 (6) on each side of midvein, base asymmetric and attenuate, margin entire, apex long-acuminate; petioles 0.5–1.0 cm long, glabrous or glabrescent with trichomes like those of the leaves; the minor leaves
<quantity id="4CA99B25FFBDFFF931AAFDF66579FD78" box="[961,1058,632,656]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="6.0" metricValueMin="2.4" pageId="5" pageNumber="6" unit="cm" value="4.2" valueMax="6.0" valueMin="2.4">2.4–6 cm</quantity>
long,
<quantity id="4CA99B25FFBDFFF93608FDF66598FD78" box="[1123,1219,632,656]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="4.0" metricValueMin="1.7" pageId="5" pageNumber="6" unit="cm" value="2.85" valueMax="4.0" valueMin="1.7">1.7–4 cm</quantity>
wide, ovate or elliptic, major veins 3–4 on each side of midvein, base rounded, asymmetric, margin entire, apex acute or rounded; petioles
<quantity id="4CA99B25FFBDFFF930BFFD33626EFD3D" box="[724,821,701,725]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="10.0" metricValueMin="1.0" pageId="5" pageNumber="6" unit="cm" value="0.55" valueMax="1.0" valueMin="0.1">0.1–1 cm</quantity>
long, glabrescent or pubescent. Flowers in fascicles of 3–6; flowering pedicels filiform, striate, pendent, not geniculate at anthesis,
<quantity id="4CA99B25FFBDFFF936D5FD6E6445FD10" box="[1214,1310,736,760]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.0" metricValueMin="1.3" pageId="5" pageNumber="6" unit="cm" value="1.65" valueMax="2.0" valueMin="1.3">1.3–2 cm</quantity>
long, moderately to densely pubescent, the trichomes simple, non-glandular, multicellular, antrorse,
<quantity id="4CA99B25FFBDFFF93702FC8D631AFCD5" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="7.5" metricValueMin="3.0" pageId="5" pageNumber="6" unit="mm" value="0.525" valueMax="0.75" valueMin="0.3">0.30–0.75 mm</quantity>
long. Flower buds ovoid. Calyx
<quantity id="4CA99B25FFBDFFF931FEFCAB655AFCD5" box="[917,1025,805,829]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="5" pageNumber="6" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2–2.5 mm</quantity>
long, ca.
<quantity id="4CA99B25FFBDFFF9360FFCAB65F8FCD5" box="[1124,1187,805,829]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="mm" value="5.0">5 mm</quantity>
wide, cup-shaped, thick, the margin truncate, pubescent with the same trichomes as pedicels, with 5 appendages
<quantity id="4CA99B25FFBDFFF937E5FCC6631AFC6B" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="5" pageNumber="6" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.5–3.5 mm</quantity>
long, ca.
<quantity id="4CA99B25FFBDFFF930CEFCE563ADFC6B" box="[677,758,875,899]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="mm" value="0.5">0.5 mm</quantity>
wide, thick, erect, subulate, inserted close to the margin, pubescent with the same trichomes as calyx tube. Corolla ca.
<quantity id="4CA99B25FFBDFFF93198FC03653EFC4D" box="[1011,1125,909,933]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="5" pageNumber="6" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
long, deeply stellate, thick, without interpetalar tissue; tube ca.
<quantity id="4CA99B25FFBDFFF9315DFC3E622FFC20" box="[822,884,944,968]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="mm" value="3.0">3 mm</quantity>
long, glabrous inside and outside; lobes ca.
<quantity id="4CA99B25FFBDFFF93728FC3E64DAFC20" box="[1347,1409,944,968]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="6" unit="mm" value="9.0">9 mm</quantity>
long, ca.
<quantity id="4CA99B25FFBDFFF9378FFC3E631AFC03" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="6" unit="mm" value="2.0">2 mm</quantity>
wide, narrowly triangular, erect, glabrous adaxially and abaxially, the tips and margins pubescent. Stamens 5, equal, filaments equal,
<quantity id="4CA99B25FFBDFFF9319CFC7B6539FBE5" box="[1015,1122,1013,1037]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" pageId="5" pageNumber="6" unit="mm" value="3.1" valueMax="3.2" valueMin="3.0">3–3.2 mm</quantity>
long, glabrous, inserted on the corolla
<quantity id="4CA99B25FFBDFFF9307EFB96633CFBD8" box="[533,615,1048,1072]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="6" unit="mm" value="1.5">1.5 mm</quantity>
from the base, with inconspicuous auricles at point of insertion; anthers ca.
<quantity id="4CA99B25FFBDFFF937E0FB966491FBD8" box="[1419,1482,1048,1072]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="mm" value="3.0">3 mm</quantity>
long, not connivent, elliptic. Ovary
<quantity id="4CA99B25FFBDFFF931E7FBB56550FBBB" box="[908,1035,1083,1107]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.7" metricValueMin="1.3" pageId="5" pageNumber="6" unit="mm" value="1.5" valueMax="1.7" valueMin="1.3">1.3–1.7 mm</quantity>
long,
<quantity id="4CA99B25FFBDFFF93627FBB56591FBBB" box="[1100,1226,1083,1107]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="5" pageNumber="6" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.2–1.5 mm</quantity>
diam, subglobose, glabrous; nectary ca.
<quantity id="4CA99B25FFBDFFF930E6FBD36385FB9D" box="[653,734,1117,1141]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="mm" value="0.3">0.3 mm</quantity>
long, inconspicuous; long style ca.
<quantity id="4CA99B25FFBDFFF93639FBD365F8FB9D" box="[1106,1187,1117,1141]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="6" unit="mm" value="6.5">6.5 mm</quantity>
long, short style ca.
<quantity id="4CA99B25FFBDFFF93712FBD36491FB9D" box="[1401,1482,1117,1141]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="6" unit="mm" value="3.6">3.6 mm</quantity>
long, widening distally, glabrous; stigma
<quantity id="4CA99B25FFBDFFF931AEFB0E654DFB70" box="[965,1046,1152,1176]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="mm" value="0.3">0.3 mm</quantity>
long,
<quantity id="4CA99B25FFBDFFF9363DFB0E65F3FB70" box="[1110,1192,1152,1176]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="mm" value="0.5">0.5 mm</quantity>
wide, globose. Berry unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34B654BFFBDFFFA305EFB2D6265F950" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8BEE36C0FFBDFFF9305EFB2D64AFFB36" blockId="5.[533,1536,355,1869]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF9305EFB2D6211FB53" bold="true" box="[565,842,1187,1211]" pageId="5" pageNumber="6">Distribution and ecology</emphasis>
. Endemic to a restricted area in central-southern
<collectingCountry id="F3467650FFBDFFF9373CFB2D64EAFB53" box="[1367,1457,1187,1211]" name="Ecuador" pageId="5" pageNumber="6">Ecuador</collectingCountry>
(
<collectingRegion id="4995F822FFBDFFF937D4FB2D6334FB36" country="Ecuador" name="Tungurahua" pageId="5" pageNumber="6">Tungurahua</collectingRegion>
,
<collectingRegion id="4995F822FFBDFFF93012FB4863F1FB36" box="[633,682,1222,1246]" country="Ecuador" name="Loja" pageId="5" pageNumber="6">Loja</collectingRegion>
,
<figureCitation id="136A2A45FFBDFFF930D8FB4863B3FB36" box="[691,744,1222,1246]" captionStart="Fig 2" captionStartId="6.[533,561,1150,1169]" captionTargetBox="[533,1320,208,1128]" captionTargetId="figure-278@6.[533,1320,208,1129]" captionTargetPageId="6" captionText="Fig 2. Distribution of Capsicum species. https://doi.org/10.1371/journal.pone.0209792.g002" figureDoi="http://doi.org/10.5281/zenodo.12630923" httpUri="https://zenodo.org/record/12630923/files/figure.png" pageId="5" pageNumber="6">Fig 2</figureCitation>
) growing in thickets in montane forests, between
<quantity id="4CA99B25FFBDFFF93692FB4864D3FB36" box="[1273,1416,1222,1246]" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.6" metricValueMin="1.5" pageId="5" pageNumber="6" unit="m" value="2050.0" valueMax="2600.0" valueMin="1500.0">1500–2600 m</quantity>
elevation.
</paragraph>
<paragraph id="8BEE36C0FFBDFFF9305EFB66645CFAE8" blockId="5.[533,1536,355,1869]" box="[565,1287,1256,1280]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF9305EFB6663F1FAE8" bold="true" box="[565,682,1256,1280]" pageId="5" pageNumber="6">Phenology</emphasis>
. Flowering from March to May. Fruiting time unknown.
</paragraph>
<paragraph id="8BEE36C0FFBDFFF9305EFA8562EFFA80" blockId="5.[533,1536,355,1869]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF9305EFA8563F7FACB" bold="true" box="[565,684,1291,1315]" pageId="5" pageNumber="6">Etymology</emphasis>
—The new species is named in honor to Raymond Benoist (1881–1970), a French botanist, who collected in French
<collectingCountry id="F3467650FFBDFFF931A7FAA0657AFAAE" box="[972,1057,1326,1350]" name="Guyana" pageId="5" pageNumber="6">Guyana</collectingCountry>
,
<collectingCountry id="F3467650FFBDFFF93640FAA065D7FAAE" box="[1067,1164,1326,1350]" name="Morocco" pageId="5" pageNumber="6">Morocco</collectingCountry>
and
<collectingCountry id="F3467650FFBDFFF936ABFAA06447FAAE" box="[1216,1308,1326,1350]" name="Ecuador" pageId="5" pageNumber="6">Ecuador</collectingCountry>
; the
<typeStatus id="54EA8862FFBDFFF93726FAA064F1FAAE" box="[1357,1450,1326,1350]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
is a nice specimen collected by her in 1931.
</paragraph>
<paragraph id="8BEE36C0FFBDFFF9305EFAFD64ECFA38" blockId="5.[533,1536,355,1869]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF9305EFAFD62F3FA63" bold="true" box="[565,936,1395,1419]" pageId="5" pageNumber="6">Species Conservation Assessment</emphasis>
. Following the IUCN Red List Criteria (IUCN 2017), this species is proposed as Endangered (
<collectionCode id="ED40AE05FFBDFFF931D7FA1862BFFA46" box="[956,996,1430,1454]" pageId="5" pageNumber="6">EN</collectionCode>
). The extent of occurrence is calculated to be
<quantity id="4CA99B25FFBDFFF937AFFA186363FA38" metricMagnitude="6" metricUnit="m" metricValue="2.05" pageId="5" pageNumber="6" unit="km" value="2050.0">2050 km</quantity>
2 (Criterion B1 &lt;
<quantity id="4CA99B25FFBDFFF93092FA36620CFA38" box="[761,855,1464,1488]" metricMagnitude="6" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="km" value="5000.0">5000 km</quantity>
2, Endangered), the area of occupancy,
<quantity id="4CA99B25FFBDFFF9369DFA366463FA38" box="[1270,1336,1464,1488]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="6" unit="km" value="12.0">12 km</quantity>
2 (Criterion
</paragraph>
<paragraph id="8BEE36C0FFBDFFF9307EFA55648BFA1B" blockId="5.[533,1536,355,1869]" box="[533,1488,1499,1523]" pageId="5" pageNumber="6">
B2 &lt;
<quantity id="4CA99B25FFBDFFF93039FA5563F8FA1B" box="[594,675,1499,1523]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="km" value="500.0">500 km</quantity>
2, Endangered) and the species is known from only three localities (Criterion
</paragraph>
<paragraph id="8BEE36C0FFBDFFF9307EFA7362D3F9B3" blockId="5.[533,1536,355,1869]" pageId="5" pageNumber="6">
B1a
<emphasis id="B925EAD2FFBDFFF93028FA736303F9FD" bold="true" box="[579,600,1533,1557]" pageId="5" pageNumber="6"><EFBFBD></emphasis>
5, Endangered). It is possible that its geographic range has declined (EOO and AOO, Criterion B2b i &amp; ii) because the species has not been collected since 1978 despite recent intensive searches in the same locations.
</paragraph>
<paragraph id="8BEE36C0FFBDFFF9305EF9EB65F7F92B" blockId="5.[533,1536,355,1869]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF9305EF9EB62C3F995" bold="true" box="[565,920,1637,1661]" pageId="5" pageNumber="6">Additional specimens examined</emphasis>
.
<materialsCitation id="3B393C9DFFBDFFF931CFF9EB644AF948" collectingDate="1978-05-21" collectorName="F. Vivar &amp; Estudiantes" country="Ecuador" elevation="2580" location="Pueblo Nuevo" pageId="5" pageNumber="6" specimenCount="1" stateProvince="Loja">
<emphasis id="B925EAD2FFBDFFF931CFF9EB6539F995" bold="true" box="[932,1122,1637,1661]" pageId="5" pageNumber="6">
<collectingCountry id="F3467650FFBDFFF931CFF9EB6573F995" box="[932,1064,1637,1661]" name="Ecuador" pageId="5" pageNumber="6">ECUADOR</collectingCountry>
.
<collectingRegion id="4995F822FFBDFFF93659F9EB6539F995" box="[1074,1122,1637,1661]" country="Ecuador" name="Loja" pageId="5" pageNumber="6">Loja</collectingRegion>
</emphasis>
:
<location id="8E8E601BFFBDFFF93604F9EB645CF995" LSID="urn:lsid:plazi:treatment:03F887D6FFBBFFFA307EF9486265F950:8E8E601BFFBDFFF93604F9EB645CF995" box="[1135,1287,1637,1661]" country="Ecuador" name="Pueblo Nuevo" pageId="5" pageNumber="6" stateProvince="Loja">Pueblo Nuevo</location>
, 04˚05’51’’S, 79˚ 11’55’’W,
<quantity id="4CA99B25FFBDFFF93014F9066394F948" box="[639,719,1672,1696]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="5" pageNumber="6" unit="m" value="2580.0">
<elevation id="007CD1F3FFBDFFF93014F9066394F948" box="[639,719,1672,1696]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="5" pageNumber="6" unit="m" value="2580.0">2580 m</elevation>
</quantity>
,
<date id="FFEF1000FFBDFFF930B2F906623AF948" box="[729,865,1672,1696]" pageId="5" pageNumber="6" value="1978-05-21">
<collectingDate id="EFABE9E8FFBDFFF930B2F906623AF948" box="[729,865,1672,1696]" pageId="5" pageNumber="6" value="1978-05-21">21 May 1978</collectingDate>
</date>
(fl),
<collectorName id="26A45316FFBDFFF931F8F90762B3F948" box="[915,1000,1672,1696]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF931F8F90762FAF948" box="[915,929,1673,1696]" italics="true" pageId="5" pageNumber="6">F</emphasis>
.
<emphasis id="B925EAD2FFBDFFF931C7F90765E1F948" box="[940,1210,1672,1696]" italics="true" pageId="5" pageNumber="6">
Vivar &amp;
<collectorName id="26A45316FFBDFFF93663F90665DAF948" box="[1032,1153,1672,1696]" pageId="5" pageNumber="6">Estudiantes</collectorName>
1066
</emphasis>
</collectorName>
(LOJA)
</materialsCitation>
;
<materialsCitation id="3B393C9DFFBDFFF93770F90665F2F92B" collectingDate="1956-03-30" collectorName="E. Asplund" country="Ecuador" elevation="1500" location="Rio Verde Grande" pageId="5" pageNumber="6" specimenCount="1" stateProvince="Tungurahua">
<emphasis id="B925EAD2FFBDFFF93770F90664FDF948" bold="true" box="[1307,1446,1672,1696]" pageId="5" pageNumber="6">
<collectingRegion id="4995F822FFBDFFF93770F90664FDF948" box="[1307,1446,1672,1696]" country="Ecuador" name="Tungurahua" pageId="5" pageNumber="6">Tungurahua</collectingRegion>
</emphasis>
:
<location id="8E8E601BFFBDFFF937DAF90663F4F92B" LSID="urn:lsid:plazi:treatment:03F887D6FFBBFFFA307EF9486265F950:8E8E601BFFBDFFF937DAF90663F4F92B" country="Ecuador" name="Rio Verde Grande" pageId="5" pageNumber="6" stateProvince="Tungurahua">Río Verde Grande</location>
,
<quantity id="4CA99B25FFBDFFF930D2F9256252F92B" box="[697,777,1707,1731]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="6" unit="m" value="1500.0">
<elevation id="007CD1F3FFBDFFF930D2F9256252F92B" box="[697,777,1707,1731]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="6" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="FFEF1000FFBDFFF93178F92562C3F92B" box="[787,920,1707,1731]" pageId="5" pageNumber="6" value="1956-03-30">
<collectingDate id="EFABE9E8FFBDFFF93178F92562C3F92B" box="[787,920,1707,1731]" pageId="5" pageNumber="6" value="1956-03-30">30 Mar 1956</collectingDate>
</date>
(fl),
<collectorName id="26A45316FFBDFFF931A1F9256566F92A" box="[970,1085,1706,1731]" pageId="5" pageNumber="6">
<emphasis id="B925EAD2FFBDFFF931A1F9256282F92A" box="[970,985,1707,1730]" italics="true" pageId="5" pageNumber="6">E</emphasis>
.
<emphasis id="B925EAD2FFBDFFF9318EF92465DAF92B" box="[997,1153,1706,1731]" italics="true" pageId="5" pageNumber="6">Asplund 20070</emphasis>
</collectorName>
(S)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36C0FFBDFFFA305EF940640FFA68" blockId="5.[533,1536,355,1869]" lastBlockId="6.[533,1536,1245,1893]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<taxonomicName id="4C514D43FFBDFFF9305EF94063A7F90D" box="[565,764,1741,1765]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="benoistii">
<emphasis id="B925EAD2FFBDFFF9305EF94063A7F90D" box="[565,764,1741,1765]" italics="true" pageId="5" pageNumber="6">Capsicum benoistii</emphasis>
</taxonomicName>
was identified as a new species by the late
<taxonomicName id="4C514D43FFBDFFF936D2F9436476F90D" box="[1209,1325,1741,1765]" class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" order="Solanales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="family">Solanaceae</taxonomicName>
specialist Armando T. Hunziker (
<collectionCode id="ED40AE05FFBDFFF930CFF97E63AAF8E0" box="[676,753,1776,1800]" country="Argentina" lsid="urn:lsid:biocol.org:col:15636" name="Universidad Nacional de Córdoba" pageId="5" pageNumber="6" type="Herbarium">CORD</collectionCode>
) who annotated the epithet name
<taxonomicName id="4C514D43FFBDFFF93630F97E65EFF8E0" box="[1115,1204,1776,1800]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="benoistii">
<emphasis id="B925EAD2FFBDFFF93630F97E65EFF8E0" box="[1115,1204,1776,1800]" italics="true" pageId="5" pageNumber="6">benoistii</emphasis>
</taxonomicName>
on the specimen housed at P (Benoist 4204), but this name was never published. It is a poorly known species collected only three times in
<collectingCountry id="F3467650FFBDFFF930C6F8BB6252F8A5" box="[685,777,1845,1869]" name="Ecuador" pageId="5" pageNumber="6">Ecuador</collectingCountry>
; none of these collections have fruits. Extensive recent field explorations in
<collectingRegion id="4995F822FFBEFFFA30D0FB53621BFB1D" box="[699,832,1245,1269]" country="Ecuador" name="Tungurahua" pageId="6" pageNumber="7">Tungurahua</collectingRegion>
were unsuccessful in finding this species. It is distinctive in its deeply lobed stellate corolla (lobes three times longer than the tube,
<figureCitation id="136A2A45FFBEFFFA368BFA8E647FFAF0" box="[1248,1316,1280,1304]" captionStart="Fig 1" captionStartId="5.[454,482,203,222]" captionTargetId="figure-1@4.[454,1536,208,1872]" captionTargetPageId="4" captionText="Fig 1. Capsicum benoistii Hunz. ex Barboza. (A) Flowering branch. (B) Flower. (C) Calyx. (D) Section of the calyx showing venation in tube and appendages. (E) Trichome of the calyx. (F) Opened corolla. (G, H) Gynoecium, with long and short style, respectively. Drawn by N. de Flury." figureDoi="http://doi.org/10.5281/zenodo.13326484" httpUri="https://zenodo.org/record/13326484/files/figure.png" pageId="6" pageNumber="7">Fig 1B</figureCitation>
) and in the presence of heterostylous flowers (
<figureCitation id="136A2A45FFBEFFFA314BFAAC62E0FAD2" box="[800,955,1314,1338]" captionStart="Fig 1" captionStartId="5.[454,482,203,222]" captionTargetId="figure-1@4.[454,1536,208,1872]" captionTargetPageId="4" captionText="Fig 1. Capsicum benoistii Hunz. ex Barboza. (A) Flowering branch. (B) Flower. (C) Calyx. (D) Section of the calyx showing venation in tube and appendages. (E) Trichome of the calyx. (F) Opened corolla. (G, H) Gynoecium, with long and short style, respectively. Drawn by N. de Flury." figureDoi="http://doi.org/10.5281/zenodo.13326484" httpUri="https://zenodo.org/record/13326484/files/figure.png" pageId="6" pageNumber="7">Fig 1G and 1H</figureCitation>
). These features plus the short flowering pedicels (
<quantity id="4CA99B25FFBEFFFA3076FACB6324FAB5" box="[541,639,1349,1373]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="2.0" metricValueMin="1.3" pageId="6" pageNumber="7" unit="cm" value="1.65" valueMax="2.0" valueMin="1.3">1.3–2 cm</quantity>
long) distinguish
<taxonomicName id="4C514D43FFBEFFFA3154FACB62EEFAB5" box="[831,949,1349,1373]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="benoistii">
<emphasis id="B925EAD2FFBEFFFA3154FACB620BFAB4" box="[831,848,1349,1372]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="B925EAD2FFBEFFFA3137FACB62EEFAB5" box="[860,949,1349,1373]" italics="true" pageId="6" pageNumber="7">benoistii</emphasis>
</taxonomicName>
from
<taxonomicName id="4C514D43FFBEFFFA319EFACB65C7FAB5" box="[1013,1180,1349,1373]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="geminifolium">
<emphasis id="B925EAD2FFBEFFFA319EFACB655DFAB4" box="[1013,1030,1349,1372]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="B925EAD2FFBEFFFA367AFACB65C7FAB5" box="[1041,1180,1349,1373]" italics="true" pageId="6" pageNumber="7">geminifolium</emphasis>
</taxonomicName>
, which has funnel-shaped corollas lobed about halfway, homostylous flowers, and longer pedicels (
<quantity id="4CA99B25FFBEFFFA36B0FAE66454FA68" box="[1243,1295,1384,1408]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="7" unit="cm" value="5.0">5 cm</quantity>
long).
</paragraph>
<caption id="DF2E6648FFBEFFFA307EFBF062F6FB53" ID-DOI="http://doi.org/10.5281/zenodo.12630923" ID-Zenodo-Dep="12630923" httpUri="https://zenodo.org/record/12630923/files/figure.png" pageId="6" pageNumber="7" startId="6.[533,561,1150,1169]" targetBox="[533,1320,208,1128]" targetPageId="6" targetType="figure">
<paragraph id="8BEE36C0FFBEFFFA307EFBF0622EFB79" blockId="6.[533,941,1150,1211]" box="[533,885,1150,1169]" pageId="6" pageNumber="7">
<emphasis id="B925EAD2FFBEFFFA307EFBF0622EFB79" bold="true" box="[533,885,1150,1169]" pageId="6" pageNumber="7">
Fig 2. Distribution of
<taxonomicName id="4C514D43FFBEFFFA30BCFBF0622EFB79" box="[727,885,1150,1169]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B925EAD2FFBEFFFA30BCFBF06276FB79" bold="true" box="[727,813,1150,1169]" italics="true" pageId="6" pageNumber="7">Capsicum</emphasis>
species.
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BEE36C0FFBEFFFA307EFB2662F6FB53" blockId="6.[533,941,1150,1211]" box="[533,941,1192,1211]" pageId="6" pageNumber="7">https://doi.org/10.1371/journal.pone.0209792.g002</paragraph>
</caption>
<paragraph id="8BEE36C0FFBEFFFA305EFA046584F9C5" blockId="6.[533,1536,1245,1893]" pageId="6" pageNumber="7">
The presence of heterostylous flowers as in
<taxonomicName id="4C514D43FFBEFFFA366BFA04652EFA4A" box="[1024,1141,1418,1442]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="benoistii">
<emphasis id="B925EAD2FFBEFFFA366BFA04654AFA49" box="[1024,1041,1418,1441]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="B925EAD2FFBEFFFA3677FA04652EFA4A" box="[1052,1141,1418,1442]" italics="true" pageId="6" pageNumber="7">benoistii</emphasis>
</taxonomicName>
is unusual among
<taxonomicName id="4C514D43FFBEFFFA3757FA0464A1FA4A" box="[1340,1530,1418,1442]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B925EAD2FFBEFFFA3757FA0464FFFA49" box="[1340,1444,1418,1441]" italics="true" pageId="6" pageNumber="7">Capsicum</emphasis>
species.
</taxonomicName>
It has been reported in
<taxonomicName id="4C514D43FFBEFFFA3160FA2362D5FA2D" box="[779,910,1453,1477]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="baccatum">
<emphasis id="B925EAD2FFBEFFFA3160FA236247FA2C" box="[779,796,1453,1476]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="B925EAD2FFBEFFFA3143FA2362D5FA2D" box="[808,910,1453,1477]" italics="true" pageId="6" pageNumber="7">baccatum</emphasis>
</taxonomicName>
L. varieties:
<taxonomicName id="4C514D43FFBEFFFA3679FA2365FEFA2D" box="[1042,1189,1453,1477]" pageId="6" pageNumber="7" rank="variety" variety="baccatum">
var.
<emphasis id="B925EAD2FFBEFFFA3654FA2365FEFA2D" box="[1087,1189,1453,1477]" italics="true" pageId="6" pageNumber="7">baccatum</emphasis>
</taxonomicName>
[
<bibRefCitation id="EFC04B31FFBEFFFA36DFFA236596FA2D" author="Barboza GE" box="[1204,1229,1453,1477]" pageId="6" pageNumber="7" pagination="21 - 25" refId="ref13912" refString="29. Barboza GE. Capsicum. In Barboza GE, coordinator. Solanaceae. Flora Argentina. Buenos Aires: IBODA-IMBIV, CONICET; 2013; 13: 21 - 25." type="journal article" year="2013">29</bibRefCitation>
] and
<taxonomicName id="4C514D43FFBEFFFA3760FA23624DFA00" authority="(Vell.) Hunz. &amp; Barboza" authorityName="Hunz. &amp; Barboza" baseAuthorityName="Vell." pageId="6" pageNumber="7" rank="variety" variety="umbilicatum">
var.
<emphasis id="B925EAD2FFBEFFFA3753FA2364E4FA2D" box="[1336,1471,1453,1477]" italics="true" pageId="6" pageNumber="7">umbilicatum</emphasis>
(Vell.) Hunz. &amp; Barboza
</taxonomicName>
[
<bibRefCitation id="EFC04B31FFBEFFFA314EFA5E6265FA00" author="Hunziker AT &amp; Barboza GE" box="[805,830,1488,1512]" pageId="6" pageNumber="7" pagination="23 - 31" refId="ref13945" refString="30. Hunziker AT, Barboza GE. Estudios sobre Solanaceae XLV. Sobre la presencia de Exodeconus en Argentina y una novedad en Capsicum baccatum. Kurtziana. 1998; 26: 23 - 31." type="journal article" year="1998">30</bibRefCitation>
,
<bibRefCitation id="EFC04B31FFBEFFFA3121FA5E6238FA00" author="Hunziker AT" box="[842,867,1488,1512]" pageId="6" pageNumber="7" refId="ref13982" refString="31. Hunziker AT. Genera Solanacearum. The genera of Solanaceae illustrated, arranged according to a new system. Ruggell: A. R. G. Gantner Verlag K. - G; 2001." type="book" year="2001">31</bibRefCitation>
], and observed in other species (
<taxonomicName id="4C514D43FFBEFFFA36D4FA5E63D6F9E2" authority="Eshbaugh, P. G. Sm. &amp; Nickrent" authorityName="Eshbaugh, P. G. Sm. &amp; Nickrent" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="tovarii">
<emphasis id="B925EAD2FFBEFFFA36D4FA5E658BFA0F" box="[1215,1232,1488,1511]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="B925EAD2FFBEFFFA36B0FA5E647BFA0F" box="[1243,1312,1488,1511]" italics="true" pageId="6" pageNumber="7">tovarii</emphasis>
Eshbaugh, P.G.Sm. &amp; Nickrent
</taxonomicName>
and
<taxonomicName id="4C514D43FFBEFFFA30AAFA7C6572F9E2" authority="Ruiz &amp; Pav., Barboza" authorityName="Ruiz &amp; Pav., Barboza" box="[705,1065,1522,1546]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="species" species="pubescens">
<emphasis id="B925EAD2FFBEFFFA30AAFA7C6389F9E1" box="[705,722,1522,1545]" italics="true" pageId="6" pageNumber="7">C</emphasis>
.
<emphasis id="B925EAD2FFBEFFFA30B5FA7C621EF9E2" box="[734,837,1522,1546]" italics="true" pageId="6" pageNumber="7">pubescens</emphasis>
Ruiz &amp; Pav., Barboza
</taxonomicName>
pers. obs.). This character deserves careful field observations to ascertain if short-styled flowers produce fruits.
</paragraph>
<paragraph id="8BEE36C0FFBEFFFA305EF9B66265F950" blockId="6.[533,1536,1245,1893]" pageId="6" pageNumber="7">
As some data are still unknown (e.g. corolla color, fruit and seed characters, and chromosome number) for this species and freshly collected leaf material is not available for DNA extraction, we cannot suggest in which of the different clades of the current phylogeny of
<taxonomicName id="4C514D43FFBEFFFA37ADF9F36309F95F" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="genus">
<emphasis id="B925EAD2FFBEFFFA37ADF9F36309F95F" italics="true" pageId="6" pageNumber="7">Capsicum</emphasis>
</taxonomicName>
[
<bibRefCitation id="EFC04B31FFBEFFFA300AF92E6335F950" author="Carrizo Garcia C &amp; Sehr EM &amp; Barfuss MHJ &amp; Barboza GE &amp; Samuel R &amp; Moscone EA" box="[609,622,1696,1720]" pageId="6" pageNumber="7" pagination="35 - 51" refId="ref12646" refString="1. Carrizo Garcia C, Sehr EM, Barfuss MHJ, Barboza GE, Samuel R, Moscone EA et al. Phylogenetic relationships, diversification and expansion of chili peppers (Capsicum, Solanaceae). Ann Bot. 2016; 118: 35 - 51. https: // doi. org / 10.1093 / aob / mcw 079 PMID: 27245634" type="journal article" year="2016">1</bibRefCitation>
] it could be placed.
</paragraph>
</subSubSection>
</treatment>
</document>
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